345 lines
44 KiB
XML
345 lines
44 KiB
XML
<document id="CE5349947CD9BF0B9A59DE749D510BD1" ID-DOI="10.5281/zenodo.5167725" ID-ISSN="1942-1354" ID-Zenodo-Dep="5167725" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628270427885" checkinUser="felipe" docAuthor="Austin, George T. & Warren, Andrew D." docDate="2009" docId="03B687432D49856A3DDDF913236D10B0" docLanguage="en" docName="InsectaMundi.0089.1-55.pdf" docOrigin="Insecta Mundi 2009 (89)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Onespa nakamura Austin and A. Warren 2009, new species" docType="treatment" docVersion="3" lastPageNumber="14" masterDocId="FF8FFF3B2D4485653D1DFFF62651142B" masterDocTitle="New looks at and for Onespa, Buzyges, and Librita (Lepidoptera: Hesperiidae: Hesperiinae), with new combinations and descriptions of a new genus and six new species" masterLastPageNumber="55" masterPageNumber="1" pageNumber="12" updateTime="1699038508896" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="3DF2FD4D102640C1D2AD38B4D625E678">
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<mods:title id="D85172FE5D4678656EC3266FFED4D060">New looks at and for Onespa, Buzyges, and Librita (Lepidoptera: Hesperiidae: Hesperiinae), with new combinations and descriptions of a new genus and six new species</mods:title>
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<mods:name id="0708F6A2D705763A20E29AE0774919E6" type="personal">
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<mods:roleTerm id="C9222F4F29A33599822C7C87D359102D">Author</mods:roleTerm>
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<mods:namePart id="72C9D45272CB71BB70AEA942255BE255">Austin, George T.</mods:namePart>
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<mods:name id="F680EE160CE2A333F9BA9E0357C9119D" type="personal">
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<mods:roleTerm id="1A1A6403989232C3F9FF6A64FE7E46E0">Author</mods:roleTerm>
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<mods:namePart id="9157402D8C2691EA26CFD3081E593BDF">Warren, Andrew D.</mods:namePart>
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<mods:typeOfResource id="803AA6D93AA8AB3258C230AC160FAA73">text</mods:typeOfResource>
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<mods:title id="2F41D40E96ADBEF71F6EAD6A3DD464A2">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="B28176F256E35997A7AD546A266D8B3C">
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<mods:date id="B244BF3E404BA2E503F24A6B3B2FA0AF">2009</mods:date>
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<mods:number id="FD770E996469931E350DD7FDEC2625EF">2009-07-24</mods:number>
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<mods:detail id="93DCD987F0A543E26D1DDBBDA3B9D4A8" type="volume">
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<mods:number id="87AD24621214FCF1F11F1F792262982A">2009</mods:number>
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<mods:detail id="C70F99437AAD38524CC0204727E9069C" type="issue">
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<mods:number id="DDA92ED4873002E2701659A4D8B13FF6">89</mods:number>
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<mods:classification id="AA465C6EC0C17425057DCAF5119B9FF6">journal article</mods:classification>
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<mods:identifier id="B1C8863F62A12BD8D136128475FA9F43" type="DOI">10.5281/zenodo.5167725</mods:identifier>
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<mods:identifier id="D8AF935201BCCBA6AD981C557C5A4FE9" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="200B2335F31BAE79A37EAAD9689A7D76" type="Zenodo-Dep">5167725</mods:identifier>
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<treatment id="03B687432D49856A3DDDF913236D10B0" LSID="urn:lsid:plazi:treatment:03B687432D49856A3DDDF913236D10B0" httpUri="http://treatment.plazi.org/id/03B687432D49856A3DDDF913236D10B0" lastPageId="15" lastPageNumber="14" pageId="13" pageNumber="12">
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<subSubSection id="C30565DE2D4985683DDDF91325CF12D0" box="[192,926,1763,1788]" pageId="13" pageNumber="12" type="nomenclature">
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<paragraph id="8BA036552D4985683DDDF91325CF12D0" blockId="13.[192,926,1763,1819]" box="[192,926,1763,1788]" pageId="13" pageNumber="12">
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<heading id="D0E881392D4985683DDDF91325CF12D0" bold="true" box="[192,926,1763,1788]" fontSize="10" level="2" pageId="13" pageNumber="12" reason="3">
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<emphasis id="B96BEA472D4985683DDDF91325CF12D0" bold="true" box="[192,926,1763,1788]" pageId="13" pageNumber="12">
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<taxonomicName id="4C1F4DD62D4985683DDDF91324BF12D0" authority="Austin and A. Warren" authorityName="Austin and A. Warren" authorityYear="2009" box="[192,750,1763,1788]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="nakamura" status="sp. nov.">
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<emphasis id="B96BEA472D4985683DDDF91327EE12D7" bold="true" box="[192,447,1765,1788]" italics="true" pageId="13" pageNumber="12">Onespa nakamura</emphasis>
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Austin and A. Warren
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</taxonomicName>
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,
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<taxonomicNameLabel id="A258573C2D4985683FE7F91525CF12D0" box="[762,926,1763,1787]" pageId="13" pageNumber="12" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30565DE2D49856B3DDDF8F527B41350" lastPageId="14" lastPageNumber="13" pageId="13" pageNumber="12" type="description">
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<paragraph id="8BA036552D4985683DDDF8F527F11330" blockId="13.[192,926,1763,1819]" box="[192,416,1795,1819]" pageId="13" pageNumber="12">
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(
