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<mods:title id="F46D3072EC195FE2F71202753BB3A2C3">Revision of the Palaearctic and Oriental representatives of Lachnocrepis LeConte and Oodes Bonelli (Coleoptera: Carabidae), with special account on Chinese species</mods:title>
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<mods:namePart id="42A8F2A19D20ADE095D5233C76B268EC">Guéorguiev, Borislav</mods:namePart>
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<mods:namePart id="5FEB117CE834C9CB1C399CFAA4F08D8D">Liang, Hongbin</mods:namePart>
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<treatment id="03BC5E5B296BFF91FF4BF982ED8DFA1B" ID-DOI="http://doi.org/10.5281/zenodo.4480029" ID-GBIF-Taxon="177139770" ID-Zenodo-Dep="4480029" LSID="urn:lsid:plazi:treatment:03BC5E5B296BFF91FF4BF982ED8DFA1B" httpUri="http://treatment.plazi.org/id/03BC5E5B296BFF91FF4BF982ED8DFA1B" lastPageId="46" lastPageNumber="47" pageId="43" pageNumber="44">
<subSubSection id="C30FBCC6296BFF94FF4BF982EF8BF94A" pageId="43" pageNumber="44" type="diagnosis">
<subSubSection id="C30FBCC6296BFF94FF4BF982EED9F9AD" box="[151,759,1590,1617]" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="8BAAEF4D296BFF94FF4BF982EED9F9AD" blockId="43.[151,759,1590,1653]" box="[151,759,1590,1617]" pageId="43" pageNumber="44">
<emphasis id="B961335F296BFF94FF4BF982EED9F9AD" bold="true" box="[151,759,1590,1617]" pageId="43" pageNumber="44">
13.
<taxonomicName id="4C1594CE296BFF94FF1FF982EED9F9AD" ID-CoL="49S3D" authority="Habu &amp; Baba, 1960" authorityName="Habu &amp; Baba" authorityYear="1960" box="[195,759,1590,1617]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="echigonus" subGenus="Oodes">
<emphasis id="B961335F296BFF94FF1FF982ED20F9AC" bold="true" box="[195,270,1590,1617]" italics="true" pageId="43" pageNumber="44">Oodes</emphasis>
(
<emphasis id="B961335F296BFF94FEC3F982ED44F9AC" bold="true" box="[287,362,1590,1617]" italics="true" pageId="43" pageNumber="44">Oodes</emphasis>
)
<emphasis id="B961335F296BFF94FEA7F982EDD9F9AC" bold="true" box="[379,503,1590,1617]" italics="true" pageId="43" pageNumber="44">echigonus</emphasis>
<bibRefCitation id="EF8492BC296BFF94FE22F983EED9F9AD" author="Habu, A. &amp; Baba, K." box="[510,759,1590,1617]" pageId="43" pageNumber="44" pagination="31 - 33" refId="ref61814" refString="Habu, A. &amp; Baba, K. (1960) One new species of Oodes from Niigata Prefecture, Japan (Coleoptera, Carabidae). Akitu, 9, 31 - 33." type="journal article" year="1960">Habu &amp; Baba, 1960</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC6296BFF94FF4BF9EFEE0AF988" box="[151,548,1627,1653]" pageId="43" pageNumber="44" type="description">
<paragraph id="8BAAEF4D296BFF94FF4BF9EFEE0AF988" blockId="43.[151,759,1590,1653]" box="[151,548,1627,1653]" pageId="43" pageNumber="44">
(
<figureCitation id="132EF3C8296BFF94FF43F9EFECDCF988" box="[159,242,1627,1653]" captionStart="FIGURE 16" captionStartId="44.[151,250,1896,1920]" captionTargetBox="[259,1335,198,1838]" captionTargetId="figure-20@44.[219,1369,181,1872]" captionTargetPageId="44" captionText="FIGURE 16. Oodes echigonus Habu &amp; Baba, 1960 (A: topotype male specimen, Japan, Niigata Pref., Katamachi; BC:topotype female specimen, same locality; DE: male specimen, China, imprecise locality; FG: female specimen, China, imprecise locality). AB, D, F: habitus; C: mentum; E: prosternal process; G: left metepisternum and metacoxa. Scale lines = 2 mm (Figs AB, D, F); = 0.5 mm (Figs C); = 1 mm (Figs E, G)." figureDoi="http://doi.org/10.5281/zenodo.4407108" httpUri="https://zenodo.org/record/4407108/files/figure.png" pageId="43" pageNumber="44">Figs 16</figureCitation>
AG,
<figureCitation id="132EF3C8296BFF94FEE4F9EFEDA6F988" box="[312,392,1627,1653]" captionStart="FIGURE 17" captionStartId="45.[151,250,1910,1934]" captionTargetBox="[223,1364,191,1881]" captionTargetId="figure-17@45.[214,1374,181,1886]" captionTargetPageId="45" captionText="FIGURE 17. Oodes echigonus Habu &amp; Baba, 1960 (AB: topotype male specimen, Japan, Niigata Pref., Katamachi; CD: male specimen, China, imprecise locality; EF: topotype female specimen, same locality as in the male topotype).A, C: median lobe of aedeagus, left lateral view; B: same, dorsal view; D: same, ventral view; E: left gonocoxite, ventral view; F: spermathecal complex and gonocoxites, ventral view. Scale lines = 0.5 mm (Figs AD, F); = 0.2 mm (Fig. E)." figureDoi="http://doi.org/10.5281/zenodo.4407112" httpUri="https://zenodo.org/record/4407112/files/figure.png" pageId="43" pageNumber="44">Figs 17</figureCitation>
AF,
