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<document id="BE84A24A001D5EF293F171FEEA14B033" ID-DOI="10.5281/zenodo.196907" ID-GBIF-Dataset="75a8fe9a-5028-4a6a-ba3c-aa404344d072" ID-ISSN="1175-5326" ID-Zenodo-Dep="196907" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460257646372" checkinUser="plazi" docAuthor="Komai, Tomoyuki, Higashiji, Takuo &amp; Castro, Peter" docDate="2010" docId="03B710750315CD4CFF154EEEFB9202F9" docLanguage="en" docName="zt02555p068.pdf" docOrigin="Zootaxa 2555" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hexagonaloides bathyalis Komai, Higashiji &amp; Castro, 2010, n. sp." docType="treatment" docVersion="10" lastPageNumber="67" masterDocId="FF8E680D0317CD49FF824E79FFC2042B" masterDocTitle="A new genus and new species of deep-water trapeziid crab (Crustacea: Decapoda: Brachyura: Trapezioidea) from the Ryukyu Islands, Japan" masterLastPageNumber="68" masterPageNumber="62" pageNumber="64" updateTime="1698243741369" updateUser="plazi">
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<mods:title id="078D3513D9A85F5B56680AD2F717835A">A new genus and new species of deep-water trapeziid crab (Crustacea: Decapoda: Brachyura: Trapezioidea) from the Ryukyu Islands, Japan</mods:title>
</mods:titleInfo>
<mods:name id="A736736FB652AA49D06F70F123D723D1" type="personal">
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<mods:namePart id="12F1BBC424E6677E4D5356BB876AFDC7">Komai, Tomoyuki</mods:namePart>
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<mods:name id="CED8FA4337DC9BF3085EE7D2ADF8C3B9" type="personal">
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<mods:roleTerm id="AA402F266A5F062BF6120501ACCEB1CB">Author</mods:roleTerm>
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<mods:namePart id="6BE8354445809CDF7E07125A438A6B86">Higashiji, Takuo</mods:namePart>
</mods:name>
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<mods:namePart id="AFC6908DA9E4117B53B73A7833DD4CB5">Castro, Peter</mods:namePart>
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<mods:typeOfResource id="51DD3024A86344545743E40EAA7B5E54">text</mods:typeOfResource>
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<mods:title id="33592CB13183468567FD067ADDA1CB68">Zootaxa</mods:title>
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<mods:part id="786BFD180ADA0C37AABFEE648CCEEE97">
<mods:date id="4967955B89AD03367703624D2AC60C44">2010</mods:date>
<mods:detail id="82410ABEAF8885B6DA95A6164DF223EA" type="volume">
<mods:number id="A47E036E790656466412EA15E7B761A7">2555</mods:number>
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<mods:classification id="80C284BD836CB703070857FDEC69A874">journal article</mods:classification>
<mods:identifier id="C969C0470A033DA447BECC32C6D4CF0F" type="DOI">10.5281/zenodo.196907</mods:identifier>
<mods:identifier id="B0E1E3F4780FD6A54CF75AA5DDDC139A" type="GBIF-Dataset">75a8fe9a-5028-4a6a-ba3c-aa404344d072</mods:identifier>
<mods:identifier id="E05CC0486192787B46F62F7D36F38A8F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E0F5A365403F16DCD104C59BF839CFCB" type="Zenodo-Dep">196907</mods:identifier>
</mods:mods>
<treatment id="03B710750315CD4CFF154EEEFB9202F9" ID-DOI="http://doi.org/10.5281/zenodo.6195936" ID-GBIF-Taxon="119392684" ID-Zenodo-Dep="6195936" LSID="urn:lsid:plazi:treatment:03B710750315CD4CFF154EEEFB9202F9" httpUri="http://treatment.plazi.org/id/03B710750315CD4CFF154EEEFB9202F9" lastPageId="5" lastPageNumber="67" pageId="2" pageNumber="64">
<subSubSection id="C304F2E80315CD4BFF154EEEFEDA04D7" pageId="2" pageNumber="64" type="nomenclature">
<paragraph id="8BA1A1630315CD4BFF154EEEFDDA049A" blockId="2.[151,644,151,252]" box="[151,536,151,177]" pageId="2" pageNumber="64">
<heading id="D0E9160F0315CD4BFF154EEEFDDA049A" bold="true" box="[151,536,151,177]" fontSize="11" level="1" pageId="2" pageNumber="64" reason="1">
<emphasis id="B96A7D710315CD4BFF154EEEFDDA049A" bold="true" box="[151,536,151,177]" pageId="2" pageNumber="64">
<taxonomicName id="4C1EDAE00315CD4BFF154EEEFE0D049A" ID-CoL="3LG9L" box="[151,463,151,177]" class="Malacostraca" family="Trapeziidae" genus="Hexagonaloides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="bathyalis" status="sp. nov.">
<emphasis id="B96A7D710315CD4BFF154EEEFE0D049A" bold="true" box="[151,463,151,177]" italics="true" pageId="2" pageNumber="64">Hexagonaloides bathyalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A259C00A0315CD4BFE544EEEFDDA049A" box="[470,536,151,177]" pageId="2" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1A1630315CD4BFF154EC4FEDA04D7" blockId="2.[151,644,151,252]" pageId="2" pageNumber="64">
