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<document id="CB03E2677BB3261D6335596D0D61457E" ID-DOI="10.11646/zootaxa.5567.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14703735" ID-ZooBank="AF6254F6-CE26-41FE-BB85-A3FFD16FAA39" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737371447294" checkinUser="plazi" docAuthor="VIRAKTAMATH, C. A., WEBB, M. D. &amp; YESHWANTH, H. M." docDate="2025" docId="03B75F43F06CFFB9FF487530321624EC" docLanguage="en" docName="zootaxa.5567.1.1.pdf" docOrigin="Zootaxa 5567 (1)" docSource="https://doi.org/10.11646/zootaxa.5567.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Panghoella maculosa VIRAKTAMATH &amp; WEBB &amp; YESHWANTH 2025, comb. nov." docType="treatment" docVersion="2" lastPageNumber="38" masterDocId="FF8E273BF049FF9CFFDF7756364D2343" masterDocTitle="Review of the Monobazus-group leafhopper genera (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of five new genera and twenty-eight new species" masterLastPageNumber="105" masterPageNumber="1" pageNumber="38" updateTime="1737389185710" updateUser="ExternalLinkService">
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<mods:title id="D11BF37062C0FD4C0ED420C4F611BE5B">Review of the Monobazus-group leafhopper genera (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of five new genera and twenty-eight new species</mods:title>
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<mods:namePart id="1753FA654DE99B48AD23E69E18ACF331">VIRAKTAMATH, C. A.</mods:namePart>
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<mods:namePart id="764110BD05D70A768E223D33CEB04232">WEBB, M. D.</mods:namePart>
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<subSubSection id="C304BDDEF06CFFB9FF48753034EC21C2" box="[151,673,614,641]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="8BA1EE55F06CFFB9FF48753034EC21C2" blockId="37.[151,698,614,677]" box="[151,673,614,641]" pageId="37" pageNumber="38">
<heading id="D0E95939F06CFFB9FF48753034EC21C2" bold="true" box="[151,673,614,641]" fontSize="11" level="2" pageId="37" pageNumber="38" reason="4">
<emphasis id="B96A3247F06CFFB9FF48753034EC21C2" bold="true" box="[151,673,614,641]" pageId="37" pageNumber="38">
<taxonomicName id="4C1E95D6F06CFFB9FF487530345E21C3" authority="(Distant)" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" baseAuthorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" baseAuthorityYear="2025" box="[151,531,614,641]" class="Insecta" family="Cicadellidae" genus="Panghoella" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="maculosa" status="comb. nov.">
<emphasis id="B96A3247F06CFFB9FF48753037D021C2" bold="true" box="[151,413,614,641]" italics="true" pageId="37" pageNumber="38">Panghoella maculosa</emphasis>
(Distant)
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<taxonomicNameLabel id="A2598F3CF06CFFB9FDC5753134EC21C2" box="[538,673,615,641]" pageId="37" pageNumber="38" rank="species">comb. nov.</taxonomicNameLabel>
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<subSubSection id="C304BDDEF06CFFB9FF4875DD34F721E6" box="[151,698,650,677]" pageId="37" pageNumber="38" type="description">
<paragraph id="8BA1EE55F06CFFB9FF4875DD34F721E6" blockId="37.[151,698,614,677]" box="[151,698,650,677]" pageId="37" pageNumber="38">
