treatments-xml/data/03/BD/87/03BD87810E6FBD1AFF28FCAB82C0FC49.xml

386 lines
63 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="B54B5D5EFCEE256BE71EBCC8ADBBC55D" ID-DOI="10.11646/zootaxa.5061.2.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="5649671" ID-ZooBank="EE0C41ED-C10C-41FA-B913-73CFD910DADF" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1636145868530" checkinUser="plazi" docAuthor="Polizei, Thiago T. S. &amp; Hamada, Neusa" docDate="2021" docId="03BD87810E6FBD1AFF28FCAB82C0FC49" docLanguage="en" docName="zootaxa.5061.2.9.pdf" docOrigin="Zootaxa 5061 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pagelmis mura Polizei &amp; Hamada 2021, sp. nov." docType="treatment" docUuid="255E923D-08A8-417B-8D3E-FB8A998397E6" docUuidSource="ZooBank" docVersion="1" lastPageNumber="374" masterDocId="FF84FFF90E6EBD1FFFBFFFE78214FF81" masterDocTitle="A new minute riffle beetle (Coleoptera: Elmidae: Pagelmis) from the Brazilian Amazon" masterLastPageNumber="376" masterPageNumber="369" pageNumber="370" updateTime="1732213088795" updateUser="guilherme" zenodo-license-document="CLOSED">
<mods:mods id="C7970978ABABC588B30FD71B1A025CE1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0D08A20FEB745198DD261DB98F5DACDC">
<mods:title id="A5A438B2C6F843F185273582854EE53F">A new minute riffle beetle (Coleoptera: Elmidae: Pagelmis) from the Brazilian Amazon</mods:title>
</mods:titleInfo>
<mods:name id="304E57F84F2543B109EB997BC47475EA" type="personal">
<mods:role id="93146DCE8FC249C2C5E3712D18D8F9F2">
<mods:roleTerm id="29BE392B98108DE3D7B4169B0732ACE8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E734E9E0B146AB0EC81750439F39D235">Polizei, Thiago T. S.</mods:namePart>
<mods:affiliation id="5CABA7EB712174501D5A18B1F01EB3E6">Museu de Zoologia da Universidade de São Paulo (MZSP). Laboratório de Coleoptera (LaC). Av. Nazaré 481, CEP: 04263 - 000, São Paulo, São Paulo State, Brazil.</mods:affiliation>
</mods:name>
<mods:name id="63ED6F586D12D52CF538FA08B198D8BF" type="personal">
<mods:role id="92110A207F7C9777E1C629C4DE344758">
<mods:roleTerm id="A3DE2CD9A39BEC808F9DA9357E132E67">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9090ADBEA8968135DD644D993F2080C2">Hamada, Neusa</mods:namePart>
<mods:affiliation id="0E372202F0F25037318B56F740DA3370">Instituto Nacional de Pesquisas da Amazônia (INPA), Laboratório de Citotaxonomia e Insetos Aquáticos (LACIA). Av. André Araújo 2936, CEP: 69067 - 375, Manaus, Amazonas State, Brazil.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="00D0FF56B7695D6FB2A7230FA69DDB7D">text</mods:typeOfResource>
<mods:relatedItem id="E2ED6F1BE18F972AC4FF520E22919948" type="host">
<mods:titleInfo id="3DA3A9E76B463A0F40A3C53D505EA2D8">
<mods:title id="2F13BA47381422EEFA5DC874D2806318">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="62C853DE11ABDD4C4AE5E53DDBCD1229">
<mods:date id="21936E39282AA8CE717DDC44D7962FDB">2021</mods:date>
<mods:detail id="88DE0C875BE0EDDCDD9B76473B45B4A4" type="pubDate">
<mods:number id="E871AE723977D9B5411D147E9BBC4A3E">2021-11-04</mods:number>
</mods:detail>
<mods:detail id="0E7C5E63CF46657B71F13F74AF1C41DC" type="volume">
<mods:number id="1827D79F77F0F308C770F675E5BF60B0">5061</mods:number>
</mods:detail>
<mods:detail id="91D075AA58CCBF5C0CACE465288CED85" type="issue">
<mods:number id="C99418D663C7B8E0637E279F4ED51F37">2</mods:number>
</mods:detail>
<mods:extent id="357780059553D9576BB5F637303D63F1" unit="page">
<mods:start id="0EC3CEED73467904DCFE6BC1B103AF1A">369</mods:start>
<mods:end id="529D2DCF6C8EF3423B887572914A7E95">376</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3F2063FD79501F0940DE9ABBC92C02B6">journal article</mods:classification>
<mods:identifier id="A7306F65E3FC9DAF34EE3A2CB8EA6103" type="DOI">10.11646/zootaxa.5061.2.9</mods:identifier>
<mods:identifier id="8642491FF475F0EF1A68B92E565B9163" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="80287536CA7E45EC9632F8ADE6CF962D" type="Zenodo-Dep">5649671</mods:identifier>
<mods:identifier id="2B306B9CF5A391D6FF0DC7D1817968A1" type="ZooBank">EE0C41ED-C10C-41FA-B913-73CFD910DADF</mods:identifier>
</mods:mods>
<treatment id="03BD87810E6FBD1AFF28FCAB82C0FC49" LSID="urn:lsid:zoobank.org:act:255E923D-08A8-417B-8D3E-FB8A998397E6" httpUri="http://treatment.plazi.org/id/03BD87810E6FBD1AFF28FCAB82C0FC49" lastPageId="5" lastPageNumber="374" pageId="1" pageNumber="370">
<subSubSection id="C30E651C0E6FBD1EFF28FCAB83BBFCE7" box="[151,431,844,870]" pageId="1" pageNumber="370" type="nomenclature">
<paragraph id="8BAB36970E6FBD1EFF28FCAB83BBFCE7" blockId="1.[151,924,844,939]" box="[151,431,844,870]" pageId="1" pageNumber="370">
<taxonomicName id="4C144D140E6FBD1EFF28FCAB835EFCE7" authority="Polizei &amp; Hamada, 2021" authorityName="Polizei &amp; Hamada" authorityYear="2021" box="[151,330,844,870]" class="Insecta" family="Elmidae" genus="Pagelmis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="mura" status="sp. nov.">
<emphasis id="B960EA850E6FBD1EFF28FCAB835EFCE7" box="[151,330,844,870]" italics="true" pageId="1" pageNumber="370">Pagelmis mura</emphasis>
</taxonomicName>
<emphasis id="B960EA850E6FBD1EFEEEFCAB83BBFCE7" bold="true" box="[337,431,844,870]" pageId="1" pageNumber="370">
<taxonomicNameLabel id="A25357FE0E6FBD1EFEEEFCAB83BBFCE7" box="[337,431,844,870]" pageId="1" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30E651C0E6FBD1EFF28FC888188FC2A" pageId="1" pageNumber="370" type="description">
<paragraph id="8BAB36970E6FBD1EFF28FC888379FC08" blockId="1.[151,924,844,939]" box="[151,365,878,905]" pageId="1" pageNumber="370">
(