[new Japanese name: Shinkai-sango-gani] (
<figureCitation id="1325BDE60315CD4BFF1D4E9BFECC04D7" box="[159,270,226,252]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,255,1980,2004]" captionStartId-1="3.[151,255,1871,1895]" captionStartId-2="4.[151,257,1878,1902]" captionTargetBox-0="[151,1434,894,1960]" captionTargetBox-1="[186,1390,229,1870]" captionTargetBox-2="[224,1363,185,1845]" captionTargetId-0="figure@2.[151,1436,894,1962]" captionTargetId-1="figure@3.[184,1418,194,1870]" captionTargetId-2="figure@4.[194,1409,162,1880]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. Entire animal in dorsal view." captionText-1="FIGURE 2. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, carapace and cephalic appendages, dorsal view; B, frontal region and cephalic appendages, left side, anterior view; C, left third maxilliped, outer view; D, third to fifth thoracic sternites, ventral view; E, details of median part of fifth to seventh thoracic sternites, ventral view; F, abdomen, outer view; F, left first gonopod, ventral (sternal) view; G, left second pleopod, ventral (sternal) view; I, same, different view. Scale bars: 1 mm for A F; 0.5 mm for G I." captionText-2="FIGURE 3. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, left major cheliped, dorsal view; B, same, chela, outer view; C, right minor cheliped, dorsal view; D, same, chela, outer view; E, left fourth pereopod (third ambulatory leg), posterior view; F, same, dactylus, posterior view; G, same, obliquely flexor view, showing flexor surface; H, left fifth pereopod (fourth ambulatory leg), posterior view; I, same, dactylus, posterior view. Scale bars: 2 mm for A, C; 1 mm for B, D, E, H; 0.5 mm for F, G, I." httpUri-0="https://zenodo.org/record/196908/files/figure.png" httpUri-1="https://zenodo.org/record/196909/files/figure.png" httpUri-2="https://zenodo.org/record/196910/files/figure.png" pageId="2" pageNumber="64">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C304F2E80315CD4BFF154F54FBD9059C" pageId="2" pageNumber="64" type="materials_examined">
<paragraph id="8BA1A1630315CD4BFF154F54FF1B05B9" blockId="2.[151,1437,301,850]" pageId="2" pageNumber="64">
<emphasis id="B96A7D710315CD4BFF154F54FE55056C" bold="true" box="[151,407,301,327]" pageId="2" pageNumber="64">Material examined.</emphasis>
<typeStatus id="54A51FC10315CD4BFE224F54FDD5056C" box="[416,535,301,327]" pageId="2" pageNumber="64" type="holotype">Holotype</typeStatus>
: male (3.8 ×
<quantity id="4CE60C860315CD4BFD394F54FCDC056C" box="[699,798,301,327]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="64" unit="mm" value="5.5">5.5 mm</quantity>
), off Ie
<collectingCountry id="F309E1F30315CD4BFC034F54FC12056C" box="[897,976,301,327]" name="Iceland" pageId="2" pageNumber="64">Island</collectingCountry>
, Okinawa, Ryukyu Islands (Okinawa Prefecture),
<geoCoordinate id="EE2AC7A40315CD4BFEAB4F2BFE070547" box="[297,453,338,364]" direction="north" orientation="latitude" pageId="2" pageNumber="64" precision="1" value="26.615784">26°36.947N</geoCoordinate>
,
<geoCoordinate id="EE2AC7A40315CD4BFE504F2BFDBB0547" box="[466,633,338,364]" direction="east" orientation="longitude" pageId="2" pageNumber="64" precision="1" value="127.73486">127°44.092E</geoCoordinate>
,
<quantity id="4CE60C860315CD4BFD044F2BFD160547" box="[646,724,338,364]" metricMagnitude="2" metricUnit="m" metricValue="2.4699999999999998" pageId="2" pageNumber="64" unit="m" value="247.0">247 m</quantity>
, on a rock entangled in gill net, coll. T. Higashiji, CBM-ZC 9935.
</paragraph>
<paragraph id="8BA1A1630315CD4BFF474FE4FBD9059C" blockId="2.[151,1437,301,850]" box="[197,1051,413,439]" pageId="2" pageNumber="64">
<typeStatus id="54A51FC10315CD4BFF474FE4FEF2059C" box="[197,304,413,439]" pageId="2" pageNumber="64" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D186AEA0315CD4BFEB94FE4FE48059C" box="[315,394,413,439]" pageId="2" pageNumber="64" type="male">1 male</specimenCount>
(3.7 ×
<quantity id="4CE60C860315CD4BFE5C4FE4FDFE059C" box="[478,572,413,439]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="2" pageNumber="64" unit="mm" value="5.1">5.1 mm</quantity>
), same data as
<typeStatus id="54A51FC10315CD4BFD6E4FE4FC94059C" box="[748,854,413,439]" pageId="2" pageNumber="64" type="holotype">holotype</typeStatus>
, CBM-ZC 9936.
