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<document id="725BDDEC07277663E2D9A31CFF1301D3" ID-CLB-Dataset="8579" ID-DOI="10.11646/zootaxa.4850.1.1" ID-GBIF-Dataset="686337a2-1528-43d0-ae32-68ca023d8df3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4407072" ID-ZooBank="18AA0411-0E54-4922-84C7-608EAC68D281" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609428297648" checkinUser="plazi" docAuthor="Guéorguiev, Borislav &amp; Liang, Hongbin" docDate="2020" docId="03BC5E5B2976FF86FF4BFE4AE93DFA18" docLanguage="en" docName="zootaxa.4850.1.1.pdf" docOrigin="Zootaxa 4850 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Oodes (Oodes) gracilis A. Villa &amp; G. B. Villa 1833" docType="treatment" docVersion="7" lastPageNumber="58" masterDocId="FF8526232940FFBFFFDCFFB4EC2EFFFD" masterDocTitle="Revision of the Palaearctic and Oriental representatives of Lachnocrepis LeConte and Oodes Bonelli (Coleoptera: Carabidae), with special account on Chinese species" masterLastPageNumber="89" masterPageNumber="1" pageNumber="55" updateTime="1698852686469" updateUser="ExternalLinkService">
<mods:mods id="774F9E9272DB51A98B83707A275394C7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="FBB20BE0B4CAACA2690B9514D289C849">Revision of the Palaearctic and Oriental representatives of Lachnocrepis LeConte and Oodes Bonelli (Coleoptera: Carabidae), with special account on Chinese species</mods:title>
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<mods:name id="6FB9129ADC5C717E935E12D3A58E2584" type="personal">
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<mods:namePart id="74BE2B75852B60ABE7837DB5A27888D7">Guéorguiev, Borislav</mods:namePart>
<mods:nameIdentifier id="1DC9D839BADCDDA806CFEEC896C6E0C9" type="ORCID">0000-0002-8532-0657</mods:nameIdentifier>
<mods:nameIdentifier id="D92DD0A50B00210CFC1A23E9DC8B524F" type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
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<mods:namePart id="35528395132FB1CE5FC9BA0979561BE2">Liang, Hongbin</mods:namePart>
<mods:nameIdentifier id="380E53C5C4BEE7F992EC27C0CEA48AE8" type="ORCID">0000-0002-9668-1167</mods:nameIdentifier>
<mods:nameIdentifier id="EBBEBFF89945BEF785274F5ADE67AAED" type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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<mods:title id="9E0727A1E4E41DE5BC1E62709C805293">Zootaxa</mods:title>
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<mods:part id="B59A9817EE3C26C3DA515BF896CD6F8A">
<mods:date id="228B12B43E6FF8914C15703638D5538C">2020</mods:date>
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<mods:number id="C619CE5C20D74FF4D5CA2CD3F7AF8B22">2020-09-08</mods:number>
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<mods:detail id="DAE293AF5E885EC5FA7DCD8586AD62AD" type="volume">
<mods:number id="32CDCC079F5833B50D997140D60DABB7">4850</mods:number>
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<mods:classification id="FFBEC122C5A06318A9AB956B2E755EBA">journal article</mods:classification>
<mods:identifier id="0B6B200A9C71F4779818661014AE3950" type="CLB-Dataset">8579</mods:identifier>
<mods:identifier id="A37E949182819FF974B272A3322EBD91" type="DOI">10.11646/zootaxa.4850.1.1</mods:identifier>
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<treatment id="03BC5E5B2976FF86FF4BFE4AE93DFA18" ID-DOI="http://doi.org/10.5281/zenodo.4480076" ID-GBIF-Taxon="177139763" ID-Zenodo-Dep="4480076" LSID="urn:lsid:plazi:treatment:03BC5E5B2976FF86FF4BFE4AE93DFA18" httpUri="http://treatment.plazi.org/id/03BC5E5B2976FF86FF4BFE4AE93DFA18" lastPageId="57" lastPageNumber="58" pageId="54" pageNumber="55">
<subSubSection id="C30FBCC62976FF89FF4BFE4AEF09FDE5" box="[151,807,510,537]" pageId="54" pageNumber="55" type="nomenclature">
<paragraph id="8BAAEF4D2976FF89FF4BFE4AEF09FDE5" blockId="54.[151,807,510,573]" box="[151,807,510,537]" pageId="54" pageNumber="55">
<emphasis id="B961335F2976FF89FF4BFE4AEF09FDE5" bold="true" box="[151,807,510,537]" pageId="54" pageNumber="55">
17.
<taxonomicName id="4C1594CE2976FF89FF1FFE4AEF09FDE5" ID-CoL="49S3Z" authority="A. Villa &amp; G. B. Villa, 1833" authorityName="A. Villa &amp; G. B. Villa" authorityYear="1833" box="[195,807,510,537]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilis" subGenus="Oodes">
<emphasis id="B961335F2976FF89FF1FFE4AED20FDE4" bold="true" box="[195,270,510,537]" italics="true" pageId="54" pageNumber="55">Oodes</emphasis>
(
<emphasis id="B961335F2976FF89FEC3FE4AED44FDE4" bold="true" box="[287,362,510,537]" italics="true" pageId="54" pageNumber="55">Oodes</emphasis>
)
<emphasis id="B961335F2976FF89FEA7FE4AEDFBFDE4" bold="true" box="[379,469,510,537]" italics="true" pageId="54" pageNumber="55">gracilis</emphasis>
A. Villa &amp; G.B.
<bibRefCitation id="EF8492BC2976FF89FD7EFE4AEF09FDE5" author="Villa, A. &amp; Villa, G. B." box="[674,807,510,536]" pageId="54" pageNumber="55" pagination="32 - 36" refId="ref64409" refString="Villa, A. &amp; Villa, G. B. (1833) Coleopterorum species novae in catalogo dupletorum extantes. Diagnosibus, adumbrationibusatque observationibus illustratae. Coleoptera Europae dupleta in collectione Villa quae pro mutua commutatione offerri possunt, Mediolani, 1833, 32 - 36." type="journal article" year="1833">Villa, 1833</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62976FF89FF4BFD97EE05FDC0" box="[151,555,547,573]" pageId="54" pageNumber="55" type="description">
<paragraph id="8BAAEF4D2976FF89FF4BFD97EE05FDC0" blockId="54.[151,807,510,573]" box="[151,555,547,573]" pageId="54" pageNumber="55">
(
<figureCitation id="132EF3C82976FF89FF43FD97ECDCFDC0" box="[159,242,547,573]" captionStart="FIGURE 21" captionStartId="55.[151,250,1890,1914]" captionTargetBox="[272,1292,206,1823]" captionTargetId="figure-17@55.[221,1367,181,1866]" captionTargetPageId="55" captionText="FIGURE 21. Oodes gracilis A. Villa &amp; G.B. Villa, 1833 (A–B: neotype and its labels; C–D: female specimen, Bulgaria, Pomorie-See; E–G: male specimen, Albania, Mali i Tabaroshit).A, C: habitus; B: left protarsomeres; D: pronotum and head; E: mentum; F: prosternal process; G: left metepisternum and metacoxa (arrow shows lateral end of metacoxal basal sulcus). Scale lines = 2 mm (Figs A, C); = 0.5 mm (Figs B, D, F–G); = 0.2 mm (Fig. E)." figureDoi="http://doi.org/10.5281/zenodo.4407122" httpUri="https://zenodo.org/record/4407122/files/figure.png" pageId="54" pageNumber="55">Figs 21</figureCitation>
A–G,
<figureCitation id="132EF3C82976FF89FEE4FD97EDA5FDC0" box="[312,395,547,573]" captionStart="FIGURE 22" captionStartId="56.[151,250,1892,1916]" captionTargetBox="[226,1363,194,1866]" captionTargetId="figure-20@56.[220,1368,181,1869]" captionTargetPageId="56" captionText="FIGURE 22. Oodes gracilis A. Villa &amp; G.B. Villa, 1833 (A–B: neotype; C: male specimen, Bulgaria, Dragoman Marsh; D–E: male specimen, Bulgaria, Srebarna Lake; F–G: female specimens, Bulgaria, Dragoman Marsh). A, C: median lobe of aedeagus, left lateral view; B, D: same, dorsal view; E: same, ventral view; F: left gonocoxite, ventral view; G: spermathecal complex and gonocoxites, ventral view. Scale lines = 0.5 mm (Figs A–E, G); = 0.2 mm (Fig. F)." figureDoi="http://doi.org/10.5281/zenodo.4407124" httpUri="https://zenodo.org/record/4407124/files/figure.png" pageId="54" pageNumber="55">Figs 22</figureCitation>
A–G,
<tableCitation id="C697DAF62976FF89FE13FD97EE0DFDC0" box="[463,547,547,573]" captionStart="TABLE 4" captionStartId="66.[151,239,1520,1544]" captionText="TABLE 4. Morphometric data for species of the genus Oodes Bonelli." pageId="54" pageNumber="55">Table 4</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62976FF89FF4BFDD3ED82FCE4" pageId="54" pageNumber="55" type="reference_group">
<paragraph id="8BAAEF4D2976FF89FF4BFDD3EE95FD63" blockId="54.[151,1436,615,793]" pageId="54" pageNumber="55">
<taxonomicName id="4C1594CE2976FF89FF4BFDD3EE43FD82" ID-CoL="49S3Z" authority="A. Villa &amp; G. B. Villa, 1833: 33" authorityName="A. Villa &amp; G. B. Villa" authorityPageNumber="33" authorityYear="1833" box="[151,621,615,639]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B961335F2976FF89FF4BFDD3ED05FD82" box="[151,299,615,639]" italics="true" pageId="54" pageNumber="55">Oodes gracilis</emphasis>
A. Villa &amp; G.B.
<bibRefCitation id="EF8492BC2976FF89FE0AFDD3EE43FD82" author="Villa, A. &amp; Villa, G. B." box="[470,621,615,639]" pageId="54" pageNumber="55" pagination="32 - 36" refId="ref64409" refString="Villa, A. &amp; Villa, G. B. (1833) Coleopterorum species novae in catalogo dupletorum extantes. Diagnosibus, adumbrationibusatque observationibus illustratae. Coleoptera Europae dupleta in collectione Villa quae pro mutua commutatione offerri possunt, Mediolani, 1833, 32 - 36." type="journal article" year="1833">Villa, 1833: 33</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54AE51EF2976FF89FDA6FDD3EE89FD82" box="[634,679,615,639]" pageId="54" pageNumber="55">type</typeStatus>
locality: ‘Italia’). Later, the locality was more precisely defined: ‘Alle rive dei laghi in Lombardia’ (
<bibRefCitation id="EF8492BC2976FF89FE1EFD32EE81FD63" author="Villa, A. &amp; Villa, G. B." box="[450,687,646,670]" pageId="54" pageNumber="55" pagination="58 - 82" refId="ref64458" refString="Villa, A. &amp; Villa, G. B. (1868) Riproduzione delle diagnosi di coleotteri e delle osservazioni pubblicate nel catalogo dei duplicati e supplementi, coll' aggiunta di nuove annotazioni. Atti della Societa italiana di Scienze naturali, Milano, 11, 58 - 82." type="journal article" year="1868">Villa &amp; Villa, 1868: 61</bibRefCitation>
).
</paragraph>
<paragraph id="8BAAEF4D2976FF89FF4BFD11ED6BFD26" blockId="54.[151,1436,615,793]" pageId="54" pageNumber="55">
=
<taxonomicName id="4C1594CE2976FF89FF77FD10EDD3FD40" ID-CoL="49S75" authority="Chaudoir, 1837: 20" authorityName="Chaudoir" authorityPageNumber="20" authorityYear="1837" box="[171,509,676,701]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B961335F2976FF89FF77FD10ED1AFD41" box="[171,308,676,700]" italics="true" pageId="54" pageNumber="55">Oodes similis</emphasis>
<bibRefCitation id="EF8492BC2976FF89FEE6FD10EDD3FD40" author="Chaudoir, M. de" box="[314,509,676,701]" pageId="54" pageNumber="55" pagination="3 - 20" refId="ref60416" refString="Chaudoir, M. de (1837) Description de quelques genres nouveaux et de quelques especes nouvelles ou inedites de carabiques. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 10 (3), 3 - 20." type="journal article" year="1837">Chaudoir, 1837: 20</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54AE51EF2976FF89FDD6FD11EE19FD40" box="[522,567,677,701]" pageId="54" pageNumber="55">type</typeStatus>
locality: ‘d’Allemagne’ [
<collectingCountry id="F302AFDD2976FF89FCE9FD11EFB2FD40" box="[821,924,677,701]" name="Germany" pageId="54" pageNumber="55">Germany</collectingCountry>
]). Synonymy established by
<bibRefCitation id="EF8492BC2976FF89FB61FD10ED6FFD26" author="Gemminger, M. &amp; Harold, E. von" pageId="54" pageNumber="55" pagination="8" refId="ref61462" refString="Gemminger, M. &amp; Harold, E. von (1868) Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. I. Cicindelidae-Carabidae. E. H. Gummi, Monachii, xxxvi + 424 + [8] pp. https: // doi. org / 10.5962 / bhl. title. 9089" type="journal article" year="1868">Gemminger &amp; Harold (1868: 232)</bibRefCitation>
.
