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<document id="324A96226FE99EF3ED6A2E67761078AA" ID-DOI="10.5281/zenodo.188586" ID-GBIF-Dataset="ddfbe6a3-4ba0-477c-82ba-0190603b6c15" ID-ISSN="1175-5326" ID-Zenodo-Dep="188586" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461095488965" checkinUser="plazi" docAuthor="Bosselaers, Jan" docDate="2009" docId="03B9291AFFD7F16BFF64F65EFD84D17D" docLanguage="en" docName="Bosselaers_2009.imd" docOrigin="Zootaxa 2141" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Agraecina rutilia Simon 1897, new combination" docType="treatment" docVersion="13" lastPageNumber="46" masterDocId="FF805162FFD1F162FFF3F327FF8FD505" masterDocTitle="Studies in Liocranidae (Araneae): redescriptions and transfers in Apostenus Westring and Brachyanillus Simon, as well as description of a new genus" masterLastPageNumber="55" masterPageNumber="37" pageNumber="43" updateTime="1698589017302" updateUser="plazi">
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<mods:title id="9B371073B28D32954AE197B987E3B4B1">Studies in Liocranidae (Araneae): redescriptions and transfers in Apostenus Westring and Brachyanillus Simon, as well as description of a new genus</mods:title>
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<mods:namePart id="AF6A93FBAA1A4289213E65C757961D91">Bosselaers, Jan</mods:namePart>
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<mods:date id="B253094137C8F2C8BD54B8D603013B56">2009</mods:date>
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<mods:classification id="0CCC2B40E89FB6F35D87BD6CF54FE4AB">journal article</mods:classification>
<mods:identifier id="B43E77BB8D3391B3AC744E106F8C595C" type="DOI">10.5281/zenodo.188586</mods:identifier>
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<treatment id="03B9291AFFD7F16BFF64F65EFD84D17D" ID-DOI="http://doi.org/10.5281/zenodo.5576005" ID-GBIF-Taxon="119620241" ID-Zenodo-Dep="5576005" LSID="urn:lsid:plazi:treatment:03B9291AFFD7F16BFF64F65EFD84D17D" httpUri="http://treatment.plazi.org/id/03B9291AFFD7F16BFF64F65EFD84D17D" lastPageId="9" lastPageNumber="46" pageId="6" pageNumber="43">
<subSubSection id="C30ACB87FFD7F164FF64F65EFE13D0B3" pageId="6" pageNumber="43" type="nomenclature">
<paragraph id="8BAF980CFFD7F164FF64F65EFD77D097" blockId="6.[151,760,1400,1462]" box="[151,760,1400,1427]" pageId="6" pageNumber="43">
<heading id="D0E72F60FFD7F164FF64F65EFD77D097" bold="true" box="[151,760,1400,1427]" fontSize="11" level="1" pageId="6" pageNumber="43" reason="1">
<emphasis id="B964441EFFD7F164FF64F65EFD77D097" bold="true" box="[151,760,1400,1427]" pageId="6" pageNumber="43">
<taxonomicName id="4C10E38FFFD7F164FF64F65EFD95D097" ID-CoL="B34F" authority="Simon, 1897" authorityName="Simon" authorityYear="1897" box="[151,538,1400,1427]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="rutilia" status="comb. nov.">
<emphasis id="B964441EFFD7F164FF64F65EFEE8D096" bold="true" box="[151,359,1401,1427]" italics="true" pageId="6" pageNumber="43">Agraecina rutilia</emphasis>
(Simon, 1897)
</taxonomicName>
<taxonomicNameLabel id="A257F965FFD7F164FDD1F65FFD77D097" box="[546,760,1400,1426]" pageId="6" pageNumber="43">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF980CFFD7F164FF64F6BBFE13D0B3" blockId="6.[151,760,1400,1462]" box="[151,412,1436,1462]" pageId="6" pageNumber="43">
<figureCitation id="132B8489FFD7F164FF64F6BBFEA7D0B3" box="[151,296,1436,1462]" captionStart="FIGURE 3 A – B" captionStartId="7.[151,256,1838,1862]" captionTargetBox="[210,1374,209,1799]" captionTargetId="figure@7.[198,1390,197,1813]" captionTargetPageId="7" captionText="FIGURE 3 A – B. Agraecina rutilia. C – G. Cybaeodes liocraninus, holotype. A, epigyne of holotype; B, vulva MRAC 148.466; C, carapace, do; D, carapace, lat; E, sternum and mouth parts, ve; F, carapace, fr; G, leg IV, tarsal tip. Scale bars: A, B, G: 0.1; C – F: 0.5." httpUri="https://zenodo.org/record/188589/files/figure.png" pageId="6" pageNumber="43">Figs. 3 A, B</figureCitation>
; 2 E; 8 C.
