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<document id="8FC698E07C55DA1743B7DD9258C8D8C8" ID-DOI="10.5281/zenodo.175790" ID-GBIF-Dataset="2a6512b7-5bd9-4682-b5db-79c72373b68c" ID-ISSN="1175-5326" ID-Zenodo-Dep="175790" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459820893039" checkinUser="plazi" docAuthor="Freitag, Hendrik &amp; Jäch, Manfred A." docDate="2007" docId="03B9F24A7C0BFF9EFF30530A6793278F" docLanguage="en" docName="zt01431p044.pdf" docOrigin="Zootaxa 1431" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hydraena (Hydraenopsis) jojoorculloi Freitag &amp; Jach, sp. n." docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="FF808A327C00FF90FFA75435641C2645" masterDocTitle="Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d’Orchymont, 1925" masterLastPageNumber="44" masterPageNumber="1" pageNumber="12" updateTime="1698221773297" updateUser="plazi">
<mods:mods id="0D9ECB323C84FCFB5C557140148FFB84" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FD99BA6A0D54089CC24B8EECE0A8413E">
<mods:title id="4B79D44536212AC910B1992558861985">Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d’Orchymont, 1925</mods:title>
</mods:titleInfo>
<mods:name id="97CA6F0323E32E70073A56330C57A4B8" type="personal">
<mods:role id="77CD7D8638A4F9FE45BD78539C9CA013">
<mods:roleTerm id="C5D204A8B1C505A62DE65C793F39EB55">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C299AF73A82ADE8BDB25644879C6E908">Freitag, Hendrik</mods:namePart>
</mods:name>
<mods:name id="819C3FFFA8CDC5D82E6F743CCC2258A9" type="personal">
<mods:role id="BB4FC307074F7D32607D8B259DA4302F">
<mods:roleTerm id="A9BD5C92D77A5C2C6A881C78FE5A68BE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="05153C356DE3DC91DABCD320B53CA879">Jäch, Manfred A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="8EFEC7C86304E5B456DCA6BAA34C31BE">text</mods:typeOfResource>
<mods:relatedItem id="DCD953A35793411D24CDB97DBAEA8654" type="host">
<mods:titleInfo id="3A39CFA623E2CD3C8E0C4934A9F5BB3A">
<mods:title id="8E3CA8AFAF8A34FED8381A03DC7C902B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2790BE4B69BF2D72DA3F25DFACFCFC16">
<mods:date id="344CB4AC3DFB7CCCEAD707FD6E910600">2007</mods:date>
<mods:detail id="73EF1B3FEBE7587D02468F967C837020" type="volume">
<mods:number id="6319D871DCAEFA336EAA21BCA21CF134">1431</mods:number>
</mods:detail>
<mods:extent id="B9EA0B77EB25A5EC6750163AFCAC7645" unit="page">
<mods:start id="97121D231773BD7920B30799F1282D6F">1</mods:start>
<mods:end id="1AED0926CA19E814C03D22DBEDD1E865">44</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="874901441D95C6ECB3EFDF8177C5BBAC">journal article</mods:classification>
<mods:identifier id="31B96C52A0FB29C97F772CE3736F9DDC" type="DOI">10.5281/zenodo.175790</mods:identifier>
<mods:identifier id="B776EB0D2936392C055F0E02AE323065" type="GBIF-Dataset">2a6512b7-5bd9-4682-b5db-79c72373b68c</mods:identifier>
<mods:identifier id="EB437D2591D99583B068D1456ECAC998" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E6DC18FFC1A9FD64A6268DDD39384DEF" type="Zenodo-Dep">175790</mods:identifier>
</mods:mods>
<treatment id="03B9F24A7C0BFF9EFF30530A6793278F" ID-DOI="http://doi.org/10.5281/zenodo.6248069" ID-GBIF-Taxon="119355662" ID-Zenodo-Dep="6248069" LSID="urn:lsid:plazi:treatment:03B9F24A7C0BFF9EFF30530A6793278F" httpUri="http://treatment.plazi.org/id/03B9F24A7C0BFF9EFF30530A6793278F" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
<subSubSection id="C30A10D77C0BFF9BFF30530A654C2139" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BAF435C7C0BFF9BFF30530A676B211C" blockId="11.[151,887,1855,1916]" box="[151,887,1855,1881]" pageId="11" pageNumber="12">