<figureCitation id="132F2A120E6FBD1EFF20FC888306FC08" box="[159,274,879,905]" captionStart-0="FIGURES 16" captionStart-1="FIGURES 711" captionStart-2="FIGURES 1214" captionStartId-0="3.[151,264,1944,1968]" captionStartId-1="4.[151,264,1943,1967]" captionStartId-2="5.[151,264,763,787]" captionTargetBox-0="[196,1391,179,1918]" captionTargetBox-1="[247,1334,193,1918]" captionTargetBox-2="[170,1421,196,726]" captionTargetId-0="figure-19@3.[196,1391,179,1918]" captionTargetId-1="figure-25@4.[228,1360,181,1920]" captionTargetId-2="figure-19@5.[151,1437,181,739]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." captionText-1="FIGURES 711. Pagelmis mura sp. nov. Male. Holotype. (7) Sternite VIII. (8) Tergite VIII. (9) Segment IX. 1011 Aedeagus. (10) Ventral view. (11) Dorsal view. Scale bar = 0.1mm." captionText-2="FIGURES 1214. Pattern of the median shiny longitudinal area without plastron on pronotum. 1213 Pagelmis mura sp. nov. (12) Male. (13) Female. (14) Pagelmis amazonica Spangler, 1981, adapted from the original description." figureDoi-0="http://doi.org/10.5281/zenodo.5649673" figureDoi-1="http://doi.org/10.5281/zenodo.5649675" figureDoi-2="http://doi.org/10.5281/zenodo.5649677" httpUri-0="https://zenodo.org/record/5649673/files/figure.png" httpUri-1="https://zenodo.org/record/5649675/files/figure.png" httpUri-2="https://zenodo.org/record/5649677/files/figure.png" pageId="1" pageNumber="370">Figs 113</figureCitation>
,
<figureCitation id="132F2A120E6FBD1EFEA1FC898371FC08" box="[286,357,878,905]" captionStart="FIGURES 1516" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[169,1418,200,1904]" captionTargetId="figure-25@6.[151,1435,181,1919]" captionTargetPageId="6" captionText="FIGURES 1516. (15) Geographical distribution map of Pagelmis (Coleoptera: Elmidae). (16) Stream in Manacapuru municipality, Amazonas state, Brazil, where the holotype of P. mura sp. nov. was collected." figureDoi="http://doi.org/10.5281/zenodo.5649679" httpUri="https://zenodo.org/record/5649679/files/figure.png" pageId="1" pageNumber="370">1516</figureCitation>
)
</paragraph>
<paragraph id="8BAB36970E6FBD1EFF28FC748188FC2A" blockId="1.[151,924,844,939]" box="[151,924,915,939]" pageId="1" pageNumber="370">
<uri id="FF853A950E6FBD1EFF28FC748188FC2A" box="[151,924,915,939]" pageId="1" pageNumber="370">
urn:lsid:zoobank.org:act:
<uuid id="FFB20C420E6FBD1EFE22FC748188FC2A" box="[413,924,915,939]" pageId="1" pageNumber="370">255E923D-08A8-417B-8D3E-FB8A998397E6</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30E651C0E6FBD1EFF28FC0B80DFFB10" pageId="1" pageNumber="370" type="materials_examined">
<paragraph id="8BAB36970E6FBD1EFF28FC0B80DFFB10" blockId="1.[151,1437,1004,2040]" pageId="1" pageNumber="370">
<emphasis id="B960EA850E6FBD1EFF28FC0B8399FB87" bold="true" box="[151,397,1004,1030]" pageId="1" pageNumber="370">Material Examined:</emphasis>
<materialsCitation id="3B7C3CCA0E6FBD1EFE28FC0B8160FBCA" collectingDate="2008-03-30" collectionCode="INPA" collectorName="Jorge L. Nessimian" country="Brazil" elevation="45" latitude="-3.0640278" location="Manacapuru" longLatPrecision="1" longitude="-60.722668" municipality="Manacapuru" pageId="1" pageNumber="370" specimenCount="1" stateProvince="Amazonas" typeStatus="holotype">
<emphasis id="B960EA850E6FBD1EFE28FC0B83EAFB87" box="[407,510,1004,1030]" italics="true" pageId="1" pageNumber="370">
<typeStatus id="54AF88350E6FBD1EFE28FC0B83EAFB87" box="[407,510,1004,1030]" pageId="1" pageNumber="370" type="holotype">Holotype</typeStatus>
</emphasis>
. Male. Pinned, genitalia, abdominal segments VIII and IX mounted on a slide. “
<collectingCountry id="F30376070E6FBD1EFF1EFBE882FFFBA8" box="[161,235,1039,1065]" name="Brazil" pageId="1" pageNumber="370">Brazil</collectingCountry>
.
<collectingRegion id="49D0F8750E6FBD1EFF4AFBE8837BFBA8" box="[245,367,1039,1065]" pageId="1" pageNumber="370">Amazonas</collectingRegion>
.
<collectingMunicipality id="6BCFACED0E6FBD1EFEC6FBE88011FBA8" box="[377,517,1039,1065]" pageId="1" pageNumber="370">Manacapuru</collectingMunicipality>
/ AM 352 - Km 75. Igarapé / pequeno no roçado.
<date id="FFAA10570E6FBD1EFB88FBE886ADFBA8" box="[1079,1209,1039,1065]" pageId="1" pageNumber="370" value="2008-03-30">
<collectingDate id="EFEEE9BF0E6FBD1EFB88FBE886ADFBA8" box="[1079,1209,1039,1065]" pageId="1" pageNumber="370" value="2008-03-30">30.III.2008</collectingDate>
</date>
.
<collectorName id="26E153410E6FBD1EFB7BFBE88788FBA8" box="[1220,1436,1039,1065]" pageId="1" pageNumber="370">Jorge L. Nessimian</collectorName>
<emphasis id="B960EA850E6FBD1EFF28FBD582C5FBCA" box="[151,209,1073,1099]" italics="true" pageId="1" pageNumber="370">et al.</emphasis>
leg.
<geoCoordinate id="EE2050500E6FBD1EFEB5FBD683B0FBCD" box="[266,420,1073,1100]" degrees="03" direction="south" minutes="03" orientation="latitude" pageId="1" pageNumber="370" precision="1" seconds="50.5" value="-3.0640278">03°0350.5”S</geoCoordinate>
<geoCoordinate id="EE2050500E6FBD1EFE13FBD68044FBCA" box="[428,592,1073,1099]" degrees="60" direction="west" minutes="43" orientation="longitude" pageId="1" pageNumber="370" precision="1" seconds="21.6" value="-60.722668">60°4321.6”W</geoCoordinate>
,
<quantity id="4CEC9B720E6FBD1EFDE3FBD68083FBCD" box="[604,663,1073,1100]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="370" unit="m" value="45.0">
<elevation id="0039D1A40E6FBD1EFDE3FBD68083FBCD" box="[604,663,1073,1100]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="370" unit="m" value="45.0">45 m</elevation>
</quantity>
above s.l.” (
<collectionCode id="ED05AE520E6FBD1EFC98FBD68179FBCA" box="[807,877,1073,1099]" pageId="1" pageNumber="370">INPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CCA0E6FBD1EFCC0FBD680D3FB10" collectingDate="2008-05-26" collectionCode="MZSP" collectorName="Carlos A. S. Azevedo" country="Brazil" elevation="75" latitude="-2.9873612" location="Igarape Bolivia" longLatPrecision="1" longitude="-59.941807" municipality="Manaus" pageId="1" pageNumber="370" specimenCode="MZSP 32999" specimenCount="1" stateProvince="Amazonas" typeStatus="allotype">
<emphasis id="B960EA850E6FBD1EFCC0FBD681C8FBCA" box="[895,988,1073,1099]" italics="true" pageId="1" pageNumber="370">
<typeStatus id="54AF88350E6FBD1EFCC0FBD681C8FBCA" box="[895,988,1073,1099]" pageId="1" pageNumber="370" type="allotype">Allotype</typeStatus>
</emphasis>
. Pinned. “
<collectingCountry id="F30376070E6FBD1EFBECFBD68689FBCA" box="[1107,1181,1073,1099]" name="Brazil" pageId="1" pageNumber="370">Brazil</collectingCountry>
.
<collectingRegion id="49D0F8750E6FBD1EFB18FBD58735FBCD" box="[1191,1313,1074,1100]" pageId="1" pageNumber="370">Amazonas</collectingRegion>
.