</paragraph>
</subSubSection>
<subSubSection id="C30ACB87FFD7F164FF64F6CEFDBDD305" box="[151,562,1513,1536]" pageId="6" pageNumber="43" type="reference_group">
<paragraph id="8BAF980CFFD7F164FF64F6CEFDBDD305" blockId="6.[151,562,1513,1536]" box="[151,562,1513,1536]" pageId="6" pageNumber="43">
<treatmentCitationGroup id="AB00BF22FFD7F164FF64F6CEFDBDD305" box="[151,562,1513,1536]" pageId="6" pageNumber="43">
<taxonomicName id="4C10E38FFFD7F164FF64F6CEFDA2D305" authority="Simon, 1897b: 498" authorityName="Simon" authorityPageNumber="498" authorityYear="1897" box="[151,557,1513,1536]" class="Arachnida" family="Liocranidae" genus="Apostenus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="rutilius">
<emphasis id="B964441EFFD7F164FF64F6CEFED7D305" box="[151,344,1513,1536]" italics="true" pageId="6" pageNumber="43">Apostenus rutilius</emphasis>
<treatmentCitation id="0AB1BE1DFFD7F164FEACF6CEFDA2D305" author="Simon" box="[351,557,1513,1536]" page="498" pageId="6" pageNumber="43" year="1897">
<bibRefCitation id="EF81E5FDFFD7F164FEACF6CEFE77D305" author="Simon" box="[351,504,1513,1536]" pageId="6" pageNumber="54" refString="Simon, E. (1897 b). Etudes arachnologiques. 27 e Memoire. XLII. Descriptions d'especes nouvelles de l'ordre des Araneae. Annales de la Societe entomologique de France 65, 465 - 510." type="journal article" year="1897" yearSuffix="b">Simon, 1897b</bibRefCitation>
: 498
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<paragraph id="8BAF980CFFD7F164FF64F508FB43D34F" blockId="6.[151,1437,1583,2035]" box="[151,1228,1583,1610]" pageId="6" pageNumber="43">
<materialsCitation id="3B789251FFD7F164FF64F508FB46D34F" ID-GBIF-Occurrence="1503373073" box="[151,1225,1583,1610]" collectionCode="MNHN" country="Sierra Leone" location="Sierra Leone" pageId="6" pageNumber="43" specimenCode="8149" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
Type material.
<typeStatus id="54AB26AEFFD7F164FEA7F517FE4ED34F" box="[340,449,1584,1610]" pageId="6" pageNumber="43" type="holotype">Holotype</typeStatus>
female, &quot;
<taxonomicName id="4C10E38FFFD7F164FDC0F517FCE4D34F" authority="E.S. Freetown" authorityName="E.S. Freetown" box="[563,875,1584,1610]" class="Arachnida" family="Liocranidae" genus="Apostenus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="rutilius">
<emphasis id="B964441EFFD7F164FDC0F517FD39D34C" box="[563,694,1584,1609]" italics="true" pageId="6" pageNumber="43">Ap. rutilius</emphasis>
E.S. Freetown
</taxonomicName>
&quot;,
<collectingCountry id="F307D89CFFD7F164FC8CF517FB9AD34F" box="[895,1045,1584,1610]" name="Sierra Leone" pageId="6" pageNumber="43">Sierra Leone</collectingCountry>
[
<collectionCode id="ED0100C9FFD7F164FBD7F517FB0AD34F" LSID="urn:lsid:biocol.org:col:34988" box="[1060,1157,1584,1610]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="43">MNHN</collectionCode>
8149]
</materialsCitation>
.