<heading id="D0E7F4307C0BFF9BFF30530A676B211C" bold="true" box="[151,887,1855,1881]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B9649F4E7C0BFF9BFF30530A676B211C" bold="true" box="[151,887,1855,1881]" pageId="11" pageNumber="12">
<taxonomicName id="4C1038DF7C0BFF9BFF30530A6734211C" ID-CoL="3N3C7" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[151,808,1855,1881]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="jojoorculloi" status="sp. nov." subGenus="Hydraenopsis">
<emphasis id="B9649F4E7C0BFF9BFF30530A650C211C" bold="true" box="[151,272,1855,1881]" italics="true" pageId="11" pageNumber="12">Hydraena</emphasis>
(
<emphasis id="B9649F4E7C0BFF9BFE87530A65DA211C" bold="true" box="[288,454,1855,1881]" italics="true" pageId="11" pageNumber="12">Hydraenopsis</emphasis>
)
<emphasis id="B9649F4E7C0BFF9BFE70530A667F211C" bold="true" box="[471,611,1855,1881]" italics="true" pageId="11" pageNumber="12">jojoorculloi</emphasis>
Freitag &amp; Jäch
</taxonomicName>
,
<taxonomicNameLabel id="A25722357C0BFF9BFC92530A676B211C" box="[821,887,1855,1881]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF435C7C0BFF9BFF305357654C2139" blockId="11.[151,887,1855,1916]" box="[151,336,1890,1916]" pageId="11" pageNumber="12">(Figs. 4, 16a–h)</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C0BFF9DFF30538465F627E7" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BAF435C7C0BFF9BFF30538464C021B1" blockId="11.[151,1436,1969,2036]" pageId="11" pageNumber="12">
<emphasis id="B9649F4E7C0BFF9BFF3053846521218E" bold="true" box="[151,317,1969,1995]" pageId="11" pageNumber="12">
<typeStatus id="54ABFDFE7C0BFF9BFF30538464C8218E" box="[151,212,1969,1995]" pageId="11" pageNumber="12">Type</typeStatus>
locality.
</emphasis>
<taxonomicName id="4C1038DF7C0BFF9BFEE3538765AF2189" box="[324,435,1970,1996]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
Stream (Lake
<taxonomicName id="4C1038DF7C0BFF9BFDC7538766D22189" box="[608,718,1970,1996]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
tributary), Municipality of Taytay, northern Palawan,
<collectingCountry id="F30703CC7C0BFF9BFAEB538764C521B1" name="Philippines" pageId="11" pageNumber="12">Philippines</collectingCountry>
.
</paragraph>
<caption id="DF6F13D47C0CFF9CFF30524E60F620F1" httpUri="https://zenodo.org/record/175794/files/figure.png" pageId="12" pageNumber="13" targetBox="[270,1322,190,1625]" targetPageId="12">
<paragraph id="8BAF435C7C0CFF9CFF30524E60F620F1" blockId="12.[151,1436,1659,1716]" pageId="12" pageNumber="13">
<emphasis id="B9649F4E7C0CFF9CFF30524E653B20D6" bold="true" box="[151,295,1659,1683]" pageId="12" pageNumber="13">FIGURE 16.</emphasis>
<taxonomicName id="4C1038DF7C0CFF9CFE89524E660120D7" box="[302,541,1659,1682]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="jojoorculloi" status="sp. nov.">
<emphasis id="B9649F4E7C0CFF9CFE89524E660120D7" box="[302,541,1659,1682]" italics="true" pageId="12" pageNumber="13">Hydraena jojoorculloi</emphasis>
</taxonomicName>
<emphasis id="B9649F4E7C0CFF9CFD82524E667D20D6" bold="true" box="[549,609,1659,1683]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="A25722357C0CFF9CFD82524E667D20D6" box="[549,609,1659,1683]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
a–c) aedeagus (holotype) in ventral (setae of right paramere omitted), dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f–g) spermatheca; h) female tergite X.
</paragraph>
</caption>
<paragraph id="8BAF435C7C0CFF9DFF60532565F627E7" blockId="12.[151,1437,1808,2034]" lastBlockId="13.[151,1437,151,2018]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="B9649F4E7C0CFF9CFF6053256618216F" bold="true" box="[199,516,1808,1834]" pageId="12" pageNumber="13">
<typeStatus id="54ABFDFE7C0CFF9CFF605325651E216F" box="[199,258,1808,1834]" pageId="12" pageNumber="13">Type</typeStatus>
material.