</paragraph>
<paragraph id="8BAAEF4D2976FF89FF4BFD56ED82FCE4" blockId="54.[151,1436,615,793]" pageId="54" pageNumber="55">
=
<taxonomicName id="4C1594CE2976FF89FF76FD56EE3DFD07" ID-CoL="49S3Y" authority="Lambert, 1853: 199" authorityName="Lambert" authorityPageNumber="199" authorityYear="1853" box="[170,531,738,762]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilior">
<emphasis id="B961335F2976FF89FF76FD56ED64FD07" box="[170,330,738,762]" italics="true" pageId="54" pageNumber="55">Oodes gracilior</emphasis>
<bibRefCitation id="EF8492BC2976FF89FE92FD56EE3DFD07" author="Lambert, P." box="[334,531,738,762]" pageId="54" pageNumber="55" pagination="199" refId="ref62723" refString="Lambert, P. (1853) Note sur l'Oodes gracilior Fairm. Annales de la Societe Linneenne de Maine-et-Loire, 1, 199." type="journal article" year="1853">Lambert, 1853: 199</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54AE51EF2976FF89FDC3FD56EE62FD07" box="[543,588,738,762]" pageId="54" pageNumber="55">type</typeStatus>
locality: ‘bord de l'étang de Marson’,
<collectingCountry id="F302AFDD2976FF89FC19FD56E821FD07" box="[965,1039,738,762]" name="France" pageId="54" pageNumber="55">France</collectingCountry>
). Synonymy established by
<bibRefCitation id="EF8492BC2976FF89FAFFFD56ED89FCE4" author="Gemminger, M. &amp; Harold, E. von" pageId="54" pageNumber="55" pagination="8" refId="ref61462" refString="Gemminger, M. &amp; Harold, E. von (1868) Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. I. Cicindelidae-Carabidae. E. H. Gummi, Monachii, xxxvi + 424 + [8] pp. https: // doi. org / 10.5962 / bhl. title. 9089" type="journal article" year="1868">Gemminger &amp; Harold (1868: 232)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62976FF89FF4BFC8AED3DFCA8" box="[151,275,830,854]" pageId="54" pageNumber="55" type="multiple">
<paragraph id="8BAAEF4D2976FF89FF4BFC8AED3DFCA8" blockId="54.[151,1437,830,1253]" box="[151,275,830,854]" pageId="54" pageNumber="55">
<emphasis id="B961335F2976FF89FF4BFC8AED23FCAB" bold="true" box="[151,269,830,854]" pageId="54" pageNumber="55">References</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62976FF89FF4BFCE8E8A1FB18" pageId="54" pageNumber="55" type="reference_group">
<paragraph id="8BAAEF4D2976FF89FF4BFCE8EEC0FC6E" blockId="54.[151,1437,830,1253]" pageId="54" pageNumber="55">
<taxonomicName id="4C1594CE2976FF89FF4BFCE8EE24FC89" ID-CoL="49S75" authority=": Chaudoir 1857: 34 - 35" authorityName="Chaudoir" authorityPageNumber="34 - 35" authorityYear="1857" box="[151,522,860,884]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B961335F2976FF89FF4BFCE8ED30FC89" box="[151,286,860,884]" italics="true" pageId="54" pageNumber="55">Oodes similis</emphasis>
:
<bibRefCitation id="EF8492BC2976FF89FEF6FCE8EE24FC89" author="Chaudoir, M. de" box="[298,522,860,884]" pageId="54" pageNumber="55" pagination="1 - 64" refId="ref60458" refString="Chaudoir, M. de (1857) Memoire sur la famille des carabiques. 6 - e partie (Continuation). Bulletin de la Societe Imperiale des Naturalistes de Moscou, 30 (3), 1 - 64." type="journal article" year="1857">Chaudoir 1857: 34–35</bibRefCitation>
</taxonomicName>
(comparison with
<taxonomicName id="4C1594CE2976FF89FD18FCE9EF78FC89" ID-CoL="49S45" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[708,854,860,884]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="helopioides">
<emphasis id="B961335F2976FF89FD18FCE9EEF8FC89" box="[708,726,861,884]" italics="true" pageId="54" pageNumber="55">O</emphasis>
.
<emphasis id="B961335F2976FF89FD3DFCE8EF78FC89" box="[737,854,860,884]" italics="true" pageId="54" pageNumber="55">helopioides</emphasis>
</taxonomicName>
);
<bibRefCitation id="EF8492BC2976FF89FCB6FCE8E87BFC88" author="Motschulsky, V. de" box="[874,1109,860,885]" pageId="54" pageNumber="55" pagination="171 - 178" refId="ref63618" refString="Motschulsky, V. de (1858) Litterature. Etudes Entomologiques, 7, 171 - 178." type="journal article" year="1858">Motschulsky 1858: 172</bibRefCitation>
(comparisons with
<taxonomicName id="4C1594CE2976FF89FAC4FCE9ED64FC6E" ID-CoL="8LFDG" authority="Motschulsky" authorityName="Motschulsky" authorityYear="1858" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B961335F2976FF89FAC4FCE9E904FC89" box="[1304,1322,861,884]" italics="true" pageId="54" pageNumber="55">O</emphasis>
.
<emphasis id="B961335F2976FF89FAE9FCE8E9B2FC89" box="[1333,1436,860,884]" italics="true" pageId="54" pageNumber="55">parallelus</emphasis>
Motschulsky
</taxonomicName>
and
<taxonomicName id="4C1594CE2976FF89FEA7FCC8EECAFC6E" ID-CoL="8LL2F" authority="Motschulsky" authorityName="Motschulsky" authorityYear="1858" box="[379,740,891,915]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="parallelogrammus">
<emphasis id="B961335F2976FF89FEA7FCC8EDA3FC6E" box="[379,397,892,915]" italics="true" pageId="54" pageNumber="55">O</emphasis>
.
<emphasis id="B961335F2976FF89FE46FCCFEE7BFC6E" box="[410,597,891,915]" italics="true" pageId="54" pageNumber="55">parallelogrammus</emphasis>
Motschulsky
</taxonomicName>
).
</paragraph>
<paragraph id="8BAAEF4D2976FF89FF4BFC2EEF65FC4F" blockId="54.[151,1437,830,1253]" box="[151,843,922,946]" pageId="54" pageNumber="55">
<taxonomicName id="4C1594CE2976FF89FF4BFC2EEE8EFC4F" ID-CoL="49S3Y" authority=": Fairmaire &amp; Laboulbene 1854: 60" authorityName="Fairmaire &amp; Laboulbene" authorityPageNumber="60" authorityYear="1854" box="[151,672,922,946]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilior">
<emphasis id="B961335F2976FF89FF4BFC2EED16FC4F" box="[151,312,922,946]" italics="true" pageId="54" pageNumber="55">Oodes gracilior</emphasis>
:
<bibRefCitation id="EF8492BC2976FF89FE9AFC2EEE8EFC4F" author="Fairmaire, L. &amp; Laboulbene, A." box="[326,672,922,946]" pageId="54" pageNumber="55" pagination="1 - 180" refId="ref61189" refString="Fairmaire, L. &amp; Laboulbene, A. (1854) Livraison 1. In: Faune entomologique francaise ou description des insectes qui se trouvent en France. Coleopteres. Tome Premier. Deyrolle, Paris, pp. i-xxxv + 1 - 180." type="book chapter" year="1854">Fairmaire &amp; Laboulbène 1854: 60</bibRefCitation>
</taxonomicName>
(re-description).
</paragraph>
<paragraph id="8BAAEF4D2976FF89FF4BFC0CE856FB5A" blockId="54.[151,1437,830,1253]" pageId="54" pageNumber="55">
<taxonomicName id="4C1594CE2976FF89FF4BFC0CEEBCFC2C" ID-CoL="49S3Z" authority=": Gemminger &amp; Harold 1868: 232" authorityName="Gemminger &amp; Harold" authorityPageNumber="232" authorityYear="1868" box="[151,658,952,977]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B961335F2976FF89FF4BFC0CED03FC2D" box="[151,301,952,976]" italics="true" pageId="54" pageNumber="55">Oodes gracilis</emphasis>
:
<bibRefCitation id="EF8492BC2976FF89FEE0FC0CEEBCFC2C" author="Gemminger, M. &amp; Harold, E. von" box="[316,658,952,977]" pageId="54" pageNumber="55" pagination="8" refId="ref61462" refString="Gemminger, M. &amp; Harold, E. von (1868) Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. I. Cicindelidae-Carabidae. E. H. Gummi, Monachii, xxxvi + 424 + [8] pp. https: // doi. org / 10.5962 / bhl. title. 9089" type="journal article" year="1868">Gemminger &amp; Harold 1868: 232</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EF8492BC2976FF89FD42FC0CEF12FC2D" author="Bedel, L." box="[670,828,952,977]" pageId="54" pageNumber="55" pagination="1 - 128" refId="ref59945" refString="Bedel, L. (1879) Faune des coleopteres du bassin de la Seine et de ses bassins secondaires. 1 er Sous-ordre. Carnivora. Annales de la Societe Entomologique de France, Series 5, 9, 1 - 128. [sep. pagin.] https: // doi. org / 10.5962 / bhl. title. 8757" type="journal article" year="1879">Bedel 1879: 54</bibRefCitation>
(identification key);
<bibRefCitation id="EF8492BC2976FF89FBC4FC0CE8E8FC2D" author="Bedel, L." box="[1048,1222,952,977]" pageId="54" pageNumber="55" pagination="129 - 256" refId="ref60009" refString="Bedel, L. (1880) Faune des coleopteres du bassin de la Seine et de ses bassins secondaires. 1 er Sous-ordre. Carnivora. Annales de la Societe Entomologique de France, (5) 10, 129 - 256. [sep. pagin.] https: // doi. org / 10.5962 / bhl. title. 8757" type="journal article" year="1880">Bedel 1880: 161</bibRefCitation>
;
<bibRefCitation id="EF8492BC2976FF89FB0EFC0CE9B6FC2C" author="Marseul, S. A. de" box="[1234,1432,952,977]" pageId="54" pageNumber="55" pagination="1 - 526" refId="ref63452" refString="Marseul, S. A. de (1880) Nouveau repertoire contenant les descriptions des especes de coleopteres de l'ancien monde publiees isolement ou en langues etrangeres, en dehors des monographies ou traites speciaux et de l'Abeille. L'Abeille, Journal d'Entomologie, 19, 1 - 526." type="journal article" year="1880">Marseul 1880: 190</bibRefCitation>
;
<bibRefCitation id="EF8492BC2976FF89FF1BFC63EDB4FC12" author="Chaudoir, M. de" box="[199,410,983,1007]" pageId="54" pageNumber="55" pagination="317 - 378" refId="ref60500" refString="Chaudoir, M. de (1882) Monographie des oodides 1 re partie. Annales de la Societe Entomologique de France, Series 6, 2, 317 - 378." type="journal article" year="1882">Chaudoir 1882: 345</bibRefCitation>
(re-description);
<bibRefCitation id="EF8492BC2976FF89FD88FC63EF3CFC12" author="Marseul, S. A. de" box="[596,786,983,1007]" pageId="54" pageNumber="55" pagination="1 - 196" refId="ref63503" refString="Marseul, S. A. de (1882) Nouveau repertoire contenant les descriptions des especes de coleopteres de l'ancien monde publiees isolement ou en langues etrangeres, en dehors des monographies ou traites speciaux et de l'Abeille. L'Abeille, Journal d'Entomologie, 20, 1 - 196." type="journal article" year="1882">Marseul 1882: 28</bibRefCitation>
;
<bibRefCitation id="EF8492BC2976FF89FCFDFC63E835FC12" author="Ganglbauer, L." box="[801,1051,983,1007]" pageId="54" pageNumber="55" refId="ref61349" refString="Ganglbauer, L. (1891 a) Die Kafer von Mitteleuropa. Die Kafer der osterreichisch-ungarischen Monarchie, Deutschlands, der Schweiz, sowie des franzosischen und italienischen Alpengebietes. Erster Band. Familienreihe Caraboidea. C. Gerold's Sohn, Wien, iii + 557 pp. https: // doi. org / 10.5962 / bhl. title. 8764" type="book" year="1891">Ganglbauer 1891a: 384</bibRefCitation>
;
<bibRefCitation id="EF8492BC2976FF89FBF5FC63E8E1FC12" author="Fiori, A." box="[1065,1231,983,1007]" pageId="54" pageNumber="55" pagination="198 - 206" refId="ref61236" refString="Fiori, A. (1903) Note di corologia e di caccia. Nuove indicazioni topografiche. Rivista Coleotterologica Italiana, 1, 198 - 206." type="journal article" year="1903">Fiori 1903: 200</bibRefCitation>