<figureCitation id="1325F2D0F06CFFB9FF4875DD375B21E6" box="[151,278,651,677]" captionStart="FIGURES 6" captionStartId="47.[151,264,1889,1913]" captionTargetBox="[156,1423,181,1860]" captionTargetId="figure-17@47.[151,1435,181,1865]" captionTargetPageId="47" captionText="FIGURES 6AR. Monobazus group of genera, habitus, dorsal and lateral view. AD, Panghoella maculosa (Distant) male (AB) and female (CD); EF, Phlogotettix indicus Rao, male; GJ, P. subhimalayanus Meshram &amp; Ramamurthy, male (GH), and female (IJ); KL, Phlogothamnus maculiceps Ishihara, male; MN, P. umiamensis sp. nov. male; OR, Wanritettix wanrianus (Matsumura), male (OP), and female (QR)." figureDoi="http://doi.org/10.5281/zenodo.14703761" httpUri="https://zenodo.org/record/14703761/files/figure.png" pageId="37" pageNumber="38">Figs 6AD</figureCitation>
,
<figureCitation id="1325F2D0F06CFFB9FEFE75DD377021E6" box="[289,317,651,677]" captionStart="FIGURES 10" captionStartId="51.[151,264,1761,1785]" captionTargetBox="[156,1421,186,1732]" captionTargetId="figure-17@51.[151,1435,181,1737]" captionTargetPageId="51" captionText="FIGURES 10AQ. Monobazus group of genera, faces. AB. Monobazus sikkimensis sp. nov., male (A), female (B); CD, M. sahyadricus sp. nov., male (C), female (D); E, M. sudindicus sp. nov., male; F, Panghoella sinuata sp. nov., male; GH, P. maculosa (Distant), male (G), female (H); I, Phlogotettix indicus Rao, male; J, P. subhimalayanus Meshram &amp; Ramamurthy, male; KL, Phlogothamnus maculiceps Ishihara, male (K), female (L); M, P. umiamensis sp. nov., male; NO, Wanritettix wanrianus (Matsumura), male (N), female (O); PQ, Longicornus grandis (Pruthi), male (P), female (Q)." figureDoi="http://doi.org/10.5281/zenodo.14703773" httpUri="https://zenodo.org/record/14703773/files/figure.png" pageId="37" pageNumber="38">10</figureCitation>
GH, 11I, 13MP, 47AM, 58DG
</paragraph>
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<subSubSection id="C304BDDEF06CFFB9FF487586346921A4" box="[151,548,719,743]" pageId="37" pageNumber="38" type="reference_group">
<paragraph id="8BA1EE55F06CFFB9FF487586346921A4" blockId="37.[151,548,719,743]" box="[151,548,719,743]" pageId="37" pageNumber="38">
<treatmentCitationGroup id="AB0EC97BF06CFFB9FF487586346921A4" box="[151,548,719,743]" pageId="37" pageNumber="38">
<treatmentCitation id="0ABFC844F06CFFB9FF487586346C21A4" author="Distant, W. L." box="[151,545,719,743]" page="56" pageId="37" pageNumber="38" year="1918">
<taxonomicName id="4C1E95D6F06CFFB9FF487586346C21A4" ID-CoL="KLF6" authority="Distant 1918: 55 - 56" authorityName="Distant" authorityPageNumber="55 - 56" authorityYear="1918" box="[151,545,719,743]" class="Insecta" family="Cicadellidae" genus="Banus" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="maculosus">
<emphasis id="B96A3247F06CFFB9FF487586370521A4" box="[151,328,719,743]" italics="true" pageId="37" pageNumber="38">Banus maculosus</emphasis>
<bibRefCitation id="EF8F93A4F06CFFB9FE917599346C21A4" author="Distant, W. L." box="[334,545,719,743]" pageId="37" pageNumber="38" refId="ref49986" refString="Distant, W. L. (1918) Rhynchota - Homoptera: Appendix. Heteroptera: Addenda. In: Shipley, A. E. &amp; Marshall, G. A. K. (Eds.), The Fauna of British India including Ceylon and Burma. Vol. 7. Taylor &amp; Francis, London, viii + 210 pp." type="book" year="1918">Distant 1918: 5556</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
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<subSubSection id="C304BDDEF06CFFB9FF487440348B2594" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="8BA1EE55F06CFFB9FF487440359D20FC" blockId="37.[151,1437,789,1968]" pageId="37" pageNumber="38">
<emphasis id="B96A3247F06CFFB9FF4874403744206C" box="[151,265,790,815]" italics="true" pageId="37" pageNumber="38">Diagnosis</emphasis>
. The species may be identified by the following combination of characters. Male pygofer with dorsal marginal process directed ventrally, not attaining ventral margin. Subgenital plates without macrosetae but with narrow hair-like setae. Aedeagal shaft curved, narrowed apically, apex in ventral view either flared (