<tableCitation id="C697DAF6296BFF94FE14F9EFEE32F988" box="[456,540,1627,1653]" captionStart="TABLE 4" captionStartId="66.[151,239,1520,1544]" captionText="TABLE 4. Morphometric data for species of the genus Oodes Bonelli." pageId="43" pageNumber="44">Table 4</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC6296BFF94FF4BF92BEF8BF94A" box="[151,933,1695,1719]" pageId="43" pageNumber="44" type="reference_group">
<paragraph id="8BAAEF4D296BFF94FF4BF92BEF8BF94A" blockId="43.[151,933,1695,1719]" box="[151,933,1695,1719]" pageId="43" pageNumber="44">
<taxonomicName id="4C1594CE296BFF94FF4BF92BEE38F94A" ID-CoL="49S3D" authority="Habu &amp; Baba, 1960" authorityName="Habu &amp; Baba" authorityYear="1960" box="[151,534,1695,1719]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="echigonus">
<emphasis id="B961335F296BFF94FF4BF92BED6BF94A" box="[151,325,1695,1719]" italics="true" pageId="43" pageNumber="44">Oodes echigonus</emphasis>
<materialsCitation id="3B7DE510296BFF94FE97F92BEE38F94A" ID-GBIF-Occurrence="3027186301" box="[331,534,1695,1719]" location="Katamachi" pageId="43" pageNumber="44" specimenCount="1" stateProvince="Niigata">
<bibRefCitation id="EF8492BC296BFF94FE97F92BEE38F94A" author="Habu, A. &amp; Baba, K." box="[331,534,1695,1719]" pageId="43" pageNumber="44" pagination="31 - 33" refId="ref61814" refString="Habu, A. &amp; Baba, K. (1960) One new species of Oodes from Niigata Prefecture, Japan (Coleoptera, Carabidae). Akitu, 9, 31 - 33." type="journal article" year="1960">Habu &amp; Baba, 1960</bibRefCitation>
</materialsCitation>
</taxonomicName>
(
<typeStatus id="54AE51EF296BFF94FDFFF92BEE7EF94A" box="[547,592,1695,1719]" pageId="43" pageNumber="44">type</typeStatus>
loc.:
<location id="8ECAB996296BFF94FD51F92BEF2CF94A" LSID="urn:lsid:plazi:treatment:03BC5E5B296BFF91FF4BF982ED8DFA1B:8ECAB996296BFF94FD51F92BEF2CF94A" box="[653,770,1695,1719]" name="Katamachi" pageId="43" pageNumber="44" stateProvince="Niigata">Katamachi</location>
,
<collectingRegion id="49D121AF296BFF94FCD0F92BEFB8F94A" box="[780,918,1695,1719]" country="Japan" name="Niigata" pageId="43" pageNumber="44">Niigata Pref.</collectingRegion>
).
</paragraph>
</subSubSection>
</subSubSection>
<subSubSection id="C30FBCC6296BFF94FF4BF968ED3DF909" box="[151,275,1756,1780]" pageId="43" pageNumber="44" type="multiple">
<paragraph id="8BAAEF4D296BFF94FF4BF968ED3DF909" blockId="43.[151,1436,1756,1873]" box="[151,275,1756,1780]" pageId="43" pageNumber="44">
<heading id="D0E25821296BFF94FF4BF968ED3DF909" bold="true" box="[151,275,1756,1780]" fontSize="10" level="2" pageId="43" pageNumber="44" reason="2">
<emphasis id="B961335F296BFF94FF4BF968ED23F909" bold="true" box="[151,269,1756,1780]" pageId="43" pageNumber="44">References</emphasis>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC6296BFF94FF4BF94FEF13F8AD" pageId="43" pageNumber="44" type="reference_group">
<paragraph id="8BAAEF4D296BFF94FF4BF94FEF13F8AD" blockId="43.[151,1436,1756,1873]" pageId="43" pageNumber="44">
<taxonomicName id="4C1594CE296BFF94FF4BF94FEE0EF8EE" ID-CoL="49S3D" authority=": Kasahara 1994: 94" authorityName="Kasahara" authorityPageNumber="94" authorityYear="1994" box="[151,544,1787,1811]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="echigonus">
<emphasis id="B961335F296BFF94FF4BF94FED67F8EE" box="[151,329,1787,1811]" italics="true" pageId="43" pageNumber="44">Oodes echigonus</emphasis>
:
<bibRefCitation id="EF8492BC296BFF94FE87F94FEE0EF8EE" author="Kasahara, S." box="[347,544,1787,1811]" pageId="43" pageNumber="44" pagination="94" refId="ref62348" refString="Kasahara, S. (1994) Oodes echigonus (Coleoptera, Carabidae) Found in Kwanto District, Central Honshu, Japan. Elytra, 22 (1), 94." type="journal article" year="1994">Kasahara 1994: 94</bibRefCitation>
</taxonomicName>
(distribution in
<collectingCountry id="F302AFDD296BFF94FD0DF94FEF21F8EE" box="[721,783,1787,1811]" name="Japan" pageId="43" pageNumber="44">Japan</collectingCountry>
);
<bibRefCitation id="EF8492BC296BFF94FCF8F948EFD7F8EE" author="Bousquet, Y." box="[804,1017,1787,1811]" pageId="43" pageNumber="44" pagination="443 - 537" refId="ref60231" refString="Bousquet, Y. (1996) Taxonomic revision of Nearctic, Mexican, and West Indian Oodini (Coleoptera: Carabidae). The Canadian Entomologist, 128, 443 - 537. https: // doi. org / 10.4039 / Ent 128443 - 3" type="journal article" year="1996">Bousquet 1996: 474</bibRefCitation>
(relationships with Nearctic congeners, distribution in
<collectingCountry id="F302AFDD296BFF94FE87F8AEEDB5F8CF" box="[347,411,1818,1842]" name="China" pageId="43" pageNumber="44">China</collectingCountry>
: Nanking, Kiangsu province);
<bibRefCitation id="EF8492BC296BFF94FD03F8AEEFBBF8CF" author="Lorenz, W." box="[735,917,1818,1842]" pageId="43" pageNumber="44" refId="ref63123" refString="Lorenz, W. (1998) Systematic list of extant ground beetles of the world (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 1 st Edition. Published by the Author, Tutzing, [ii] + 502 pp." type="book" year="1998">Lorenz 1998: 305</bibRefCitation>