<collectingMunicipality id="6BCFACED0E6FBD1EFA92FBD5879DFBCD" box="[1325,1417,1074,1100]" pageId="1" pageNumber="370">Manaus</collectingMunicipality>
. / Reserva Ducke AM010 Km 26 /
<location id="8ECB604C0E6FBD1EFDA8FBB380D3FBEF" LSID="urn:lsid:plazi:treatment:03BD87810E6FBD1AFF28FCAB82C0FC49:8ECB604C0E6FBD1EFDA8FBB380D3FBEF" box="[535,711,1108,1134]" country="Brazil" latitude="-2.9873612" longLatPrecision="1" longitude="-59.941807" municipality="Manaus" name="Igarape Bolivia" pageId="1" pageNumber="370" stateProvince="Amazonas">Igarapé Bolívia</location>
.
<date id="FFAA10570E6FBD1EFD6FFBB38156FBEF" box="[720,834,1108,1134]" pageId="1" pageNumber="370" value="2008-05-26">
<collectingDate id="EFEEE9BF0E6FBD1EFD6FFBB38156FBEF" box="[720,834,1108,1134]" pageId="1" pageNumber="370" value="2008-05-26">26.V.2008</collectingDate>
</date>
/
<geoCoordinate id="EE2050500E6FBD1EFCE8FBB381E4FBEE" box="[855,1008,1108,1135]" degrees="02" direction="south" minutes="59" orientation="latitude" pageId="1" pageNumber="370" precision="1" seconds="14.5" value="-2.9873612">02°5914.5”S</geoCoordinate>
<geoCoordinate id="EE2050500E6FBD1EFC49FBB3868EFBEE" box="[1014,1178,1108,1135]" degrees="59" direction="west" minutes="56" orientation="longitude" pageId="1" pageNumber="370" precision="1" seconds="30.5" value="-59.941807">59°5630.5”W</geoCoordinate>
,
<quantity id="4CEC9B720E6FBD1EFB1AFBB386C9FBEE" box="[1189,1245,1108,1135]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="1" pageNumber="370" unit="m" value="75.0">
<elevation id="0039D1A40E6FBD1EFB1AFBB386C9FBEE" box="[1189,1245,1108,1135]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="1" pageNumber="370" unit="m" value="75.0">75 m</elevation>
</quantity>
above s.l.
<collectorName id="26E153410E6FBD1EFAECFBB38326FB10" pageId="1" pageNumber="370">Carlos A.S. Azevedo</collectorName>
<emphasis id="B960EA850E6FBD1EFE86FB908365FB10" box="[313,369,1143,1169]" italics="true" pageId="1" pageNumber="370">et al.</emphasis>
leg. //
<specimenCode id="DBB29EEC0E6FBD1EFE00FB908043FB10" box="[447,599,1143,1169]" pageId="1" pageNumber="370">MZSP 32999</specimenCode>
” (
<collectionCode id="ED05AE520E6FBD1EFDD0FB9080AAFB10" box="[623,702,1143,1169]" pageId="1" pageNumber="370">MZSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E651C0E6FBD1EFF78FB7E81B4FAE3" pageId="1" pageNumber="370" type="diagnosis">
<paragraph id="8BAB36970E6FBD1EFF78FB7E81B4FAE3" blockId="1.[151,1437,1004,2040]" pageId="1" pageNumber="370">
<emphasis id="B960EA850E6FBD1EFF78FB7E8356FB32" bold="true" box="[199,322,1177,1203]" pageId="1" pageNumber="370">Diagnosis.</emphasis>
The new species can be distinguished by having 1) body densely covered by distinct elongate granules; 2) pronotum with median area shiny and longitudinal, without plastron, creating a pattern in the middle third of posterior margin, that is almost straight from the base to posterior 1/3 of pronotum, then strongly narrowed into apical portion that is slightly widened into arrowhead-like shape, reaching slightly at anterior 1/3; 3) elytra with the intervals III without plastron; 4) anterior tibiae with a cleaning fringe; 5) males with an apicoventral tubercle on meso- and metatrochanter; 6) smaller size (
<quantity id="4CEC9B720E6FBD1EFD21FAA08156FAE0" box="[670,834,1351,1377]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" metricValueMax="1.33" metricValueMin="1.31" pageId="1" pageNumber="370" unit="mm" value="1.32" valueMax="1.33" valueMin="1.31">1.311.33 mm</quantity>
length).
</paragraph>
</subSubSection>
<subSubSection id="C30E651C0E6FBD1DFF78FA8E81C7FAD4" lastPageId="2" lastPageNumber="371" pageId="1" pageNumber="370" type="description">
<paragraph id="8BAB36970E6FBD1EFF78FA8E868FFA6D" blockId="1.[151,1437,1004,2040]" pageId="1" pageNumber="370">
<emphasis id="B960EA850E6FBD1EFF78FA8E834FFA02" bold="true" box="[199,347,1385,1411]" pageId="1" pageNumber="370">Description:</emphasis>
<emphasis id="B960EA850E6FBD1EFED8FA8E83B5FA02" box="[359,417,1385,1411]" italics="true" pageId="1" pageNumber="370">Male</emphasis>
. (
<figureCitation id="132F2A120E6FBD1EFE04FA8E8026FA02" box="[443,562,1385,1411]" captionStart-0="FIGURES 16" captionStart-1="FIGURES 711" captionStart-2="FIGURES 1214" captionStartId-0="3.[151,264,1944,1968]" captionStartId-1="4.[151,264,1943,1967]" captionStartId-2="5.[151,264,763,787]" captionTargetBox-0="[196,1391,179,1918]" captionTargetBox-1="[247,1334,193,1918]" captionTargetBox-2="[170,1421,196,726]" captionTargetId-0="figure-19@3.[196,1391,179,1918]" captionTargetId-1="figure-25@4.[228,1360,181,1920]" captionTargetId-2="figure-19@5.[151,1437,181,739]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." captionText-1="FIGURES 711. Pagelmis mura sp. nov. Male. Holotype. (7) Sternite VIII. (8) Tergite VIII. (9) Segment IX. 1011 Aedeagus. (10) Ventral view. (11) Dorsal view. Scale bar = 0.1mm." captionText-2="FIGURES 1214. Pattern of the median shiny longitudinal area without plastron on pronotum. 1213 Pagelmis mura sp. nov. (12) Male. (13) Female. (14) Pagelmis amazonica Spangler, 1981, adapted from the original description." figureDoi-0="http://doi.org/10.5281/zenodo.5649673" figureDoi-1="http://doi.org/10.5281/zenodo.5649675" figureDoi-2="http://doi.org/10.5281/zenodo.5649677" httpUri-0="https://zenodo.org/record/5649673/files/figure.png" httpUri-1="https://zenodo.org/record/5649675/files/figure.png" httpUri-2="https://zenodo.org/record/5649677/files/figure.png" pageId="1" pageNumber="370">Figs 312</figureCitation>
) Body elongate, subparallel, flat. Cuticle reddish-brown, fully covered by opaque plastron except for the parts of head, median longitudinal area of pronotum, a longitudinal band on elytra near the suture, scutellum, and the legs. Surface with dense elongate granules and micropunctures. Total length (anterior margin of the pronotum to elytral apex)
<quantity id="4CEC9B720E6FBD1EFD7FFA368132FA6D" box="[704,806,1489,1516]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="1" pageNumber="370" unit="mm" value="1.31">1.31 mm</quantity>
, greatest width (elytra)
<quantity id="4CEC9B720E6FBD1EFB8FFA368683FA6D" box="[1072,1175,1489,1516]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="370" unit="mm" value="0.48">0.48 mm</quantity>
.