(‘fiume Lamato in Calabria’,
<collectingCountry id="F302AFDD2976FF89FEEEFC42ED4BFBF3" box="[306,357,1014,1038]" name="Italy" pageId="54" pageNumber="55">Italy</collectingCountry>
);
<bibRefCitation id="EF8492BC2976FF89FEAAFC42EE05FBF3" author="Reitter, E." box="[374,555,1014,1038]" pageId="54" pageNumber="55" refId="ref63987" refString="Reitter, E. (1908) Fauna Germanica. Die Kafer des Deutschen Reiches. Nach der analytischen Methode bearbeitet. I. Band. Schriften des Deutschen Lehrervereins fur Naturkunde 22. K. G. Lutz, Stuttgart, viii + 248 pp., 40 pls." type="book" year="1908">Reitter 1908: 186</bibRefCitation>
;
<bibRefCitation id="EF8492BC2976FF89FDE9FC42EEF4FBF3" author="Porta, A." box="[565,730,1014,1038]" pageId="54" pageNumber="55" refId="ref63945" refString="Porta, A. (1923) Fauna Coleopterorum Italica. Vol. I. Adephaga. Cicindelidae, Carabidae, Haliplidae, Hygrobiidae, Dyticidae, Gyrinidae, Rhisodidae, Paussidae. Stabilimento Tipographico Piacentino, Piacenza, 285 pp." type="book" year="1923">Porta 1923: 213</bibRefCitation>
;
<bibRefCitation id="EF8492BC2976FF89FD39FC42EFECFBF3" author="Burmeister, F." box="[741,962,1014,1038]" pageId="54" pageNumber="55" refId="ref60386" refString="Burmeister, F. (1939) Biologie, Okologie und Verbreitung der europaischen Kafer, I Adephaga, Caraboidea. E. Goecke, Krefeld, 307 pp." type="book" year="1939">Burmeister 1939: 165</bibRefCitation>
(distribution and ecology);
<bibRefCitation id="EF8492BC2976FF89FB01FC42E9B6FBF3" author="Jeannel, R." box="[1245,1432,1014,1038]" pageId="54" pageNumber="55" pagination="572 - 1173" refId="ref62241" refString="Jeannel, R. (1942) Coleopteres carabiques. Deuxieme Partie. In: Faune de France. 40. Librairie de la Faculte des Sciences, Paris, pp. 572 - 1173." type="book chapter" year="1942">Jeannel 1942: 982</bibRefCitation>
;
<bibRefCitation id="EF8492BC2976FF89FF1BFBA0ED50FBD1" author="Focarile, A." box="[199,382,1044,1068]" pageId="54" pageNumber="55" pagination="17 - 114" refId="ref61264" refString="Focarile, A. (1959) Ricerche coleotterologiche sul litorale ionico della Puglia, Lucania e Calabria campagna 1956 - I. Notizie untroduttive Coleoptera, Carabidae. Memorie de la Societa Entomologica Italiana, Fasc. Speciale, 38, 17 - 114." type="journal article" year="1959">Focarile 1959: 75</bibRefCitation>
(taxonomic notes and comparison with
<taxonomicName id="4C1594CE2976FF89FCF8FBA1EF95FBD1" ID-CoL="49S45" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[804,955,1044,1068]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="helopioides">
<emphasis id="B961335F2976FF89FCF8FBA1EF18FBD1" box="[804,822,1045,1068]" italics="true" pageId="54" pageNumber="55">O</emphasis>
.
<emphasis id="B961335F2976FF89FC9AFBA0EF95FBD1" box="[838,955,1044,1068]" italics="true" pageId="54" pageNumber="55">helopioides</emphasis>
</taxonomicName>
);
<bibRefCitation id="EF8492BC2976FF89FC0FFBA0E8FFFBD0" author="Fontolan, P." box="[979,1233,1044,1069]" pageId="54" pageNumber="55" pagination="118 - 121" refId="ref61310" refString="Fontolan, P. (1959) Sugli Oodes helopioides Fabr. e gracilis Villa nella Laguna di Venezia (Col. Carabidae). Bollettino della Societa Entomologica Italiana, Genova, 91, 118 - 121." type="journal article" year="1959">Fontolan 1959: 120–121</bibRefCitation>
(re-description and comparison with
<taxonomicName id="4C1594CE2976FF89FEA9FB80EE29FBB6" ID-CoL="49S45" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[373,519,1075,1099]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="helopioides">
<emphasis id="B961335F2976FF89FEA9FB80EDA9FBB6" box="[373,391,1076,1099]" italics="true" pageId="54" pageNumber="55">O</emphasis>
.
<emphasis id="B961335F2976FF89FE4EFB87EE29FBB6" box="[402,519,1075,1099]" italics="true" pageId="54" pageNumber="55">helopioides</emphasis>
</taxonomicName>
);
<bibRefCitation id="EF8492BC2976FF89FDC0FB87EED6FBB6" author="Magistretti, M." box="[540,760,1075,1099]" pageId="54" pageNumber="55" refId="ref63422" refString="Magistretti, M. (1965) Coleoptera: Cicindelidae, Carabidae. Catalogo topografico. Fauna d'Italia. 8. Calderini, Bologna, xv + 512 pp." type="book" year="1965">Magistretti 1965: 257</bibRefCitation>
;
<bibRefCitation id="EF8492BC2976FF89FCDCFB87EFBBFBB6" author="Allen, R. T." box="[768,917,1075,1099]" pageId="54" pageNumber="55" pagination="1 - 46" refId="ref59448" refString="Allen, R. T. (1973) Species of Carabidae (Insecta: Coleoptera) in the University of Arkansas Insect Collection. University of Arkansas (Fayetteville) Agricultural Experiment Station Special Report, 19, 1 - 46." type="journal article" year="1973">Allen 1973: 32</bibRefCitation>
(Arkansas Insect Collection);
<bibRefCitation id="EF8492BC2976FF89FB1AFB80ECC3FB97" author="Bousquet, Y." pageId="54" pageNumber="55" pagination="443 - 537" refId="ref60231" refString="Bousquet, Y. (1996) Taxonomic revision of Nearctic, Mexican, and West Indian Oodini (Coleoptera: Carabidae). The Canadian Entomologist, 128, 443 - 537. https: // doi. org / 10.4039 / Ent 128443 - 3" type="journal article" year="1996">Bousquet 1996: 449– 450</bibRefCitation>
(various data, incl. bibliography), 472 (relationships with
<taxonomicName id="4C1594CE2976FF89FCE0FBE6EFEAFB97" ID-CoL="49S2B" authorityName="Dejean" authorityYear="1831" box="[828,964,1106,1130]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="amaroides">
<emphasis id="B961335F2976FF89FCE0FBE6EF60FB94" box="[828,846,1106,1129]" italics="true" pageId="54" pageNumber="55">O</emphasis>
.
<emphasis id="B961335F2976FF89FC86FBE6EFEAFB97" box="[858,964,1106,1130]" italics="true" pageId="54" pageNumber="55">amaroides</emphasis>
</taxonomicName>
);
<bibRefCitation id="EF8492BC2976FF89FC06FBE6E8A0FB97" author="Ortuno, V. M." box="[986,1166,1106,1130]" pageId="54" pageNumber="55" pagination="3 - 10" refId="ref63811" refString="Ortuno, V. M. (1998) La genitalia femenina en los Oodinae (Caraboidea: Callistidae) de la region paleartica occidental. Elytron, 12, 3 - 10." type="journal article" year="1998">Ortuño 1998: 5–6</bibRefCitation>
(female genitalia);
<bibRefCitation id="EF8492BC2976FF89FA89FBE6ED1FFB75" author="Lorenz, W." pageId="54" pageNumber="55" refId="ref63123" refString="Lorenz, W. (1998) Systematic list of extant ground beetles of the world (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 1 st Edition. Published by the Author, Tutzing, [ii] + 502 pp." type="book" year="1998">Lorenz 1998: 305</bibRefCitation>
;
<bibRefCitation id="EF8492BC2976FF89FEE7FBC5EE25FB75" author="Bousquet, Y." box="[315,523,1136,1161]" pageId="54" pageNumber="55" pagination="1851" refId="ref60278" refString="Bousquet, Y. (2003) Tribe Oodini LaFerte-Senectere, 1851. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 444 - 445." type="book chapter" year="2003">Bousquet 2003: 445</bibRefCitation>
;
<bibRefCitation id="EF8492BC2976FF89FDCAFBC5EEE3FB75" author="Lorenz, W." box="[534,717,1136,1160]" pageId="54" pageNumber="55" refId="ref63176" refString="Lorenz, W. (2005) Systematic list of extant ground beetles of the world (Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Published by the author, Tutzing, [ii] + 530 pp." type="book" year="2005">Lorenz 2005: 325</bibRefCitation>
;
<bibRefCitation id="EF8492BC2976FF89FD04FBC4EFFBFB75" author="Elderkhanova, Z. M." box="[728,981,1136,1161]" pageId="54" pageNumber="55" pagination="27" refId="ref60844" refString="Elderkhanova, Z. M. (2012) Zhuzhelitsy (Coleoptera, Carabidae) ostrova Tyuleniy v severozapadnom Kaspii [Ground beetles (Coleoptera, Carabidae) of the Tyuleniy Island in the northwestern Caspian Sea]. In: Materialy XIV syezda Russkogo entomologicheskogo obstchestva, Sankt-Peterburg, 27 avgusta- 1 sentyabrya 2012 [XIV Congress of the Russian Entomological Society, Saint-Petersburg, 27 August- 1 September, 2012], pp. 495. [in Russian]" type="journal article" year="2012">Elderkhanova 2012: 495</bibRefCitation>
;
<bibRefCitation id="EF8492BC2976FF89FC3CFBC4E937FB75" author="Nakhibasheva, G. M. &amp; Belousov, I. A. &amp; Kabak, I. I. &amp; Abdurakhmanov, G. M." box="[992,1305,1136,1161]" pageId="54" pageNumber="55" pagination="1" refId="ref63709" refString="Nakhibasheva, G. M., Belousov, I. A., Kabak, I. I. &amp; Abdurakhmanov, G. M. (2012) Materialy po rasprostraneniyu nekotorykh vidov zhuzhelits (Coleoptera, Carabidae) v Dagestane [Materials on the distribution of some ground-beetle species in Dagestan (Coleoptera, Carabidae)]. In: Materialy XIV syezda Russkogo entomologicheskogo obstchestva, Sankt-Peterburg, 27 Avgusta- 1 Sentyabrya 2012 [XIV Congress of the Russian Entomological Society, Saint-Petersburg, 27 August- 1 September 1 2012], 305. [in Russian]" type="journal article" year="2012">
Nakhibasheva
<emphasis id="B961335F2976FF89FBAAFBC5E88DFB75" box="[1142,1187,1136,1160]" italics="true" pageId="54" pageNumber="55">et al</emphasis>
. 2012: 305
</bibRefCitation>
;
<bibRefCitation id="EF8492BC2976FF89FAF8FBC4ED6AFB5A" author="Belousov, I. A. &amp; Kabak, I. I. &amp; Nakhibasheva, G. M. &amp; Mukhtarova, G. M. &amp; Rabadanov, M." pageId="54" pageNumber="55" pagination="93 - 98" refId="ref60137" refString="Belousov, I. A., Kabak, I. I., Nakhibasheva, G. M., Mukhtarova, G. M. &amp; Rabadanov, M. (2014) Carabid beetles fauna (Coleoptera, Carabidae) of the Tshetshen island in the Caspian sea. The South of Russia: ecology, development, 9 (3), 93 - 98. [in Russian] https: // doi. org / 10.18470 / 1992 - 1098 - 2014 - 3 - 93 - 98" type="journal article" year="2014">
Belousov
<emphasis id="B961335F2976FF89FA56FBC5ECF5FB5A" italics="true" pageId="54" pageNumber="55">et al</emphasis>
. 2014: 96
</bibRefCitation>
;
<bibRefCitation id="EF8492BC2976FF89FE92FB3BEE14FB5A" author="Ibanez Orrico, M. A." box="[334,570,1167,1191]" pageId="54" pageNumber="55" pagination="77 - 88" refId="ref62045" refString="Ibanez Orrico, M. A. (2014) Contribucion al conocimiento de la fauna valenciana de coleopteros (E de la Peninsula Iberica). I. Coleoptera, Carabidae. Arquivos Entomoloxicos, 10, 77 - 88." type="journal article" year="2014">Ibáñez Orrico 2014: 83</bibRefCitation>
(distribution in Iberian Peninsula);
<bibRefCitation id="EF8492BC2976FF89FC78FB24E85DFB5A" author="Bousquet, Y." box="[932,1139,1167,1191]" pageId="54" pageNumber="55" pagination="1851" refId="ref60329" refString="Bousquet, Y. (2017) Tribe Oodini LaFerte-Senectere, 1851. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Revised and Updated Edition. Archostemata-Myxophaga-Adephaga. Brill, Leiden and Boston, pp. 635 - 636." type="book chapter" year="2017">Bousquet 2017: 636</bibRefCitation>
.