;
<bibRefCitation id="EF8492BC296BFF94FC42F8AEE842F8CF" author="Bousquet, Y." box="[926,1132,1818,1842]" pageId="43" pageNumber="44" pagination="1851" refId="ref60278" refString="Bousquet, Y. (2003) Tribe Oodini LaFerte-Senectere, 1851. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 444 - 445." type="book chapter" year="2003">Bousquet 2003: 445</bibRefCitation>
;
<bibRefCitation id="EF8492BC296BFF94FBA9F8AEE905F8CF" author="Lorenz, W." box="[1141,1323,1818,1842]" pageId="43" pageNumber="44" refId="ref63176" refString="Lorenz, W. (2005) Systematic list of extant ground beetles of the world (Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Published by the author, Tutzing, [ii] + 530 pp." type="book" year="2005">Lorenz 2005: 325</bibRefCitation>
;
<bibRefCitation id="EF8492BC296BFF94FAE8F8AEED77F8AC" author="Hasegawa, M. &amp; Kanie, N. &amp; Toda, N." pageId="43" pageNumber="44" pagination="21 - 24" refId="ref61948" refString="Hasegawa, M., Kanie, N. &amp; Toda, N. (2015) Oodine carabid beetles (Coleoptera, Carabidae) from Aichi Prefecture, central Japan. Science Report of the Toyohashi Museum of Natural History, 25, 21 - 24. [in Japanese]" type="journal article" year="2015">
Hasegawa
<emphasis id="B961335F296BFF94FF1BF88DECDAF8AD" box="[199,244,1848,1872]" italics="true" pageId="43" pageNumber="44">et al</emphasis>
. 2015: 22
</bibRefCitation>
(
<collectingCountry id="F302AFDD296BFF94FEBBF88DED8BF8AC" box="[359,421,1849,1873]" name="Japan" pageId="43" pageNumber="44">Japan</collectingCountry>
,
<collectingRegion id="49D121AF296BFF94FE71F88CEE77F8AD" box="[429,601,1848,1872]" country="Japan" name="Aichi" pageId="43" pageNumber="44">Aichi Prefecture</collectingRegion>
);
<bibRefCitation id="EF8492BC296BFF94FDB6F88DEF17F8AD" author="Bousquet, Y." box="[618,825,1848,1873]" pageId="43" pageNumber="44" pagination="1851" refId="ref60329" refString="Bousquet, Y. (2017) Tribe Oodini LaFerte-Senectere, 1851. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Revised and Updated Edition. Archostemata-Myxophaga-Adephaga. Brill, Leiden and Boston, pp. 635 - 636." type="book chapter" year="2017">Bousquet 2017: 636</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC6296BFF91FF4BF8CBEE42FE98" lastPageId="46" lastPageNumber="47" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="8BAAEF4D296BFF94FF4BF8CBE944F843" blockId="43.[151,1437,1918,1982]" pageId="43" pageNumber="44">
<emphasis id="B961335F296BFF94FF4BF8CBED6DF865" bold="true" box="[151,323,1918,1945]" pageId="43" pageNumber="44">Type material</emphasis>
. Not examined.
<typeStatus id="54AE51EF296BFF94FDD7F8CBEE5DF864" box="[523,627,1919,1945]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
preserved in the Insect Museum of the National Institute for AgroEnvironmental Science, Tsukuba (https://www.naro.affrc.go.jp/org/niaes/type/dbcarabidae/o_echigonus.html).
</paragraph>
<caption id="DF6ABFC5296CFF93FF4BF8DCEE80F810" ID-DOI="http://doi.org/10.5281/zenodo.4407108" ID-Zenodo-Dep="4407108" httpUri="https://zenodo.org/record/4407108/files/figure.png" pageId="44" pageNumber="45" startId="44.[151,250,1896,1920]" targetBox="[259,1335,198,1838]" targetPageId="44">
<paragraph id="8BAAEF4D296CFF93FF4BF8DCEE80F810" blockId="44.[151,1436,1896,2029]" pageId="44" pageNumber="45">
<emphasis id="B961335F296CFF93FF4BF8DCED30F87D" bold="true" box="[151,286,1896,1920]" pageId="44" pageNumber="45">FIGURE 16.</emphasis>
<taxonomicName id="4C1594CE296CFF93FEFFF8DDEEB7F87D" authority="Habu &amp; Baba, 1960" authorityName="Habu &amp; Baba" authorityYear="1960" box="[291,665,1896,1921]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="echigonus">
<emphasis id="B961335F296CFF93FEFFF8DDEDE1F87C" box="[291,463,1897,1921]" italics="true" pageId="44" pageNumber="45">Oodes echigonus</emphasis>
<bibRefCitation id="EF8492BC296CFF93FE0FF8DDEEB7F87D" author="Habu, A. &amp; Baba, K." box="[467,665,1896,1921]" pageId="44" pageNumber="45" pagination="31 - 33" refId="ref61814" refString="Habu, A. &amp; Baba, K. (1960) One new species of Oodes from Niigata Prefecture, Japan (Coleoptera, Carabidae). Akitu, 9, 31 - 33." type="journal article" year="1960">Habu &amp; Baba, 1960</bibRefCitation>
</taxonomicName>
(A: topotype male specimen, Japan, Niigata Pref., Katamachi; BC: topotype female specimen, same locality; DE: male specimen, China, imprecise locality; FG: female specimen, China, imprecise locality). AB, D, F: habitus; C: mentum; E: prosternal process; G: left metepisternum and metacoxa. Scale lines = 2 mm (Figs AB, D, F); = 0.5 mm (Figs C); = 1 mm (Figs E, G).