</paragraph>
<paragraph id="8BAF980CFFD7F164FF36F571FEE7D392" blockId="6.[151,1437,1583,2035]" pageId="6" pageNumber="43">
<materialsCitation id="3B789251FFD7F164FF36F571FEEBD392" ID-GBIF-Occurrence="1503373078" collectingDate="1976-09" collectionCode="MRAC" collectorName="D. Olu-Pitt" country="Sierra Leone" location="Mt. Aureol" municipality="Freetown" pageId="6" pageNumber="43" specimenCode="148.466" specimenCount="1" specimenCount-female="1">
<emphasis id="B964441EFFD7F164FF36F571FD82D375" bold="true" box="[197,525,1622,1648]" pageId="6" pageNumber="43">Other material examined.</emphasis>
1 ♀,
<collectingCountry id="F307D89CFFD7F164FDA0F571FD60D375" box="[595,751,1622,1648]" name="Sierra Leone" pageId="6" pageNumber="43">Sierra Leone</collectingCountry>
, Freetown, Mt. Aureol,
<date id="FFAEBECCFFD7F164FBE2F571FB59D375" box="[1041,1238,1622,1648]" pageId="6" pageNumber="43" value="1976-09">September 1976</date>
, D. Olu-Pitt leg. [
<collectionCode id="ED0100C9FFD7F164FF6EF55AFF76D392" box="[157,249,1661,1687]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="43">MRAC</collectionCode>
148.466]
</materialsCitation>
.
</paragraph>
<subSubSection id="C30ACB87FFD7F164FF36F584FDFDD20E" pageId="6" pageNumber="43" type="diagnosis">
<paragraph id="8BAF980CFFD7F164FF36F584FDFDD20E" blockId="6.[151,1437,1583,2035]" pageId="6" pageNumber="43">
<emphasis id="B964441EFFD7F164FF36F584FEC8D3B8" bold="true" box="[197,327,1699,1725]" pageId="6" pageNumber="43">Diagnosis.</emphasis>
<taxonomicName id="4C10E38FFFD7F164FEBCF583FE33D3B8" box="[335,444,1700,1725]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="rutilia">
<emphasis id="B964441EFFD7F164FEBCF583FE33D3B8" box="[335,444,1700,1725]" italics="true" pageId="6" pageNumber="43">A. rutilia</emphasis>
</taxonomicName>
differs from the other four
<taxonomicName id="4C10E38FFFD7F164FCFAF583FC0AD3B8" box="[777,901,1700,1725]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B964441EFFD7F164FCFAF583FC0AD3B8" box="[777,901,1700,1725]" italics="true" pageId="6" pageNumber="43">Agraecina</emphasis>
</taxonomicName>
species known by the long, parallel, closely adjacent ID which are curled and connected to small, clearly defined subglobular ST1 and ST2 at their posterior and anterior ends, respectively.
</paragraph>
</subSubSection>
<subSubSection id="C30ACB87FFD7F16BFF36F430FBC8D7CA" lastPageId="9" lastPageNumber="46" pageId="6" pageNumber="43" type="description">
<paragraph id="8BAF980CFFD7F164FF36F430FD93D237" blockId="6.[151,1437,1583,2035]" box="[197,540,1815,1842]" pageId="6" pageNumber="43">
<emphasis id="B964441EFFD7F164FF36F430FED2D234" bold="true" box="[197,349,1815,1841]" pageId="6" pageNumber="43">Description.</emphasis>
<collectingCountry id="F307D89CFFD7F164FE97F43FFE2FD237" box="[356,416,1816,1842]" name="Malta" pageId="6" pageNumber="43">Male</collectingCountry>
unknown.
