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<document ID-DOI="10.11646/zootaxa.5264.4.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7837222" ID-ZooBank="A65E722C-C9D8-43DA-9668-D7338CAA55CA" approvalRequired="9" approvalRequired_for_taxonomicNames="2" approvalRequired_for_textStreams="7" checkinTime="1681730831263" checkinUser="plazi" docAuthor="Ranjan, Rahul, Singh, Navneet &amp; Kirti, Jagbir Singh" docDate="2023" docId="03BAA304FFD0FFCDFF0B483CFBDB1BAB" docLanguage="en" docName="zootaxa.5264.4.9.pdf" docOrigin="Zootaxa 5264 (4)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coenodomus wangi Ranjan, Singh &amp; Kirti 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="594" masterDocId="FF83DB7CFFD1FFCCFF9C4B0FFFCD1C66" masterDocTitle="A new species of Coenodomus Walsingham, 1888 (Epipaschiinae, Pyralidae, Lepidoptera) from India" masterLastPageNumber="598" masterPageNumber="593" pageNumber="594" updateTime="1681731933916" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Coenodomus Walsingham, 1888 (Epipaschiinae, Pyralidae, Lepidoptera) from India</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ranjan, Rahul</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-4983-0403</mods:nameIdentifier>
<mods:affiliation>Zoological Survey of India, M-Block New, Alipore, Kolkata 700 053, West Bengal, India. &amp; Department of Zoology &amp; Environmental Sciences, Punjabi University, Patiala 147 002, Punjab, India. &amp; https: // orcid. org / 0000 - 0003 - 4983 - 0403 &amp; https: // orcid. org / 0000 - 0002 - 6657 - 7983</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Singh, Navneet</mods:namePart>
<mods:affiliation>Zoological Survey of India, M-Block New, Alipore, Kolkata 700 053, West Bengal, India.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kirti, Jagbir Singh</mods:namePart>
<mods:affiliation>Department of Zoology &amp; Environmental Sciences, Punjabi University, Patiala 147 002, Punjab, India. &amp; https: // orcid. org / 0000 - 0002 - 9670 - 5931</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-17</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5264</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>593</mods:start>
<mods:end>598</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5264.4.9</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7837222</mods:identifier>
<mods:identifier type="ZooBank">A65E722C-C9D8-43DA-9668-D7338CAA55CA</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7837184" ID-Zenodo-Dep="7837184" LSID="urn:lsid:plazi:treatment:03BAA304FFD0FFCDFF0B483CFBDB1BAB" httpUri="http://treatment.plazi.org/id/03BAA304FFD0FFCDFF0B483CFBDB1BAB" lastPageNumber="594" pageId="1" pageNumber="594">
<subSubSection box="[151,786,819,846]" pageId="1" pageNumber="594" type="nomenclature">
<paragraph blockId="1.[151,786,819,881]" box="[151,786,819,846]" pageId="1" pageNumber="594">
<heading bold="true" box="[151,786,819,846]" fontSize="11" level="1" pageId="1" pageNumber="594" reason="1">
<emphasis bold="true" box="[151,786,819,846]" pageId="1" pageNumber="594">
<taxonomicName authority="Ranjan, Singh &amp; Kirti" authorityName="Ranjan, Singh &amp; Kirti" authorityYear="2023" box="[151,678,819,846]" class="Insecta" family="Pyralidae" genus="Coenodomus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="wangi" status="sp. nov.">
<emphasis bold="true" box="[151,390,819,846]" italics="true" pageId="1" pageNumber="594">Coenodomus wangi</emphasis>
Ranjan, Singh &amp; Kirti
</taxonomicName>
,
<taxonomicNameLabel box="[692,786,819,845]" pageId="1" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,267,855,881]" pageId="1" pageNumber="594" type="description">
<paragraph blockId="1.[151,786,819,881]" box="[151,267,855,881]" pageId="1" pageNumber="594">
(
<figureCitation box="[159,259,855,881]" captionStart="FIGURES 15" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[175,1411,200,1919]" captionTargetId="figure-29@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 15. Imago and male genitalia of Coenodomus spp.: 1, 4, C. wangi sp. nov., male; 2, 5, C. dudgeoni, male (photo credit: Wang et al. 2017); 3, C. dudgeoni, male (© NMHUK)." figureDoi="http://doi.org/10.5281/zenodo.7837224" httpUri="https://zenodo.org/record/7837224/files/figure.png" pageId="1" pageNumber="594">Figs 1, 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="594" type="materials_examined">
<paragraph blockId="1.[151,1437,927,1998]" box="[151,639,927,954]" pageId="1" pageNumber="594">
