180 lines
21 KiB
XML
180 lines
21 KiB
XML
<document id="66FE9D1E79E06A5AE8C37F656CEBCC16" ID-DOI="10.11646/zootaxa.3857.1.3" ID-GBIF-Dataset="23e68cbb-17b7-4dea-aad0-7360b8766cc0" ID-ISSN="1175-5326" ID-Zenodo-Dep="228181" ID-ZooBank="A26948F7-39C6-4858-B7FD-380E12F9BD34" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460895922835" checkinUser="plazi" docAuthor="Siddiqui, Pirzada Jamal, Amir, Shabir Ali & Masroor, Rafaqat" docDate="2014" docId="03B5EA0A0F11CB0BFF341AFEFBB2FF79" docLanguage="en" docName="zt03857p100.pdf" docOrigin="Zootaxa 3857 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acanthopagrus sheim Iwatsuki 2013" docType="treatment" docVersion="8" lastPageNumber="78" masterDocId="FF8C92720F17CB03FFA31C27FFBAFF81" masterDocTitle="The sparid fishes of Pakistan, with new distribution records" masterLastPageNumber="100" masterPageNumber="71" pageNumber="77" updateTime="1698360089480" updateUser="plazi">
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<mods:titleInfo id="D6A2DEA8C3CD07EE7ACB6B98FA9B2C98">
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<mods:title id="9CAD164D7F37E73266E57527111E4B75">The sparid fishes of Pakistan, with new distribution records</mods:title>
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<mods:roleTerm id="75C0EBC3133DDB80FB1158611A2BC5C2">Author</mods:roleTerm>
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<mods:namePart id="064EA2F49FA0EA3F97D20048910D12F2">Siddiqui, Pirzada Jamal</mods:namePart>
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<mods:namePart id="736A3B7D92B5072B5BD3A6E53B0B74BD">Amir, Shabir Ali</mods:namePart>
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<mods:namePart id="0EAF7B2F174C241DA7ABC9ED0C84A773">Masroor, Rafaqat</mods:namePart>
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<mods:title id="8DA833DF1B1B9B72F91E1484B9E3F86B">Zootaxa</mods:title>
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<mods:part id="07BC18E451013EFC3BC2C0E70F8BEECD">
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<mods:date id="E7529CD76D166730AC65F62B22CD6271">2014</mods:date>
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<mods:number id="D6F850A12C832AC0E6CA97D8330668D6">3857</mods:number>
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<mods:identifier id="AE8A28C64265867EE5C65B29A55F25FC" type="DOI">10.11646/zootaxa.3857.1.3</mods:identifier>
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<treatment id="03B5EA0A0F11CB0BFF341AFEFBB2FF79" ID-DOI="http://doi.org/10.5281/zenodo.6139397" ID-GBIF-Taxon="119587517" ID-Zenodo-Dep="6139397" LSID="urn:lsid:plazi:treatment:03B5EA0A0F11CB0BFF341AFEFBB2FF79" httpUri="http://treatment.plazi.org/id/03B5EA0A0F11CB0BFF341AFEFBB2FF79" lastPageId="8" lastPageNumber="78" pageId="6" pageNumber="77">
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<subSubSection id="C30608970F11CB05FF341AFEFE54F8DC" pageId="6" pageNumber="77" type="nomenclature">
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<paragraph id="8BA35B1C0F11CB05FF341AFEFDE7F972" blockId="6.[151,807,1753,1885]" box="[151,605,1753,1779]" pageId="6" pageNumber="77">
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<heading id="D0EBEC700F11CB05FF341AFEFDE7F972" bold="true" box="[151,605,1753,1779]" fontSize="11" level="1" pageId="6" pageNumber="77" reason="1">
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<taxonomicName id="4C1C209F0F11CB05FF341AFEFDE7F972" ID-CoL="8YVS" authority="Iwatsuki, 2013" authorityName="Iwatsuki" authorityYear="2013" box="[151,605,1753,1779]" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="77" phylum="Chordata" rank="species" species="sheim">
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<emphasis id="B968870E0F11CB05FF341AFEFDE7F972" bold="true" box="[151,605,1753,1779]" pageId="6" pageNumber="77">