</paragraph>
<paragraph id="8BAF980CFFD7F16BFF36F419FC01D75E" blockId="6.[151,1437,1583,2035]" lastBlockId="9.[151,1437,152,1144]" lastPageId="9" lastPageNumber="46" pageId="6" pageNumber="43">
Female (
<typeStatus id="54AB26AEFFD7F164FEDFF419FE15D25D" box="[300,410,1854,1880]" pageId="6" pageNumber="43" type="holotype">holotype</typeStatus>
). Total length 3.68. Carapace length 1.47,
<geoCoordinate id="EE24FECBFFD7F164FC63F419FC6BD25D" box="[912,996,1854,1880]" direction="west" orientation="longitude" pageId="6" pageNumber="43" precision="555" value="-1.1">w 1.10</geoCoordinate>
, pale ochre with dark striae radiating from fovea. Fovea brown, pronounced, length 0.24, anterior end 0.97 from front end of carapace. Both eye rows procurved from front and recurved from above. AER
<geoCoordinate id="EE24FECBFFD7F164FCBCF4ABFC2DD2A3" box="[847,930,1932,1958]" direction="west" orientation="longitude" pageId="6" pageNumber="43" precision="555" value="-0.32">w 0.32</geoCoordinate>
, PER
<geoCoordinate id="EE24FECBFFD7F164FC1FF4ABFBCFD2A3" box="[1004,1088,1932,1958]" direction="west" orientation="longitude" pageId="6" pageNumber="43" precision="555" value="-0.46">w 0.46</geoCoordinate>
, four straight hairs in front of AER. All eyes subequal, except AME whose diameter is 1/3 of diameter of others. AME separated by a distance equal to their diameter, and separated from ALE by 1/2 of AME diameter. PE separated by slightly less than their diameter, PME slightly closer to PLE than to each other. MOQ depth 0.16, anterior
<geoCoordinate id="EE24FECBFFD9F16AFAB1F4ACFA17D2A0" box="[1346,1432,1931,1957]" direction="west" orientation="longitude" pageId="8" pageNumber="45" precision="555" value="-0.12">w 0.12</geoCoordinate>
, posterior
<geoCoordinate id="EE24FECBFFD9F16AFEF4F496FED5D2CE" box="[263,346,1969,1995]" direction="west" orientation="longitude" pageId="8" pageNumber="45" precision="555" value="-0.24">w 0.24</geoCoordinate>
. Clypeus slightly larger than diameter of AME, chilum absent. Chelicerae yellow, with three teeth along promarginal cheliceral rim, largest one in the middle and smallest one furthest from fang base, and two teeth on retromarginal rim, smallest one closest to fang base. A large curved seta in front of fang base. Sternum yellow, smooth, shield-shaped, length 0.79,
<geoCoordinate id="EE24FECBFFD8F16BFCC2F399FC05D5DD" box="[817,906,190,216]" direction="west" orientation="longitude" pageId="9" pageNumber="46" precision="555" value="-0.71">w 0.71</geoCoordinate>
. No PCT, no ICS, PLB inconspicuous and isolated. Labium slightly wider than long. Endites long, parallel sided, not notched, no apical hair tuft. Serrula present. Abdomen do unicolorous grey, sparsely covered with thin brown hairs, ve pale greyish white. ALS conical, with short terminal article, PMS short, triangular, with three large cylindrical gland spigots, PLS subcylindrical, stout, separated by a distance equal to their length. Trochanters notched, retrocoxal hymen small, hyaline, feathery hairs present on legs. Legs unicolorous yellow, leg formula 4123, leg length I 3.81 II 3.66 III 3.34 IV 4.68. No vt preening brush on mt III or IV. Tarsi without claw tufts or tenent hairs, claws with 5–10 teeth. Leg spination (
<figureCitation id="132B8489FFD8F16BFE22F2EAFDBED4E2" box="[465,561,461,487]" captionStart="FIGURE 2 A – K" captionStartId="4.[151,257,1819,1843]" captionTargetBox="[168,1425,219,1763]" captionTargetId="figure@4.[151,1436,194,1795]" captionTargetPageId="4" captionText="FIGURE 2 A – K. Leg spination schematics. A, legend; B, Apostenus humilis, male holotype; C, Apostenus humilis, female, MNHN 25; D, Apostenus ochraceus, female; E, Agraecina rutilia, female holotype; F, Agraecina lineata, female, MNHN 5653; G, Cybaeodes liocraninus, holotype; H, Cybaeodes testaceus, female holotype, MNHN 3107; I, Cybaeodes madidus, female holotype, MNHN 1677; J, Cybaeodes marinae, female, Italy, Castelporziano, AMNH; K, Arabelia pheidoleicomes, female holotype." httpUri="https://zenodo.org/record/188588/files/figure.png" pageId="9" pageNumber="46">Fig. 2 E</figureCitation>
) fe: I pl 0-0-1 do 1-1-1; II pl 0-0-1 do 1-1-1; III do 1-1-3; IV do 1-1-3; pa: III dt 1; IV dt 1; ti: I plv 1-1-0 rlv 1-1-0; II rlv 1-1-0; III pl 1-1-0 do 1-1-0 rl 1-1-0 plv 1-1-1 rlv 0-1-1; IV pl 1- 1-0 do 1-0-1 rl 1-1-0 plv 1-1-1 rlv 0-1-1; mt: I plv 1-1-0 rlv 1-1-0; II plv 1-1-0 rlv 1-1-0; III pl 0-2-2 rl 0-2-2 plv 1-1-1 rlv 1-1-1; IV pl 1-2-2 rl 1-2-2 plv 1-1-0 rlv 1-1-0 vts 1.