<figureCitation id="1325F2D0F06CFFB9FB2F740B33012034" box="[1264,1356,861,887]" captionStart="FIGURES 47" captionStartId="88.[151,264,1859,1883]" captionTargetBox="[163,1426,186,1830]" captionTargetId="figure-26@88.[158,1430,181,1835]" captionTargetPageId="88" captionText="FIGURES 47AM. Panghoella maculosa (Distant), male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; DE, style, dorsal view and magnified view of apophysis; F, aedeagus ventral view; G, connective and aedeagus, lateral view; HJ, syntype male from Kurseong: H, aedeagus lateral view; I, apex of aedeagal shaft lateral view magnified; J, aedeagus, ventral view; male from Nepal: KL aedeagus, lateral and ventral view, M, style dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14703873" httpUri="https://zenodo.org/record/14703873/files/figure.png" pageId="37" pageNumber="38">Fig. 47J</figureCitation>
) or not (
<figureCitation id="1325F2D0F06CFFB9FF4074D736BB20D8" box="[159,246,897,923]" captionStart="FIGURES 47" captionStartId="88.[151,264,1859,1883]" captionTargetBox="[163,1426,186,1830]" captionTargetId="figure-26@88.[158,1430,181,1835]" captionTargetPageId="88" captionText="FIGURES 47AM. Panghoella maculosa (Distant), male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; DE, style, dorsal view and magnified view of apophysis; F, aedeagus ventral view; G, connective and aedeagus, lateral view; HJ, syntype male from Kurseong: H, aedeagus lateral view; I, apex of aedeagal shaft lateral view magnified; J, aedeagus, ventral view; male from Nepal: KL aedeagus, lateral and ventral view, M, style dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14703873" httpUri="https://zenodo.org/record/14703873/files/figure.png" pageId="37" pageNumber="38">Figs 47</figureCitation>
FL), gonopore apical on ventral surface, basal processes articulated with base of atrium and longer (
<figureCitation id="1325F2D0F06CFFB9FAB574D736FE20FC" captionStart="FIGURES 47" captionStartId="88.[151,264,1859,1883]" captionTargetBox="[163,1426,186,1830]" captionTargetId="figure-26@88.[158,1430,181,1835]" captionTargetPageId="88" captionText="FIGURES 47AM. Panghoella maculosa (Distant), male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; DE, style, dorsal view and magnified view of apophysis; F, aedeagus ventral view; G, connective and aedeagus, lateral view; HJ, syntype male from Kurseong: H, aedeagus lateral view; I, apex of aedeagal shaft lateral view magnified; J, aedeagus, ventral view; male from Nepal: KL aedeagus, lateral and ventral view, M, style dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14703873" httpUri="https://zenodo.org/record/14703873/files/figure.png" pageId="37" pageNumber="38">Figs 47</figureCitation>
FGH) or shorter than shaft (
<figureCitation id="1325F2D0F06CFFB9FE3474F3340C20FC" box="[491,577,933,959]" captionStart="FIGURES 47" captionStartId="88.[151,264,1859,1883]" captionTargetBox="[163,1426,186,1830]" captionTargetId="figure-26@88.[158,1430,181,1835]" captionTargetPageId="88" captionText="FIGURES 47AM. Panghoella maculosa (Distant), male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; DE, style, dorsal view and magnified view of apophysis; F, aedeagus ventral view; G, connective and aedeagus, lateral view; HJ, syntype male from Kurseong: H, aedeagus lateral view; I, apex of aedeagal shaft lateral view magnified; J, aedeagus, ventral view; male from Nepal: KL aedeagus, lateral and ventral view, M, style dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14703873" httpUri="https://zenodo.org/record/14703873/files/figure.png" pageId="37" pageNumber="38">Figs 47</figureCitation>
KL), atrium not produced ventrally.