</paragraph>
</subSubSection>
<subSubSection id="C304F2E80315CD4CFF474FBBFE9E0249" lastPageId="5" lastPageNumber="67" pageId="2" pageNumber="64" type="description">
<paragraph id="8BA1A1630315CD4BFF474FBBFF130779" blockId="2.[151,1437,301,850]" pageId="2" pageNumber="64">
<emphasis id="B96A7D710315CD4BFF474FBBFE9805F7" bold="true" box="[197,346,450,476]" pageId="2" pageNumber="64">Description</emphasis>
.
<typeStatus id="54A51FC10315CD4BFEE84FBBFE1B05F0" box="[362,473,450,475]" pageId="2" pageNumber="64" type="holotype">
<emphasis id="B96A7D710315CD4BFEE84FBBFE1B05F0" box="[362,473,450,475]" italics="true" pageId="2" pageNumber="64">Holotype</emphasis>
</typeStatus>
. Carapace (
<figureCitation id="1325BDE60315CD4BFDE94FBBFD0205F7" box="[619,704,450,476]" captionStart="FIGURE 1" captionStartId="2.[151,255,1980,2004]" captionTargetBox="[151,1434,894,1960]" captionTargetId="figure@2.[151,1436,894,1962]" captionTargetPageId="2" captionText="FIGURE 1. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. Entire animal in dorsal view." httpUri="https://zenodo.org/record/196908/files/figure.png" pageId="2" pageNumber="64">Figs. 1</figureCitation>
,
<figureCitation id="1325BDE60315CD4BFD524FBBFD1D05F7" box="[720,735,450,476]" captionStart="FIGURE 2" captionStartId="3.[151,255,1871,1895]" captionTargetBox="[186,1390,229,1870]" captionTargetId="figure@3.[184,1418,194,1870]" captionTargetPageId="3" captionText="FIGURE 2. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, carapace and cephalic appendages, dorsal view; B, frontal region and cephalic appendages, left side, anterior view; C, left third maxilliped, outer view; D, third to fifth thoracic sternites, ventral view; E, details of median part of fifth to seventh thoracic sternites, ventral view; F, abdomen, outer view; F, left first gonopod, ventral (sternal) view; G, left second pleopod, ventral (sternal) view; I, same, different view. Scale bars: 1 mm for A F; 0.5 mm for G I." httpUri="https://zenodo.org/record/196909/files/figure.png" pageId="2" pageNumber="64">2</figureCitation>
A) hexagonal in outline, about 1.4 times wider than long; dorsal surface slightly convex longitudinally and transversely, microscopically granular, smooth. Front about half of greatest width of carapace, nearly straight medially, sloping slightly laterally in dorsal view. Orbit lacking fissures; supraorbital margin concave in dorsal view, lateral angle slightly produced, unarmed; suborbital margin weakly granular, inner suborbital angle bluntly triangular, not reaching front, antennular flagellum entering orbit. Lateral margin convex with small acute tooth at midlength, anterolateral margin very slightly arcuate, posterolateral margin nearly straight or very slightly convex; posterior margin subequal to width between outer orbital angles, with very shallow excavation laterally. Pterygostomial region (
<figureCitation id="1325BDE60315CD4BFAB54CB1FA4206C9" box="[1335,1408,712,738]" captionStart="FIGURE 2" captionStartId="3.[151,255,1871,1895]" captionTargetBox="[186,1390,229,1870]" captionTargetId="figure@3.[184,1418,194,1870]" captionTargetPageId="3" captionText="FIGURE 2. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, carapace and cephalic appendages, dorsal view; B, frontal region and cephalic appendages, left side, anterior view; C, left third maxilliped, outer view; D, third to fifth thoracic sternites, ventral view; E, details of median part of fifth to seventh thoracic sternites, ventral view; F, abdomen, outer view; F, left first gonopod, ventral (sternal) view; G, left second pleopod, ventral (sternal) view; I, same, different view. Scale bars: 1 mm for A F; 0.5 mm for G I." httpUri="https://zenodo.org/record/196909/files/figure.png" pageId="2" pageNumber="64">Fig. 2</figureCitation>
B) nearly smooth, a shallow groove extending from anterolateral angle of buccal frame. Buccal frame not narrowed anteriorly; epistome very narrow, with 2 rounded lobes either side of broadly triangular median lobe.
</paragraph>
<caption id="DF61F1EB0315CD4BFF1549C5FEDA03D8" httpUri="https://zenodo.org/record/196908/files/figure.png" pageId="2" pageNumber="64" targetBox="[151,1434,894,1960]" targetPageId="2">
<paragraph id="8BA1A1630315CD4BFF1549C5FEDA03D8" blockId="2.[151,1436,1980,2035]" pageId="2" pageNumber="64">
<emphasis id="B96A7D710315CD4BFF1549C5FED103FF" bold="true" box="[151,275,1980,2004]" pageId="2" pageNumber="64">FIGURE 1</emphasis>
.