</paragraph>
<caption id="DF6FC884FFD6F165FF64F409FE03D28E" httpUri="https://zenodo.org/record/188589/files/figure.png" pageId="7" pageNumber="44" targetBox="[210,1374,209,1799]" targetPageId="7">
<paragraph id="8BAF980CFFD6F165FF64F409FE03D28E" blockId="7.[151,1437,1838,1931]" pageId="7" pageNumber="44">
<emphasis id="B964441EFFD6F165FF64F409FEDAD243" bold="true" box="[151,341,1838,1862]" pageId="7" pageNumber="44">FIGURE 3 A–B.</emphasis>
<taxonomicName id="4C10E38FFFD6F165FEAEF408FD95D243" box="[349,538,1839,1862]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="rutilia">
<emphasis id="B964441EFFD6F165FEAEF408FD95D243" box="[349,538,1839,1862]" italics="true" pageId="7" pageNumber="44">Agraecina rutilia</emphasis>
</taxonomicName>
.
<emphasis id="B964441EFFD6F165FDDBF409FDD1D243" bold="true" box="[552,606,1838,1862]" pageId="7" pageNumber="44">C–G.</emphasis>
<taxonomicName id="4C10E38FFFD6F165FD9AF408FCEDD243" box="[617,866,1839,1862]" class="Arachnida" family="Liocranidae" genus="Cybaeodes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="liocraninus">
<emphasis id="B964441EFFD6F165FD9AF408FCEDD243" box="[617,866,1839,1862]" italics="true" pageId="7" pageNumber="44">Cybaeodes liocraninus</emphasis>
</taxonomicName>
, holotype. A, epigyne of holotype; B, vulva MRAC 148.466; C, carapace, do; D, carapace, lat; E, sternum and mouth parts, ve; F, carapace, fr; G, leg IV, tarsal tip. Scale bars: A, B, G: 0.1; C–F: 0.5.
</paragraph>
</caption>
<caption id="DF6FC884FFD9F16AFF64F405FB68D259" httpUri="https://zenodo.org/record/188590/files/figure.png" pageId="8" pageNumber="45" targetBox="[155,1429,217,1755]" targetPageId="8">
<paragraph id="8BAF980CFFD9F16AFF64F405FB68D259" blockId="8.[151,1435,1826,1884]" pageId="8" pageNumber="45">
<emphasis id="B964441EFFD9F16AFF64F405FEDED23F" bold="true" box="[151,337,1826,1850]" pageId="8" pageNumber="45">FIGURE 4 A–G.</emphasis>
<taxonomicName id="4C10E38FFFD9F16AFEA8F404FDB5D23F" box="[347,570,1827,1850]" class="Arachnida" family="Liocranidae" genus="Cybaeodes" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="B964441EFFD9F16AFEA8F404FDB5D23F" box="[347,570,1827,1850]" italics="true" pageId="8" pageNumber="45">Cybaeodes testaceus</emphasis>
</taxonomicName>
, holotype. A, carapace, do; B, carapace, lat; C, sternum and mouth parts, ve; D, carapace, fr; E, epigyne; F, leg I, tarsal tip; G, leg III, tarsal tip. Scale bars: A–D: 0.5; E: 0.25; F–G: 0.1.
