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<document id="D8BED8DC3EB779D70C9268856ED24667" ID-CLB-Dataset="29895" ID-DOI="10.11646/zootaxa.4434.2.5" ID-GBIF-Dataset="1445ab9f-f210-4f20-9163-ee875a213214" ID-ISSN="1175-5326" ID-Zenodo-Dep="1290584" ID-ZooBank="D1264EF3-FBC1-4F5D-8A3A-545BE4BE924D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1529062106843" checkinUser="plazi" docAuthor="Deepak, V., Khandekar, Akshay, Chaitanya, R. &amp; Karanth, Praveen" docDate="2018" docId="03BC7748FFBACC51A1B1FF4DFB6AFEF4" docLanguage="en" docName="zootaxa.4434.2.5.pdf.imd" docOrigin="Zootaxa 4434 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sitana gokakensis Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth 2018, sp. nov." docType="treatment" docUuid="975BBC83-D5D2-4F41-9551-2454463DABE0" docUuidSource="ZooBank" docVersion="8" lastPageNumber="343" masterDocId="FF850F30FFB3CC41A126FF93FFCFFFE8" masterDocTitle="Descriptions of two new endemic and cryptic species of Sitana Cuvier, 1829 from peninsular India" masterLastPageNumber="365" masterPageNumber="327" pageNumber="336" updateTime="1698549085721" updateUser="ExternalLinkService">
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<mods:title id="D813DA8A8E2E6879EBA34D6885C9C093">Descriptions of two new endemic and cryptic species of Sitana Cuvier, 1829 from peninsular India</mods:title>
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<mods:namePart id="608C387203BF3467F15BBBB4F073B283">Chaitanya, R.</mods:namePart>
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<mods:namePart id="76B7539F8F63EDDC7ED1B5E4B6859189">Karanth, Praveen</mods:namePart>
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<treatment id="03BC7748FFBACC51A1B1FF4DFB6AFEF4" ID-DOI="http://doi.org/10.5281/zenodo.5958922" ID-GBIF-Taxon="145953172" ID-Zenodo-Dep="5958922" LSID="urn:lsid:zoobank.org:act:975BBC83-D5D2-4F41-9551-2454463DABE0" httpUri="http://treatment.plazi.org/id/03BC7748FFBACC51A1B1FF4DFB6AFEF4" lastPageId="16" lastPageNumber="343" pageId="9" pageNumber="336">
<subSubSection id="C30F95D5FFBACC48A1B1FF4DFD45FEF1" pageId="9" pageNumber="336" type="nomenclature">
<paragraph id="8BAAC65EFFBACC48A1B1FF4DFE19FF10" blockId="9.[151,650,222,282]" box="[151,470,222,248]" pageId="9" pageNumber="336">
<heading id="D0E27132FFBACC48A1B1FF4DFE19FF10" bold="true" box="[151,470,222,248]" fontSize="11" level="1" pageId="9" pageNumber="336" reason="1">
<emphasis id="B9611A4CFFBACC48A1B1FF4DFE19FF10" bold="true" box="[151,470,222,248]" pageId="9" pageNumber="336">
<taxonomicName id="4C15BDDDFFBACC48A1B1FF4DFEBEFF10" authority="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth, 2018" authorityName="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth" authorityYear="2018" box="[151,369,222,248]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="species" species="gokakensis" status="sp. nov.">
<emphasis id="B9611A4CFFBACC48A1B1FF4DFEBEFF10" bold="true" box="[151,369,222,248]" italics="true" pageId="9" pageNumber="336">Sitana gokakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A252A737FFBACC48A05FFF4DFE19FF10" box="[377,470,222,248]" pageId="9" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAAC65EFFBACC48A1B1FE92FD45FEF1" blockId="9.[151,650,222,282]" box="[151,650,257,282]" pageId="9" pageNumber="336">
<figureCitation id="132EDADBFFBACC48A1B1FE92FF3BFEF1" box="[151,244,257,281]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStartId-0="9.[151,249,1949,1971]" captionStartId-1="10.[151,249,1967,1989]" captionStartId-2="11.[151,249,1976,1998]" captionTargetBox-0="[175,1395,1035,1917]" captionTargetBox-1="[199,1381,226,1914]" captionTargetBox-2="[193,1398,214,1932]" captionTargetId-0="figure@9.[151,1436,1018,1927]" captionTargetId-1="figure@10.[176,1418,193,1949]" captionTargetId-2="figure@11.[170,1417,193,1955]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="FIGURE 7. Holotype of Sitana gokakensis sp. nov. (BNHS 2490) from Gokak, Karnataka. (A) full body dorsal (B) flank region (C) dorsal view of head (D) lateral view of head. Scale bar:10 mm." captionText-1="FIGURE 8. Sitana gokakensis sp. nov. colouration in life. A. Holotype (BNHS 2490) in life, B. another adult male collected during 2016 from the type locality." captionText-2="FIGURE 9. Sitana gokakensis sp. nov. colouration in life. A. An adult male from type locality, B. paratype BNHS 2491, C &amp; D. two different uncollected adult females from the type locality." httpUri-0="https://zenodo.org/record/1290598/files/figure.png" httpUri-1="https://zenodo.org/record/1290600/files/figure.png" httpUri-2="https://zenodo.org/record/1290602/files/figure.png" pageId="9" pageNumber="336">Fig. 79</figureCitation>
&amp;
<figureCitation id="132EDADBFFBACC48A03EFE92FEFAFEF1" box="[280,309,257,281]" captionStart="FIGURE 11" captionStartId="14.[151,249,1874,1896]" captionTargetBox="[151,1436,409,1860]" captionTargetId="figure@14.[151,1436,409,1860]" captionTargetPageId="14" captionText="FIGURE 11. Hemipenis of Sitana gokakensis sp. nov. (CES 141136): (A) sulcal view; (B) asulcal view; C) lateral view; (D) apical view. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290606/files/figure.png" pageId="9" pageNumber="336">11</figureCitation>
;
<tableCitation id="C697F3E5FFBACC48A064FE92FE5BFEF1" box="[322,404,257,282]" captionStart="TABLE 3" captionStartId="23.[151,239,152,175]" captionText="TABLE 3. Range of select scale counts of the two new Sitana species including the type series, sample size in brackets. Abbreviations, “_” = NA. 0 = absent. Lam4 = lamellae on fourth toe; SL = supralabials; IL = infralabial; MBS = mid body scales; ENL = enlarged scales on the flank region; ESD = enlarged scale rows in dewlap; VEN = number of ventral scales, VEND= number of ventral scales on dewlap; VENB= number of ventral scales on belly; VS = vertebral scales" pageId="9" pageNumber="336">Table 3</tableCitation>
&amp;
<tableCitation id="C697F3E5FFBACC48A09EFE92FE06FEF1" box="[440,457,257,281]" captionStart="TABLE 4" captionStartId="24.[147,229,487,508]" captionText="TABLE 4. Meristic anđ morphometric đata of 25 inđiviđuals of the two new species useđ in the multivariate analyses. Note: Damageđ tails were not measuređ anđ TL was not" pageId="9" pageNumber="336">4</tableCitation>
; Appendix 7 &amp; 8
</paragraph>
</subSubSection>
<subSubSection id="C30F95D5FFBACC48A1B1FEDAFE99FE19" pageId="9" pageNumber="336" type="materials_examined">
<paragraph id="8BAAC65EFFBACC48A1B1FEDAFAD3FE6E" blockId="9.[151,1436,329,966]" pageId="9" pageNumber="336">
<materialsCitation id="3B7DCC03FFBACC48A1B1FEDAFAD7FE6E" ID-GBIF-Occurrence="1849038459" collectionCode="adult, male, from, Gokak, plateau, district, collected, and, Aparna, Lajmi" collectorName="V. Deepak" country="India" elevation="255" latitude="16.18618" location="Karnataka" longLatPrecision="1" longitude="74.75952" pageId="9" pageNumber="336" specimenCode="BNHS 2490" specimenCount="1" stateProvince="Karnataka" typeStatus="holotype">
<emphasis id="B9611A4CFFBACC48A1B1FEDAFEC4FE8A" bold="true" box="[151,267,329,354]" pageId="9" pageNumber="336">
<typeStatus id="54AE78FCFFBACC48A1B1FEDAFEC9FE8A" box="[151,262,329,354]" pageId="9" pageNumber="336" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBB36E25FFBACC48A03FFEDAFE63FE89" box="[281,428,329,353]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="9" pageNumber="336">BNHS 2490</specimenCode>
,
<collectionCode id="ED045E9BFFBACC48A098FEDAFDF5FE89" box="[446,570,329,353]" pageId="9" pageNumber="336">adult male</collectionCode>
(
<figureCitation id="132EDADBFFBACC48A369FEDAFD56FE89" box="[591,665,329,353]" captionStart="FIGURE 7" captionStartId="9.[151,249,1949,1971]" captionTargetBox="[175,1395,1035,1917]" captionTargetId="figure@9.[151,1436,1018,1927]" captionTargetPageId="9" captionText="FIGURE 7. Holotype of Sitana gokakensis sp. nov. (BNHS 2490) from Gokak, Karnataka. (A) full body dorsal (B) flank region (C) dorsal view of head (D) lateral view of head. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290598/files/figure.png" pageId="9" pageNumber="336">Fig. 7</figureCitation>
&amp;
<figureCitation id="132EDADBFFBACC48A3EDFEDAFD21FE89" box="[715,750,329,353]" captionStart="FIGURE 8" captionStartId="10.[151,249,1967,1989]" captionTargetBox="[199,1381,226,1914]" captionTargetId="figure@10.[176,1418,193,1949]" captionTargetPageId="10" captionText="FIGURE 8. Sitana gokakensis sp. nov. colouration in life. A. Holotype (BNHS 2490) in life, B. another adult male collected during 2016 from the type locality." httpUri="https://zenodo.org/record/1290600/files/figure.png" pageId="9" pageNumber="336">8A</figureCitation>
)
<collectionCode id="ED045E9BFFBACC48A222FEDAFC39FE89" box="[772,1014,329,353]" pageId="9" pageNumber="336">from Gokak plateau</collectionCode>
, Belagavi
<collectionCode id="ED045E9BFFBACC48A55EFEDAFB05FE89" box="[1144,1226,329,353]" pageId="9" pageNumber="336">district</collectionCode>
,
<collectingRegion id="49D108BCFFBACC48A5FDFEDAFA9DFE89" box="[1243,1362,329,353]" country="India" name="Karnataka" pageId="9" pageNumber="336">Karnataka</collectingRegion>
,
<collectingCountry id="F30286CEFFBACC48A444FEDAFA53FE89" box="[1378,1436,329,353]" name="India" pageId="9" pageNumber="336">India</collectingCountry>
(
<geoCoordinate id="EE21A099FFBACC48A1B9FEFEFEE4FE6E" box="[159,299,365,390]" direction="north" orientation="latitude" pageId="9" pageNumber="336" precision="1" value="16.18618">16.18618°N</geoCoordinate>
,
<geoCoordinate id="EE21A099FFBACC48A010FEFEFE0EFE6E" box="[310,449,365,390]" direction="east" orientation="longitude" pageId="9" pageNumber="336" precision="1" value="74.75952">74.75952°E</geoCoordinate>
),
<elevation id="0038216DFFBACC48A0F5FEFEFDD5FE6D" box="[467,538,365,390]" metricMagnitude="2" metricUnit="m" metricValue="2.55" pageId="9" pageNumber="336" unit="m" value="255.0">
<quantity id="4CED6BBBFFBACC48A0F5FEFEFDD5FE6D" box="[467,538,365,390]" metricMagnitude="2" metricUnit="m" metricValue="2.55" pageId="9" pageNumber="336" unit="m" value="255.0">255 m</quantity>
</elevation>
elevation,
<collectionCode id="ED045E9BFFBACC48A3B0FEFDFD34FE6E" box="[662,763,366,390]" pageId="9" pageNumber="336">collected</collectionCode>
on 0 8.08.2013 by
<collectorName id="26E0A388FFBACC48A2EBFEFDFB8EFE6E" box="[973,1089,366,390]" pageId="9" pageNumber="336">V. Deepak</collectorName>
<collectionCode id="ED045E9BFFBACC48A56EFEFDFAD7FE6E" box="[1096,1304,366,390]" pageId="9" pageNumber="336">and Aparna Lajmi</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAAC65EFFBACC48A1E1FE02FE99FE19" blockId="9.[151,1436,329,966]" pageId="9" pageNumber="336">
<emphasis id="B9611A4CFFBACC48A1E1FE02FE89FE42" bold="true" box="[199,326,401,426]" pageId="9" pageNumber="336">Paratypes.</emphasis>
BNHS 2491, adult female (
<figureCitation id="132EDADBFFBACC48A359FE02FD1BFE42" box="[639,724,401,426]" captionStart="FIGURE 9" captionStartId="11.[151,249,1976,1998]" captionTargetBox="[193,1398,214,1932]" captionTargetId="figure@11.[170,1417,193,1955]" captionTargetPageId="11" captionText="FIGURE 9. Sitana gokakensis sp. nov. colouration in life. A. An adult male from type locality, B. paratype BNHS 2491, C &amp; D. two different uncollected adult females from the type locality." httpUri="https://zenodo.org/record/1290602/files/figure.png" pageId="9" pageNumber="336">Fig. 9b</figureCitation>
; Appendix 7b) collected by V. Deepak and Aparna Lajmi; CES
<date id="FFABE09EFFBACC48A1B1FE26FF22FE26" box="[151,237,437,462]" pageId="9" pageNumber="336" value="36-11-14">141136</date>
, an adult male (Appendix 7a) collected by V. Deepak and Kunal Arekar on
<date id="FFABE09EFFBACC48A568FE26FB69FE26" box="[1102,1190,437,462]" pageId="9" pageNumber="336" value="15-12-05">12.5.15</date>
; both same collection data as holotype.