<taxonomicName id="4C1EDAE00315CD4BFEA349C4FDFA03FF" box="[289,568,1981,2004]" class="Malacostraca" family="Trapeziidae" genus="Hexagonaloides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="bathyalis" status="gen. nov.">
<emphasis id="B96A7D710315CD4BFEA349C4FDFA03FF" box="[289,568,1981,2004]" italics="true" pageId="2" pageNumber="64">Hexagonaloides bathyalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A259C00A0315CD4BFDC249C5FD4E03FF" box="[576,652,1980,2004]" pageId="2" pageNumber="64" rank="genus">
<emphasis id="B96A7D710315CD4BFDC249C5FD4E03FF" bold="true" box="[576,652,1980,2004]" pageId="2" pageNumber="64">n. gen.</emphasis>
</taxonomicNameLabel>
,
<emphasis id="B96A7D710315CD4BFD1849C5FD1403FF" bold="true" box="[666,726,1980,2004]" pageId="2" pageNumber="64">n. sp.</emphasis>
, holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. Entire animal in dorsal view.
</paragraph>
</caption>
<caption id="DF61F1EB0314CD4AFF154936FB7B03CC" httpUri="https://zenodo.org/record/196909/files/figure.png" pageId="3" pageNumber="65" targetBox="[186,1390,229,1870]" targetPageId="3">
<paragraph id="8BA1A1630314CD4AFF154936FB7B03CC" blockId="3.[151,1436,1871,2023]" pageId="3" pageNumber="65">
<emphasis id="B96A7D710314CD4AFF154936FED8034C" bold="true" box="[151,282,1871,1895]" pageId="3" pageNumber="65">FIGURE 2.</emphasis>
<taxonomicName id="4C1EDAE00314CD4AFEA04929FDFB034C" box="[290,569,1872,1895]" class="Malacostraca" family="Trapeziidae" genus="Hexagonaloides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="65" phylum="Arthropoda" rank="species" species="bathyalis" status="gen. nov.">
<emphasis id="B96A7D710314CD4AFEA04929FDFB034C" box="[290,569,1872,1895]" italics="true" pageId="3" pageNumber="65">Hexagonaloides bathyalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A259C00A0314CD4AFDC24936FD4E034C" box="[576,652,1871,1895]" pageId="3" pageNumber="65" rank="genus">
<emphasis id="B96A7D710314CD4AFDC24936FD4E034C" bold="true" box="[576,652,1871,1895]" pageId="3" pageNumber="65">n. gen.</emphasis>
</taxonomicNameLabel>
,
<emphasis id="B96A7D710314CD4AFD184936FD15034C" bold="true" box="[666,727,1871,1895]" pageId="3" pageNumber="65">n. sp.</emphasis>
, holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, carapace and cephalic appendages, dorsal view; B, frontal region and cephalic appendages, left side, anterior view; C, left third maxilliped, outer view; D, third to fifth thoracic sternites, ventral view; E, details of median part of fifth to seventh thoracic sternites, ventral view; F, abdomen, outer view; F, left first gonopod, ventral (sternal) view; G, left second pleopod, ventral (sternal) view; I, same, different view. Scale bars: 1 mm for AF; 0.5 mm for GI.
</paragraph>
</caption>
<caption id="DF61F1EB0313CD4DFF15492FFC6A03C6" httpUri="https://zenodo.org/record/196910/files/figure.png" pageId="4" pageNumber="66" targetBox="[224,1363,185,1845]" targetPageId="4">
<paragraph id="8BA1A1630313CD4DFF15492FFC6A03C6" blockId="4.[151,1436,1878,2029]" pageId="4" pageNumber="66">
<emphasis id="B96A7D710313CD4DFF15492FFEDF0345" bold="true" box="[151,285,1878,1902]" pageId="4" pageNumber="66">FIGURE 3.</emphasis>
<taxonomicName id="4C1EDAE00313CD4DFEA6492FFD810346" box="[292,579,1878,1901]" class="Malacostraca" family="Trapeziidae" genus="Hexagonaloides" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="bathyalis" status="gen. nov.">
<emphasis id="B96A7D710313CD4DFEA6492FFD810346" box="[292,579,1878,1901]" italics="true" pageId="4" pageNumber="66">Hexagonaloides bathyalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A259C00A0313CD4DFDC9492FFD5B0345" box="[587,665,1878,1902]" pageId="4" pageNumber="66" rank="genus">
<emphasis id="B96A7D710313CD4DFDC9492FFD5B0345" bold="true" box="[587,665,1878,1902]" pageId="4" pageNumber="66">n. gen.</emphasis>
</taxonomicNameLabel>
,
<emphasis id="B96A7D710313CD4DFD2A492FFD240345" bold="true" box="[680,742,1878,1902]" pageId="4" pageNumber="66">n. sp.</emphasis>
, holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, left major cheliped, dorsal view; B, same, chela, outer view; C, right minor cheliped, dorsal view; D, same, chela, outer view; E, left fourth pereopod (third ambulatory leg), posterior view; F, same, dactylus, posterior view; G, same, obliquely flexor view, showing flexor surface; H, left fifth pereopod (fourth ambulatory leg), posterior view; I, same, dactylus, posterior view. Scale bars: 2 mm for A, C; 1 mm for B, D, E, H; 0.5 mm for F, G, I.
