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<document id="84B7272228278681000ABB81C3185F61" ID-DOI="10.11646/zootaxa.5556.1.14" ID-ISSN="1175-5326" ID-Zenodo-Dep="14595072" ID-ZooBank="7FA510AD-15BA-4E08-B789-8C58B296D8C8" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1735930822596" checkinUser="plazi" docAuthor="Miko, Ladislav" docDate="2024" docId="03BD87BCF862FFC4EBFAFC11AD8F7454" docLanguage="en" docName="zootaxa.5556.1.14.pdf" docOrigin="Zootaxa 5556 (1)" docSource="https://doi.org/10.11646/zootaxa.5556.1.14" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sternoppia (Luisternoppia) subiasi Miko, 2024, sp.nov." docType="treatment" docVersion="1" lastPageNumber="204" masterDocId="FF84FFC4F863FFC0EB6DFF99A8137751" masterDocTitle="Oribatid mites (Acarina, Oribatida) from French Guyana III. A new subgenus and its type species, Sternoppia (Luisternoppia) subiasi sp. nov. from family Sternoppiidae Balogh et Mahunka, 1969" masterLastPageNumber="204" masterPageNumber="200" pageNumber="201" updateTime="1736262198873" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="3D0DB02500F3A2CC828ADEC07ADC916A">Oribatid mites (Acarina, Oribatida) from French Guyana III. A new subgenus and its type species, Sternoppia (Luisternoppia) subiasi sp. nov. from family Sternoppiidae Balogh et Mahunka, 1969</mods:title>
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<mods:namePart id="4CF3AD69FD8A33367BBE9F1BCE72538C">Miko, Ladislav</mods:namePart>
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<treatment id="03BD87BCF862FFC4EBFAFC11AD8F7454" LSID="urn:lsid:plazi:treatment:03BD87BCF862FFC4EBFAFC11AD8F7454" httpUri="http://treatment.plazi.org/id/03BD87BCF862FFC4EBFAFC11AD8F7454" lastPageId="4" lastPageNumber="204" pageId="1" pageNumber="201">
<subSubSection id="C30E6521F862FFC1EBFAFC11AAC974CE" box="[151,730,903,928]" pageId="1" pageNumber="201" type="nomenclature">
<paragraph id="8BAB36AAF862FFC1EBFAFC11AAC974CE" blockId="1.[151,730,903,928]" box="[151,730,903,928]" pageId="1" pageNumber="201">
Type species:
<taxonomicName id="4C144D29F862FFC1EA4BFC1EAA9A74CE" authorityName="Miko" authorityYear="2024" box="[294,649,903,927]" class="Arachnida" family="Sternoppiidae" genus="Sternoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="201" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov." subGenus="Luisternoppia">
<emphasis id="B960EAB8F862FFC1EA4BFC1EAA9A74CE" box="[294,649,903,927]" italics="true" pageId="1" pageNumber="201">Sternoppia (Luisternoppia) subiasi</emphasis>
</taxonomicName>
<emphasis id="B960EAB8F862FFC1E9E2FC1EAAC974CE" bold="true" box="[655,730,903,927]" pageId="1" pageNumber="201">
<taxonomicNameLabel id="A25357C3F862FFC1E9E2FC1EAAC974CE" box="[655,730,903,927]" pageId="1" pageNumber="201" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30E6521F862FFC1EBFAFC54AD2473CA" pageId="1" pageNumber="201" type="diagnosis">
<paragraph id="8BAB36AAF862FFC1EBFAFC54AD2473CA" blockId="1.[151,1437,973,1432]" pageId="1" pageNumber="201">
<emphasis id="B960EAB8F862FFC1EBFAFC54A98A74B6" bold="true" box="[151,409,973,999]" pageId="1" pageNumber="201">Subgeneric diagnosis.</emphasis>
With character states of
<taxonomicName id="4C144D29F862FFC1E9C4FC57AC5874B6" authority="Balogh &amp; Mahunka, 1968" authorityName="Balogh &amp; Mahunka" authorityYear="1968" box="[681,1099,973,1000]" class="Arachnida" family="Sternoppiidae" genus="Sternoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="201" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAB8F862FFC1E9C4FC57AB3774B9" box="[681,804,974,1000]" italics="true" pageId="1" pageNumber="201">Sternoppia</emphasis>
<bibRefCitation id="EF854B5BF862FFC1E847FC54AC5874B6" author="Balogh, J. &amp; Mahunka, S." box="[810,1099,973,999]" pageId="1" pageNumber="201" pagination="31 - 69" refId="ref3154" refString="Balogh, J. &amp; Mahunka, S. (1968) The zoological results of the Hungarian soil zoological expeditions to South America. 5. Acari: Data to the oribatid fauna of the environment of Cordoba, Argentina. Opuscula Zoologica Budapest, VIII (2), 31 - 69." type="journal article" year="1968">Balogh &amp; Mahunka, 1968</bibRefCitation>
</taxonomicName>
(also see
<bibRefCitation id="EF854B5BF862FFC1EFDAFC54AD8374B6" author="Ermilov, S. G. &amp; Subias, L. S. &amp; Shtanchaeva, U. Y. &amp; Friedrich, A." box="[1207,1424,973,1000]" pageId="1" pageNumber="201" pagination="373 - 384" refId="ref3326" refString="Ermilov, S. G., Subias, L. S., Shtanchaeva, U. Y. &amp; Friedrich, A. (2022) A new subgenus and three new species of Sternoppia (Acari, Oribatida, Sternoppiidae) from Peru. Zootaxa, 5195 (4), 373 - 384. https: // doi. org / 10.11646 / zootaxa. 5195.4.4" type="journal article" year="2022">Ermilov et al, 2022</bibRefCitation>
), with the following: Distinct transcostula present. Bothridial seta„dendritiform“, with well-developed fusiform head, ciliated distally with 56 long, mostly distally furcate or trifurcate cilia. Notogastral setae of
<emphasis id="B960EAB8F862FFC1EFD7FB8CACD8737E" box="[1210,1227,1045,1071]" italics="true" pageId="1" pageNumber="201">l-</emphasis>
and
<emphasis id="B960EAB8F862FFC1EE69FB8CAD08737E" box="[1284,1307,1045,1071]" italics="true" pageId="1" pageNumber="201">h-</emphasis>
series long, reaching beyond the insertions of subsequent setae, insertions of
<emphasis id="B960EAB8F862FFC1E8E9FBA0ABB37302" box="[900,928,1081,1107]" italics="true" pageId="1" pageNumber="201">lm</emphasis>
clearly anteriomediad to insertions of
<emphasis id="B960EAB8F862FFC1EE32FBA0AD667302" box="[1375,1397,1081,1107]" italics="true" pageId="1" pageNumber="201">la</emphasis>
. In addition to epimeral seta
<emphasis id="B960EAB8F862FFC1EADAFBC4A9C07326" box="[439,467,1117,1143]" italics="true" pageId="1" pageNumber="201">3a</emphasis>
, also
<emphasis id="B960EAB8F862FFC1E975FBC4AA277326" box="[536,564,1117,1143]" italics="true" pageId="1" pageNumber="201">4a</emphasis>
strongly modified, penicillate to ramose, thick, with long and sharp ciliae or rami, directed anteriad. Similarly modified setae present on ventral side of tarsi II and III (seta
<emphasis id="B960EAB8F862FFC1EFD0FB1BAD3573CA" box="[1213,1318,1154,1179]" italics="true" pageId="1" pageNumber="201">pv, s, a, u</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E6521F862FFC1EBAAFB3CAC8A72C6" pageId="1" pageNumber="201" type="discussion">
<paragraph id="8BAB36AAF862FFC1EBAAFB3CAC6E73EE" blockId="1.[151,1437,973,1432]" box="[199,1149,1189,1216]" pageId="1" pageNumber="201">
<emphasis id="B960EAB8F862FFC1EBAAFB3CA95D73EE" bold="true" box="[199,334,1189,1215]" pageId="1" pageNumber="201">Etymology.</emphasis>
The subgeneric name prefix refers to given name of Luis Santos Subías.