</paragraph>
</subSubSection>
<subSubSection id="C30F95D5FFBACC48A1E1FE6EFBFBFC2E" pageId="9" pageNumber="336" type="diagnosis">
<paragraph id="8BAAC65EFFBACC48A1E1FE6EFBFBFC2E" blockId="9.[151,1436,329,966]" pageId="9" pageNumber="336">
<emphasis id="B9611A4CFFBACC48A1E1FE6EFE8EFDFE" bold="true" box="[199,321,509,534]" pageId="9" pageNumber="336">Diagnosis.</emphasis>
A large sized
<taxonomicName id="4C15BDDDFFBACC48A0CCFE6DFDFDFDFD" authorityName="Cuvier" authorityYear="1829" box="[490,562,510,533]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="genus">
<emphasis id="B9611A4CFFBACC48A0CCFE6DFDFDFDFD" box="[490,562,510,533]" italics="true" pageId="9" pageNumber="336">Sitana</emphasis>
</taxonomicName>
with a maximum SVL of
<quantity id="4CED6BBBFFBACC48A242FE6EFC02FDFD" box="[868,973,509,534]" metricMagnitude="-2" metricUnit="m" metricValue="5.31" pageId="9" pageNumber="336" unit="mm" value="53.1">53.1 mm</quantity>
, distinguished from its congeners by the following combination of characters: 1) dewlap feebly serrated without bright orange patches in breeding males (vs well serrated with bright orange patches in breeding males in
<taxonomicName id="4C15BDDDFFBACC48A242FDD5FBC0FDB5" box="[868,1039,582,605]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="species" species="ponticeriana">
<emphasis id="B9611A4CFFBACC48A242FDD5FBC0FDB5" box="[868,1039,582,605]" italics="true" pageId="9" pageNumber="336">S. ponticeriana</emphasis>
</taxonomicName>
,
<taxonomicName id="4C15BDDDFFBACC48A53AFDD5FBBEFDB5" authorityName="Deepak" authorityYear="2016" box="[1052,1137,582,605]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="species" species="visiri">
<emphasis id="B9611A4CFFBACC48A53AFDD5FBBEFDB5" box="[1052,1137,582,605]" italics="true" pageId="9" pageNumber="336">S. visiri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C15BDDDFFBACC48A559FDD5FAA2FDB5" authorityName="Deepak, Khandekar, Varma &amp; Chaitanya" authorityYear="2016" box="[1151,1389,581,605]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="species" species="marudhamneydhal">
<emphasis id="B9611A4CFFBACC48A559FDD5FAA2FDB5" box="[1151,1389,581,605]" italics="true" pageId="9" pageNumber="336">S. marudhamneydhal</emphasis>
</taxonomicName>
and
<taxonomicName id="4C15BDDDFFBACC48A1B1FDF9FEC3FD69" box="[151,268,617,641]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="species" species="devakai">
<emphasis id="B9611A4CFFBACC48A1B1FDF9FEC3FD69" box="[151,268,617,641]" italics="true" pageId="9" pageNumber="336">S. devakai</emphasis>
</taxonomicName>
); 2) dewlap extending beyond forearm insertion (vs not extending in
<emphasis id="B9611A4CFFBACC48A509FDF9FA82FD69" box="[1071,1357,617,641]" italics="true" pageId="9" pageNumber="336">
<taxonomicName id="4C15BDDDFFBACC48A509FDF9FB72FD69" authorityName="Schleich, Kästle &amp; Shah" authorityYear="1998" box="[1071,1213,617,641]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="species" species="sivalensis">S. sivalensis</taxonomicName>
,
<taxonomicName id="4C15BDDDFFBACC48A5EDFDF9FA82FD69" authorityName="Anders &amp; Kästle" authorityYear="2002" box="[1227,1357,617,641]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="species" species="schleichi">S. schleichi</taxonomicName>
</emphasis>
and
<taxonomicName id="4C15BDDDFFBACC48A4A1FDF9FF1DFD4D" authorityName="Schleich &amp; Kästle" authorityYear="1998" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="species" species="fusca">
<emphasis id="B9611A4CFFBACC48A4A1FDF9FF1DFD4D" italics="true" pageId="9" pageNumber="336">S. fusca</emphasis>
</taxonomicName>
); 3) four prominent enlarged non spine like scales bordering occipital region (vs enlarged spine like scales in
<taxonomicName id="4C15BDDDFFBACC48A1B1FD21FEAEFD21" authorityName="Deepak, Vyas &amp; Giri in Deepak, Giri, Asif, Dutta, Vyas, Zambre, Bhosale &amp; Karanth" authorityYear="2016" box="[151,353,689,713]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="species" species="spinaecephalus">
<emphasis id="B9611A4CFFBACC48A1B1FD21FEAEFD21" box="[151,353,689,713]" italics="true" pageId="9" pageNumber="336">S. spinaecephalus</emphasis>
</taxonomicName>
); 4) dewlap large sized extending up to 55% of trunk (vs up to 44 % of trunk in
<taxonomicName id="4C15BDDDFFBACC48A42EFD21FAB2FD21" box="[1288,1405,689,713]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="species" species="laticeps">
<emphasis id="B9611A4CFFBACC48A42EFD21FAB2FD21" box="[1288,1405,689,713]" italics="true" pageId="9" pageNumber="336">S. laticeps</emphasis>
</taxonomicName>
in
<collectingRegion id="49D108BCFFBACC48A1B1FD45FF37FD06" box="[151,248,726,750]" country="Sri Lanka" name="Southern" pageId="9" pageNumber="336">southern</collectingRegion>
most population which appears to be an outlier for the species; up to 29% of trunk in remaining population (
<figureCitation id="132EDADBFFBACC48A039FD69FEBCFCFA" box="[287,371,762,786]" captionStart="FIGURE 10" captionStartId="13.[151,249,1701,1723]" captionTargetBox="[157,1428,848,1659]" captionTargetId="figure@13.[151,1436,844,1680]" captionTargetPageId="13" captionText="FIGURE 10. Box plot showing proportion of dewlap in trunk. Note red circle highlighing the southern most population (Bagalkot &amp; Tummargudi) of S. laticeps which have relatively larger dewlaps compared to all other populations." httpUri="https://zenodo.org/record/1290604/files/figure.png" pageId="9" pageNumber="336">Fig. 10</figureCitation>
) and vs up to 42% of trunk in
<taxonomicName id="4C15BDDDFFBACC48A3FDFD69FCECFCF9" authorityName="Cuvier" authorityYear="1829" box="[731,803,762,785]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="genus">
<emphasis id="B9611A4CFFBACC48A3FDFD69FCECFCF9" box="[731,803,762,785]" italics="true" pageId="9" pageNumber="336">Sitana</emphasis>
</taxonomicName>
sp1 (
<figureCitation id="132EDADBFFBACC48A245FD69FC76FCFA" box="[867,953,762,786]" captionStart="FIGURE 10" captionStartId="13.[151,249,1701,1723]" captionTargetBox="[157,1428,848,1659]" captionTargetId="figure@13.[151,1436,844,1680]" captionTargetPageId="13" captionText="FIGURE 10. Box plot showing proportion of dewlap in trunk. Note red circle highlighing the southern most population (Bagalkot &amp; Tummargudi) of S. laticeps which have relatively larger dewlaps compared to all other populations." httpUri="https://zenodo.org/record/1290604/files/figure.png" pageId="9" pageNumber="336">Fig. 10</figureCitation>
); 5) body size (male &amp; females averaged) relatively large (SVL: mean 48.8 +/- 3.6 SE; range 42.453.1; n=14) vs (body size relatively small in
<taxonomicName id="4C15BDDDFFBACC48A401FC8DFA53FCDD" box="[1319,1436,797,821]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="species" species="laticeps">
<emphasis id="B9611A4CFFBACC48A401FC8DFA53FCDD" box="[1319,1436,797,821]" italics="true" pageId="9" pageNumber="336">S. laticeps</emphasis>
</taxonomicName>
SVL: mean 47.8 +/- 3.2 SE; range 40.7 54.6; n=63 and in
<emphasis id="B9611A4CFFBACC48A262FCD1FC95FCB1" box="[836,858,834,857]" italics="true" pageId="9" pageNumber="336">S.</emphasis>
sp1 SVL: mean 46.2 +/- 3.2 SE; range 38.954.1; n=31 and
<taxonomicName id="4C15BDDDFFBACC48A02FFCF5FE1CFC95" authorityName="Deepak, Vyas &amp; Giri in Deepak, Giri, Asif, Dutta, Vyas, Zambre, Bhosale &amp; Karanth" authorityYear="2016" box="[265,467,869,893]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="species" species="spinaecephalus">
<emphasis id="B9611A4CFFBACC48A02FFCF5FE1CFC95" box="[265,467,869,893]" italics="true" pageId="9" pageNumber="336">S. spinaecephalus</emphasis>
</taxonomicName>
SVL: mean 47.3 +/- 3.7 SE; range 39.5 56.6; n=25). It has to be noted here that the southernmost population of
<taxonomicName id="4C15BDDDFFBACC48A0F3FC19FD86FC49" box="[469,585,905,929]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="species" species="laticeps">
<emphasis id="B9611A4CFFBACC48A0F3FC19FD86FC49" box="[469,585,905,929]" italics="true" pageId="9" pageNumber="336">S. laticeps</emphasis>
</taxonomicName>
appears to have some relatively large bodied individuals (SVL: mean 50.1 +/- 4.2 SE; range 45.254.6; n=6) but need a larger sample size to further confirm.
</paragraph>
</subSubSection>
<caption id="DF6A96D6FFBACC48A1B1F80EFC4BF839" httpUri="https://zenodo.org/record/1290598/files/figure.png" pageId="9" pageNumber="336" startId="9.[151,249,1949,1971]" targetBox="[175,1395,1035,1917]" targetPageId="9">
<paragraph id="8BAAC65EFFBACC48A1B1F80EFC4BF839" blockId="9.[151,1435,1948,2001]" pageId="9" pageNumber="336">
<emphasis id="B9611A4CFFBACC48A1B1F80EFEDAF85A" bold="true" box="[151,277,1949,1971]" pageId="9" pageNumber="336">FIGURE 7.</emphasis>
Holotype of
<taxonomicName id="4C15BDDDFFBACC48A082F80EFD92F85A" authority="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth, 2018" authorityName="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth" authorityYear="2018" box="[420,605,1948,1970]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="336" phylum="Chordata" rank="species" species="gokakensis" status="sp. nov.">
<emphasis id="B9611A4CFFBACC48A082F80EFD92F85A" box="[420,605,1948,1970]" italics="true" pageId="9" pageNumber="336">Sitana gokakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A252A737FFBACC48A340F80EFD76F85A" box="[614,697,1949,1970]" pageId="9" pageNumber="336" rank="species">
<emphasis id="B9611A4CFFBACC48A340F80EFD76F85A" bold="true" box="[614,697,1949,1970]" pageId="9" pageNumber="336">sp. nov.</emphasis>
</taxonomicNameLabel>
(BNHS 2490) from Gokak, Karnataka. (A) full body dorsal (B) flank region (C) dorsal view of head (D) lateral view of head. Scale bar:10 mm.
</paragraph>
</caption>
<caption id="DF6A96D6FFB9CC4BA1B1F83CFE37F80B" httpUri="https://zenodo.org/record/1290600/files/figure.png" pageId="10" pageNumber="337" startId="10.[151,249,1967,1989]" targetBox="[199,1381,226,1914]" targetPageId="10">
<paragraph id="8BAAC65EFFB9CC4BA1B1F83CFE37F80B" blockId="10.[151,1435,1967,2019]" pageId="10" pageNumber="337">
<emphasis id="B9611A4CFFB9CC4BA1B1F83CFEDCF82C" bold="true" box="[151,275,1967,1989]" pageId="10" pageNumber="337">FIGURE 8.</emphasis>
<taxonomicName id="4C15BDDDFFB9CC4BA03CF823FE1EF82D" authority="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth, 2018" authorityName="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth" authorityYear="2018" box="[282,465,1967,1989]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="337" phylum="Chordata" rank="species" species="gokakensis" status="sp. nov.">
<emphasis id="B9611A4CFFB9CC4BA03CF823FE1EF82D" box="[282,465,1967,1989]" italics="true" pageId="10" pageNumber="337">Sitana gokakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A252A737FFB9CC4BA0FFF83CFDE5F82C" box="[473,554,1967,1988]" pageId="10" pageNumber="337" rank="species">
<emphasis id="B9611A4CFFB9CC4BA0FFF83CFDE5F82C" bold="true" box="[473,554,1967,1988]" pageId="10" pageNumber="337">sp. nov.</emphasis>
</taxonomicNameLabel>
colouration in life. A. Holotype (BNHS 2490) in life, B. another adult male collected during 2016 from the type locality.