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<figureCitation id="13242AD02D4985683DD4F8F527141330" box="[201,325,1795,1819]" captionStart="Figure 1-18" captionStartId="5.[192,276,1691,1713]" captionTargetBox="[233,1406,197,1670]" captionTargetId="figure-16@5.[221,1418,189,1680]" captionTargetPageId="5" captionText="Figure 1-18. Onespa (full data in text; ds = dorsal surface, vs = ventral surface; scale bar = 10 mm). 1) O. nubis, holotype male, El Salvador, 25 March 1972, ds; 2) same, vs; 3) O. nubis, male, El Triunfo, Chiapas, Mexico, 22 March 2007, ds; 4) same, vs; 5) O. nubis, paratype female, El Salvador, 28 December 1971, ds; 6) same, vs; 7) O. gala, holotype male, Veracruz, Mexico, ds, reproduced from photograph at MGCL; 8) same, vs; 9) O. gala, male, Michoacán, Mexico, 22 July 1996, ds; 10) same, vs; 11) O. gala, female, Michoacán, Mexico, 27 July 1996, ds; 12) same, vs; 13) O. brockorum, holotype male, Sonora, Mexico, 24 July 1997, ds; 14) same, vs; 15) O. brockorum, paratype female, Sonora, Mexico, 22 July 1985, ds; 16) same, vs; 17) O. nakamura, holotype male, Costa Rica, 13 June 2005, ds; 18) same, vs." figureDoi="http://doi.org/10.5281/zenodo.5167727" httpUri="https://zenodo.org/record/5167727/files/figure.png" pageId="13" pageNumber="12">Fig. 17-18</figureCitation>
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,
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<figureCitation id="13242AD02D4985683C52F8F5273E1330" box="[335,367,1795,1819]" captionStart="Figure 63-66" captionStartId="35.[192,276,1215,1237]" captionTargetBox="[263,1362,195,1189]" captionTargetId="figure-293@35.[263,1362,192,1189]" captionTargetPageId="35" captionText="Figure 63-66. Male genitalia of Onespa (full data in text; shown are lateral view of uncus, gnathos, tegumen, and saccus; dorsal view of uncus and tegumen; ventral view of gnathos uncus; internal view of right valva; ventral view of saccus; ventral, dorsal, right lateral and/or left lateral views of aedeagus; scale bar = 1 mm). 63) O. nubis, El Salvador (SRS #4803); 64) O. brockorum, Sonora, Mexico, paratype (SRS #1852); 65) O. gala, Michoacán, Mexico (GTA #14065); 66) O. nakamura, Costa Rica, holotype (GTA #14033)." figureDoi="http://doi.org/10.5281/zenodo.5167739" httpUri="https://zenodo.org/record/5167739/files/figure.png" pageId="13" pageNumber="12">66</figureCitation>
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,
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<figureCitation id="13242AD02D4985683C64F8F527C61330" box="[377,407,1795,1819]" captionStart="Figure 75-86" captionStartId="38.[192,276,1030,1052]" captionTargetBox="[447,1192,193,1003]" captionTargetId="figure-298@38.[447,1192,193,1003]" captionTargetPageId="38" captionText="Figure 75-86. Rigid cornuti of Onespa, Buzyges, Librita, and Neposa (full data in text; shown are lateral and dorsal views). 75) O. nubis, El Salvador (SRS #4803); 76) O. gala, Michoacán, Mexico (GTA #14065); 77) O. brockorum, Sonora, Mexico, paratype (SRS #1852); 78) O. nakamura, Costa Rica, holotype (GTA #14033); 79) B. idothea, Costa Rica (ADW #97-102); 80) B. rolla, Costa Rica (GTA #14055); 81) B. benito, Oaxaca, Mexico (SRS #4468); 82) B. mellanaformis, Costa Rica, paratype (GTA #14031); 83) L. librita, Veracruz, Mexico (GTA #14135); 84) N. heras, Guerrero, Mexico (ADW #97-112); 85) N. armandoi, Oaxaca, Mexico, holotype (GTA #14143); 86) N. isabelae, Guatemala, paratype (ADW #98-15)." figureDoi="http://doi.org/10.5281/zenodo.5167749" httpUri="https://zenodo.org/record/5167749/files/figure.png" pageId="13" pageNumber="12">78</figureCitation>
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)
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</paragraph>
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<paragraph id="8BA036552D49856B3DDDF8B524931610" blockId="13.[192,1440,1859,1915]" lastBlockId="14.[192,1441,195,1915]" lastPageId="14" lastPageNumber="13" pageId="13" pageNumber="12">
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<emphasis id="B96BEA472D4985683DDDF8B527E81370" bold="true" box="[192,441,1859,1883]" pageId="13" pageNumber="12">Description. Male</emphasis>
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(
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<figureCitation id="13242AD02D4985683CD6F8B524181370" box="[459,585,1859,1883]" captionStart="Figure 1-18" captionStartId="5.[192,276,1691,1713]" captionTargetBox="[233,1406,197,1670]" captionTargetId="figure-16@5.[221,1418,189,1680]" captionTargetPageId="5" captionText="Figure 1-18. Onespa (full data in text; ds = dorsal surface, vs = ventral surface; scale bar = 10 mm). 1) O. nubis, holotype male, El Salvador, 25 March 1972, ds; 2) same, vs; 3) O. nubis, male, El Triunfo, Chiapas, Mexico, 22 March 2007, ds; 4) same, vs; 5) O. nubis, paratype female, El Salvador, 28 December 1971, ds; 6) same, vs; 7) O. gala, holotype male, Veracruz, Mexico, ds, reproduced from photograph at MGCL; 8) same, vs; 9) O. gala, male, Michoacán, Mexico, 22 July 1996, ds; 10) same, vs; 11) O. gala, female, Michoacán, Mexico, 27 July 1996, ds; 12) same, vs; 13) O. brockorum, holotype male, Sonora, Mexico, 24 July 1997, ds; 14) same, vs; 15) O. brockorum, paratype female, Sonora, Mexico, 22 July 1985, ds; 16) same, vs; 17) O. nakamura, holotype male, Costa Rica, 13 June 2005, ds; 18) same, vs." figureDoi="http://doi.org/10.5281/zenodo.5167727" httpUri="https://zenodo.org/record/5167727/files/figure.png" pageId="13" pageNumber="12">Fig. 17-18</figureCitation>
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) - forewing length =
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<quantity id="4CE79BB02D4985683E48F8B525931370" box="[853,962,1859,1883]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="13" pageNumber="12" unit="mm" value="18.2">18.2 mm</quantity>
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(n=1,
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<typeStatus id="54A488F72D4985683908F8B522D21370" box="[1045,1155,1859,1883]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
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); forewing produced to pointed apex, termen slightly convex, no stigma or brand; hindwing convex, weakly lobate at tornus; dorsal forewing dark brown with red-brown iridescence; proximal 1/2 costa bright red-brown, this color extending as overscaling of setiform scales into adjacent anterior portion and base of discal cell; sparse paler (more yellowish) red-brown overscaling in base of CuA 2 -2A; yellow-orange overscaling of setiform scales in middle 1/3 of anal cell; translucent pale yellow-orange macules (edged distad with shining gold color) as follows: subapical near bases of R
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<subScript id="179B34102D4A856B3FDAFEA224811549" attach="right" box="[711,720,340,354]" fontSize="6" pageId="14" pageNumber="13">4</subScript>