</paragraph>
</caption>
<caption id="DF6ABFC5296DFF92FF4BF8C2E84AF807" ID-DOI="http://doi.org/10.5281/zenodo.4407112" ID-Zenodo-Dep="4407112" httpUri="https://zenodo.org/record/4407112/files/figure.png" pageId="45" pageNumber="46" startId="45.[151,250,1910,1934]" targetBox="[223,1364,191,1881]" targetPageId="45">
<paragraph id="8BAAEF4D296DFF92FF4BF8C2E84AF807" blockId="45.[151,1437,1910,2043]" pageId="45" pageNumber="46">
<emphasis id="B961335F296DFF92FF4BF8C2ED30F873" bold="true" box="[151,286,1910,1934]" pageId="45" pageNumber="46">FIGURE 17.</emphasis>
<taxonomicName id="4C1594CE296DFF92FEF8F8C2EEB2F873" authority="Habu &amp; Baba, 1960" authorityName="Habu &amp; Baba" authorityYear="1960" box="[292,668,1910,1934]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="echigonus">
<emphasis id="B961335F296DFF92FEF8F8C2EDFEF873" box="[292,464,1910,1934]" italics="true" pageId="45" pageNumber="46">Oodes echigonus</emphasis>
<bibRefCitation id="EF8492BC296DFF92FE09F8C2EEB2F873" author="Habu, A. &amp; Baba, K." box="[469,668,1910,1934]" pageId="45" pageNumber="46" pagination="31 - 33" refId="ref61814" refString="Habu, A. &amp; Baba, K. (1960) One new species of Oodes from Niigata Prefecture, Japan (Coleoptera, Carabidae). Akitu, 9, 31 - 33." type="journal article" year="1960">Habu &amp; Baba, 1960</bibRefCitation>
</taxonomicName>
(AB: topotype male specimen, Japan, Niigata Pref., Katamachi; CD: male specimen, China, imprecise locality; EF: topotype female specimen, same locality as in the male topotype). A, C: median lobe of aedeagus, left lateral view; B: same, dorsal view; D: same, ventral view; E: left gonocoxite, ventral view; F: spermathecal complex and gonocoxites, ventral view. Scale lines = 0.5 mm (Figs AD, F); = 0.2 mm (Fig. E).
</paragraph>
</caption>
<paragraph id="8BAAEF4D296EFF91FF1BFF23EDD5FF04" blockId="46.[151,1437,150,1510]" pageId="46" pageNumber="47">
<emphasis id="B961335F296EFF91FF1BFF23ED8EFF4D" bold="true" box="[199,416,150,177]" pageId="46" pageNumber="47">Topotype material</emphasis>
.
<collectingCountry id="F302AFDD296EFF91FE77FF23EE2AFF4C" box="[427,516,151,177]" name="Japan" pageId="46" pageNumber="47">JAPAN</collectingCountry>
:
<emphasis id="B961335F296EFF91FDD0FF23EED6FF4D" bold="true" box="[524,760,150,177]" pageId="46" pageNumber="47">N i i g a t a Prefecture</emphasis>
:
<specimenCount id="9D1324C4296EFF91FCD8FF23EF11FF4C" box="[772,831,151,177]" pageId="46" pageNumber="47" type="male">2♂♂</specimenCount>
,
<specimenCount id="9D1324C4296EFF91FC94FF23EF45FF4C" box="[840,875,151,177]" pageId="46" pageNumber="47" type="female">1♀</specimenCount>
,
<collectingRegion id="49D121AF296EFF91FCA7FF23E83EFF4C" box="[891,1040,151,177]" country="Japan" name="Niigata" pageId="46" pageNumber="47">Niigata Pref.</collectingRegion>
, Jyôetsu-shi, Ookata-ku, Katamachi
<date id="FFABC98D296EFF91FF4BFF0FED25FF28" box="[151,267,187,213]" pageId="46" pageNumber="47" value="1976-09-03">3.IX.1976</date>
, Seiji Morita leg. (
<collectionCode id="ED047788296EFF91FE31FF0FEE77FF28" box="[493,601,187,213]" pageId="46" pageNumber="47">NMNHS</collectionCode>
);
<specimenCount id="9D1324C4296EFF91FDB2FF0FEEBCFF28" box="[622,658,187,213]" pageId="46" pageNumber="47" type="male">1♂</specimenCount>
, same data, but collected
<date id="FFABC98D296EFF91FC63FF0EE81FFF28" box="[959,1073,186,213]" pageId="46" pageNumber="47" value="1991-05-25">25.V.1991</date>
(
<collectionCode id="ED047788296EFF91FB9EFF0FE880FF28" box="[1090,1198,187,213]" pageId="46" pageNumber="47">NMNHS</collectionCode>
);
<specimenCount id="9D1324C4296EFF91FB1FFF0FE8C8FF28" box="[1219,1254,187,213]" pageId="46" pageNumber="47" type="female">1♀</specimenCount>
, same data, but collected
<date id="FFABC98D296EFF91FEDFFF6BED5BFF04" box="[259,373,223,249]" pageId="46" pageNumber="47" value="1991-06-02">2.VI.1991</date>
(
<collectionCode id="ED047788296EFF91FE58FF6BEDDEFF04" box="[388,496,223,249]" pageId="46" pageNumber="47">NMNHS</collectionCode>
).
</paragraph>
<paragraph id="8BAAEF4D296EFF91FF1BFEB6EF75FEBC" blockId="46.[151,1437,150,1510]" pageId="46" pageNumber="47">
<emphasis id="B961335F296EFF91FF1BFEB6EDD6FEE1" bold="true" box="[199,504,258,284]" pageId="46" pageNumber="47">Other material examined</emphasis>
.