</paragraph>
</caption>
<caption id="DF6A96D6FFB8CC4AA1B1F82BFCE7F805" httpUri="https://zenodo.org/record/1290602/files/figure.png" pageId="11" pageNumber="338" startId="11.[151,249,1976,1998]" targetBox="[193,1398,214,1932]" targetPageId="11">
<paragraph id="8BAAC65EFFB8CC4AA1B1F82BFCE7F805" blockId="11.[151,1436,1976,2029]" pageId="11" pageNumber="338">
<emphasis id="B9611A4CFFB8CC4AA1B1F82BFEDDF825" bold="true" box="[151,274,1976,1998]" pageId="11" pageNumber="338">FIGURE 9.</emphasis>
<taxonomicName id="4C15BDDDFFB8CC4AA03FF82AFE1FF826" authority="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth, 2018" authorityName="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth" authorityYear="2018" box="[281,464,1976,1998]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="338" phylum="Chordata" rank="species" species="gokakensis" status="sp. nov.">
<emphasis id="B9611A4CFFB8CC4AA03FF82AFE1FF826" box="[281,464,1976,1998]" italics="true" pageId="11" pageNumber="338">Sitana gokakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A252A737FFB8CC4AA0F1F82AFDE7F826" box="[471,552,1977,1998]" pageId="11" pageNumber="338" rank="species">
<emphasis id="B9611A4CFFB8CC4AA0F1F82AFDE7F826" bold="true" box="[471,552,1977,1998]" pageId="11" pageNumber="338">sp. nov.</emphasis>
</taxonomicNameLabel>
colouration in life. A. An adult male from type locality, B. paratype BNHS 2491, C &amp; D. two different uncollected adult females from the type locality.
</paragraph>
</caption>
<subSubSection id="C30F95D5FFBFCC4FA1E1FF04FA52F823" lastPageId="14" lastPageNumber="341" pageId="12" pageNumber="339" type="description">
<paragraph id="8BAAC65EFFBFCC4DA1E1FF04FC7CF898" blockId="12.[151,1438,151,2013]" pageId="12" pageNumber="339">
<emphasis id="B9611A4CFFBFCC4DA1E1FF04FE2AFF58" bold="true" box="[199,485,151,176]" pageId="12" pageNumber="339">
Description of
<typeStatus id="54AE78FCFFBFCC4DA051FF04FE2FFF58" box="[375,480,151,176]" pageId="12" pageNumber="339" type="holotype">holotype</typeStatus>
.
</emphasis>
The
<typeStatus id="54AE78FCFFBFCC4DA338FF0BFD4FFF58" box="[542,640,152,176]" pageId="12" pageNumber="339" type="holotype">holotype</typeStatus>
is in good condition; hemipenis partially everted, exposed and seen on both sides when viewed dorsally (
<figureCitation id="132EDADBFFBFCC4DA330FF2FFDA2FF3C" box="[534,621,188,212]" captionStart="FIGURE 7" captionStartId="9.[151,249,1949,1971]" captionTargetBox="[175,1395,1035,1917]" captionTargetId="figure@9.[151,1436,1018,1927]" captionTargetPageId="9" captionText="FIGURE 7. Holotype of Sitana gokakensis sp. nov. (BNHS 2490) from Gokak, Karnataka. (A) full body dorsal (B) flank region (C) dorsal view of head (D) lateral view of head. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290598/files/figure.png" pageId="12" pageNumber="339">Fig. 7a</figureCitation>
). Tail entire, curved slightly towards the right; loose folds of skin on the dorsum and a small incision of
<quantity id="4CED6BBBFFBFCC4DA0D3FF73FDE0FF10" box="[501,559,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="339" unit="mm" value="4.0">4mm</quantity>
to extricate tissue, are artefacts of preservation. An adult male, SVL
<quantity id="4CED6BBBFFBFCC4DA414FF4CFA55FF1F" box="[1330,1434,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="5.12" pageId="12" pageNumber="339" unit="mm" value="51.2">51.2 mm</quantity>
. Head relatively long (HL/SVL ratio 0.29), wide (HW/HL ratio 0.70), not depressed (HH/HL ratio 0.51), distinct from neck. Snout short (SE/HL ratio 0.39), longer than diameter of orbit (OD/SE ratio 0.73), obtusely pointed in profile when viewed dorsally (
<figureCitation id="132EDADBFFBFCC4DA0DEFEDFFD9FFE8C" box="[504,592,332,356]" captionStart="FIGURE 7" captionStartId="9.[151,249,1949,1971]" captionTargetBox="[175,1395,1035,1917]" captionTargetId="figure@9.[151,1436,1018,1927]" captionTargetPageId="9" captionText="FIGURE 7. Holotype of Sitana gokakensis sp. nov. (BNHS 2490) from Gokak, Karnataka. (A) full body dorsal (B) flank region (C) dorsal view of head (D) lateral view of head. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290598/files/figure.png" pageId="12" pageNumber="339">Fig. 7c</figureCitation>
); rostral much wider than high, contacted laterally on either side by first supralabial, a prenasal and dorsally by two smaller, keeled scales (
<figureCitation id="132EDADBFFBFCC4DA25DFEE3FC1FFE60" box="[891,976,368,392]" captionStart="FIGURE 7" captionStartId="9.[151,249,1949,1971]" captionTargetBox="[175,1395,1035,1917]" captionTargetId="figure@9.[151,1436,1018,1927]" captionTargetPageId="9" captionText="FIGURE 7. Holotype of Sitana gokakensis sp. nov. (BNHS 2490) from Gokak, Karnataka. (A) full body dorsal (B) flank region (C) dorsal view of head (D) lateral view of head. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290598/files/figure.png" pageId="12" pageNumber="339">Fig. 7e</figureCitation>
). Canthus rostralis and supraciliary edge, sharp. Nostril roughly circular, laterally positioned and placed in the centre of a large, undivided nasal. Nasal scale bordered by eight scales on either side: one supranasal, three postnasals, one prenasal, the last two scales form a series of enlarged scales bordering the supralabials and a small scale separates them from the first supralabial. Ten supralabials on the right side (11 on left side), first slightly higher than others, broader than long, roughly rectangular, rest more elongate, weakly keeled, bordered above by a row of slightly smaller, rectangular, weakly keeled scales, which start at posterior margin of first supralabial, decreasing in size posteriorly and terminating after the posterior margin of orbit. Twelve infralabials on the right side (13 on left), first slightly smaller, the rest elongate, keeled increasing in size posteriorly. Loreal region concave, with scales of heterogeneous shape and size. Canthals enlarged, overlapping, slightly protruding on supraorbital ridge laterally. Eye large (ED/HL ratio 0.27); pupil rounded, covered under the eyelids; eyelids covered with scales that are heterogeneous in shape and size; larger scales on the upper eyelids, keeled, elongate and bluntly pointed, rest smooth; supraciliaries longer than broad. Orbital scales small but not granular. Scales on postorbital and temporal region, heterogeneous, sub- imbricate, strongly keeled, directed backward and upwards. Tympanum naked. Scales on dorsal surface of snout, forehead, interorbital and occipital region highly heterogeneous in size, shape, mostly elongate, strongly keeled longitudinally (
<figureCitation id="132EDADBFFBFCC4DA065FC1FFE56FC4C" box="[323,409,908,932]" captionStart="FIGURE 7" captionStartId="9.[151,249,1949,1971]" captionTargetBox="[175,1395,1035,1917]" captionTargetId="figure@9.[151,1436,1018,1927]" captionTargetPageId="9" captionText="FIGURE 7. Holotype of Sitana gokakensis sp. nov. (BNHS 2490) from Gokak, Karnataka. (A) full body dorsal (B) flank region (C) dorsal view of head (D) lateral view of head. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290598/files/figure.png" pageId="12" pageNumber="339">Fig. 7c</figureCitation>
); scales on snout large in size, those on forehead slightly larger, interorbital region largest; occipital region with much smaller imbricate scales relative to other dorsal head scales; four relatively enlarged scales on a transverse row in the occiput region; nine scales anterior and 14 scales posterior of eyelids in the interorbital region; supraorbital scales along the supraciliary edge elongate, keeled, decreasing in size posteriorly, following curvature of orbit. Parietals larger than surrounding scales, longer than broad, tricarinate, separated from each other by two inter-parietal scales; anterior large, roughly triangular, tricarinate; posterior roughly pentagonal, bicarinate, with distinctly visible pineal eye. Parietals and the posterior inter-parietal part of a series of nine large, strongly keeled scales traversing the forehead. Mental shield narrower than rostral, roughly pentagonal, pointed posteriorly, a pair of roughly hexagonal postmentals, slightly shorter than mental, completely divided by a smaller gular scale (
<figureCitation id="132EDADBFFBFCC4DA006FB43FEB7FB00" box="[288,376,1232,1256]" captionStart="FIGURE 7" captionStartId="9.[151,249,1949,1971]" captionTargetBox="[175,1395,1035,1917]" captionTargetId="figure@9.[151,1436,1018,1927]" captionTargetPageId="9" captionText="FIGURE 7. Holotype of Sitana gokakensis sp. nov. (BNHS 2490) from Gokak, Karnataka. (A) full body dorsal (B) flank region (C) dorsal view of head (D) lateral view of head. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290598/files/figure.png" pageId="12" pageNumber="339">Fig. 7d</figureCitation>
); scales on the gular region homogenous in shape, those behind mental smooth, increasing in size and carination posteriorly. Dewlap large (DEWL/SVL ratio 0.69), extends posteriorly over 55% of trunk, extending much beyond axilla (
<figureCitation id="132EDADBFFBFCC4DA321FA8BFDAFFAD8" box="[519,608,1303,1328]" captionStart="FIGURE 7" captionStartId="9.[151,249,1949,1971]" captionTargetBox="[175,1395,1035,1917]" captionTargetId="figure@9.[151,1436,1018,1927]" captionTargetPageId="9" captionText="FIGURE 7. Holotype of Sitana gokakensis sp. nov. (BNHS 2490) from Gokak, Karnataka. (A) full body dorsal (B) flank region (C) dorsal view of head (D) lateral view of head. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290598/files/figure.png" pageId="12" pageNumber="339">Fig. 7b</figureCitation>
); about nine rows of anterior dewlap scales smaller, elongate, pointed, keeled; remainder of scales much enlarged, keeled, pointed, gradually increasing in size towards margin; single marginal row largest with lanceolate scales. Enlarged scales on dewlap in 23 rows. Nuchal and dorsal crest absent (
<figureCitation id="132EDADBFFBFCC4DA1B8FA17FF3BFA74" box="[158,244,1412,1436]" captionStart="FIGURE 7" captionStartId="9.[151,249,1949,1971]" captionTargetBox="[175,1395,1035,1917]" captionTargetId="figure@9.[151,1436,1018,1927]" captionTargetPageId="9" captionText="FIGURE 7. Holotype of Sitana gokakensis sp. nov. (BNHS 2490) from Gokak, Karnataka. (A) full body dorsal (B) flank region (C) dorsal view of head (D) lateral view of head. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290598/files/figure.png" pageId="12" pageNumber="339">Fig. 7a</figureCitation>
). Scales on nuchal region smaller, less than half the size of those on interorbital region, imbricate, strongly keeled. Body slender, 60 rows of scales around midbody; vertebral scale row with 46 scales, vertebral scale row partially paired and alternating. Five enlarged dorsal scale rows, on either side of the vertebral scale row (except in 3 places the five rows on either side are in contact with each other), the 5 dorsal scale rows starts from back of neck until groin, sub equal in size and shape, imbricate, pointed, keeled, directed backwards forming regularly arranged longitudinal rows; scales on flanks heterogeneous in size, shape, smaller than those on back, pointed, keeled, upper rows directed backwards and upwards, lower rows backwards and downwards; ventral scales subimbricate, keeled, homogenous in size and shape, arranged in 101 rows. Fore and hindlimbs relatively slender, tibia short (CL/SVL ratio 0.33); digits moderately long, ending in strong, elongate, slightly recurved claw; inter-digital webbing absent; subdigital lamellae entire, bi-mucronate, 22 subdigital lamellae on toe IV including claw sheath; relative length of right fingers 3&gt; 4&gt; 2&gt; 5&gt; 1, right toes 4&gt; 3&gt; 2&gt; 1. Fore and hindlimbs covered above and below with regularly arranged, enlarged, pointed, strongly keeled, scales. Tail long (TL/SVL ratio 2.35), entire, base swollen, uniformly covered with similar sized, keeled, pointed, regularly arranged, backwardly directed imbricate scales; subcaudal scales keeled, weakly pointed near base, becoming pointed posteriorly.
</paragraph>
<paragraph id="8BAAC65EFFBFCC4CA1E1F8EFFC5FFEA8" blockId="12.[151,1438,151,2013]" lastBlockId="13.[151,1438,151,791]" lastPageId="13" lastPageNumber="340" pageId="12" pageNumber="339">
<materialsCitation id="3B7DCC03FFBFCC4DA1E1F8EFFEAAF850" ID-GBIF-Occurrence="1849038564" collectionCode="Head, than, the, torso, dark, flank, and, tail" location="Head" municipality="Colour" pageId="12" pageNumber="339" specimenCount="1" typeStatus="holotype">
<emphasis id="B9611A4CFFBFCC4DA1E1F8EFFE35F87D" bold="true" box="[199,506,1916,1941]" pageId="12" pageNumber="339">
<collectingMunicipality id="6BCE5C24FFBFCC4DA1E1F8EFFED4F87D" box="[199,283,1916,1941]" pageId="12" pageNumber="339">Colour</collectingMunicipality>
of
<typeStatus id="54AE78FCFFBFCC4DA067F8EFFE68F87D" box="[321,423,1916,1941]" pageId="12" pageNumber="339" type="holotype">holotype</typeStatus>
in life.