<materialsCitation box="[151,639,927,954]" country="India" location="Umtasor" pageId="1" pageNumber="594" specimenCount="1" stateProvince="Meghalaya" typeStatus="holotype">
<emphasis bold="true" box="[151,313,927,953]" pageId="1" pageNumber="594">
<typeStatus box="[151,210,927,953]" pageId="1" pageNumber="594">Type</typeStatus>
locality:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:03BAA304FFD0FFCDFF0B483CFBDB1BAB:8ECC44C9FFD0FFCDFEDC48AFFE6D1FDC" box="[320,416,928,954]" country="India" name="Umtasor" pageId="1" pageNumber="594" stateProvince="Meghalaya">Umtasor</location>
(
<collectingRegion box="[431,560,927,953]" country="India" name="Meghalaya" pageId="1" pageNumber="594">Meghalaya</collectingRegion>
,
<collectingCountry box="[571,630,927,953]" name="India" pageId="1" pageNumber="594">India</collectingCountry>
)
</materialsCitation>
</paragraph>
<paragraph blockId="1.[151,1437,927,1998]" box="[199,1252,963,992]" pageId="1" pageNumber="594">
<emphasis bold="true" box="[199,435,963,989]" pageId="1" pageNumber="594">Material examined:</emphasis>
<materialsCitation box="[442,1252,963,992]" collectingDate="2014-09-15" collectionCode="R" country="India" location="Umtasor" pageId="1" pageNumber="594" specimenCount="1" stateProvince="Meghalaya" typeStatus="holotype">
<emphasis bold="true" box="[442,551,963,989]" pageId="1" pageNumber="594">
<typeStatus box="[442,551,963,989]" pageId="1" pageNumber="594">Holotype</typeStatus>
</emphasis>
(♁).
<collectingCountry box="[611,671,963,989]" name="India" pageId="1" pageNumber="594">India</collectingCountry>
,
<emphasis bold="true" box="[683,815,963,989]" pageId="1" pageNumber="594">
<collectingRegion box="[683,815,963,989]" country="India" name="Meghalaya" pageId="1" pageNumber="594">Meghalaya</collectingRegion>
</emphasis>
,
<location LSID="urn:lsid:plazi:treatment:03BAA304FFD0FFCDFF0B483CFBDB1BAB:8ECC44C9FFD0FFCDFCA148CBFC521FB8" box="[829,927,964,990]" country="India" name="Umtasor" pageId="1" pageNumber="594" stateProvince="Meghalaya">Umtasor</location>
,
<date box="[938,1058,963,990]" pageId="1" pageNumber="594" value="2014-09-15">
<collectingDate box="[938,1058,963,990]" pageId="1" pageNumber="594" value="2014-09-15">15.ix.2014</collectingDate>
</date>
(
<emphasis box="[1074,1108,963,989]" italics="true" pageId="1" pageNumber="594">leg</emphasis>
.
<collectionCode box="[1123,1145,964,990]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="594">R</collectionCode>
. Ranjan).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="594" type="diagnosis">
<paragraph blockId="1.[151,1437,927,1998]" pageId="1" pageNumber="594">
<emphasis bold="true" box="[199,324,999,1025]" pageId="1" pageNumber="594">Diagnosis:</emphasis>
Morphologically,
<taxonomicName authority="Ranjan, Singh &amp; Kirti" authorityName="Ranjan, Singh &amp; Kirti" authorityYear="2023" box="[535,901,999,1026]" class="Insecta" family="Pyralidae" genus="Coenodomus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="wangi" status="sp. nov.">
<emphasis box="[535,638,1000,1026]" italics="true" pageId="1" pageNumber="594">C. wangi</emphasis>
Ranjan, Singh &amp; Kirti
</taxonomicName>
,
<emphasis bold="true" box="[917,1009,999,1025]" pageId="1" pageNumber="594">
<taxonomicNameLabel box="[917,1009,999,1025]" pageId="1" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[1026,1093,999,1025]" captionStart="FIGURES 15" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[175,1411,200,1919]" captionTargetId="figure-29@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 15. Imago and male genitalia of Coenodomus spp.: 1, 4, C. wangi sp. nov., male; 2, 5, C. dudgeoni, male (photo credit: Wang et al. 2017); 3, C. dudgeoni, male (© NMHUK)." figureDoi="http://doi.org/10.5281/zenodo.7837224" httpUri="https://zenodo.org/record/7837224/files/figure.png" pageId="1" pageNumber="594">Fig. 1</figureCitation>
) is most closely similar to
<taxonomicName authorityName="Hampson" authorityYear="1896" class="Insecta" family="Pyralidae" genus="Coenodomus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="dudgeoni">
<emphasis italics="true" pageId="1" pageNumber="594">C. dudgeoni</emphasis>
</taxonomicName>
(