<collectingCountry id="F302AFDD296EFF91FDD4FEB7EE4BFEE0" box="[520,613,259,285]" name="China" pageId="46" pageNumber="47">CHINA</collectingCountry>
:
<emphasis id="B961335F296EFF91FDAEFEB7EF5CFEE1" bold="true" box="[626,882,258,285]" pageId="46" pageNumber="47">I m p re c i s e localities</emphasis>
:
<specimenCount id="9D1324C4296EFF91FC5FFEB7EF93FEE0" box="[899,957,259,285]" pageId="46" pageNumber="47" type="male">8♂♂</specimenCount>
,
<specimenCount id="9D1324C4296EFF91FC16FEB7E82DFEE0" box="[970,1027,259,285]" pageId="46" pageNumber="47" type="female">8♀♀</specimenCount>
, China (
<collectionCode id="ED047788296EFF91FBBAFEB7E886FEE0" box="[1126,1192,259,285]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="46" pageNumber="47">IZAS</collectionCode>
,
<collectionCode id="ED047788296EFF91FB6AFEB7E90EFEE0" box="[1206,1312,259,285]" pageId="46" pageNumber="47">NMNHS</collectionCode>
).
<emphasis id="B961335F296EFF91FAE8FEB7ED2EFEBD" bold="true" pageId="46" pageNumber="47">J i a n g x i Province</emphasis>
:
<specimenCount id="9D1324C4296EFF91FED3FE93ED1CFEBC" box="[271,306,295,321]" pageId="46" pageNumber="47" type="female">1♀</specimenCount>
,
<collectingRegion id="49D121AF296EFF91FE9CFE93EDBAFEBC" box="[320,404,295,321]" country="China" name="Jiangxi" pageId="46" pageNumber="47">Jiangxi</collectingRegion>
, Liantang, field margin, 1953.8. (
<collectionCode id="ED047788296EFF91FCD7FE93EF61FEBC" box="[779,847,295,321]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="46" pageNumber="47">IZAS</collectionCode>
).
</paragraph>
<paragraph id="8BAAEF4D296EFF91FF1BFEFFEE42FE98" blockId="46.[151,1437,150,1510]" box="[199,620,331,357]" pageId="46" pageNumber="47">
TME:
<specimenCount id="9D1324C4296EFF91FECDFEFFED82FE98" box="[273,428,331,357]" pageId="46" pageNumber="47" type="generic">22 specimens</specimenCount>
. TGE:
<specimenCount id="9D1324C4296EFF91FE27FEFFEE18FE98" box="[507,566,331,357]" pageId="46" pageNumber="47" type="male">3♂♂</specimenCount>
,
<specimenCount id="9D1324C4296EFF91FD9DFEFFEE48FE98" box="[577,614,331,357]" pageId="46" pageNumber="47" type="female">1♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC6296EFF91FF1BFEDAEF4FFE50" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="8BAAEF4D296EFF91FF1BFEDAEF4FFE50" blockId="46.[151,1437,150,1510]" pageId="46" pageNumber="47">
<emphasis id="B961335F296EFF91FF1BFEDAED14FE75" bold="true" box="[199,314,366,392]" pageId="46" pageNumber="47">Diagnosis</emphasis>
. Members of this species are readily distinguished from those of other Palaearctic
<taxonomicName id="4C1594CE296EFF91FB2FFEDBE915FE74" box="[1267,1339,367,393]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F296EFF91FB2FFEDBE915FE74" box="[1267,1339,367,393]" italics="true" pageId="46" pageNumber="47">Oodes</emphasis>
</taxonomicName>
by their larger size, more than
<quantity id="4CED42A8296EFF91FE4CFE26EDD8FE50" box="[400,502,402,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="46" pageNumber="47" unit="mm" value="12.5">12.5 mm</quantity>
, and very low PA/PW (
<tableCitation id="C697DAF6296EFF91FD20FE27EF7AFE50" box="[764,852,403,429]" captionStart="TABLE 4" captionStartId="66.[151,239,1520,1544]" captionText="TABLE 4. Morphometric data for species of the genus Oodes Bonelli." pageId="46" pageNumber="47">Table 4</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC6296EFF91FF1BFE02EFFBFB5C" pageId="46" pageNumber="47" type="description">
<paragraph id="8BAAEF4D296EFF91FF1BFE02EFFBFB5C" blockId="46.[151,1437,150,1510]" pageId="46" pageNumber="47">
<emphasis id="B961335F296EFF91FF1BFE02ED7FFE2D" bold="true" box="[199,337,438,464]" pageId="46" pageNumber="47">Description</emphasis>
.
<emphasis id="B961335F296EFF91FEBBFE02EDEDFE2C" bold="true" box="[359,451,438,465]" italics="true" pageId="46" pageNumber="47">Habitus</emphasis>
. Specimens of large size (BL: 12.815.0 mm, BW:
<quantity id="4CED42A8296EFF91FB9AFE02E8F6FE2C" box="[1094,1240,438,465]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="6.2" metricValueMin="5.3" pageId="46" pageNumber="47" unit="mm" value="5.75" valueMax="6.2" valueMin="5.3">5.36.2 mm</quantity>
), with subovate, moderately convex body (
<figureCitation id="132EF3C8296EFF91FE18FE6FEEB3FE08" box="[452,669,475,501]" captionStart="FIGURE 16" captionStartId="44.[151,250,1896,1920]" captionTargetBox="[259,1335,198,1838]" captionTargetId="figure-20@44.[219,1369,181,1872]" captionTargetPageId="44" captionText="FIGURE 16. Oodes echigonus Habu &amp; Baba, 1960 (A: topotype male specimen, Japan, Niigata Pref., Katamachi; BC:topotype female specimen, same locality; DE: male specimen, China, imprecise locality; FG: female specimen, China, imprecise locality). AB, D, F: habitus; C: mentum; E: prosternal process; G: left metepisternum and metacoxa. Scale lines = 2 mm (Figs AB, D, F); = 0.5 mm (Figs C); = 1 mm (Figs E, G)." figureDoi="http://doi.org/10.5281/zenodo.4407108" httpUri="https://zenodo.org/record/4407108/files/figure.png" pageId="46" pageNumber="47">Figs 16A, B, D, F</figureCitation>
).