</paragraph>
<paragraph id="8BAAEF4D2976FF89FF4BFB1AE8A1FB18" blockId="54.[151,1437,830,1253]" pageId="54" pageNumber="55">
<taxonomicName id="4C1594CE2976FF89FF4BFB1AEEB6FB3B" ID-CoL="49S3Z" authority=": Ganglbauer 1891 b: 51" authorityName="Ganglbauer" authorityPageNumber="51" authorityYear="1891" box="[151,664,1198,1222]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilis" subGenus="Oodes">
<emphasis id="B961335F2976FF89FF4BFB1AECF6FB3B" box="[151,216,1198,1222]" italics="true" pageId="54" pageNumber="55">Oodes</emphasis>
(
<emphasis id="B961335F2976FF89FF31FB1AED00FB3B" box="[237,302,1198,1222]" italics="true" pageId="54" pageNumber="55">Oodes</emphasis>
)
<emphasis id="B961335F2976FF89FE98FB1AEDBFFB3B" box="[324,401,1198,1222]" italics="true" pageId="54" pageNumber="55">gracilis</emphasis>
:
<bibRefCitation id="EF8492BC2976FF89FE79FB1AEEB6FB3B" author="Ganglbauer, L." box="[421,664,1198,1222]" pageId="54" pageNumber="55" pagination="1 - 58" refId="ref61412" refString="Ganglbauer, L. (1891 b) Cicindelidae, Carabidae. In: Heyden, L. von, Reitter, E. &amp; Weise J., Catalogus Coleopterorum Europae, Caucasi et Armeniae rossicae. E. Reitter, Modling, pp. 1 - 58." type="book chapter" year="1891">Ganglbauer 1891b: 51</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EF8492BC2976FF89FD76FB1AEF61FB3B" author="Csiki, E." box="[682,847,1198,1222]" pageId="54" pageNumber="55" pagination="1 - 110" refId="ref60532" refString="Csiki, E. (1906) Cicindelidae, Carabidae. In: Heyden, L. F. J. D. von, Reitter, E. &amp; Weise, J., Catalogus Coleopterorum Europae, Caucasi et Armeniae Rossicae. Editio Secunda. R. Friedlander &amp; Sohn, Modling, Berlin and E. Reitter, Caen, pp. 1 - 110." type="book chapter" year="1906">Csiki 1906: 57</bibRefCitation>
;
<bibRefCitation id="EF8492BC2976FF89FCBDFB1AE817FB3B" author="Jakobson, G. G." box="[865,1081,1198,1222]" pageId="54" pageNumber="55" pagination="241 - 320" refId="ref62206" refString="Jakobson, G. G. (1906) Fasc. 4. In: Zhuki Rossii i Zapadnoi Evropy. A. F. Devrien, Sankt-Petersburg, pp. 241 - 320." type="book chapter" year="1906">Jakobson 1906: 310</bibRefCitation>
(distribution);
<bibRefCitation id="EF8492BC2976FF89FB3CFB1AE9B3FB3B" author="Csiki, E." box="[1248,1437,1198,1222]" pageId="54" pageNumber="55" pagination="739 - 1022" refId="ref60601" refString="Csiki, E. (1931) Carabidae: Harpalinae V. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum catalogus. Pars 115. W. Junk, Berlin, pp. 739 - 1022." type="book chapter" year="1931">Csiki 1931: 1007</bibRefCitation>
(‘Mitteleuropa, Mittelmeer–gebiet’);
<bibRefCitation id="EF8492BC2976FF89FD9DFB78EF55FB19" author="Kryzhanovskij, O. L. &amp; Belousov, I. A. &amp; Kabak, I. I. &amp; Kataev, M. B. &amp; Makarov, K. V. &amp; Shilenkov, V. G." box="[577,891,1228,1252]" pageId="54" pageNumber="55" refId="ref62520" refString="Kryzhanovskij, O. L., Belousov, I. A., Kabak, I. I., Kataev, M. B., Makarov, K. V. &amp; Shilenkov, V. G. (1995). A Checklist of the Ground-beetles of Russia and Adjacent Lands (Insecta, Coleoptera, Carabidae). Pensoft, Sofia &amp; Moscow, 271 pp." type="book" year="1995">
Kryzhanovskij
<emphasis id="B961335F2976FF89FD00FB79EF27FB19" box="[732,777,1228,1252]" italics="true" pageId="54" pageNumber="55">et al</emphasis>
. 1995: 158
</bibRefCitation>
(distribution in ex-USSR).
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62976FF86FF4BFAA7EE42FE08" lastPageId="57" lastPageNumber="58" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="8BAAEF4D2976FF89FF4BFAA7ECD6FA88" blockId="54.[151,1437,1298,2009]" pageId="54" pageNumber="55">
<emphasis id="B961335F2976FF89FF4BFAA7ED15FAD1" bold="true" box="[151,315,1298,1325]" pageId="54" pageNumber="55">Type material</emphasis>
.
<taxonomicName id="4C1594CE2976FF89FE9BFAA7EEFAFAD0" authority="A. Villa &amp; G. B. Villa" authorityName="A. Villa &amp; G. B. Villa" authorityYear="1833" box="[327,724,1299,1325]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B961335F2976FF89FE9BFAA7EDC7FAD0" box="[327,489,1299,1325]" italics="true" pageId="54" pageNumber="55">Oodes gracilis</emphasis>
A. Villa &amp; G.B. Villa
</taxonomicName>
:
<typeStatus id="54AE51EF2976FF89FD00FAA7EF1BFAD0" box="[732,821,1299,1325]" pageId="54" pageNumber="55" type="neotype">neotype</typeStatus>
<specimenCount id="9D1324C42976FF89FCE5FAA7EF7EFAD0" box="[825,848,1299,1325]" pageId="54" pageNumber="55" type="male">♂</specimenCount>
, ‘Oasi Torbiere diAlbate—Bassone (CO)
<date id="FFABC98D2976FF89FAC1FAA7E9B2FAD0" box="[1309,1436,1299,1325]" pageId="54" pageNumber="55" value="1978-06-31">31.VI.1978</date>
Leg. Sciaky—Pavesi [w, h] //
<typeStatus id="54AE51EF2976FF89FE2DFA83EE72FAAC" box="[497,604,1335,1361]" pageId="54" pageNumber="55" type="neotype">Neotypus</typeStatus>
<taxonomicName id="4C1594CE2976FF89FDBAFA83E812FAAC" authority="Villa Det. Sciaky e Pavesi" authorityName="Villa Det. Sciaky e Pavesi" box="[614,1084,1335,1361]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilis">Oodes gracilis Villa Det. Sciaky e Pavesi</taxonomicName>
[r, h]’ (MSNM) (
<figureCitation id="132EF3C82976FF89FAD4FA83E9B9FAAC" box="[1288,1431,1335,1361]" captionStart="FIGURE 21" captionStartId="55.[151,250,1890,1914]" captionTargetBox="[272,1292,206,1823]" captionTargetId="figure-17@55.[221,1367,181,1866]" captionTargetPageId="55" captionText="FIGURE 21. Oodes gracilis A. Villa &amp; G.B. Villa, 1833 (A–B: neotype and its labels; C–D: female specimen, Bulgaria, Pomorie-See; E–G: male specimen, Albania, Mali i Tabaroshit).A, C: habitus; B: left protarsomeres; D: pronotum and head; E: mentum; F: prosternal process; G: left metepisternum and metacoxa (arrow shows lateral end of metacoxal basal sulcus). Scale lines = 2 mm (Figs A, C); = 0.5 mm (Figs B, D, F–G); = 0.2 mm (Fig. E)." figureDoi="http://doi.org/10.5281/zenodo.4407122" httpUri="https://zenodo.org/record/4407122/files/figure.png" pageId="54" pageNumber="55">Figs 21A, B</figureCitation>
,
<figureCitation id="132EF3C82976FF89FF4BFAEFECC4FA88" box="[151,234,1371,1397]" captionStart="FIGURE 22" captionStartId="56.[151,250,1892,1916]" captionTargetBox="[226,1363,194,1866]" captionTargetId="figure-20@56.[220,1368,181,1869]" captionTargetPageId="56" captionText="FIGURE 22. Oodes gracilis A. Villa &amp; G.B. Villa, 1833 (A–B: neotype; C: male specimen, Bulgaria, Dragoman Marsh; D–E: male specimen, Bulgaria, Srebarna Lake; F–G: female specimens, Bulgaria, Dragoman Marsh). A, C: median lobe of aedeagus, left lateral view; B, D: same, dorsal view; E: same, ventral view; F: left gonocoxite, ventral view; G: spermathecal complex and gonocoxites, ventral view. Scale lines = 0.5 mm (Figs A–E, G); = 0.2 mm (Fig. F)." figureDoi="http://doi.org/10.5281/zenodo.4407124" httpUri="https://zenodo.org/record/4407124/files/figure.png" pageId="54" pageNumber="55">22A, B</figureCitation>
).
</paragraph>
<paragraph id="8BAAEF4D2976FF89FF1BFACBEF87F96B" blockId="54.[151,1437,1298,2009]" pageId="54" pageNumber="55">
<bibRefCitation id="EF8492BC2976FF89FF1BFACBED8CFA64" author="Focarile, A." box="[199,418,1406,1433]" pageId="54" pageNumber="55" pagination="17 - 114" refId="ref61264" refString="Focarile, A. (1959) Ricerche coleotterologiche sul litorale ionico della Puglia, Lucania e Calabria campagna 1956 - I. Notizie untroduttive Coleoptera, Carabidae. Memorie de la Societa Entomologica Italiana, Fasc. Speciale, 38, 17 - 114." type="journal article" year="1959">Focarile (1959: 75)</bibRefCitation>
considered that the description of
<bibRefCitation id="EF8492BC2976FF89FCF0FACBE83FFA64" author="Villa, A. &amp; Villa, G. B." box="[812,1041,1407,1433]" pageId="54" pageNumber="55" pagination="32 - 36" refId="ref64409" refString="Villa, A. &amp; Villa, G. B. (1833) Coleopterorum species novae in catalogo dupletorum extantes. Diagnosibus, adumbrationibusatque observationibus illustratae. Coleoptera Europae dupleta in collectione Villa quae pro mutua commutatione offerri possunt, Mediolani, 1833, 32 - 36." type="journal article" year="1833">Villa &amp; Villa (1833)</bibRefCitation>
refers to
<taxonomicName id="4C1594CE2976FF89FBA3FACBE9BBFA64" authorityName="Porta" authorityYear="1923" box="[1151,1429,1407,1433]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="variety" species="helopioides" variety="fiorii">
<emphasis id="B961335F2976FF89FBA3FACBE8BDFA64" box="[1151,1171,1407,1433]" italics="true" pageId="54" pageNumber="55">O</emphasis>
.
<emphasis id="B961335F2976FF89FB7EFACBE90DFA64" box="[1186,1315,1407,1433]" italics="true" pageId="54" pageNumber="55">helopioides</emphasis>
var.