</emphasis>
<location id="8ECA9085FFBFCC4DA327F8EFFDF3F87C" LSID="urn:lsid:plazi:treatment:03BC7748FFBACC51A1B1FF4DFB6AFEF4:8ECA9085FFBFCC4DA327F8EFFDF3F87C" box="[513,572,1916,1940]" municipality="Colour" name="Head" pageId="12" pageNumber="339">
<collectionCode id="ED045E9BFFBFCC4DA327F8EFFDF3F87C" box="[513,572,1916,1940]" pageId="12" pageNumber="339">Head</collectionCode>
</location>
darker
<collectionCode id="ED045E9BFFBFCC4DA3B5F8EFFD0CF87C" box="[659,707,1916,1940]" pageId="12" pageNumber="339">than</collectionCode>
body, dorsum of
<collectionCode id="ED045E9BFFBFCC4DA2AFF8EEFBEBF87C" box="[905,1060,1916,1940]" pageId="12" pageNumber="339">the torso dark</collectionCode>
brown, darker compared to
<collectionCode id="ED045E9BFFBFCC4DA444F8EFFF3FF851" pageId="12" pageNumber="339">flank and tail</collectionCode>
(
<figureCitation id="132EDADBFFBFCC4DA025F833FE91F850" box="[259,350,1952,1976]" captionStart="FIGURE 7" captionStartId="9.[151,249,1949,1971]" captionTargetBox="[175,1395,1035,1917]" captionTargetId="figure@9.[151,1436,1018,1927]" captionTargetPageId="9" captionText="FIGURE 7. Holotype of Sitana gokakensis sp. nov. (BNHS 2490) from Gokak, Karnataka. (A) full body dorsal (B) flank region (C) dorsal view of head (D) lateral view of head. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290598/files/figure.png" pageId="12" pageNumber="339">Fig. 7a</figureCitation>
)
</materialsCitation>
.
<materialsCitation id="3B7DCC03FFBFCC4DA053F833FC70F850" ID-GBIF-Occurrence="1849038563" box="[373,959,1952,1977]" collectionCode="Neck, region, with, and, dark, pink" location="Neck region" pageId="12" pageNumber="339" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED045E9BFFBFCC4DA053F833FD8DF850" box="[373,578,1952,1976]" pageId="12" pageNumber="339">
<location id="8ECA9085FFBFCC4DA053F833FDCCF850" LSID="urn:lsid:plazi:treatment:03BC7748FFBACC51A1B1FF4DFB6AFEF4:8ECA9085FFBFCC4DA053F833FDCCF850" box="[373,515,1952,1976]" name="Neck region" pageId="12" pageNumber="339">Neck region</location>
with
</collectionCode>
blue
<collectionCode id="ED045E9BFFBFCC4DA3ACF832FCE0F850" box="[650,815,1953,1977]" pageId="12" pageNumber="339">and dark pink</collectionCode>
colouration
</materialsCitation>
.
<materialsCitation id="3B7DCC03FFBFCC4DA2E8F833FCB2F834" ID-GBIF-Occurrence="1849038561" collectionCode="Iris, colour, cream, with, five, the, one, neck" location="Dorsum" municipality="Iris" pageId="12" pageNumber="339" specimenCount="1" typeStatus="holotype">
<collectingMunicipality id="6BCE5C24FFBFCC4DA2E8F833FC3CF850" box="[974,1011,1952,1976]" pageId="12" pageNumber="339">
<collectionCode id="ED045E9BFFBFCC4DA2E8F833FC3CF850" box="[974,1011,1952,1976]" pageId="12" pageNumber="339">Iris</collectionCode>
</collectingMunicipality>
yellow in
<collectionCode id="ED045E9BFFBFCC4DA55CF833FB0AF850" box="[1146,1221,1952,1976]" pageId="12" pageNumber="339">colour</collectionCode>
, tympanum
<collectionCode id="ED045E9BFFBFCC4DA47EF833FA52F850" box="[1368,1437,1952,1976]" pageId="12" pageNumber="339">cream</collectionCode>
coloured.
<location id="8ECA9085FFBFCC4DA021F857FEAEF834" LSID="urn:lsid:plazi:treatment:03BC7748FFBACC51A1B1FF4DFB6AFEF4:8ECA9085FFBFCC4DA021F857FEAEF834" box="[263,353,1988,2012]" municipality="Iris" name="Dorsum" pageId="12" pageNumber="339">Dorsum</location>
<collectionCode id="ED045E9BFFBFCC4DA04EF856FE02F834" box="[360,461,1989,2012]" pageId="12" pageNumber="339">with five</collectionCode>
black blotches
<collectionCode id="ED045E9BFFBFCC4DA35DF856FD03F834" box="[635,716,1988,2012]" pageId="12" pageNumber="339">the one</collectionCode>
on
<collectionCode id="ED045E9BFFBFCC4DA3D1F857FCE3F834" box="[759,812,1988,2012]" pageId="12" pageNumber="339">neck</collectionCode>
darker
</materialsCitation>
.
<materialsCitation id="3B7DCC03FFBFCC4CA2A0F857FC14FF58" ID-GBIF-Occurrence="1849038558" collectionCode="Flank, region, red, colour, with, mottled, dark, scales, the, flank, and, thigh, cream" lastPageId="13" lastPageNumber="340" location="Enlarged" pageId="12" pageNumber="339" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED045E9BFFBFCC4DA2A0F857FBDBF834" box="[902,1044,1988,2012]" pageId="12" pageNumber="339">Flank region</collectionCode>
brick
<collectionCode id="ED045E9BFFBFCC4DA57AF856FB4FF834" box="[1116,1152,1989,2012]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14967" name="University of Redlands" pageId="12" pageNumber="339">red</collectionCode>
<collectionCode id="ED045E9BFFBFCC4DA5A1F857FA52F834" box="[1159,1437,1988,2012]" pageId="12" pageNumber="339">colour with mottled dark</collectionCode>
brown patches.
<location id="8ECA9085FFBECC4CA076FF0BFE7BFF58" LSID="urn:lsid:plazi:treatment:03BC7748FFBACC51A1B1FF4DFB6AFEF4:8ECA9085FFBECC4CA076FF0BFE7BFF58" box="[336,436,152,176]" name="Enlarged" pageId="13" pageNumber="340">Enlarged</location>
<collectionCode id="ED045E9BFFBECC4CA099FF0BFDCDFF58" box="[447,514,152,176]" pageId="13" pageNumber="340">scales</collectionCode>
on
<collectionCode id="ED045E9BFFBECC4CA313FF0BFCABFF58" box="[565,868,152,176]" pageId="13" pageNumber="340">the flank and thigh cream</collectionCode>
in
<collectionCode id="ED045E9BFFBECC4CA2B6FF0BFC14FF58" box="[912,987,152,176]" pageId="13" pageNumber="340">colour</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7DCC03FFBECC4CA2CEFF0BFC20FF10" ID-GBIF-Occurrence="1849038460" collectionCode="Belly, white, colour, with, most, scales, and, dark, suffused, red, dorsal, side, cream, patch, the, thigh, extending, region, pale" location="Forelimbs" municipality="Belly" pageId="13" pageNumber="340" specimenCount="1" typeStatus="holotype">
<collectingMunicipality id="6BCE5C24FFBECC4CA2CEFF0BFBEAFF47" box="[1000,1061,152,175]" pageId="13" pageNumber="340">
<collectionCode id="ED045E9BFFBECC4CA2CEFF0BFBEAFF47" box="[1000,1061,152,175]" pageId="13" pageNumber="340">Belly</collectionCode>
</collectingMunicipality>
off
<collectionCode id="ED045E9BFFBECC4CA57DFF0BFB56FF47" box="[1115,1177,152,175]" pageId="13" pageNumber="340">white</collectionCode>
in
<collectionCode id="ED045E9BFFBECC4CA5E3FF0BFA85FF47" box="[1221,1354,152,176]" pageId="13" pageNumber="340">colour with</collectionCode>
brown speckles on
<collectionCode id="ED045E9BFFBECC4CA006FF2FFE6AFF3C" box="[288,421,188,212]" pageId="13" pageNumber="340">most scales</collectionCode>
.
<location id="8ECA9085FFBECC4CA088FF2FFDEFFF3D" LSID="urn:lsid:plazi:treatment:03BC7748FFBACC51A1B1FF4DFB6AFEF4:8ECA9085FFBECC4CA088FF2FFDEFFF3D" box="[430,544,188,213]" municipality="Belly" name="Forelimbs" pageId="13" pageNumber="340">Forelimbs</location>
<collectionCode id="ED045E9BFFBECC4CA301FF2FFD9FFF3C" box="[551,592,188,212]" pageId="13" pageNumber="340">and</collectionCode>
hindlimbs
<collectionCode id="ED045E9BFFBECC4CA3E8FF2FFCCFFF3C" box="[718,768,188,212]" pageId="13" pageNumber="340">dark</collectionCode>
brown
<collectionCode id="ED045E9BFFBECC4CA273FF2FFC23FF3C" box="[853,1004,188,212]" pageId="13" pageNumber="340">suffused with</collectionCode>
<collectionCode id="ED045E9BFFBECC4CA2D5FF2EFBD8FF3C" box="[1011,1047,189,212]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14967" name="University of Redlands" pageId="13" pageNumber="340">red</collectionCode>
on
<collectionCode id="ED045E9BFFBECC4CA567FF2FFB72FF3C" box="[1089,1213,188,212]" pageId="13" pageNumber="340">dorsal side</collectionCode>
,
<collectionCode id="ED045E9BFFBECC4CA5EEFF2FFA9FFF3C" box="[1224,1360,188,212]" pageId="13" pageNumber="340">cream patch</collectionCode>
on
<collectionCode id="ED045E9BFFBECC4CA45CFF2EFA53FF3C" box="[1402,1436,189,212]" pageId="13" pageNumber="340">the</collectionCode>
back of
<collectionCode id="ED045E9BFFBECC4CA1D3FF73FE69FF10" box="[245,422,224,248]" pageId="13" pageNumber="340">thigh extending</collectionCode>
to
<collectionCode id="ED045E9BFFBECC4CA0E8FF73FE3FFF1F" box="[462,496,224,247]" pageId="13" pageNumber="340">the</collectionCode>
saccaral
<collectionCode id="ED045E9BFFBECC4CA37BFF73FCC1FF10" box="[605,782,224,248]" pageId="13" pageNumber="340">region and pale</collectionCode>
on
<collectionCode id="ED045E9BFFBECC4CA21DFF73FC92FF1F" box="[827,861,224,247]" pageId="13" pageNumber="340">the</collectionCode>
ventral
<collectionCode id="ED045E9BFFBECC4CA298FF73FC20FF10" box="[958,1007,224,248]" pageId="13" pageNumber="340">side</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7DCC03FFBECC4CA2DDFF73FAE9FF10" ID-GBIF-Occurrence="1849038463" box="[1019,1318,224,248]" collectionCode="Lower, jaws, cream" location="Lower" pageId="13" pageNumber="340" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED045E9BFFBECC4CA2DDFF73FB01FF10" box="[1019,1230,224,248]" pageId="13" pageNumber="340">
<location id="8ECA9085FFBECC4CA2DDFF73FB8BFF10" LSID="urn:lsid:plazi:treatment:03BC7748FFBACC51A1B1FF4DFB6AFEF4:8ECA9085FFBECC4CA2DDFF73FB8BFF10" box="[1019,1092,224,248]" name="Lower" pageId="13" pageNumber="340">Lower</location>
jaws cream
</collectionCode>
mostly
</materialsCitation>
,
<materialsCitation id="3B7DCC03FFBECC4CA417FF73FA56FEF4" ID-GBIF-Occurrence="1849038650" collectionCode="dark, from, the, scale, rows, and, along, dewlap" location="mentum" pageId="13" pageNumber="340" specimenCount="1" stateProvince="mentum" typeStatus="holotype">
<collectionCode id="ED045E9BFFBECC4CA417FF73FAACFF10" box="[1329,1379,224,248]" pageId="13" pageNumber="340">dark</collectionCode>
blue colouration starts
<collectionCode id="ED045E9BFFBECC4CA044FE97FE0DFEF4" box="[354,450,260,284]" pageId="13" pageNumber="340">from the</collectionCode>
<collectingRegion id="49D108BCFFBECC4CA0EDFE96FDE8FEF4" box="[459,551,261,284]" country="Myanmar" name="Chin" pageId="13" pageNumber="340">mentum</collectingRegion>
(on 78 horizontal
<collectionCode id="ED045E9BFFBECC4CA22EFE97FCB1FEF4" box="[776,894,260,284]" pageId="13" pageNumber="340">scale rows</collectionCode>
(
<figureCitation id="132EDADBFFBECC4CA2A8FE97FC2CFEF4" box="[910,995,260,284]" captionStart="FIGURE 7" captionStartId="9.[151,249,1949,1971]" captionTargetBox="[175,1395,1035,1917]" captionTargetId="figure@9.[151,1436,1018,1927]" captionTargetPageId="9" captionText="FIGURE 7. Holotype of Sitana gokakensis sp. nov. (BNHS 2490) from Gokak, Karnataka. (A) full body dorsal (B) flank region (C) dorsal view of head (D) lateral view of head. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290598/files/figure.png" pageId="13" pageNumber="340">Fig. 7d</figureCitation>
)
<collectionCode id="ED045E9BFFBECC4CA2D4FE97FBD5FEF4" box="[1010,1050,260,284]" pageId="13" pageNumber="340">and</collectionCode>
extends
<collectionCode id="ED045E9BFFBECC4CA5A7FE97FB70FEF4" box="[1153,1215,260,284]" pageId="13" pageNumber="340">along</collectionCode>
midline of
<collectionCode id="ED045E9BFFBECC4CA462FE97FA56FEF4" box="[1348,1433,260,284]" pageId="13" pageNumber="340">dewlap</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7DCC03FFBECC4CA1B1FEBBFC5FFEA8" ID-GBIF-Occurrence="1849038464" box="[151,912,295,320]" collectionCode="Dewlap, white, each, scale, dewlap, with" location="Dewlap" pageId="13" pageNumber="340" specimenCount="1" stateProvince="mentum" typeStatus="holotype">
<collectionCode id="ED045E9BFFBECC4CA1B1FEBBFEF8FED7" box="[151,311,296,320]" pageId="13" pageNumber="340">
<location id="8ECA9085FFBECC4CA1B1FEBBFF21FEA8" LSID="urn:lsid:plazi:treatment:03BC7748FFBACC51A1B1FF4DFB6AFEF4:8ECA9085FFBECC4CA1B1FEBBFF21FEA8" box="[151,238,296,320]" name="Dewlap" pageId="13" pageNumber="340" stateProvince="mentum">Dewlap</location>
white
</collectionCode>
,
<collectionCode id="ED045E9BFFBECC4CA067FEBBFE7CFEA8" box="[321,435,296,320]" pageId="13" pageNumber="340">each scale</collectionCode>
on
<collectionCode id="ED045E9BFFBECC4CA0F8FEBBFDA8FED7" box="[478,615,296,320]" pageId="13" pageNumber="340">dewlap with</collectionCode>
clustered brown speckles.