<emphasis id="B961335F296EFF91FD6FFE6FEFCCFE08" bold="true" box="[691,994,474,501]" italics="true" pageId="46" pageNumber="47">Ratios and measurements</emphasis>
. See
<tableCitation id="C697DAF6296EFF91FBFAFE6FE850FE08" box="[1062,1150,475,501]" captionStart="TABLE 4" captionStartId="66.[151,239,1520,1544]" captionText="TABLE 4. Morphometric data for species of the genus Oodes Bonelli." pageId="46" pageNumber="47">Table 4</tableCitation>
.
<emphasis id="B961335F296EFF91FB50FE6EE978FE08" bold="true" box="[1164,1366,474,501]" pageId="46" pageNumber="47">
<emphasis id="B961335F296EFF91FB50FE6EE960FE08" bold="true" box="[1164,1358,474,501]" italics="true" pageId="46" pageNumber="47">Color and luster</emphasis>
.
</emphasis>
Body including palpi and legs black; antennae piceous to rufopiceous. Integument dorsally and ventrally moderately shiny, without iridescence.
<emphasis id="B961335F296EFF91FE19FD97EE7AFDC0" bold="true" box="[453,596,547,573]" italics="true" pageId="46" pageNumber="47">Punctuation</emphasis>
. Dorsal surface without punctuation; sides of prosternum and metasternum, meso- and metepisternum densely punctate; proepisternum finely rugose; abdominal ventrites finely rugose and punctured.
<emphasis id="B961335F296EFF91FEC4FDDEED78FD78" bold="true" box="[280,342,618,645]" italics="true" pageId="46" pageNumber="47">Head</emphasis>
. Half or less than half as wide as pronotum (PW/HW: 2.002.16).
<collectingRegion id="49D121AF296EFF91FBBFFDDFE8ECFD78" box="[1123,1218,619,645]" country="Myanmar" name="Chin" pageId="46" pageNumber="47">Mentum</collectingRegion>
tooth with distinct paramedial border (
<figureCitation id="132EF3C8296EFF91FEAFFD3BEDF4FD57" box="[371,474,655,682]" captionStart="FIGURE 16" captionStartId="44.[151,250,1896,1920]" captionTargetBox="[259,1335,198,1838]" captionTargetId="figure-20@44.[219,1369,181,1872]" captionTargetPageId="44" captionText="FIGURE 16. Oodes echigonus Habu &amp; Baba, 1960 (A: topotype male specimen, Japan, Niigata Pref., Katamachi; BC:topotype female specimen, same locality; DE: male specimen, China, imprecise locality; FG: female specimen, China, imprecise locality). AB, D, F: habitus; C: mentum; E: prosternal process; G: left metepisternum and metacoxa. Scale lines = 2 mm (Figs AB, D, F); = 0.5 mm (Figs C); = 1 mm (Figs E, G)." figureDoi="http://doi.org/10.5281/zenodo.4407108" httpUri="https://zenodo.org/record/4407108/files/figure.png" pageId="46" pageNumber="47">Fig. 16C</figureCitation>
).
<emphasis id="B961335F296EFF91FE31FD3AEE6DFD54" bold="true" box="[493,579,654,681]" italics="true" pageId="46" pageNumber="47">Thorax</emphasis>
. Pronotum with sides rounded toward posterior angles (PW/PB: 1.021.06); maximum width in posterior third; laterobasal impressions indistinct; base moderately sinuate; anterior angles rounded, moderately projected anteriorly. Prosternum with median longitudinal sulcus shallow; prosternal process with apex rounded and border nearly complete, reduced at apex (
<figureCitation id="132EF3C8296EFF91FC43FD4FE827FCEB" box="[927,1033,763,790]" captionStart="FIGURE 16" captionStartId="44.[151,250,1896,1920]" captionTargetBox="[259,1335,198,1838]" captionTargetId="figure-20@44.[219,1369,181,1872]" captionTargetPageId="44" captionText="FIGURE 16. Oodes echigonus Habu &amp; Baba, 1960 (A: topotype male specimen, Japan, Niigata Pref., Katamachi; BC:topotype female specimen, same locality; DE: male specimen, China, imprecise locality; FG: female specimen, China, imprecise locality). AB, D, F: habitus; C: mentum; E: prosternal process; G: left metepisternum and metacoxa. Scale lines = 2 mm (Figs AB, D, F); = 0.5 mm (Figs C); = 1 mm (Figs E, G)." figureDoi="http://doi.org/10.5281/zenodo.4407108" httpUri="https://zenodo.org/record/4407108/files/figure.png" pageId="46" pageNumber="47">Fig. 16E</figureCitation>
). Metepisternum longer than wide (MA/MM: 0.871.00), with lateral margin convex and coadunation with epipleuron rather long, located anteriorly (
<figureCitation id="132EF3C8296EFF91FF7CFCF7ED24FCA3" box="[160,266,835,862]" captionStart="FIGURE 16" captionStartId="44.[151,250,1896,1920]" captionTargetBox="[259,1335,198,1838]" captionTargetId="figure-20@44.[219,1369,181,1872]" captionTargetPageId="44" captionText="FIGURE 16. Oodes echigonus Habu &amp; Baba, 1960 (A: topotype male specimen, Japan, Niigata Pref., Katamachi; BC:topotype female specimen, same locality; DE: male specimen, China, imprecise locality; FG: female specimen, China, imprecise locality). AB, D, F: habitus; C: mentum; E: prosternal process; G: left metepisternum and metacoxa. Scale lines = 2 mm (Figs AB, D, F); = 0.5 mm (Figs C); = 1 mm (Figs E, G)." figureDoi="http://doi.org/10.5281/zenodo.4407108" httpUri="https://zenodo.org/record/4407108/files/figure.png" pageId="46" pageNumber="47">Fig. 16G</figureCitation>
).
