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<document id="261CBC23DBB5466762C42FE50A61B4F5" ID-CLB-Dataset="54828" ID-DOI="10.5733/afin.054.0101" ID-GBIF-Dataset="753510a6-cffa-4637-a1bd-e9ae7d5d1660" ID-ISSN="2305-2562" ID-Zenodo-Dep="7670100" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676639734503" checkinUser="felipe" docAuthor="Griffiths, O. L. &amp; Herbert, D. G." docDate="2013" docId="03BC1F3CE52ED91D0788FC7E4234B40E" docLanguage="en" docName="AfrInvertebr.54.1.1-48.pdf" docOrigin="African Invertebrates 54 (1)" docStyle="DocumentStyle:0201127D758FBEB889983D376DC13FCB.4:AfrInvertebr.2008-2015.journal_article.1cover" docStyleId="0201127D758FBEB889983D376DC13FCB" docStyleName="AfrInvertebr.2008-2015.journal_article.1cover" docStyleVersion="4" docTitle="Tropidophora humbug Griffiths &amp; Herbert 2013, sp. n." docType="treatment" docUuid="5A3982CB-F82D-47D0-A1E7-416E7A0659D5" docUuidSource="ZooBank" docVersion="8" lastPageNumber="10" masterDocId="FF856744E529D917064DFF964621B743" masterDocTitle="New species of land snails (Mollusca: Gastropoda) from two isolated karst formations in central western Madagascar: Tsingy Beanka and Antsingimavo, with additional notes on other regional endemics" masterLastPageNumber="48" masterPageNumber="1" pageNumber="7" updateTime="1699475339093" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FEDEBC211A71757B44DE080430915D46">New species of land snails (Mollusca: Gastropoda) from two isolated karst formations in central western Madagascar: Tsingy Beanka and Antsingimavo, with additional notes on other regional endemics</mods:title>
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<mods:namePart id="328D1389E74743C6E6E7885F7DB94F0A">Griffiths, O. L.</mods:namePart>
<mods:affiliation id="653364148F8D9A790C65C2D956FBAD87">Research associate of the Australian Museum, Sydney, Australia and Biodiversity Conservation Madagascar (BCM) Ltd, Rivière des Anguilles, Mauritius</mods:affiliation>
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<mods:namePart id="6C6102E1714D1875E6471B7AD6C3CBE4">Herbert, D. G.</mods:namePart>
<mods:affiliation id="A7BBA2AA7BA76895F244E70A28EC3143">KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg, 3200 South Africa and School of Life Sciences, University of KwaZulu-Natal, Pietermaritzburg, 3206 South Africa</mods:affiliation>
<mods:nameIdentifier id="9201035360FFBB597794AD4E9D1929EA" type="email">dherbert@nmsa.org.za</mods:nameIdentifier>
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<mods:title id="AA1D79E7DAF0E0EBD65745E060B6534D">African Invertebrates</mods:title>
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<treatment id="03BC1F3CE52ED91D0788FC7E4234B40E" ID-DOI="http://doi.org/10.5281/zenodo.7670246" ID-GBIF-Taxon="209341513" ID-Zenodo-Dep="7670246" LSID="urn:lsid:zoobank.org:act:5A3982CB-F82D-47D0-A1E7-416E7A0659D5" httpUri="http://treatment.plazi.org/id/03BC1F3CE52ED91D0788FC7E4234B40E" lastPageId="10" lastPageNumber="10" pageId="7" pageNumber="7">
<subSubSection id="C30FFDA1E52ED9100788FC7E453BB340" box="[453,794,1000,1027]" pageId="7" pageNumber="7" type="nomenclature">
<paragraph id="8BAAAE2AE52ED9100788FC7E453BB340" blockId="7.[144,1104,1000,1676]" box="[453,794,1000,1027]" pageId="7" pageNumber="7">
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<taxonomicName id="4C15D5A9E52ED9100788FC7E44F6B341" ID-CoL="59C7Q" authority="Griffiths &amp; Herbert, 2013" authorityName="Griffiths &amp; Herbert" authorityYear="2013" box="[453,727,1000,1026]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="7" phylum="Mollusca" rank="species" species="humbug" status="sp. nov.">
<emphasis id="B9617238E52ED9100788FC7E44F6B341" bold="true" box="[453,727,1000,1026]" pageId="7" pageNumber="7">Tropidophora humbug</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A252CF43E52ED9100493FC7F453BB340" box="[734,794,1001,1027]" pageId="7" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30FFDA1E52ED91D046CFB844234B40E" lastPageId="10" lastPageNumber="10" pageId="7" pageNumber="7" type="description">
<paragraph id="8BAAAE2AE52ED910046CFB84449FB36F" blockId="7.[144,1104,1000,1676]" box="[545,702,1042,1068]" pageId="7" pageNumber="7">
<figureCitation id="132EB2AFE52ED910046CFB844449B36F" box="[545,616,1042,1068]" captionStart="Fig" captionStartId="8.[180,212,760,781]" captionTargetBox="[187,1137,216,734]" captionTargetId="figure-294@8.[181,1141,204,742]" captionTargetPageId="8" captionText="Fig.5. Tropidophora humbug sp.n.: (AC) holotype,Tsingy Beanka,max.diameter 31.5mm,height 16.3 mm, AMS C.474165; (DF) paratype, heavily banded specimen, Tsingy de Bemaraha, max. diameter 30.4 mm, height 14.1 mm, NMSA L8444/T2658; (G) operculum of holotype, external surface, max. diameter 11.0 mm, AMS C.474165." figureDoi="http://doi.org/10.5281/zenodo.7670110" httpUri="https://zenodo.org/record/7670110/files/figure.png" pageId="7" pageNumber="7">Figs 5</figureCitation>
,
<figureCitation id="132EB2AFE52ED9100439FB8444A5B36F" box="[628,644,1042,1068]" captionStart="Fig" captionStartId="9.[144,177,1052,1073]" captionTargetBox="[151,1096,205,1034]" captionTargetId="figure-239@9.[151,1097,204,1034]" captionTargetPageId="9" captionText="Fig.6. Tropidophora humbug sp.n., radula, paratype NMSA L7042/T2961:(A) entire width of radula,scale bar = 250 μm; (B) rachidian and left inner lateral teeth, scale bar = 100 μm; (C) right outer lateral and marginal teeth, scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7670112" httpUri="https://zenodo.org/record/7670112/files/figure.png" pageId="7" pageNumber="7">6</figureCitation>
,
<figureCitation id="132EB2AFE52ED91004DDFB84449FB36F" box="[656,702,1042,1068]" captionStart="Fig" captionStartId="36.[181,214,1462,1483]" captionTargetBox="[190,1134,203,1445]" captionTargetId="figure-127@36.[189,1134,203,1445]" captionTargetPageId="36" captionText="Fig. 25. Living animals from Tsingy Beanka: (A) Acroptychia bathiei Fischer-Piette &amp; Bedoucha, 1965, stn 13/06, shell diameter 23.2 mm, NMSA L7315; (B) Tropidophora humbug sp. n., stn 12/06, shell diameter 28.3 mm, paratype,NMSA L7204/T2984; (C) Tropidophora secunda Fischer-Piette &amp; Bedoucha, 1965, stn 03/06, shell diameter approx.19.0 mm; (D) Clavator griffithsjonesi Emberton, 1999, stn 12/06, shell height 94.8 mm, NMSA L7192; (E) Ampelita milloti Fischer-Piette,1952,stn 07/09, shell diameter 19.2 mm, NMSA L8502; (F) Ampelita namerokoensis Fischer-Piette, 1952, stn 12/06, shell diameter 20.3 mm, NMSA L8442; (G) Kalidos griffithshauchleri Emberton, 2002, stn 12/06, shell diameter approx. 60 mm; (H) Kalidos maryannae sp. n., stn 12/06, shell diameter 22.5 mm, paratype, NMSA L7193/T2944." figureDoi="http://doi.org/10.5281/zenodo.7670161" httpUri="https://zenodo.org/record/7670161/files/figure.png" pageId="7" pageNumber="7">25B</figureCitation>
</paragraph>
<paragraph id="8BAAAE2AE52ED91006DDFBAC44B1B337" blockId="7.[144,1104,1000,1676]" pageId="7" pageNumber="7">Etymology: The colour pattern of bold stripes is reminiscent of that of old-fashioned humbug sweets; used as a noun in apposition.</paragraph>
<paragraph id="8BAAAE2AE52ED91006DDFB1447C3B3BF" blockId="7.[144,1104,1000,1676]" pageId="7" pageNumber="7">Diagnosis: Spire very low, shell almost planorboid, body whorl not conspicuously tumescent, umbilicus very wide; columella lip not expanded and not reflected over umbilicus; surface virtually smooth to the naked eye; boldly marked with brown spiral bands on a near white ground.</paragraph>
<paragraph id="8BAAAE2AE52ED91006DDFA9C473AB267" blockId="7.[144,1104,1000,1676]" box="[144,283,1290,1316]" pageId="7" pageNumber="7">Description:</paragraph>
<paragraph id="8BAAAE2AE52ED91F06DDFAA24458B4F7" blockId="7.[144,1104,1000,1676]" lastBlockId="8.[181,1142,890,1677]" lastPageId="8" lastPageNumber="8" pageId="7" pageNumber="7">
<emphasis id="B9617238E52ED91006DDFAA246E9B20F" box="[144,200,1332,1356]" italics="true" pageId="7" pageNumber="7">Shell</emphasis>
: Medium sized, depressed-discoidal to planorboid; spire very low (H:D=0.46 0.62), often with only embryonic whorls projecting in apertural view; final part of last adult whorl descending prior to aperture, but not steeply so; whorls more or less evenly rounded, suture indented; umbilicus very wide, its margin evenly rounded, underside of embryonic whorls clearly visible. Protoconch of 1¼1½ whorls, essentially smooth, but microscopically shagreened. Teleoconch of a further 2½2¾ whorls; the first with approx. 6 weak spiral ridges crossed by numerous fine, close-set, axial threads; spirals becoming more numerous but less distinct on subsequent whorls and axials somewhat coarser; axials resembling fine, uneven, close-set growth-lines on last half-whorl of adult, these extending onto base and into umbilicus; base lacking spiral sculpture. Aperture subcircular, strongly oblique to vertical axis of shell; peristome incomplete, interrupted briefly in parietal region; rim of peristome reflected forming a flaring lip, but this not noticeably enlarged in columella region.
</paragraph>
<caption id="DF6AFEA2E521D91F06F9FD6E441AB416" ID-DOI="http://doi.org/10.5281/zenodo.7670110" ID-Zenodo-Dep="7670110" httpUri="https://zenodo.org/record/7670110/files/figure.png" pageId="8" pageNumber="8" startId="8.[180,212,760,781]" targetBox="[187,1137,216,734]" targetPageId="8">
<paragraph id="8BAAAE2AE521D91F06F9FD6E441AB416" blockId="8.[180,1143,760,853]" pageId="8" pageNumber="8">
Fig. 5.
<taxonomicName id="4C15D5A9E521D91F06BCFD6C4796B44E" authority="Griffiths &amp; Herbert, 2013" authorityName="Griffiths &amp; Herbert" authorityYear="2013" box="[241,439,762,781]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="8" phylum="Mollusca" rank="species" species="humbug" status="sp. nov.">
<emphasis id="B9617238E521D91F06BCFD6C4796B44E" box="[241,439,762,781]" italics="true" pageId="8" pageNumber="8">Tropidophora humbug</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A252CF43E521D91F07F7FD6E47CAB44E" box="[442,491,760,781]" pageId="8" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
: (AC) holotype,
<taxonomicName id="4C15D5A9E521D91F04C8FD6E452BB44E" authorityName=", Antsingimavo" authorityYear="2013" box="[645,778,760,781]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="8" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
,max.diameter 31.5 mm, height 16.3 mm, AMS C.474165; (DF) paratype, heavily banded specimen, Tsingy de Bemaraha, max. diameter 30.4 mm, height 14.1 mm, NMSA L8444/T2658; (G) operculum of holotype, external surface, max. diameter 11.0 mm, AMS C.474165.
</paragraph>
</caption>
<paragraph id="8BAAAE2AE521D91F069DFC2C422EB3D6" blockId="8.[181,1142,890,1677]" pageId="8" pageNumber="8">Ground colour bone-white, boldly marked with dark brown spiral bands, the one at and extending just below periphery usually broadest, commonly with 2 or 3 additional bands above and a further 2 or 3 below this; an additional fine orange-brown intermediary spiral line sometimes present in intervals between bands; umbilical region with additional colour bands, often fine, such that base may have up to 7 bands in total; exceptionally, base may have only 1 or 2 colour bands; precise position of colour bands somewhat variable, but uppermost band generally not in contact with suture.</paragraph>
<paragraph id="8BAAAE2AE521D91F069DFB0C47E9B396" blockId="8.[181,1142,890,1677]" pageId="8" pageNumber="8">
Dimensions:
<typeStatus id="54AE1088E521D91F0728FB0C47F1B3F7" box="[357,464,1178,1204]" pageId="8" pageNumber="8" type="holotype">Holotype</typeStatus>
, max. diameter
<quantity id="4CED03CFE521D91F04CFFB0C44C9B3F6" box="[642,744,1178,1205]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" pageId="8" pageNumber="8" unit="mm" value="31.5">31.5 mm</quantity>
, height
<quantity id="4CED03CFE521D91F050CFB0C4589B3F6" box="[833,936,1178,1205]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="8" pageNumber="8" unit="mm" value="16.3">16.3 mm</quantity>
; largest specimen, max. diameter 32.0 mm.
</paragraph>
<paragraph id="8BAAAE2AE521D91F06F8FB724794B27F" blockId="8.[181,1142,890,1677]" pageId="8" pageNumber="8">
<emphasis id="B9617238E521D91F06F8FB724756B3BF" box="[181,375,1252,1276]" italics="true" pageId="8" pageNumber="8">External features</emphasis>
(
<figureCitation id="132EB2AFE521D91F07CAFB7447CEB3BF" box="[391,495,1250,1276]" captionStart="Fig" captionStartId="36.[181,214,1462,1483]" captionTargetBox="[190,1134,203,1445]" captionTargetId="figure-127@36.[189,1134,203,1445]" captionTargetPageId="36" captionText="Fig. 25. Living animals from Tsingy Beanka: (A) Acroptychia bathiei Fischer-Piette &amp; Bedoucha, 1965, stn 13/06, shell diameter 23.2 mm, NMSA L7315; (B) Tropidophora humbug sp. n., stn 12/06, shell diameter 28.3 mm, paratype,NMSA L7204/T2984; (C) Tropidophora secunda Fischer-Piette &amp; Bedoucha, 1965, stn 03/06, shell diameter approx.19.0 mm; (D) Clavator griffithsjonesi Emberton, 1999, stn 12/06, shell height 94.8 mm, NMSA L7192; (E) Ampelita milloti Fischer-Piette,1952,stn 07/09, shell diameter 19.2 mm, NMSA L8502; (F) Ampelita namerokoensis Fischer-Piette, 1952, stn 12/06, shell diameter 20.3 mm, NMSA L8442; (G) Kalidos griffithshauchleri Emberton, 2002, stn 12/06, shell diameter approx. 60 mm; (H) Kalidos maryannae sp. n., stn 12/06, shell diameter 22.5 mm, paratype, NMSA L7193/T2944." figureDoi="http://doi.org/10.5281/zenodo.7670161" httpUri="https://zenodo.org/record/7670161/files/figure.png" pageId="8" pageNumber="8">Fig. 25B</figureCitation>
): Head-foot more or less uniformly dark grey-brown, but eyestalks and tentacle bases paler; tip of snout conspicuously indented in mid-line; skin texture finely granular.
</paragraph>
<paragraph id="8BAAAE2AE521D91F06F8FADA47F3B147" blockId="8.[181,1142,890,1677]" pageId="8" pageNumber="8">
<emphasis id="B9617238E521D91F06F8FADA4711B227" box="[181,304,1356,1380]" italics="true" pageId="8" pageNumber="8">Operculum</emphasis>
(
<figureCitation id="132EB2AFE521D91F0773FADC47B7B227" box="[318,406,1354,1380]" captionStart="Fig" captionStartId="8.[180,212,760,781]" captionTargetBox="[187,1137,216,734]" captionTargetId="figure-294@8.[181,1141,204,742]" captionTargetPageId="8" captionText="Fig.5. Tropidophora humbug sp.n.: (AC) holotype,Tsingy Beanka,max.diameter 31.5mm,height 16.3 mm, AMS C.474165; (DF) paratype, heavily banded specimen, Tsingy de Bemaraha, max. diameter 30.4 mm, height 14.1 mm, NMSA L8444/T2658; (G) operculum of holotype, external surface, max. diameter 11.0 mm, AMS C.474165." figureDoi="http://doi.org/10.5281/zenodo.7670110" httpUri="https://zenodo.org/record/7670110/files/figure.png" pageId="8" pageNumber="8">Fig. 5G</figureCitation>
): Oligospiral; exterior portion a calcareous disc, attached to an inner and slightly larger corneous layer; external surface off-white (usually encrusted with detritus particles), lacking colour pattern, but with a broad convex spiral ridge more or less at mid-whorl; thinner toward periphery; edge of disc concave, with numerous, closeset, rather unevenly spaced transverse partitions, except along the growing (parietal) margin, which is smooth.
</paragraph>
<paragraph id="8BAAAE2AE521D91E06F8F983458BB262" blockId="8.[181,1142,890,1677]" lastBlockId="9.[144,1104,1159,1677]" lastPageId="9" lastPageNumber="9" pageId="8" pageNumber="8">
<emphasis id="B9617238E521D91F06F8F9834727B16E" box="[181,262,1557,1581]" italics="true" pageId="8" pageNumber="8">Radula</emphasis>
(
<figureCitation id="132EB2AFE521D91F075BF984477DB16F" box="[278,348,1554,1580]" captionStart="Fig" captionStartId="9.[144,177,1052,1073]" captionTargetBox="[151,1096,205,1034]" captionTargetId="figure-239@9.[151,1097,204,1034]" captionTargetPageId="9" captionText="Fig.6. Tropidophora humbug sp.n., radula, paratype NMSA L7042/T2961:(A) entire width of radula,scale bar = 250 μm; (B) rachidian and left inner lateral teeth, scale bar = 100 μm; (C) right outer lateral and marginal teeth, scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7670112" httpUri="https://zenodo.org/record/7670112/files/figure.png" pageId="8" pageNumber="8">Fig. 6</figureCitation>
): Formula 1+2+1+2+1; length
<quantity id="4CED03CFE521D91F04F6F984452DB16E" box="[699,780,1554,1581]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="8" unit="mm" value="11.0">11 mm</quantity>
,
<emphasis id="B9617238E521D91F0555F9834513B16E" box="[792,818,1557,1581]" italics="true" pageId="8" pageNumber="8">ca</emphasis>
35 rows/mm; dentition fine. Rachidian with seven rounded cusps, central one largest, the outermost pair small; occasionally with very small intermediary teeth between the larger ones. Inner laterals with four cusps, the second of which is bluntly rounded and consistently the largest, the others more pointed; outer lateral with five cusps, the inner one usually slightly larger, the others progressively smaller. Marginal teeth broad, the edge comprising three regions; an inner coarsely dentate element with
<emphasis id="B9617238E520D91E04DCFB5C448AB3A1" box="[657,683,1226,1250]" italics="true" pageId="9" pageNumber="9">ca</emphasis>
9 denticles; a more finely denticulate central region, and a smooth outer portion. Such a radula conforms to the pattern seen in
<taxonomicName id="4C15D5A9E520D91E06E1FA9C4451B262" authority="Martens, 1880" authorityName="Martens" authorityYear="1880" box="[172,624,1287,1314]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="9" phylum="Mollusca" rank="subGenus" subGenus="Ligatella">
<emphasis id="B9617238E520D91E06E1FA9C47E7B261" box="[172,454,1290,1314]" italics="true" pageId="9" pageNumber="9">Tropidophora (Ligatella)</emphasis>
Martens, 1880
</taxonomicName>
(Fischer-Piette
<emphasis id="B9617238E520D91E056FFA9C4572B261" box="[802,851,1290,1314]" italics="true" pageId="9" pageNumber="9">et al</emphasis>
. 1969).
</paragraph>
<caption id="DF6AFEA2E520D91E06DDFB8A4416B322" ID-DOI="http://doi.org/10.5281/zenodo.7670112" ID-Zenodo-Dep="7670112" httpUri="https://zenodo.org/record/7670112/files/figure.png" pageId="9" pageNumber="9" startId="9.[144,177,1052,1073]" targetBox="[151,1096,205,1034]" targetPageId="9">
<paragraph id="8BAAAE2AE520D91E06DDFB8A4416B322" blockId="9.[144,1104,1052,1121]" pageId="9" pageNumber="9">
Fig. 6.
<taxonomicName id="4C15D5A9E520D91E0680FB8847B9B372" authority="Griffiths &amp; Herbert, 2013" authorityName="Griffiths &amp; Herbert" authorityYear="2013" box="[205,408,1054,1073]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="9" phylum="Mollusca" rank="species" species="humbug" status="sp. nov.">
<emphasis id="B9617238E520D91E0680FB8847B9B372" box="[205,408,1054,1073]" italics="true" pageId="9" pageNumber="9">Tropidophora humbug</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A252CF43E520D91E07D1FB8A47ECB372" box="[412,461,1052,1073]" pageId="9" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
, radula, paratype NMSA L7042/T2961:(A) entire width of radula, scale bar = 250 μm; (B) rachidian and left inner lateral teeth, scale bar = 100 μm; (C) right outer lateral and marginal teeth, scale bar = 100 μm.
</paragraph>
</caption>
<paragraph id="8BAAAE2AE520D91E06DDFAB84458B230" blockId="9.[144,1104,1159,1677]" pageId="9" pageNumber="9">
<materialsCitation id="3B7DA477E520D91E06DDFAB84458B230" collectingDate="2009-04" collectionCode="AMS" collectorName="Central W Madagascar &amp; R. Randalana" country="Madagascar" elevation="380" latitude="-18.05028" location="Andohanandranogedro" longLatPrecision="1" longitude="44.53786" pageId="9" pageNumber="9" specimenCount="1" typeStatus="holotype">
<typeStatus id="54AE1088E520D91E06DDFAB846C6B200" box="[144,231,1326,1347]" pageId="9" pageNumber="9" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F302EEBAE520D91E06A0FAB847B0B200" box="[237,401,1326,1347]" name="Madagascar" pageId="9" pageNumber="9">MADAGASCAR</collectingCountry>
:
<collectorName id="26E0CBFCE520D91E07DAFAB84445B200" box="[407,612,1326,1347]" pageId="9" pageNumber="9">Central W Madagascar</collectorName>
,
<emphasis id="B9617238E520D91E0427FAA6445EB200" box="[618,639,1328,1347]" italics="true" pageId="9" pageNumber="9">ca</emphasis>
<locationDeviation id="9A18B708E520D91E04C9FAB8456FB200" box="[644,846,1326,1347]" location="Maintirano" pageId="9" pageNumber="9">
<quantity id="4CED03CFE520D91E04C9FAB8449AB200" box="[644,699,1326,1347]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="9" unit="km" value="60.0">60 km</quantity>
E of Maintirano
</locationDeviation>
, tall dense dry forest growing above cliffs on E side of southern
<taxonomicName id="4C15D5A9E520D91E07F7FAD0441EB218" authorityName=", Antsingimavo" authorityYear="2013" box="[442,575,1350,1371]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
at
<location id="8ECAF8F1E520D91E041AFAD04508B218" LSID="urn:lsid:plazi:treatment:03BC1F3CE52ED91D0788FC7E4234B40E:8ECAF8F1E520D91E041AFAD04508B218" box="[599,809,1350,1371]" country="Madagascar" latitude="-18.05028" longLatPrecision="1" longitude="44.53786" name="Andohanandranogedro" pageId="9" pageNumber="9">Andohanandranogedro</location>
,
<geoCoordinate id="EE21C8EDE520D91E057DFAD045BBB218" box="[816,922,1350,1371]" degrees="18.05028" direction="south" orientation="latitude" pageId="9" pageNumber="9" precision="1" value="-18.05028">18.05028°S</geoCoordinate>
<geoCoordinate id="EE21C8EDE520D91E05D3FAD0422DB218" box="[926,1036,1350,1371]" degrees="44.53786" direction="east" orientation="longitude" pageId="9" pageNumber="9" precision="1" value="44.53786">44.53786°E</geoCoordinate>
,
<quantity id="4CED03CFE520D91E0259FAD0426DB218" box="[1044,1100,1350,1371]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="9" pageNumber="9" unit="m" value="380.0">
<elevation id="00384919E520D91E0259FAD0426DB218" box="[1044,1100,1350,1371]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="9" pageNumber="9" unit="m" value="380.0">380m</elevation>
</quantity>
,
<date id="FFAB88EAE520D91E06DDFAC846F4B230" box="[144,213,1374,1395]" pageId="9" pageNumber="9" value="2009-04">
<collectingDate id="EFEF7102E520D91E06DDFAC846F4B230" box="[144,213,1374,1395]" pageId="9" pageNumber="9" value="2009-04">iv.2009</collectingDate>
</date>
,
<collectorName id="26E0CBFCE520D91E0693FAC8477AB230" box="[222,347,1374,1395]" pageId="9" pageNumber="9">R. Randalana</collectorName>
, stn R04/09 (
<collectionCode id="ED0436EFE520D91E0797FAC8442DB230" box="[474,524,1374,1395]" pageId="9" pageNumber="9">AMS</collectionCode>
C.474165).
</materialsCitation>
</paragraph>
<paragraph id="8BAAAE2AE520D91E06DDFAE8450BB148" blockId="9.[144,1104,1159,1677]" pageId="9" pageNumber="9">
<materialsCitation id="3B7DA477E520D91E06DDFAE8452EB2D0" box="[144,783,1406,1427]" collectingDate="2009-04" collectionCode="NMSA" collectorName="Central W Madagascar &amp; R. Randalana" country="Madagascar" elevation="380" latitude="-18.05028" location="Andohanandranogedro" longLatPrecision="1" longitude="44.53786" pageId="9" pageNumber="9" specimenCode="L8547" specimenCount="2" typeStatus="paratype">
<typeStatus id="54AE1088E520D91E06DDFAE846CDB2D0" box="[144,236,1406,1427]" pageId="9" pageNumber="9" type="paratype">Paratypes</typeStatus>
: Same data as holotype (
<collectionCode id="ED0436EFE520D91E0785FAE8442BB2D0" box="[456,522,1406,1427]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="9" pageNumber="9" type="Museum">NMSA</collectionCode>
<specimenCode id="DBB30651E520D91E0442FAE8446BB2D0" box="[527,586,1406,1427]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="9" pageNumber="9" type="Museum">L8547</specimenCode>
/T2962,
<specimenCount id="9D1365A3E520D91E04DBFAE8452BB2D0" box="[662,778,1406,1427]" pageId="9" pageNumber="9" type="generic">2 specimens</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B7DA477E520D91E0555FAE846FBB2E8" collectionCode="AMS" country="Madagascar" pageId="9" pageNumber="9" specimenCount="5" typeStatus="paratype">
stn 14/95 (
<collectionCode id="ED0436EFE520D91E0533FAE84591B2D0" box="[894,944,1406,1427]" pageId="9" pageNumber="9">AMS</collectionCode>
C.469588,
<specimenCount id="9D1365A3E520D91E0254FAE846F5B2E8" pageId="9" pageNumber="9" type="generic">5 specimens</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B7DA477E520D91E06A9FA004443B2E8" box="[228,610,1430,1451]" collectionCode="AMS" country="Madagascar" pageId="9" pageNumber="9" specimenCount="5" typeStatus="paratype">
stn 10/96 (
<collectionCode id="ED0436EFE520D91E0706FA00475CB2E8" box="[331,381,1430,1451]" pageId="9" pageNumber="9">AMS</collectionCode>
C.469587,
<specimenCount id="9D1365A3E520D91E07A5FA00447CB2E8" box="[488,605,1430,1451]" pageId="9" pageNumber="9" type="generic">5 specimens</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B7DA477E520D91E0421FA0045CBB2E8" box="[620,1002,1430,1451]" collectionCode="AMS" country="Madagascar" pageId="9" pageNumber="9" specimenCount="2" typeStatus="paratype">
stn 14/96 (
<collectionCode id="ED0436EFE520D91E0499FA004527B2E8" box="[724,774,1430,1451]" pageId="9" pageNumber="9">AMS</collectionCode>
C.469589,
<specimenCount id="9D1365A3E520D91E053DFA0045C4B2E8" box="[880,997,1430,1451]" pageId="9" pageNumber="9" type="generic">2 specimens</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B7DA477E520D91E05B9FA0047C5B280" collectionCode="NMSA" country="Madagascar" pageId="9" pageNumber="9" specimenCode="L8444" specimenCount="6" typeStatus="paratype">
stn 15/96 (
<collectionCode id="ED0436EFE520D91E06DBFA3846F9B280" box="[150,216,1454,1475]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="9" pageNumber="9" type="Museum">NMSA</collectionCode>
<specimenCode id="DBB30651E520D91E0692FA38473AB280" box="[223,283,1454,1475]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="9" pageNumber="9" type="Museum">L8444</specimenCode>
/T2658,
<specimenCount id="9D1365A3E520D91E0724FA3847FFB280" box="[361,478,1454,1475]" pageId="9" pageNumber="9" type="generic">6 specimens</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B7DA477E520D91E07A2FA38453FB298" collectionCode="NMSA, MNHN" country="Madagascar" pageId="9" pageNumber="9" specimenCode="L7042, NHMUK 20120013" specimenCount="49" typeStatus="paratype">
stn 03/06 (
<collectionCode id="ED0436EFE520D91E0417FA3844BDB280" box="[602,668,1454,1475]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="9" pageNumber="9" type="Museum">NMSA</collectionCode>
<specimenCode id="DBB30651E520D91E04EEFA3844C1B280" box="[675,736,1454,1475]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="9" pageNumber="9" type="Museum">L7042</specimenCode>
/T2961,
<specimenCount id="9D1365A3E520D91E0560FA38458DB280" box="[813,940,1454,1475]" pageId="9" pageNumber="9" type="generic">45 specimens</specimenCount>
, some in alcohol;
<collectionCode id="ED0436EFE520D91E06DDFA5046F4B298" box="[144,213,1478,1499]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="9">MNHN</collectionCode>
IM-2010-20067,
<specimenCount id="9D1365A3E520D91E073FFA5047C3B298" box="[370,482,1478,1499]" pageId="9" pageNumber="9" type="generic">2 specimens</specimenCount>
;
<specimenCode id="DBB30651E520D91E07A4FA5044BEB298" box="[489,671,1478,1499]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="9" type="Museum">NHMUK 20120013</specimenCode>
,
<specimenCount id="9D1365A3E520D91E04EBFA504539B298" box="[678,792,1478,1499]" pageId="9" pageNumber="9" type="generic">2 specimens</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B7DA477E520D91E0568FA504726B2B0" collectionCode="NMSA" country="Madagascar" pageId="9" pageNumber="9" specimenCode="L7204" specimenCount="2" typeStatus="paratype">
stn 12/06 (
<collectionCode id="ED0436EFE520D91E05C5FA5045E8B298" box="[904,969,1478,1499]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="9" pageNumber="9" type="Museum">NMSA</collectionCode>
<specimenCode id="DBB30651E520D91E0583FA50422BB298" box="[974,1034,1478,1499]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="9" pageNumber="9" type="Museum">L7204</specimenCode>
/T2984,
<specimenCount id="9D1365A3E520D91E06DDFA484720B2B0" box="[144,257,1502,1523]" pageId="9" pageNumber="9" type="generic">2 specimens</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B7DA477E520D91E0742FA4845AFB2B0" box="[271,910,1502,1523]" collectionCode="AMS, TMAM" country="Madagascar" pageId="9" pageNumber="9" specimenCode="T166" specimenCount="13" typeStatus="paratype">
stn R04/09 (
<collectionCode id="ED0436EFE520D91E07CFFA484795B2B0" box="[386,436,1502,1523]" pageId="9" pageNumber="9">AMS</collectionCode>
C.469590,
<specimenCount id="9D1365A3E520D91E0456FA4844AAB2B0" box="[539,651,1502,1523]" pageId="9" pageNumber="9" type="generic">6 specimens</specimenCount>
;
<collectionCode id="ED0436EFE520D91E04DFFA4844F9B2B0" box="[658,728,1502,1523]" pageId="9" pageNumber="9">TMAM</collectionCode>
<specimenCode id="DBB30651E520D91E0491FA48452FB2B0" box="[732,782,1502,1523]" collectionCode="TMAM" pageId="9" pageNumber="9">T166</specimenCode>
,
<specimenCount id="9D1365A3E520D91E055BFA4845A6B2B0" box="[790,903,1502,1523]" pageId="9" pageNumber="9" type="generic">7 specimens</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B7DA477E520D91E05D8FA484765B148" collectionCode="TMAM" country="Madagascar" pageId="9" pageNumber="9" specimenCode="T160" specimenCount="5" typeStatus="paratype">
stn R01/10 (
<collectionCode id="ED0436EFE520D91E0245FA484271B2B0" box="[1032,1104,1502,1523]" pageId="9" pageNumber="9">TMAM</collectionCode>
<specimenCode id="DBB30651E520D91E06DDFA6046E0B148" box="[144,193,1526,1547]" collectionCode="TMAM" pageId="9" pageNumber="9">T160</specimenCode>
,
<specimenCount id="9D1365A3E520D91E0686FA60471EB148" box="[203,319,1526,1547]" pageId="9" pageNumber="9" type="generic">5 specimens</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B7DA477E520D91E0703FA604507B148" box="[334,806,1526,1547]" collectionCode="MNHN" country="Madagascar" pageId="9" pageNumber="9" specimenCount="7" typeStatus="paratype">
stn R02/10 (
<collectionCode id="ED0436EFE520D91E0789FA60442AB148" box="[452,523,1526,1547]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="9">MNHN</collectionCode>
IM-2010-20068,
<specimenCount id="9D1365A3E520D91E04E1FA604501B148" box="[684,800,1526,1547]" pageId="9" pageNumber="9" type="generic">7 specimens</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAAAE2AE520D91E06DDF980445CB100" blockId="9.[144,1104,1159,1677]" pageId="9" pageNumber="9">
Additional locality data:
<taxonomicName id="4C15D5A9E520D91E0723F98E47D0B168" authorityName=", Antsingimavo" authorityYear="2013" box="[366,497,1560,1579]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="9" phylum="Mollusca" rank="species" species="beanka">
<emphasis id="B9617238E520D91E0723F98E47D0B168" box="[366,497,1560,1579]" italics="true" pageId="9" pageNumber="9">Tsingy Beanka</emphasis>
</taxonomicName>
: stns 05/06, 11/06, 12/06, 13/06, 16/06, 18/06, 01/09, 03/09, 11/09, 03/10, 10/10.
<emphasis id="B9617238E520D91E0743F9A647E6B100" box="[270,455,1584,1603]" italics="true" pageId="9" pageNumber="9">Tsingy de Bemaraha</emphasis>
: stns 08/96, 09/96.
</paragraph>
<paragraph id="8BAAAE2AE520D91E06DDF9C444E7B1CF" blockId="9.[144,1104,1159,1677]" pageId="9" pageNumber="9">
Distribution:A narrow-range endemic; known only from the
<taxonomicName id="4C15D5A9E520D91E0561F9C445F0B12F" authorityName=", Antsingimavo" authorityYear="2013" box="[812,977,1618,1644]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
and Tsingy de Bemaraha, but common at both these localities.
</paragraph>
<paragraph id="8BAAAE2AE523D91D06F8FF5D445DB646" blockId="10.[181,1143,203,846]" pageId="10" pageNumber="10">Habitat: Tall semi-deciduous and deciduous western dry forest. Often found on tree trunks after rain. Aestivates in leaf-litter.</paragraph>
<paragraph id="8BAAAE2AE523D91D06F8FE85443BB5AE" blockId="10.[181,1143,203,846]" pageId="10" pageNumber="10">
Remarks: Fischer-Piette
<emphasis id="B9617238E523D91D0787FE8047DAB66D" box="[458,507,278,302]" italics="true" pageId="10" pageNumber="10">et al</emphasis>
. (1993) grouped medium-sized, low-spired Madagascan
<taxonomicName id="4C15D5A9E523D91D06F8FEA04788B60E" box="[181,425,307,334]" pageId="10" pageNumber="10">
<emphasis id="B9617238E523D91D06F8FEA04770B60D" box="[181,337,310,334]" italics="true" pageId="10" pageNumber="10">Tropidophora</emphasis>
species
</taxonomicName>
together in their groupe despèces du
<taxonomicName id="4C15D5A9E523D91D0513FEA047AFB62E" authority="(Petit de la Saussaye, 1844)" baseAuthorityName="Petit de la Saussaye" baseAuthorityYear="1844" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="10" phylum="Mollusca" rank="species" species="deshayesiana">
<emphasis id="B9617238E523D91D0513FEA04230B60D" box="[862,1041,310,334]" italics="true" pageId="10" pageNumber="10">T. deshayesiana</emphasis>
(Petit de la Saussaye, 1844)
</taxonomicName>
, but with the exception of
<taxonomicName id="4C15D5A9E523D91D04F2FEC045C5B62E" authority="(Sowerby, 1843)" baseAuthorityName="Sowerby" baseAuthorityYear="1843" box="[703,996,339,366]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="10" phylum="Mollusca" rank="species" species="vittata">
<emphasis id="B9617238E523D91D04F2FEC04502B62D" box="[703,803,342,366]" italics="true" pageId="10" pageNumber="10">T. vittata</emphasis>
(Sowerby, 1843)
</taxonomicName>
all members of this group have much stronger spiral sculpture than
<taxonomicName id="4C15D5A9E523D91D055CFEE045A4B6CD" authorityName="Griffiths &amp; Herbert" authorityYear="2013" box="[785,901,374,398]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="10" phylum="Mollusca" rank="species" species="humbug">
<emphasis id="B9617238E523D91D055CFEE045A4B6CD" box="[785,901,374,398]" italics="true" pageId="10" pageNumber="10">T. humbug</emphasis>
</taxonomicName>
, including
<taxonomicName id="4C15D5A9E523D91D024FFEE047B9B6EE" authority="Fischer-Piette, 1949" authorityName=": Fischer-Piette" authorityYear="1949" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="10" phylum="Mollusca" rank="species" species="chavani">
<emphasis id="B9617238E523D91D024FFEE04257B6CD" box="[1026,1142,374,398]" italics="true" pageId="10" pageNumber="10">T. chavani</emphasis>
Fischer-Piette, 1949
</taxonomicName>