<emphasis id="B961335F2976FF89FA81FACBE9BBFA64" box="[1373,1429,1407,1433]" italics="true" pageId="54" pageNumber="55">fiorii</emphasis>
</taxonomicName>
, and not to
<taxonomicName id="4C1594CE2976FF89FEC9FA17EDA1FA40" box="[277,399,1443,1469]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B961335F2976FF89FEC9FA17ED07FA40" box="[277,297,1443,1469]" italics="true" pageId="54" pageNumber="55">O</emphasis>
.
<emphasis id="B961335F2976FF89FEE5FA17EDA1FA40" box="[313,399,1443,1469]" italics="true" pageId="54" pageNumber="55">gracilis</emphasis>
</taxonomicName>
(sensu
<bibRefCitation id="EF8492BC2976FF89FE34FA17EEB9FA40" author="Lindroth, C. H." box="[488,663,1443,1469]" pageId="54" pageNumber="55" pagination="109 - 157" refId="ref62860" refString="Lindroth, C. H. (1943) Oodes gracilis Villa. Eine thermophile Carabide Schwedens. Notulae Entomologicae, 22, 109 - 157." type="journal article" year="1943">Lindroth, 1943</bibRefCitation>
). M. Pavesi (pers. comm.) wrote us: “
<emphasis id="B961335F2976FF89FB92FA17E8CDF9D4" italics="true" pageId="54" pageNumber="55">
Several specialists (including me), having not checked carefully enough the collection materials, suspected that the original description of
<taxonomicName id="4C1594CE2976FF89FA5DFA73ECC3F9F8" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilis">O. gracilis</taxonomicName>
(type locality “rive dei laghi lombardi” = shores of
<collectingRegion id="49D121AF2976FF89FC9EFA5FEF9AF9F8" box="[834,948,1515,1541]" country="Italy" name="Lombardia" pageId="54" pageNumber="55">Lombardy</collectingRegion>
lakes) would indeed refer to small, slender individuals of
<taxonomicName id="4C1594CE2976FF89FEE7F9BBED91F9D4" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[315,447,1551,1577]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="helopioides">helopioides</taxonomicName>
, since
<taxonomicName id="4C1594CE2976FF89FDD2F9BBEE4AF9D4" box="[526,612,1551,1577]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilis">gracilis</taxonomicName>
appeared to be restricted to coastal, brackish biotopes.
</emphasis>
”. However, this assumption could not be verified, as the type series of the species together with entire collection of the brothers Villa had been destroyed in the fire of the Museum of Milan in
<date id="FFABC98D2976FF89FCC8F9E3EF88F98C" box="[788,934,1623,1649]" pageId="54" pageNumber="55" value="1943-08">August 1943</date>
. In order to maintain the status of
<taxonomicName id="4C1594CE2976FF89FAC2F9E3E9B8F98C" box="[1310,1430,1623,1649]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B961335F2976FF89FAC2F9E3E91CF98C" box="[1310,1330,1623,1649]" italics="true" pageId="54" pageNumber="55">O</emphasis>
.
<emphasis id="B961335F2976FF89FA9CF9E3E9B8F98C" box="[1344,1430,1623,1649]" italics="true" pageId="54" pageNumber="55">gracilis</emphasis>
</taxonomicName>
, a
<typeStatus id="54AE51EF2976FF89FF77F9CFED2AF968" box="[171,260,1659,1685]" pageId="54" pageNumber="55" type="neotype">neotype</typeStatus>
of this species has been designated (
<bibRefCitation id="EF8492BC2976FF89FD42F9CFEFB2F96B" author="Sciaky, R. &amp; Pavesi, M." box="[670,924,1659,1686]" pageId="54" pageNumber="55" pagination="13 - 26" refId="ref64118" refString="Sciaky, R. &amp; Pavesi, M. (1986) Nuovi dati geonemici su Carabidae italiani. Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 127 (1 - 2), 13 - 26." type="journal article" year="1986">Sciaky &amp; Pavesi 1986</bibRefCitation>
).
</paragraph>
<paragraph id="8BAAEF4D2976FF89FF1BF92BEF19F944" blockId="54.[151,1437,1298,2009]" box="[199,823,1694,1721]" pageId="54" pageNumber="55">
<emphasis id="B961335F2976FF89FF1BF92BED40F945" bold="true" box="[199,366,1694,1721]" pageId="54" pageNumber="55">
<typeStatus id="54AE51EF2976FF89FF1BF92BED2CF944" box="[199,258,1695,1721]" pageId="54" pageNumber="55">Type</typeStatus>
material
</emphasis>
.
<taxonomicName id="4C1594CE2976FF89FEA0F92BEEA6F944" authority="Chaudoir" authorityName="Chaudoir" authorityYear="1837" box="[380,648,1695,1721]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B961335F2976FF89FEA0F92BEE3DF944" box="[380,531,1695,1721]" italics="true" pageId="54" pageNumber="55">Oodes similis</emphasis>
Chaudoir
</taxonomicName>
: Not examined.
</paragraph>
<paragraph id="8BAAEF4D2976FF89FF1BF977EF66F920" blockId="54.[151,1437,1298,2009]" box="[199,840,1730,1757]" pageId="54" pageNumber="55">
<emphasis id="B961335F2976FF89FF1BF977ED40F921" bold="true" box="[199,366,1730,1757]" pageId="54" pageNumber="55">
<typeStatus id="54AE51EF2976FF89FF1BF977ED2CF920" box="[199,258,1731,1757]" pageId="54" pageNumber="55">Type</typeStatus>
material
</emphasis>
.
<taxonomicName id="4C1594CE2976FF89FEA0F977EEB6F920" authority="Lambert" authorityName="Lambert" authorityYear="1853" box="[380,664,1731,1757]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilior">
<emphasis id="B961335F2976FF89FEA0F977EE00F920" box="[380,558,1731,1757]" italics="true" pageId="54" pageNumber="55">Oodes gracilior</emphasis>
Lambert
</taxonomicName>
: Not examined.
</paragraph>
<paragraph id="8BAAEF4D2976FF89FF1BF953EFC9F890" blockId="54.[151,1437,1298,2009]" pageId="54" pageNumber="55">
Paul
<bibRefCitation id="EF8492BC2976FF89FF23F953ED81F8FC" author="Lambert, P." box="[255,431,1766,1793]" pageId="54" pageNumber="55" pagination="199" refId="ref62723" refString="Lambert, P. (1853) Note sur l'Oodes gracilior Fairm. Annales de la Societe Linneenne de Maine-et-Loire, 1, 199." type="journal article" year="1853">Lambert (1853)</bibRefCitation>
ascribed the authorship of
<taxonomicName id="4C1594CE2976FF89FD07F953EF7CF8FC" box="[731,850,1767,1793]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B961335F2976FF89FD07F953EEC1F8FC" box="[731,751,1767,1793]" italics="true" pageId="54" pageNumber="55">O</emphasis>
.
<emphasis id="B961335F2976FF89FD20F953EF7CF8FC" box="[764,850,1767,1793]" italics="true" pageId="54" pageNumber="55">gracilis</emphasis>
</taxonomicName>
to Leon Faimaire perhaps because the former author used original notes of the latter author. Soon after, L. Fairmaire (
<bibRefCitation id="EF8492BC2976FF89FCB0F8BFE8C1F8DB" author="Fairmaire, L. &amp; Laboulbene, A." box="[876,1263,1802,1830]" pageId="54" pageNumber="55" pagination="1 - 180" refId="ref61189" refString="Fairmaire, L. &amp; Laboulbene, A. (1854) Livraison 1. In: Faune entomologique francaise ou description des insectes qui se trouvent en France. Coleopteres. Tome Premier. Deyrolle, Paris, pp. i-xxxv + 1 - 180." type="book chapter" year="1854">Fairmaire &amp; Laboulbène 1854: 60</bibRefCitation>
) gave a second description using the same descriptive terminology.
<bibRefCitation id="EF8492BC2976FF89FD06F89BEF99F8B4" author="Jeannel, R." box="[730,951,1839,1865]" pageId="54" pageNumber="55" pagination="572 - 1173" refId="ref62241" refString="Jeannel, R. (1942) Coleopteres carabiques. Deuxieme Partie. In: Faune de France. 40. Librairie de la Faculte des Sciences, Paris, pp. 572 - 1173." type="book chapter" year="1942">Jeannel (1942: 982)</bibRefCitation>
also attributed
<taxonomicName id="4C1594CE2976FF89FBBDF89BE899F8B4" box="[1121,1207,1839,1865]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B961335F2976FF89FBBDF89BE899F8B4" box="[1121,1207,1839,1865]" italics="true" pageId="54" pageNumber="55">gracilis</emphasis>
</taxonomicName>
to L. Fairmaire. The correct authorship of this form has been indicated by
<bibRefCitation id="EF8492BC2976FF89FD31F8E7EFCDF890" author="Chaudoir, M. de" box="[749,995,1874,1901]" pageId="54" pageNumber="55" pagination="317 - 378" refId="ref60500" refString="Chaudoir, M. de (1882) Monographie des oodides 1 re partie. Annales de la Societe Entomologique de France, Series 6, 2, 317 - 378." type="journal article" year="1882">Chaudoir (1882: 345)</bibRefCitation>
.
</paragraph>
<paragraph id="8BAAEF4D2976FF89FF1BF8C2E94DF86C" blockId="54.[151,1437,1298,2009]" box="[199,1379,1910,1937]" pageId="54" pageNumber="55">
<emphasis id="B961335F2976FF89FF1BF8C2EDDAF86D" bold="true" box="[199,500,1910,1936]" pageId="54" pageNumber="55">Other material examined</emphasis>
.
<collectingCountry id="F302AFDD2976FF89FDDEF8C3EE5BF86C" box="[514,629,1911,1937]" name="France" pageId="54" pageNumber="55">FRANCE</collectingCountry>
:
<emphasis id="B961335F2976FF89FD5CF8C2EFFFF86C" bold="true" box="[640,977,1910,1937]" pageId="54" pageNumber="55">A u v e r g n e –R h ô n e–A l p e s</emphasis>
:
<specimenCount id="9D1324C42976FF89FC3EF8C3E82BF86C" box="[994,1029,1911,1937]" pageId="54" pageNumber="55" type="female">1♀</specimenCount>
, ‘Dr Jacquet Lyon’ (
<collectionCode id="ED0477882976FF89FB30F8C3E976F86C" box="[1260,1368,1911,1937]" pageId="54" pageNumber="55">NMNHS</collectionCode>
).
</paragraph>
<paragraph id="8BAAEF4D2976FF89FF1BF82FEED9F824" blockId="54.[151,1437,1298,2009]" pageId="54" pageNumber="55">
<collectingCountry id="F302AFDD2976FF89FF1BF82FED67F848" box="[199,329,1947,1973]" name="Albania" pageId="54" pageNumber="55">ALBANIA</collectingCountry>
:
<emphasis id="B961335F2976FF89FE8AF82EEE1CF849" bold="true" box="[342,562,1946,1972]" pageId="54" pageNumber="55">
<collectingRegion id="49D121AF2976FF89FE8AF82EEE1CF849" box="[342,562,1946,1972]" country="Albania" name="Shkoder" pageId="54" pageNumber="55">Shkodër County</collectingRegion>
</emphasis>
:
<specimenCount id="9D1324C42976FF89FD9EF82FEE53F848" box="[578,637,1947,1973]" pageId="54" pageNumber="55" type="male">3♂♂</specimenCount>
, ‘ALB.,
<collectingRegion id="49D121AF2976FF89FD3EF82FEFB2F848" box="[738,924,1947,1973]" country="Albania" name="Shkoder" pageId="54" pageNumber="55">Shkodra District</collectingRegion>
Mali Tarabosh Mtn [Mali i Tabaroshit] karst terrain,
<quantity id="4CED42A82976FF89FF32F80BED1BF824" box="[238,309,1983,2009]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="55" unit="m" value="100.0">100 m</quantity>
<date id="FFABC98D2976FF89FEE0F80BED90F824" box="[316,446,1983,2009]" pageId="54" pageNumber="55" value="2000-11-24">24.XI.2000</date>
, A. Zhalov leg.’ (
<collectionCode id="ED0477882976FF89FD5CF80BEEC2F824" box="[640,748,1983,2009]" pageId="54" pageNumber="55">NMNHS</collectionCode>
).