<typeStatus id="54ABFDFE7C0CFF9CFE3553256618216F" box="[402,516,1808,1834]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
</emphasis>
ɗ (
<collectionCode id="ED01DB997C0CFF9CFD9A5325668C216F" box="[573,656,1808,1834]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="13">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C0CFF9CFD125325671C216F" box="[693,768,1808,1834]" pageId="12" pageNumber="13">PHIL</collectionCode>
.: Palawan, Taytay; Lake
<taxonomicName id="4C1038DF7C0CFF9CFB99532560B1216F" box="[1086,1197,1808,1834]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
tributary, Alipuran Stream, semi-prim. forest c.
<quantity id="4CE8EEB97C0CFF9CFE70530D660C2117" box="[471,528,1848,1874]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="13" unit="m" value="40.0">40m</quantity>
asl,
<geoCoordinate id="EE24259B7C0CFF9CFDE2530D66BA2117" box="[581,678,1848,1874]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="10.75">10°45'N</geoCoordinate>
<geoCoordinate id="EE24259B7C0CFF9CFD0B530D670B2117" box="[684,791,1848,1874]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="119.53333">
119°32'
<geoCoordinate id="EE24259B7C0CFF9CFCA3530D670B2117" box="[772,791,1848,1874]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="5555" value="29.4">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C0CFF9CFCB9530D678A2117" box="[798,918,1848,1874]" pageId="12" pageNumber="13" value="1995-04-29">29.4.1995</date>
, leg. Freitag (15)M”, terminal parts of abdomen and aedeagus glued separately.
<emphasis id="B9649F4E7C0CFF9CFDEC535566CD213F" bold="true" box="[587,721,1888,1914]" pageId="12" pageNumber="13">
<typeStatus id="54ABFDFE7C0CFF9CFDEC535566D1213F" box="[587,717,1888,1914]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
:
</emphasis>
2 ΨΨ (
<collectionCode id="ED01DB997C0CFF9CFC8B53556762213F" box="[812,894,1888,1914]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="13">NMW</collectionCode>
): same locality data as
<typeStatus id="54ABFDFE7C0CFF9CFB315355611C213F" box="[1174,1280,1888,1914]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
; 1 ɗ (
<collectionCode id="ED01DB997C0CFF9CFAED5355618B213F" box="[1354,1431,1888,1914]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="13">NMW</collectionCode>
, aedeagus lacking), 2 ΨΨ (
<collectionCode id="ED01DB997C0CFF9CFE6B53BD660521E7" box="[460,537,1928,1954]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="13">NMW</collectionCode>
, mutilated): “
<collectionCode id="ED01DB997C0CFF9CFD1C53BD671A21E7" box="[699,774,1928,1954]" pageId="12" pageNumber="13">PHIL</collectionCode>
.: Palawan, Taytay; 1/2way high-way to Lake
<taxonomicName id="4C1038DF7C0CFF9CFA8153BD618421E7" box="[1318,1432,1928,1954]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
,
<taxonomicName id="4C1038DF7C0CFF9CFF305385651A218F" box="[151,262,1968,1994]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
Stream trib., prim.forest c.
<quantity id="4CE8EEB97C0CFF9CFDE553856665218F" box="[578,633,1968,1994]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="13" unit="m" value="30.0">30m</quantity>
asl,
<geoCoordinate id="EE24259B7C0CFF9CFD0E53856717218F" box="[681,779,1968,1994]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="10.783334">10°47'N</geoCoordinate>
<geoCoordinate id="EE24259B7C0CFF9CFCB553856761218F" box="[786,893,1968,1994]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="119.51667">119°31'E</geoCoordinate>
03.4.1995,leg. Freitag(13a)M”; 2 ΨΨ (
<collectionCode id="ED01DB997C0CFF9CFAED5385618B218F" box="[1354,1431,1968,1994]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="13">NMW</collectionCode>
, 1 ex. mutilated): “
<collectionCode id="ED01DB997C0CFF9CFEDB53ED65DB21B7" box="[380,455,2008,2034]" pageId="12" pageNumber="13">PHIL</collectionCode>
.: Palawan, Taytay; 1/3way high-way to Lake
<taxonomicName id="4C1038DF7C0CFF9CFBA253ED606B21B7" box="[1029,1143,2008,2034]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
,
<taxonomicName id="4C1038DF7C0CFF9CFB2E53ED60E421B7" box="[1161,1272,2008,2034]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
Stream trib., prim.forest c.