</paragraph>
<paragraph id="8BA1EE55F06CFFB9FF18749C371527D4" blockId="37.[151,1437,789,1968]" pageId="37" pageNumber="38">
<emphasis id="B96A3247F06CFFB9FF18749C370620A0" box="[199,331,970,995]" italics="true" pageId="37" pageNumber="38">Description</emphasis>
. Pale ochraceous.Crown with brown transverse markings enclosing pale areas (
<figureCitation id="1325F2D0F06CFFB9FB6D749F32BB20A0" box="[1202,1270,969,995]" captionStart="FIGURES 6" captionStartId="47.[151,264,1889,1913]" captionTargetBox="[156,1423,181,1860]" captionTargetId="figure-17@47.[151,1435,181,1865]" captionTargetPageId="47" captionText="FIGURES 6AR. Monobazus group of genera, habitus, dorsal and lateral view. AD, Panghoella maculosa (Distant) male (AB) and female (CD); EF, Phlogotettix indicus Rao, male; GJ, P. subhimalayanus Meshram &amp; Ramamurthy, male (GH), and female (IJ); KL, Phlogothamnus maculiceps Ishihara, male; MN, P. umiamensis sp. nov. male; OR, Wanritettix wanrianus (Matsumura), male (OP), and female (QR)." figureDoi="http://doi.org/10.5281/zenodo.14703761" httpUri="https://zenodo.org/record/14703761/files/figure.png" pageId="37" pageNumber="38">Figs 6</figureCitation>
AC). Face with one spot dorsad of antennal base, lateral margins of frontoclypeus, one spot on each side of base of clypellus, brown. Pronotum with anterior submargin piceous stripe sometimes broken into transverse spots, Exposed mesonotum with basal triangles brown in darker specimens. Forewing venation brown with dark brown spots on apices of claval veins, claval apex and veins reaching costa in apical half. Legs ochraceous with bases of setae on meta tibiae spotted with dark brown.
</paragraph>
<paragraph id="8BA1EE55F06CFFB9FF1873F732D0279C" blockId="37.[151,1437,789,1968]" pageId="37" pageNumber="38">Crown 0.5× as long as interocular distance, subconically rounded. Pronotum 2.02.1× as wide as long medially. Female sternite VII about 1.7× as wide as long, posterior margin concave with median lobe.</paragraph>
<paragraph id="8BA1EE55F06CFFB9FF1873BF33412698" blockId="37.[151,1437,789,1968]" pageId="37" pageNumber="38">
<emphasis id="B96A3247F06CFFB9FF1873BF373D2640" box="[199,368,1257,1283]" italics="true" pageId="37" pageNumber="38">Male genitalia</emphasis>
. Pygofer with anterior marginal apodeme, 1.2× as long as wide in lateral view, posteriorly subconically rounded, posterior margin with short row of setae near ventral margin, dorsal process directed ventrally and not attaining ventral margin, apex not bifid. Subgenital plate 3.0× as long as basal width, without macrosetae but with marginal long, narrow hair-like setae. Aedeagus with dorsal apodeme half as long as shaft, shaft curved anteriorly in distal 0.33, apex in lateral view narrowed, apex in ventral view either flared (
<figureCitation id="1325F2D0F06CFFB9FB79722F334926D0" box="[1190,1284,1401,1427]" captionStart="FIGURES 47" captionStartId="88.[151,264,1859,1883]" captionTargetBox="[163,1426,186,1830]" captionTargetId="figure-26@88.[158,1430,181,1835]" captionTargetPageId="88" captionText="FIGURES 47AM. Panghoella maculosa (Distant), male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; DE, style, dorsal view and magnified view of apophysis; F, aedeagus ventral view; G, connective and aedeagus, lateral view; HJ, syntype male from Kurseong: H, aedeagus lateral view; I, apex of aedeagal shaft lateral view magnified; J, aedeagus, ventral view; male from Nepal: KL aedeagus, lateral and ventral view, M, style dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14703873" httpUri="https://zenodo.org/record/14703873/files/figure.png" pageId="37" pageNumber="38">Fig. 47J</figureCitation>
) or not (