</paragraph>
<paragraph id="8BAB36AAF862FFC1EBAAFB50AC8A72C6" blockId="1.[151,1437,973,1432]" pageId="1" pageNumber="201">
<emphasis id="B960EAB8F862FFC1EBAAFB50A92873B2" bold="true" box="[199,315,1225,1251]" pageId="1" pageNumber="201">Remarks.</emphasis>
The presence of transcostula (or translamella) in higher oribatids is often used as a generic character and could probably justify establishing a separate genus within
<taxonomicName id="4C144D29F862FFC1E81EFB74AC017256" authorityName="Balogh &amp; Mahunka" authorityYear="1969" box="[883,1042,1261,1287]" class="Arachnida" family="Sternoppiidae" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="201" phylum="Arthropoda" rank="family">Sternoppiidae</taxonomicName>
. The new species is however very similar in general morphology to species from the genus
<taxonomicName id="4C144D29F862FFC1E878FA8BAB83727D" authorityName="Balogh &amp; Mahunka" authorityYear="1968" box="[789,912,1298,1324]" class="Arachnida" family="Sternoppiidae" genus="Sternoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="201" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAB8F862FFC1E878FA8BAB83727D" box="[789,912,1298,1324]" italics="true" pageId="1" pageNumber="201">Sternoppia</emphasis>
</taxonomicName>
in most morphological characters, particularly in the development of the bothridial seta and ventral part of the body. As this is the first and single species with these distinctive characters, placement within
<taxonomicName id="4C144D29F862FFC1E930FAC3AACB7225" authorityName="Balogh &amp; Mahunka" authorityYear="1968" box="[605,728,1370,1396]" class="Arachnida" family="Sternoppiidae" genus="Sternoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="201" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAB8F862FFC1E930FAC3AACB7225" box="[605,728,1370,1396]" italics="true" pageId="1" pageNumber="201">Sternoppia</emphasis>
</taxonomicName>
as a separate subgenus is proposed. Further discovery of other species with these subgenus characters may eventually lead to its elevation to generic level.
</paragraph>
</subSubSection>
<subSubSection id="C30E6521F862FFC2EBFAFA70ABF97680" lastPageId="2" lastPageNumber="202" pageId="1" pageNumber="201" type="description">
<paragraph id="8BAB36AAF862FFC1EBFAFA70A9837152" blockId="1.[151,400,1513,1575]" box="[151,400,1513,1539]" pageId="1" pageNumber="201">
<emphasis id="B960EAB8F862FFC1EBFAFA70A9837152" bold="true" box="[151,400,1513,1539]" pageId="1" pageNumber="201">Description of adult</emphasis>
</paragraph>
<paragraph id="8BAB36AAF862FFC1EBFAF994A91F7176" blockId="1.[151,400,1513,1575]" box="[151,268,1549,1575]" pageId="1" pageNumber="201">
(
<figureCitation id="132F2A2FF862FFC1EBF2F994A8F57176" box="[159,230,1549,1575]" captionStart="FIGURE 1" captionStartId="2.[152,255,1982,2006]" captionTargetBox="[157,1430,998,1953]" captionTargetId="figure-402@2.[151,1435,993,1958]" captionTargetPageId="2" captionText="FIGURE 1. Sternoppia (Luisternoppia) subiasi sp. nov., adult (holotype). A—dorsal view; B—ventral view; C—detail of genital shield and setae (enlarged, not to scale). Scale bar indicating 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14595074" httpUri="https://zenodo.org/record/14595074/files/figure.png" pageId="1" pageNumber="201">Figs 1</figureCitation>
,
<figureCitation id="132F2A2FF862FFC1EB99F994A9107176" box="[244,259,1549,1575]" captionStart="FIGURE 2" captionStartId="3.[152,255,1123,1147]" captionTargetBox="[156,1430,186,1094]" captionTargetId="figure-25@3.[151,1435,181,1099]" captionTargetPageId="3" captionText="FIGURE 2. Sternoppia (Luisternoppia) subiasi sp. nov., adult (holotype). A—infracapitulum; B—bothridial area with base of costulae, bothridial and interlamellar seta; C—selected epimeral setae; D—prodorsal setae; E—selected notogastral setae; F—tibia and tarsus of leg III. Scale bars indicating: 50 µm (bar near D for A, C, D, E), 50 µm (B), and 100 µm (F)." figureDoi="http://doi.org/10.5281/zenodo.14595076" httpUri="https://zenodo.org/record/14595076/files/figure.png" pageId="1" pageNumber="201">2</figureCitation>
)
</paragraph>
<paragraph id="8BAB36AAF862FFC1EBFAF9CFAC92713E" blockId="1.[151,1437,1621,2008]" box="[151,1153,1621,1648]" pageId="1" pageNumber="201">
<emphasis id="B960EAB8F862FFC1EBFAF9CFA952713E" box="[151,321,1622,1647]" italics="true" pageId="1" pageNumber="201">Measurements.</emphasis>
<typeStatus id="54AF8808F862FFC1EA2AF9CCA9BC713E" box="[327,431,1621,1647]" pageId="1" pageNumber="201" type="holotype">Holotype</typeStatus>
body length: 335; notogaster length: 215; notogaster width: 228
</paragraph>
<paragraph id="8BAB36AAF862FFC1EBAAF9E3ABAA71AE" blockId="1.[151,1437,1621,2008]" pageId="1" pageNumber="201">
<emphasis id="B960EAB8F862FFC1EBAAF9E3A95E71C2" box="[199,333,1658,1683]" italics="true" pageId="1" pageNumber="201">Integument.</emphasis>
Body color brown. Body surface smooth, lateral parts of prodorsum anterior and laterad to bothridium and between bothridium and acetabula I and II, as well as on lateral parts of epimeres I with fine and dense granulation. Lateral part of pedotectum I with reticulate pattern, dorsal part of podosoma above acetabula III and IV and discidium with larger flat tubercles, covered by granulation.