</paragraph>
</caption>
<paragraph id="8BA1A1630312CD4CFF474EE1FB9E05B9" blockId="5.[151,1437,152,1746]" pageId="5" pageNumber="67">
Eye (
<figureCitation id="1325BDE60312CD4CFE804EE1FE850499" box="[258,327,152,178]" captionStart="FIGURE 2" captionStartId="3.[151,255,1871,1895]" captionTargetBox="[186,1390,229,1870]" captionTargetId="figure@3.[184,1418,194,1870]" captionTargetPageId="3" captionText="FIGURE 2. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, carapace and cephalic appendages, dorsal view; B, frontal region and cephalic appendages, left side, anterior view; C, left third maxilliped, outer view; D, third to fifth thoracic sternites, ventral view; E, details of median part of fifth to seventh thoracic sternites, ventral view; F, abdomen, outer view; F, left first gonopod, ventral (sternal) view; G, left second pleopod, ventral (sternal) view; I, same, different view. Scale bars: 1 mm for A F; 0.5 mm for G I." httpUri="https://zenodo.org/record/196909/files/figure.png" pageId="5" pageNumber="67">Fig. 2</figureCitation>
A, B) not concealed in orbit when retracted; cornea slightly narrower than eyestalk. Antennular peduncle (
<figureCitation id="1325BDE60312CD4CFE924EC4FE9704FC" box="[272,341,189,215]" captionStart="FIGURE 2" captionStartId="3.[151,255,1871,1895]" captionTargetBox="[186,1390,229,1870]" captionTargetId="figure@3.[184,1418,194,1870]" captionTargetPageId="3" captionText="FIGURE 2. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, carapace and cephalic appendages, dorsal view; B, frontal region and cephalic appendages, left side, anterior view; C, left third maxilliped, outer view; D, third to fifth thoracic sternites, ventral view; E, details of median part of fifth to seventh thoracic sternites, ventral view; F, abdomen, outer view; F, left first gonopod, ventral (sternal) view; G, left second pleopod, ventral (sternal) view; I, same, different view. Scale bars: 1 mm for A F; 0.5 mm for G I." httpUri="https://zenodo.org/record/196909/files/figure.png" pageId="5" pageNumber="67">Fig. 2</figureCitation>
A, B) with first segment transversely oval. Antennal peduncle (
<figureCitation id="1325BDE60312CD4CFBB44EC4FBBE04FC" box="[1078,1148,189,215]" captionStart="FIGURE 2" captionStartId="3.[151,255,1871,1895]" captionTargetBox="[186,1390,229,1870]" captionTargetId="figure@3.[184,1418,194,1870]" captionTargetPageId="3" captionText="FIGURE 2. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, carapace and cephalic appendages, dorsal view; B, frontal region and cephalic appendages, left side, anterior view; C, left third maxilliped, outer view; D, third to fifth thoracic sternites, ventral view; E, details of median part of fifth to seventh thoracic sternites, ventral view; F, abdomen, outer view; F, left first gonopod, ventral (sternal) view; G, left second pleopod, ventral (sternal) view; I, same, different view. Scale bars: 1 mm for A F; 0.5 mm for G I." httpUri="https://zenodo.org/record/196909/files/figure.png" pageId="5" pageNumber="67">Fig. 2</figureCitation>
A, B) standing obliquely in frontal view. Interantennular septum broadly triangular, depressed. Third maxilliped (
<figureCitation id="1325BDE60312CD4CFB294E9BFB3004D7" box="[1195,1266,226,252]" captionStart="FIGURE 2" captionStartId="3.[151,255,1871,1895]" captionTargetBox="[186,1390,229,1870]" captionTargetId="figure@3.[184,1418,194,1870]" captionTargetPageId="3" captionText="FIGURE 2. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, carapace and cephalic appendages, dorsal view; B, frontal region and cephalic appendages, left side, anterior view; C, left third maxilliped, outer view; D, third to fifth thoracic sternites, ventral view; E, details of median part of fifth to seventh thoracic sternites, ventral view; F, abdomen, outer view; F, left first gonopod, ventral (sternal) view; G, left second pleopod, ventral (sternal) view; I, same, different view. Scale bars: 1 mm for A F; 0.5 mm for G I." httpUri="https://zenodo.org/record/196909/files/figure.png" pageId="5" pageNumber="67">Fig. 2</figureCitation>
C) moderately wide; ischium widened proximally, with prominent projection distally on mesial margin, broadly rounded proximomesial margin; merus smaller than ischium, with small triangular process on distal margin, forming articulation with carpus; dactylus slightly longer than propodus, tapering distally; exopod moderately stout, inner mesial margin with triangular process subdistally; flagellum well developed.