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-R
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<subScript id="179B34102D4A856B3FF1FEA524A4154A" attach="left" box="[748,757,339,353]" fontSize="6" pageId="14" pageNumber="13">5</subScript>
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and R
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<subScript id="179B34102D4A856B3E5EFEA5251D154A" attach="left" box="[835,844,339,353]" fontSize="6" pageId="14" pageNumber="13">5</subScript>
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-M
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<subScript id="179B34102D4A856B3E73FEA225261549" attach="left" box="[878,887,340,354]" fontSize="6" pageId="14" pageNumber="13">1</subScript>
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, more or less quadrate, that in R
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<subScript id="179B34102D4A856B3816FEA52345154A" attach="right" box="[1291,1300,339,353]" fontSize="6" pageId="14" pageNumber="13">5</subScript>
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-M
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<subScript id="179B34102D4A856B382BFEA2236E1549" attach="left" box="[1334,1343,340,354]" fontSize="6" pageId="14" pageNumber="13">1</subScript>
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about 2 times size of that in R
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<subScript id="179B34102D4A856B3CCCFE82278B15A9" attach="right" box="[465,474,372,386]" fontSize="6" pageId="14" pageNumber="13">4</subScript>
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-R
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<subScript id="179B34102D4A856B3CEBFE8527AE15AA" attach="right" box="[502,511,371,385]" fontSize="6" pageId="14" pageNumber="13">5</subScript>
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; postmedial in M
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<subScript id="179B34102D4A856B3FCAFE8224B115A9" attach="left" box="[727,736,372,386]" fontSize="6" pageId="14" pageNumber="13">3</subScript>
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-CuA
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<subScript id="179B34102D4A856B3E3DFE82257815A9" attach="left" box="[800,809,372,386]" fontSize="6" pageId="14" pageNumber="13">1</subScript>
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, trapezoidal (horizontally), 1/3 distance from base; subquadrate in CuA 1 -CuA 2, largest, distal edge slightly concave, centered under origin of CuA 1, overlapping proximal 1/3 of macule in M 3 -CuA 1; discal cell, hourglass shaped, about 1/4 distance from distal end, overlapping proximal 1/3 of macule in CuA 1 -CuA 2 (just proximad of origin of vein CuA 1); opaque yelloworange macule just distad of center of posterior 1/2 of CuA
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<subScript id="179B34102D4A856B3E86FE0225F51629" attach="left" box="[923,932,500,514]" fontSize="6" pageId="14" pageNumber="13">2</subScript>
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-2A; much smaller opaque yellow-orange macule in anterior 1/2 of CuA 2 -2A, just distad of center of macule in CuA 1 -CuA 2; right wing with minute opaque yellow-orange macule in mid-cell R
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</paragraph>
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<paragraph id="8BA036552D4A856B3FDFFDC2249A1669" blockId="14.[192,1441,195,1915]" box="[706,715,564,578]" pageId="14" pageNumber="13">3</paragraph>
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<paragraph id="8BA036552D4A856B3FD6FDD224B71610" blockId="14.[192,1441,195,1915]" box="[715,742,548,571]" pageId="14" pageNumber="13">-R</paragraph>
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<paragraph id="8BA036552D4A856B3FFBFDC224BE1669" blockId="14.[192,1441,195,1915]" box="[742,751,564,578]" pageId="14" pageNumber="13">4</paragraph>
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<paragraph id="8BA036552D4A856B3FF2FDD2234D1610" blockId="14.[192,1441,195,1915]" box="[751,1308,547,571]" pageId="14" pageNumber="13">; fringe dark gray-brown, slightly paler distad.</paragraph>
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<paragraph id="8BA036552D4A856B3DEDFDB527B31690" blockId="14.[192,1441,195,1915]" pageId="14" pageNumber="13">
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Dorsal hindwing dark brown; proximal 2/3 overscaled with long brown and a few ochreous scales caudad of vein Sc+R 1, nearly reaching termen along vein 2A; opaque, small, irregularly-shaped yelloworange postmedial macules in Rs-M
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<subScript id="179B34102D4A856B3F6CFD62242B1689" attach="left" box="[625,634,660,674]" fontSize="6" pageId="14" pageNumber="13">1</subScript>
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, M
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<subScript id="179B34102D4A856B3FBDFD6224F81689" attach="left" box="[672,681,660,674]" fontSize="6" pageId="14" pageNumber="13">1</subScript>
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-M
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<subScript id="179B34102D4A856B3FD7FD6224821689" attach="left" box="[714,723,660,674]" fontSize="6" pageId="14" pageNumber="13">3</subScript>
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(a few scales), M
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<subScript id="179B34102D4A856B3E82FD6225F91689" attach="left" box="[927,936,660,674]" fontSize="6" pageId="14" pageNumber="13">3</subScript>
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-CuA
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<subScript id="179B34102D4A856B3EFAFD6225A11689" attach="left" box="[999,1008,660,674]" fontSize="6" pageId="14" pageNumber="13">1</subScript>
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, and CuA
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<subScript id="179B34102D4A856B3975FD6222201689" attach="left" box="[1128,1137,660,674]" fontSize="6" pageId="14" pageNumber="13">1</subScript>
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-CuA
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<subScript id="179B34102D4A856B39ADFD6222E81689" attach="left" box="[1200,1209,660,674]" fontSize="6" pageId="14" pageNumber="13">2</subScript>
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; fringes dark graybrown, pale gray distad.