</paragraph>
</caption>
<paragraph id="8BAF980CFFD8F16BFF36F14FFBC8D7CA" blockId="9.[151,1437,152,1144]" pageId="9" pageNumber="46">
Epigyne (
<figureCitation id="132B8489FFD8F16BFEC5F14FFE15D787" box="[310,410,616,642]" captionStart="FIGURE 3 A – B" captionStartId="7.[151,256,1838,1862]" captionTargetBox="[210,1374,209,1799]" captionTargetId="figure@7.[198,1390,197,1813]" captionTargetPageId="7" captionText="FIGURE 3 A – B. Agraecina rutilia. C – G. Cybaeodes liocraninus, holotype. A, epigyne of holotype; B, vulva MRAC 148.466; C, carapace, do; D, carapace, lat; E, sternum and mouth parts, ve; F, carapace, fr; G, leg IV, tarsal tip. Scale bars: A, B, G: 0.1; C – F: 0.5." httpUri="https://zenodo.org/record/188589/files/figure.png" pageId="9" pageNumber="46">Fig. 3 A</figureCitation>
) only weakly sclerotised, semitransparent, without a clear median septum. CO anterior. Vulva (
<figureCitation id="132B8489FFD8F16BFF1DF1A9FEDED7AD" box="[238,337,654,680]" captionStart="FIGURE 3 A – B" captionStartId="7.[151,256,1838,1862]" captionTargetBox="[210,1374,209,1799]" captionTargetId="figure@7.[198,1390,197,1813]" captionTargetPageId="7" captionText="FIGURE 3 A – B. Agraecina rutilia. C – G. Cybaeodes liocraninus, holotype. A, epigyne of holotype; B, vulva MRAC 148.466; C, carapace, do; D, carapace, lat; E, sternum and mouth parts, ve; F, carapace, fr; G, leg IV, tarsal tip. Scale bars: A, B, G: 0.1; C – F: 0.5." httpUri="https://zenodo.org/record/188589/files/figure.png" pageId="9" pageNumber="46">Fig. 3 B</figureCitation>
) with anterior CO, long, parallel, closely adjacent ID curled at both ends. Anteriorly, the ID are connected to small, subglobular ST2, posteriorly to almost isodiametric ST1.
</paragraph>
</subSubSection>
<subSubSection id="C30ACB87FFD8F16BFF36F1FCFBD4D157" pageId="9" pageNumber="46" type="discussion">
<paragraph id="8BAF980CFFD8F16BFF36F1FCFBD4D157" blockId="9.[151,1437,152,1144]" pageId="9" pageNumber="46">
<emphasis id="B964441EFFD8F16BFF36F1FCFEDBD7F0" bold="true" box="[197,340,731,757]" pageId="9" pageNumber="46">Discussion.</emphasis>
Due to its notched trochanters, long and narrow carapace, ti I, mt I and mt II with two ve spine pairs, ti II with only two rlv spines, as well as its vulvar structure and an epigyne without a clear median septum,
<taxonomicName id="4C10E38FFFD8F16BFF0EF00EFEE5D647" box="[253,362,809,834]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="rutilia">
<emphasis id="B964441EFFD8F16BFF0EF00EFEE5D647" box="[253,362,809,834]" italics="true" pageId="9" pageNumber="46">A. rutilia</emphasis>
</taxonomicName>
does not belong in
<taxonomicName id="4C10E38FFFD8F16BFDAAF00EFD5BD647" box="[601,724,809,834]" class="Arachnida" family="Liocranidae" genus="Apostenus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B964441EFFD8F16BFDAAF00EFD5BD647" box="[601,724,809,834]" italics="true" pageId="9" pageNumber="46">Apostenus</emphasis>
</taxonomicName>
. However, all these characters, especially the leg spination (compare
<figureCitation id="132B8489FFD8F16BFEFEF077FE54D66F" box="[269,475,848,874]" captionStart="FIGURE 2 A – K" captionStartId="4.[151,257,1819,1843]" captionTargetBox="[168,1425,219,1763]" captionTargetId="figure@4.[151,1436,194,1795]" captionTargetPageId="4" captionText="FIGURE 2 A – K. Leg spination schematics. A, legend; B, Apostenus humilis, male holotype; C, Apostenus humilis, female, MNHN 25; D, Apostenus ochraceus, female; E, Agraecina rutilia, female holotype; F, Agraecina lineata, female, MNHN 5653; G, Cybaeodes liocraninus, holotype; H, Cybaeodes testaceus, female holotype, MNHN 3107; I, Cybaeodes madidus, female holotype, MNHN 1677; J, Cybaeodes marinae, female, Italy, Castelporziano, AMNH; K, Arabelia pheidoleicomes, female holotype." httpUri="https://zenodo.org/record/188588/files/figure.png" pageId="9" pageNumber="46">Figs. 2 E and 2 F</figureCitation>
), are compatible with a place in the genus
<taxonomicName id="4C10E38FFFD8F16BFC26F077FBC0D66C" box="[981,1103,848,873]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B964441EFFD8F16BFC26F077FBC0D66C" box="[981,1103,848,873]" italics="true" pageId="9" pageNumber="46">Agraecina</emphasis>
</taxonomicName>
(
<figureCitation id="132B8489FFD8F16BFB93F077FB48D66F" box="[1120,1223,848,874]" captionStart="FIGURE 8 A – H" captionStartId="15.[151,259,1348,1372]" captionTargetBox="[151,1435,313,1323]" captionTargetId="figure@15.[151,1436,313,1324]" captionTargetPageId="15" captionText="FIGURE 8 A – H. Dorsal views of specimens. A, Apostenus humilis, holotype; B, Apostenus ochraceus, Lesbos, Kratigos; C, Agraecina rutilia, holotype; D, Agraecina lineata, female, MNHN 5653; E, Cybaeodes liocraninus, holotype; F, Cybaeodes testaceus, holotype; G, Cybaeodes madidus, holotype; H, Arabelia pheidoleicomes sp. n., Lesbos, Agiasos, Megali Limni. Scale bars: 0.5." httpUri="https://zenodo.org/record/188593/files/figure.png" pageId="9" pageNumber="46">Fig. 8 D</figureCitation>
). Based on vulvar structure,
<taxonomicName id="4C10E38FFFD8F16BFEFEF050FEF6D695" box="[269,377,887,912]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="rutilia">
<emphasis id="B964441EFFD8F16BFEFEF050FEF6D695" box="[269,377,887,912]" italics="true" pageId="9" pageNumber="46">A. rutilia</emphasis>
</taxonomicName>
displays the highest level of vulvar complexity of all known
<taxonomicName id="4C10E38FFFD8F16BFBA0F050FB41D695" box="[1107,1230,887,912]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B964441EFFD8F16BFBA0F050FB41D695" box="[1107,1230,887,912]" italics="true" pageId="9" pageNumber="46">Agraecina</emphasis>
</taxonomicName>
species. One can hypothesise a series starting with the simple vulva of
<taxonomicName id="4C10E38FFFD8F16BFCFCF0BAFC38D6B3" box="[783,951,925,950]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="canariensis">
<emphasis id="B964441EFFD8F16BFCFCF0BAFC38D6B3" box="[783,951,925,950]" italics="true" pageId="9" pageNumber="46">A. canariensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C10E38FFFD8F16BFC02F0BAFBD4D6B3" box="[1009,1115,925,950]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="hodna">
<emphasis id="B964441EFFD8F16BFC02F0BAFBD4D6B3" box="[1009,1115,925,950]" italics="true" pageId="9" pageNumber="46">A. hodna</emphasis>
</taxonomicName>
, where the wide, divergent ID are gradually merging into inconspicuous ST1 and ST2 at their anterior and posterior ends, respectively, resulting in a dumbbell shape (
<bibRefCitation id="EF81E5FDFFD8F16BFDE2F0CDFD64D101" author="Wunderlich" box="[529,747,1002,1028]" pageId="9" pageNumber="55" refString="Wunderlich, J. (1992) The Spider fauna of the Macaronesian Islands. Taxonomy, Ecology, Biogeography and Evolution. Beitrage Zur Araneologie 1, Straubenhardt, 619 pp." type="book" year="1992">Wunderlich 1992</bibRefCitation>