</materialsCitation>
</paragraph>
<paragraph id="8BAAC65EFFBECC4CA1E1FEDFFED4FCFF" blockId="13.[151,1438,151,791]" pageId="13" pageNumber="340">
<materialsCitation id="3B7DCC03FFBECC4CA1E1FEDFFA91FE38" ID-GBIF-Occurrence="1849038461" collectionCode="Head, dark, than, the, neck, Two, start, that, and, after, cream, scales, with, small, light, two" location="Dorsum" municipality="Tympanum" pageId="13" pageNumber="340" specimenCount="1" typeStatus="holotype">
<emphasis id="B9611A4CFFBECC4CA1E1FEDFFDA6FE8D" bold="true" box="[199,617,332,357]" pageId="13" pageNumber="340">
Colour of
<typeStatus id="54AE78FCFFBECC4CA067FEDFFE67FE8D" box="[321,424,332,357]" pageId="13" pageNumber="340" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
<collectionCode id="ED045E9BFFBECC4CA357FEDFFD2AFE8C" box="[625,741,332,356]" pageId="13" pageNumber="340">Head dark</collectionCode>
brown, darker
<collectionCode id="ED045E9BFFBECC4CA2B4FEDFFC23FE8C" box="[914,1004,332,356]" pageId="13" pageNumber="340">than the</collectionCode>
body—with scattered blue patches on
<collectionCode id="ED045E9BFFBECC4CA1B1FEE3FF32FE60" box="[151,253,368,392]" pageId="13" pageNumber="340">the neck</collectionCode>
.
<collectionCode id="ED045E9BFFBECC4CA02AFEE3FEF2FE60" box="[268,317,368,392]" pageId="13" pageNumber="340">Two</collectionCode>
broken black stripes
<collectionCode id="ED045E9BFFBECC4CA319FEE3FDA0FE60" box="[575,623,368,392]" pageId="13" pageNumber="340">start</collectionCode>
below
<collectionCode id="ED045E9BFFBECC4CA3ECFEE3FD23FE6F" box="[714,748,368,391]" pageId="13" pageNumber="340">the</collectionCode>
eye approximately in
<collectionCode id="ED045E9BFFBECC4CA2DFFEE3FBD4FE6F" box="[1017,1051,368,391]" pageId="13" pageNumber="340">the</collectionCode>
middle directed downwards
<collectionCode id="ED045E9BFFBECC4CA454FEE3FA53FE60" box="[1394,1436,368,392]" pageId="13" pageNumber="340">that</collectionCode>
broaden
<collectionCode id="ED045E9BFFBECC4CA1DAFE07FEEBFE44" box="[252,292,404,428]" pageId="13" pageNumber="340">and</collectionCode>
end
<collectionCode id="ED045E9BFFBECC4CA042FE07FE0AFE44" box="[356,453,404,428]" pageId="13" pageNumber="340">after the</collectionCode>
forearm insertion.
<collectingMunicipality id="6BCE5C24FFBECC4CA38FFE07FCE7FE44" box="[681,808,404,428]" pageId="13" pageNumber="340">Tympanum</collectingMunicipality>
<collectionCode id="ED045E9BFFBECC4CA215FE07FCB7FE44" box="[819,888,404,428]" pageId="13" pageNumber="340">cream</collectionCode>
coloured, slightly lighter
<collectionCode id="ED045E9BFFBECC4CA58DFE07FAC6FE44" box="[1195,1289,404,428]" pageId="13" pageNumber="340">than the</collectionCode>
surrounding
<collectionCode id="ED045E9BFFBECC4CA1B1FE2BFE58FE38" box="[151,407,440,464]" pageId="13" pageNumber="340">scales with small light</collectionCode>
brown speckles.
<location id="8ECA9085FFBECC4CA347FE2BFD74FE38" LSID="urn:lsid:plazi:treatment:03BC7748FFBACC51A1B1FF4DFB6AFEF4:8ECA9085FFBECC4CA347FE2BFD74FE38" box="[609,699,440,464]" municipality="Tympanum" name="Dorsum" pageId="13" pageNumber="340">Dorsum</location>
<collectionCode id="ED045E9BFFBECC4CA3E3FE2BFD38FE38" box="[709,759,440,464]" pageId="13" pageNumber="340">dark</collectionCode>
brown
<collectionCode id="ED045E9BFFBECC4CA277FE2BFC79FE38" box="[849,950,440,464]" pageId="13" pageNumber="340">with two</collectionCode>
black blotches on
<collectionCode id="ED045E9BFFBECC4CA5B7FE2BFB38FE38" box="[1169,1271,440,464]" pageId="13" pageNumber="340">the neck</collectionCode>
; 3 broad
</materialsCitation>
,
<materialsCitation id="3B7DCC03FFBECC4CA44DFE2BFC09FDD7" ID-GBIF-Occurrence="1849038559" collectionCode="dark, the, tail, and, found, The, five, rows, scales, scale, pale, white, from, neck, ends, near" location="The" pageId="13" pageNumber="340" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED045E9BFFBECC4CA44DFE2BFA52FE38" box="[1387,1437,440,464]" pageId="13" pageNumber="340">dark</collectionCode>
brown bars at
<collectionCode id="ED045E9BFFBECC4CA011FE4EFE4BFE1C" box="[311,388,476,500]" pageId="13" pageNumber="340">the tail</collectionCode>
base
<collectionCode id="ED045E9BFFBECC4CA0E2FE4FFE22FE1C" box="[452,493,476,500]" pageId="13" pageNumber="340">and</collectionCode>
multiple smaller bars
<collectionCode id="ED045E9BFFBECC4CA3C1FE4FFCE7FE1C" box="[743,808,476,500]" pageId="13" pageNumber="340">found</collectionCode>
throughout
<collectionCode id="ED045E9BFFBECC4CA294FE4EFBCBFE1C" box="[946,1028,476,500]" pageId="13" pageNumber="340">the tail</collectionCode>
.
<collectionCode id="ED045E9BFFBECC4CA528FE4EFBA3FE1C" box="[1038,1132,477,500]" pageId="13" pageNumber="340">
<location id="8ECA9085FFBECC4CA528FE4EFBF5FE1C" LSID="urn:lsid:plazi:treatment:03BC7748FFBACC51A1B1FF4DFB6AFEF4:8ECA9085FFBECC4CA528FE4EFBF5FE1C" box="[1038,1082,477,500]" name="The" pageId="13" pageNumber="340">The</location>
five
</collectionCode>
enlarged
<collectionCode id="ED045E9BFFBECC4CA5FCFE4FFADFFE1C" box="[1242,1296,476,500]" pageId="13" pageNumber="340">rows</collectionCode>
of
<collectionCode id="ED045E9BFFBECC4CA410FE4FFAB6FE1C" box="[1334,1401,476,500]" pageId="13" pageNumber="340">scales</collectionCode>
on
<collectionCode id="ED045E9BFFBECC4CA1B1FD93FF76FDFF" box="[151,185,512,535]" pageId="13" pageNumber="340">the</collectionCode>
dorsum flanking
<collectionCode id="ED045E9BFFBECC4CA0A4FD93FE6BFDFF" box="[386,420,512,535]" pageId="13" pageNumber="340">the</collectionCode>
vertebral
<collectionCode id="ED045E9BFFBECC4CA331FD93FD08FDF0" box="[535,711,512,536]" pageId="13" pageNumber="340">scale rows dark</collectionCode>
brown coloured, a
<collectionCode id="ED045E9BFFBECC4CA285FD93FC1DFDF0" box="[931,978,512,536]" pageId="13" pageNumber="340">pale</collectionCode>
brown broken
<collectionCode id="ED045E9BFFBECC4CA558FD93FB73FDFF" box="[1150,1212,512,535]" pageId="13" pageNumber="340">white</collectionCode>
stripe starting
<collectionCode id="ED045E9BFFBECC4CA440FD93FF03FDD7" pageId="13" pageNumber="340">from neck</collectionCode>
bordering enlarged
<collectionCode id="ED045E9BFFBECC4CA088FDB4FE29FDD7" box="[430,486,551,575]" pageId="13" pageNumber="340">scale</collectionCode>
row on
<collectionCode id="ED045E9BFFBECC4CA362FDBBFDA9FDD7" box="[580,614,552,575]" pageId="13" pageNumber="340">the</collectionCode>
dorsum
<collectionCode id="ED045E9BFFBECC4CA3EFFDB4FC46FDD7" box="[713,905,551,575]" pageId="13" pageNumber="340">ends near the tail</collectionCode>
base
</materialsCitation>
,
<materialsCitation id="3B7DCC03FFBECC4CA2F6FDBBFA86FD43" ID-GBIF-Occurrence="1849038458" collectionCode="the, dark, with, mixed, light, scales, lower, paler, mottled, white, Brown, present, dorsal, side, Tail, dewlap, spots, some, spot" location="Enlarged" municipality="Venter" pageId="13" pageNumber="340" specimenCode="upper 3" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBB36E25FFBECC4CA2F6FDBBFBECFDD7" box="[976,1059,551,575]" collectionCode="upper" pageId="13" pageNumber="340">upper 3</specimenCode>
/4th of
<collectionCode id="ED045E9BFFBECC4CA54BFDBBFB40FDD7" box="[1133,1167,552,575]" pageId="13" pageNumber="340">the</collectionCode>
flanks
<collectionCode id="ED045E9BFFBECC4CA5C4FDB4FADBFDD7" box="[1250,1300,551,575]" pageId="13" pageNumber="340">dark</collectionCode>
brown
<collectionCode id="ED045E9BFFBECC4CA44CFDBBFED8FD8B" pageId="13" pageNumber="340">with mixed light</collectionCode>
brown
<collectionCode id="ED045E9BFFBECC4CA04AFDD8FE7CFD8B" box="[364,435,587,611]" pageId="13" pageNumber="340">scales</collectionCode>
,
<collectionCode id="ED045E9BFFBECC4CA09BFDD8FDE9FD8B" box="[445,550,587,611]" pageId="13" pageNumber="340">the lower</collectionCode>
end of flanks
<collectionCode id="ED045E9BFFBECC4CA3E2FDD8FD30FD8B" box="[708,767,587,611]" pageId="13" pageNumber="340">paler</collectionCode>
.
<collectingMunicipality id="6BCE5C24FFBECC4CA22EFDD8FC9EFD8A" box="[776,849,587,610]" pageId="13" pageNumber="340">Venter</collectingMunicipality>
<collectionCode id="ED045E9BFFBECC4CA27EFDD8FC28FD8A" box="[856,999,587,611]" pageId="13" pageNumber="340">mottled with</collectionCode>
brown on off
<collectionCode id="ED045E9BFFBECC4CA5A3FDD8FADDFD8B" box="[1157,1298,587,611]" pageId="13" pageNumber="340">white scales</collectionCode>
.
<collectionCode id="ED045E9BFFBECC4CA43CFDD8FAA9FD8B" box="[1306,1382,587,611]" pageId="13" pageNumber="340">Brown</collectionCode>
bars
<collectionCode id="ED045E9BFFBECC4CA1B1FDFCFF27FD6F" box="[151,232,623,647]" pageId="13" pageNumber="340">present</collectionCode>
on
<collectionCode id="ED045E9BFFBECC4CA033FDE3FE76FD6F" box="[277,441,623,647]" pageId="13" pageNumber="340">the dorsal side</collectionCode>
of limbs.
<collectionCode id="ED045E9BFFBECC4CA309FDFCFD01FD6F" box="[559,718,623,647]" pageId="13" pageNumber="340">Tail with dark</collectionCode>
brown bands throughout.