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</paragraph>
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<paragraph id="8BA036552D4A856B3DEDFD35274E1790" blockId="14.[192,1441,195,1915]" pageId="14" pageNumber="13">Ventral forewing brown, paler than on dorsum; costa, apex, and outer margin overscaled with bright red-brown, entirely filling costal, subcostal, radial, and medial cells, latter occurring distad of macule in M 3 -CuA 1, narrowing to termen at vein CuA 2; anterior central 1/3 of discal cell overscaled with yelloworange flat and setiform scales, these mixed with a few red-brown scales; anal cell gray; macules as on dorsum except macule in discal cell and CuA 1 -CuA 2 outlined with yellow-orange and yellow macule in CuA 2 -2A expanded across entire cell, making macule largest in venter, extending slightly proximad and distad of edges of macule in CuA 1 -CuA 2 but with anterioproximal portion excavate for about 1/4 width of macule.</paragraph>
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<paragraph id="8BA036552D4A856B3DEDFC3522C71030" blockId="14.[192,1441,195,1915]" pageId="14" pageNumber="13">Ventral hindwing entirely bright red-brown except much sparser on both sides of vein 2A, and as vaguely paler (yellow overscaled with red-brown) macules including medial in Sc+R 1 -Rs, a series of small postmedial macules from Rs to 2A, and a small macule at distal end of discal cell.</paragraph>
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<paragraph id="8BA036552D4A856B3DEDFBD525031150" blockId="14.[192,1441,195,1915]" pageId="14" pageNumber="13">Dorsal head mixture of gray and red-brown scales, ventral head whitish becoming dull red-brown caudad; dorsal palpi dark gray, red-brown on outer surface, black on inner surface, ventral palpi gray distad and red-brown proximad; antennae 52% of costal length, shaft black on dorsum, checkered with yellow-orange on venter, yellow-orange becoming broader distad, club 29% of length of shaft, dorsal club black, ventral club yellow-orange, nudum red-brown, 13 segments; dorsal thorax red-brown on sides, gray with green iridescence centrally cephalad and pale yellow-gray caudad; ventral thorax mixture of gray and red-brown; legs red-brown with long setiform scales, ochreous proximad, red-brown distad, protibia not spined, red-brown epiphysis extending distad to barely overlap proximal portion of tarsus, mesotibia with medium length spines, pair of spurs distad, outer about 1/2 length of inner, metatibia spined, two pairs of spurs, outer about 1/2 length of inner; dorsal abdomen black, sparse ochreous at segments; ventral abdomen and caudal end red-brown.</paragraph>
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<paragraph id="8BA036552D4A856B3DEDFA7527231370" blockId="14.[192,1441,195,1915]" pageId="14" pageNumber="13">
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Genitalia (
|
||
<figureCitation id="13242AD02D4A856B3C6DFA75279411B0" box="[368,453,1411,1435]" captionStart="Figure 63-66" captionStartId="35.[192,276,1215,1237]" captionTargetBox="[263,1362,195,1189]" captionTargetId="figure-293@35.[263,1362,192,1189]" captionTargetPageId="35" captionText="Figure 63-66. Male genitalia of Onespa (full data in text; shown are lateral view of uncus, gnathos, tegumen, and saccus; dorsal view of uncus and tegumen; ventral view of gnathos uncus; internal view of right valva; ventral view of saccus; ventral, dorsal, right lateral and/or left lateral views of aedeagus; scale bar = 1 mm). 63) O. nubis, El Salvador (SRS #4803); 64) O. brockorum, Sonora, Mexico, paratype (SRS #1852); 65) O. gala, Michoacán, Mexico (GTA #14065); 66) O. nakamura, Costa Rica, holotype (GTA #14033)." figureDoi="http://doi.org/10.5281/zenodo.5167739" httpUri="https://zenodo.org/record/5167739/files/figure.png" pageId="14" pageNumber="13">Fig. 66</figureCitation>
|
||
,
|
||
<figureCitation id="13242AD02D4A856B3CD2FA7527BC11B0" box="[463,493,1411,1435]" captionStart="Figure 75-86" captionStartId="38.[192,276,1030,1052]" captionTargetBox="[447,1192,193,1003]" captionTargetId="figure-298@38.[447,1192,193,1003]" captionTargetPageId="38" captionText="Figure 75-86. Rigid cornuti of Onespa, Buzyges, Librita, and Neposa (full data in text; shown are lateral and dorsal views). 75) O. nubis, El Salvador (SRS #4803); 76) O. gala, Michoacán, Mexico (GTA #14065); 77) O. brockorum, Sonora, Mexico, paratype (SRS #1852); 78) O. nakamura, Costa Rica, holotype (GTA #14033); 79) B. idothea, Costa Rica (ADW #97-102); 80) B. rolla, Costa Rica (GTA #14055); 81) B. benito, Oaxaca, Mexico (SRS #4468); 82) B. mellanaformis, Costa Rica, paratype (GTA #14031); 83) L. librita, Veracruz, Mexico (GTA #14135); 84) N. heras, Guerrero, Mexico (ADW #97-112); 85) N. armandoi, Oaxaca, Mexico, holotype (GTA #14143); 86) N. isabelae, Guatemala, paratype (ADW #98-15)." figureDoi="http://doi.org/10.5281/zenodo.5167749" httpUri="https://zenodo.org/record/5167749/files/figure.png" pageId="14" pageNumber="13">78</figureCitation>
|
||
) - uncus relatively long, hooked caudad in lateral view, entire and narrowing to weakly lobed caudal end in dorsal view; gnathos robust, well-separated from and shorter than uncus in lateral view, divided with arms slender, widely apart cephalad and approaching caudad in ventral view, much broader than uncus in middle; tegumen thin in lateral view, broad in dorsal view and flaring cephalad, ventral arm combining with dorsal arm of saccus, this combined structure broadest and bent ventrad of its middle; anterior arm of saccus long and thin, mostly straight, about 1.1 times length of uncus and dorsal portion of tegumen, moderately broad in ventral view and tapering gradually to pointed cephalic end; valva simple, little differentiation between costa and ampulla, latter produced dorsally to small pointed triangular process near juncture with harpe, harpe tapered to elongate, triangular, and sharply pointed caudal end, sacculus narrow, ventral edge of valva concave in middle; aedeagus straight, tubular, moderately long, about 1.1 times length of valva, caudal end slightly expanded with a short triangular projection caudad from its ventral surface, elongate triangular titillator on right side cephalad of caudal end, ventral surface with still larger and more elongate triangular keel-like titillator; vesica with cornuti including an elongate flexible spinulose pad and two large heavily sclerotized and sharply pointed spikes.