<figureCitation box="[273,376,1035,1061]" captionStart="FIGURES 15" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[175,1411,200,1919]" captionTargetId="figure-29@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 15. Imago and male genitalia of Coenodomus spp.: 1, 4, C. wangi sp. nov., male; 2, 5, C. dudgeoni, male (photo credit: Wang et al. 2017); 3, C. dudgeoni, male (© NMHUK)." figureDoi="http://doi.org/10.5281/zenodo.7837224" httpUri="https://zenodo.org/record/7837224/files/figure.png" pageId="1" pageNumber="594">Figs 23</figureCitation>
) and is best distinguished on the basis of male genitalia (
<figureCitation box="[1044,1116,1035,1061]" captionStart="FIGURES 15" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[175,1411,200,1919]" captionTargetId="figure-29@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 15. Imago and male genitalia of Coenodomus spp.: 1, 4, C. wangi sp. nov., male; 2, 5, C. dudgeoni, male (photo credit: Wang et al. 2017); 3, C. dudgeoni, male (© NMHUK)." figureDoi="http://doi.org/10.5281/zenodo.7837224" httpUri="https://zenodo.org/record/7837224/files/figure.png" pageId="1" pageNumber="594">Fig. 4</figureCitation>
): the mediolateral spines of uncus are absent (whereas in
<taxonomicName authorityName="Hampson" authorityYear="1896" box="[481,618,1071,1098]" class="Insecta" family="Pyralidae" genus="Coenodomus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="dudgeoni">
<emphasis box="[481,618,1071,1098]" italics="true" pageId="1" pageNumber="594">C. dudgeoni</emphasis>
</taxonomicName>
, the lateral sides of uncus bear a prominent medio lateral spine (
<figureCitation box="[1344,1411,1071,1098]" captionStart="FIGURES 15" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[175,1411,200,1919]" captionTargetId="figure-29@2.[166,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 15. Imago and male genitalia of Coenodomus spp.: 1, 4, C. wangi sp. nov., male; 2, 5, C. dudgeoni, male (photo credit: Wang et al. 2017); 3, C. dudgeoni, male (© NMHUK)." figureDoi="http://doi.org/10.5281/zenodo.7837224" httpUri="https://zenodo.org/record/7837224/files/figure.png" pageId="1" pageNumber="594">Fig. 5</figureCitation>
)), and the small, dorsally directed process present on the tip of basal part of sacculus, which is absent in
<taxonomicName authorityName="Hampson" authorityYear="1896" box="[1292,1429,1107,1134]" class="Insecta" family="Pyralidae" genus="Coenodomus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="dudgeoni">
<emphasis box="[1292,1429,1107,1134]" italics="true" pageId="1" pageNumber="594">C. dudgeoni</emphasis>
</taxonomicName>
. Another closely related species is
<taxonomicName authorityName="Hampson" authorityYear="1891" box="[541,717,1143,1170]" class="Insecta" family="Pyralidae" genus="Coenodomus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="rotundinidus">
<emphasis box="[541,717,1143,1170]" italics="true" pageId="1" pageNumber="594">C. rotundinidus</emphasis>
</taxonomicName>
(see
<bibRefCitation author="Wang, M. Q. &amp; Chen, F. Q. &amp; Wu, C. S." box="[779,979,1143,1170]" pageId="1" pageNumber="594" pagination="55 - 68" refId="ref3459" refString="Wang, M. Q., Chen, F. Q. &amp; Wu, C. S. (2017) A review of the genus Coenodomus Walsingham, 1888 in China (Lepidoptera: Pyralidae: Epipaschiinae), with descriptions of four new species. Annales Zoologici, Warszawa, 67 (1), 55 - 68. https: // doi. org / 10.3161 / 00034541 ANZ 2017.67.1.007" type="journal article" year="2017">
Wang
<emphasis box="[853,911,1143,1169]" italics="true" pageId="1" pageNumber="594">et al.</emphasis>
2017
</bibRefCitation>
: Fig. 22), but
<taxonomicName authorityName="Ranjan, Singh &amp; Kirti" authorityYear="2023" box="[1142,1244,1144,1170]" class="Insecta" family="Pyralidae" genus="Coenodomus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="wangi">
<emphasis box="[1142,1244,1144,1170]" italics="true" pageId="1" pageNumber="594">C. wangi</emphasis>
</taxonomicName>
is distinct in the distal costa extending the valva, the lack of the lateral spines of the uncus, and the juxta split from apex to base to form two elongate thin processes (whereas in
<taxonomicName authorityName="Hampson" authorityYear="1891" box="[674,850,1215,1242]" class="Insecta" family="Pyralidae" genus="Coenodomus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="species" species="rotundinidus">
<emphasis box="[674,850,1215,1242]" italics="true" pageId="1" pageNumber="594">C. rotundinidus</emphasis>
</taxonomicName>
, the valva apex is extended to form a rounded lobe together with the distal costa, lateral spines are present at the uncus sides, and the juxta is less deeply bifurcated, with a heart-shaped base).