<figureCitation id="1325F2D0F06CFFB9FAB5722F36F926F4" captionStart="FIGURES 47" captionStartId="88.[151,264,1859,1883]" captionTargetBox="[163,1426,186,1830]" captionTargetId="figure-26@88.[158,1430,181,1835]" captionTargetPageId="88" captionText="FIGURES 47AM. Panghoella maculosa (Distant), male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; DE, style, dorsal view and magnified view of apophysis; F, aedeagus ventral view; G, connective and aedeagus, lateral view; HJ, syntype male from Kurseong: H, aedeagus lateral view; I, apex of aedeagal shaft lateral view magnified; J, aedeagus, ventral view; male from Nepal: KL aedeagus, lateral and ventral view, M, style dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14703873" httpUri="https://zenodo.org/record/14703873/files/figure.png" pageId="37" pageNumber="38">Figs 47</figureCitation>
FL); gonopore apical on ventral surface; basal processes arising at base of atrium ventrally, articulated and exceed shaft length; and longer (
<figureCitation id="1325F2D0F06CFFB9FE707297344E2698" box="[431,515,1473,1499]" captionStart="FIGURES 47" captionStartId="88.[151,264,1859,1883]" captionTargetBox="[163,1426,186,1830]" captionTargetId="figure-26@88.[158,1430,181,1835]" captionTargetPageId="88" captionText="FIGURES 47AM. Panghoella maculosa (Distant), male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; DE, style, dorsal view and magnified view of apophysis; F, aedeagus ventral view; G, connective and aedeagus, lateral view; HJ, syntype male from Kurseong: H, aedeagus lateral view; I, apex of aedeagal shaft lateral view magnified; J, aedeagus, ventral view; male from Nepal: KL aedeagus, lateral and ventral view, M, style dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14703873" httpUri="https://zenodo.org/record/14703873/files/figure.png" pageId="37" pageNumber="38">Figs 47</figureCitation>
FG) or shorter than shaft (
<figureCitation id="1325F2D0F06CFFB9FCF7729735302698" box="[808,893,1473,1499]" captionStart="FIGURES 47" captionStartId="88.[151,264,1859,1883]" captionTargetBox="[163,1426,186,1830]" captionTargetId="figure-26@88.[158,1430,181,1835]" captionTargetPageId="88" captionText="FIGURES 47AM. Panghoella maculosa (Distant), male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; DE, style, dorsal view and magnified view of apophysis; F, aedeagus ventral view; G, connective and aedeagus, lateral view; HJ, syntype male from Kurseong: H, aedeagus lateral view; I, apex of aedeagal shaft lateral view magnified; J, aedeagus, ventral view; male from Nepal: KL aedeagus, lateral and ventral view, M, style dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14703873" httpUri="https://zenodo.org/record/14703873/files/figure.png" pageId="37" pageNumber="38">Figs 47</figureCitation>
KL), atrium not produced ventrally.
</paragraph>
<paragraph id="8BA1EE55F06CFFB9FF1872B03767250B" blockId="37.[151,1437,789,1968]" pageId="37" pageNumber="38">
<emphasis id="B96A3247F06CFFB9FF1872B0372426BC" box="[199,361,1510,1535]" italics="true" pageId="37" pageNumber="38">Measurements</emphasis>
. Male
<quantity id="4CE643B0F06CFFB9FE6472B3345E2543" box="[443,531,1509,1536]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="37" pageNumber="38" unit="mm" value="5.9">5.9 mm</quantity>
long,
<quantity id="4CE643B0F06CFFB9FD8372B334A82543" box="[604,741,1509,1536]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="37" pageNumber="38" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.31.4 mm</quantity>
wide across eyes and
<quantity id="4CE643B0F06CFFB9FC3972B332232543" box="[998,1134,1509,1536]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="37" pageNumber="38" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.41.5 mm</quantity>
wide across posterolateral angels of pronotum. Female
<quantity id="4CE643B0F06CFFB9FE04715F347C2567" box="[475,561,1545,1572]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="37" pageNumber="38" unit="mm" value="6.1">6.1 mm</quantity>
long,
<quantity id="4CE643B0F06CFFB9FDA6715F34822567" box="[633,719,1545,1572]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="37" pageNumber="38" unit="mm" value="1.4">1.4 mm</quantity>
wide across eyes and
<quantity id="4CE643B0F06CFFB9FC14715F326F2567" box="[971,1058,1545,1572]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="37" pageNumber="38" unit="mm" value="1.6">1.6 mm</quantity>
wide across posterolateral angels of pronotum.