|
||
</paragraph>
|
||
<paragraph id="8BA036552D4A856B3DEDF89527B41350" blockId="14.[192,1441,195,1915]" box="[240,485,1891,1915]" pageId="14" pageNumber="13">
|
||
<emphasis id="B96BEA472D4A856B3DEDF89527041350" bold="true" box="[240,341,1891,1915]" pageId="14" pageNumber="13">Female</emphasis>
|
||
- unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30565DE2D4B856A3DDDFF3523F115B0" pageId="15" pageNumber="14" type="materials_examined">
|
||
<paragraph id="8BA036552D4B856A3DDDFF3523661570" blockId="15.[192,1440,195,347]" pageId="15" pageNumber="14">
|
||
<emphasis id="B96BEA472D4B856A3DDDFF35275D14F0" bold="true" box="[192,268,195,219]" pageId="15" pageNumber="14">Type.</emphasis>
|
||
<materialsCitation id="3B773C082D4B856A3C0FFF3522871510" collectingDate="2005-06-13" collectorName="I. Nakamura & Palpus & Not Turesis & Genitalic Vial" country="Costa Rica" elevation="1300" latitude="10.060278" location="Bajo la Hondura" longLatPrecision="21" longitude="-83.98194" pageId="15" pageNumber="14" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
|
||
<typeStatus id="54A488F72D4B856A3C0FFF3527D014F0" box="[274,385,195,219]" pageId="15" pageNumber="14" type="holotype">Holotype</typeStatus>
|
||
male with the following labels: white, printed - /
|
||
<collectingCountry id="F30876C52D4B856A3ECDFF35222C14F0" box="[976,1149,195,219]" name="Costa Rica" pageId="15" pageNumber="14">COSTA RICA</collectingCountry>
|
||
: Prov.
|
||
<collectingRegion id="49DBF8B72D4B856A39D0FF35236814F0" box="[1229,1337,195,219]" country="Costa Rica" name="San Jose" pageId="15" pageNumber="14">San José</collectingRegion>
|
||
/
|
||
<location id="8EC0608E2D4B856A3851FF35276314D0" LSID="urn:lsid:plazi:treatment:03B687432D49856A3DDDF913236D10B0:8EC0608E2D4B856A3851FF35276314D0" country="Costa Rica" latitude="10.060278" longLatPrecision="21" longitude="-83.98194" name="Bajo la Hondura" pageId="15" pageNumber="14" stateProvince="San Jose">Bajo la Hondura</location>
|
||
,
|
||
<quantity id="4CE79BB02D4B856A3C21FF15278514D0" box="[316,468,227,251]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.45" metricValueMin="1.15" pageId="15" pageNumber="14" unit="m" value="1300.0" valueMax="1450.0" valueMin="1150.0">
|
||
<elevation id="0032D1662D4B856A3C21FF15278514D0" box="[316,468,227,251]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.45" metricValueMin="1.15" pageId="15" pageNumber="14" unit="m" value="1300.0" valueMax="1450.0" valueMin="1150.0">1150-1450m</elevation>
|
||
</quantity>
|
||
/
|
||
<geoCoordinate id="EE2B50922D4B856A3CFAFF15242014D0" box="[487,625,226,251]" degrees="10" direction="north" minutes="03" orientation="latitude" pageId="15" pageNumber="14" precision="15" seconds="37" value="10.060278">
|
||
10
|
||
<superScript id="7C6A9B1D2D4B856A3F18FF14245C14DB" attach="none" box="[517,525,226,240]" fontSize="6" pageId="15" pageNumber="14">o</superScript>
|
||
03’37”N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2B50922D4B856A3F61FF15255614D0" box="[636,775,226,251]" degrees="83" direction="west" minutes="58" orientation="longitude" pageId="15" pageNumber="14" precision="15" seconds="55" value="-83.98194">
|
||
83
|
||
<superScript id="7C6A9B1D2D4B856A3F87FF1424F314DB" attach="none" box="[666,674,226,240]" fontSize="6" pageId="15" pageNumber="14">o</superScript>
|
||
58’55”W
|
||
</geoCoordinate>
|
||
/
|
||
<date id="FFA110952D4B856A3E06FF1525F714D0" box="[795,934,227,251]" pageId="15" pageNumber="14" value="2005-06-13">
|
||
<collectingDate id="EFE5E97D2D4B856A3E06FF1525F714D0" box="[795,934,227,251]" pageId="15" pageNumber="14" value="2005-06-13">13. vi. 2005</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="26EA53832D4B856A3EA4FF12220914D0" box="[953,1112,227,251]" pageId="15" pageNumber="14">I. Nakamura</collectorName>
|
||
leg. /; white, handprinted - /
|
||
<collectorName id="26EA53832D4B856A3DCDFEF527771530" box="[208,294,259,283]" pageId="15" pageNumber="14">Palpus</collectorName>
|
||
attached /; white, handprinted - / n=7/7 / not spined / no brand /; white, handprinted /
|
||
<collectorName id="26EA53832D4B856A386FFEF2274C1510" pageId="15" pageNumber="14">Not Turesis</collectorName>
|
||
/; white, printed and handprinted - /
|
||
<collectorName id="26EA53832D4B856A3FC0FED525D91510" box="[733,904,291,315]" pageId="15" pageNumber="14">Genitalic Vial</collectorName>
|
||
/ GTA-14033 /; red, printed
|
||
</materialsCitation>
|
||
- /
|
||
<materialsCitation id="3B773C082D4B856A39EAFED523631570" collectionCode="MGCL" collectorName="Austin & A. Warren" country="Costa Rica" pageId="15" pageNumber="14" specimenCount="2" stateProvince="San Jose" typeStatus="holotype">