</paragraph>
<paragraph id="8BAB36AAF862FFC2EBAAF890AD5277A8" blockId="1.[151,1437,1621,2008]" lastBlockId="2.[151,1437,151,933]" lastPageId="2" lastPageNumber="202" pageId="1" pageNumber="201">
<emphasis id="B960EAB8F862FFC1EBAAF890A95F7072" box="[199,332,1801,1827]" italics="true" pageId="1" pageNumber="201">Prodorsum.</emphasis>
Rostrum rounded or very slightly angulate. Costula well developed, with slightly arched proximal part, well developed postcostular tubercle and another subcostular cuticular thickening present around insertion of interlamellar seta, reaching bothridium. Bothridium quite small, with small postbothridial tubercle, another, large and elongated tubercle present anterolateral to bothridium. Transcostula present, well developed and quite broad, slightly concave flatly U-shaped, forming together with anterior part of costulae blunt ceratiform projections. Tectopedial field with distinctly thickened, costular border anteriad and anteriolaterad, with row of irregular sigillae laterad and anteriolaterad to costulae. Bothridial seta with well-developed fusiform head, ciliated distally with 56 long, mostly distally furcate or trifurcate cilia. Rostral (44), lamellar (62), and interlamellar (71) setae long, setiform,
<emphasis id="B960EAB8F861FFC2EA6FFF46A90977A9" box="[258,282,223,248]" italics="true" pageId="2" pageNumber="202">ro</emphasis>
densely,
<emphasis id="B960EAB8F861FFC2EAEEFF46A98A77A9" box="[387,409,223,248]" italics="true" pageId="2" pageNumber="202">in</emphasis>
and
<emphasis id="B960EAB8F861FFC2EAA2FF46A9F077A8" box="[463,483,223,249]" italics="true" pageId="2" pageNumber="202">le</emphasis>
sparsely, shortly ciliate. Exobothridial seta (45) rectangularly bent, setiform.
</paragraph>
<paragraph id="8BAB36AAF861FFC2EBAAFE9AABF97680" blockId="2.[151,1437,151,933]" pageId="2" pageNumber="202">
<emphasis id="B960EAB8F861FFC2EBAAFE9AA950764D" box="[199,323,259,284]" italics="true" pageId="2" pageNumber="202">Notogaster</emphasis>
. Roughly circular, broader than its length. Ten pairs of notogastral setae,
<emphasis id="B960EAB8F861FFC2EF13FE9AAC99764D" box="[1150,1162,259,284]" italics="true" pageId="2" pageNumber="202">c</emphasis>
(6) short setiform, setae of
<emphasis id="B960EAB8F861FFC2EBD5FEBEA8DA7610" box="[184,201,295,321]" italics="true" pageId="2" pageNumber="202">l-</emphasis>
and
<emphasis id="B960EAB8F861FFC2EA69FEBEA9087610" box="[260,283,295,321]" italics="true" pageId="2" pageNumber="202">h-</emphasis>
series setiform, sparsely ciliate, very long (
<emphasis id="B960EAB8F861FFC2E866FEBEAB327610" box="[779,801,295,321]" italics="true" pageId="2" pageNumber="202">la</emphasis>
93;
<emphasis id="B960EAB8F861FFC2E834FEBEAB667610" box="[857,885,295,321]" italics="true" pageId="2" pageNumber="202">lm</emphasis>
87;
<emphasis id="B960EAB8F861FFC2E8C1FEBEABD17610" box="[940,962,295,321]" italics="true" pageId="2" pageNumber="202">lp</emphasis>
81;
<emphasis id="B960EAB8F861FFC2E897FEBEAC037616" box="[1018,1040,295,327]" italics="true" pageId="2" pageNumber="202">
h
<subScript id="179034EFF861FFC2EF65FEA0AC037616" attach="left" box="[1032,1040,313,327]" fontSize="6" pageId="2" pageNumber="202">1</subScript>
</emphasis>
111;
<emphasis id="B960EAB8F861FFC2EF39FEBEAC797616" box="[1108,1130,295,327]" italics="true" pageId="2" pageNumber="202">
h
<subScript id="179034EFF861FFC2EF0FFEA0AC797616" attach="left" box="[1122,1130,313,327]" fontSize="6" pageId="2" pageNumber="202">2</subScript>
</emphasis>
90;
<emphasis id="B960EAB8F861FFC2EFCFFEBEACAB7616" box="[1186,1208,295,327]" italics="true" pageId="2" pageNumber="202">
h
<subScript id="179034EFF861FFC2EFDDFEA0ACAB7616" attach="left" box="[1200,1208,313,327]" fontSize="6" pageId="2" pageNumber="202">3</subScript>
</emphasis>
132),
<emphasis id="B960EAB8F861FFC2EE68FEBEAD087616" box="[1285,1307,295,327]" italics="true" pageId="2" pageNumber="202">
h
<subScript id="179034EFF861FFC2EE7EFEA0AD087616" attach="left" box="[1299,1307,313,327]" fontSize="6" pageId="2" pageNumber="202">3</subScript>
</emphasis>
clearly the longest, all of them reaching distinctly beyond insertions of subsequent seta. Setae
<emphasis id="B960EAB8F861FFC2EF5EFED2AC527634" box="[1075,1089,331,357]" italics="true" pageId="2" pageNumber="202">h</emphasis>
distally attenuated and slightly flagelliform. Setae
<emphasis id="B960EAB8F861FFC2EA00FEF6A96876D9" box="[365,379,367,392]" italics="true" pageId="2" pageNumber="202">p</emphasis>
much shorter (3235), setiform, sparsely but distinctly ciliated. Seta
<emphasis id="B960EAB8F861FFC2EF16FEF6AC8476D8" box="[1147,1175,367,393]" italics="true" pageId="2" pageNumber="202">lm</emphasis>
inserted anteriomediad to
<emphasis id="B960EAB8F861FFC2EBDFFE0AA8DB76FC" box="[178,200,403,429]" italics="true" pageId="2" pageNumber="202">la</emphasis>
. Lyrifissure
<emphasis id="B960EAB8F861FFC2EA3FFE0AA97B76FD" box="[338,360,403,428]" italics="true" pageId="2" pageNumber="202">ia</emphasis>
and
<emphasis id="B960EAB8F861FFC2EAF7FE0AA9A576FD" box="[410,438,403,428]" italics="true" pageId="2" pageNumber="202">im</emphasis>
visible, quite long (612), other lyrifissures not observed.Two small round spots (diameter around 2) present, one posterolaterad to
<emphasis id="B960EAB8F861FFC2E936FE2EAA627680" box="[603,625,439,465]" italics="true" pageId="2" pageNumber="202">lp</emphasis>
insertion, one anterolaterad to
<emphasis id="B960EAB8F861FFC2E8A0FE2EABF07686" box="[973,995,439,471]" italics="true" pageId="2" pageNumber="202">
h
<subScript id="179034EFF861FFC2E8B6FE50ABF07686" attach="left" box="[987,995,457,471]" fontSize="6" pageId="2" pageNumber="202">2</subScript>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E6521F861FFC2EBAAFE42AD9374F4" pageId="2" pageNumber="202" type="reference_group">
<paragraph id="8BAB36AAF861FFC2EBAAFE42ACA2756C" blockId="2.[151,1437,151,933]" pageId="2" pageNumber="202">
<taxonomicName id="4C144D29F861FFC2EBAAFE42A94576A4" authorityName="Krefft" authorityYear="1871" box="[199,342,475,501]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="2" pageNumber="202" phylum="Nematoda" rank="genus">
<emphasis id="B960EAB8F861FFC2EBAAFE42A94576A4" box="[199,342,475,501]" italics="true" pageId="2" pageNumber="202">Gnathosoma</emphasis>
</taxonomicName>
. Subcapitulum size 86 × 53; mentum length 33; rutellum distally broadened, hyaline, pointed anteriad. Subcapitular setae setiform,
<emphasis id="B960EAB8F861FFC2E953FE66AA5F7549" box="[574,588,511,536]" italics="true" pageId="2" pageNumber="202">a</emphasis>
(10) smooth,
<emphasis id="B960EAB8F861FFC2E984FE66AAEE7549" box="[745,765,511,536]" italics="true" pageId="2" pageNumber="202">m</emphasis>
(27) and
<emphasis id="B960EAB8F861FFC2E804FE66AB647548" box="[873,887,511,537]" italics="true" pageId="2" pageNumber="202">h</emphasis>
(34) with long, sparse ciliae, palp and chelicerae as in
<taxonomicName id="4C144D29F861FFC2EBBFFDBAAA52756C" authority="(Ermilov et al. 2022)" baseAuthorityName="Ermilov" baseAuthorityYear="2022" box="[210,577,547,573]" class="Arachnida" family="Sternoppiidae" genus="Sternoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAB8F861FFC2EBBFFDBAA95E756C" box="[210,333,547,573]" italics="true" pageId="2" pageNumber="202">Sternoppia</emphasis>
(
<bibRefCitation id="EF854B5BF861FFC2EA31FDBAAA2A756C" author="Ermilov, S. G. &amp; Subias, L. S. &amp; Shtanchaeva, U. Y. &amp; Friedrich, A." box="[348,569,547,573]" pageId="2" pageNumber="202" pagination="373 - 384" refId="ref3326" refString="Ermilov, S. G., Subias, L. S., Shtanchaeva, U. Y. &amp; Friedrich, A. (2022) A new subgenus and three new species of Sternoppia (Acari, Oribatida, Sternoppiidae) from Peru. Zootaxa, 5195 (4), 373 - 384. https: // doi. org / 10.11646 / zootaxa. 5195.4.4" type="journal article" year="2022">
Ermilov
<emphasis id="B960EAB8F861FFC2EAADFDBAA9EA756C" box="[448,505,547,573]" italics="true" pageId="2" pageNumber="202">et al.</emphasis>
2022
</bibRefCitation>
)
</taxonomicName>
(not studied in detail due to limited material available).