</paragraph>
<paragraph id="8BA1EE55F06CFFB9FF187107348B2594" blockId="37.[151,1437,789,1968]" pageId="37" pageNumber="38">
<emphasis id="B96A3247F06CFFB9FF18710737D22528" box="[199,415,1617,1643]" italics="true" pageId="37" pageNumber="38">Material examined</emphasis>
.
<materialsCitation id="3B76E408F06CFFB9FE6F7104344725CC" collectionCode="BMNH" collectorName="E. Himalayas" country="India" elevation="1478" location="INDIA" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" typeStatus="syntype">
<collectingCountry id="F309AEC5F06CFFB9FE6F71043449252F" box="[432,516,1618,1644]" name="India" pageId="37" pageNumber="38">INDIA</collectingCountry>
:
<typeStatus id="54A550F7F06CFFB9FDCD71043424252F" box="[530,617,1618,1644]" pageId="37" pageNumber="38" type="syntype">syntype</typeStatus>
<specimenCount id="9D1825DCF06CFFB9FDAC710434C7252F" box="[627,650,1618,1644]" count="1" pageId="37" pageNumber="38" type="male"></specimenCount>
, “
<collectorName id="26EB8B83F06CFFB9FD7A7107350E2528" box="[677,835,1617,1643]" pageId="37" pageNumber="38">E. Himalayas</collectorName>
, Kurseong, alt.
<quantity id="4CE643B0F06CFFB9FC22710732D42528" box="[1021,1177,1617,1644]" metricMagnitude="3" metricUnit="m" metricValue="1.47828" metricValueMax="1.524" metricValueMin="1.43256" pageId="37" pageNumber="38" unit="ft" value="4850.0" valueMax="5000.0" valueMin="4700.0">
<elevation id="00330966F06CFFB9FC22710732D42528" box="[1021,1177,1617,1644]" metricMagnitude="3" metricUnit="m" metricValue="1.47828" metricValueMax="1.524" metricValueMin="1.43256" pageId="37" pageNumber="38" unit="ft" value="4850.0" valueMax="5000.0" valueMin="4700.0">47005000 ft</elevation>
</quantity>
, 28.vi 10. Annandale” “NHMUK 013590030” (
<collectionCode id="ED0F7690F06CFFB9FE777123344F25CC" box="[424,514,1653,1679]" pageId="37" pageNumber="38">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B76E408F06CFFB9FDCC7123324B25CC" box="[531,1030,1653,1680]" collectionCode="BMNH" collectorName="Simla" country="India" pageId="37" pageNumber="38" specimenCount="1" typeStatus="syntype">
<specimenCount id="9D1825DCF06CFFB9FDCC712334C025CC" box="[531,653,1653,1679]" count="1" pageId="37" pageNumber="38" type="generic">1specimen</specimenCount>
,
<collectorName id="26EB8B83F06CFFB9FD4A7123349425CC" box="[661,729,1653,1679]" pageId="37" pageNumber="38">Simla</collectorName>
, c. 7000, 12.v.08 (
<collectionCode id="ED0F7690F06CFFB9FC7B712335B325CC" box="[932,1022,1653,1679]" pageId="37" pageNumber="38">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B76E408F06CFFB9FBD0712334DA25F0" collectionCode="BMNH" collectorName="Gopaldhara &amp; Darjeeling &amp; H. Stevens" country="India" elevation="1859" latitude="4.72" longLatPrecision="55" longitude="-6.1" pageId="37" pageNumber="38" specimenCount="1" typeStatus="syntype">
<specimenCount id="9D1825DCF06CFFB9FBD0712332C425CC" box="[1039,1161,1653,1679]" count="1" pageId="37" pageNumber="38" type="generic">1specimen</specimenCount>
,
<collectorName id="26EB8B83F06CFFB9FB4E7123335A25CC" box="[1169,1303,1653,1679]" pageId="37" pageNumber="38">Gopaldhara</collectorName>
,
<collectorName id="26EB8B83F06CFFB9FAC0712333D525CC" box="[1311,1432,1653,1679]" pageId="37" pageNumber="38">Darjeeling</collectorName>