|
||
<typeStatus id="54A488F72D4B856A39EAFED523C21510" box="[1271,1427,291,315]" pageId="15" pageNumber="14" type="holotype">HOLOTYPE</typeStatus>
|
||
/
|
||
<taxonomicName id="4C1F4DD62D4B856A3DDDFEB227F51570" authorityName="Austin and A. Warren" authorityYear="2009" box="[192,420,323,347]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nakamura">
|
||
<emphasis id="B96BEA472D4B856A3DDDFEB227F51570" box="[192,420,323,347]" italics="true" pageId="15" pageNumber="14">Onespa nakamura</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<collectorName id="26EA53832D4B856A3CABFEB524581570" box="[438,521,323,347]" pageId="15" pageNumber="14">Austin</collectorName>
|
||
&
|
||
<collectorName id="26EA53832D4B856A3F36FEB524FB1570" box="[555,682,323,347]" pageId="15" pageNumber="14">A. Warren</collectorName>
|
||
/, deposited at
|
||
<collectionCode id="ED0EAE902D4B856A3E42FEB525E71570" box="[863,950,323,347]" pageId="15" pageNumber="14">MGCL</collectionCode>
|
||
. Known only from the
|
||
<typeStatus id="54A488F72D4B856A39D8FEB523631570" box="[1221,1330,323,347]" pageId="15" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA036552D4B856A3DDDFE7523F115B0" blockId="15.[192,1440,386,411]" box="[192,1440,386,411]" pageId="15" pageNumber="14">
|
||
<materialsCitation id="3B773C082D4B856A3DDDFE75255715B0" box="[192,774,387,411]" country="Costa Rica" location="San Jose Province" pageId="15" pageNumber="14" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
|
||
<emphasis id="B96BEA472D4B856A3DDDFE75272715B0" bold="true" box="[192,374,387,411]" pageId="15" pageNumber="14">
|
||
<typeStatus id="54A488F72D4B856A3DDDFE75275515B0" box="[192,260,387,411]" pageId="15" pageNumber="14">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F30876C52D4B856A3C67FE75247515B0" box="[378,548,387,411]" name="Costa Rica" pageId="15" pageNumber="14">COSTA RICA</collectingCountry>
|
||
:
|
||
<collectingRegion id="49DBF8B72D4B856A3F31FE75255715B0" box="[556,774,387,411]" country="Costa Rica" name="San Jose" pageId="15" pageNumber="14">San José Province</collectingRegion>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B773C082D4B856A3E0DFE7523CC15B0" box="[784,1437,386,411]" country="Costa Rica" elevation="1300" latitude="10.060278" location="Bajo la Hondura" longLatPrecision="21" longitude="-83.98194" pageId="15" pageNumber="14" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
|
||
<location id="8EC0608E2D4B856A3E0DFE75258715B0" LSID="urn:lsid:plazi:treatment:03B687432D49856A3DDDF913236D10B0:8EC0608E2D4B856A3E0DFE75258715B0" box="[784,982,387,411]" country="Costa Rica" latitude="10.060278" longLatPrecision="21" longitude="-83.98194" name="Bajo la Hondura" pageId="15" pageNumber="14" stateProvince="San Jose">Bajo la Hondura</location>
|
||
,
|
||
<quantity id="4CE79BB02D4B856A3EFDFE75222615B0" box="[992,1143,387,411]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.45" metricValueMin="1.15" pageId="15" pageNumber="14" unit="m" value="1300.0" valueMax="1450.0" valueMin="1150.0">
|
||
<elevation id="0032D1662D4B856A3EFDFE75222615B0" box="[992,1143,387,411]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.45" metricValueMin="1.15" pageId="15" pageNumber="14" unit="m" value="1300.0" valueMax="1450.0" valueMin="1150.0">1150-1450m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="EE2B50922D4B856A399DFE75235615B0" box="[1152,1287,386,411]" degrees="10" direction="north" minutes="03" orientation="latitude" pageId="15" pageNumber="14" precision="15" seconds="37" value="10.060278">
|
||
10
|
||
<superScript id="7C6A9B1D2D4B856A3983FE7422F715BB" attach="right" box="[1182,1190,386,400]" fontSize="6" pageId="15" pageNumber="14">o</superScript>
|
||
03’37”N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2B50922D4B856A380CFE7523CC15B0" box="[1297,1437,386,411]" degrees="83" direction="west" minutes="58" orientation="longitude" pageId="15" pageNumber="14" precision="15" seconds="55" value="-83.98194">
|
||
83
|
||
<superScript id="7C6A9B1D2D4B856A3833FE74236715BB" attach="none" box="[1326,1334,386,400]" fontSize="6" pageId="15" pageNumber="14">o</superScript>
|
||
58’55”W
|
||
</geoCoordinate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30565DE2D4B856A3DDDFE35258715D0" pageId="15" pageNumber="14" type="etymology">
|
||
<paragraph id="8BA036552D4B856A3DDDFE35258715D0" blockId="15.[192,1440,451,507]" pageId="15" pageNumber="14">
|
||
<emphasis id="B96BEA472D4B856A3DDDFE35270C15F0" bold="true" box="[192,349,451,475]" pageId="15" pageNumber="14">Etymology.</emphasis>
|
||