</paragraph>
<paragraph id="8BAB36970E6FBD1EFF78FA1386B2F880" blockId="1.[151,1437,1004,2040]" pageId="1" pageNumber="370">
<emphasis id="B960EA850E6FBD1EFF78FA138310F98F" box="[199,260,1524,1550]" italics="true" pageId="1" pageNumber="370">Head</emphasis>
(
<figureCitation id="132F2A120E6FBD1EFEAEFA138360F98E" box="[273,372,1524,1551]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="1" pageNumber="370">Figs 36</figureCitation>
) partly retractable into prothorax; dorsal surface with micropunctures. Eyes protruding laterally. Antennae filiform, 11-segmented, antennomere VIXI with few, very short thin setae on apex. Frontoclypeal suture straight. Clypeus 4x wider than long. Labrum (
<figureCitation id="132F2A120E6FBD1EFD4DF9DE8120F9D2" box="[754,820,1593,1619]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="1" pageNumber="370">Fig. 3</figureCitation>
) subrectangular; anterolateral angles rounded; anterior margin bi-sinuate. Maxilla: maxillary palpus 4-segmented, palpomere IV subequal in length to the two-preceding ones combined. Labium: gula narrower than submentum and mentum, subrectangular, glabrous, with punctures; submentum and mentum flat, subrectangular, glabrous, with punctures; labial palpus 3-segmented; apical palpomere subequal in length to the two preceding ones combined; ligula weakly sclerotized, almost 2x longer than wide, anterior margin sinuous, anterolateral angles arcuate and raised. Gena covered by tomentum.
</paragraph>
<paragraph id="8BAB36970E6FBD1DFF78F8EE81D7FC94" blockId="1.[151,1437,1004,2040]" lastBlockId="2.[151,1437,151,1978]" lastPageId="2" lastPageNumber="371" pageId="1" pageNumber="370">
<emphasis id="B960EA850E6FBD1EFF78F8EE830CF8A2" box="[199,280,1801,1827]" italics="true" pageId="1" pageNumber="370">Thorax</emphasis>
(
<figureCitation id="132F2A120E6FBD1EFE98F8EE8399F8A2" box="[295,397,1801,1827]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="1" pageNumber="370">Figs 36</figureCitation>
,
<figureCitation id="132F2A120E6FBD1EFE27F8EE83A3F8A2" box="[408,439,1801,1827]" captionStart="FIGURES 1214" captionStartId="5.[151,264,763,787]" captionTargetBox="[170,1421,196,726]" captionTargetId="figure-19@5.[151,1437,181,739]" captionTargetPageId="5" captionText="FIGURES 1214. Pattern of the median shiny longitudinal area without plastron on pronotum. 1213 Pagelmis mura sp. nov. (12) Male. (13) Female. (14) Pagelmis amazonica Spangler, 1981, adapted from the original description." figureDoi="http://doi.org/10.5281/zenodo.5649677" httpUri="https://zenodo.org/record/5649677/files/figure.png" pageId="1" pageNumber="370">12</figureCitation>
). Pronotum (
<figureCitation id="132F2A120E6FBD1EFDF6F8EE80DDF8A5" box="[585,713,1801,1828]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="1" pageNumber="370">Figs 3, 56</figureCitation>
,
<figureCitation id="132F2A120E6FBD1EFD6AF8EE80E6F8A2" box="[725,754,1801,1827]" captionStart="FIGURES 1214" captionStartId="5.[151,264,763,787]" captionTargetBox="[170,1421,196,726]" captionTargetId="figure-19@5.[151,1437,181,739]" captionTargetPageId="5" captionText="FIGURES 1214. Pattern of the median shiny longitudinal area without plastron on pronotum. 1213 Pagelmis mura sp. nov. (12) Male. (13) Female. (14) Pagelmis amazonica Spangler, 1981, adapted from the original description." figureDoi="http://doi.org/10.5281/zenodo.5649677" httpUri="https://zenodo.org/record/5649677/files/figure.png" pageId="1" pageNumber="370">12</figureCitation>
) 1.2x longer than wide; anterior margin strongly arcuate and raised medially, with a distinct carinae not reaching the lateral margins (
<figureCitation id="132F2A120E6FBD1EFC0EF8CB8600F8C6" box="[945,1044,1836,1863]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="1" pageNumber="370">Figs 3, 5</figureCitation>
); sides broadly convex, crenulated; posterior angles slightly acute; posterior margin tri-sinuate, without prescutellar fovea; pronotal surface (
<figureCitation id="132F2A120E6FBD1EFA8DF8A88783F8E8" box="[1330,1431,1870,1897]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="1" pageNumber="370">Figs 3, 5</figureCitation>
,
<figureCitation id="132F2A120E6FBD1EFF28F89682A0F80A" box="[151,180,1905,1931]" captionStart="FIGURES 1214" captionStartId="5.[151,264,763,787]" captionTargetBox="[170,1421,196,726]" captionTargetId="figure-19@5.[151,1437,181,739]" captionTargetPageId="5" captionText="FIGURES 1214. Pattern of the median shiny longitudinal area without plastron on pronotum. 1213 Pagelmis mura sp. nov. (12) Male. (13) Female. (14) Pagelmis amazonica Spangler, 1981, adapted from the original description." figureDoi="http://doi.org/10.5281/zenodo.5649677" httpUri="https://zenodo.org/record/5649677/files/figure.png" pageId="1" pageNumber="370">12</figureCitation>
) with median area shiny and longitudinal, without plastron, creating a pattern in the middle third of posterior margin, that is almost straight from the base to posterior 1/3 of pronotum, then strongly narrowed into apical portion that is slightly widened into arrowhead-like shape, reaching slightly at anterior 1/3; a median longitudinal impression distinct, from base to the posterior half, medially attenuated; integument with row of elongate granules, and micropunctures distanced by the equivalent of one or two times their diameter organized circular like. Scutellum (
<figureCitation id="132F2A120E6CBD1DFF1FFF5C82F6FF54" box="[160,226,187,213]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="2" pageNumber="371">Fig. 3</figureCitation>
) shield-like, glabrous. Elytra (
<figureCitation id="132F2A120E6CBD1DFD87FF5C80ACFF54" box="[568,696,186,213]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="2" pageNumber="371">Figs 3, 56</figureCitation>
) 1.7x longer than wide; wider than pronotum; almost flat dorsally (
<figureCitation id="132F2A120E6CBD1DFF1FFF3882F1FF7B" box="[160,229,223,250]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="2" pageNumber="371">Fig. 6</figureCitation>
); anterior margin smooth, without plastron; subparallel in anterior 2/3, margins slightly crenulated, apices rounded; humeri rounded; elytra with the intervals III without plastron, intervals IIIVII well marked by distinct rows of elongate granules, with micropunctures distanced by the equivalent of one or two times their diameters; first row of granules (between intervals I and II), less visible, and covered by plastron (better visible on female (
<figureCitation id="132F2A120E6CBD1DFF1FFE8882F6FE08" box="[160,226,367,393]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="2" pageNumber="371">Fig. 1</figureCitation>
)). Epipleura narrow, wider anteriorly, covered by plastron and micropunctures (
<figureCitation id="132F2A120E6CBD1DFB3AFE8886E4FE0B" box="[1157,1264,367,394]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="2" pageNumber="371">Figs 4, 6</figureCitation>
). Metathoracic wings absent (apterous). Hypomeron (
<figureCitation id="132F2A120E6CBD1DFDF3FE7480A7FE2F" box="[588,691,403,430]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="2" pageNumber="371">Figs 4, 6</figureCitation>
) broad, 2x longer than wide, with rows of elongate granules and covered by micropunctures. Prosternum 2x wider than long, anterior margin arcuate, disc covered by granules and micropunctures (
<figureCitation id="132F2A120E6CBD1DFEE8FE3C838AFE74" box="[343,414,475,501]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="2" pageNumber="371">Fig. 4</figureCitation>
). Prosternal process narrow, with parallel sides, about 1/3 as wide as long, apex rounded (
<figureCitation id="132F2A120E6CBD1DFF1FFE1882FCFD98" box="[160,232,511,537]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="2" pageNumber="371">Fig. 4</figureCitation>
). Proepimeron and proepisternum covered by plastron. Mesoventrite with mesal cavity for the reception of prosternal process; mesepimeron covered by plastron. Metaventrite longer than mesoventrite (
<figureCitation id="132F2A120E6CBD1DFB51FDC48721FDBC" box="[1262,1333,547,573]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="2" pageNumber="371">Fig. 4</figureCitation>
), mesaly strongly depressed; discrimen inconspicuous, surface with few setae, plastron and micropunctures, sides similar, but granulose. Legs (
<figureCitation id="132F2A120E6CBD1DFE35FD8C83E4FD07" box="[394,496,619,646]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="2" pageNumber="371">Figs 36</figureCitation>
) without plastron. Pro and mesocoxae rounded, metacoxae transverse. Trochanter subtriangular; meso and metatrochanter with an apicoventral tubercle. Femora strongly narrowed basally, 3.5x longer than wide. Tibiae thin, 7x longer than wide, with spines on distal ¾; protibiae with short cleaning fringe on distal half, not reaching the apex (
<figureCitation id="132F2A120E6CBD1DFDF8FD3080A4FD70" box="[583,688,726,753]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="2" pageNumber="371">Figs 3, 5</figureCitation>
). Tarsus elongated, with few thin setae on ventral surface; apical tarsomere shorter than the four preceding ones combined. Claws simple.