,
<geoCoordinate id="EE2A8892F06CFFB9FF4871CF36A925F0" box="[151,228,1689,1715]" degrees="4,7200" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="5" value="4.72">4,7200</geoCoordinate>
<geoCoordinate id="EE2A8892F06CFFB9FF3A71CF377C25F0" box="[229,305,1689,1715]" degrees="6,100" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-6.1">
<quantity id="4CE643B0F06CFFB9FF2C71CF377C25F0" box="[243,305,1689,1715]" metricMagnitude="3" metricUnit="m" metricValue="1.85928" pageId="37" pageNumber="38" unit="ft" value="6100.0">
<elevation id="00330966F06CFFB9FF2C71CF377C25F0" box="[243,305,1689,1715]" metricMagnitude="3" metricUnit="m" metricValue="1.85928" pageId="37" pageNumber="38" unit="ft" value="6100.0">6,100</elevation>
</quantity>
</geoCoordinate>
ft,
<collectorName id="26EB8B83F06CFFB9FE8271CC379125F0" box="[349,476,1689,1716]" pageId="37" pageNumber="38">H. Stevens</collectorName>
, ix.17 (
<collectionCode id="ED0F7690F06CFFB9FDEA71CF34C225F0" box="[565,655,1689,1715]" pageId="37" pageNumber="38">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B76E408F06CFFB9FD7B71CC36AB2594" collectingDate="1967-07-15" collectorName="Ktmd &amp; Godavari &amp; Can Nepal Exped" country="Nepal" location="NEPAL" pageId="37" pageNumber="38" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="syntype">
<collectingCountry id="F309AEC5F06CFFB9FD7B71CC354D25F7" box="[676,768,1690,1716]" name="Nepal" pageId="37" pageNumber="38">NEPAL</collectingCountry>
:
<specimenCount id="9D1825DCF06CFFB9FCD171CF357E25F0" box="[782,819,1689,1715]" count="2" pageId="37" pageNumber="38" type="male">2♂</specimenCount>
,
<specimenCount id="9D1825DCF06CFFB9FC9E71CF352925F0" box="[833,868,1689,1715]" count="1" pageId="37" pageNumber="38" type="female">1♀</specimenCount>
,
<collectorName id="26EB8B83F06CFFB9FCAA71CF35F425F0" box="[885,953,1689,1715]" pageId="37" pageNumber="38">Ktmd</collectorName>
:
<collectorName id="26EB8B83F06CFFB9FC1871CF327F25F0" box="[967,1074,1689,1715]" pageId="37" pageNumber="38">Godavari</collectorName>
, 6000,
<date id="FFA0C895F06CFFB9FB4D71CF335925F7" box="[1170,1300,1689,1716]" pageId="37" pageNumber="38" value="1967-07-15">
<collectingDate id="EFE4317DF06CFFB9FB4D71CF335925F7" box="[1170,1300,1689,1716]" pageId="37" pageNumber="38" value="1967-07-15">15.vii.1967</collectingDate>
</date>
,
<collectorName id="26EB8B83F06CFFB9FAFD71CF36AB2594" pageId="37" pageNumber="38">Can Nepal Exped</collectorName>
</materialsCitation>
.;
<materialsCitation id="3B76E408F06CFFB9FF2A71EB348F2594" box="[245,706,1725,1752]" collectingDate="1967-07-23" collectionCode="CNC" collectorName="Ktmd &amp; Godavari &amp; Can Nepal Exped" country="Nepal" location="NEPAL" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" typeStatus="syntype">
<specimenCount id="9D1825DCF06CFFB9FF2A71EB37542594" box="[245,281,1725,1751]" count="1" pageId="37" pageNumber="38" type="male">1♂</specimenCount>
, same data except
<date id="FFA0C895F06CFFB9FE3371EB34262594" box="[492,619,1725,1751]" pageId="37" pageNumber="38" value="1967-07-23">
<collectingDate id="EFE4317DF06CFFB9FE3371EB34262594" box="[492,619,1725,1751]" pageId="37" pageNumber="38" value="1967-07-23">23.vii.1967</collectingDate>