<quantity id="4CE8EEB97C0DFF9DFE9354AD657026F7" box="[308,364,152,178]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="14" unit="m" value="35.0">35m</quantity>
asl,
<geoCoordinate id="EE24259B7C0DFF9DFE0E54AD661526F7" box="[425,521,152,178]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="10.783334">10°47'N</geoCoordinate>
<geoCoordinate id="EE24259B7C0DFF9DFDB254AD669C26F7" box="[533,640,152,178]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="925" value="119.51667">
119°31'
<geoCoordinate id="EE24259B7C0DFF9DFDCB54AD669C26F7" box="[620,640,152,178]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="5555" value="29.4">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C0DFF9DFD2C54AD671E26F7" box="[651,770,152,178]" pageId="13" pageNumber="14" value="1995-04-29">29.4.1995</date>
, leg. Freitag (13b)M”; 1 Ψ (
<collectionCode id="ED01DB997C0DFF9DFBC754AD60AE26F7" box="[1120,1202,152,178]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="14">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C0DFF9DFB7354AD610326F7" box="[1236,1311,152,178]" pageId="13" pageNumber="14">PHIL</collectionCode>
.: Palawan, P.Princesa; S. Vincente Taranaban Riv;
<quantity id="4CE8EEB97C0DFF9DFDDC54F566DA269F" box="[635,710,192,218]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="14" unit="km" value="0.5">0.5km</quantity>
N of Highway, mountain riv. resid. pool,
<quantity id="4CE8EEB97C0DFF9DFB7754F56118269F" box="[1232,1284,192,218]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="m" value="10.0">10m</quantity>
asl
<geoCoordinate id="EE24259B7C0DFF9DFA9C54F56180269F" box="[1339,1436,192,218]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="10.016666">10°01'N</geoCoordinate>
<geoCoordinate id="EE24259B7C0DFF9DFF3054DD65182747" box="[151,260,232,258]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="925" value="119.01667">119°01'E</geoCoordinate>
,
<date id="FFAE659C7C0DFF9DFEB754DD65992747" box="[272,389,232,258]" pageId="13" pageNumber="14" value="1995-04-30">30.4.1995</date>
leg. Freitag (16)M”; 1 Ψ (
<collectionCode id="ED01DB997C0DFF9DFD1F54DD67162747" box="[696,778,232,258]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="14">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C0DFF9DFC8F54DD676F2747" box="[808,883,232,258]" pageId="13" pageNumber="14">PHIL</collectionCode>
.: Palawan, P.Princesa; Junction to Napsan,
<quantity id="4CE8EEB97C0DFF9DFACF54DD61802747" box="[1384,1436,232,258]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="14" unit="km" value="8.0">8km</quantity>
SW PPC;
<quantity id="4CE8EEB97C0DFF9DFEB55525655A276F" box="[274,326,272,298]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="m" value="20.0">20m</quantity>
S Binuan Bridge, sec.veget. c.
<quantity id="4CE8EEB97C0DFF9DFD12552566F1276F" box="[693,749,272,298]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="m" value="20.0">20m</quantity>
asl
<geoCoordinate id="EE24259B7C0DFF9DFC875525676F276F" box="[800,883,272,298]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="9.716666">9°43'N</geoCoordinate>
<geoCoordinate id="EE24259B7C0DFF9DFCDB552567F6276F" box="[892,1002,272,298]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="925" value="118.666664">118°40'E</geoCoordinate>
,
<date id="FFAE659C7C0DFF9DFC5055256071276F" box="[1015,1133,272,298]" pageId="13" pageNumber="14" value="1995-03-28">28.3.1995</date>
leg. Freitag (22)M”; 1 Ψ (
<collectionCode id="ED01DB997C0DFF9DFF38550D64ED2717" box="[159,241,312,338]" httpUri="http://grbio.org/cool/134h-pm2f" name="Museum of Natural History, University of the Philippines" pageId="13" pageNumber="14">UPLB</collectionCode>
): “
<collectionCode id="ED01DB997C0DFF9DFEB5550D65412717" box="[274,349,312,338]" pageId="13" pageNumber="14">PHIL</collectionCode>
.: Palawan, P. Princesa Panaguman R.PR1, 1.Drift
<geoCoordinate id="EE24259B7C0DFF9DFC1E550D605D2717" box="[953,1089,312,338]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="15" value="10.2525">10°15'09''N</geoCoordinate>
<geoCoordinate id="EE24259B7C0DFF9DFBEC550D60C22717" box="[1099,1246,312,338]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="15" value="118.9675">118°58'03''E</geoCoordinate>