</paragraph>
<paragraph id="8BAB36AAF861FFC2EBAAFDDEAD9374F4" blockId="2.[151,1437,151,933]" pageId="2" pageNumber="202">
<emphasis id="B960EAB8F861FFC2EBAAFDDEAA847531" box="[199,663,583,609]" italics="true" pageId="2" pageNumber="202">Epimeral and lateral podosomal regions.</emphasis>
Epimeral plates with complicated tecta, these tecta on epimere I and partly also epimere II overlapping, with distinct angular lobes anteriomediad, bearing insertion of seta
<emphasis id="B960EAB8F861FFC2EF91FDF2AD0A75D4" box="[1276,1305,619,645]" italics="true" pageId="2" pageNumber="202">1a</emphasis>
. Tecta from epimeres II and III not touching medially, surrounding slightly depressed anterogenital area and bearing distinct and modified epimeral setae
<emphasis id="B960EAB8F861FFC2EAC2FD2AA9D8759C" box="[431,459,691,717]" italics="true" pageId="2" pageNumber="202">3a</emphasis>
, tectum of epimere 4 framing epimeral posterior margin, well developed and distinct. Epimeral setae of various shapes and sizes. Setae
<emphasis id="B960EAB8F861FFC2E9A3FD4EAAFA75A0" box="[718,745,727,753]" italics="true" pageId="2" pageNumber="202">3a</emphasis>
(33) strong, thickened, ramose with rami of different length, pointed mediad, directed against each-other. Setae
<emphasis id="B960EAB8F861FFC2E9BCFD62AAFD7444" box="[721,750,763,789]" italics="true" pageId="2" pageNumber="202">4a</emphasis>
(27) strong, penicillate, standing close to each-other, parallel and oriented anteriad, inserted on thickened cuticular arch anterior to genital opening. Other epimeral setae long, setiform, sparsely but distinctly ciliated, seta
<emphasis id="B960EAB8F861FFC2E9F7FCDAAAA5740C" box="[666,694,835,861]" italics="true" pageId="2" pageNumber="202">1a</emphasis>
shorter (25), strongly bend and slightly thickened at the base;
<emphasis id="B960EAB8F861FFC2EEEDFCDAAD8E740C" box="[1408,1437,835,861]" italics="true" pageId="2" pageNumber="202">1b</emphasis>
longer (44), also with thickened proximal (basal) part. Other epimeral setae setiform and sparsely ciliated,
<emphasis id="B960EAB8F861FFC2EE5DFCFEAD8F74D0" box="[1328,1436,871,897]" italics="true" pageId="2" pageNumber="202">2a, 3b, 3c</emphasis>
shorter (4144) and
<emphasis id="B960EAB8F861FFC2EA16FC12A9D374F4" box="[379,448,907,933]" italics="true" pageId="2" pageNumber="202">1c, 4b</emphasis>
and
<emphasis id="B960EAB8F861FFC2EA9BFC12AA0374F4" box="[502,528,907,933]" italics="true" pageId="2" pageNumber="202">4c</emphasis>
much longer (60, 66, and 54, respectively). Discidium rather large, dentiform.
</paragraph>
</subSubSection>
<caption id="DF6B6622F861FFC2EBF5F827ABB070AB" ID-DOI="http://doi.org/10.5281/zenodo.14595074" ID-Zenodo-Dep="14595074" httpUri="https://zenodo.org/record/14595074/files/figure.png" pageId="2" pageNumber="202" startId="2.[152,255,1982,2006]" targetBox="[157,1430,998,1953]" targetPageId="2" targetType="figure">
<paragraph id="8BAB36AAF861FFC2EBF5F827ABB070AB" blockId="2.[151,1436,1982,2042]" pageId="2" pageNumber="202">
<emphasis id="B960EAB8F861FFC2EBF5F827A9087087" bold="true" box="[152,283,1982,2006]" pageId="2" pageNumber="202">FIGURE 1.</emphasis>
<taxonomicName id="4C144D29F861FFC2EA49F827AA9E7087" authorityName="Miko" authorityYear="2024" box="[292,653,1982,2006]" class="Arachnida" family="Sternoppiidae" genus="Sternoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="202" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov." subGenus="Luisternoppia">
<emphasis id="B960EAB8F861FFC2EA49F827AA9E7087" box="[292,653,1982,2006]" italics="true" pageId="2" pageNumber="202">Sternoppia (Luisternoppia) subiasi</emphasis>
</taxonomicName>
<emphasis id="B960EAB8F861FFC2E9FBF827AAFA7087" bold="true" box="[662,745,1982,2006]" pageId="2" pageNumber="202">
<taxonomicNameLabel id="A25357C3F861FFC2E9FBF827AAFA7087" box="[662,745,1982,2006]" pageId="2" pageNumber="202" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult (holotype). A—dorsal view; B—ventral view; C—detail of genital shield and setae (enlarged, not to scale). Scale bar indicating 100 µm.