</date>
(
<collectionCode id="ED0F7690F06CFFB9FDA571EB34F42594" box="[634,697,1725,1751]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="37" pageNumber="38">CNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C304BDDEF06CFFB9FF1871B7321624EC" pageId="37" pageNumber="38" type="discussion">
<paragraph id="8BA1EE55F06CFFB9FF1871B7321624EC" blockId="37.[151,1437,789,1968]" pageId="37" pageNumber="38">
<emphasis id="B96A3247F06CFFB9FF1871B7377925B8" bold="true" box="[199,308,1761,1787]" pageId="37" pageNumber="38">Remarks</emphasis>
. This species was described from an unspecified number of specimens (syntypic) with data: “E. Himalayas; Kurseong (
<emphasis id="B96A3247F06CFFB9FE4070533455245C" box="[415,536,1797,1823]" italics="true" pageId="37" pageNumber="38">Annandale</emphasis>
)”. The aedeagal apex of the
<typeStatus id="54A550F7F06CFFB9FCB9705035DB2463" box="[870,918,1798,1824]" pageId="37" pageNumber="38">type</typeStatus>
differs slightly in lateral view from the male examined in having its anterior margin slightly expanded (
<figureCitation id="1325F2D0F06CFFB9FCE1707F35DC2400" box="[830,913,1833,1859]" captionStart="FIGURES 47" captionStartId="88.[151,264,1859,1883]" captionTargetBox="[163,1426,186,1830]" captionTargetId="figure-26@88.[158,1430,181,1835]" captionTargetPageId="88" captionText="FIGURES 47AM. Panghoella maculosa (Distant), male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; DE, style, dorsal view and magnified view of apophysis; F, aedeagus ventral view; G, connective and aedeagus, lateral view; HJ, syntype male from Kurseong: H, aedeagus lateral view; I, apex of aedeagal shaft lateral view magnified; J, aedeagus, ventral view; male from Nepal: KL aedeagus, lateral and ventral view, M, style dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14703873" httpUri="https://zenodo.org/record/14703873/files/figure.png" pageId="37" pageNumber="38">Fig. 47</figureCitation>
HI) rather than tapered (
<figureCitation id="1325F2D0F06CFFB9FB73707F32B32400" box="[1196,1278,1833,1859]" captionStart="FIGURES 47" captionStartId="88.[151,264,1859,1883]" captionTargetBox="[163,1426,186,1830]" captionTargetId="figure-26@88.[158,1430,181,1835]" captionTargetPageId="88" captionText="FIGURES 47AM. Panghoella maculosa (Distant), male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; DE, style, dorsal view and magnified view of apophysis; F, aedeagus ventral view; G, connective and aedeagus, lateral view; HJ, syntype male from Kurseong: H, aedeagus lateral view; I, apex of aedeagal shaft lateral view magnified; J, aedeagus, ventral view; male from Nepal: KL aedeagus, lateral and ventral view, M, style dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14703873" httpUri="https://zenodo.org/record/14703873/files/figure.png" pageId="37" pageNumber="38">Fig. 47</figureCitation>
FG). The two BMNH non-types are currently mislaid after return from loan. This species may prove to be a species complex considering the variability in the curvature of aedeagal shaft in lateral view, apex of shaft and relative length of basal processes and shaft. One segregate of this complex is described as new species below.
</paragraph>
</subSubSection>
</treatment>
</document>