</paragraph>
<paragraph id="8BAB36970E6CBD1DFF78FCF88714FC48" blockId="2.[151,1437,151,1978]" pageId="2" pageNumber="371">
<emphasis id="B960EA850E6CBD1DFF78FCF88325FCB8" box="[199,305,799,825]" italics="true" pageId="2" pageNumber="371">Abdomen</emphasis>
(
<figureCitation id="132F2A120E6CBD1DFEFFFCF883D8FCBB" box="[320,460,799,826]" captionStart-0="FIGURES 16" captionStart-1="FIGURES 711" captionStartId-0="3.[151,264,1944,1968]" captionStartId-1="4.[151,264,1943,1967]" captionTargetBox-0="[196,1391,179,1918]" captionTargetBox-1="[247,1334,193,1918]" captionTargetId-0="figure-19@3.[196,1391,179,1918]" captionTargetId-1="figure-25@4.[228,1360,181,1920]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." captionText-1="FIGURES 711. Pagelmis mura sp. nov. Male. Holotype. (7) Sternite VIII. (8) Tergite VIII. (9) Segment IX. 1011 Aedeagus. (10) Ventral view. (11) Dorsal view. Scale bar = 0.1mm." figureDoi-0="http://doi.org/10.5281/zenodo.5649673" figureDoi-1="http://doi.org/10.5281/zenodo.5649675" httpUri-0="https://zenodo.org/record/5649673/files/figure.png" httpUri-1="https://zenodo.org/record/5649675/files/figure.png" pageId="2" pageNumber="371">Figs 4, 611</figureCitation>
) with five ventrites (
<figureCitation id="132F2A120E6CBD1DFD09FCF880EEFCB8" box="[694,762,799,825]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="2" pageNumber="371">Fig. 4</figureCitation>
), covered by plastron, micropunctures and few granules and setae; intercoxal process of ventrite 1 depressed; posterolateral margins not produced. Tergite VIII (
<figureCitation id="132F2A120E6CBD1DFB51FCA48725FCDC" box="[1262,1329,835,861]" captionStart="FIGURES 711" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[247,1334,193,1918]" captionTargetId="figure-25@4.[228,1360,181,1920]" captionTargetPageId="4" captionText="FIGURES 711. Pagelmis mura sp. nov. Male. Holotype. (7) Sternite VIII. (8) Tergite VIII. (9) Segment IX. 1011 Aedeagus. (10) Ventral view. (11) Dorsal view. Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.5649675" httpUri="https://zenodo.org/record/5649675/files/figure.png" pageId="2" pageNumber="371">Fig. 7</figureCitation>
) rounded posteriorly; anterior margin slightly arcuate medially; surface covered by few setae. Sternite VIII (
<figureCitation id="132F2A120E6CBD1DFAAEFC808743FC00" box="[1297,1367,871,897]" captionStart="FIGURES 711" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[247,1334,193,1918]" captionTargetId="figure-25@4.[228,1360,181,1920]" captionTargetPageId="4" captionText="FIGURES 711. Pagelmis mura sp. nov. Male. Holotype. (7) Sternite VIII. (8) Tergite VIII. (9) Segment IX. 1011 Aedeagus. (10) Ventral view. (11) Dorsal view. Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.5649675" httpUri="https://zenodo.org/record/5649675/files/figure.png" pageId="2" pageNumber="371">Fig. 8</figureCitation>
) with anterior apodeme about as long as rest of sternite, posterior margin arcuate, with long setae. Segment IX (
<figureCitation id="132F2A120E6CBD1DFAF0FC6C8780FC24" box="[1359,1428,907,933]" captionStart="FIGURES 711" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[247,1334,193,1918]" captionTargetId="figure-25@4.[228,1360,181,1920]" captionTargetPageId="4" captionText="FIGURES 711. Pagelmis mura sp. nov. Male. Holotype. (7) Sternite VIII. (8) Tergite VIII. (9) Segment IX. 1011 Aedeagus. (10) Ventral view. (11) Dorsal view. Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.5649675" httpUri="https://zenodo.org/record/5649675/files/figure.png" pageId="2" pageNumber="371">Fig. 9</figureCitation>
) forming a subrectangular genital capsule surrounding aedeagus with two long anterior median struts.
</paragraph>
<paragraph id="8BAB36970E6CBD1DFF78FC34832EFB44" blockId="2.[151,1437,151,1978]" pageId="2" pageNumber="371">
<emphasis id="B960EA850E6CBD1DFF78FC34837FFC6C" box="[199,363,979,1005]" italics="true" pageId="2" pageNumber="371">Male genitalia</emphasis>
(
<figureCitation id="132F2A120E6CBD1DFEC7FC3483E2FC6C" box="[376,502,979,1005]" captionStart="FIGURES 711" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[247,1334,193,1918]" captionTargetId="figure-25@4.[228,1360,181,1920]" captionTargetPageId="4" captionText="FIGURES 711. Pagelmis mura sp. nov. Male. Holotype. (7) Sternite VIII. (8) Tergite VIII. (9) Segment IX. 1011 Aedeagus. (10) Ventral view. (11) Dorsal view. Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.5649675" httpUri="https://zenodo.org/record/5649675/files/figure.png" pageId="2" pageNumber="371">Figs 1011</figureCitation>
) symmetrical. Phallobase tubular, approximately 3.7x longer than wide, 1.7x longer than penis. Parameres fused, narrowed towards the apex, wider at the base, approximately 5x longer than wide, apex slightly divergent; surface with sensorial pores on posterior ¾ and tufts of setae on apical 1/3; in ventral view inner margin arcuate (
<figureCitation id="132F2A120E6CBD1DFEEEFBD883B2FBD8" box="[337,422,1087,1113]" captionStart="FIGURES 711" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[247,1334,193,1918]" captionTargetId="figure-25@4.[228,1360,181,1920]" captionTargetPageId="4" captionText="FIGURES 711. Pagelmis mura sp. nov. Male. Holotype. (7) Sternite VIII. (8) Tergite VIII. (9) Segment IX. 1011 Aedeagus. (10) Ventral view. (11) Dorsal view. Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.5649675" httpUri="https://zenodo.org/record/5649675/files/figure.png" pageId="2" pageNumber="371">Fig. 10</figureCitation>
), in dorsal view straight (
<figureCitation id="132F2A120E6CBD1DFD71FBD88135FBD8" box="[718,801,1087,1113]" captionStart="FIGURES 711" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[247,1334,193,1918]" captionTargetId="figure-25@4.[228,1360,181,1920]" captionTargetPageId="4" captionText="FIGURES 711. Pagelmis mura sp. nov. Male. Holotype. (7) Sternite VIII. (8) Tergite VIII. (9) Segment IX. 1011 Aedeagus. (10) Ventral view. (11) Dorsal view. Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.5649675" httpUri="https://zenodo.org/record/5649675/files/figure.png" pageId="2" pageNumber="371">Fig. 11</figureCitation>
). Penis shorter than parameres, approximately reaching ¾ of the length of the parameres, 3x longer than wide at the base, constricted in the basal and apical 1/3, lateral margins sinuous, rounded apex, three basolateral apophyses; fibula narrow, approximately as long as 1/3 of penis; corona absent.