<date id="FFAE659C7C0DFF9DFB4F550D617E2717" box="[1256,1378,312,338]" pageId="13" pageNumber="14" value="2001-05-17">17.V.2001</date>
, leg. Freitag (47)”; 1 Ψ (
<collectionCode id="ED01DB997C0DFF9DFEDD555565D1273F" box="[378,461,352,378]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="14">NMW</collectionCode>
): “
<collectingCountry id="F30703CC7C0DFF9DFE4D55556668273F" box="[490,628,352,378]" name="Philippines" pageId="13" pageNumber="14">Philippines</collectingCountry>
, Palawan centr. Ulangoan river,
<quantity id="4CE8EEB97C0DFF9DFC5455556022273F" box="[1011,1086,352,378]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="13" pageNumber="14" unit="km" value="18.0">18 km</quantity>
NE San Rafael,
<date id="FFAE659C7C0DFF9DFAA35555618B273F" box="[1284,1431,352,378]" pageId="13" pageNumber="14" value="1995-12-06">6. XII. 1995</date>
, J. Kodada &amp; B. Rigová lgt.”.
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C0DFF9EFF60559A663E269F" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
<paragraph id="8BAF435C7C0DFF9DFF60559A607527B7" blockId="13.[151,1437,151,2018]" pageId="13" pageNumber="14">
<emphasis id="B9649F4E7C0DFF9DFF60559A6543278C" bold="true" box="[199,351,431,457]" pageId="13" pageNumber="14">Description.</emphasis>
Body approximately
<quantity id="4CE8EEB97C0DFF9DFDC4558566D2278F" box="[611,718,432,458]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="13" pageNumber="14" unit="mm" value="1.25">1.25 mm</quantity>
long. Habitus as in Fig. 4. Elytra brown; frons dark brown; pronotum, legs and maxillary palpi distinctly paler yellowish brown to gold-brown.
</paragraph>
<paragraph id="8BAF435C7C0DFF9DFF605635678824FF" blockId="13.[151,1437,151,2018]" pageId="13" pageNumber="14">Pronotum wider than long, broadest at anterior 0.3; entire pronotum densely punctate; punctures small and moderately deeply impressed, interstices glabrous; anterior margin slightly concave; anterior angles rounded; lateral rim denticulate; lateral margins anteriorly convergent, posteriorly sinuously convergent; pronoto-elytral angle obtuse; posterior margin slightly biconcave; lateral hypomeron posteriorly slightly broader than profemur, mesal hypomeron narrow, inconspicuous.</paragraph>
<paragraph id="8BAF435C7C0DFF9DFF6056FD604D25EF" blockId="13.[151,1437,151,2018]" pageId="13" pageNumber="14">Elytra elongately oval, not conjointly rounded apically; each elytron with a tiny excision next to sutural keel, with ca. 16 longitudinal, more or less regular rows of punctures (approximately nine between suture and shoulder); punctures small and moderately deeply (apically shallowly) impressed, interstices and intervals convex to microreticulate; lateral portion moderately explanate, not reaching elytral apex, anteriorly and posteriorly denticulate; inflexed lateral portion (pseudepipleuron) anteriorly slightly broader than metafemur, reaching posterior 0.2, row of punctures inconspicuous; epipleuron narrow, short.</paragraph>
<paragraph id="8BAF435C7C0DFF9DFF60578D6740220F" blockId="13.[151,1437,151,2018]" pageId="13" pageNumber="14">Mesoventrite with well developed longitudinal ridges; mesoventral intercoxal process moderately broad, slightly narrower than pseudepipleuron; metaventral disc distinctly impressed; metaventral plaques inconspicuous, pubescent; intercoxal sternite moderately large, approximately as broad as metafemur and as long as broad; metaventrite and ventrites 1–4 moderately pubescent.</paragraph>
<paragraph id="8BAF435C7C0DFF9DFF60506D664722DF" blockId="13.[151,1437,151,2018]" pageId="13" pageNumber="14">Femora with inner margin straight to concave, outer margin conspicuously convex; tibiae with inner margin straight, outer one slightly convex.</paragraph>
<paragraph id="8BAF435C7C0DFF9DFF60509D603523F7" blockId="13.[151,1437,151,2018]" pageId="13" pageNumber="14">
Aedeagus (
<figureCitation id="132B5FD97C0DFF9DFEEB509D65B72287" box="[332,427,1192,1218]" captionStart="FIGURE 16" captionStartId="12.[151,255,1659,1683]" captionTargetBox="[270,1322,190,1625]" captionTargetId="figure@12.[264,1323,182,1634]" captionTargetPageId="12" captionText="FIGURE 16. Hydraena jojoorculloi sp. n. a – c) aedeagus (holotype) in ventral (setae of right paramere omitted), dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175794/files/figure.png" pageId="13" pageNumber="14">Figs. 16</figureCitation>