<location id="8ECA9085FFBECC4CA2DCFDFCFB91FD6F" LSID="urn:lsid:plazi:treatment:03BC7748FFBACC51A1B1FF4DFB6AFEF4:8ECA9085FFBECC4CA2DCFDFCFB91FD6F" box="[1018,1118,623,647]" municipality="Venter" name="Enlarged" pageId="13" pageNumber="340">Enlarged</location>
<collectionCode id="ED045E9BFFBECC4CA541FDFCFB65FD6F" box="[1127,1194,623,647]" pageId="13" pageNumber="340">scales</collectionCode>
on
<collectionCode id="ED045E9BFFBECC4CA5F1FDFCFA52FD6F" box="[1239,1437,623,647]" pageId="13" pageNumber="340">dewlap with dark</collectionCode>
brown/black
<collectionCode id="ED045E9BFFBECC4CA00FFD00FEA8FD43" box="[297,359,659,683]" pageId="13" pageNumber="340">spots</collectionCode>
,
<collectionCode id="ED045E9BFFBECC4CA057FD00FE04FD43" box="[369,459,659,683]" pageId="13" pageNumber="340">mottled</collectionCode>
,
<collectionCode id="ED045E9BFFBECC4CA0F2FD00FD5EFD43" box="[468,657,659,683]" pageId="13" pageNumber="340">with some scales</collectionCode>
having
<collectionCode id="ED045E9BFFBECC4CA3CDFD00FCD2FD43" box="[747,797,659,683]" pageId="13" pageNumber="340">dark</collectionCode>
brown/black
<collectionCode id="ED045E9BFFBECC4CA290FD00FBE7FD42" box="[950,1064,659,683]" pageId="13" pageNumber="340">spots with</collectionCode>
a
<collectionCode id="ED045E9BFFBECC4CA564FD00FB7AFD43" box="[1090,1205,659,683]" pageId="13" pageNumber="340">white spot</collectionCode>
in
<collectionCode id="ED045E9BFFBECC4CA5FEFD00FB35FD42" box="[1240,1274,659,682]" pageId="13" pageNumber="340">the</collectionCode>
center
</materialsCitation>
.
<materialsCitation id="3B7DCC03FFBECC4CA474FD00FED4FCFF" ID-GBIF-Occurrence="1849038465" collectionCode="region, side, the, scale, dewlap, white, with, dark, gray, and, scales, after, its, rest, are, faint" location="mentum" pageId="13" pageNumber="340" specimenCount="1" stateProvince="mentum" typeStatus="holotype">
Throat
<collectionCode id="ED045E9BFFBECC4CA1B1FD24FF10FD27" box="[151,223,695,719]" pageId="13" pageNumber="340">region</collectionCode>
on either
<collectionCode id="ED045E9BFFBECC4CA074FD24FEB0FD27" box="[338,383,695,719]" pageId="13" pageNumber="340">side</collectionCode>
of
<collectionCode id="ED045E9BFFBECC4CA080FD2BFE07FD27" box="[422,456,696,719]" pageId="13" pageNumber="340">the</collectionCode>
enlarged
<collectionCode id="ED045E9BFFBECC4CA31EFD24FDBFFD27" box="[568,624,695,719]" pageId="13" pageNumber="340">scale</collectionCode>
on
<collectionCode id="ED045E9BFFBECC4CA3BAFD24FD22FD27" box="[668,749,695,719]" pageId="13" pageNumber="340">dewlap</collectionCode>
off
<collectionCode id="ED045E9BFFBECC4CA23BFD2BFC5AFD27" box="[797,917,696,719]" pageId="13" pageNumber="340">white with</collectionCode>
bluish
<collectionCode id="ED045E9BFFBECC4CA2CFFD24FB9AFD27" box="[1001,1109,695,719]" pageId="13" pageNumber="340">dark gray</collectionCode>
/black speckles
<collectionCode id="ED045E9BFFBECC4CA436FD24FA52FD27" box="[1296,1437,695,719]" pageId="13" pageNumber="340">and the dark</collectionCode>
blue colouration on
<collectionCode id="ED045E9BFFBECC4CA05DFD48FE52FD1A" box="[379,413,731,754]" pageId="13" pageNumber="340">the</collectionCode>
throat is prominent 5
<collectionCode id="ED045E9BFFBECC4CA3BDFD48FC8CFD1A" box="[667,835,731,755]" pageId="13" pageNumber="340">scales after the</collectionCode>
<collectingRegion id="49D108BCFFBECC4CA26DFD48FC68FD1A" box="[843,935,731,754]" country="Myanmar" name="Chin" pageId="13" pageNumber="340">mentum</collectingRegion>
<collectionCode id="ED045E9BFFBECC4CA289FD48FC35FD1B" box="[943,1018,731,755]" pageId="13" pageNumber="340">and its</collectionCode>
surrounding
<collectionCode id="ED045E9BFFBECC4CA5B4FD48FB16FD1B" box="[1170,1241,731,755]" pageId="13" pageNumber="340">scales</collectionCode>
,
<collectionCode id="ED045E9BFFBECC4CA5C2FD48FA53FD1B" box="[1252,1436,731,755]" pageId="13" pageNumber="340">the rest are faint</collectionCode>
bluish
<collectionCode id="ED045E9BFFBECC4CA1C4FD6CFED8FCFF" box="[226,279,767,791]" pageId="13" pageNumber="340">gray</collectionCode>
.
</materialsCitation>
</paragraph>
<caption id="DF6A96D6FFBECC4CA1B1F936FAC3F932" httpUri="https://zenodo.org/record/1290604/files/figure.png" pageId="13" pageNumber="340" startId="13.[151,249,1701,1723]" targetBox="[157,1428,848,1659]" targetPageId="13">
<paragraph id="8BAAC65EFFBECC4CA1B1F936FAC3F932" blockId="13.[151,1436,1701,1754]" pageId="13" pageNumber="340">
<emphasis id="B9611A4CFFBECC4CA1B1F936FEECF953" bold="true" box="[151,291,1701,1723]" pageId="13" pageNumber="340">FIGURE 10.</emphasis>
Box plot showing proportion of dewlap in trunk. Note red circle highlighing the southern most population (Bagalkot &amp; Tummargudi) of
<taxonomicName id="4C15BDDDFFBECC4CA0EEF956FDFFF932" box="[456,560,1732,1754]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="340" phylum="Chordata" rank="species" species="laticeps">
<emphasis id="B9611A4CFFBECC4CA0EEF956FDFFF932" box="[456,560,1732,1754]" italics="true" pageId="13" pageNumber="340">S. laticeps</emphasis>
</taxonomicName>
which have relatively larger dewlaps compared to all other populations.
</paragraph>
</caption>
<paragraph id="8BAAC65EFFBECC4FA1E1F894FC0FFE8D" blockId="13.[151,1438,1799,2004]" lastBlockId="14.[151,1438,151,357]" lastPageId="14" lastPageNumber="341" pageId="13" pageNumber="340">
<emphasis id="B9611A4CFFBECC4CA1E1F894FE15F8C8" bold="true" box="[199,474,1799,1824]" pageId="13" pageNumber="340">Variation in paratypes.</emphasis>
The two paratypes agree with the holotype in overall scalation with some exceptions. CES
<date id="FFABE09EFFBECC4CA1F2F8B8FEE8F8AC" box="[212,295,1835,1860]" pageId="13" pageNumber="340" value="36-11-14">141136</date>
has 101 ventral scales and 32 belly ventral scales, 58 mid-body scales, 49 dorsal scales, one less infralabial on each side and one additional supralabial on the left. The female BNHS 2491 agrees with the holotype in overall scalation except that it lacks a dewlap, and has 67 ventral scales, one less infralabial on each side. CES
<date id="FFABE09EFFBECC4CA1B1F804FF25F858" box="[151,234,1943,1968]" pageId="13" pageNumber="340" value="36-11-14">141136</date>
differs in colouration from the holotype in having a light brown dorsum (an artefact of preservation),
<materialsCitation id="3B7DCC03FFBECC4CA44BF804FB65F83C" ID-GBIF-Occurrence="1849038560" collectionCode="four, dark, the, first, one, neck" location="No" pageId="13" pageNumber="340" specimenCount="1">
<collectionCode id="ED045E9BFFBECC4CA44BF804FA53F847" box="[1389,1436,1943,1967]" pageId="13" pageNumber="340">four</collectionCode>
distinct
<collectionCode id="ED045E9BFFBECC4CA1D7F82FFEECF83C" box="[241,291,1980,2004]" pageId="13" pageNumber="340">dark</collectionCode>
brown patches on
<collectionCode id="ED045E9BFFBECC4CA0DCF82FFDD3F83B" box="[506,540,1980,2003]" pageId="13" pageNumber="340">the</collectionCode>
dorsum
<collectionCode id="ED045E9BFFBECC4CA3A6F82FFCC8F83C" box="[640,775,1980,2004]" pageId="13" pageNumber="340">the first one</collectionCode>
darker.
<location id="8ECA9085FFBECC4CA242F82FFC49F83C" LSID="urn:lsid:plazi:treatment:03BC7748FFBACC51A1B1FF4DFB6AFEF4:8ECA9085FFBECC4CA242F82FFC49F83C" box="[868,902,1980,2004]" name="No" pageId="13" pageNumber="340">No</location>
blue patches on
<collectionCode id="ED045E9BFFBECC4CA560F82FFB65F83C" box="[1094,1194,1980,2004]" pageId="13" pageNumber="340">the neck</collectionCode>
</materialsCitation>
. The stripe below the eye, indistinguishable compared to the surrounding scale, only a short light brown stripe ends near the forearm insertion. Blue line on the dewlap/throat darker than the holotype. Broad bands are present throughout the tail. BNHS 2491 differs in colouration from the holotype in having head colouration similar to body, three rhomboidal/ oval patterns on the dorsum, pale white stripe on the dorsum absent. Head brown with patches of brick red and grey (likely due to shedding scales), no blue patches on the neck. Throat region pale white in colour with dark black spots and speckles. One rhomboidal marking on the dorsum of tail base.
</paragraph>
<caption id="DF6A96D6FFBDCC4FA1B1F8C1FE07F86E" httpUri="https://zenodo.org/record/1290606/files/figure.png" pageId="14" pageNumber="341" startId="14.[151,249,1874,1896]" targetBox="[151,1436,409,1860]" targetPageId="14">
<paragraph id="8BAAC65EFFBDCC4FA1B1F8C1FE07F86E" blockId="14.[151,1436,1874,1926]" pageId="14" pageNumber="341">
<emphasis id="B9611A4CFFBDCC4FA1B1F8C1FED1F88F" bold="true" box="[151,286,1874,1896]" pageId="14" pageNumber="341">FIGURE 11.</emphasis>
Hemipenis of
<taxonomicName id="4C15BDDDFFBDCC4FA091F8C1FDA1F880" authority="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth, 2018" authorityName="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth" authorityYear="2018" box="[439,622,1874,1896]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="341" phylum="Chordata" rank="species" species="gokakensis" status="sp. nov.">
<emphasis id="B9611A4CFFBDCC4FA091F8C1FDA1F880" box="[439,622,1874,1896]" italics="true" pageId="14" pageNumber="341">Sitana gokakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A252A737FFBDCC4FA350F8C1FD08F88F" box="[630,711,1874,1895]" pageId="14" pageNumber="341" rank="species">
<emphasis id="B9611A4CFFBDCC4FA350F8C1FD08F88F" bold="true" box="[630,711,1874,1895]" pageId="14" pageNumber="341">sp. nov.</emphasis>
</taxonomicNameLabel>
(CES 141136): (A) sulcal view; (B) asulcal view; C) lateral view; (D) apical view. Scale bar:10 mm.
</paragraph>
</caption>
<paragraph id="8BAAC65EFFBDCC4FA1E1F820FA52F823" blockId="14.[151,1437,1971,2032]" box="[199,1437,1971,1996]" pageId="14" pageNumber="341">
<emphasis id="B9611A4CFFBDCC4FA1E1F820FDE3F824" bold="true" box="[199,556,1971,1996]" pageId="14" pageNumber="341">Variation in live colouration.</emphasis>
Most of the male specimens collected during this study matched with the
</paragraph>
</subSubSection>
<subSubSection id="C30F95D5FFBDCC4EA1B1F84BFBDAFE60" lastPageId="15" lastPageNumber="342" pageId="14" pageNumber="341" type="materials_examined">
<paragraph id="8BAAC65EFFBDCC4FA1B1F84BFA53F818" blockId="14.[151,1437,1971,2032]" box="[151,1436,2007,2032]" pageId="14" pageNumber="341">
<collectionCode id="ED045E9BFFBDCC4FA1B1F84BFF36F818" box="[151,249,2008,2032]" pageId="14" pageNumber="341">
<typeStatus id="54AE78FCFFBDCC4FA1B1F84BFF36F818" box="[151,249,2008,2032]" pageId="14" pageNumber="341" type="holotype">holotype</typeStatus>
</collectionCode>
in
<collectionCode id="ED045E9BFFBDCC4FA004F84BFE83F807" box="[290,332,2008,2031]" pageId="14" pageNumber="341">live</collectionCode>
colouration.
<collectionCode id="ED045E9BFFBDCC4FA0C3F84BFD65F818" box="[485,682,2008,2032]" pageId="14" pageNumber="341">There were some</collectionCode>
specimens
<collectionCode id="ED045E9BFFBDCC4FA214F84BFAC3F818" box="[818,1292,2008,2032]" pageId="14" pageNumber="341">which were different from the rest which</collectionCode>
we describe
</paragraph>
<paragraph id="8BAAC65EFFBCCC4EA1B1FF0BFBDAFE60" blockId="15.[151,1438,151,933]" pageId="15" pageNumber="342">
herein.