</paragraph>
<paragraph id="8BA1A1630312CD4CFF474FE4FC720697" blockId="5.[151,1437,152,1746]" pageId="5" pageNumber="67">
Chelipeds (
<figureCitation id="1325BDE60312CD4CFEC94FE4FE5F059C" box="[331,413,413,439]" captionStart="FIGURE 1" captionStartId="2.[151,255,1980,2004]" captionTargetBox="[151,1434,894,1960]" captionTargetId="figure@2.[151,1436,894,1962]" captionTargetPageId="2" captionText="FIGURE 1. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. Entire animal in dorsal view." httpUri="https://zenodo.org/record/196908/files/figure.png" pageId="5" pageNumber="67">Figs. 1</figureCitation>
,
<figureCitation id="1325BDE60312CD4CFE2E4FE4FE7B059C" box="[428,441,413,439]" captionStart="FIGURE 3" captionStartId="4.[151,257,1878,1902]" captionTargetBox="[224,1363,185,1845]" captionTargetId="figure@4.[194,1409,162,1880]" captionTargetPageId="4" captionText="FIGURE 3. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, left major cheliped, dorsal view; B, same, chela, outer view; C, right minor cheliped, dorsal view; D, same, chela, outer view; E, left fourth pereopod (third ambulatory leg), posterior view; F, same, dactylus, posterior view; G, same, obliquely flexor view, showing flexor surface; H, left fifth pereopod (fourth ambulatory leg), posterior view; I, same, dactylus, posterior view. Scale bars: 2 mm for A, C; 1 mm for B, D, E, H; 0.5 mm for F, G, I." httpUri="https://zenodo.org/record/196910/files/figure.png" pageId="5" pageNumber="67">3</figureCitation>
AD) unequal, slightly dissimilar, surface smooth to naked eye, but microscopically granular, sparsely setose. Larger (left) cheliped about 1.9 carapace length, not massive; merus with blunt, minutely granular anterior ridge, largely visible in dorsal view; dorsodistal margin spinulose in inner half. Carpus short, lacking tooth at inner angle. Chela 2.7 times as long as deep. Palm elongate, depth slightly increasing distally; dorsal and ventral surfaces rounded; ventral margin in outer view slightly sinuous; fixed finger slightly deflexed, terminating in curved, acute tip, cutting edge thin, sinuous, minutely dentate. Dactylus 0.7 times as long as palm, weakly curved, minutely granular on dorsal margin, terminating in acute tip crossing tip of fixed finger; cutting edge thin, minutely granular.
</paragraph>
<paragraph id="8BA1A1630312CD4CFF474CB1FCC60779" blockId="5.[151,1437,152,1746]" pageId="5" pageNumber="67">Smaller (right) cheliped about 1.7 of carapace length; merus bearing clearly granular anterior ridge. Chela 3.0 times as long as deep. Fixed finger with cutting edge nearly straight except for curved tip, bearing 3 nearly indistinguishable teeth proximally. Dactylus 0.9 times as long as palm, terminating in curved, acute tip, crossing tip of fixed finger, cutting edge non dentate.</paragraph>
<paragraph id="8BA1A1630312CD4CFF474D24FCD70019" blockId="5.[151,1437,152,1746]" pageId="5" pageNumber="67">
Ambulatory legs (
<figureCitation id="1325BDE60312CD4CFE194D24FE2C075C" box="[411,494,861,887]" captionStart="FIGURE 1" captionStartId="2.[151,255,1980,2004]" captionTargetBox="[151,1434,894,1960]" captionTargetId="figure@2.[151,1436,894,1962]" captionTargetPageId="2" captionText="FIGURE 1. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. Entire animal in dorsal view." httpUri="https://zenodo.org/record/196908/files/figure.png" pageId="5" pageNumber="67">Figs. 1</figureCitation>
,
<figureCitation id="1325BDE60312CD4CFE7F4D24FDC8075C" box="[509,522,861,887]" captionStart="FIGURE 3" captionStartId="4.[151,257,1878,1902]" captionTargetBox="[224,1363,185,1845]" captionTargetId="figure@4.[194,1409,162,1880]" captionTargetPageId="4" captionText="FIGURE 3. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, left major cheliped, dorsal view; B, same, chela, outer view; C, right minor cheliped, dorsal view; D, same, chela, outer view; E, left fourth pereopod (third ambulatory leg), posterior view; F, same, dactylus, posterior view; G, same, obliquely flexor view, showing flexor surface; H, left fifth pereopod (fourth ambulatory leg), posterior view; I, same, dactylus, posterior view. Scale bars: 2 mm for A, C; 1 mm for B, D, E, H; 0.5 mm for F, G, I." httpUri="https://zenodo.org/record/196910/files/figure.png" pageId="5" pageNumber="67">3</figureCitation>
EI) long, slender, smooth to naked eye, sparsely setose; similar in structure, third leg longest; dactylo-propodal lock well developed. Propodi each with flexor margin bearing row of slender movable spinules on distal half. Dactyli subequal in length to propodi, weakly curved, each terminating in minute corneous spinule; extensor surfaces each with slender short spiniform setae arranged in 3 irregular rows in distal 0.40.5 portion; flexor surface narrowly channeled, flanked by thin, crenulated edges, each edge with 811 slender corneous spinules.