It is our pleasure to name this species after Ichiro Nakamura who generously and graciously allowed us to study his collections of
|
||
<taxonomicName id="4C1F4DD62D4B856A3FCCFE15253415D0" box="[721,869,483,507]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="family">Hesperiidae</taxonomicName>
|
||
in depth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30565DE2D4B856A3DDDFDD523CD1610" box="[192,1436,547,571]" pageId="15" pageNumber="14" type="distribution">
|
||
<paragraph id="8BA036552D4B856A3DDDFDD523CD1610" blockId="15.[192,1436,547,571]" box="[192,1436,547,571]" pageId="15" pageNumber="14">
|
||
<emphasis id="B96BEA472D4B856A3DDDFDD524171610" bold="true" box="[192,582,547,571]" pageId="15" pageNumber="14">Distribution and phenology.</emphasis>
|
||
The species is known at present only from the
|
||
<typeStatus id="54A488F72D4B856A3960FDD522B61610" box="[1149,1255,547,571]" pageId="15" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
taken in June.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30565DE2D4B856A3DDDFD95236D10B0" pageId="15" pageNumber="14" type="diagnosis">
|
||
<paragraph id="8BA036552D4B856A3DDDFD9527B117F7" blockId="15.[192,1440,611,1179]" pageId="15" pageNumber="14">
|
||
<emphasis id="B96BEA472D4B856A3DDDFD9524771650" bold="true" box="[192,550,611,635]" pageId="15" pageNumber="14">Diagnosis and discussion.</emphasis>
|
||
<taxonomicName id="4C1F4DD62D4B856A3F30FD9225421650" authorityName="Austin and A. Warren" authorityYear="2009" box="[557,787,611,635]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nakamura">
|
||
<emphasis id="B96BEA472D4B856A3F30FD9225421650" box="[557,787,611,635]" italics="true" pageId="15" pageNumber="14">Onespa nakamura</emphasis>
|
||
</taxonomicName>
|
||
, discovered as a single male among a collection from
|
||
<collectingCountry id="F30876C52D4B856A3DDDFD75271516B0" box="[192,324,643,667]" name="Costa Rica" pageId="15" pageNumber="14">Costa Rica</collectingCountry>
|
||
, prompted this re-examination of
|
||
<taxonomicName id="4C1F4DD62D4B856A3FCAFD72256316B0" authorityName="Steinhauser" authorityYear="1974" box="[727,818,644,667]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96BEA472D4B856A3FCAFD72256316B0" box="[727,818,644,667]" italics="true" pageId="15" pageNumber="14">Onespa</emphasis>
|
||
</taxonomicName>
|
||
. Color and pattern of
|
||
<taxonomicName id="4C1F4DD62D4B856A3929FD72228416B0" authorityName="Austin and A. Warren" authorityYear="2009" box="[1076,1237,643,667]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nakamura">
|
||
<emphasis id="B96BEA472D4B856A3929FD72228416B0" box="[1076,1237,643,667]" italics="true" pageId="15" pageNumber="14">O. nakamura</emphasis>
|
||
</taxonomicName>
|
||
are immediately reminiscent of
|
||
<taxonomicName id="4C1F4DD62D4B856A3C6AFD52244F1690" authorityName="Steinhasuser" authorityYear="1974" box="[375,542,675,699]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nubis">
|
||
<emphasis id="B96BEA472D4B856A3C6AFD52244F1690" box="[375,542,675,699]" italics="true" pageId="15" pageNumber="14">Onespa nubis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1F4DD62D4B856A3F44FD5224AC1690" authorityName="Austin and A. Warren" authorityYear="2009" box="[601,765,675,699]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nakamura">
|
||
<emphasis id="B96BEA472D4B856A3F44FD5224AC1690" box="[601,765,675,699]" italics="true" pageId="15" pageNumber="14">O. nakamura</emphasis>
|
||
</taxonomicName>
|
||
is the largest species of the genus. Affinity was further strengthened by the absence of a stigma and morphology of the male genitalia. These latter, while within the general gestalt of its congeners, differ from all by the more elongate uncus, gnathos, and valva and the prominent keel-like titillator on the ventral surface of the aedeagus.
|
||
<taxonomicName id="4C1F4DD62D4B856A3918FCF222B81730" authorityName="Austin and A. Warren" authorityYear="2009" box="[1029,1257,771,795]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nakamura">
|
||
<emphasis id="B96BEA472D4B856A3918FCF222B81730" box="[1029,1257,771,795]" italics="true" pageId="15" pageNumber="14">Onespa nakamura</emphasis>
|
||
</taxonomicName>
|
||
shares with
|
||
<taxonomicName id="4C1F4DD62D4B856A3899FCF227521710" authorityName="Steinhasuser" authorityYear="1974" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nubis">
|
||
<emphasis id="B96BEA472D4B856A3899FCF227521710" italics="true" pageId="15" pageNumber="14">O. nubis</emphasis>
|
||
</taxonomicName>
|
||
the presence of two prominent cornutal spikes and an elongate flexible cornutus.