</paragraph>
<caption id="DF6ABFC52977FF88FF4BF8D6EFACF81A" ID-DOI="http://doi.org/10.5281/zenodo.4407122" ID-Zenodo-Dep="4407122" httpUri="https://zenodo.org/record/4407122/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,250,1890,1914]" targetBox="[272,1292,206,1823]" targetPageId="55">
<paragraph id="8BAAEF4D2977FF88FF4BF8D6EFACF81A" blockId="55.[151,1437,1890,2023]" pageId="55" pageNumber="56">
<emphasis id="B961335F2977FF88FF4BF8D6ED0DF887" bold="true" box="[151,291,1890,1914]" pageId="55" pageNumber="56">FIGURE 21.</emphasis>
<taxonomicName id="4C1594CE2977FF88FEF1F8D7EED8F886" authority="A. Villa &amp; G. B. Villa, 1833" authorityName="A. Villa &amp; G. B. Villa" authorityYear="1833" box="[301,758,1891,1915]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B961335F2977FF88FEF1F8D7EDEBF886" box="[301,453,1891,1915]" italics="true" pageId="55" pageNumber="56">Oodes gracilis</emphasis>
A. Villa &amp; G.B.
<bibRefCitation id="EF8492BC2977FF88FD5EF8D7EED8F886" author="Villa, A. &amp; Villa, G. B." box="[642,758,1891,1915]" pageId="55" pageNumber="56" pagination="32 - 36" refId="ref64409" refString="Villa, A. &amp; Villa, G. B. (1833) Coleopterorum species novae in catalogo dupletorum extantes. Diagnosibus, adumbrationibusatque observationibus illustratae. Coleoptera Europae dupleta in collectione Villa quae pro mutua commutatione offerri possunt, Mediolani, 1833, 32 - 36." type="journal article" year="1833">Villa, 1833</bibRefCitation>
</taxonomicName>
(A–B: neotype and its labels; C–D: female specimen, Bulgaria, Pomorie-See; E–G: male specimen, Albania, Mali i Tabaroshit). A, C: habitus; B: left protarsomeres; D: pronotum and head; E: mentum; F: prosternal process; G: left metepisternum and metacoxa (arrow shows lateral end of metacoxal basal sulcus). Scale lines = 2 mm (Figs A, C); = 0.5 mm (Figs B, D, F–G); = 0.2 mm (Fig. E).
</paragraph>
</caption>
<caption id="DF6ABFC52978FF87FF4BF8D0EFF2F814" ID-DOI="http://doi.org/10.5281/zenodo.4407124" ID-Zenodo-Dep="4407124" httpUri="https://zenodo.org/record/4407124/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,250,1892,1916]" targetBox="[226,1363,194,1866]" targetPageId="56">
<paragraph id="8BAAEF4D2978FF87FF4BF8D0EFF2F814" blockId="56.[151,1437,1892,2025]" pageId="56" pageNumber="57">
<emphasis id="B961335F2978FF87FF4BF8D0ED0EF881" bold="true" box="[151,288,1892,1916]" pageId="56" pageNumber="57">FIGURE 22.</emphasis>
<taxonomicName id="4C1594CE2978FF87FEFAF8D1EEF9F880" authority="A. Villa &amp; G. B. Villa, 1833" authorityName="A. Villa &amp; G. B. Villa" authorityYear="1833" box="[294,727,1893,1917]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B961335F2978FF87FEFAF8D1ED94F880" box="[294,442,1893,1917]" italics="true" pageId="56" pageNumber="57">Oodes gracilis</emphasis>
A. Villa &amp; G.B.
<bibRefCitation id="EF8492BC2978FF87FDBAF8D1EEF9F880" author="Villa, A. &amp; Villa, G. B." box="[614,727,1893,1917]" pageId="56" pageNumber="57" pagination="32 - 36" refId="ref64409" refString="Villa, A. &amp; Villa, G. B. (1833) Coleopterorum species novae in catalogo dupletorum extantes. Diagnosibus, adumbrationibusatque observationibus illustratae. Coleoptera Europae dupleta in collectione Villa quae pro mutua commutatione offerri possunt, Mediolani, 1833, 32 - 36." type="journal article" year="1833">Villa, 1833</bibRefCitation>
</taxonomicName>
(A–B: neotype; C: male specimen, Bulgaria, Dragoman Marsh; D–E: male specimen, Bulgaria, Srebarna Lake; F–G: female specimens, Bulgaria, Dragoman Marsh). A, C: median lobe of aedeagus, left lateral view; B, D: same, dorsal view; E: same, ventral view; F: left gonocoxite, ventral view; G: spermathecal complex and gonocoxites, ventral view. Scale lines = 0.5 mm (Figs A–E, G); = 0.2 mm (Fig. F).
</paragraph>
</caption>
<paragraph id="8BAAEF4D2979FF86FF1BFF23EF7FFE50" blockId="57.[151,1437,150,1509]" pageId="57" pageNumber="58">
<collectingCountry id="F302AFDD2979FF86FF1BFF23ED72FF4C" box="[199,348,151,177]" name="Bulgaria" pageId="57" pageNumber="58">BULGARIA</collectingCountry>
:
<emphasis id="B961335F2979FF86FEB9FF22EE6CFF4D" bold="true" box="[357,578,150,177]" pageId="57" pageNumber="58">S i l i s t r a Province</emphasis>
:
<specimenCount id="9D1324C42979FF86FD93FF23EE5AFF4C" box="[591,628,151,177]" pageId="57" pageNumber="58" type="male">1♂</specimenCount>
, ‘BG, Nova Cherna Vill. Ornithological station
<date id="FFABC98D2979FF86FB5EFF23E900FF4C" box="[1154,1326,151,177]" pageId="57" pageNumber="58" value="2007-06-19" valueMax="2007-06-20" valueMin="2007-06-19">19–20.VI.2007</date>
, light trap B. Guéorguiev leg.’ (
<collectionCode id="ED0477882979FF86FE5EFF0FEDC0FF28" box="[386,494,187,213]" pageId="57" pageNumber="58">NMNHS</collectionCode>
);
<specimenCount id="9D1324C42979FF86FDDDFF0FEE12FF28" box="[513,572,187,213]" pageId="57" pageNumber="58" type="male">3♂♂</specimenCount>
, ‘Srebarna Lake, wharf near the lake,
<geoCoordinate id="EE21898A2979FF86FC34FF0FE87AFF28" box="[1000,1108,187,213]" degrees="44.10331" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="1" value="44.10331">44.10331</geoCoordinate>
,
<geoCoordinate id="EE21898A2979FF86FB83FF0FE8E5FF28" box="[1119,1227,187,213]" degrees="27.06392" direction="east" orientation="longitude" pageId="57" pageNumber="58" precision="1" value="27.06392">27.06392</geoCoordinate>
,
<date id="FFABC98D2979FF86FB0AFF0FE95BFF28" box="[1238,1397,187,213]" pageId="57" pageNumber="58" value="2017-05-28" valueMax="2017-05-29" valueMin="2017-05-28">28–29.V.2017</date>
, G. Georgiev, W. Rossi &amp; D. Stoianova’ (
<collectionCode id="ED0477882979FF86FDE9FF6BEE8FFF04" box="[565,673,223,249]" pageId="57" pageNumber="58">NMNHS</collectionCode>
).
<emphasis id="B961335F2979FF86FD6EFF6BEF55FF05" bold="true" box="[690,891,222,249]" pageId="57" pageNumber="58">Va r n a Province</emphasis>
:
<specimenCount id="9D1324C42979FF86FC56FF6BEF82FF04" box="[906,940,223,249]" pageId="57" pageNumber="58" type="female">1♀</specimenCount>
, ‘БГ Крапец, Дуранкулашкο блатο; Залив! мοрски брЯГ пοд вοдοр. и камъни [Bulgaria, Krapets, Durankulak Marsh; bay! seashore under algae and stones], 23.7.92 LEG. L. Penev’ (IBER).
<emphasis id="B961335F2979FF86FDC0FE92EEF0FEBD" bold="true" box="[540,734,294,320]" pageId="57" pageNumber="58">
<collectingRegion id="49D121AF2979FF86FDC0FE92EEF0FEBD" box="[540,734,294,320]" country="Bulgaria" name="Sofiya" pageId="57" pageNumber="58">Sofia Province</collectingRegion>
</emphasis>
:
<specimenCount id="9D1324C42979FF86FD2DFE93EF3BFEBC" box="[753,789,295,321]" pageId="57" pageNumber="58" type="male">1♂</specimenCount>
,
<specimenCount id="9D1324C42979FF86FCF9FE93EF66FEBC" box="[805,840,295,321]" pageId="57" pageNumber="58" type="female">1♀</specimenCount>
, ‘BG,
<collectingRegion id="49D121AF2979FF86FC40FE93E80DFEBC" box="[924,1059,295,321]" country="Bulgaria" name="Sofiya-Grad" pageId="57" pageNumber="58">Sofia Distr.</collectingRegion>
, Dragomansko blato [Dragoman Marsh],
<geoCoordinate id="EE21898A2979FF86FF27FEFFED49FE98" box="[251,359,331,357]" degrees="42.93712" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="1" value="42.93712">42.93712</geoCoordinate>
,
<geoCoordinate id="EE21898A2979FF86FEABFEFEEDCDFE98" box="[375,483,330,357]" degrees="22.95200" direction="east" orientation="longitude" pageId="57" pageNumber="58" precision="1" value="22.952">22.95200</geoCoordinate>
,
<date id="FFABC98D2979FF86FE2EFEFFEE4AFE98" box="[498,612,331,357]" pageId="57" pageNumber="58" value="2017-06-02">2.VI.2017</date>
W. Rossi &amp; B. Guéorguiev leg.’ (
<collectionCode id="ED0477882979FF86FC22FEFFE844FE98" box="[1022,1130,331,357]" pageId="57" pageNumber="58">NMNHS</collectionCode>
).
<emphasis id="B961335F2979FF86FB5DFEFFE970FE99" bold="true" box="[1153,1374,330,357]" pageId="57" pageNumber="58">
<collectingRegion id="49D121AF2979FF86FB5DFEFFE970FE99" box="[1153,1374,330,357]" country="Bulgaria" name="Burgas" pageId="57" pageNumber="58">Burgas Province</collectingRegion>
</emphasis>
:
<specimenCount id="9D1324C42979FF86FAAEFEFFE9BBFE98" box="[1394,1429,331,357]" pageId="57" pageNumber="58" type="female">1♀</specimenCount>
, ‘Страндж еЗерο [Strandza Mt., lake], 2.VI. 9…’ (
<collectionCode id="ED0477882979FF86FD60FEDBEF06FE74" box="[700,808,367,393]" pageId="57" pageNumber="58">NMNHS</collectionCode>
);
<specimenCount id="9D1324C42979FF86FCE7FEDBEF73FE74" box="[827,861,367,393]" pageId="57" pageNumber="58" type="female">1♀</specimenCount>
, ‘BG: Pomorie bei
<collectingRegion id="49D121AF2979FF86FBE7FEDBE8A4FE74" box="[1083,1162,367,393]" country="Bulgaria" name="Burgas" pageId="57" pageNumber="58">Burgas</collectingRegion>
SO-Ufer Pomorie–See (6)
<date id="FFABC98D2979FF86FF63FE27ED1FFE50" box="[191,305,403,429]" pageId="57" pageNumber="58" value="2000-05-09">09.V.2000</date>
Handfang leg. Wolfg. Beier (Germany)’ (cWB).