</paragraph>
</caption>
<caption id="DF6B6622F860FFC3EBF5FBFAAD357392" ID-DOI="http://doi.org/10.5281/zenodo.14595076" ID-Zenodo-Dep="14595076" httpUri="https://zenodo.org/record/14595076/files/figure.png" pageId="3" pageNumber="203" startId="3.[152,255,1123,1147]" targetBox="[156,1430,186,1094]" targetPageId="3" targetType="figure">
<paragraph id="8BAB36AAF860FFC3EBF5FBFAAD357392" blockId="3.[151,1436,1123,1220]" pageId="3" pageNumber="203">
<emphasis id="B960EAB8F860FFC3EBF5FBFAA909732A" bold="true" box="[152,282,1123,1147]" pageId="3" pageNumber="203">FIGURE 2.</emphasis>
<taxonomicName id="4C144D29F860FFC3EA4FFBFAAA9A732A" authorityName="Miko" authorityYear="2024" box="[290,649,1123,1147]" class="Arachnida" family="Sternoppiidae" genus="Sternoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="203" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov." subGenus="Luisternoppia">
<emphasis id="B960EAB8F860FFC3EA4FFBFAAA9A732A" box="[290,649,1123,1147]" italics="true" pageId="3" pageNumber="203">Sternoppia (Luisternoppia) subiasi</emphasis>
</taxonomicName>
<emphasis id="B960EAB8F860FFC3E9FCFBFAAAF0732A" bold="true" box="[657,739,1123,1147]" pageId="3" pageNumber="203">
<taxonomicNameLabel id="A25357C3F860FFC3E9FCFBFAAAF0732A" box="[657,739,1123,1147]" pageId="3" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult (holotype). A—infracapitulum; B—bothridial area with base of costulae, bothridial and interlamellar seta; C—selected epimeral setae; D—prodorsal setae; E—selected notogastral setae; F—tibia and tarsus of leg III. Scale bars indicating: 50 µm (bar near D for A, C, D, E), 50 µm (B), and 100 µm (F).
</paragraph>
</caption>
<subSubSection id="C30E6521F860FFC3EBAAFA8CA91771C6" pageId="3" pageNumber="203" type="description">
<paragraph id="8BAB36AAF860FFC3EBAAFA8CAA4172EE" blockId="3.[151,1437,1301,2012]" pageId="3" pageNumber="203">
<emphasis id="B960EAB8F860FFC3EBAAFA8CA982727E" box="[199,401,1301,1327]" italics="true" pageId="3" pageNumber="203">Anogenital region</emphasis>
. Genital opening (39 x 39) and anal opening (66 x 66) rather small, separated by broad strip of ventral plate. Genito-anal formula
<quantity id="4CEC9B4FF860FFC3E92DFAA0AA4D7202" box="[576,606,1337,1363]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="3" pageNumber="203" unit="g" value="6.0">
6
<emphasis id="B960EAB8F860FFC3E923FAA3AA4D7202" box="[590,606,1338,1363]" italics="true" pageId="3" pageNumber="203">g</emphasis>
</quantity>
, 1
<emphasis id="B960EAB8F860FFC3E914FAA3AA8F7202" box="[633,668,1338,1363]" italics="true" pageId="3" pageNumber="203">ag,</emphasis>
2
<emphasis id="B960EAB8F860FFC3E9DFFAA3AAC67202" box="[690,725,1338,1363]" italics="true" pageId="3" pageNumber="203">an,</emphasis>
<specimenCount id="9D12FD23F860FFC3E9B3FAA0AB1C7205" box="[734,783,1337,1364]" count="3" pageId="3" pageNumber="203" type="generic">
3
<emphasis id="B960EAB8F860FFC3E981FAA0AB1B7202" box="[748,776,1337,1363]" italics="true" pageId="3" pageNumber="203">ad</emphasis>
.
</specimenCount>
Genital setae (1317) setiform, with 12 bristles distally. Anal setae (1925) setiform, distinctly ciliated. Aggenital and adanal setae of similar length (3136), setiform and ciliated,
<emphasis id="B960EAB8F860FFC3EB9AFA18A90872F3" box="[247,283,1409,1442]" italics="true" pageId="3" pageNumber="203">
ad
<subScript id="179034EFF860FFC3EA7EFA0DA90872F3" attach="left" box="[275,283,1428,1442]" fontSize="6" pageId="3" pageNumber="203">3</subScript>
</emphasis>
strongly shifted laterad, distance
<emphasis id="B960EAB8F860FFC3E9FEFA18AAF972F3" box="[659,746,1409,1442]" italics="true" pageId="3" pageNumber="203">
ad
<subScript id="179034EFF860FFC3E9C2FA0DAAA472F3" attach="left" box="[687,695,1428,1442]" fontSize="6" pageId="3" pageNumber="203">3</subScript>
ad
<subScript id="179034EFF860FFC3E98FFA0DAAF972F3" attach="left" box="[738,746,1428,1442]" fontSize="6" pageId="3" pageNumber="203">3</subScript>
</emphasis>
roughly 1,6 times more than distance
<emphasis id="B960EAB8F860FFC3EFF8FA18ACF872F3" box="[1173,1259,1409,1442]" italics="true" pageId="3" pageNumber="203">
ad
<subScript id="179034EFF860FFC3EFDCFA0DACAA72F3" attach="left" box="[1201,1209,1428,1442]" fontSize="6" pageId="3" pageNumber="203">1</subScript>
ad
<subScript id="179034EFF860FFC3EF8EFA0DACF872F3" attach="left" box="[1251,1259,1428,1442]" fontSize="6" pageId="3" pageNumber="203">1</subScript>
</emphasis>
. Lyrifissure
<emphasis id="B960EAB8F860FFC3EE15FA18AD8F72CA" box="[1400,1436,1409,1435]" italics="true" pageId="3" pageNumber="203">iad</emphasis>
(1819) close and parallel to anal plate.
</paragraph>
<paragraph id="8BAB36AAF860FFC3EBAAFA53A91771C6" blockId="3.[151,1437,1301,2012]" pageId="3" pageNumber="203">
<emphasis id="B960EAB8F860FFC3EBAAFA53A8EF72B2" box="[199,252,1482,1507]" italics="true" pageId="3" pageNumber="203">Legs</emphasis>
. Leg setation and solenidia as in other
<taxonomicName id="4C144D29F860FFC3E9DEFA53AC3672B2" authority="(Ermilov et al. 2022)" baseAuthorityName="Ermilov" baseAuthorityYear="2022" box="[691,1061,1481,1508]" class="Arachnida" family="Sternoppiidae" genus="Sternoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAB8F860FFC3E9DEFA53AB3D72B5" box="[691,814,1482,1508]" italics="true" pageId="3" pageNumber="203">Sternoppia</emphasis>
(
<bibRefCitation id="EF854B5BF860FFC3E851FA50AC0F72B2" author="Ermilov, S. G. &amp; Subias, L. S. &amp; Shtanchaeva, U. Y. &amp; Friedrich, A." box="[828,1052,1481,1508]" pageId="3" pageNumber="203" pagination="373 - 384" refId="ref3326" refString="Ermilov, S. G., Subias, L. S., Shtanchaeva, U. Y. &amp; Friedrich, A. (2022) A new subgenus and three new species of Sternoppia (Acari, Oribatida, Sternoppiidae) from Peru. Zootaxa, 5195 (4), 373 - 384. https: // doi. org / 10.11646 / zootaxa. 5195.4.4" type="journal article" year="2022">
Ermilov
<emphasis id="B960EAB8F860FFC3E8CCFA53ABC072B2" box="[929,979,1481,1507]" italics="true" pageId="3" pageNumber="203">et al</emphasis>
. 2022
</bibRefCitation>
)
</taxonomicName>
, formulas: I (1-5-2-4-20) [1-2-2], II (1-5-2-4-16) [1-1-2], III (2-3-1-3-15) [1-1-0], IV (1-2-2-3-12) [0-1-0]. Famulus setiform, tarsal solenidia blunt, other setiform, proral setae on legs II-IV short, thorn-like, some ventral setae on tarsi IIII (
<emphasis id="B960EAB8F860FFC3EFCFF98BAD3A717D" box="[1186,1321,1554,1580]" italics="true" pageId="3" pageNumber="203">u, a, s, pv“)</emphasis>
modified, stronger, with long and acute cilia, penicillate. Seta
<emphasis id="B960EAB8F860FFC3E985F9AFAAE87101" box="[744,763,1590,1616]" italics="true" pageId="3" pageNumber="203">v</emphasis>
of trochanter I and II very long, setiform, densely ciliated. Remaining setae mostly setiform and ciliated. More detailed study of legs not performed due to limited material available.