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="594" type="description">
<paragraph blockId="1.[151,1437,927,1998]" pageId="1" pageNumber="594">
<emphasis bold="true" box="[199,347,1323,1349]" pageId="1" pageNumber="594">Description:</emphasis>
Adults. Labial palpi upturned. Antennae bipectinated, pectination shortened towards tip; scape bearing a long projection, densely covered with long hair-like scales and reaching up to end of metathorax. Patagia with long hairs, reaching mesothorax. Forewing dark ash colour; costal area dominated by olive scales; fuscous tufts of hairs basal of pale sinuous antemedial line; another tuft on discocellular; postmedial line interrupted, slightly S-shaped, olive patch beyond it on inner margin, outer area with tufts of fuscous and white scales. Ante- and postmedial lines bordered with olive bands. Hindwing concolorous to forewing with an indistinct, sinuous, white medial line, an olive patch beyond it at anal angle; tuft of fuscous hairs near base, inner margin, end of cell and on outer area. Underside of both wings with markings extremely reduced. Male genitalia with uncus almost rounded, broad, setose dorsally, folded laterally, concave ventrally, slightly notched at apex. Gnathos short, reaching up to 1/4
<superScript attach="left" box="[187,200,1647,1661]" fontSize="6" pageId="1" pageNumber="594">th</superScript>
of uncus, lateral arms sclerotised, conjoined apically to form a spined apex. Tegumen broad, shorter than uncus. Vinculum longer than tegumen, forming small U-shape, with notched saccus. Transtilla membranous, slightly sclerotised at base. Juxta reduced to two thin elongate lateral processes connected in U-shape at ventral end, each process at half the length bearing series of minute spines. Valva simple, broad, setose with a triangular window at base, costa slightly concave, sclerotised, distal costa conspicuously produced beyond valva, sacculus swollen at base with a small dorsally directed process at distal end of swollen part. Aedeagus bent ca. 80˚ at the entry of ductus ejaculatorius, coecum cylindrical, vesica with a small crescent-shaped sclerotisation.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="594" type="etymology">
<paragraph blockId="1.[151,1437,927,1998]" pageId="1" pageNumber="594">
<emphasis bold="true" box="[199,338,1899,1925]" pageId="1" pageNumber="594">Etymology:</emphasis>
The species is named in honour of Dr. MingQiang Wang (
<collectingCountry box="[998,1068,1899,1925]" name="China" pageId="1" pageNumber="594">China</collectingCountry>
) for his valuable contributions to the taxonomy of the genus
<taxonomicName authority="Walsingham." authorityName="Walsingham." box="[455,755,1935,1961]" class="Insecta" family="Pyralidae" genus="Coenodomus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis box="[455,601,1935,1961]" italics="true" pageId="1" pageNumber="594">Coenodomus</emphasis>
Walsingham.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection box="[199,1046,1971,1998]" pageId="1" pageNumber="594" type="distribution">
<paragraph blockId="1.[151,1437,927,1998]" box="[199,1046,1971,1998]" pageId="1" pageNumber="594">
<emphasis bold="true" box="[199,355,1971,1997]" pageId="1" pageNumber="594">Distribution:</emphasis>
The new species is so far reported from its
<typeStatus box="[844,892,1972,1998]" pageId="1" pageNumber="594">type</typeStatus>
locality only.
</paragraph>
</subSubSection>
</treatment>
</document>