<emphasis id="B961335F296EFF91FEC3FCF6ED43FCA0" bold="true" box="[287,365,834,861]" pageId="46" pageNumber="47">
<emphasis id="B961335F296EFF91FEC3FCF6ED49FCA0" bold="true" box="[287,359,834,861]" italics="true" pageId="46" pageNumber="47">Elytra</emphasis>
.
</emphasis>
Apical sinuation weak, but distinct. Basal margin distinct laterally, forming a minute denticle at shoulder, disappearing medially at level between striae 2 and 3. Granulation in marginal furrow continuous. Parascutellar striola and stria 1 shallower than other striae, striola punctate as well as striae 14 anteriorly; stria 7 as distinct as stria 6. Intervals 17 rather flat, interval 8 convex.
<emphasis id="B961335F296EFF91FCACFC1BEF88FC34" bold="true" box="[880,934,943,969]" italics="true" pageId="46" pageNumber="47">Legs</emphasis>
. Metacoxal basal sulcus extending to lateral fourth. Male mesotibia not modified. Protarsomeres 13 of male moderately dilated, with second segment longer than wide (W/Lp2: 0.820.88).
<emphasis id="B961335F296EFF91FE26FC42EE8FFBEC" bold="true" box="[506,673,1014,1041]" italics="true" pageId="46" pageNumber="47">Male genitalia</emphasis>
. Median lobe (
<figureCitation id="132EF3C8296EFF91FC96FC43E838FBEC" box="[842,1046,1015,1041]" captionStart="FIGURE 17" captionStartId="45.[151,250,1910,1934]" captionTargetBox="[223,1364,191,1881]" captionTargetId="figure-17@45.[214,1374,181,1886]" captionTargetPageId="45" captionText="FIGURE 17. Oodes echigonus Habu &amp; Baba, 1960 (AB: topotype male specimen, Japan, Niigata Pref., Katamachi; CD: male specimen, China, imprecise locality; EF: topotype female specimen, same locality as in the male topotype).A, C: median lobe of aedeagus, left lateral view; B: same, dorsal view; D: same, ventral view; E: left gonocoxite, ventral view; F: spermathecal complex and gonocoxites, ventral view. Scale lines = 0.5 mm (Figs AD, F); = 0.2 mm (Fig. E)." figureDoi="http://doi.org/10.5281/zenodo.4407112" httpUri="https://zenodo.org/record/4407112/files/figure.png" pageId="46" pageNumber="47">Figs 17A, B, C, D</figureCitation>
) with basal bulb short and narrow; angle between basal bulb and shaft nearly straight; shaft long and swollen; apex short, tapered and curved ventrally; apical lamella widely rounded; ostium long, but shorter than in
<taxonomicName id="4C1594CE296EFF91FCA8FB8BE834FBA4" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[884,1050,1087,1113]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="helopioides">
<emphasis id="B961335F296EFF91FCA8FB8BEFA6FBA4" box="[884,904,1087,1113]" italics="true" pageId="46" pageNumber="47">O</emphasis>
.
<emphasis id="B961335F296EFF91FC45FB8BE834FBA4" box="[921,1050,1087,1113]" italics="true" pageId="46" pageNumber="47">helopioides</emphasis>
</taxonomicName>
, not reaching basal bulb.
<emphasis id="B961335F296EFF91FA99FB8AECD2FB80" bold="true" italics="true" pageId="46" pageNumber="47">Female genitalia</emphasis>
. Basal gonocoxite with ten or so lateroapical setae. Apical gonocoxite without dorsolateral ensiform setae, with nematiform setae (
<figureCitation id="132EF3C8296EFF91FE7DFB33EE28FB5C" box="[417,518,1159,1185]" captionStart="FIGURE 17" captionStartId="45.[151,250,1910,1934]" captionTargetBox="[223,1364,191,1881]" captionTargetId="figure-17@45.[214,1374,181,1886]" captionTargetPageId="45" captionText="FIGURE 17. Oodes echigonus Habu &amp; Baba, 1960 (AB: topotype male specimen, Japan, Niigata Pref., Katamachi; CD: male specimen, China, imprecise locality; EF: topotype female specimen, same locality as in the male topotype).A, C: median lobe of aedeagus, left lateral view; B: same, dorsal view; D: same, ventral view; E: left gonocoxite, ventral view; F: spermathecal complex and gonocoxites, ventral view. Scale lines = 0.5 mm (Figs AD, F); = 0.2 mm (Fig. E)." figureDoi="http://doi.org/10.5281/zenodo.4407112" httpUri="https://zenodo.org/record/4407112/files/figure.png" pageId="46" pageNumber="47">Fig. 17E</figureCitation>
). Spermatheca coiled apically (
<figureCitation id="132EF3C8296EFF91FCB9FB33EFE6FB5C" box="[869,968,1159,1185]" captionStart="FIGURE 17" captionStartId="45.[151,250,1910,1934]" captionTargetBox="[223,1364,191,1881]" captionTargetId="figure-17@45.[214,1374,181,1886]" captionTargetPageId="45" captionText="FIGURE 17. Oodes echigonus Habu &amp; Baba, 1960 (AB: topotype male specimen, Japan, Niigata Pref., Katamachi; CD: male specimen, China, imprecise locality; EF: topotype female specimen, same locality as in the male topotype).A, C: median lobe of aedeagus, left lateral view; B: same, dorsal view; D: same, ventral view; E: left gonocoxite, ventral view; F: spermathecal complex and gonocoxites, ventral view. Scale lines = 0.5 mm (Figs AD, F); = 0.2 mm (Fig. E)." figureDoi="http://doi.org/10.5281/zenodo.4407112" httpUri="https://zenodo.org/record/4407112/files/figure.png" pageId="46" pageNumber="47">Fig. 17F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC6296EFF91FF1BFB1EEF29FB38" box="[199,775,1194,1221]" pageId="46" pageNumber="47" type="biology_ecology">