</paragraph>
</subSubSection>
<subSubSection id="C30E6521F860FFC3EBAAF938AB5C703E" pageId="3" pageNumber="203" type="materials_examined">
<paragraph id="8BAB36AAF860FFC3EBAAF938AB487076" blockId="3.[151,1437,1301,2012]" pageId="3" pageNumber="203">
<emphasis id="B960EAB8F860FFC3EBAAF938A96471EA" bold="true" box="[199,375,1697,1723]" pageId="3" pageNumber="203">Type material.</emphasis>
<materialsCitation id="3B7C3CF7F860FFC3EA12F938AA0B71EA" box="[383,536,1697,1723]" collectedFrom="litter, dead leaves, rotting wood on the soil surface" collectingDate="2012-03-10" collectionCode="GSM" collectorName="Ladislav Miko" country="French Guyana" county="Montagne de Kaw" elevation="220" latitude="4.5545135" location="South America" longLatPrecision="1" longitude="-52.171684" municipality="French" pageId="3" pageNumber="203" specimenCount="1" typeStatus="holotype">
<typeStatus id="54AF8808F860FFC3EA12F938A9F471EA" box="[383,487,1697,1723]" pageId="3" pageNumber="203" type="holotype">Holotype</typeStatus>
and
</materialsCitation>
<materialsCitation id="3B7C3CF7F860FFC3E94CF93BAB487076" collectedFrom="litter, dead leaves, rotting wood on the soil surface" collectingDate="2012-03-10" collectionCode="GSM" collectorName="Ladislav Miko" country="French Guyana" county="Montagne de Kaw" elevation="220" latitude="4.5545135" location="South America" longLatPrecision="1" longitude="-52.171684" municipality="French" pageId="3" pageNumber="203" specimenCount="1" typeStatus="paratype">
one damaged paratype (sex not studied) from
<location id="8ECB6071F860FFC3EF42F938ACC871EA" LSID="urn:lsid:plazi:treatment:03BD87BCF862FFC4EBFAFC11AD8F7454:8ECB6071F860FFC3EF42F938ACC871EA" box="[1071,1243,1697,1723]" country="French Guyana" county="Montagne de Kaw" latitude="4.5545135" longLatPrecision="1" longitude="-52.171684" municipality="French" name="South America" pageId="3" pageNumber="203">South America</location>
,
<collectingCountry id="F303763AF860FFC3EF85F938AD8B71EA" box="[1256,1432,1697,1723]" pageId="3" pageNumber="203">French Guyana</collectingCountry>
, sample LM-2012-008:
<collectingCounty id="62CA4E26F860FFC3EAF2F95FAA6771B1" box="[415,628,1733,1760]" pageId="3" pageNumber="203">Montagne de Kaw</collectingCounty>
,
<collectionCode id="ED05AE6FF860FFC3E9ECF95CAAAD718E" box="[641,702,1733,1759]" country="United Kingdom" httpUri="http://grbio.org/cool/fzze-3s97" name="Geologic Museum" pageId="3" pageNumber="203">GSM</collectionCode>
<geoCoordinate id="EE20506DF860FFC3E9AAF95CAB65718E" box="[711,886,1733,1759]" degrees="4" direction="north" minutes="33" orientation="latitude" pageId="3" pageNumber="203" precision="1" seconds="16.248" value="4.5545135">4°3316.248”N</geoCoordinate>
,
<geoCoordinate id="EE20506DF860FFC3E8EEF95CAC5771B1" box="[899,1092,1733,1760]" degrees="52" direction="west" minutes="10" orientation="longitude" pageId="3" pageNumber="203" precision="1" seconds="18.061" value="-52.171684">52°1018.061”W</geoCoordinate>
,
<location id="8ECB6071F860FFC3EF3CF95FAD8B718E" LSID="urn:lsid:plazi:treatment:03BD87BCF862FFC4EBFAFC11AD8F7454:8ECB6071F860FFC3EF3CF95FAD8B718E" box="[1105,1432,1733,1760]" country="French Guyana" county="Montagne de Kaw" latitude="4.5545135" longLatPrecision="1" longitude="-52.171684" municipality="French" name="near Auberge Camp Caiman" pageId="3" pageNumber="203">near Auberge Camp Caiman</location>
,
<location id="8ECB6071F860FFC3EBFAF970AA327052" LSID="urn:lsid:plazi:treatment:03BD87BCF862FFC4EBFAFC11AD8F7454:8ECB6071F860FFC3EBFAF970AA327052" box="[151,545,1769,1796]" country="French Guyana" county="Montagne de Kaw" latitude="4.5545135" longLatPrecision="1" longitude="-52.171684" municipality="French" name="close to Reserve for Coq du Roche" pageId="3" pageNumber="203">close to Reserve for Coq du Roche</location>
,
<elevation id="0039D199F860FFC3E941F970AB3A7052" box="[556,809,1769,1796]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="203" unit="m" value="220.0">
<quantity id="4CEC9B4FF860FFC3E941F970AA677055" box="[556,628,1769,1796]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="203" unit="m" value="220.0">220 m</quantity>
above sea level
</elevation>
. Tropical rainforest
<collectedFrom id="38087557F860FFC3EF66F970AC517052" box="[1035,1090,1769,1795]" pageId="3" pageNumber="203">litter</collectedFrom>
,
<collectedFrom id="38087557F860FFC3EF20F970ACC17052" box="[1101,1234,1769,1795]" pageId="3" pageNumber="203">dead leaves</collectedFrom>
, and
<collectedFrom id="38087557F860FFC3EE61F970A9747076" pageId="3" pageNumber="203">rotting wood on the soil surface</collectedFrom>
, moist.
<date id="FFAA106AF860FFC3EAD3F894AA937076" box="[446,640,1805,1832]" pageId="3" pageNumber="203" value="2012-03-10">
<collectingDate id="EFEEE982F860FFC3EAD3F894AA937076" box="[446,640,1805,1832]" pageId="3" pageNumber="203" value="2012-03-10">
March 10
<superScript id="7C619BE2F860FFC3E947F894AA24704A" attach="none" box="[554,567,1805,1819]" fontSize="6" pageId="3" pageNumber="203">th</superScript>
, 2012
</collectingDate>
</date>
,
<collectorName id="26E1537CF860FFC3E9E6F894AB3C7076" box="[651,815,1805,1831]" pageId="3" pageNumber="203">Ladislav Miko</collectorName>
lgt.