</paragraph>
<paragraph id="8BAB36970E6CBD1DFF78FB2881C7FAD4" blockId="2.[151,1437,151,1978]" pageId="2" pageNumber="371">
<emphasis id="B960EA850E6CBD1DFF78FB28830FFB68" box="[199,283,1231,1257]" italics="true" pageId="2" pageNumber="371">Female</emphasis>
. (
<figureCitation id="132F2A120E6CBD1DFE8CFB28838DFB68" box="[307,409,1231,1257]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="2" pageNumber="371">Figs 12</figureCitation>
,
<figureCitation id="132F2A120E6CBD1DFE18FB2883D7FB68" box="[423,451,1231,1257]" captionStart="FIGURES 1214" captionStartId="5.[151,264,763,787]" captionTargetBox="[170,1421,196,726]" captionTargetId="figure-19@5.[151,1437,181,739]" captionTargetPageId="5" captionText="FIGURES 1214. Pattern of the median shiny longitudinal area without plastron on pronotum. 1213 Pagelmis mura sp. nov. (12) Male. (13) Female. (14) Pagelmis amazonica Spangler, 1981, adapted from the original description." figureDoi="http://doi.org/10.5281/zenodo.5649677" httpUri="https://zenodo.org/record/5649677/files/figure.png" pageId="2" pageNumber="371">13</figureCitation>
)
<quantity id="4CEC9B720E6CBD1DFE6AFB28802FFB68" box="[469,571,1231,1257]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="2" pageNumber="371" unit="mm" value="1.33">1.33 mm</quantity>
long, greatest width (elytra)
<quantity id="4CEC9B720E6CBD1DFC36FB2981F0FB68" box="[905,996,1230,1257]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="371" unit="mm" value="0.5">0.5 mm</quantity>
. External morphology similar to male, except for absence of deep depression on metaventrite and on intercoxal process of abdominal ventrite 1, and the lack of the apicoventral tubercle on meso- and metatrochanter. On the female specimen, shiny pronotal median area reaching from base to middle, not strongly narrowed on apex (
<figureCitation id="132F2A120E6CBD1DFCEDFADC818DFAD4" box="[850,921,1339,1365]" captionStart="FIGURES 16" captionStartId="3.[151,264,1944,1968]" captionTargetBox="[196,1391,179,1918]" captionTargetId="figure-19@3.[196,1391,179,1918]" captionTargetPageId="3" captionText="FIGURES 16. Pagelmis mura sp. nov. Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="2" pageNumber="371">Figs 1</figureCitation>
,
<figureCitation id="132F2A120E6CBD1DFC18FADC81D2FAD4" box="[935,966,1339,1365]" captionStart="FIGURES 1214" captionStartId="5.[151,264,763,787]" captionTargetBox="[170,1421,196,726]" captionTargetId="figure-19@5.[151,1437,181,739]" captionTargetPageId="5" captionText="FIGURES 1214. Pattern of the median shiny longitudinal area without plastron on pronotum. 1213 Pagelmis mura sp. nov. (12) Male. (13) Female. (14) Pagelmis amazonica Spangler, 1981, adapted from the original description." figureDoi="http://doi.org/10.5281/zenodo.5649677" httpUri="https://zenodo.org/record/5649677/files/figure.png" pageId="2" pageNumber="371">13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E651C0E6CBD1DFF78FAB88781FA40" pageId="2" pageNumber="371" type="etymology">
<paragraph id="8BAB36970E6CBD1DFF78FAB88781FA40" blockId="2.[151,1437,151,1978]" pageId="2" pageNumber="371">
<emphasis id="B960EA850E6CBD1DFF78FAB8835AFAF8" bold="true" box="[199,334,1375,1401]" pageId="2" pageNumber="371">Etymology.</emphasis>
The specific epithet,
<taxonomicName id="4C144D140E6CBD1DFDF4FAB88092FAF9" authorityName="Polizei &amp; Hamada" authorityYear="2021" box="[587,646,1375,1400]" class="Insecta" family="Elmidae" genus="Pagelmis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="mura">
<emphasis id="B960EA850E6CBD1DFDF4FAB88092FAF9" box="[587,646,1375,1400]" italics="true" pageId="2" pageNumber="371">mura</emphasis>
</taxonomicName>
is in honor of the Mura ethnic group that inhabits the hydrographic basins of the Madeira,
<collectingRegion id="49D0F8750E6CBD1DFE2EFA648013FA1C" box="[401,519,1411,1437]" country="Venezuela" name="Amazonas" pageId="2" pageNumber="371">Amazonas</collectingRegion>
and Purus Rivers, in the
<collectingRegion id="49D0F8750E6CBD1DFCA6FA6481D8FA1C" box="[793,972,1411,1437]" country="Venezuela" name="Amazonas" pageId="2" pageNumber="371">Amazonas state</collectingRegion>
. The Mura were known for their excellent knowledge of navigation in the different aquatic systems of the Amazon region. The name is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C30E651C0E6CBD1DFF78FA2D8700F960" pageId="2" pageNumber="371" type="biology_ecology">
<paragraph id="8BAB36970E6CBD1DFF78FA2D8700F960" blockId="2.[151,1437,151,1978]" pageId="2" pageNumber="371">
<emphasis id="B960EA850E6CBD1DFF78FA2D833EFA65" bold="true" box="[199,298,1482,1508]" pageId="2" pageNumber="371">Habitat.</emphasis>
Both specimens were found in typical slow-flowing Amazon streams, with black water and sandy streambed, with patches of leaf litter and roots of riparian vegetation (
<figureCitation id="132F2A120E6CBD1DFC79FA088609F98B" box="[966,1053,1519,1546]" captionStart="FIGURES 1516" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[169,1418,200,1904]" captionTargetId="figure-25@6.[151,1435,181,1919]" captionTargetPageId="6" captionText="FIGURES 1516. (15) Geographical distribution map of Pagelmis (Coleoptera: Elmidae). (16) Stream in Manacapuru municipality, Amazonas state, Brazil, where the holotype of P. mura sp. nov. was collected." figureDoi="http://doi.org/10.5281/zenodo.5649679" httpUri="https://zenodo.org/record/5649679/files/figure.png" pageId="2" pageNumber="371">Fig. 16</figureCitation>
). The
<typeStatus id="54AF88350E6CBD1DFBD6FA0886DFF988" box="[1129,1227,1519,1545]" pageId="2" pageNumber="371" type="holotype">holotype</typeStatus>
locality is a firstorder stream (
<emphasis id="B960EA850E6CBD1DFE8AF9F48367F9AD" box="[309,371,1555,1580]" italics="true" pageId="2" pageNumber="371">sensu</emphasis>
<bibRefCitation id="EF854B660E6CBD1DFEC4F9F48003F9AC" author="Strahler, A. N." box="[379,535,1554,1581]" pageId="2" pageNumber="371" pagination="913 - 920" refId="ref4545" refString="Strahler, A. N. (1957) Quantitative analysis of watershed geomorphology. Eos, Transactions American Geophysical Union, 38 (6), 913 - 920. https: // doi. org / 10.1029 / TR 038 i 006 p 00913" type="journal article" year="1957">Strahler 1957</bibRefCitation>