;
<bibRefCitation id="EF81E5FDFFD8F16BFD0BF0CDFC3ED101" author="Bosmans" box="[760,945,1002,1028]" pageId="9" pageNumber="54" refString="Bosmans, R. (1999) The genera Agroeca, Agraecina, Apostenus and Scotina in the Maghreb countries (Araneae: Liocranidae). Bulletin de l' Institut royal des Sciences naturelles de Belgique 69, 25 - 34." type="journal article" year="1999">Bosmans 1999</bibRefCitation>
), leading over the terminally curled and enlarged combination of ID and ST1-ST
<quantity id="4CE835E9FFD8F16BFD83F736FD11D12E" box="[624,670,1041,1067]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="9" pageNumber="46" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4C10E38FFFD8F16BFD56F736FCA7D12F" box="[677,808,1041,1066]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="cristiani">
<emphasis id="B964441EFFD8F16BFD56F736FCA7D12F" box="[677,808,1041,1066]" italics="true" pageId="9" pageNumber="46">A. cristiani</emphasis>
</taxonomicName>
and
<taxonomicName id="4C10E38FFFD8F16BFC92F736FC5DD12F" box="[865,978,1041,1066]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="lineata">
<emphasis id="B964441EFFD8F16BFC92F736FC5DD12F" box="[865,978,1041,1066]" italics="true" pageId="9" pageNumber="46">A. lineata</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF81E5FDFFD8F16BFC11F736FB57D12E" author="Weiss" box="[994,1240,1041,1067]" pageId="9" pageNumber="55" refString="Weiss, I. &amp; Sarbu, S. (1994) Die Hohlenspinne Agraecina cristiani (Georgescu, 1989) n. comb. (Arachnida, Araneae, Liocranidae) Verhandlungen des naturwissenschaftliches Vereins Hamburg, Neue Folge 34, 249 - 257." type="journal article" year="1994">Weiss &amp; Sarbu 1994</bibRefCitation>
;
<bibRefCitation id="EF81E5FDFFD8F16BFB16F736FA1BD12E" author="Bosmans" box="[1253,1428,1041,1067]" pageId="9" pageNumber="54" refString="Bosmans, R. (1999) The genera Agroeca, Agraecina, Apostenus and Scotina in the Maghreb countries (Araneae: Liocranidae). Bulletin de l' Institut royal des Sciences naturelles de Belgique 69, 25 - 34." type="journal article" year="1999">Bosmans 1999</bibRefCitation>
) to the terminally curled ID connected to clearly defined ST1 and ST2 of
<taxonomicName id="4C10E38FFFD8F16BFC18F71FFBDBD154" box="[1003,1108,1080,1105]" class="Arachnida" family="Liocranidae" genus="Agraecina" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="rutilia">
<emphasis id="B964441EFFD8F16BFC18F71FFBDBD154" box="[1003,1108,1080,1105]" italics="true" pageId="9" pageNumber="46">A. rutilia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30ACB87FFD8F16BFF36F779FD84D17D" box="[197,523,1118,1144]" pageId="9" pageNumber="46" type="distribution">
<paragraph id="8BAF980CFFD8F16BFF36F779FD84D17D" blockId="9.[151,1437,152,1144]" box="[197,523,1118,1144]" pageId="9" pageNumber="46">
<emphasis id="B964441EFFD8F16BFF36F779FEEAD17D" bold="true" box="[197,357,1118,1144]" pageId="9" pageNumber="46">Distribution.</emphasis>
<collectingCountry id="F307D89CFFD8F16BFE9FF779FD88D17D" box="[364,519,1118,1144]" name="Sierra Leone" pageId="9" pageNumber="46">Sierra Leone</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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