</materialsCitation>
</paragraph>
<paragraph id="8BAB36AAF860FFC3EBAAF8A8AB5C703E" blockId="3.[151,1437,1301,2012]" pageId="3" pageNumber="203">
The holotype is deposited in the Acarological collection of the Senckenberg Museum in Gorlitz, Germany,
<typeStatus id="54AF8808F860FFC3EBFAF8CFA8E47021" box="[151,247,1878,1904]" pageId="3" pageNumber="203" type="paratype">paratype</typeStatus>
in collection of the author (
<collectingRegion id="49D0F848F860FFC3E942F8CCAA92703E" box="[559,641,1877,1903]" country="Czech Republic" name="Praha" pageId="3" pageNumber="203">Prague</collectingRegion>
,
<collectingCountry id="F303763AF860FFC3E9E1F8CCAB50703E" box="[652,835,1877,1903]" name="Czech Republic" pageId="3" pageNumber="203">Czech Republic</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E6521F860FFC3EBAAF8E0A92D708D" pageId="3" pageNumber="203" type="discussion">
<paragraph id="8BAB36AAF860FFC3EBAAF8E0A92D708D" blockId="3.[151,1437,1301,2012]" pageId="3" pageNumber="203">
<emphasis id="B960EAB8F860FFC3EBAAF8E0A92870C2" bold="true" box="[199,315,1913,1939]" pageId="3" pageNumber="203">Remarks.</emphasis>
The new species differs from all known
<taxonomicName id="4C144D29F860FFC3E86FF8E3ABC770C2" box="[770,980,1913,1940]" pageId="3" pageNumber="203">
<emphasis id="B960EAB8F860FFC3E86FF8E3AB6E70C5" box="[770,893,1914,1940]" italics="true" pageId="3" pageNumber="203">Sternoppia</emphasis>
species
</taxonomicName>
by the presence of a transcostula (versus transcostula absent), very long notogastral setae (versus setae of notogaster relatively shorter) and specific form of bothridial seta.
</paragraph>
</subSubSection>
<subSubSection id="C30E6521F867FFC4EBAAFF0FABA07785" pageId="4" pageNumber="204" type="etymology">
<paragraph id="8BAB36AAF867FFC4EBAAFF0FABA07785" blockId="4.[151,1436,150,213]" pageId="4" pageNumber="204">
<emphasis id="B960EAB8F867FFC4EBAAFF0FA95D77E1" bold="true" box="[199,334,150,176]" pageId="4" pageNumber="204">Etymology.</emphasis>
The species is named in honour of outstanding Spanish acarologist Luis Santos Subías, a reminder of his extensive work in taxonomy and biogeography of oribatid mites.
</paragraph>
</subSubSection>
<subSubSection id="C30E6521F867FFC4EBFAFEBFAD8F7454" pageId="4" pageNumber="204" type="key">
<paragraph id="8BAB36AAF867FFC4EBFAFEBFAA0E7611" blockId="4.[151,840,294,357]" box="[151,541,294,320]" pageId="4" pageNumber="204">
<heading id="D0E381C6F867FFC4EBFAFEBFAA0E7611" bold="true" box="[151,541,294,320]" fontSize="11" level="1" pageId="4" pageNumber="204" reason="1">
<emphasis id="B960EAB8F867FFC4EBFAFEBFAA0E7611" bold="true" box="[151,541,294,320]" pageId="4" pageNumber="204">
Key to subgenera of
<taxonomicName id="4C144D29F867FFC4EAF4FEBFAA0E7611" ID-CoL="645ZB" box="[409,541,294,320]" class="Arachnida" family="Sternoppiidae" genus="Sternoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAB8F867FFC4EAF4FEBFAA0E7611" bold="true" box="[409,541,294,320]" italics="true" pageId="4" pageNumber="204">Sternoppia</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAB36AAF867FFC4EBFAFED3AB5B7635" blockId="4.[151,840,294,357]" box="[151,840,330,357]" pageId="4" pageNumber="204">
A key to all other species is available in
<bibRefCitation id="EF854B5BF867FFC4E936FED3AB5B7635" author="Ermilov, S. G. &amp; Subias, L. S. &amp; Shtanchaeva, U. Y. &amp; Friedrich, A." box="[603,840,330,356]" pageId="4" pageNumber="204" pagination="373 - 384" refId="ref3326" refString="Ermilov, S. G., Subias, L. S., Shtanchaeva, U. Y. &amp; Friedrich, A. (2022) A new subgenus and three new species of Sternoppia (Acari, Oribatida, Sternoppiidae) from Peru. Zootaxa, 5195 (4), 373 - 384. https: // doi. org / 10.11646 / zootaxa. 5195.4.4" type="journal article" year="2022">
Ermilov
<emphasis id="B960EAB8F867FFC4E9D3FED2AAE57635" box="[702,758,330,356]" italics="true" pageId="4" pageNumber="204">et al.</emphasis>
(2022)
</bibRefCitation>
</paragraph>
<keyStep id="30E02E0FF867FFC4EBFAFE16AD8F754B" pageId="4" pageNumber="204">
<paragraph id="8BAB36AAF867FFC4EBFAFE16AD8F768E" blockId="4.[151,1437,399,773]" pageId="4" pageNumber="204">
<keyLead id="30E5959FF867FFC4EBFAFE16AD8F768E" pageId="4" pageNumber="204">
1. Bothridial seta bacilliform, with slightly developed, not ciliated head; interbothridial region with two pairs of tubercles; adanal setae not ciliated; body length: 660720.................................