|
||
<taxonomicName id="4C1F4DD62D4B856A39D5FCD2230D1710" authorityName="Austin & Warren" authorityYear="2009" baseAuthorityName="Godman" baseAuthorityYear="1900" box="[1224,1372,803,827]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="gala">
|
||
<emphasis id="B96BEA472D4B856A39D5FCD2230D1710" box="[1224,1372,803,827]" italics="true" pageId="15" pageNumber="14">Onespa gala</emphasis>
|
||
</taxonomicName>
|
||
has a single spike and
|
||
<taxonomicName id="4C1F4DD62D4B856A3C94FCB2247E1770" authorityName="Austin and A. Warren" authorityYear="2009" box="[393,559,835,859]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="brockorum">
|
||
<emphasis id="B96BEA472D4B856A3C94FCB2247E1770" box="[393,559,835,859]" italics="true" pageId="15" pageNumber="14">O. brockorum</emphasis>
|
||
</taxonomicName>
|
||
has a small spike in addition to the robust one; both have a much smaller flexible cornutus than either
|
||
<taxonomicName id="4C1F4DD62D4B856A3F36FC9224C51750" authorityName="Steinhasuser" authorityYear="1974" box="[555,660,867,891]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nubis">
|
||
<emphasis id="B96BEA472D4B856A3F36FC9224C51750" box="[555,660,867,891]" italics="true" pageId="15" pageNumber="14">O. nubis</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C1F4DD62D4B856A3FA1FC9225331750" authorityName="Austin and A. Warren" authorityYear="2009" box="[700,866,867,891]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nakamura">
|
||
<emphasis id="B96BEA472D4B856A3FA1FC9225331750" box="[700,866,867,891]" italics="true" pageId="15" pageNumber="14">O. nakamura</emphasis>
|
||
</taxonomicName>
|
||
. The right lateral titillator on
|
||
<taxonomicName id="4C1F4DD62D4B856A39C3FC9223D51750" authorityName="Austin and A. Warren" authorityYear="2009" box="[1246,1412,867,891]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nakamura">
|
||
<emphasis id="B96BEA472D4B856A39C3FC9223D51750" box="[1246,1412,867,891]" italics="true" pageId="15" pageNumber="14">O. nakamura</emphasis>
|
||
</taxonomicName>
|
||
is large. This structure is much smaller on
|
||
<taxonomicName id="4C1F4DD62D4B856A3FB3FC72255717B0" authorityName="Austin & Warren" authorityYear="2009" baseAuthorityName="Godman" baseAuthorityYear="1900" box="[686,774,899,923]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="gala">
|
||
<emphasis id="B96BEA472D4B856A3FB3FC72255717B0" box="[686,774,899,923]" italics="true" pageId="15" pageNumber="14">O. gala</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1F4DD62D4B856A3E5DFC7225B617B0" authorityName="Austin and A. Warren" authorityYear="2009" box="[832,999,899,923]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="brockorum">
|
||
<emphasis id="B96BEA472D4B856A3E5DFC7225B617B0" box="[832,999,899,923]" italics="true" pageId="15" pageNumber="14">O. brockorum</emphasis>
|
||
</taxonomicName>
|
||
and absent on
|
||
<taxonomicName id="4C1F4DD62D4B856A3980FC72235217B0" authorityName="Steinhasuser" authorityYear="1974" box="[1181,1283,899,923]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nubis">
|
||
<emphasis id="B96BEA472D4B856A3980FC72235217B0" box="[1181,1283,899,923]" italics="true" pageId="15" pageNumber="14">O. nubis</emphasis>
|
||
</taxonomicName>
|
||
. In addition,
|
||
<taxonomicName id="4C1F4DD62D4B856A3DDDFC5227F41790" authorityName="Austin and A. Warren" authorityYear="2009" box="[192,421,931,955]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nakamura">
|
||
<emphasis id="B96BEA472D4B856A3DDDFC5227F41790" box="[192,421,931,955]" italics="true" pageId="15" pageNumber="14">Onespa nakamura</emphasis>
|
||
</taxonomicName>
|
||
has a triangular keel-like titillator on ventral surface of the aedeagus, a structure absent on its congeners.
|
||
</paragraph>
|
||
<paragraph id="8BA036552D4B856A3DEDFC12236D10B0" blockId="15.[192,1440,611,1179]" pageId="15" pageNumber="14">
|
||
In the presence of tibial spines,
|
||
<taxonomicName id="4C1F4DD62D4B856A3F64FC12254E17D0" authorityName="Austin and A. Warren" authorityYear="2009" box="[633,799,995,1019]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nakamura">
|
||
<emphasis id="B96BEA472D4B856A3F64FC12254E17D0" box="[633,799,995,1019]" italics="true" pageId="15" pageNumber="14">O. nakamura</emphasis>
|
||
</taxonomicName>
|
||
differs from the other three species included within
|
||
<taxonomicName id="4C1F4DD62D4B856A3DDDFBF2274D1030" authorityName="Steinhauser" authorityYear="1974" box="[192,284,1028,1051]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96BEA472D4B856A3DDDFBF2274D1030" box="[192,284,1028,1051]" italics="true" pageId="15" pageNumber="14">Onespa</emphasis>
|
||
</taxonomicName>
|
||
. That character, generally a useful taxonomic trait among hesperiines (
|
||
<emphasis id="B96BEA472D4B856A399EFBF222FD1030" box="[1155,1196,1028,1051]" italics="true" pageId="15" pageNumber="14">e.g.</emphasis>
|
||
,
|
||
<bibRefCitation id="EF8E4BA42D4B856A39A4FBF2231B1030" author="Evans, W. H." box="[1209,1354,1027,1051]" pageId="15" pageNumber="14" refId="ref46128" refString="Evans, W. H. 1955. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV (groups H to P), Hesperiinae and Megathyminae. British Museum (Natural History): London. 499 p." type="book" year="1955">Evans 1955</bibRefCitation>
|
||
), is not here considered to exclude
|
||
<taxonomicName id="4C1F4DD62D4B856A3F17FBD224FE1010" authorityName="Austin and A. Warren" authorityYear="2009" box="[522,687,1059,1083]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nakamura">
|
||
<emphasis id="B96BEA472D4B856A3F17FBD224FE1010" box="[522,687,1059,1083]" italics="true" pageId="15" pageNumber="14">O. nakamura</emphasis>
|
||
</taxonomicName>
|
||
from the genus. Other traits, including color and pattern, absence of a stigma, and especially genital morphology are consistently within the concept of
|
||
<taxonomicName id="4C1F4DD62D4B856A39FAFBB223101070" authorityName="Steinhauser" authorityYear="1974" box="[1255,1345,1092,1115]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96BEA472D4B856A39FAFBB223101070" box="[1255,1345,1092,1115]" italics="true" pageId="15" pageNumber="14">Onespa</emphasis>
|
||
</taxonomicName>
|
||
and not with other potentially related genera (see general discussion below). Discovery of a female of
|
||
<taxonomicName id="4C1F4DD62D4B856A381CFB9223F11050" authorityName="Austin and A. Warren" authorityYear="2009" box="[1281,1440,1123,1147]" class="Insecta" family="Hesperiidae" genus="Onespa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="nakamura">
|
||
<emphasis id="B96BEA472D4B856A381CFB9223F11050" box="[1281,1440,1123,1147]" italics="true" pageId="15" pageNumber="14">O. nakamura</emphasis>
|
||
</taxonomicName>
|
||
and examination of her genitalia, however, are essential for its unequivocal generic association.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |