treatments-xml/data/03/BB/0D/03BB0D181F0EFFC8718BFE4BFAC2FD8B.xml
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<document id="7F01DC0B733937123BB366F61FE6B52D" ID-CLB-Dataset="27727" ID-DOI="10.11646/zootaxa.4535.1.1" ID-GBIF-Dataset="a047bcd9-ab65-4e3f-b07a-c830c7af1072" ID-ISSN="1175-5326" ID-Zenodo-Dep="2615773" ID-ZooBank="ABE9DB1F-7378-4571-90C4-A3FDB66527F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553788891023" checkinUser="plazi" docAuthor="Freyhof, Jörg, Bayçelebi, Esra &amp; Geiger, Matthias" docDate="2018" docId="03BB0D181F0EFFC8718BFE4BFAC2FD8B" docLanguage="en" docName="zootaxa.4535.1.1.pdf" docOrigin="Zootaxa 4535 (1)" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Cobitis erkakanae Freyhof &amp; Bayçelebi &amp; Geiger 2018, new species" docType="treatment" docVersion="11" lastPageNumber="58" masterDocId="FF8275601F38FFF1711CFFFCFFABFF93" masterDocTitle="Review of the genus Cobitis in the Middle East, with the description of eight new species (Teleostei: Cobitidae)" masterLastPageNumber="75" masterPageNumber="1" pageNumber="55" updateTime="1698715239854" updateUser="ExternalLinkService">
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<mods:title id="9D183775B8A1AFC4DF7FC7CE029F591E">Review of the genus Cobitis in the Middle East, with the description of eight new species (Teleostei: Cobitidae)</mods:title>
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<mods:namePart id="D7F3A4DA49B7FCC04A52ACA38FA06F7C">Freyhof, Jörg</mods:namePart>
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<mods:namePart id="F053E54299B56C1628917EBAC10C1349">Bayçelebi, Esra</mods:namePart>
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<mods:namePart id="B2EAF17FC720443F7A6723A87A817FBD">Geiger, Matthias</mods:namePart>
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<mods:title id="0F477A714C6C4C83D8DACA4059C2CB52">Zootaxa</mods:title>
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<mods:date id="36F356FACA7A219ED490BD273C1C8589">2018</mods:date>
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<mods:number id="C97B4A6582BB92235814C0116B0B8E18">2018-12-21</mods:number>
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<treatment id="03BB0D181F0EFFC8718BFE4BFAC2FD8B" ID-DOI="http://doi.org/10.5281/zenodo.5699585" ID-GBIF-Taxon="154521903" ID-Zenodo-Dep="5699585" LSID="urn:lsid:plazi:treatment:03BB0D181F0EFFC8718BFE4BFAC2FD8B" httpUri="http://treatment.plazi.org/id/03BB0D181F0EFFC8718BFE4BFAC2FD8B" lastPageId="57" lastPageNumber="58" pageId="54" pageNumber="55">
<subSubSection id="C308EF851F0EFFC7718BFE4BFDBBFE41" box="[151,528,439,466]" pageId="54" pageNumber="55" type="nomenclature">
<paragraph id="8BADBC0E1F0EFFC7718BFE4BFDBBFE41" blockId="54.[151,528,439,499]" box="[151,528,439,466]" pageId="54" pageNumber="55">
<heading id="D0E50B621F0EFFC7718BFE4BFDBBFE41" bold="true" box="[151,528,439,466]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
<emphasis id="B966601C1F0EFFC7718BFE4BFDBBFE41" bold="true" box="[151,528,439,466]" pageId="54" pageNumber="55">
<taxonomicName id="4C12C78D1F0EFFC7718BFE4BFED9FE42" authority="Freyhof &amp; Bayçelebi &amp; Geiger, 2018" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[151,370,439,465]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="erkakanae" status="sp. nov.">
<emphasis id="B966601C1F0EFFC7718BFE4BFED9FE42" bold="true" box="[151,370,439,465]" italics="true" pageId="54" pageNumber="55">Cobitis erkakanae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A255DD671F0EFFC7709CFE44FDBBFE41" box="[384,528,440,466]" pageId="54" pageNumber="55" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C308EF851F0EFFC7718BFE26FE88FE60" box="[151,291,474,499]" pageId="54" pageNumber="55" type="description">
<paragraph id="8BADBC0E1F0EFFC7718BFE26FE88FE60" blockId="54.[151,528,439,499]" box="[151,291,474,499]" pageId="54" pageNumber="55">
(
<figureCitation id="1329A08B1F0EFFC77183FE26FEB0FE60" box="[159,283,474,499]" captionStart-0="FIGURE 42" captionStart-1="FIGURE 43" captionStart-2="FIGURE 44" captionStartId-0="55.[151,250,813,835]" captionStartId-1="55.[151,250,1387,1409]" captionStartId-2="55.[151,250,1931,1953]" captionTargetBox-0="[190,1411,335,752]" captionTargetBox-1="[187,1398,964,1342]" captionTargetBox-2="[151,1436,1520,1909]" captionTargetId-0="figure@55.[151,1436,301,792]" captionTargetId-1="figure@55.[151,1436,947,1366]" captionTargetId-2="figure@55.[151,1436,1520,1909]" captionTargetPageId-0="55" captionTargetPageId-1="55" captionTargetPageId-2="55" captionText-0="FIGURE 42. Cobitis erkakanae, left column from the top: males, ZFMK ICH-98958, holotype, 46 mm SL; paratypes, FSJF 2510, 49 mm SL, 49 mm SL; right column from the top: females: FSJF 2427, 67 mm SL, 68 mm SL, 60 mm SL; Turkey: river connecting Lakes Gölbaşı and Azaplı." captionText-1="FIGURE 43. Cobitis erkakanae, left column from the top: males, ZFMK ICH-98958, holotype, 46 mm SL; paratypes, FSJF 2510, 49 mm SL, 49 mm SL; right column from the top: females: FSJF 2427, 67 mm SL, 68 mm SL, 60 mm SL; Turkey: river connecting Lakes Gölbaşı and Azaplı." captionText-2="FIGURE 44. Cobitis erkakanae, FSJF 2510, female, about 65 mm SL; Turkey: river connecting Lakes Gölbaşı and Azaplı." httpUri-0="https://zenodo.org/record/2615859/files/figure.png" httpUri-1="https://zenodo.org/record/2615861/files/figure.png" httpUri-2="https://zenodo.org/record/2615863/files/figure.png" pageId="54" pageNumber="55">Fig. 4244</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C308EF851F0EFFC7718BFDDEFEB0FD7C" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="8BADBC0E1F0EFFC7718BFDDEFE56FDCC" blockId="54.[151,1437,546,2018]" pageId="54" pageNumber="55">
<materialsCitation id="3B7AB6531F0EFFC7718BFDDEFE56FDCC" ID-GBIF-Occurrence="2120487013" collectionCode="ZFMK" country="Turkey" county="Lakes Golbasi" location="Golbasi" municipality="Azapli" pageId="54" pageNumber="55" specimenCode="ICH-98958, 49" specimenCount="1" stateProvince="Adiyaman" typeStatus="holotype">
<emphasis id="B966601C1F0EFFC7718BFDDEFEA0FDA8" bold="true" box="[151,267,546,571]" pageId="54" pageNumber="55">
<typeStatus id="54A902AC1F0EFFC7718BFDDEFEADFDA8" box="[151,262,546,571]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED0324CB1F0EFFC77008FDDFFEC9FDA9" box="[276,354,547,570]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="54" pageNumber="55" type="Museum">ZFMK</collectionCode>
<specimenCode id="DBB414751F0EFFC77077FDDEFE40FDA8" box="[363,491,546,571]" collectionCode="ICH-" pageId="54" pageNumber="55">ICH-98958</specimenCode>
,
<specimenCode id="DBB414751F0EFFC770E7FDDFFDBCFDA8" box="[507,535,547,571]" collectionCode="ICH-" pageId="54" pageNumber="55">49</specimenCode>
mm SL;
<collectingCountry id="F305FC9E1F0EFFC7739BFDDEFD70FDA8" box="[647,731,546,571]" name="Turkey" pageId="54" pageNumber="55">Turkey</collectingCountry>
:
<collectingRegion id="49D672EC1F0EFFC773F4FDDEFCF6FDA8" box="[744,861,546,571]" country="Turkey" name="Adiyaman" pageId="54" pageNumber="55">Adıyaman</collectingRegion>
prov.: river connecting
<collectingCounty id="62CCC4821F0EFFC7756EFDDEFABDFDA8" box="[1138,1302,546,571]" pageId="54" pageNumber="55">Lakes Gölbaşı</collectingCounty>
and
<collectingMunicipality id="6BC926741F0EFFC7744DFDDEFA37FDA8" box="[1361,1436,546,571]" pageId="54" pageNumber="55">Azaplı</collectingMunicipality>
west of
<location id="8ECDEAD51F0EFFC771F3FDBAFEE2FDCC" LSID="urn:lsid:plazi:treatment:03BB0D181F0EFFC8718BFE4BFAC2FD8B:8ECDEAD51F0EFFC771F3FDBAFEE2FDCC" box="[239,329,582,607]" country="Turkey" county="Lakes Golbasi" municipality="Azapli" name="Golbasi" pageId="54" pageNumber="55" stateProvince="Adiyaman">Gölbaşı</location>
, 37.790 37.626.
</materialsCitation>
</paragraph>
<paragraph id="8BADBC0E1F0EFFC771DBFD96FC00FD10" blockId="54.[151,1437,546,2018]" box="[199,939,618,643]" pageId="54" pageNumber="55">
<materialsCitation id="3B7AB6531F0EFFC771DBFD96FC0CFD10" ID-GBIF-Occurrence="2120485282" box="[199,935,618,643]" collectionCode="FSJF" country="Turkey" county="Lakes Golbasi" location="Golbasi" municipality="Azapli" pageId="54" pageNumber="55" specimenCode="FSJF 2510, 16, 34-65" specimenCount="1" stateProvince="Adiyaman" typeStatus="paratype">
<emphasis id="B966601C1F0EFFC771DBFD96FEEDFD10" bold="true" box="[199,326,618,643]" pageId="54" pageNumber="55">
<typeStatus id="54A902AC1F0EFFC771DBFD96FEEAFD10" box="[199,321,618,643]" pageId="54" pageNumber="55" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="DBB414751F0EFFC77050FD97FE63FD10" box="[332,456,618,643]" collectionCode="FSJF" pageId="54" pageNumber="55">FSJF 2510</specimenCode>
,
<specimenCode id="DBB414751F0EFFC770CFFD96FE5AFD10" box="[467,497,618,643]" collectionCode="FSJF" pageId="54" pageNumber="55">16</specimenCode>
,
<quantity id="4CEA11EB1F0EFFC770E1FD97FDDDFD11" box="[509,630,618,643]" metricMagnitude="-2" metricUnit="m" metricValue="4.95" metricValueMax="6.5" metricValueMin="3.4" pageId="54" pageNumber="55" unit="mm" value="49.5" valueMax="65.0" valueMin="34.0">
<specimenCode id="DBB414751F0EFFC770E1FD97FDE8FD10" box="[509,579,618,643]" collectionCode="FSJF" pageId="54" pageNumber="55">3465</specimenCode>
mm
</quantity>
SL; same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BADBC0E1F0EFFC771DBFD72FEB0FD7C" blockId="54.[151,1437,546,2018]" pageId="54" pageNumber="55">
<emphasis id="B966601C1F0EFFC771DBFD72FD4FFD34" bold="true" box="[199,740,654,679]" pageId="54" pageNumber="55">Material used in molecular genetic analysis.</emphasis>
<materialsCitation id="3B7AB6531F0EFFC773F2FD73FEBCFD7C" ID-GBIF-Occurrence="2120487405" collectionCode="FSJF, KJ" country="Turkey" county="Lakes Golbasi" location="Golbasi" municipality="Azapli" pageId="54" pageNumber="55" specimenCode="DNA-982" specimenCount="1" stateProvince="Adiyaman">
<collectionCode id="ED0324CB1F0EFFC773F2FD73FC83FD34" box="[750,808,655,679]" pageId="54" pageNumber="55">FSJF</collectionCode>
<specimenCode id="DBB414751F0EFFC7722FFD73FC0DFD34" box="[819,934,655,679]" collectionCode="DNA-" pageId="54" pageNumber="55">DNA-982</specimenCode>
;
<collectingCountry id="F305FC9E1F0EFFC772A9FD72FBA2FD34" box="[949,1033,654,679]" name="Turkey" pageId="54" pageNumber="55">Turkey</collectingCountry>
:
<collectingRegion id="49D672EC1F0EFFC7750BFD72FB27FD34" box="[1047,1164,654,679]" country="Turkey" name="Adiyaman" pageId="54" pageNumber="55">Adıyaman</collectingRegion>
prov.: river connecting
<collectingCounty id="62CCC4821F0EFFC7718BFD4EFE91FD58" box="[151,314,690,715]" pageId="54" pageNumber="55">Lakes Gölbaşı</collectingCounty>
and
<collectingMunicipality id="6BC926741F0EFFC77068FD4EFE14FD58" box="[372,447,690,715]" pageId="54" pageNumber="55">Azaplı</collectingMunicipality>
west of
<location id="8ECDEAD51F0EFFC77338FD4EFDD5FD58" LSID="urn:lsid:plazi:treatment:03BB0D181F0EFFC8718BFE4BFAC2FD8B:8ECDEAD51F0EFFC77338FD4EFDD5FD58" box="[548,638,690,715]" country="Turkey" county="Lakes Golbasi" municipality="Azapli" name="Golbasi" pageId="54" pageNumber="55" stateProvince="Adiyaman">Gölbaşı</location>
, 37.790 37.626. (GenBank accession numbers:
<accessionNumber id="944121ED1F0EFFC77583FD4EFABEFD58" box="[1183,1301,690,715]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KJ552949" pageId="54" pageNumber="55">
<collectionCode id="ED0324CB1F0EFFC77583FD4EFB15FD58" box="[1183,1214,690,715]" pageId="54" pageNumber="55">KJ</collectionCode>
552949
</accessionNumber>
,
<accessionNumber id="944121ED1F0EFFC7743EFD4EFA33FD58" box="[1314,1432,690,715]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KJ552977" pageId="54" pageNumber="55">KJ552977</accessionNumber>
,
<accessionNumber id="944121ED1F0EFFC7718BFD2AFEA4FD7C" box="[151,271,726,751]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KJ553235" pageId="54" pageNumber="55">
<collectionCode id="ED0324CB1F0EFFC7718BFD2AFF1CFD7C" box="[151,183,726,751]" pageId="54" pageNumber="55">KJ</collectionCode>
553235
</accessionNumber>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C308EF851F0EFFC771DBFD06FAD6FBEC" pageId="54" pageNumber="55" type="diagnosis">
<paragraph id="8BADBC0E1F0EFFC771DBFD06FBC9FC31" blockId="54.[151,1437,546,2018]" pageId="54" pageNumber="55">
<emphasis id="B966601C1F0EFFC771DBFD06FEEAFC80" bold="true" box="[199,321,762,787]" pageId="54" pageNumber="55">Diagnosis.</emphasis>
<taxonomicName id="4C12C78D1F0EFFC77054FD07FDBFFC80" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[328,532,763,787]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="erkakanae">
<emphasis id="B966601C1F0EFFC77054FD07FDBFFC80" box="[328,532,763,787]" italics="true" pageId="54" pageNumber="55">Cobitis erkakanae</emphasis>
</taxonomicName>
is distinguished from other
<taxonomicName id="4C12C78D1F0EFFC77251FD07FC35FC80" box="[845,926,763,787]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F0EFFC77251FD07FC35FC80" box="[845,926,763,787]" italics="true" pageId="54" pageNumber="55">Cobitis</emphasis>
</taxonomicName>
species in the Anatolian Mediterranean basin by a combination of characters, none of them unique. It is distinguished from
<taxonomicName id="4C12C78D1F0EFFC77516FCE3FBDAFCA5" authorityName="Erk'akan, Özeren &amp; Nalbant" authorityYear="2008" box="[1034,1137,799,822]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="evreni">
<emphasis id="B966601C1F0EFFC77516FCE3FBDAFCA5" box="[1034,1137,799,822]" italics="true" pageId="54" pageNumber="55">C. evreni</emphasis>
</taxonomicName>
from the adjacent Ceyhan River drainage by having a small, roundish or comma-shaped black spot at the upper caudal-fin base (vs. absent) and the blotches in Z2 and Z4 being well separated, horizontally elongate and often fused to adjacent once but not forming a stripe (vs. Z2 and Z4 formed by one continuous stripe on the anterior flank).
</paragraph>
<paragraph id="8BADBC0E1F0EFFC771DBFC52FAD6FBEC" blockId="54.[151,1437,546,2018]" pageId="54" pageNumber="55">
<taxonomicName id="4C12C78D1F0EFFC771DBFC52FE3DFC54" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[199,406,942,967]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="erkakanae">Cobitis erkakanae</taxonomicName>
is distinguished from
<taxonomicName id="4C12C78D1F0EFFC773BBFC53FCB4FC54" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[679,799,942,967]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="aliyeae">C. aliyeae</taxonomicName>
,
<taxonomicName id="4C12C78D1F0EFFC7722CFC53FC13FC54" box="[816,952,942,967]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="anabelae">C. anabelae</taxonomicName>
and
<taxonomicName id="4C12C78D1F0EFFC772E6FC53FB2DFC54" authorityName="Krupp &amp; Moubayed" authorityYear="1992" box="[1018,1158,942,967]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="levantina">C. levantina</taxonomicName>
by having horizontally elongated blotches in Z2 and Z4, often fused to adjacent once, (vs. blotches in Z2 and Z4 usually well distinct from each other, roundish, or squarish, not elongate), a yellow background colour (vs. silvery or whitish), pigmentation in Z1 well separated from Z2 along its complete length (vs. Z1 and Z2 usually fused into a mottled pattern in
<taxonomicName id="4C12C78D1F0EFFC77014FBBFFE3BFBC8" authorityName="Krupp &amp; Moubayed" authorityYear="1992" box="[264,400,1091,1115]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="levantina">
<emphasis id="B966601C1F0EFFC77014FBBFFE3BFBC8" box="[264,400,1091,1115]" italics="true" pageId="54" pageNumber="55">C. levantina</emphasis>
</taxonomicName>
), the flank yellowish in life (vs. silvery greyish) and the caudal fin with 46 wide, regularlyshaped bands (vs. many, very narrow and irregularly-shaped bands in
<taxonomicName id="4C12C78D1F0EFFC772BAFB9BFBB3FBED" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[934,1048,1126,1150]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="aliyeae">
<emphasis id="B966601C1F0EFFC772BAFB9BFBB3FBED" box="[934,1048,1126,1150]" italics="true" pageId="54" pageNumber="55">C. aliyeae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C12C78D1F0EFFC7753AFB9BFB06FBED" box="[1062,1197,1126,1150]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="anabelae">
<emphasis id="B966601C1F0EFFC7753AFB9BFB06FBED" box="[1062,1197,1126,1150]" italics="true" pageId="54" pageNumber="55">C. anabelae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C12C78D1F0EFFC775F8FB9BFAC7FBED" authorityName="Krupp &amp; Moubayed" authorityYear="1992" box="[1252,1388,1126,1150]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="levantina">
<emphasis id="B966601C1F0EFFC775F8FB9BFAC7FBED" box="[1252,1388,1126,1150]" italics="true" pageId="54" pageNumber="55">C. levantina</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C308EF851F0EFFC971DBFB77FA22F801" lastPageId="56" lastPageNumber="57" pageId="54" pageNumber="55" type="description">