<collectionCode id="ED045E9BFFBCCC4EA1CCFF0BFEE4FF47" box="[234,299,152,175]" pageId="15" pageNumber="342">Three</collectionCode>
<collectionCode id="ED045E9BFFBCCC4EA014FF0BFE99FF58" box="[306,342,152,176]" country="Japan" httpUri="http://grbio.org/cool/ybcf-zh2i" name="Department of Biotechnology" pageId="15" pageNumber="342">out</collectionCode>
of
<collectionCode id="ED045E9BFFBCCC4EA05DFF0BFE09FF47" box="[379,454,152,175]" pageId="15" pageNumber="342">the ten</collectionCode>
breeding
<collectionCode id="ED045E9BFFBCCC4EA313FF0BFCFBFF47" box="[565,820,152,176]" pageId="15" pageNumber="342">males were found with</collectionCode>
blue colouration on
<collectionCode id="ED045E9BFFBCCC4EA53CFF0BFB4CFF47" box="[1050,1155,152,175]" pageId="15" pageNumber="342">the upper</collectionCode>
eyelids.
<collectionCode id="ED045E9BFFBCCC4EA5C0FF0BFAE4FF58" box="[1254,1323,152,176]" pageId="15" pageNumber="342">Seven</collectionCode>
<collectionCode id="ED045E9BFFBCCC4EA417FF0BFA9AFF58" box="[1329,1365,152,176]" country="Japan" httpUri="http://grbio.org/cool/ybcf-zh2i" name="Department of Biotechnology" pageId="15" pageNumber="342">out</collectionCode>
of
<collectionCode id="ED045E9BFFBCCC4EA45CFF0BFECBFF3C" pageId="15" pageNumber="342">the ten males</collectionCode>
observed in-situ
<collectionCode id="ED045E9BFFBCCC4EA0F6FF2FFE37FF3C" box="[464,504,188,212]" pageId="15" pageNumber="342">had</collectionCode>
mostly
<collectionCode id="ED045E9BFFBCCC4EA37FFF2FFCF4FF3C" box="[601,827,188,212]" pageId="15" pageNumber="342">dark pink and tinge</collectionCode>
of blue on
<collectionCode id="ED045E9BFFBCCC4EA2E1FF2EFC26FF3C" box="[967,1001,189,212]" pageId="15" pageNumber="342">the</collectionCode>
nuchal
<collectionCode id="ED045E9BFFBCCC4EA56FFF2FFB5AFF3C" box="[1097,1173,188,212]" pageId="15" pageNumber="342">region</collectionCode>
.
<collectionCode id="ED045E9BFFBCCC4EA587FF2EFA86FF3C" box="[1185,1353,188,212]" pageId="15" pageNumber="342">The fifth scale</collectionCode>
on
<collectionCode id="ED045E9BFFBCCC4EA45CFF2EFA53FF3C" box="[1402,1436,189,212]" pageId="15" pageNumber="342">the</collectionCode>
enlarged
<collectionCode id="ED045E9BFFBCCC4EA1D8FF73FEBCFF10" box="[254,371,224,248]" pageId="15" pageNumber="342">scale rows</collectionCode>
flanking vertebral
<collectionCode id="ED045E9BFFBCCC4EA36CFF73FD42FF10" box="[586,653,224,248]" pageId="15" pageNumber="342">scales</collectionCode>
on
<collectionCode id="ED045E9BFFBCCC4EA391FF73FD16FF1F" box="[695,729,224,247]" pageId="15" pageNumber="342">the</collectionCode>
dorsum
<collectionCode id="ED045E9BFFBCCC4EA21AFF73FCA7FF10" box="[828,872,224,248]" pageId="15" pageNumber="342">was</collectionCode>
bordered
<collectionCode id="ED045E9BFFBCCC4EA2FFFF73FBC4FF1F" box="[985,1035,224,247]" pageId="15" pageNumber="342">with</collectionCode>
continuous
<collectionCode id="ED045E9BFFBCCC4EA5B2FF73FB16FF10" box="[1172,1241,224,248]" pageId="15" pageNumber="342">cream</collectionCode>
coloured/reddish stripe starting
<collectionCode id="ED045E9BFFBCCC4EA01AFE97FD8FFEF4" box="[316,576,260,284]" pageId="15" pageNumber="342">from neck till the vent</collectionCode>
(e.g, see
<figureCitation id="132EDADBFFBCCC4EA395FE97FCF3FEF5" box="[691,828,260,285]" captionStart="FIGURE 9" captionStartId="11.[151,249,1976,1998]" captionTargetBox="[193,1398,214,1932]" captionTargetId="figure@11.[170,1417,193,1955]" captionTargetPageId="11" captionText="FIGURE 9. Sitana gokakensis sp. nov. colouration in life. A. An adult male from type locality, B. paratype BNHS 2491, C &amp; D. two different uncollected adult females from the type locality." httpUri="https://zenodo.org/record/1290602/files/figure.png" pageId="15" pageNumber="342">Fig. 9 a,b,d</figureCitation>
), in
<collectionCode id="ED045E9BFFBCCC4EA257FE97FBB9FEF4" box="[881,1142,260,284]" pageId="15" pageNumber="342">some individuals these</collectionCode>
stripes
<collectionCode id="ED045E9BFFBCCC4EA5F4FE96FAC7FEF4" box="[1234,1288,261,284]" pageId="15" pageNumber="342">were</collectionCode>
broken.
<collectionCode id="ED045E9BFFBCCC4EA457FE96FA52FEF4" box="[1393,1437,261,284]" pageId="15" pageNumber="342">The</collectionCode>
rhomboidal
<collectionCode id="ED045E9BFFBCCC4EA004FEBBFE90FEA8" box="[290,351,296,320]" pageId="15" pageNumber="342">patch</collectionCode>
on
<collectionCode id="ED045E9BFFBCCC4EA0A9FEBBFDE8FEA8" box="[399,551,296,320]" pageId="15" pageNumber="342">the neck was</collectionCode>
single in
<collectionCode id="ED045E9BFFBCCC4EA3B8FEBBFCACFEA8" box="[670,867,296,320]" pageId="15" pageNumber="342">many individuals</collectionCode>
but
<collectionCode id="ED045E9BFFBCCC4EA2BDFEBBFC72FED7" box="[923,957,296,319]" pageId="15" pageNumber="342">the</collectionCode>
5 rhomboid patches on
<collectionCode id="ED045E9BFFBCCC4EA5FDFEBBFB32FED7" box="[1243,1277,296,319]" pageId="15" pageNumber="342">the</collectionCode>
dorsum
<collectionCode id="ED045E9BFFBCCC4EA440FEBBFA53FED7" box="[1382,1436,296,319]" pageId="15" pageNumber="342">were</collectionCode>
separated by a
<collectionCode id="ED045E9BFFBCCC4EA019FEDFFE7AFE8C" box="[319,437,332,356]" pageId="15" pageNumber="342">cream line</collectionCode>
on
<collectionCode id="ED045E9BFFBCCC4EA0F9FEDEFDCEFE8C" box="[479,513,333,356]" pageId="15" pageNumber="342">the</collectionCode>
vertebral
<collectionCode id="ED045E9BFFBCCC4EA355FEDFFD74FE8C" box="[627,699,332,356]" pageId="15" pageNumber="342">region</collectionCode>
(e.g, see
<figureCitation id="132EDADBFFBCCC4EA201FEDFFCB6FE8C" box="[807,889,332,356]" captionStart="FIGURE 9" captionStartId="11.[151,249,1976,1998]" captionTargetBox="[193,1398,214,1932]" captionTargetId="figure@11.[170,1417,193,1955]" captionTargetPageId="11" captionText="FIGURE 9. Sitana gokakensis sp. nov. colouration in life. A. An adult male from type locality, B. paratype BNHS 2491, C &amp; D. two different uncollected adult females from the type locality." httpUri="https://zenodo.org/record/1290602/files/figure.png" pageId="15" pageNumber="342">Fig. 9c</figureCitation>
).
<collectionCode id="ED045E9BFFBCCC4EA2AAFEDFFC74FE8C" box="[908,955,332,356]" pageId="15" pageNumber="342">One</collectionCode>
of
<collectionCode id="ED045E9BFFBCCC4EA2C7FEDEFB75FE8C" box="[993,1210,332,356]" pageId="15" pageNumber="342">the adult males had</collectionCode>
blue colouration on
<collectionCode id="ED045E9BFFBCCC4EA1B1FEE3FEE0FE60" box="[151,303,368,392]" pageId="15" pageNumber="342">the tail which</collectionCode>
became prominent immediately
<collectionCode id="ED045E9BFFBCCC4EA386FEE3FD1BFE60" box="[672,724,368,392]" pageId="15" pageNumber="342">after</collectionCode>
euthanization (Appendix 8).
</paragraph>
</subSubSection>
<subSubSection id="C30F95D5FFBCCC4EA1E1FE07FA96FD24" pageId="15" pageNumber="342" type="description">
<paragraph id="8BAAC65EFFBCCC4EA1E1FE07FA96FD24" blockId="15.[151,1438,151,933]" pageId="15" pageNumber="342">
<emphasis id="B9611A4CFFBCCC4EA1E1FE07FE21FE45" bold="true" box="[199,494,404,429]" pageId="15" pageNumber="342">Hemipenial morphology.</emphasis>
Four samples were examined (CES
<date id="FFABE09EFFBCCC4EA2A0FE07FC14FE45" box="[902,987,404,429]" pageId="15" pageNumber="342" value="36-11-14">141136</date>
<date id="FFABE09EFFBCCC4EA2CCFE07FBF0FE45" box="[1002,1087,404,429]" pageId="15" pageNumber="342" value="39-11-14">141139</date>
). Hemipenis bilobed, relatively small, longer than wide and shallowly forked. Sulcus spermaticus bifurcated and the fork continues onto the apical lobes (
<figureCitation id="132EDADBFFBCCC4EA1C4FE4FFE89FE1C" box="[226,326,476,500]" captionStart="FIGURE 11" captionStartId="14.[151,249,1874,1896]" captionTargetBox="[151,1436,409,1860]" captionTargetId="figure@14.[151,1436,409,1860]" captionTargetPageId="14" captionText="FIGURE 11. Hemipenis of Sitana gokakensis sp. nov. (CES 141136): (A) sulcal view; (B) asulcal view; C) lateral view; (D) apical view. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290606/files/figure.png" pageId="15" pageNumber="342">Fig. 11a</figureCitation>
). Sulcal lips raised and papillate, sulcus smooth originating from the side of the base. Apex with small serrated row of calyces and the sulcal region of apex nude (
<figureCitation id="132EDADBFFBCCC4EA2C0FD93FB9DFDF0" box="[998,1106,512,536]" captionStart="FIGURE 11" captionStartId="14.[151,249,1874,1896]" captionTargetBox="[151,1436,409,1860]" captionTargetId="figure@14.[151,1436,409,1860]" captionTargetPageId="14" captionText="FIGURE 11. Hemipenis of Sitana gokakensis sp. nov. (CES 141136): (A) sulcal view; (B) asulcal view; C) lateral view; (D) apical view. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290606/files/figure.png" pageId="15" pageNumber="342">Fig. 11d</figureCitation>
), medial projection absent. Ornamentation is differentiated and combination of flounces and calyces observed. Papillae present between the apical lobes. Apical regions on the lobes of sulcal side calyces are serrated and continuous; calyces are relatively larger and non-serrated at the base of the lobes (
<figureCitation id="132EDADBFFBCCC4EA39DFDFFFCFBFD6D" box="[699,820,620,645]" captionStart="FIGURE 11" captionStartId="14.[151,249,1874,1896]" captionTargetBox="[151,1436,409,1860]" captionTargetId="figure@14.[151,1436,409,1860]" captionTargetPageId="14" captionText="FIGURE 11. Hemipenis of Sitana gokakensis sp. nov. (CES 141136): (A) sulcal view; (B) asulcal view; C) lateral view; (D) apical view. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290606/files/figure.png" pageId="15" pageNumber="342">Fig. 11a,b</figureCitation>
). Calyces are deep regular pits on the asulcal side and become shallow at basal region (
<figureCitation id="132EDADBFFBCCC4EA335FD03FDB7FD40" box="[531,632,655,680]" captionStart="FIGURE 11" captionStartId="14.[151,249,1874,1896]" captionTargetBox="[151,1436,409,1860]" captionTargetId="figure@14.[151,1436,409,1860]" captionTargetPageId="14" captionText="FIGURE 11. Hemipenis of Sitana gokakensis sp. nov. (CES 141136): (A) sulcal view; (B) asulcal view; C) lateral view; (D) apical view. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290606/files/figure.png" pageId="15" pageNumber="342">Fig. 11b</figureCitation>
). Ridges between the calyces are thin and show micro-ornamentation which are scalloped. Flounces present, eight to ten in number, all prominent on the asulcal side (
<figureCitation id="132EDADBFFBCCC4EA5EAFD27FA83FD24" box="[1228,1356,692,716]" captionStart="FIGURE 11" captionStartId="14.[151,249,1874,1896]" captionTargetBox="[151,1436,409,1860]" captionTargetId="figure@14.[151,1436,409,1860]" captionTargetPageId="14" captionText="FIGURE 11. Hemipenis of Sitana gokakensis sp. nov. (CES 141136): (A) sulcal view; (B) asulcal view; C) lateral view; (D) apical view. Scale bar:10 mm." httpUri="https://zenodo.org/record/1290606/files/figure.png" pageId="15" pageNumber="342">Fig. 11B,C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30F95D5FFBCCC4EA1E1FD44FCE2FCD0" pageId="15" pageNumber="342" type="etymology">
<paragraph id="8BAAC65EFFBCCC4EA1E1FD44FD6AFCFC" blockId="15.[151,1438,151,933]" pageId="15" pageNumber="342">
<emphasis id="B9611A4CFFBCCC4EA1E1FD44FE81FD18" bold="true" box="[199,334,727,752]" pageId="15" pageNumber="342">Etymology.</emphasis>
The specific epithet is an adjectival toponym and refers to the Gokak plateau of Belagavi district in
<collectingRegion id="49D108BCFFBCCC4EA192FD6FFEE3FCFC" box="[180,300,764,788]" country="India" name="Karnataka" pageId="15" pageNumber="342">Karnataka</collectingRegion>
, to which this species is endemic.