a–c) elongate and slender; main piece rather straight in ventral view, slightly sinuous in lateral view; without setae (three micropores present); phallobase asymmetric. Distal lobe intricately shaped, not clearly delimited from main piece; ejaculatory duct distally ending in a tapered stout flagellum, which is dorsally embedded in a less sclerotized ovoid structure. Parameres firmly fused to main piece; left paramere short, moderately wide, inserted at about basal 0.50 of aedeagus, latero-apically with ca. seven moderately long setae; right paramere inserted at about basal 0.55 of aedeagus, longer, wide, apically attenuately rounded, subapically with two parallel rows of ca. four moderately long setae.
</paragraph>
<paragraph id="8BAF435C7C0DFF9DFF6051F560F1206F" blockId="13.[151,1437,151,2018]" pageId="13" pageNumber="14">
Gonocoxite (
<figureCitation id="132B5FD97C0DFF9DFEC651F565AA239F" box="[353,438,1472,1498]" captionStart="FIGURE 16" captionStartId="12.[151,255,1659,1683]" captionTargetBox="[270,1322,190,1625]" captionTargetId="figure@12.[264,1323,182,1634]" captionTargetPageId="12" captionText="FIGURE 16. Hydraena jojoorculloi sp. n. a – c) aedeagus (holotype) in ventral (setae of right paramere omitted), dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175794/files/figure.png" pageId="13" pageNumber="14">Fig. 16</figureCitation>
e) subsemicircular, more or less evenly rounded; lateral margin only slightly convex subbasally; setae of subapical tufts straight; basal area of ventral plate with very few small scattered setae; condyles small and inconspicuous; dorsal plate asymmetric, simple, not surpassing outer plate.
</paragraph>
<paragraph id="8BAF435C7C0DFF9DFF60520D663C203F" blockId="13.[151,1437,151,2018]" pageId="13" pageNumber="14">
Spermatheca as in
<figureCitation id="132B5FD97C0DFF9DFE06520D65E22017" box="[417,510,1592,1618]" captionStart="FIGURE 16" captionStartId="12.[151,255,1659,1683]" captionTargetBox="[270,1322,190,1625]" captionTargetId="figure@12.[264,1323,182,1634]" captionTargetPageId="12" captionText="FIGURE 16. Hydraena jojoorculloi sp. n. a – c) aedeagus (holotype) in ventral (setae of right paramere omitted), dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175794/files/figure.png" pageId="13" pageNumber="14">Figs. 16</figureCitation>
f–g; proximal portion distally globular, proximately tubular, bent; distal portion wide and tubular, slightly curved.
</paragraph>
<paragraph id="8BAF435C7C0DFF9DFF6052BD606C212F" blockId="13.[151,1437,151,2018]" pageId="13" pageNumber="14">
Secondary sexual characters. Male profemur with tiny denticle near base of ventral face (dissection of leg recommended). Male terminal sternite elongately subpentagonal, apical margin widely rounded; base slender, very slightly produced laterad; spiculum gastrale almost twice as long as terminal sternite, slightly bent (
<figureCitation id="132B5FD97C0DFF9DFACA52ED64AA215F" captionStart="FIGURE 16" captionStartId="12.[151,255,1659,1683]" captionTargetBox="[270,1322,190,1625]" captionTargetId="figure@12.[264,1323,182,1634]" captionTargetPageId="12" captionText="FIGURE 16. Hydraena jojoorculloi sp. n. a – c) aedeagus (holotype) in ventral (setae of right paramere omitted), dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175794/files/figure.png" pageId="13" pageNumber="14">Fig. 16</figureCitation>
d). Male tergite X apically distinctly excised. Female tergite X (
<figureCitation id="132B5FD97C0DFF9DFC0B5335601D215F" box="[940,1025,1792,1818]" captionStart="FIGURE 16" captionStartId="12.[151,255,1659,1683]" captionTargetBox="[270,1322,190,1625]" captionTargetId="figure@12.[264,1323,182,1634]" captionTargetPageId="12" captionText="FIGURE 16. Hydraena jojoorculloi sp. n. a – c) aedeagus (holotype) in ventral (setae of right paramere omitted), dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175794/files/figure.png" pageId="13" pageNumber="14">Fig. 16</figureCitation>
h) subsemicircular; lateral margins evenly rounded; disc sparsely covered with setae; subapical fringe of vermiform setae widely interrupted medially, laterally with short trichoid setae; hyaline apical margin notched medially.