<paragraph id="8BAAEF4D296EFF91FF1BFB1EEF29FB38" blockId="46.[151,1437,150,1510]" box="[199,775,1194,1221]" pageId="46" pageNumber="47">
<emphasis id="B961335F296EFF91FF1BFB1EED0DFB39" bold="true" box="[199,291,1194,1220]" pageId="46" pageNumber="47">Habitat</emphasis>
. See “Habitat” under
<taxonomicName id="4C1594CE296EFF91FDC5FB1FEF2EFB39" authorityName="Bates" authorityYear="1873" box="[537,768,1195,1221]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="vicarius">
<emphasis id="B961335F296EFF91FDC5FB1FEF2EFB39" box="[537,768,1195,1221]" italics="true" pageId="46" pageNumber="47">Pseudoodes vicarius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC6296EFF91FF1BFB7AE830FB17" box="[199,1054,1230,1258]" pageId="46" pageNumber="47" type="distribution">
<paragraph id="8BAAEF4D296EFF91FF1BFB7AE830FB17" blockId="46.[151,1437,150,1510]" box="[199,1054,1230,1258]" pageId="46" pageNumber="47">
<emphasis id="B961335F296EFF91FF1BFB7AED77FB15" bold="true" box="[199,345,1230,1256]" pageId="46" pageNumber="47">Distribution</emphasis>
. Palaearctic Region:
<collectingCountry id="F302AFDD296EFF91FD95FB7BEEA7FB14" box="[585,649,1231,1257]" name="Japan" pageId="46" pageNumber="47">Japan</collectingCountry>
(Honshu),
<collectingCountry id="F302AFDD296EFF91FCDBFB7BEF64FB14" box="[775,842,1231,1257]" name="China" pageId="46" pageNumber="47">China</collectingCountry>
(
<collectingRegion id="49D121AF296EFF91FC85FB7BEF9FFB14" box="[857,945,1231,1257]" country="China" name="Jiangsu" pageId="46" pageNumber="47">Jiangsu</collectingRegion>
,
<collectingRegion id="49D121AF296EFF91FC61FB7BE83DFB17" box="[957,1043,1231,1258]" country="China" name="Jiangxi" pageId="46" pageNumber="47">Jiangxi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC6296EFF91FF1BFB47ED8DFA1B" pageId="46" pageNumber="47" type="discussion">
<paragraph id="8BAAEF4D296EFF91FF1BFB47ED8DFA1B" blockId="46.[151,1437,150,1510]" pageId="46" pageNumber="47">
<emphasis id="B961335F296EFF91FF1BFB47ED24FAF0" bold="true" box="[199,266,1267,1293]" pageId="46" pageNumber="47">Notes</emphasis>
. Describing the male protarsomeres,
<bibRefCitation id="EF8492BC296EFF91FD6BFB47EF87FAF3" author="Habu, A. &amp; Baba, K." box="[695,937,1267,1294]" pageId="46" pageNumber="47" pagination="31 - 33" refId="ref61814" refString="Habu, A. &amp; Baba, K. (1960) One new species of Oodes from Niigata Prefecture, Japan (Coleoptera, Carabidae). Akitu, 9, 31 - 33." type="journal article" year="1960">Habu &amp; Baba (1960)</bibRefCitation>
stated that “
<emphasis id="B961335F296EFF91FBE0FB47EF37FACC" italics="true" pageId="46" pageNumber="47">segment 3 is almost as long as segment 2, slightly narrower than this, with apex oblique</emphasis>
” (ibid.: 32) and “
<emphasis id="B961335F296EFF91FC34FAA3E924FACC" box="[1000,1290,1303,1329]" italics="true" pageId="46" pageNumber="47">with lateral sides parallel</emphasis>
” (ibid.: 33). We measured the lengths of male protarsomeres 2 and
<quantity id="4CED42A8296EFF91FD25FA8FEF0AFAA8" box="[761,804,1339,1365]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="46" pageNumber="47" unit="in" value="3.0">3 in</quantity>
four Chinese specimens and two topotypical specimens. It was established that protarsomere 3 is about 1.151.25 times longer than protarsomere 2, and thus, the above indication by Habu &amp; Baba is not correct. It should be noted also that specimens from
<collectingCountry id="F302AFDD296EFF91FB89FA37E8B6FA60" box="[1109,1176,1411,1437]" name="China" pageId="46" pageNumber="47">China</collectingCountry>
have smaller body size (
<quantity id="4CED42A8296EFF91FF43FA13ED69FA3C" box="[159,327,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.385" metricValueMax="1.49" metricValueMin="1.28" pageId="46" pageNumber="47" unit="mm" value="13.85" valueMax="14.9" valueMin="12.8">12.814.9 mm</quantity>
, original data), while those from
<collectingCountry id="F302AFDD296EFF91FD1EFA13EF2CFA3C" box="[706,770,1447,1473]" name="Japan" pageId="46" pageNumber="47">Japan</collectingCountry>
have body length 14.0
<quantity id="4CED42A8296EFF91FBC9FA12E857FA3C" box="[1045,1145,1446,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="46" pageNumber="47" unit="mm" value="15.2">15.2 mm</quantity>
(data from
<bibRefCitation id="EF8492BC296EFF91FADCFA13ECE1FA18" author="Habu, A. &amp; Baba, K." pageId="46" pageNumber="47" pagination="31 - 33" refId="ref61814" refString="Habu, A. &amp; Baba, K. (1960) One new species of Oodes from Niigata Prefecture, Japan (Coleoptera, Carabidae). Akitu, 9, 31 - 33." type="journal article" year="1960">Habu &amp; Baba 1960</bibRefCitation>
and original data).
</paragraph>
</subSubSection>
</treatment>
</document>