</paragraph>
<paragraph id="8BAAEF4D2979FF86FF1BFE03E9B4FE2C" blockId="57.[151,1437,150,1509]" box="[199,1434,438,465]" pageId="57" pageNumber="58">
<collectingCountry id="F302AFDD2979FF86FF1BFE03ED03FE2C" box="[199,301,439,465]" name="Israel" pageId="57" pageNumber="58">ISRAEL</collectingCountry>
:
<emphasis id="B961335F2979FF86FEE4FE03EDDFFE2D" bold="true" box="[312,497,438,465]" pageId="57" pageNumber="58">North District</emphasis>
:
<specimenCount id="9D1324C42979FF86FDDCFE03EE0BFE2C" box="[512,549,439,465]" pageId="57" pageNumber="58" type="male">1♂</specimenCount>
, ‘Israel: Ne’ot Mordekhay
<date id="FFABC98D2979FF86FC8AFE02EFE7FE2C" box="[854,969,438,465]" pageId="57" pageNumber="58" value="2005-05-30">30.V.2005</date>
L. Zarabi V. Chikatunov’ (SMNH-TAU).
</paragraph>
<paragraph id="8BAAEF4D2979FF86FF1BFE6FEE42FE08" blockId="57.[151,1437,150,1509]" box="[199,620,474,501]" pageId="57" pageNumber="58">
TME:
<specimenCount id="9D1324C42979FF86FECDFE6EED82FE08" box="[273,428,474,501]" pageId="57" pageNumber="58" type="generic">15 specimens</specimenCount>
. TGE:
<specimenCount id="9D1324C42979FF86FE27FE6FEE18FE08" box="[507,566,475,501]" pageId="57" pageNumber="58" type="male">2♂♂</specimenCount>
,
<specimenCount id="9D1324C42979FF86FD9DFE6FEE48FE08" box="[577,614,475,501]" pageId="57" pageNumber="58" type="female">1♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62979FF86FF1BFE4AEEAAFDC0" pageId="57" pageNumber="58" type="diagnosis">
<paragraph id="8BAAEF4D2979FF86FF1BFE4AEEAAFDC0" blockId="57.[151,1437,150,1509]" pageId="57" pageNumber="58">
<emphasis id="B961335F2979FF86FF1BFE4AED14FDE5" bold="true" box="[199,314,510,536]" pageId="57" pageNumber="58">Diagnosis</emphasis>
. Distinguished from other
<taxonomicName id="4C1594CE2979FF86FDB7FE4BEE9DFDE4" box="[619,691,511,537]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F2979FF86FDB7FE4BEE9DFDE4" box="[619,691,511,537]" italics="true" pageId="57" pageNumber="58">Oodes</emphasis>
</taxonomicName>
species by features given in the key to species of
<taxonomicName id="4C1594CE2979FF86FB35FE4BE91FFDE4" box="[1257,1329,511,537]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F2979FF86FB35FE4BE91FFDE4" box="[1257,1329,511,537]" italics="true" pageId="57" pageNumber="58">Oodes</emphasis>
</taxonomicName>
. See also “Diagnosis” under “
<taxonomicName id="4C1594CE2979FF86FEA6FD97EDFEFDC0" authorityName="A. Villa &amp; G. B. Villa" authorityYear="1833" box="[378,464,547,573]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B961335F2979FF86FEA6FD97EDFEFDC0" box="[378,464,547,573]" italics="true" pageId="57" pageNumber="58">gracilis</emphasis>
</taxonomicName>
” species group.
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62979FF86FF1BFDF2E92AFA84" pageId="57" pageNumber="58" type="description">
<paragraph id="8BAAEF4D2979FF86FF1BFDF2E9B3FD54" blockId="57.[151,1437,150,1509]" pageId="57" pageNumber="58">
<emphasis id="B961335F2979FF86FF1BFDF2ED7FFD9D" bold="true" box="[199,337,582,608]" pageId="57" pageNumber="58">Description</emphasis>
.
<emphasis id="B961335F2979FF86FEBDFDF2ED93FD9C" bold="true" box="[353,445,582,609]" italics="true" pageId="57" pageNumber="58">Habitus</emphasis>
. Specimens of middle size (BL:
<quantity id="4CED42A82979FF86FCE9FDF3EF91FD9C" box="[821,959,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="8.85" metricValueMax="9.8" metricValueMin="7.9" pageId="57" pageNumber="58" unit="mm" value="8.85" valueMax="9.8" valueMin="7.9">7.9–9.8 mm</quantity>
, BW: 3.0–
<quantity id="4CED42A82979FF86FBE6FDF3E8BBFD9C" box="[1082,1173,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="57" pageNumber="58" unit="mm" value="3.9">3.9 mm</quantity>
), with subelongate, not very convex body (
<figureCitation id="132EF3C82979FF86FEB3FDDFEDD2FD78" box="[367,508,619,645]" captionStart="FIGURE 21" captionStartId="55.[151,250,1890,1914]" captionTargetBox="[272,1292,206,1823]" captionTargetId="figure-17@55.[221,1367,181,1866]" captionTargetPageId="55" captionText="FIGURE 21. Oodes gracilis A. Villa &amp; G.B. Villa, 1833 (A–B: neotype and its labels; C–D: female specimen, Bulgaria, Pomorie-See; E–G: male specimen, Albania, Mali i Tabaroshit).A, C: habitus; B: left protarsomeres; D: pronotum and head; E: mentum; F: prosternal process; G: left metepisternum and metacoxa (arrow shows lateral end of metacoxal basal sulcus). Scale lines = 2 mm (Figs A, C); = 0.5 mm (Figs B, D, F–G); = 0.2 mm (Fig. E)." figureDoi="http://doi.org/10.5281/zenodo.4407122" httpUri="https://zenodo.org/record/4407122/files/figure.png" pageId="57" pageNumber="58">Figs 21A, C</figureCitation>
).
<emphasis id="B961335F2979FF86FDCCFDDFEF16FD78" bold="true" box="[528,824,618,645]" italics="true" pageId="57" pageNumber="58">Ratios and measurements</emphasis>
. See
<tableCitation id="C697DAF62979FF86FCAAFDDFEFE2FD78" box="[886,972,619,645]" captionStart="TABLE 4" captionStartId="66.[151,239,1520,1544]" captionText="TABLE 4. Morphometric data for species of the genus Oodes Bonelli." pageId="57" pageNumber="58">Table 4</tableCitation>
.
<emphasis id="B961335F2979FF86FC0BFDDEE8B4FD78" bold="true" box="[983,1178,618,645]" pageId="57" pageNumber="58">
<emphasis id="B961335F2979FF86FC0BFDDEE8BDFD78" bold="true" box="[983,1171,618,645]" italics="true" pageId="57" pageNumber="58">Color and luster</emphasis>
.
</emphasis>
Body black, pronotum posterolaterally rufous, appendages piceous to rufopiceous. Integument moderately shiny, without iridescence.
</paragraph>
<paragraph id="8BAAEF4D2979FF86FF4BFD07E92AFA84" blockId="57.[151,1437,150,1509]" pageId="57" pageNumber="58">
<emphasis id="B961335F2979FF86FF4BFD07ED08FD30" bold="true" box="[151,294,691,717]" italics="true" pageId="57" pageNumber="58">Punctuation</emphasis>
. Dorsal surface without punctuation; prosternum and mesoepisternum nearly smooth, proepisternum and abdominal ventrites at sides finely rugose; abdominal ventrite 6 at apex both finely punctate and rugose; sides of metasternum and metepisternum superficially punctate.
<emphasis id="B961335F2979FF86FCF9FD4EEF4DFCE8" bold="true" box="[805,867,762,789]" italics="true" pageId="57" pageNumber="58">Head</emphasis>
. More than half as wide as pronotum (see
<tableCitation id="C697DAF62979FF86FAE6FD4FE9A1FCE8" box="[1338,1423,763,789]" captionStart="TABLE 4" captionStartId="66.[151,239,1520,1544]" captionText="TABLE 4. Morphometric data for species of the genus Oodes Bonelli." pageId="57" pageNumber="58">Table 4</tableCitation>
).
<collectingRegion id="49D121AF2979FF86FF4BFCABECD8FCC4" box="[151,246,799,825]" country="Myanmar" name="Chin" pageId="57" pageNumber="58">Mentum</collectingRegion>
tooth with distinct paramedial border (
<figureCitation id="132EF3C82979FF86FD75FCABEF20FCC4" box="[681,782,799,825]" captionStart="FIGURE 21" captionStartId="55.[151,250,1890,1914]" captionTargetBox="[272,1292,206,1823]" captionTargetId="figure-17@55.[221,1367,181,1866]" captionTargetPageId="55" captionText="FIGURE 21. Oodes gracilis A. Villa &amp; G.B. Villa, 1833 (A–B: neotype and its labels; C–D: female specimen, Bulgaria, Pomorie-See; E–G: male specimen, Albania, Mali i Tabaroshit).A, C: habitus; B: left protarsomeres; D: pronotum and head; E: mentum; F: prosternal process; G: left metepisternum and metacoxa (arrow shows lateral end of metacoxal basal sulcus). Scale lines = 2 mm (Figs A, C); = 0.5 mm (Figs B, D, F–G); = 0.2 mm (Fig. E)." figureDoi="http://doi.org/10.5281/zenodo.4407122" httpUri="https://zenodo.org/record/4407122/files/figure.png" pageId="57" pageNumber="58">Fig. 21E</figureCitation>
).
<emphasis id="B961335F2979FF86FCFDFCAAEF59FCC4" bold="true" box="[801,887,798,825]" italics="true" pageId="57" pageNumber="58">Thorax</emphasis>
. Pronotum (
<figureCitation id="132EF3C82979FF86FBDDFCABE849FCC4" box="[1025,1127,799,825]" captionStart="FIGURE 21" captionStartId="55.[151,250,1890,1914]" captionTargetBox="[272,1292,206,1823]" captionTargetId="figure-17@55.[221,1367,181,1866]" captionTargetPageId="55" captionText="FIGURE 21. Oodes gracilis A. Villa &amp; G.B. Villa, 1833 (A–B: neotype and its labels; C–D: female specimen, Bulgaria, Pomorie-See; E–G: male specimen, Albania, Mali i Tabaroshit).A, C: habitus; B: left protarsomeres; D: pronotum and head; E: mentum; F: prosternal process; G: left metepisternum and metacoxa (arrow shows lateral end of metacoxal basal sulcus). Scale lines = 2 mm (Figs A, C); = 0.5 mm (Figs B, D, F–G); = 0.2 mm (Fig. E)." figureDoi="http://doi.org/10.5281/zenodo.4407122" httpUri="https://zenodo.org/record/4407122/files/figure.png" pageId="57" pageNumber="58">Fig. 21D</figureCitation>
) with sides rounded toward posterior angles (PW/PB: 1.06–1.13); maximum width in posterior third; laterobasal impressions indistinct; base moderately sinuate; anterior angles rounded, moderately projected anteriorly. Prosternum with median longitudinal sulcus distinct, deeper than in
<taxonomicName id="4C1594CE2979FF86FE28FC3FEEB7FC58" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[500,665,907,933]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="helopioides">
<emphasis id="B961335F2979FF86FE28FC3FEE26FC58" box="[500,520,907,933]" italics="true" pageId="57" pageNumber="58">O</emphasis>
.
<emphasis id="B961335F2979FF86FDC4FC3FEEB7FC58" box="[536,665,907,933]" italics="true" pageId="57" pageNumber="58">helopioides</emphasis>
</taxonomicName>
; prosternal process with rounded apex and nearly complete border, reduced only at sides (
<figureCitation id="132EF3C82979FF86FE51FC1BEDC1FC34" box="[397,495,943,969]" captionStart="FIGURE 21" captionStartId="55.[151,250,1890,1914]" captionTargetBox="[272,1292,206,1823]" captionTargetId="figure-17@55.[221,1367,181,1866]" captionTargetPageId="55" captionText="FIGURE 21. Oodes gracilis A. Villa &amp; G.B. Villa, 1833 (A–B: neotype and its labels; C–D: female specimen, Bulgaria, Pomorie-See; E–G: male specimen, Albania, Mali i Tabaroshit).A, C: habitus; B: left protarsomeres; D: pronotum and head; E: mentum; F: prosternal process; G: left metepisternum and metacoxa (arrow shows lateral end of metacoxal basal sulcus). Scale lines = 2 mm (Figs A, C); = 0.5 mm (Figs B, D, F–G); = 0.2 mm (Fig. E)." figureDoi="http://doi.org/10.5281/zenodo.4407122" httpUri="https://zenodo.org/record/4407122/files/figure.png" pageId="57" pageNumber="58">Fig. 21F</figureCitation>
). Metepisternum longer than wide (MA/MM: 0.84–0.90), with lateral margin convex and coadunation with epipleuron very short, located anteriorly (
<figureCitation id="132EF3C82979FF86FCB3FC67EFF7FC10" box="[879,985,979,1005]" captionStart="FIGURE 21" captionStartId="55.[151,250,1890,1914]" captionTargetBox="[272,1292,206,1823]" captionTargetId="figure-17@55.[221,1367,181,1866]" captionTargetPageId="55" captionText="FIGURE 21. Oodes gracilis A. Villa &amp; G.B. Villa, 1833 (A–B: neotype and its labels; C–D: female specimen, Bulgaria, Pomorie-See; E–G: male specimen, Albania, Mali i Tabaroshit).A, C: habitus; B: left protarsomeres; D: pronotum and head; E: mentum; F: prosternal process; G: left metepisternum and metacoxa (arrow shows lateral end of metacoxal basal sulcus). Scale lines = 2 mm (Figs A, C); = 0.5 mm (Figs B, D, F–G); = 0.2 mm (Fig. E)." figureDoi="http://doi.org/10.5281/zenodo.4407122" httpUri="https://zenodo.org/record/4407122/files/figure.png" pageId="57" pageNumber="58">Fig. 21G</figureCitation>
).