</paragraph>
<paragraph id="8BAF435C7C0DFF9EFF605342663E269F" blockId="13.[151,1437,151,2018]" lastBlockId="14.[151,1436,152,458]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="B9649F4E7C0DFF9DFF60534265C021D4" bold="true" box="[199,476,1911,1937]" pageId="13" pageNumber="14">Differential diagnosis.</emphasis>
This species can easily be distinguished from
<taxonomicName id="4C1038DF7C0DFF9DFBB1534D60FA21D4" box="[1046,1254,1912,1937]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="scabra">
<emphasis id="B9649F4E7C0DFF9DFBB1534D60FA21D4" box="[1046,1254,1912,1937]" italics="true" pageId="13" pageNumber="14">Hydraena scabra</emphasis>
</taxonomicName>
by the smaller size, by the conspicuously yellowish pronotum, and by its smaller and only moderately deeply impressed elytral and pronotal punctures. Furthermore, it can easily be recognized by its aedeagus (main piece rather straight, distal lobe ovoid with flagellum-like structure), the shape of the gonocoxite, and the presence of vermiform setae on female tergite X.
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C0EFF9EFF6054D2653F273F" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8BAF435C7C0EFF9EFF6054D260F92747" blockId="14.[151,1436,152,458]" box="[199,1253,231,258]" pageId="14" pageNumber="15">
<emphasis id="B9649F4E7C0EFF9EFF6054D2657C2744" bold="true" box="[199,352,231,257]" pageId="14" pageNumber="15">Distribution</emphasis>
(
<figureCitation id="132B5FD97C0EFF9EFED754DD65D42747" box="[368,456,232,258]" captionStart="FIGURE 30" captionStartId="41.[151,255,1142,1166]" captionTargetBox="[267,1314,183,1127]" captionTargetId="figure@41.[264,1323,183,1135]" captionTargetPageId="41" captionText="FIGURE 30. Collecting sites and geographical distribution of Hydraena species in Palawan and Busuanga, including enlarged map of St. Paul National Park: Hydraena scabra d’Orchymont, H. jojoorculloi sp. n., H. nielshaggei sp. n., H. zetteli sp. n., H. castanescens sp. n., H. claudia sp. n., H. kodadai sp. n." httpUri="https://zenodo.org/record/175808/files/figure.png" pageId="14" pageNumber="15">Fig. 30</figureCitation>
). Known only from three localities in northern and central Palawan.
</paragraph>
<paragraph id="8BAF435C7C0EFF9EFF60553A653F273F" blockId="14.[151,1436,152,458]" pageId="14" pageNumber="15">
<emphasis id="B9649F4E7C0EFF9EFF60553A652C276C" bold="true" box="[199,304,271,297]" pageId="14" pageNumber="15">Ecology.</emphasis>
This species occurs in small and medium-sized creeks draining hilly lowlands with forest cover. The species was collected manually in shallow, slowly flowing, unshaded reaches with sand, gravel, FPOM and CPOM.
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C0EFF9EFF6055B26793278F" pageId="14" pageNumber="15" type="etymology">
<paragraph id="8BAF435C7C0EFF9EFF6055B26793278F" blockId="14.[151,1436,152,458]" pageId="14" pageNumber="15">
<emphasis id="B9649F4E7C0EFF9EFF6055B2654827E4" bold="true" box="[199,340,391,417]" pageId="14" pageNumber="15">Etymology.</emphasis>
Named for Jojo Orcullo (Puerto Princesa City), Palawan artist, for his support during the field work in Palawan and his personal loyalty to the first author.
</paragraph>
</subSubSection>
</treatment>
</document>
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