), situated in the municipality of Manacapuru, Amazonas State, in a private rural property. The stream had a mean width of
<quantity id="4CEC9B720E6CBD1DFDD0F9D080A5F9D0" box="[623,689,1591,1617]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="371" unit="m" value="1.2">
<elevation id="0039D1A40E6CBD1DFDD0F9D080A5F9D0" box="[623,689,1591,1617]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="371" unit="m" value="1.2">1.2 m</elevation>
</quantity>
, a mean depth of
<quantity id="4CEC9B720E6CBD1DFCC9F9D081A9F9D0" box="[886,957,1591,1617]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="371" unit="cm" value="11.0">11 cm</quantity>
; its water temperature was 26.5°C, pH 4.6, and electrical conductivity 21.9 µS/cm. The
<typeStatus id="54AF88350E6CBD1DFD2CF9BC80F9F9F4" box="[659,749,1627,1653]" pageId="2" pageNumber="371" type="allotype">allotype</typeStatus>
locality,
<collectingCountry id="F30376070E6CBD1DFCE7F9BC81BEF9F4" box="[856,938,1627,1653]" name="Bolivia" pageId="2" pageNumber="371">Bolivia</collectingCountry>
stream, is a second-order stream located in the
<location id="8ECB604C0E6CBD1DFF7FF99883C2F918" LSID="urn:lsid:plazi:treatment:03BD87810E6FBD1AFF28FCAB82C0FC49:8ECB604C0E6CBD1DFF7FF99883C2F918" box="[192,470,1663,1689]" country="Bolivia" county="Manaus" municipality="Instituto Nacional de Pesquisas da Amazonia" name="Reserva Florestal Ducke" pageId="2" pageNumber="371">Reserva Florestal Ducke</location>
, a biological reserve that belongs to the
<emphasis id="B960EA850E6CBD1DFC27F9988781F918" box="[920,1429,1663,1689]" italics="true" pageId="2" pageNumber="371">
<collectingMunicipality id="6BCFACED0E6CBD1DFC27F9988781F918" box="[920,1429,1663,1689]" pageId="2" pageNumber="371">Instituto Nacional de Pesquisas da Amazônia</collectingMunicipality>
</emphasis>
, located adjacent to the city of
<collectingCounty id="62CA4E1B0E6CBD1DFE4BF944805BF93C" box="[500,591,1699,1725]" pageId="2" pageNumber="371">Manaus</collectingCounty>
, Amazonas. This stream was characterized by having mean width of
<quantity id="4CEC9B720E6CBD1DFAD6F9448783F93C" box="[1385,1431,1699,1725]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="371" unit="m" value="2.0">
<elevation id="0039D1A40E6CBD1DFAD6F9448783F93C" box="[1385,1431,1699,1725]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="371" unit="m" value="2.0">2 m</elevation>
</quantity>
, mean depth of
<quantity id="4CEC9B720E6CBD1DFE81F92183BDF960" box="[318,425,1734,1761]" metricMagnitude="-1" metricUnit="m" metricValue="2.516" pageId="2" pageNumber="371" unit="cm" value="25.16">25.16 cm</quantity>
, water temperature 24.7°C; pH 4.92, and electrical conductivity 13.72 µS/cm.
</paragraph>
</subSubSection>
<subSubSection id="C30E651C0E6CBD1DFF78F90D808CF884" box="[199,664,1770,1797]" pageId="2" pageNumber="371" type="distribution">
<paragraph id="8BAB36970E6CBD1DFF78F90D808CF884" blockId="2.[151,1437,151,1978]" box="[199,664,1770,1797]" pageId="2" pageNumber="371">
<emphasis id="B960EA850E6CBD1DFF78F90D8377F885" bold="true" box="[199,355,1770,1796]" pageId="2" pageNumber="371">Distribution:</emphasis>
<collectingCountry id="F30376070E6CBD1DFED5F90C83C7F884" box="[362,467,1771,1797]" name="Brazil" pageId="2" pageNumber="371">BRAZIL</collectingCountry>
. Amazonas State.
</paragraph>
</subSubSection>
<subSubSection id="C30E651C0E6CBD1DFF78F8E98102F838" pageId="2" pageNumber="371" type="discussion">
<paragraph id="8BAB36970E6CBD1DFF78F8E98102F838" blockId="2.[151,1437,151,1978]" pageId="2" pageNumber="371">
<emphasis id="B960EA850E6CBD1DFF78F8E983A6F8A8" bold="true" box="[199,434,1806,1833]" pageId="2" pageNumber="371">Comparative notes.</emphasis>
<taxonomicName id="4C144D140E6CBD1DFE04F8E88073F8A9" authority="Polizei &amp; Hamada, 2021" authorityName="Polizei &amp; Hamada" authorityYear="2021" box="[443,615,1807,1833]" class="Insecta" family="Elmidae" genus="Pagelmis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="mura" status="sp. nov.">
<emphasis id="B960EA850E6CBD1DFE04F8E88073F8A9" box="[443,615,1807,1833]" italics="true" pageId="2" pageNumber="371">Pagelmis mura</emphasis>
</taxonomicName>
<emphasis id="B960EA850E6CBD1DFDD0F8E880DFF8A8" bold="true" box="[623,715,1807,1833]" pageId="2" pageNumber="371">
<taxonomicNameLabel id="A25357FE0E6CBD1DFDD0F8E880DFF8A8" box="[623,715,1807,1833]" pageId="2" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
share to
<taxonomicName id="4C144D140E6CBD1DFC89F8E881D8F8A9" authorityName="Spangler" authorityYear="1981" box="[822,972,1807,1832]" class="Insecta" family="Elmidae" genus="Pagelmis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="amazonica">
<emphasis id="B960EA850E6CBD1DFC89F8E881D8F8A9" box="[822,972,1807,1832]" italics="true" pageId="2" pageNumber="371">P. amazonica</emphasis>
</taxonomicName>
the structure of male genitalia, in which parameres are longer than penis. However, in
<taxonomicName id="4C144D140E6CBD1DFD27F8D48139F8CD" authorityName="Spangler" authorityYear="1981" box="[664,813,1843,1868]" class="Insecta" family="Elmidae" genus="Pagelmis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="amazonica">
<emphasis id="B960EA850E6CBD1DFD27F8D48139F8CD" box="[664,813,1843,1868]" italics="true" pageId="2" pageNumber="371">P. amazonica</emphasis>
</taxonomicName>
the body is not covered by distinct elongate granulates; pronotum has the shiny, longitudinal, median area without plastron arcuate into apical portion; tibiae without cleaning fringes; metathoracic wings present; males with an apicoventral tubercle only on metatrochanter (very short in females) and the specimens are bigger (
<quantity id="4CEC9B720E6CBD1DFD11F878811EF838" box="[686,778,1951,1977]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="371" unit="mm" value="2.2">2.2 mm</quantity>
).