</paragraph>
<paragraph id="8BA1A1630312CD4CFF474A44FF3700C7" blockId="5.[151,1437,152,1746]" pageId="5" pageNumber="67">
Thoracic sternum broad. Third and fourth sternites (
<figureCitation id="1325BDE60312CD4CFCB04A44FCB8007C" box="[818,890,1085,1111]" captionStart="FIGURE 2" captionStartId="3.[151,255,1871,1895]" captionTargetBox="[186,1390,229,1870]" captionTargetId="figure@3.[184,1418,194,1870]" captionTargetPageId="3" captionText="FIGURE 2. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, carapace and cephalic appendages, dorsal view; B, frontal region and cephalic appendages, left side, anterior view; C, left third maxilliped, outer view; D, third to fifth thoracic sternites, ventral view; E, details of median part of fifth to seventh thoracic sternites, ventral view; F, abdomen, outer view; F, left first gonopod, ventral (sternal) view; G, left second pleopod, ventral (sternal) view; I, same, different view. Scale bars: 1 mm for A F; 0.5 mm for G I." httpUri="https://zenodo.org/record/196909/files/figure.png" pageId="5" pageNumber="67">Fig. 2</figureCitation>
D) fused, forming large plate, anterior margin broadly convex with median point, short suture between third and fourth sternites. Fifth sternite with tuft of short setae on either side of midline; press button very small, papilla-like (
<figureCitation id="1325BDE60312CD4CFBA04AF1FBA90089" box="[1058,1131,1160,1186]" captionStart="FIGURE 2" captionStartId="3.[151,255,1871,1895]" captionTargetBox="[186,1390,229,1870]" captionTargetId="figure@3.[184,1418,194,1870]" captionTargetPageId="3" captionText="FIGURE 2. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, carapace and cephalic appendages, dorsal view; B, frontal region and cephalic appendages, left side, anterior view; C, left third maxilliped, outer view; D, third to fifth thoracic sternites, ventral view; E, details of median part of fifth to seventh thoracic sternites, ventral view; F, abdomen, outer view; F, left first gonopod, ventral (sternal) view; G, left second pleopod, ventral (sternal) view; I, same, different view. Scale bars: 1 mm for A F; 0.5 mm for G I." httpUri="https://zenodo.org/record/196909/files/figure.png" pageId="5" pageNumber="67">Fig. 2</figureCitation>
D, E). Sixth sternite with short transverse row of setae on either side of midline (
<figureCitation id="1325BDE60312CD4CFCA74AD4FCAF00EC" box="[805,877,1197,1223]" captionStart="FIGURE 2" captionStartId="3.[151,255,1871,1895]" captionTargetBox="[186,1390,229,1870]" captionTargetId="figure@3.[184,1418,194,1870]" captionTargetPageId="3" captionText="FIGURE 2. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, carapace and cephalic appendages, dorsal view; B, frontal region and cephalic appendages, left side, anterior view; C, left third maxilliped, outer view; D, third to fifth thoracic sternites, ventral view; E, details of median part of fifth to seventh thoracic sternites, ventral view; F, abdomen, outer view; F, left first gonopod, ventral (sternal) view; G, left second pleopod, ventral (sternal) view; I, same, different view. Scale bars: 1 mm for A F; 0.5 mm for G I." httpUri="https://zenodo.org/record/196909/files/figure.png" pageId="5" pageNumber="67">Fig. 2</figureCitation>
E). Gonopore coxal, but located close to eighth sternite.
</paragraph>
<paragraph id="8BA1A1630312CD4CFF474A81FF3D0177" blockId="5.[151,1437,152,1746]" pageId="5" pageNumber="67">
Male abdomen (
<figureCitation id="1325BDE60312CD4CFE064A81FE080139" box="[388,458,1272,1298]" captionStart="FIGURE 2" captionStartId="3.[151,255,1871,1895]" captionTargetBox="[186,1390,229,1870]" captionTargetId="figure@3.[184,1418,194,1870]" captionTargetPageId="3" captionText="FIGURE 2. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, carapace and cephalic appendages, dorsal view; B, frontal region and cephalic appendages, left side, anterior view; C, left third maxilliped, outer view; D, third to fifth thoracic sternites, ventral view; E, details of median part of fifth to seventh thoracic sternites, ventral view; F, abdomen, outer view; F, left first gonopod, ventral (sternal) view; G, left second pleopod, ventral (sternal) view; I, same, different view. Scale bars: 1 mm for A F; 0.5 mm for G I." httpUri="https://zenodo.org/record/196909/files/figure.png" pageId="5" pageNumber="67">Fig. 2</figureCitation>
F) with third to fifth somites incompletely fused, sutures clearly delimited in lateral sides. Third somite widest. Telson (
<figureCitation id="1325BDE60312CD4CFDCA4B64FD52011C" box="[584,656,1309,1335]" captionStart="FIGURE 2" captionStartId="3.[151,255,1871,1895]" captionTargetBox="[186,1390,229,1870]" captionTargetId="figure@3.[184,1418,194,1870]" captionTargetPageId="3" captionText="FIGURE 2. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, carapace and cephalic appendages, dorsal view; B, frontal region and cephalic appendages, left side, anterior view; C, left third maxilliped, outer view; D, third to fifth thoracic sternites, ventral view; E, details of median part of fifth to seventh thoracic sternites, ventral view; F, abdomen, outer view; F, left first gonopod, ventral (sternal) view; G, left second pleopod, ventral (sternal) view; I, same, different view. Scale bars: 1 mm for A F; 0.5 mm for G I." httpUri="https://zenodo.org/record/196909/files/figure.png" pageId="5" pageNumber="67">Fig. 2</figureCitation>
F) triangular, nearly as long as wide, with proximolateral corner rounded.