<emphasis id="B961335F2979FF86FC32FC66E813FC10" bold="true" box="[1006,1085,978,1005]" pageId="57" pageNumber="58">
<emphasis id="B961335F2979FF86FC32FC66E818FC10" bold="true" box="[1006,1078,978,1005]" italics="true" pageId="57" pageNumber="58">Elytra</emphasis>
.
</emphasis>
Apical sinuation weak, poorly defined. Basal margin distinct laterally, forming a small denticle at shoulder, disappearing medially at level of stria 2. Granulation in elytral marginal furrow discontinuous, separated into shorter humeral, and longer apical group (interruption located in second fourth). Parascutellar striola punctate, less impressed than other striae; striae finely punctate anteriorly; stria 7 less impressed than stria 6. Intervals 1–7 somewhat flatter than in
<taxonomicName id="4C1594CE2979FF86FB44FBD7E914FB80" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[1176,1338,1123,1149]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="helopioides">
<emphasis id="B961335F2979FF86FB44FBD7E882FB80" box="[1176,1196,1123,1149]" italics="true" pageId="57" pageNumber="58">O</emphasis>
.
<emphasis id="B961335F2979FF86FB65FBD7E914FB80" box="[1209,1338,1123,1149]" italics="true" pageId="57" pageNumber="58">helopioides</emphasis>
</taxonomicName>
, interval 8 more convex than others.
<emphasis id="B961335F2979FF86FE0BFB33EE23FB5C" bold="true" box="[471,525,1159,1185]" italics="true" pageId="57" pageNumber="58">Legs</emphasis>
. Metacoxal basal sulcus highly reduced, ending at medial fifth. Male mesotibia not modified. Protarsomeres 1–3 of male (
<figureCitation id="132EF3C82979FF86FDACFB1FEEFDFB38" box="[624,723,1195,1221]" captionStart="FIGURE 21" captionStartId="55.[151,250,1890,1914]" captionTargetBox="[272,1292,206,1823]" captionTargetId="figure-17@55.[221,1367,181,1866]" captionTargetPageId="55" captionText="FIGURE 21. Oodes gracilis A. Villa &amp; G.B. Villa, 1833 (A–B: neotype and its labels; C–D: female specimen, Bulgaria, Pomorie-See; E–G: male specimen, Albania, Mali i Tabaroshit).A, C: habitus; B: left protarsomeres; D: pronotum and head; E: mentum; F: prosternal process; G: left metepisternum and metacoxa (arrow shows lateral end of metacoxal basal sulcus). Scale lines = 2 mm (Figs A, C); = 0.5 mm (Figs B, D, F–G); = 0.2 mm (Fig. E)." figureDoi="http://doi.org/10.5281/zenodo.4407122" httpUri="https://zenodo.org/record/4407122/files/figure.png" pageId="57" pageNumber="58">Fig. 21B</figureCitation>
) moderately dilated, with second tarsomere slightly longer than wide (W/Lp2: 0.86–0.93).
<emphasis id="B961335F2979FF86FE1DFB7AEE46FB14" bold="true" box="[449,616,1230,1257]" italics="true" pageId="57" pageNumber="58">Male genitalia</emphasis>
. Median lobe (
<figureCitation id="132EF3C82979FF86FCD3FB7BEFD6FB14" box="[783,1016,1231,1257]" captionStart="FIGURE 22" captionStartId="56.[151,250,1892,1916]" captionTargetBox="[226,1363,194,1866]" captionTargetId="figure-20@56.[220,1368,181,1869]" captionTargetPageId="56" captionText="FIGURE 22. Oodes gracilis A. Villa &amp; G.B. Villa, 1833 (A–B: neotype; C: male specimen, Bulgaria, Dragoman Marsh; D–E: male specimen, Bulgaria, Srebarna Lake; F–G: female specimens, Bulgaria, Dragoman Marsh). A, C: median lobe of aedeagus, left lateral view; B, D: same, dorsal view; E: same, ventral view; F: left gonocoxite, ventral view; G: spermathecal complex and gonocoxites, ventral view. Scale lines = 0.5 mm (Figs A–E, G); = 0.2 mm (Fig. F)." figureDoi="http://doi.org/10.5281/zenodo.4407124" httpUri="https://zenodo.org/record/4407124/files/figure.png" pageId="57" pageNumber="58">Figs 22A, B, C, D, E</figureCitation>
) with basal bulb large; angle between basal bulb and shaft nearly right; shaft long, slightly swollen compared with basal bulb; apex short, narrow; apical lamella short, bent to left, rounded at tip; ostium not reaching basal bulb.
<emphasis id="B961335F2979FF86FC10FAA2E8A1FACC" bold="true" box="[972,1167,1302,1329]" italics="true" pageId="57" pageNumber="58">Female genitalia</emphasis>
(see also
<bibRefCitation id="EF8492BC2979FF86FB22FAA3E9BEFACC" author="Ortuno, V. M." box="[1278,1424,1303,1329]" pageId="57" pageNumber="58" pagination="3 - 10" refId="ref63811" refString="Ortuno, V. M. (1998) La genitalia femenina en los Oodinae (Caraboidea: Callistidae) de la region paleartica occidental. Elytron, 12, 3 - 10." type="journal article" year="1998">Ortuño 1998</bibRefCitation>
). Basal gonocoxite with six to eight lateroapical setae arranged in line. Apical gonocoxite with single, very small dorsolateral ensiform seta and nematiform setae (
<figureCitation id="132EF3C82979FF86FD1CFAEBEF0AFA84" box="[704,804,1375,1401]" captionStart="FIGURE 22" captionStartId="56.[151,250,1892,1916]" captionTargetBox="[226,1363,194,1866]" captionTargetId="figure-20@56.[220,1368,181,1869]" captionTargetPageId="56" captionText="FIGURE 22. Oodes gracilis A. Villa &amp; G.B. Villa, 1833 (A–B: neotype; C: male specimen, Bulgaria, Dragoman Marsh; D–E: male specimen, Bulgaria, Srebarna Lake; F–G: female specimens, Bulgaria, Dragoman Marsh). A, C: median lobe of aedeagus, left lateral view; B, D: same, dorsal view; E: same, ventral view; F: left gonocoxite, ventral view; G: spermathecal complex and gonocoxites, ventral view. Scale lines = 0.5 mm (Figs A–E, G); = 0.2 mm (Fig. F)." figureDoi="http://doi.org/10.5281/zenodo.4407124" httpUri="https://zenodo.org/record/4407124/files/figure.png" pageId="57" pageNumber="58">Fig. 22F</figureCitation>
). Spermatheca twisted apically (
<figureCitation id="132EF3C82979FF86FB53FAEBE8D6FA84" box="[1167,1272,1375,1401]" captionStart="FIGURE 22" captionStartId="56.[151,250,1892,1916]" captionTargetBox="[226,1363,194,1866]" captionTargetId="figure-20@56.[220,1368,181,1869]" captionTargetPageId="56" captionText="FIGURE 22. Oodes gracilis A. Villa &amp; G.B. Villa, 1833 (A–B: neotype; C: male specimen, Bulgaria, Dragoman Marsh; D–E: male specimen, Bulgaria, Srebarna Lake; F–G: female specimens, Bulgaria, Dragoman Marsh). A, C: median lobe of aedeagus, left lateral view; B, D: same, dorsal view; E: same, ventral view; F: left gonocoxite, ventral view; G: spermathecal complex and gonocoxites, ventral view. Scale lines = 0.5 mm (Figs A–E, G); = 0.2 mm (Fig. F)." figureDoi="http://doi.org/10.5281/zenodo.4407124" httpUri="https://zenodo.org/record/4407124/files/figure.png" pageId="57" pageNumber="58">Fig. 22G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62979FF86FF1BFA36E93DFA18" pageId="57" pageNumber="58" type="distribution">
<paragraph id="8BAAEF4D2979FF86FF1BFA36E93DFA18" blockId="57.[151,1437,150,1509]" pageId="57" pageNumber="58">
<emphasis id="B961335F2979FF86FF1BFA36ED77FA61" bold="true" box="[199,345,1410,1436]" pageId="57" pageNumber="58">Distribution</emphasis>
. Most parts of Europe (except the northernmost regions, but including southern
<collectingCountry id="F302AFDD2979FF86FACCFA37E947FA60" box="[1296,1385,1411,1437]" name="Sweden" pageId="57" pageNumber="58">Sweden</collectingCountry>
and the Caucasus Major),
<collectingCountry id="F302AFDD2979FF86FE44FA13EDC7FA3C" box="[408,489,1447,1473]" name="Turkey" pageId="57" pageNumber="58">Turkey</collectingCountry>
, Transcaucasia (
<collectingCountry id="F302AFDD2979FF86FD7BFA13EF20FA3C" box="[679,782,1447,1473]" name="Armenia" pageId="57" pageNumber="58">Armenia</collectingCountry>
,
<collectingCountry id="F302AFDD2979FF86FCC7FA13EFB5FA3F" box="[795,923,1447,1474]" name="Azerbaijan" pageId="57" pageNumber="58">Azerbaijan</collectingCountry>
),
<collectingCountry id="F302AFDD2979FF86FC6DFA13EFDCFA3C" box="[945,1010,1447,1473]" name="Israel" pageId="57" pageNumber="58">Israel</collectingCountry>
,
<collectingCountry id="F302AFDD2979FF86FBDDFA13E81FFA3C" box="[1025,1073,1447,1473]" name="Iran" pageId="57" pageNumber="58">Iran</collectingCountry>
,
<collectingCountry id="F302AFDD2979FF86FBE3FA13E8F3FA3C" box="[1087,1245,1447,1473]" name="Turkmenistan" pageId="57" pageNumber="58">Turkmenistan</collectingCountry>
, western Siberia (
<bibRefCitation id="EF8492BC2979FF86FF43FA7FED6BFA18" author="Lindroth, C. H." box="[159,325,1483,1509]" pageId="57" pageNumber="58" pagination="1 - 709" refId="ref62956" refString="Lindroth, C. H. (1992) Ground Beetles (Carabidae) of Fenoscandia-A Zoogeographical Study. I &amp; III. Smithsonian Institution Libraries and The National Science Foundation, Washington, 630 pp. &amp; 814 pp. [translation by J. Adis, Erwin T. L. of Lindroth C. H. 1945. Die Fennoskandischen Carabidae. Eine tiergeographische Studie. I. Spezieller Teil. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles Handlingar, Sjatte FoGden, Series B, 4 (1), 1 - 709]" type="book chapter" year="1992">Lindroth 1992</bibRefCitation>
;
<bibRefCitation id="EF8492BC2979FF86FE8CFA7FEE58FA18" author="Kryzhanovskij, O. L. &amp; Belousov, I. A. &amp; Kabak, I. I. &amp; Kataev, M. B. &amp; Makarov, K. V. &amp; Shilenkov, V. G." box="[336,630,1482,1509]" pageId="57" pageNumber="58" refId="ref62520" refString="Kryzhanovskij, O. L., Belousov, I. A., Kabak, I. I., Kataev, M. B., Makarov, K. V. &amp; Shilenkov, V. G. (1995). A Checklist of the Ground-beetles of Russia and Adjacent Lands (Insecta, Coleoptera, Carabidae). Pensoft, Sofia &amp; Moscow, 271 pp." type="book" year="1995">
Kryzhanovskij
<emphasis id="B961335F2979FF86FE20FA7FEE03FA18" box="[508,557,1483,1509]" italics="true" pageId="57" pageNumber="58">et al</emphasis>
. 1995
</bibRefCitation>
;
<bibRefCitation id="EF8492BC2979FF86FD5EFA7FEF1EFA18" author="Bousquet, Y." box="[642,816,1483,1509]" pageId="57" pageNumber="58" pagination="1851" refId="ref60329" refString="Bousquet, Y. (2017) Tribe Oodini LaFerte-Senectere, 1851. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Revised and Updated Edition. Archostemata-Myxophaga-Adephaga. Brill, Leiden and Boston, pp. 635 - 636." type="book chapter" year="2017">Bousquet 2017</bibRefCitation>
). We record it for the first time from
<collectingCountry id="F302AFDD2979FF86FB11FA7FE921FA18" box="[1229,1295,1483,1509]" name="Israel" pageId="57" pageNumber="58">Israel</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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