<taxonomicName id="4C144D29F867FFC4E894FE34AD8F7693" authority="Ermilov et al., 2022" authorityName="Ermilov" authorityYear="2022" box="[1017,1436,428,450]" class="Arachnida" family="Sternoppiidae" genus="Sternoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="204" phylum="Arthropoda" rank="subGenus" subGenus="Andoppia">
<emphasis id="B960EAB8F867FFC4E894FE34ACC57690" box="[1017,1238,428,450]" italics="true" pageId="4" pageNumber="204">Sternoppia (Andoppia)</emphasis>
<bibRefCitation id="EF854B5BF867FFC4EFB1FE35AD8F7693" author="Ermilov, S. G. &amp; Subias, L. S. &amp; Shtanchaeva, U. Y. &amp; Friedrich, A." box="[1244,1436,428,450]" pageId="4" pageNumber="204" pagination="373 - 384" refId="ref3326" refString="Ermilov, S. G., Subias, L. S., Shtanchaeva, U. Y. &amp; Friedrich, A. (2022) A new subgenus and three new species of Sternoppia (Acari, Oribatida, Sternoppiidae) from Peru. Zootaxa, 5195 (4), 373 - 384. https: // doi. org / 10.11646 / zootaxa. 5195.4.4" type="journal article" year="2022">
Ermilov
<emphasis id="B960EAB8F867FFC4EE5DFE34AD727690" box="[1328,1377,428,450]" italics="true" pageId="4" pageNumber="204">et al.</emphasis>
, 2022
</bibRefCitation>
</taxonomicName>
............................................................Monotypical,
<typeStatus id="54AF8808F867FFC4EF50FE53AC75768E" box="[1085,1126,458,479]" pageId="4" pageNumber="204">type</typeStatus>
species:
<taxonomicName id="4C144D29F867FFC4EFD3FE53AD8F768E" authorityName="Ermilov, Subías, Shtanchaeva &amp; Friedrich" authorityYear="2022" box="[1214,1436,458,479]" class="Arachnida" family="Sternoppiidae" genus="Sternoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="204" phylum="Arthropoda" rank="species" species="tetratuberculata" subGenus="Andoppia">
<emphasis id="B960EAB8F867FFC4EFD3FE53AD8F768E" box="[1214,1436,458,479]" italics="true" pageId="4" pageNumber="204">S. (A.) tetratuberculata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAB36AAF867FFC4EBFAFE71AD8F754B" blockId="4.[151,1437,399,773]" pageId="4" pageNumber="204">
<keyLead id="30E5959FF867FFC4EBFAFE71AD8F754B" pageId="4" pageNumber="204">- Bothridial seta setiform or slightly fusiform, unilaterally or distally ciliated; interbothridial region without or with one pair of tubercles; adanal setae ciliated.......................................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E02E0FF867FFC4EBFAFDBBAD8F7454" pageId="4" pageNumber="204">
<paragraph id="8BAB36AAF867FFC4EBFAFDBBAD8F75DE" blockId="4.[151,1437,399,773]" pageId="4" pageNumber="204">
<keyLead id="30E5959FF867FFC4EBFAFDBBAD8F75DE" pageId="4" pageNumber="204">
2. Distinct transcostula present, costula anteriad with angular distal end. Notogastral setae very long, reaching clearly over insertions of following setae. Epimeral setae
<emphasis id="B960EAB8F867FFC4E9F4FDA6AAA27505" box="[665,689,575,596]" italics="true" pageId="4" pageNumber="204">3a</emphasis>
and
<emphasis id="B960EAB8F867FFC4E98FFDA6AAE87505" box="[738,763,575,596]" italics="true" pageId="4" pageNumber="204">4a</emphasis>
modified, thick, penicillate or ramose, with long and dense rami and cilia...............................................................
<taxonomicName id="4C144D29F867FFC4EF7FFDC4AD0B7523" authorityName="Miko" authorityYear="2024" box="[1042,1304,605,626]" class="Arachnida" family="Sternoppiidae" genus="Sternoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="204" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Luisternoppia">
<emphasis id="B960EAB8F867FFC4EF7FFDC4AD0B7523" box="[1042,1304,605,626]" italics="true" pageId="4" pageNumber="204">Sternoppia (Luisternoppia)</emphasis>
</taxonomicName>
<emphasis id="B960EAB8F867FFC4EE73FDC4AD8F7523" bold="true" box="[1310,1436,605,626]" pageId="4" pageNumber="204">
<taxonomicNameLabel id="A25357C3F867FFC4EE73FDC4AD8F7523" box="[1310,1436,605,626]" pageId="4" pageNumber="204" rank="subGenus">subgen. nov.</taxonomicNameLabel>
</emphasis>
.............................................................Monotypical,
<typeStatus id="54AF8808F867FFC4EF27FDE3AC6075DE" box="[1098,1139,634,655]" pageId="4" pageNumber="204">type</typeStatus>
species
<taxonomicName id="4C144D29F867FFC4EFA9FDE3AD5A75DE" authorityName="Miko" authorityYear="2024" box="[1220,1353,634,655]" class="Arachnida" family="Sternoppiidae" genus="Sternoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="204" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov." subGenus="Luisternoppia">
<emphasis id="B960EAB8F867FFC4EFA9FDE3AD5A75DE" box="[1220,1353,634,655]" italics="true" pageId="4" pageNumber="204">S. (L.) subiasi</emphasis>
</taxonomicName>
<emphasis id="B960EAB8F867FFC4EE3DFDE3AD8F75DE" bold="true" box="[1360,1436,634,655]" pageId="4" pageNumber="204">
<taxonomicNameLabel id="A25357C3F867FFC4EE3DFDE3AD8F75DE" box="[1360,1436,634,655]" pageId="4" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BAB36AAF867FFC4EBFAFD01AD8F7454" blockId="4.[151,1437,399,773]" pageId="4" pageNumber="204">
<keyLead id="30E5959FF867FFC4EBFAFD01AD8F7454" pageId="4" pageNumber="204">
- Transcostula absent, costula anteriorly without angular ends. Notogastral setae usually short and not reaching insertions of following seta, rarely one or few setal pairs long. None or only one pair of epimeral seta (either
<emphasis id="B960EAB8F867FFC4EF18FD2CAC9E759B" box="[1141,1165,693,714]" italics="true" pageId="4" pageNumber="204">3a</emphasis>
or
<emphasis id="B960EAB8F867FFC4EFC0FD2CACD6759B" box="[1197,1221,693,714]" italics="true" pageId="4" pageNumber="204">4a</emphasis>
) usually modified and strongly ciliated...................................................
<taxonomicName id="4C144D29F867FFC4E887FD4BAC4075B6" ID-CoL="645ZB" box="[1002,1107,722,743]" class="Arachnida" family="Sternoppiidae" genus="Sternoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAB8F867FFC4E887FD4BAC4075B6" box="[1002,1107,722,743]" italics="true" pageId="4" pageNumber="204">Sternoppia</emphasis>
</taxonomicName>
(s. str.)
<bibRefCitation id="EF854B5BF867FFC4EFCFFD4BAD8F75B9" author="Balogh, J. &amp; Mahunka, S." box="[1186,1436,722,744]" pageId="4" pageNumber="204" pagination="31 - 69" refId="ref3154" refString="Balogh, J. &amp; Mahunka, S. (1968) The zoological results of the Hungarian soil zoological expeditions to South America. 5. Acari: Data to the oribatid fauna of the environment of Cordoba, Argentina. Opuscula Zoologica Budapest, VIII (2), 31 - 69." type="journal article" year="1968">Balogh &amp; Mahunka, 1968</bibRefCitation>
.................................................. 16 species,
<typeStatus id="54AF8808F867FFC4E8DFFD69ABC87454" box="[946,987,752,773]" pageId="4" pageNumber="204">type</typeStatus>
species
<taxonomicName id="4C144D29F867FFC4EF40FD76AD8F7454" ID-CoL="52FPT" authority="Balogh &amp; Mahunka, 1968" authorityName="Balogh &amp; Mahunka" authorityYear="1968" box="[1069,1436,751,773]" class="Arachnida" family="Sternoppiidae" genus="Sternoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="204" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="B960EAB8F867FFC4EF40FD76AC8F7455" box="[1069,1180,751,772]" italics="true" pageId="4" pageNumber="204">S. mirabilis</emphasis>
<bibRefCitation id="EF854B5BF867FFC4EFCFFD76AD8F7454" author="Balogh, J. &amp; Mahunka, S." box="[1186,1436,751,773]" pageId="4" pageNumber="204" pagination="31 - 69" refId="ref3154" refString="Balogh, J. &amp; Mahunka, S. (1968) The zoological results of the Hungarian soil zoological expeditions to South America. 5. Acari: Data to the oribatid fauna of the environment of Cordoba, Argentina. Opuscula Zoologica Budapest, VIII (2), 31 - 69." type="journal article" year="1968">Balogh &amp; Mahunka, 1968</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>