</paragraph>
<paragraph id="8BAAC65EFFBCCC4EA1E1FC8CFCE2FCD0" blockId="15.[151,1438,151,933]" box="[199,813,799,824]" pageId="15" pageNumber="342">
<emphasis id="B9611A4CFFBCCC4EA1E1FC8CFE33FCD0" bold="true" box="[199,508,799,824]" pageId="15" pageNumber="342">Suggested common name.</emphasis>
Gokak fan-throated lizard.
</paragraph>
</subSubSection>
<subSubSection id="C30F95D5FFBCCC4EA1E1FCD7FA52F803" pageId="15" pageNumber="342" type="distribution">
<paragraph id="8BAAC65EFFBCCC4EA1E1FCD7FB9AFC4C" blockId="15.[151,1438,151,933]" pageId="15" pageNumber="342">
<emphasis id="B9611A4CFFBCCC4EA1E1FCD7FEAFFCB5" bold="true" box="[199,352,836,861]" pageId="15" pageNumber="342">Distribution.</emphasis>
<taxonomicName id="4C15BDDDFFBCCC4EA041FCD7FDFEFCB4" authority="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth, 2018" authorityName="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth" authorityYear="2018" box="[359,561,836,860]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="342" phylum="Chordata" rank="species" species="gokakensis" status="sp. nov.">
<emphasis id="B9611A4CFFBCCC4EA041FCD7FDFEFCB4" box="[359,561,836,860]" italics="true" pageId="15" pageNumber="342">Sitana gokakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A252A737FFBCCC4EA31FFCD7FD5CFCB5" box="[569,659,836,861]" pageId="15" pageNumber="342" rank="species">
<emphasis id="B9611A4CFFBCCC4EA31FFCD7FD5CFCB5" bold="true" box="[569,659,836,861]" pageId="15" pageNumber="342">sp. nov.</emphasis>
</taxonomicNameLabel>
is endemic to Gokak plateau in Belagavi district,
<collectingRegion id="49D108BCFFBCCC4EA5E0FCD7FAF6FCB4" box="[1222,1337,836,860]" country="India" name="Karnataka" pageId="15" pageNumber="342">Karnataka</collectingRegion>
(
<figureCitation id="132EDADBFFBCCC4EA46EFCD7FA44FCB4" box="[1352,1419,836,860]" captionStart="FIGURE 1" captionStartId="1.[151,249,1950,1972]" captionTargetBox="[182,1403,757,1910]" captionTargetId="figure@1.[151,1436,735,1928]" captionTargetPageId="1" captionText="FIGURE 1. Map showing sampling localities where we found different Sitana spp. during this study. Rivers and hills referred in this study are labelled. Locality where the two new species were collected are marked in red (S. gokakensis sp. nov.) and blue (S. thondalu sp. nov.). Localities referred in table 1." httpUri="https://zenodo.org/record/1290586/files/figure.png" pageId="15" pageNumber="342">Fig. 1</figureCitation>
). The samples collected from north and south outside this plateau (Nipani, Ramdurga, Bagalkot) were of
<taxonomicName id="4C15BDDDFFBCCC4EA404FCFBFA5AFC68" box="[1314,1429,872,896]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="342" phylum="Chordata" rank="species" species="laticeps">
<emphasis id="B9611A4CFFBCCC4EA404FCFBFA5AFC68" box="[1314,1429,872,896]" italics="true" pageId="15" pageNumber="342">S. laticeps</emphasis>
</taxonomicName>
. The Ghataprabha is a rocky river and a potential geographical barrier for this species.
</paragraph>
<caption id="DF6A96D6FFBCCC4EA1B1F8D4FBE4F8B5" box="[151,1067,1863,1885]" httpUri="https://zenodo.org/record/1290608/files/figure.png" pageId="15" pageNumber="342" startId="15.[151,249,1863,1885]" targetBox="[151,1436,1149,1842]" targetPageId="15">
<paragraph id="8BAAC65EFFBCCC4EA1B1F8D4FBE4F8B5" blockId="15.[151,1067,1863,1885]" box="[151,1067,1863,1885]" pageId="15" pageNumber="342">
<emphasis id="B9611A4CFFBCCC4EA1B1F8D4FED0F8B4" bold="true" box="[151,287,1863,1885]" pageId="15" pageNumber="342">FIGURE 12.</emphasis>
Habitat of
<taxonomicName id="4C15BDDDFFBCCC4EA0B7F8DBFD88F8B5" authority="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth, 2018" authorityName="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth" authorityYear="2018" box="[401,583,1863,1885]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="342" phylum="Chordata" rank="species" species="gokakensis" status="sp. nov.">
<emphasis id="B9611A4CFFBCCC4EA0B7F8DBFD88F8B5" box="[401,583,1863,1885]" italics="true" pageId="15" pageNumber="342">Sitana gokakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A252A737FFBCCC4EA368F8DBFD51F8B5" box="[590,670,1864,1885]" pageId="15" pageNumber="342" rank="species">
<emphasis id="B9611A4CFFBCCC4EA368F8DBFD51F8B5" bold="true" box="[590,670,1864,1885]" pageId="15" pageNumber="342">sp. nov.</emphasis>
</taxonomicNameLabel>
at the type locality, Gokak, Karnataka.
</paragraph>
</caption>
<paragraph id="8BAAC65EFFBCCC4EA1E1F819FA52F803" blockId="15.[151,1437,1930,2027]" pageId="15" pageNumber="342">
<emphasis id="B9611A4CFFBCCC4EA1E1F819FDD8F84B" bold="true" box="[199,535,1930,1955]" pageId="15" pageNumber="342">Habitat and natural history.</emphasis>
<taxonomicName id="4C15BDDDFFBCCC4EA338F818FD27F84A" authority="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth, 2018" authorityName="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth" authorityYear="2018" box="[542,744,1930,1954]" class="Reptilia" family="Agamidae" genus="Sitana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="342" phylum="Chordata" rank="species" species="gokakensis" status="sp. nov.">
<emphasis id="B9611A4CFFBCCC4EA338F818FD27F84A" box="[542,744,1930,1954]" italics="true" pageId="15" pageNumber="342">Sitana gokakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A252A737FFBCCC4EA3C9F819FC86F84B" box="[751,841,1930,1955]" pageId="15" pageNumber="342" rank="species">
<emphasis id="B9611A4CFFBCCC4EA3C9F819FC86F84B" bold="true" box="[751,841,1930,1955]" pageId="15" pageNumber="342">sp. nov.</emphasis>
</taxonomicNameLabel>
is only known from the open rocky habitat in Gokak plateau (
<figureCitation id="132EDADBFFBCCC4EA1D0F83CFE84F82F" box="[246,331,1967,1991]" captionStart="FIGURE 12" captionStartId="15.[151,249,1863,1885]" captionTargetBox="[151,1436,1149,1842]" captionTargetId="figure@15.[151,1436,985,1842]" captionTargetPageId="15" captionText="FIGURE 12. Habitat of Sitana gokakensis sp. nov. at the type locality, Gokak, Karnataka." httpUri="https://zenodo.org/record/1290608/files/figure.png" pageId="15" pageNumber="342">Fig. 12</figureCitation>
). Gokak hills receives
<quantity id="4CED6BBBFFBCCC4EA36DF83CFD67F82E" box="[587,680,1967,1991]" metricMagnitude="-1" metricUnit="m" metricValue="8.2" pageId="15" pageNumber="342" unit="mm" value="820.0">820 mm</quantity>
of average annual rainfall. The habitat is xeric and is dominated by thorny and dwarf succulent species (Malpure
<emphasis id="B9611A4CFFBCCC4EA380F840FD16F802" box="[678,729,2002,2026]" italics="true" pageId="15" pageNumber="342">et al</emphasis>
. 2016). The recently described
<taxonomicName id="4C15BDDDFFBCCC4EA56DF841FA88F802" authorityName="Deepak &amp; Khandekar &amp; Chaitanya &amp; Karanth" authorityYear="2018" box="[1099,1351,2002,2026]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageId="15" pageNumber="342" phylum="Magnoliopsida" rank="species" species="gokakensis">
<emphasis id="B9611A4CFFBCCC4EA56DF841FA88F802" box="[1099,1351,2002,2026]" italics="true" pageId="15" pageNumber="342">Euphorbia gokakensis</emphasis>
</taxonomicName>
Yadav,
</paragraph>
</subSubSection>
<subSubSection id="C30F95D5FFA3CC51A1B1FF0BFB6AFEF4" pageId="16" pageNumber="343" type="description">
<paragraph id="8BAAC65EFFA3CC51A1B1FF0BFB6AFEF4" blockId="16.[151,1436,151,284]" pageId="16" pageNumber="343">
Malpure,
<bibRefCitation id="EF84BBAFFFA3CC51A02DFF0BFE0BFF58" box="[267,452,151,176]" pageId="16" pageNumber="343" refId="ref11727" refString="Malpure, N. V., Chandore, A. N. &amp; Yadav, S. R., 2016. Euphorbia gokakensis sp. nov. (Euphorbiaceae) from sandstone formations in Karnataka, India. Nordic Journal of Botany 34, 380 - 383. https: // doi. org / 10.1111 / njb. 00997" type="journal article">Chandore, 2016</bibRefCitation>
is endemic to this plateau which is found alongside other succulents such as
<taxonomicName id="4C15BDDDFFA3CC51A4A2FF0AFED5FF3C" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageId="16" pageNumber="343" phylum="Magnoliopsida" rank="species" species="antiquorum">
<emphasis id="B9611A4CFFA3CC51A4A2FF0AFED5FF3C" italics="true" pageId="16" pageNumber="343">E. antiquorum</emphasis>
</taxonomicName>
L.,
<taxonomicName id="4C15BDDDFFA3CC51A061FF2EFE2BFF3C" authorityName="Haines" authorityYear="1914" box="[327,484,188,212]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageId="16" pageNumber="343" phylum="Magnoliopsida" rank="species" species="caducifolia">
<emphasis id="B9611A4CFFA3CC51A061FF2EFE2BFF3C" box="[327,484,188,212]" italics="true" pageId="16" pageNumber="343">E. caducifolia</emphasis>
</taxonomicName>
Haines and
<taxonomicName id="4C15BDDDFFA3CC51A356FF2FFCD1FF3C" box="[624,798,188,212]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" higherTaxonomySource="CoL" kingdom="Plantae" order="Caryophyllales" pageId="16" pageNumber="343" phylum="Magnoliopsida" rank="species" species="elatior">
<emphasis id="B9611A4CFFA3CC51A356FF2FFCD1FF3C" box="[624,798,188,212]" italics="true" pageId="16" pageNumber="343">Opuntia elatior</emphasis>
</taxonomicName>
Mill. and non-succulents like
<taxonomicName id="4C15BDDDFFA3CC51A555FF2FFA53FF3C" authority="(Willd.)" baseAuthorityName="Willd." box="[1139,1436,188,212]" class="Magnoliopsida" family="Fabaceae" genus="Mundulea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" pageId="16" pageNumber="343" phylum="Tracheophyta" rank="species" species="sericea">
<emphasis id="B9611A4CFFA3CC51A555FF2FFAF2FF3C" box="[1139,1341,188,212]" italics="true" pageId="16" pageNumber="343">Mundulea sericea</emphasis>
(Willd.)
</taxonomicName>
A.Chev. (Malpure
<emphasis id="B9611A4CFFA3CC51A057FF72FE68FF10" box="[369,423,224,248]" italics="true" pageId="16" pageNumber="343">et al</emphasis>
. 2016).
<taxonomicName id="4C15BDDDFFA3CC51A32AFF73FDB9FF10" box="[524,630,224,248]" class="Reptilia" family="Lacertidae" genus="Ophisops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="343" phylum="Chordata" rank="genus">
<emphasis id="B9611A4CFFA3CC51A32AFF73FDB9FF10" box="[524,630,224,248]" italics="true" pageId="16" pageNumber="343">Ophisops</emphasis>
</taxonomicName>
sp and
<taxonomicName id="4C15BDDDFFA3CC51A3FFFF73FB5DFF10" authority="Gleadow, 1887" authorityName="Gleadow" authorityYear="1887" box="[729,1170,224,248]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="343" phylum="Chordata" rank="species" species="murrayi">
<emphasis id="B9611A4CFFA3CC51A3FFFF73FC16FF1F" box="[729,985,224,248]" italics="true" pageId="16" pageNumber="343">Hemidactylus murrayi</emphasis>
Gleadow, 1887
</taxonomicName>
are the two sympatric lizards found in this plateau. Breeding males were recorded during May and August months.
</paragraph>
</subSubSection>
</treatment>
</document>