</paragraph>
</subSubSection>
<caption id="DF6B661F0E6DBD1CFF28F87F8659F855" ID-DOI="http://doi.org/10.5281/zenodo.5649673" ID-Zenodo-Dep="5649673" httpUri="https://zenodo.org/record/5649673/files/figure.png" pageId="3" pageNumber="372" startId="3.[151,264,1944,1968]" targetBox="[196,1391,179,1918]" targetPageId="3">
<paragraph id="8BAB36970E6DBD1CFF28F87F8659F855" blockId="3.[151,1436,1944,2005]" pageId="3" pageNumber="372">
<emphasis id="B960EA850E6DBD1CFF28F87F832FF831" bold="true" box="[151,315,1944,1968]" pageId="3" pageNumber="372">FIGURES 16.</emphasis>
<taxonomicName id="4C144D140E6DBD1CFEFCF87F83CAF831" authority="Polizei &amp; Hamada, 2021" authorityName="Polizei &amp; Hamada" authorityYear="2021" box="[323,478,1944,1968]" class="Insecta" family="Elmidae" genus="Pagelmis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="mura" status="sp. nov.">
<emphasis id="B960EA850E6DBD1CFEFCF87F83CAF831" box="[323,478,1944,1968]" italics="true" pageId="3" pageNumber="372">Pagelmis mura</emphasis>
</taxonomicName>
<emphasis id="B960EA850E6DBD1CFE5AF87F8023F831" bold="true" box="[485,567,1944,1968]" pageId="3" pageNumber="372">
<taxonomicNameLabel id="A25357FE0E6DBD1CFE5AF87F8023F831" box="[485,567,1944,1968]" pageId="3" pageNumber="372" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Habitus. 12. Female. Allotype. (1) Dorsal view. (2) Ventral view. 36. Male. Holotype. (3) Dorsal view. (4) Ventral view. (5) Frontal view. (6) Lateral view. Scale bar = 0.5mm.
</paragraph>
</caption>
<caption id="DF6B661F0E6ABD1BFF28F87080C4F855" ID-DOI="http://doi.org/10.5281/zenodo.5649675" ID-Zenodo-Dep="5649675" httpUri="https://zenodo.org/record/5649675/files/figure.png" pageId="4" pageNumber="373" startId="4.[151,264,1943,1967]" targetBox="[247,1334,193,1918]" targetPageId="4">
<paragraph id="8BAB36970E6ABD1BFF28F87080C4F855" blockId="4.[151,1437,1943,2004]" pageId="4" pageNumber="373">
<emphasis id="B960EA850E6ABD1BFF28F8708352F82E" bold="true" box="[151,326,1943,1967]" pageId="4" pageNumber="373">FIGURES 711.</emphasis>
<taxonomicName id="4C144D140E6ABD1BFEF3F87F83F1F82E" authority="Polizei &amp; Hamada, 2021" authorityName="Polizei &amp; Hamada" authorityYear="2021" box="[332,485,1944,1968]" class="Insecta" family="Elmidae" genus="Pagelmis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="mura" status="sp. nov.">
<emphasis id="B960EA850E6ABD1BFEF3F87F83F1F82E" box="[332,485,1944,1968]" italics="true" pageId="4" pageNumber="373">Pagelmis mura</emphasis>
</taxonomicName>
<emphasis id="B960EA850E6ABD1BFE53F87F8028F831" bold="true" box="[492,572,1944,1968]" pageId="4" pageNumber="373">
<taxonomicNameLabel id="A25357FE0E6ABD1BFE53F87F8028F831" box="[492,572,1944,1968]" pageId="4" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male. Holotype. (7) Sternite VIII. (8) Tergite VIII. (9) Segment IX. 10
<emphasis id="B960EA850E6ABD1BFAB7F87F8701F831" bold="true" box="[1288,1301,1944,1968]" pageId="4" pageNumber="373"></emphasis>
11 Aedeagus. (10) Ventral view. (11) Dorsal view. Scale bar = 0.1mm.
</paragraph>
</caption>
<caption id="DF6B661F0E6BBD1AFF28FD1C86AAFCB6" ID-DOI="http://doi.org/10.5281/zenodo.5649677" ID-Zenodo-Dep="5649677" httpUri="https://zenodo.org/record/5649677/files/figure.png" pageId="5" pageNumber="374" startId="5.[151,264,763,787]" targetBox="[170,1421,196,726]" targetPageId="5">
<paragraph id="8BAB36970E6BBD1AFF28FD1C86AAFCB6" blockId="5.[151,1436,763,824]" pageId="5" pageNumber="374">
<emphasis id="B960EA850E6BBD1AFF28FD1C8340FC92" bold="true" box="[151,340,763,787]" pageId="5" pageNumber="374">FIGURES 1214.</emphasis>
Pattern of the median shiny longitudinal area without plastron on pronotum. 12
<emphasis id="B960EA850E6BBD1AFBC1FD1B869FFC95" bold="true" box="[1150,1163,764,788]" pageId="5" pageNumber="374"></emphasis>
13
<taxonomicName id="4C144D140E6BBD1AFB15FD1C8750FC92" authority="Polizei &amp; Hamada, 2021" authorityName="Polizei &amp; Hamada" authorityYear="2021" box="[1194,1348,763,787]" class="Insecta" family="Elmidae" genus="Pagelmis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="mura" status="sp. nov.">
<emphasis id="B960EA850E6BBD1AFB15FD1C8750FC92" box="[1194,1348,763,787]" italics="true" pageId="5" pageNumber="374">Pagelmis mura</emphasis>
</taxonomicName>
<emphasis id="B960EA850E6BBD1AFAF4FD1B8788FC95" bold="true" box="[1355,1436,764,788]" pageId="5" pageNumber="374">
<taxonomicNameLabel id="A25357FE0E6BBD1AFAF4FD1B8788FC95" box="[1355,1436,764,788]" pageId="5" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(12) Male. (13) Female. (14)
<taxonomicName id="4C144D140E6BBD1AFE7EFCF88120FCB9" authority="Spangler, 1981" authorityName="Spangler" authorityYear="1981" box="[449,820,799,824]" class="Insecta" family="Elmidae" genus="Pagelmis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="amazonica">
<emphasis id="B960EA850E6BBD1AFE7EFCF88087FCB6" box="[449,659,799,823]" italics="true" pageId="5" pageNumber="374">Pagelmis amazonica</emphasis>
<bibRefCitation id="EF854B660E6BBD1AFD26FCF88120FCB9" author="Spangler, P. J." box="[665,820,799,824]" pageId="5" pageNumber="374" pagination="286 - 294" refId="ref4445" refString="Spangler, P. J. (1981) Pagelmis amazonica, a new genus and species of water beetle from Ecuador (Coleoptera: Elmidae). Pan- Pac Entomologist, 57, 286 - 294." type="journal article" year="1981">Spangler, 1981</bibRefCitation>
</taxonomicName>
, adapted from the original description.
</paragraph>
</caption>
<subSubSection id="C30E651C0E6BBD1AFF78FC8182C0FC49" pageId="5" pageNumber="374" type="materials_examined">
<paragraph id="8BAB36970E6BBD1AFF78FC8182C0FC49" blockId="5.[151,1436,869,968]" pageId="5" pageNumber="374">
<emphasis id="B960EA850E6BBD1AFF78FC8183A7FCFE" bold="true" box="[199,435,869,896]" pageId="5" pageNumber="374">
<typeStatus id="54AF88350E6BBD1AFF78FC818320FC01" box="[199,308,870,896]" pageId="5" pageNumber="374" type="holotype">Holotype</typeStatus>
condition.
</emphasis>
It is slightly damaged. Missing parts include tarsomeres IV-V and the claws of the right foreleg and tarsomeres IV-V on the right hind leg. Genitalia, abdominal segments VIII and IX are preserved on a slide.
</paragraph>
</subSubSection>
</treatment>
</document>