<paragraph id="8BADBC0E1F0EFFC771DBFB77FDF0FA0C" blockId="54.[151,1437,546,2018]" pageId="54" pageNumber="55">
<taxonomicName id="4C12C78D1F0EFFC771DBFB77FE3DFB30" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[199,406,1163,1187]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="erkakanae">
<emphasis id="B966601C1F0EFFC771DBFB77FE3DFB30" box="[199,406,1163,1187]" italics="true" pageId="54" pageNumber="55">Cobitis erkakanae</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C12C78D1F0EFFC77385FB77FC9EFB30" authorityName="Bacescu" authorityYear="1962" box="[665,821,1163,1187]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="battalgilae">
<emphasis id="B966601C1F0EFFC77385FB77FC9EFB30" box="[665,821,1163,1187]" italics="true" pageId="54" pageNumber="55">C. battalgilae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C12C78D1F0EFFC77273FB77FC5DFB30" authorityName="Battalgazi" authorityYear="1944" box="[879,1014,1163,1187]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F0EFFC77273FB77FC5DFB30" box="[879,1014,1163,1187]" italics="true" pageId="54" pageNumber="55">C. phrygica</emphasis>
</taxonomicName>
by having one series of horizontally elongate blotches in Z4 (vs. blotches in Z4 usually roundish, squarish or vertically elongate, often very densely set, dissociated in many individuals, especially in
<taxonomicName id="4C12C78D1F0EFFC773AEFB2FFC91FB78" authorityName="Battalgazi" authorityYear="1944" box="[690,826,1235,1259]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F0EFFC773AEFB2FFC91FB78" box="[690,826,1235,1259]" italics="true" pageId="54" pageNumber="55">C. phrygica</emphasis>
</taxonomicName>
, forming a wide band or field of small, irregularlyshaped blotches and spots), usually lacking flank pigmentation below Z4 (vs. present in most
<taxonomicName id="4C12C78D1F0EFFC775DBFB0BFAE6FA9D" authorityName="Battalgazi" authorityYear="1944" box="[1223,1357,1270,1294]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F0EFFC775DBFB0BFAE6FA9D" box="[1223,1357,1270,1294]" italics="true" pageId="54" pageNumber="55">C. phrygica</emphasis>
</taxonomicName>
and in some
<taxonomicName id="4C12C78D1F0EFFC771C5FAE7FED9FAA0" authorityName="Bacescu" authorityYear="1962" box="[217,370,1307,1331]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="battalgilae">
<emphasis id="B966601C1F0EFFC771C5FAE7FED9FAA0" box="[217,370,1307,1331]" italics="true" pageId="54" pageNumber="55">C. battalgilae</emphasis>
</taxonomicName>
), having a small, roundish or comma-shaped black spot at the upper caudal-fin base (vs. absent in
<taxonomicName id="4C12C78D1F0EFFC771A8FAC3FE93FAC5" authorityName="Battalgazi" authorityYear="1944" box="[180,312,1342,1366]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F0EFFC771A8FAC3FE93FAC5" box="[180,312,1342,1366]" italics="true" pageId="54" pageNumber="55">C. phrygica</emphasis>
</taxonomicName>
), a bifurcate suborbital spine (vs. simple in
<taxonomicName id="4C12C78D1F0EFFC7723AFAC3FC01FAC5" authorityName="Battalgazi" authorityYear="1944" box="[806,938,1342,1366]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F0EFFC7723AFAC3FC01FAC5" box="[806,938,1342,1366]" italics="true" pageId="54" pageNumber="55">C. phrygica</emphasis>
</taxonomicName>
), a greater prepelvic (5761% SL vs.
<quantity id="4CEA11EB1F0EFFC7744AFAC2FF06FAE8" metricMagnitude="0" metricUnit="m" metricValue="1.3969999999999998" metricValueMax="1.4224" metricValueMin="1.3716" pageId="54" pageNumber="55" unit="in" value="55.0" valueMax="56.0" valueMin="54.0">5456 in</quantity>
<taxonomicName id="4C12C78D1F0EFFC771A8FA9FFEE6FAE8" authorityName="Bacescu" authorityYear="1962" box="[180,333,1379,1403]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="battalgilae">
<emphasis id="B966601C1F0EFFC771A8FA9FFEE6FAE8" box="[180,333,1379,1403]" italics="true" pageId="54" pageNumber="55">C. battalgilae</emphasis>
</taxonomicName>
), preanal distance (8285% SL vs. 7480) and a narrower dorsal-fin depth (1619% SL vs.
<quantity id="4CEA11EB1F0EFFC7744AFA9FFF06FA0C" metricMagnitude="-1" metricUnit="m" metricValue="5.461" metricValueMax="5.588" metricValueMin="5.334" pageId="54" pageNumber="55" unit="in" value="21.5" valueMax="22.0" valueMin="21.0">2122 in</quantity>
<taxonomicName id="4C12C78D1F0EFFC771A8FA7BFEE6FA0D" authorityName="Bacescu" authorityYear="1962" box="[180,333,1414,1438]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="battalgilae">
<emphasis id="B966601C1F0EFFC771A8FA7BFEE6FA0D" box="[180,333,1414,1438]" italics="true" pageId="54" pageNumber="55">C. battalgilae</emphasis>
</taxonomicName>
;
<quantity id="4CEA11EB1F0EFFC77040FA7BFE14FA0C" box="[348,447,1414,1439]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" metricValueMax="5.334" metricValueMin="4.826" pageId="54" pageNumber="55" unit="in" value="20.0" valueMax="21.0" valueMin="19.0">1921 in</quantity>
<taxonomicName id="4C12C78D1F0EFFC770DAFA7BFDE0FA0D" authorityName="Battalgazi" authorityYear="1944" box="[454,587,1414,1438]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="phrygica">
<emphasis id="B966601C1F0EFFC770DAFA7BFDE0FA0D" box="[454,587,1414,1438]" italics="true" pageId="54" pageNumber="55">C. phrygica</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BADBC0E1F0EFFC771DBFA57FD40F9BC" blockId="54.[151,1437,546,2018]" pageId="54" pageNumber="55">
<taxonomicName id="4C12C78D1F0EFFC771DBFA57FE3DFA50" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[199,406,1451,1475]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="erkakanae">
<emphasis id="B966601C1F0EFFC771DBFA57FE3DFA50" box="[199,406,1451,1475]" italics="true" pageId="54" pageNumber="55">Cobitis erkakanae</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C12C78D1F0EFFC7738BFA57FC9AFA50" authorityName="Erkakan, Ozdemir &amp; Ozeren" authorityYear="2017" box="[663,817,1451,1475]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="sipahilerae">
<emphasis id="B966601C1F0EFFC7738BFA57FC9AFA50" box="[663,817,1451,1475]" italics="true" pageId="54" pageNumber="55">C sipahilerae</emphasis>
</taxonomicName>
by having 35 brown, large and roundish blotches on the back anterior to the dorsal-fin origin (vs. back plain cream-brown, except few individuals with 13 narrow, squarish, brown bars) and a series of distinct blotches in Z4 (vs. a dark-brown stripe in Z4 reaching from above the pectoral-fin base until the dorsal-fin base or beyond).
</paragraph>
<paragraph id="8BADBC0E1F0EFFC771DBF9C0FBA5F8BD" blockId="54.[151,1437,546,2018]" pageId="54" pageNumber="55">
It is distinguished from
<taxonomicName id="4C12C78D1F0EFFC770C4F9C0FD99F9C7" baseAuthorityName="Heckel" baseAuthorityYear="1848" box="[472,562,1596,1620]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="linea">
<emphasis id="B966601C1F0EFFC770C4F9C0FD99F9C7" box="[472,562,1596,1620]" italics="true" pageId="54" pageNumber="55">C. linea</emphasis>
</taxonomicName>
by having large, well separated, usually horizontally elongate blotches in Z4, often fused to each other (vs. very small and roundish or comma-shaped, very densely set, often fused into stripes or dissociated into a field of several rows of blotches).
<taxonomicName id="4C12C78D1F0EFFC77285F979FBC6F90E" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[921,1133,1669,1693]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="erkakanae">
<emphasis id="B966601C1F0EFFC77285F979FBC6F90E" box="[921,1133,1669,1693]" italics="true" pageId="54" pageNumber="55">Cobitis erkakanae</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C12C78D1F0EFFC7749EF97AFEB9F952" authorityName="Coad &amp; Sarieyyupoglu" authorityYear="1988" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="elazigensis">
<emphasis id="B966601C1F0EFFC7749EF97AFEB9F952" italics="true" pageId="54" pageNumber="55">C. elazigensis</emphasis>
</taxonomicName>
by the blotches in Z2 and Z4 being usually horizontally elongate and often fused to each other forming short stripes (vs. blotches in Z2 and Z4 roundish, very widely spaced and not fused to each other), 35 dark-brown blotches on the back anterior to the dorsal fin-origin (vs. 59) and being much smaller, no female was found to be larger than
<quantity id="4CEA11EB1F0EFFC7700AF8EAFECEF8BE" box="[278,357,1814,1838]" metricMagnitude="-2" metricUnit="m" metricValue="8.3" pageId="54" pageNumber="55" unit="mm" value="83.0">83 mm</quantity>
SL (vs. female growing up to
<quantity id="4CEA11EB1F0EFFC773A7F8E9FCB3F8BE" box="[699,792,1813,1838]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="54" pageNumber="55" unit="mm" value="160.0">160 mm</quantity>
SL in
<taxonomicName id="4C12C78D1F0EFFC7727FF8EAFC55F8BE" authorityName="Coad &amp; Sarieyyupoglu" authorityYear="1988" box="[867,1022,1813,1837]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="elazigensis">
<emphasis id="B966601C1F0EFFC7727FF8EAFCDDF8BE" box="[867,886,1814,1837]" italics="true" pageId="54" pageNumber="55">C</emphasis>
.
<emphasis id="B966601C1F0EFFC7729FF8E9FC55F8BE" box="[899,1022,1813,1837]" italics="true" pageId="54" pageNumber="55">elazigensis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BADBC0E1F0EFFC771DBF8C5FDF6F871" blockId="54.[151,1437,546,2018]" pageId="54" pageNumber="55">
<emphasis id="B966601C1F0EFFC771DBF8C5FEF2F8C1" bold="true" box="[199,345,1849,1874]" pageId="54" pageNumber="55">Description.</emphasis>
See
<figureCitation id="1329A08B1F0EFFC77091F8C5FD85F8C1" box="[397,558,1849,1874]" captionStart-0="FIGURE 42" captionStart-1="FIGURE 43" captionStart-2="FIGURE 44" captionStartId-0="55.[151,250,813,835]" captionStartId-1="55.[151,250,1387,1409]" captionStartId-2="55.[151,250,1931,1953]" captionTargetBox-0="[190,1411,335,752]" captionTargetBox-1="[187,1398,964,1342]" captionTargetBox-2="[151,1436,1520,1909]" captionTargetId-0="figure@55.[151,1436,301,792]" captionTargetId-1="figure@55.[151,1436,947,1366]" captionTargetId-2="figure@55.[151,1436,1520,1909]" captionTargetPageId-0="55" captionTargetPageId-1="55" captionTargetPageId-2="55" captionText-0="FIGURE 42. Cobitis erkakanae, left column from the top: males, ZFMK ICH-98958, holotype, 46 mm SL; paratypes, FSJF 2510, 49 mm SL, 49 mm SL; right column from the top: females: FSJF 2427, 67 mm SL, 68 mm SL, 60 mm SL; Turkey: river connecting Lakes Gölbaşı and Azaplı." captionText-1="FIGURE 43. Cobitis erkakanae, left column from the top: males, ZFMK ICH-98958, holotype, 46 mm SL; paratypes, FSJF 2510, 49 mm SL, 49 mm SL; right column from the top: females: FSJF 2427, 67 mm SL, 68 mm SL, 60 mm SL; Turkey: river connecting Lakes Gölbaşı and Azaplı." captionText-2="FIGURE 44. Cobitis erkakanae, FSJF 2510, female, about 65 mm SL; Turkey: river connecting Lakes Gölbaşı and Azaplı." httpUri-0="https://zenodo.org/record/2615859/files/figure.png" httpUri-1="https://zenodo.org/record/2615861/files/figure.png" httpUri-2="https://zenodo.org/record/2615863/files/figure.png" pageId="54" pageNumber="55">Figures 4244</figureCitation>
for general appearance and
<tableCitation id="C69089B51F0EFFC77275F8C5FC10F8C1" box="[873,955,1849,1874]" captionStart="TABLE 6" captionStartId="56.[151,239,152,175]" captionText="TABLE 6. Morphometric data of Cobitis erkakanae (holotype ZFMK ICH-98958 and paratypes FSJF 2427; n = 13). The calculations include the holotype." pageId="54" pageNumber="55">Table 6</tableCitation>
for morphometric data of the
<typeStatus id="54A902AC1F0EFFC77417F8C5FAC6F8C1" box="[1291,1389,1849,1874]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D1477871F0EFFC7718BF8A1FE80F8E5" box="[151,299,1885,1910]" pageId="54" pageNumber="55" type="generic" typeStatus="paratypes">16 paratypes</specimenCount>
. Greatest body depth at or slightly anterior to dorsal-fin origin, decreasing towards caudal-fin base. Head profile slightly convex, head length 1.21.6 times in body depth. Snout pointed, its length 0.70.9 times in postorbital length. Eye diameter 0.3 times in head depth at eye, 1.42.2 times in interorbital width. Caudal peduncle 1.01.5 times longer than deep
</paragraph>
<paragraph id="8BADBC0E1F0FFFC671DBFF6BFBA8FF6B" blockId="55.[151,1436,151,248]" pageId="55" pageNumber="56">
Pelvic axillary lobe present or absent. Margin of dorsal and anal fins convex. Caudal fin truncate or slightly rounded. A shallow dorsal and ventral keel on caudal peduncle. External part of the suborbital spine bifurcate, reaching slightly beyond centre of eye. Largest recorded specimen
<quantity id="4CEA11EB1F0FFFC6729AFF23FC7FFF64" box="[902,980,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" pageId="55" pageNumber="56" unit="mm" value="67.0">67 mm</quantity>
SL.
</paragraph>
<caption id="DF6DEC861F0FFFC6718BFCD1FDB1FC12" httpUri="https://zenodo.org/record/2615859/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,250,813,835]" targetBox="[190,1411,335,752]" targetPageId="55">
<paragraph id="8BADBC0E1F0FFFC6718BFCD1FDB1FC12" blockId="55.[151,1436,813,897]" pageId="55" pageNumber="56">
<emphasis id="B966601C1F0FFFC6718BFCD1FE8AFCD0" bold="true" box="[151,289,813,835]" pageId="55" pageNumber="56">FIGURE 42.</emphasis>
<taxonomicName id="4C12C78D1F0FFFC67034FCD1FE49FCD0" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[296,482,813,835]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="erkakanae">
<emphasis id="B966601C1F0FFFC67034FCD1FE49FCD0" box="[296,482,813,835]" italics="true" pageId="55" pageNumber="56">Cobitis erkakanae</emphasis>
</taxonomicName>
, left column from the top: males, ZFMK ICH-98958, holotype, 46 mm SL; paratypes, FSJF 2510, 49 mm SL, 49 mm SL; right column from the top: females: FSJF 2427, 67 mm SL, 68 mm SL, 60 mm SL; Turkey: river connecting Lakes Gölbaşı and Azaplı.
</paragraph>
</caption>
<caption id="DF6DEC861F0FFFC6718BFA97FDB1FA2C" httpUri="https://zenodo.org/record/2615861/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,250,1387,1409]" targetBox="[187,1398,964,1342]" targetPageId="55">
<paragraph id="8BADBC0E1F0FFFC6718BFA97FDB1FA2C" blockId="55.[151,1436,1387,1471]" pageId="55" pageNumber="56">
<emphasis id="B966601C1F0FFFC6718BFA97FE8AFA12" bold="true" box="[151,289,1387,1409]" pageId="55" pageNumber="56">FIGURE 43.</emphasis>
<taxonomicName id="4C12C78D1F0FFFC67034FA97FE49FA12" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[296,482,1387,1409]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="erkakanae">
<emphasis id="B966601C1F0FFFC67034FA97FE49FA12" box="[296,482,1387,1409]" italics="true" pageId="55" pageNumber="56">Cobitis erkakanae</emphasis>
</taxonomicName>
, left column from the top: males, ZFMK ICH-98958, holotype, 46 mm SL; paratypes, FSJF 2510, 49 mm SL, 49 mm SL; right column from the top: females: FSJF 2427, 67 mm SL, 68 mm SL, 60 mm SL; Turkey: river connecting Lakes Gölbaşı and Azaplı.
</paragraph>
</caption>
<caption id="DF6DEC861F0FFFC6718BF877FAD0F832" box="[151,1403,1931,1953]" httpUri="https://zenodo.org/record/2615863/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,250,1931,1953]" targetBox="[151,1436,1520,1909]" targetPageId="55">
<paragraph id="8BADBC0E1F0FFFC6718BF877FAD0F832" blockId="55.[151,1403,1931,1953]" box="[151,1403,1931,1953]" pageId="55" pageNumber="56">
<emphasis id="B966601C1F0FFFC6718BF877FE8BF832" bold="true" box="[151,288,1931,1953]" pageId="55" pageNumber="56">FIGURE 44.</emphasis>
<taxonomicName id="4C12C78D1F0FFFC67039F877FE75F832" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[293,478,1931,1953]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="erkakanae">
<emphasis id="B966601C1F0FFFC67039F877FE75F832" box="[293,478,1931,1953]" italics="true" pageId="55" pageNumber="56">Cobitis erkakanae</emphasis>
</taxonomicName>
, FSJF 2510, female, about 65 mm SL; Turkey: river connecting Lakes Gölbaşı and Azaplı.
</paragraph>
</caption>
<caption id="DF6DEC861F00FFC9718BFF64FE55FF5C" pageId="56" pageNumber="57">
<paragraph id="8BADBC0E1F00FFC9718BFF64FE55FF5C" blockId="56.[151,1436,151,207]" pageId="56" pageNumber="57">
<emphasis id="B966601C1F00FFC9718BFF64FEACFF3C" bold="true" box="[151,263,152,175]" pageId="56" pageNumber="57">TABLE 6.</emphasis>
Morphometric data of
<taxonomicName id="4C12C78D1F00FFC970E1FF6BFD6BFF3C" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[509,704,151,175]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="erkakanae">
<emphasis id="B966601C1F00FFC970E1FF6BFD6BFF3C" box="[509,704,151,175]" italics="true" pageId="56" pageNumber="57">Cobitis erkakanae</emphasis>
</taxonomicName>
(holotype ZFMK ICH-98958 and paratypes FSJF 2427; n = 13). The calculations include the holotype.
</paragraph>
</caption>
<paragraph id="8BADBC0E1F00FFC973B0FF16FA83FB1D" pageId="56" pageNumber="57">
<table id="F9124EAE1F00000E7183FF16FAE9FB1D" box="[159,1346,234,1166]" colsContinueFrom="53.[159,1344,234,1166]" colsContinueIn="63.[159,1342,1026,1958]" gridcols="6" gridrows="23" pageId="56" pageNumber="57">
<tr id="3522BE4C1F00000E7183FF16FAE9FE93" box="[159,1346,234,256]" gridrow="0" pageId="56" pageNumber="57" rowspan-0="1" rowspan-4="1" rowspan-5="1">
<th id="76F3D7301F00000E73B0FF16FCAEFE93" box="[684,773,234,256]" gridcol="1" gridrow="0" pageId="56" pageNumber="57">holotype</th>
<th id="76F3D7301F00000E7259FF16FBB4FE93" box="[837,1055,234,256]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="56" pageNumber="57">holotype &amp; paratypes</th>
</tr>
<tr id="3522BE4C1F00000E7183FEE8FAE9FEB9" box="[159,1346,276,298]" gridrow="1" pageId="56" pageNumber="57" rowspan-0="1" rowspan-3="1" rowspan-5="1">
<td id="76F3D7301F00000E73B0FEE8FCAEFEB9" box="[684,773,276,298]" gridcol="1" gridrow="1" pageId="56" pageNumber="57">male</td>
<td id="76F3D7301F00000E7259FEE8FC00FEB9" box="[837,939,276,298]" gridcol="2" gridrow="1" pageId="56" pageNumber="57">male</td>
<td id="76F3D7301F00000E7548FEE8FB11FEB9" box="[1108,1210,276,298]" gridcol="4" gridrow="1" pageId="56" pageNumber="57">female</td>
</tr>
<tr id="3522BE4C1F00000E7183FEC2FAE9FEC0" box="[159,1346,318,339]" gridrow="2" pageId="56" pageNumber="57" rowspan-0="1" rowspan-1="1">
<td id="76F3D7301F00000E7259FEC2FC00FEC0" box="[837,939,318,339]" gridcol="2" gridrow="2" pageId="56" pageNumber="57">range</td>
<td id="76F3D7301F00000E72FFFEC2FBB4FEC0" box="[995,1055,318,339]" gridcol="3" gridrow="2" pageId="56" pageNumber="57">mean</td>
<td id="76F3D7301F00000E7548FEC2FB11FEC0" box="[1108,1210,318,339]" gridcol="4" gridrow="2" pageId="56" pageNumber="57">range</td>
<td id="76F3D7301F00000E7414FEC2FAE9FEC0" box="[1288,1346,318,339]" gridcol="5" gridrow="2" pageId="56" pageNumber="57">mean</td>
</tr>
<tr id="3522BE4C1F00000E7183FE9AFAE9FEEF" box="[159,1346,358,380]" gridrow="3" pageId="56" pageNumber="57" rowspan-3="1" rowspan-5="1">
<th id="76F3D7301F00000E7183FE9AFE4AFEEF" box="[159,481,358,380]" gridcol="0" gridrow="3" pageId="56" pageNumber="57">SL (mm)</th>
<td id="76F3D7301F00000E73B0FE9AFCAEFEEF" box="[684,773,358,380]" gridcol="1" gridrow="3" pageId="56" pageNumber="57">49</td>
<td id="76F3D7301F00000E7259FE9AFC00FEEF" box="[837,939,358,380]" gridcol="2" gridrow="3" pageId="56" pageNumber="57">4849</td>
<td id="76F3D7301F00000E7548FE9AFB11FEEF" box="[1108,1210,358,380]" gridcol="4" gridrow="3" pageId="56" pageNumber="57">4465</td>
</tr>
<tr id="3522BE4C1F00000E7183FE6CFAE9FE35" box="[159,1346,400,422]" gridrow="4" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FE6CFAE9FE35" box="[159,1346,400,422]" colspan="6" colspanRight="5" gridcol="0" gridrow="4" pageId="56" pageNumber="57">In percent of standard length</th>
</tr>
<tr id="3522BE4C1F00000E7183FE45FAE9FE5C" box="[159,1346,441,463]" gridrow="5" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FE45FE4AFE5C" box="[159,481,441,463]" gridcol="0" gridrow="5" pageId="56" pageNumber="57">Head length</th>
<td id="76F3D7301F00000E73B0FE45FCAEFE5C" box="[684,773,441,463]" gridcol="1" gridrow="5" pageId="56" pageNumber="57">21.7</td>
<td id="76F3D7301F00000E7259FE45FC00FE5C" box="[837,939,441,463]" gridcol="2" gridrow="5" pageId="56" pageNumber="57">21.722.7</td>
<td id="76F3D7301F00000E72FFFE45FBB4FE5C" box="[995,1055,441,463]" gridcol="3" gridrow="5" pageId="56" pageNumber="57">22.3</td>
<td id="76F3D7301F00000E7548FE45FB11FE5C" box="[1108,1210,441,463]" gridcol="4" gridrow="5" pageId="56" pageNumber="57">20.523.1</td>
<td id="76F3D7301F00000E7414FE45FAE9FE5C" box="[1288,1346,441,463]" gridcol="5" gridrow="5" pageId="56" pageNumber="57">21.5</td>
</tr>
<tr id="3522BE4C1F00000E7183FE1EFAE9FE6B" box="[159,1346,482,504]" gridrow="6" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FE1EFE4AFE6B" box="[159,481,482,504]" gridcol="0" gridrow="6" pageId="56" pageNumber="57">Predorsal length</th>
<td id="76F3D7301F00000E73B0FE1EFCAEFE6B" box="[684,773,482,504]" gridcol="1" gridrow="6" pageId="56" pageNumber="57">54.4</td>
<td id="76F3D7301F00000E7259FE1EFC00FE6B" box="[837,939,482,504]" gridcol="2" gridrow="6" pageId="56" pageNumber="57">54.455.8</td>
<td id="76F3D7301F00000E72FFFE1EFBB4FE6B" box="[995,1055,482,504]" gridcol="3" gridrow="6" pageId="56" pageNumber="57">54.9</td>
<td id="76F3D7301F00000E7548FE1EFB11FE6B" box="[1108,1210,482,504]" gridcol="4" gridrow="6" pageId="56" pageNumber="57">52.656.0</td>
<td id="76F3D7301F00000E7414FE1EFAE9FE6B" box="[1288,1346,482,504]" gridcol="5" gridrow="6" pageId="56" pageNumber="57">54.5</td>
</tr>
<tr id="3522BE4C1F00000E7183FDF0FAE9FDB1" box="[159,1346,524,546]" gridrow="7" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FDF0FE4AFDB1" box="[159,481,524,546]" gridcol="0" gridrow="7" pageId="56" pageNumber="57">Prepelvic length</th>
<td id="76F3D7301F00000E73B0FDF0FCAEFDB1" box="[684,773,524,546]" gridcol="1" gridrow="7" pageId="56" pageNumber="57">61.2</td>
<td id="76F3D7301F00000E7259FDF0FC00FDB1" box="[837,939,524,546]" gridcol="2" gridrow="7" pageId="56" pageNumber="57">59.861.2</td>
<td id="76F3D7301F00000E72FFFDF0FBB4FDB1" box="[995,1055,524,546]" gridcol="3" gridrow="7" pageId="56" pageNumber="57">60.5</td>
<td id="76F3D7301F00000E7548FDF0FB11FDB1" box="[1108,1210,524,546]" gridcol="4" gridrow="7" pageId="56" pageNumber="57">56.961.3</td>
<td id="76F3D7301F00000E7414FDF0FAE9FDB1" box="[1288,1346,524,546]" gridcol="5" gridrow="7" pageId="56" pageNumber="57">59.3</td>
</tr>
<tr id="3522BE4C1F00000E7183FDC9FAE9FDD8" box="[159,1346,565,587]" gridrow="8" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FDC9FE4AFDD8" box="[159,481,565,587]" gridcol="0" gridrow="8" pageId="56" pageNumber="57">Preanal length</th>
<td id="76F3D7301F00000E73B0FDC9FCAEFDD8" box="[684,773,565,587]" gridcol="1" gridrow="8" pageId="56" pageNumber="57">83.3</td>
<td id="76F3D7301F00000E7259FDC9FC00FDD8" box="[837,939,565,587]" gridcol="2" gridrow="8" pageId="56" pageNumber="57">82.685.1</td>
<td id="76F3D7301F00000E72FFFDC9FBB4FDD8" box="[995,1055,565,587]" gridcol="3" gridrow="8" pageId="56" pageNumber="57">83.9</td>
<td id="76F3D7301F00000E7548FDC9FB11FDD8" box="[1108,1210,565,587]" gridcol="4" gridrow="8" pageId="56" pageNumber="57">81.985.0</td>
<td id="76F3D7301F00000E7414FDC9FAE9FDD8" box="[1288,1346,565,587]" gridcol="5" gridrow="8" pageId="56" pageNumber="57">83.2</td>
</tr>
<tr id="3522BE4C1F00000E7183FDA2FAE9FDE7" box="[159,1346,606,628]" gridrow="9" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FDA2FE4AFDE7" box="[159,481,606,628]" gridcol="0" gridrow="9" pageId="56" pageNumber="57">Body depth at dorsal-fin origin</th>
<td id="76F3D7301F00000E73B0FDA2FCAEFDE7" box="[684,773,606,628]" gridcol="1" gridrow="9" pageId="56" pageNumber="57">18.3</td>
<td id="76F3D7301F00000E7259FDA2FC00FDE7" box="[837,939,606,628]" gridcol="2" gridrow="9" pageId="56" pageNumber="57">15.318.3</td>
<td id="76F3D7301F00000E72FFFDA2FBB4FDE7" box="[995,1055,606,628]" gridcol="3" gridrow="9" pageId="56" pageNumber="57">16.9</td>
<td id="76F3D7301F00000E7548FDA2FB11FDE7" box="[1108,1210,606,628]" gridcol="4" gridrow="9" pageId="56" pageNumber="57">14.117.4</td>
<td id="76F3D7301F00000E7414FDA2FAE9FDE7" box="[1288,1346,606,628]" gridcol="5" gridrow="9" pageId="56" pageNumber="57">16.0</td>
</tr>
<tr id="3522BE4C1F00000E7183FD74FAE9FD0D" box="[159,1346,648,670]" gridrow="10" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FD74FE4AFD0D" box="[159,481,648,670]" gridcol="0" gridrow="10" pageId="56" pageNumber="57">Body width at dorsal-fin origin</th>
<td id="76F3D7301F00000E73B0FD74FCAEFD0D" box="[684,773,648,670]" gridcol="1" gridrow="10" pageId="56" pageNumber="57">8.6</td>
<td id="76F3D7301F00000E7259FD74FC00FD0D" box="[837,939,648,670]" gridcol="2" gridrow="10" pageId="56" pageNumber="57">8.39.0</td>
<td id="76F3D7301F00000E72FFFD74FBB4FD0D" box="[995,1055,648,670]" gridcol="3" gridrow="10" pageId="56" pageNumber="57">8.7</td>
<td id="76F3D7301F00000E7548FD74FB11FD0D" box="[1108,1210,648,670]" gridcol="4" gridrow="10" pageId="56" pageNumber="57">7.99.9</td>
<td id="76F3D7301F00000E7414FD74FAE9FD0D" box="[1288,1346,648,670]" gridcol="5" gridrow="10" pageId="56" pageNumber="57">8.7</td>
</tr>
<tr id="3522BE4C1F00000E7183FD4DFAE9FD54" box="[159,1346,689,711]" gridrow="11" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FD4DFE4AFD54" box="[159,481,689,711]" gridcol="0" gridrow="11" pageId="56" pageNumber="57">Depth of caudal peduncle</th>
<td id="76F3D7301F00000E73B0FD4DFCAEFD54" box="[684,773,689,711]" gridcol="1" gridrow="11" pageId="56" pageNumber="57">10.4</td>
<td id="76F3D7301F00000E7259FD4DFC00FD54" box="[837,939,689,711]" gridcol="2" gridrow="11" pageId="56" pageNumber="57">10.311.3</td>
<td id="76F3D7301F00000E72FFFD4DFBB4FD54" box="[995,1055,689,711]" gridcol="3" gridrow="11" pageId="56" pageNumber="57">10.7</td>
<td id="76F3D7301F00000E7548FD4DFB11FD54" box="[1108,1210,689,711]" gridcol="4" gridrow="11" pageId="56" pageNumber="57">8.610.9</td>
<td id="76F3D7301F00000E7414FD4DFAE9FD54" box="[1288,1346,689,711]" gridcol="5" gridrow="11" pageId="56" pageNumber="57">10.1</td>
</tr>
<tr id="3522BE4C1F00000E7183FD26FAE9FD63" box="[159,1346,730,752]" gridrow="12" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FD26FE4AFD63" box="[159,481,730,752]" gridcol="0" gridrow="12" pageId="56" pageNumber="57">Length of caudal peduncle</th>
<td id="76F3D7301F00000E73B0FD26FCAEFD63" box="[684,773,730,752]" gridcol="1" gridrow="12" pageId="56" pageNumber="57">11.7</td>
<td id="76F3D7301F00000E7259FD26FC00FD63" box="[837,939,730,752]" gridcol="2" gridrow="12" pageId="56" pageNumber="57">10.813.1</td>
<td id="76F3D7301F00000E72FFFD26FBB4FD63" box="[995,1055,730,752]" gridcol="3" gridrow="12" pageId="56" pageNumber="57">11.9</td>
<td id="76F3D7301F00000E7548FD26FB11FD63" box="[1108,1210,730,752]" gridcol="4" gridrow="12" pageId="56" pageNumber="57">11.514.7</td>
<td id="76F3D7301F00000E7414FD26FAE9FD63" box="[1288,1346,730,752]" gridcol="5" gridrow="12" pageId="56" pageNumber="57">12.4</td>
</tr>
<tr id="3522BE4C1F00000E7183FCF8FAE9FC89" box="[159,1346,772,794]" gridrow="13" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FCF8FE4AFC89" box="[159,481,772,794]" gridcol="0" gridrow="13" pageId="56" pageNumber="57">Dorsal-fin depth</th>
<td id="76F3D7301F00000E73B0FCF8FCAEFC89" box="[684,773,772,794]" gridcol="1" gridrow="13" pageId="56" pageNumber="57">18.1</td>
<td id="76F3D7301F00000E7259FCF8FC00FC89" box="[837,939,772,794]" gridcol="2" gridrow="13" pageId="56" pageNumber="57">16.218.6</td>
<td id="76F3D7301F00000E72FFFCF8FBB4FC89" box="[995,1055,772,794]" gridcol="3" gridrow="13" pageId="56" pageNumber="57">17.8</td>
<td id="76F3D7301F00000E7548FCF8FB11FC89" box="[1108,1210,772,794]" gridcol="4" gridrow="13" pageId="56" pageNumber="57">16.518.9</td>
<td id="76F3D7301F00000E7414FCF8FAE9FC89" box="[1288,1346,772,794]" gridcol="5" gridrow="13" pageId="56" pageNumber="57">17.4</td>
</tr>
<tr id="3522BE4C1F00000E7183FCD1FAE9FCD0" box="[159,1346,813,835]" gridrow="14" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FCD1FE4AFCD0" box="[159,481,813,835]" gridcol="0" gridrow="14" pageId="56" pageNumber="57">Caudal-fin length</th>
<td id="76F3D7301F00000E73B0FCD1FCAEFCD0" box="[684,773,813,835]" gridcol="1" gridrow="14" pageId="56" pageNumber="57">17.1</td>
<td id="76F3D7301F00000E7259FCD1FC00FCD0" box="[837,939,813,835]" gridcol="2" gridrow="14" pageId="56" pageNumber="57">16.419.2</td>
<td id="76F3D7301F00000E72FFFCD1FBB4FCD0" box="[995,1055,813,835]" gridcol="3" gridrow="14" pageId="56" pageNumber="57">17.9</td>
<td id="76F3D7301F00000E7548FCD1FB11FCD0" box="[1108,1210,813,835]" gridcol="4" gridrow="14" pageId="56" pageNumber="57">17.421.1</td>
<td id="76F3D7301F00000E7414FCD1FAE9FCD0" box="[1288,1346,813,835]" gridcol="5" gridrow="14" pageId="56" pageNumber="57">18.9</td>
</tr>
<tr id="3522BE4C1F00000E7183FCAAFAE9FCFF" box="[159,1346,854,876]" gridrow="15" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FCAAFE4AFCFF" box="[159,481,854,876]" gridcol="0" gridrow="15" pageId="56" pageNumber="57">Pectoral-fin length</th>
<td id="76F3D7301F00000E73B0FCAAFCAEFCFF" box="[684,773,854,876]" gridcol="1" gridrow="15" pageId="56" pageNumber="57">19.6</td>
<td id="76F3D7301F00000E7259FCAAFC00FCFF" box="[837,939,854,876]" gridcol="2" gridrow="15" pageId="56" pageNumber="57">17.120.0</td>
<td id="76F3D7301F00000E72FFFCAAFBB4FCFF" box="[995,1055,854,876]" gridcol="3" gridrow="15" pageId="56" pageNumber="57">18.7</td>
<td id="76F3D7301F00000E7548FCAAFB11FCFF" box="[1108,1210,854,876]" gridcol="4" gridrow="15" pageId="56" pageNumber="57">13.116.7</td>
<td id="76F3D7301F00000E7414FCAAFAE9FCFF" box="[1288,1346,854,876]" gridcol="5" gridrow="15" pageId="56" pageNumber="57">15.3</td>
</tr>
<tr id="3522BE4C1F00000E7183FC7CFAE9FC05" box="[159,1346,896,918]" gridrow="16" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FC7CFE4AFC05" box="[159,481,896,918]" gridcol="0" gridrow="16" pageId="56" pageNumber="57">Pelvic-fin length</th>
<td id="76F3D7301F00000E73B0FC7CFCAEFC05" box="[684,773,896,918]" gridcol="1" gridrow="16" pageId="56" pageNumber="57">14.4</td>
<td id="76F3D7301F00000E7259FC7CFC00FC05" box="[837,939,896,918]" gridcol="2" gridrow="16" pageId="56" pageNumber="57">13.114.8</td>
<td id="76F3D7301F00000E72FFFC7CFBB4FC05" box="[995,1055,896,918]" gridcol="3" gridrow="16" pageId="56" pageNumber="57">14.1</td>
<td id="76F3D7301F00000E7548FC7CFB11FC05" box="[1108,1210,896,918]" gridcol="4" gridrow="16" pageId="56" pageNumber="57">11.713.7</td>
<td id="76F3D7301F00000E7414FC7CFAE9FC05" box="[1288,1346,896,918]" gridcol="5" gridrow="16" pageId="56" pageNumber="57">12.5</td>
</tr>
<tr id="3522BE4C1F00000E7183FC55FAE9FC2C" box="[159,1346,937,959]" gridrow="17" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FC55FAE9FC2C" box="[159,1346,937,959]" colspan="6" colspanRight="5" gridcol="0" gridrow="17" pageId="56" pageNumber="57">In percent of head length</th>
</tr>
<tr id="3522BE4C1F00000E7183FC2EFAE9FC7B" box="[159,1346,978,1000]" gridrow="18" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FC2EFE4AFC7B" box="[159,481,978,1000]" gridcol="0" gridrow="18" pageId="56" pageNumber="57">Snout length</th>
<td id="76F3D7301F00000E73B0FC2EFCAEFC7B" box="[684,773,978,1000]" gridcol="1" gridrow="18" pageId="56" pageNumber="57">36</td>
<td id="76F3D7301F00000E7259FC2EFC00FC7B" box="[837,939,978,1000]" gridcol="2" gridrow="18" pageId="56" pageNumber="57">3639</td>
<td id="76F3D7301F00000E72FFFC2EFBB4FC7B" box="[995,1055,978,1000]" gridcol="3" gridrow="18" pageId="56" pageNumber="57">38.6</td>
<td id="76F3D7301F00000E7548FC2EFB11FC7B" box="[1108,1210,978,1000]" gridcol="4" gridrow="18" pageId="56" pageNumber="57">3744</td>
<td id="76F3D7301F00000E7414FC2EFAE9FC7B" box="[1288,1346,978,1000]" gridcol="5" gridrow="18" pageId="56" pageNumber="57">40.5</td>
</tr>
<tr id="3522BE4C1F00000E7183FC00FAE9FB81" box="[159,1346,1020,1042]" gridrow="19" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FC00FE4AFB81" box="[159,481,1020,1042]" gridcol="0" gridrow="19" pageId="56" pageNumber="57">Postorbital distance</th>
<td id="76F3D7301F00000E73B0FC00FCAEFB81" box="[684,773,1020,1042]" gridcol="1" gridrow="19" pageId="56" pageNumber="57">48</td>
<td id="76F3D7301F00000E7259FC00FC00FB81" box="[837,939,1020,1042]" gridcol="2" gridrow="19" pageId="56" pageNumber="57">4855</td>
<td id="76F3D7301F00000E72FFFC00FBB4FB81" box="[995,1055,1020,1042]" gridcol="3" gridrow="19" pageId="56" pageNumber="57">52.6</td>
<td id="76F3D7301F00000E7548FC00FB11FB81" box="[1108,1210,1020,1042]" gridcol="4" gridrow="19" pageId="56" pageNumber="57">4956</td>
<td id="76F3D7301F00000E7414FC00FAE9FB81" box="[1288,1346,1020,1042]" gridcol="5" gridrow="19" pageId="56" pageNumber="57">52.4</td>
</tr>
<tr id="3522BE4C1F00000E7183FBD9FAE9FBA8" box="[159,1346,1061,1083]" gridrow="20" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FBD9FE4AFBA8" box="[159,481,1061,1083]" gridcol="0" gridrow="20" pageId="56" pageNumber="57">Head depth at eye</th>
<td id="76F3D7301F00000E73B0FBD9FCAEFBA8" box="[684,773,1061,1083]" gridcol="1" gridrow="20" pageId="56" pageNumber="57">52</td>
<td id="76F3D7301F00000E7259FBD9FC00FBA8" box="[837,939,1061,1083]" gridcol="2" gridrow="20" pageId="56" pageNumber="57">4752</td>
<td id="76F3D7301F00000E72FFFBD9FBB4FBA8" box="[995,1055,1061,1083]" gridcol="3" gridrow="20" pageId="56" pageNumber="57">49.9</td>
<td id="76F3D7301F00000E7548FBD9FB11FBA8" box="[1108,1210,1061,1083]" gridcol="4" gridrow="20" pageId="56" pageNumber="57">4556</td>
<td id="76F3D7301F00000E7414FBD9FAE9FBA8" box="[1288,1346,1061,1083]" gridcol="5" gridrow="20" pageId="56" pageNumber="57">51.0</td>
</tr>
<tr id="3522BE4C1F00000E7183FBB2FAE9FBF7" box="[159,1346,1102,1124]" gridrow="21" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FBB2FE4AFBF7" box="[159,481,1102,1124]" gridcol="0" gridrow="21" pageId="56" pageNumber="57">Eye diameter</th>
<td id="76F3D7301F00000E73B0FBB2FCAEFBF7" box="[684,773,1102,1124]" gridcol="1" gridrow="21" pageId="56" pageNumber="57">18</td>
<td id="76F3D7301F00000E7259FBB2FC00FBF7" box="[837,939,1102,1124]" gridcol="2" gridrow="21" pageId="56" pageNumber="57">1518</td>
<td id="76F3D7301F00000E72FFFBB2FBB4FBF7" box="[995,1055,1102,1124]" gridcol="3" gridrow="21" pageId="56" pageNumber="57">16.2</td>
<td id="76F3D7301F00000E7548FBB2FB11FBF7" box="[1108,1210,1102,1124]" gridcol="4" gridrow="21" pageId="56" pageNumber="57">1317</td>
<td id="76F3D7301F00000E7414FBB2FAE9FBF7" box="[1288,1346,1102,1124]" gridcol="5" gridrow="21" pageId="56" pageNumber="57">14.9</td>
</tr>
<tr id="3522BE4C1F00000E7183FB84FAE9FB1D" box="[159,1346,1144,1166]" gridrow="22" pageId="56" pageNumber="57">
<th id="76F3D7301F00000E7183FB84FE4AFB1D" box="[159,481,1144,1166]" gridcol="0" gridrow="22" pageId="56" pageNumber="57">Interorbital width</th>
<td id="76F3D7301F00000E73B0FB84FCAEFB1D" box="[684,773,1144,1166]" gridcol="1" gridrow="22" pageId="56" pageNumber="57">10</td>
<td id="76F3D7301F00000E7259FB84FC00FB1D" box="[837,939,1144,1166]" gridcol="2" gridrow="22" pageId="56" pageNumber="57">811</td>
<td id="76F3D7301F00000E72FFFB84FBB4FB1D" box="[995,1055,1144,1166]" gridcol="3" gridrow="22" pageId="56" pageNumber="57">9.6</td>
<td id="76F3D7301F00000E7548FB84FB11FB1D" box="[1108,1210,1144,1166]" gridcol="4" gridrow="22" pageId="56" pageNumber="57">710</td>
<td id="76F3D7301F00000E7414FB84FAE9FB1D" box="[1288,1346,1144,1166]" gridcol="5" gridrow="22" pageId="56" pageNumber="57">8.6</td>
</tr>
</table>
</paragraph>
<paragraph id="8BADBC0E1F00FFC971DBFB31FAA1FA71" blockId="56.[151,1437,1229,2010]" pageId="56" pageNumber="57">
Dorsal fin with 3 unbranched and 6½ (15) branched rays. Anal fin with 3 unbranched and 5½ (14) and 6½ (1) branched rays. Caudal fin with 6+6 branched rays in the one male, 7+
<quantity id="4CEA11EB1F00FFC972DCFB0EFC46FA99" box="[960,1005,1265,1290]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="56" pageNumber="57" unit="in" value="7.0">7 in</quantity>
other males, 8+
<quantity id="4CEA11EB1F00FFC975B4FB0EFB7DFA99" box="[1192,1238,1265,1290]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="56" pageNumber="57" unit="in" value="7.0">7 in</quantity>
the two females. Pectoral fin with 7 (1), 8 (10), 9 (3) and 10 (1) branched rays and pelvic fin with 4 (1) and 5 (14) branched rays. Body completely covered by embedded scales, except on belly and breast. Scales small. Focal zone in subdorsal scales about less than 1/3 or 1/5 of vertical scale diameter. Lateral line short, with 59 pores or absent. Lips (
<figureCitation id="1329A08B1F00FFC97473FAA1FF1FFA09" captionStart="FIGURE 54" captionStartId="67.[151,250,1575,1597]" captionTargetBox="[151,1436,220,1518]" captionTargetId="figure@67.[151,1436,220,1518]" captionTargetPageId="67" captionText="FIGURE 54. Upper row: Cobitis afifeae, FSJF 3022, paratype, 76 mm SL; Turkey: Büyük Menderes River; C. aliyeae, FSJF 2427, paratype, female, 65 mm SL; Turkey: Seyhan River; C. anabelae, ZFMK ICH-98633, holotype, male, 53 mm SL; Turkey: Karasu River; C. emrei, FSJF 1960, paratype, female, 77 mm SL; Turkey: Lake Sapanca basin; Lower row: C. erkakanae, FSJF 2510, paratype, female, 68 mm SL; Turkey: canal at Lake Gölbaşı; C. joergbohleni, ZFMK ICH-97624, holotype, female, 60 mm SL; Turkey: spring Soysallı; C. pirii, FSJF 2474, paratype, female, 95 mm SL; Turkey: streams Çayköy; C. troasensis, FSJF 2383, paratype, female, 72 mm SL; Turkey: stream Tuzla." httpUri="https://zenodo.org/record/2615885/files/figure.png" pageId="56" pageNumber="57">Fig. 54</figureCitation>
) thin and mental lobes of lower lip long, usually well separated from lower lip, produced into a short barbel-like process in few individuals. Rostral barbel reaching base of mandibular barbel. Mandibular barbel reaching to or slightly beyond vertical of nostril. Maxillary barbel reaching vertical of front border or middle of eye.
</paragraph>
<paragraph id="8BADBC0E1F00FFC971DBFA11FE24F9B9" blockId="56.[151,1437,1229,2010]" pageId="56" pageNumber="57">
<emphasis id="B966601C1F00FFC971DBFA11FE12F995" bold="true" box="[199,441,1517,1542]" pageId="56" pageNumber="57">Sexual dimorphism.</emphasis>
Male have a longer pectoral fin than female (1720% SL vs. 1317) and two laminae circularis (vs. absent).
</paragraph>
<paragraph id="8BADBC0E1F00FFC971DBF9C9FA22F801" blockId="56.[151,1437,1229,2010]" pageId="56" pageNumber="57">
<emphasis id="B966601C1F00FFC971DBF9C9FEF4F9DD" bold="true" box="[199,351,1589,1614]" pageId="56" pageNumber="57">Colouration.</emphasis>
Background colour yellowish with a dark-brown pigmentation pattern organised in one middorsal and four lateral zones. Mid-dorsal pigmentation consisting in a series of 912, elongate blotches, often very closely set or slightly fused. Pigmentation in Z1 well developed, with many very small to medium sized spots narrower than Z2, not reaching dorsally to interspaces of mid-dorsal blotches, fused with Z2 on postdorsal flank. Spots in Z2 about eye size or larger, often fused to adjacent spots, fused to short stripes in two males. Spots in Z2 usually well distinguished until caudal peduncle. Pigmentation in Z4 formed by 36 predorsal, 12 subdorsal and 59 postdorsal blotches, blotches elongate, anterior most and posterior most blotches often roundish. A single, usually indistinct black spot at upper caudal-fin base, less than eye diameter. Upper part of head, opercle and snout covered by large spots. A dark-brown stripe between eye and snout. Fins hyaline. Caudal fin with 47 and dorsal fin with 46 dark-brown, sometimes irregular shaped bars. Few dark-brown spots in paired fins. Barbels whitish.
</paragraph>
</subSubSection>
<subSubSection id="C308EF851F00FFC971DBF861FD46F849" pageId="56" pageNumber="57" type="etymology">
<paragraph id="8BADBC0E1F00FFC971DBF861FD46F849" blockId="56.[151,1437,1229,2010]" pageId="56" pageNumber="57">
<emphasis id="B966601C1F00FFC971DBF861FEE5F825" bold="true" box="[199,334,1949,1974]" pageId="56" pageNumber="57">Etymology.</emphasis>
Named for Füsun Erkakan (
<collectingRegion id="49D672EC1F00FFC973A9F861FCA0F825" box="[693,779,1949,1974]" country="Turkey" name="Ankara" pageId="56" pageNumber="57">Ankara</collectingRegion>
) for her contribution to the exploration of the species diversity of
<taxonomicName id="4C12C78D1F00FFC97002F83DFEC4F84A" box="[286,367,1985,2009]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="56" pageNumber="57" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F00FFC97002F83DFEC4F84A" box="[286,367,1985,2009]" italics="true" pageId="56" pageNumber="57">Cobitis</emphasis>
</taxonomicName>
. A noun in genitive, indeclinable.
</paragraph>
</subSubSection>
<subSubSection id="C308EF851F01FFC871DBFF6BFB73FF23" box="[199,1240,151,176]" pageId="57" pageNumber="58" type="distribution">
<paragraph id="8BADBC0E1F01FFC871DBFF6BFB73FF23" blockId="57.[151,1437,151,536]" box="[199,1240,151,176]" pageId="57" pageNumber="58">
<emphasis id="B966601C1F01FFC871DBFF6BFECBFF23" bold="true" box="[199,352,151,176]" pageId="57" pageNumber="58">Distribution.</emphasis>
<taxonomicName id="4C12C78D1F01FFC8707BFF64FD9FFF23" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[359,564,152,176]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="erkakanae">
<emphasis id="B966601C1F01FFC8707BFF64FD9FFF23" box="[359,564,152,176]" italics="true" pageId="57" pageNumber="58">Cobitis erkakanae</emphasis>
</taxonomicName>
has only been found at one place in the Gölbaşı lakes basin.
</paragraph>
</subSubSection>
<subSubSection id="C308EF851F01FFC871DBFF40FAC2FD8B" pageId="57" pageNumber="58" type="discussion">
<paragraph id="8BADBC0E1F01FFC871DBFF40FDE6FE3E" blockId="57.[151,1437,151,536]" pageId="57" pageNumber="58">
<emphasis id="B966601C1F01FFC871DBFF40FE90FF46" bold="true" box="[199,315,188,213]" pageId="57" pageNumber="58">Remarks.</emphasis>
The finding of a third species of
<taxonomicName id="4C12C78D1F01FFC873A7FF40FCA7FF47" box="[699,780,188,212]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="57" pageNumber="58" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F01FFC873A7FF40FCA7FF47" box="[699,780,188,212]" italics="true" pageId="57" pageNumber="58">Cobitis</emphasis>
</taxonomicName>
in the Ceyhan River drainage is an unusual situation. All three species occur in allopatry within the Ceyhan River drainage but we are not aware of a biogeographic background explaining, why the Gölbaşı lakes have an own fish fauna, isolated from the Ceyhan River. Not only
<taxonomicName id="4C12C78D1F01FFC8749EFEF8FEA7FED3" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="erkakanae">
<emphasis id="B966601C1F01FFC8749EFEF8FEA7FED3" italics="true" pageId="57" pageNumber="58">C. erkakanae</emphasis>
</taxonomicName>
is (likely to be) endemic to the Gölbaşı lakes,
<bibRefCitation id="EF83C1FF1F01FFC87202FEDBFC55FED3" author="Geiger, M. F. &amp; Herder, F. &amp; Monaghan, M. T. &amp; Almada, V. &amp; Barbieri, R. &amp; Bariche, M. &amp; Berrebi, P. &amp; Bohlen, J. &amp; Casal-Lopez, M. &amp; Delmastro, G. B. &amp; Denys, G. P. J. &amp; Dettai, A. &amp; Doadrio, I. &amp; Kalogianni, E. &amp; Karst, H. &amp; Kottelat, M. &amp; Kovacic, M. &amp; Laporte, M. &amp; Lorenzoni, M. &amp; Marcic, Z. &amp; Ozulug, M. &amp; Perdices, A. &amp; Perea, S. &amp; Persat, H. &amp; Porcelotti, S. &amp; Puzzi, C. &amp; Robalo, J. &amp; Sanda, R. &amp; Schneider, M. &amp; Slechtova, V. &amp; Stoumboudi, M. &amp; Walter, S. &amp; Freyhof, J." box="[798,1022,295,320]" pageId="57" pageNumber="58" pagination="1210 - 1221" refId="ref47247" refString="Geiger, M. F., Herder, F., Monaghan, M. T., Almada, V., Barbieri, R., Bariche, M., Berrebi, P., Bohlen, J., Casal-Lopez, M., Delmastro, G. B., Denys, G. P. J., Dettai, A., Doadrio, I., Kalogianni, E., Karst, H., Kottelat, M., Kovacic, M., Laporte, M., Lorenzoni, M., Marcic, Z., Ozulug, M., Perdices, A., Perea, S., Persat, H., Porcelotti, S., Puzzi, C., Robalo, J., Sanda, R., Schneider, M., Slechtova, V., Stoumboudi, M., Walter, S. &amp; Freyhof, J. (2014) Spatial heterogeneity in the Mediterranean Biodiversity Hotspot affects barcoding accuracy of its freshwater fishes. Molecular Ecology Resources, 14, 1210 - 1221. https: // doi. org / 10.1111 / 1755 - 0998.12257" type="journal article" year="2014">
Geiger
<emphasis id="B966601C1F01FFC8726EFED5FC08FED3" box="[882,931,296,320]" italics="true" pageId="57" pageNumber="58">et al</emphasis>
. (2014)
</bibRefCitation>
also show molecular data indication that the populations of
<taxonomicName id="4C12C78D1F01FFC87084FEB0FE50FEF7" baseAuthorityName="Heckel" baseAuthorityYear="1843" box="[408,507,332,356]" class="Actinopterygii" family="Cyprinodontidae" genus="Aphanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="mento">
<emphasis id="B966601C1F01FFC87084FEB0FE50FEF7" box="[408,507,332,356]" italics="true" pageId="57" pageNumber="58">A. mento</emphasis>
</taxonomicName>
and
<taxonomicName id="4C12C78D1F01FFC8732DFEB0FC9EFEF7" box="[561,821,332,356]" class="Actinopterygii" family="Cyprinidae" genus="Pseudophoxinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="zekayi">
<emphasis id="B966601C1F01FFC8732DFEB0FC9EFEF7" box="[561,821,332,356]" italics="true" pageId="57" pageNumber="58">Pseudophoxinus zekayi</emphasis>
</taxonomicName>
are isolated from those in the Ceyhan. While the water from the Gölbşı lakes flows to the Ceyhan today, we postulate, that this was an endorheic basin for a while and was connected only recently to the Ceyhan.
</paragraph>
<paragraph id="8BADBC0E1F01FFC871DBFE4BFAC2FD8B" blockId="57.[151,1437,151,536]" pageId="57" pageNumber="58">
Molecular data (
<figureCitation id="1329A08B1F01FFC8709CFE4BFE6AFE43" box="[384,449,439,464]" captionStart="FIGURE 1" captionStartId="6.[151,249,1665,1687]" captionTargetBox="[164,1417,206,1642]" captionTargetId="figure@6.[155,1431,190,1647]" captionTargetPageId="6" captionText="FIGURE 1. Maximum likelihood estimation of the phylogenetic relationships based on the mitochondrial COI barcode region (TN93 model, discrete Gamma distribution for rate differences with 3 categories +G parameter = 0.3792. Nucleotide positions with less than 95% site coverage were eliminated, resulting in 590 analysed positions. Numbers of major nodes indicate bootstrap values from 1000 pseudoreplicates from the ML, NJ and MP method. A star denotes highest possible support from all three methods. Solid bars right to the specimen labels indicate species delimitation results from PTP followed by the results of the mPTP approach as dashed bars. Except to the outgroups and the branch to the right part of the figure, the tree is drawn to scale with branch lengths depicting number of substitutions per site." httpUri="https://zenodo.org/record/2615777/files/figure.png" pageId="57" pageNumber="58">Fig. 1</figureCitation>
) place
<taxonomicName id="4C12C78D1F01FFC87309FE44FD07FE43" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[533,684,440,464]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="erkakanae">
<emphasis id="B966601C1F01FFC87309FE44FD07FE43" box="[533,684,440,464]" italics="true" pageId="57" pageNumber="58">C. erkakanae</emphasis>
</taxonomicName>
in the
<taxonomicName id="4C12C78D1F01FFC873E5FE44FCF9FE43" baseAuthorityName="Heckel" baseAuthorityYear="1848" box="[761,850,440,464]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="linea">
<emphasis id="B966601C1F01FFC873E5FE44FCF9FE43" box="[761,850,440,464]" italics="true" pageId="57" pageNumber="58">C. linea</emphasis>
</taxonomicName>
species group (
<taxonomicName id="4C12C78D1F01FFC87518FE44FBF6FE43" baseAuthorityName="Heckel" baseAuthorityYear="1848" box="[1028,1117,440,464]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="linea">
<emphasis id="B966601C1F01FFC87518FE44FBF6FE43" box="[1028,1117,440,464]" italics="true" pageId="57" pageNumber="58">C. linea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C12C78D1F01FFC87570FE44FB75FE43" authorityName="Freyhof &amp; Bayçelebi &amp; Geiger" authorityYear="2018" box="[1132,1246,440,464]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="aliyeae">
<emphasis id="B966601C1F01FFC87570FE44FB75FE43" box="[1132,1246,440,464]" italics="true" pageId="57" pageNumber="58">C. aliyeae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C12C78D1F01FFC875F0FE44FADFFE43" box="[1260,1396,440,464]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="anabelae">
<emphasis id="B966601C1F01FFC875F0FE44FADFFE43" box="[1260,1396,440,464]" italics="true" pageId="57" pageNumber="58">C. anabelae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C12C78D1F01FFC8749EFE44FEB9FE67" authorityName="Coad &amp; Sarieyyupoglu" authorityYear="1988" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="elazigensis">
<emphasis id="B966601C1F01FFC8749EFE44FEB9FE67" italics="true" pageId="57" pageNumber="58">C. elazigensis</emphasis>
</taxonomicName>
,
<emphasis id="B966601C1F01FFC87003FE20FE73FE67" box="[287,472,476,500]" italics="true" pageId="57" pageNumber="58">
and
<taxonomicName id="4C12C78D1F01FFC8704CFE20FE73FE67" authorityName="Krupp &amp; Moubayed" authorityYear="1992" box="[336,472,476,500]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="levantina">C. levantina</taxonomicName>
</emphasis>
). Based on DNA barcoding it is well separated from all other included
<taxonomicName id="4C12C78D1F01FFC875EFFE20FAEFFE67" box="[1267,1348,476,500]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="57" pageNumber="58" phylum="Chordata" rank="genus">
<emphasis id="B966601C1F01FFC875EFFE20FAEFFE67" box="[1267,1348,476,500]" italics="true" pageId="57" pageNumber="58">Cobitis</emphasis>
</taxonomicName>
, and by a minimum K2P distance of 5.5% to
<taxonomicName id="4C12C78D1F01FFC8732AFDFCFD79FD8B" authorityName="Coad &amp; Sarieyyupoglu" authorityYear="1988" box="[566,722,512,536]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="elazigensis">
<emphasis id="B966601C1F01FFC8732AFDFCFD79FD8B" box="[566,722,512,536]" italics="true" pageId="57" pageNumber="58">C. elazigensis</emphasis>
</taxonomicName>
. It is also supported by the PTP approach as distinct entity.
</paragraph>
</subSubSection>
</treatment>
</document>