(see above) with which it is sympatric. Besides being relatively smooth,
<taxonomicName id="4C15D5A9E523D91D075EFE204753B68D" baseAuthorityName="Sowerby" baseAuthorityYear="1843" box="[275,370,438,462]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="10" phylum="Mollusca" rank="species" species="vittata">
<emphasis id="B9617238E523D91D075EFE204753B68D" box="[275,370,438,462]" italics="true" pageId="10" pageNumber="10">T. vittata</emphasis>
</taxonomicName>
, which is recorded only from north-eastern
<collectingCountry id="F302EEBAE523D91D0507FE2545F1B68E" box="[842,976,435,461]" name="Madagascar" pageId="10" pageNumber="10">Madagascar</collectingCountry>
(Fischer-Piette
<emphasis id="B9617238E523D91D06F8FE4046C4B6AD" box="[181,229,470,494]" italics="true" pageId="10" pageNumber="10">et al</emphasis>
. 1993), shares with
<taxonomicName id="4C15D5A9E523D91D07F2FE404413B6AD" authorityName="Griffiths &amp; Herbert" authorityYear="2013" box="[447,562,470,494]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="10" phylum="Mollusca" rank="species" species="humbug">
<emphasis id="B9617238E523D91D07F2FE404413B6AD" box="[447,562,470,494]" italics="true" pageId="10" pageNumber="10">T. humbug</emphasis>
</taxonomicName>
a similarly banded colour pattern and is undoubtedly the most similar species. However,
<taxonomicName id="4C15D5A9E523D91D0478FE6044B4B54D" baseAuthorityName="Sowerby" baseAuthorityYear="1843" box="[565,661,502,526]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="10" phylum="Mollusca" rank="species" species="vittata">
<emphasis id="B9617238E523D91D0478FE6044B4B54D" box="[565,661,502,526]" italics="true" pageId="10" pageNumber="10">T. vittata</emphasis>
</taxonomicName>
has a far more prominent spire, deeper body whorl, narrower umbilicus, and a more extensively flared aperture lip, often with a wide extension in the columella region that partially obscures the umbilicus at full maturity. Fischer-Piette
<emphasis id="B9617238E523D91D0714FDC047B3B52D" box="[345,402,598,622]" italics="true" pageId="10" pageNumber="10">et al.</emphasis>
(1993) stated that the
<typeStatus id="54AE1088E523D91D04DEFDC544D4B52E" box="[659,757,595,621]" pageId="10" pageNumber="10" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C15D5A9E523D91D0551FDC045A0B52D" baseAuthorityName="Sowerby" baseAuthorityYear="1843" box="[796,897,598,622]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="10" phylum="Mollusca" rank="species" species="vittata">
<emphasis id="B9617238E523D91D0551FDC045A0B52D" box="[796,897,598,622]" italics="true" pageId="10" pageNumber="10">T. vittata</emphasis>
</taxonomicName>
was in the NHMUK, but it could not be located there (Ablett
<emphasis id="B9617238E523D91D043AFDE04492B5CD" box="[631,691,630,654]" italics="true" pageId="10" pageNumber="10">in litt</emphasis>
.
<date id="FFAB88EAE523D91D048CFDE2451BB5CE" box="[705,826,627,654]" pageId="10" pageNumber="10" value="2011-11">Nov. 2011</date>
).
<taxonomicName id="4C15D5A9E523D91D0500FDE046D8B5EE" authority="(Pfeiffer, 1847)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1847" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="10" phylum="Mollusca" rank="species" species="semidecussata">
<emphasis id="B9617238E523D91D0500FDE0422BB5CD" box="[845,1034,630,654]" italics="true" pageId="10" pageNumber="10">T. semidecussata</emphasis>
(Pfeiffer, 1847)
</taxonomicName>
, is also similar and though a very variable species, like
<taxonomicName id="4C15D5A9E523D91D053CFD0045F4B5ED" baseAuthorityName="Sowerby" baseAuthorityYear="1843" box="[881,981,662,686]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="10" phylum="Mollusca" rank="species" species="vittata">
<emphasis id="B9617238E523D91D053CFD0045F4B5ED" box="[881,981,662,686]" italics="true" pageId="10" pageNumber="10">T. vittata</emphasis>
</taxonomicName>
, it has a more elevated spire, tumescent body whorl, narrower umbilicus and a more well developed, strongly reflected columella lip.
</paragraph>
<paragraph id="8BAAAE2AE523D91D069DFD654234B40E" blockId="10.[181,1143,203,846]" pageId="10" pageNumber="10">
Specimens of
<taxonomicName id="4C15D5A9E523D91D0725FD6047FBB44D" authorityName="Griffiths &amp; Herbert" authorityYear="2013" box="[360,474,758,782]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="10" phylum="Mollusca" rank="species" species="humbug">
<emphasis id="B9617238E523D91D0725FD6047FBB44D" box="[360,474,758,782]" italics="true" pageId="10" pageNumber="10">T. humbug</emphasis>
</taxonomicName>
from the Tsingy de Bemaraha commonly have a slightly more elevated spire than those from
<taxonomicName id="4C15D5A9E523D91D0442FC854498B46E" authorityName=", Antsingimavo" authorityYear="2013" box="[527,697,787,813]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="10" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
, but H:D ratios for the two populations overlap considerably (H:D=0.470.61 compared to 0.460.53 respectively).
</paragraph>
</subSubSection>
</treatment>
</document>