</paragraph>
<paragraph id="8BA1A1630312CD4CFF474B11FC2101D9" blockId="5.[151,1437,152,1746]" pageId="5" pageNumber="67">
First gonopod (
<figureCitation id="1325BDE60312CD4CFE034B11FE0801A9" box="[385,458,1384,1410]" captionStart="FIGURE 2" captionStartId="3.[151,255,1871,1895]" captionTargetBox="[186,1390,229,1870]" captionTargetId="figure@3.[184,1418,194,1870]" captionTargetPageId="3" captionText="FIGURE 2. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, carapace and cephalic appendages, dorsal view; B, frontal region and cephalic appendages, left side, anterior view; C, left third maxilliped, outer view; D, third to fifth thoracic sternites, ventral view; E, details of median part of fifth to seventh thoracic sternites, ventral view; F, abdomen, outer view; F, left first gonopod, ventral (sternal) view; G, left second pleopod, ventral (sternal) view; I, same, different view. Scale bars: 1 mm for A F; 0.5 mm for G I." httpUri="https://zenodo.org/record/196909/files/figure.png" pageId="5" pageNumber="67">Fig. 2</figureCitation>
G) reaching to suture between fourth and fifth sternites, curved at proximal 0.3; distal 0.6 slender, gently curved dorsally, parallel to midline, gradually tapering to acute tip; row of minute spiniform setae directed laterally on distal 0.25 of lateral margin. Second gonopod (
<figureCitation id="1325BDE60312CD4CFBE64BCBFB6B01E7" box="[1124,1193,1458,1484]" captionStart="FIGURE 2" captionStartId="3.[151,255,1871,1895]" captionTargetBox="[186,1390,229,1870]" captionTargetId="figure@3.[184,1418,194,1870]" captionTargetPageId="3" captionText="FIGURE 2. Hexagonaloides bathyalis n. gen., n. sp., holotype, male (3.8 × 5.5 mm), CBM-ZC 9935. A, carapace and cephalic appendages, dorsal view; B, frontal region and cephalic appendages, left side, anterior view; C, left third maxilliped, outer view; D, third to fifth thoracic sternites, ventral view; E, details of median part of fifth to seventh thoracic sternites, ventral view; F, abdomen, outer view; F, left first gonopod, ventral (sternal) view; G, left second pleopod, ventral (sternal) view; I, same, different view. Scale bars: 1 mm for A F; 0.5 mm for G I." httpUri="https://zenodo.org/record/196909/files/figure.png" pageId="5" pageNumber="67">Fig. 2</figureCitation>
H, I) short, about 0.3 length of first gonopod, slightly curved ventrally; tip obliquely truncate.
</paragraph>
<paragraph id="8BA1A1630312CD4CFF474B84FDEB023C" blockId="5.[151,1437,152,1746]" box="[197,553,1533,1559]" pageId="5" pageNumber="67">
<typeStatus id="54A51FC10312CD4CFF474B84FEF3023D" box="[197,305,1533,1558]" pageId="5" pageNumber="67" type="paratype">
<emphasis id="B96A7D710312CD4CFF474B84FEF3023D" box="[197,305,1533,1558]" italics="true" pageId="5" pageNumber="67">Paratype</emphasis>
</typeStatus>
. Similar to
<typeStatus id="54A51FC10312CD4CFE394B84FDE4023C" box="[443,550,1533,1559]" pageId="5" pageNumber="67" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BA1A1630312CD4CFF47485BFE9E0249" blockId="5.[151,1437,152,1746]" pageId="5" pageNumber="67">
<emphasis id="B96A7D710312CD4CFF47485BFE650217" bold="true" box="[197,423,1570,1596]" pageId="5" pageNumber="67">Coloration in life</emphasis>
. Body and appendages pale pink, bases of fingers and palms of chelipeds darker. Corneas grayish.
</paragraph>
</subSubSection>
<subSubSection id="C304F2E80312CD4CFF474814FEF70287" pageId="5" pageNumber="67" type="discussion">
<paragraph id="8BA1A1630312CD4CFF474814FEF70287" blockId="5.[151,1437,152,1746]" pageId="5" pageNumber="67">
<emphasis id="B96A7D710312CD4CFF474814FEFB02AC" bold="true" box="[197,313,1645,1671]" pageId="5" pageNumber="67">Remarks</emphasis>
. The two specimens are small, but the well-developed gonopods clearly suggest that they are not juveniles.
</paragraph>
</subSubSection>
<subSubSection id="C304F2E80312CD4CFF4748CEFB9202F9" box="[197,1104,1719,1746]" pageId="5" pageNumber="67" type="etymology">
<paragraph id="8BA1A1630312CD4CFF4748CEFB9202F9" blockId="5.[151,1437,152,1746]" box="[197,1104,1719,1746]" pageId="5" pageNumber="67">
<emphasis id="B96A7D710312CD4CFF4748CEFE8802FA" bold="true" box="[197,330,1719,1745]" pageId="5" pageNumber="67">Etymology</emphasis>
. Named in reference to the deep-water habitat of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>