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<emphasis id="B968870E0F11CB05FF341AFEFE1AF972" bold="true" box="[151,416,1753,1779]" italics="true" pageId="6" pageNumber="77">Acanthopagrus sheim</emphasis>
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Iwatsuki, 2013
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BA35B1C0F11CB05FF341ADAFE54F8DC" blockId="6.[151,807,1753,1885]" pageId="6" pageNumber="77">
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Common name: Spotted Yellowfin Seabream Local names: Dandya, Nehra (Sindh); Tintle (Balochistan) (
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<figureCitation id="132747990F11CB05FF3D1B63FEB3F8DC" box="[158,265,1860,1885]" captionStart="FIGURE 4" captionStartId="7.[151,250,1088,1110]" captionTargetBox="[329,1257,447,1066]" captionTargetId="figure@7.[329,1257,447,1066]" captionTargetPageId="7" captionText="FIGURE 4. Acanthopagrus sheim, PMNH 53008, 196 mm SL, WWFH, Karachi. Faint yellowish color in pelvic and anal fin and whitish lower margin of lower caudal lobe." httpUri="https://zenodo.org/record/228185/files/figure.png" pageId="6" pageNumber="77">Figures 4</figureCitation>
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&
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<figureCitation id="132747990F11CB05FE8E1B63FEF3F8DC" box="[301,329,1860,1885]" captionStart="FIGURE 15" captionStartId="20.[151,250,1789,1811]" captionTargetBox="[362,1223,1064,1752]" captionTargetId="figure@20.[351,1235,1056,1766]" captionTargetPageId="20" captionText="FIGURE 15. Details of dentition (A-Y) and gill rakers (Z & Z 1) in species of family Sparidae from Pakistan; Acanthopagrus catenula, A) upper jaw & B) lower jaw; A. berda, C) upper jaw & D) lower jaw; A. arabicus, E) upper jaw & F) lower jaw; A. sheim, G) upper jaw & H) lower jaw; Sparidentex hasta, I) upper jaw & J) lower jaw; Sparidentex jamalensis., K) upper jaw & L) lower jaw; Crenidens indicus, M) upper jaw & N) lower jaw; Diplodus capensis, O) upper jaw & P) lower jaw; Q) upper and lower jaws of Diplodus omanensis; Rhabdosargus sarba, R) upper jaw & S) lower jaw; Rhabdosargus haffara, T) upper jaw & U) lower jaw; V) upper and lower jaws of Agryrops spinifer; W) upper and lower jaws of Cheimerius nufar; Pagellus affinis, X) upper jaw & Y) lower jaw; Z) Argyrops spinifer, showing comparatively small gill rakers; and Z 1) Cheimerius nufar, showing elongated fan-like gill rakers." httpUri="https://zenodo.org/record/228196/files/figure.png" pageId="6" pageNumber="77">15</figureCitation>
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G, H;
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<tableCitation id="C69E6EA70F11CB05FE311B63FE58F8DC" box="[402,482,1860,1885]" captionStart="TABLE 1" captionStartId="21.[151,239,811,834]" captionTargetBox="[159,1436,893,1173]" captionTargetPageId="21" captionText="TABLE 1. Counts and morphometric data of sparid species occurring in the coastal waters of Pakistan; mean in parenthesis." httpUri="http://table.plazi.org/id/DF630B940F02CB16FF341F0CFEACFCE3" pageId="6" pageNumber="77" tableUuid="DF630B940F02CB16FF341F0CFEACFCE3">Table 1</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30608970F11CB05FF341BACFBBFF824" box="[151,1029,1931,1957]" pageId="6" pageNumber="77" type="reference_group">
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<paragraph id="8BA35B1C0F11CB05FF341BACFBBFF824" blockId="6.[151,1437,1931,2029]" box="[151,1029,1931,1957]" pageId="6" pageNumber="77">
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<taxonomicName id="4C1C209F0F11CB05FF341BACFDD4F824" ID-CoL="8YVS" authority="Iwatsuki, 2013: 85" authorityName="Iwatsuki" authorityPageNumber="85" authorityYear="2013" box="[151,622,1931,1957]" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="77" phylum="Chordata" rank="species" species="sheim">
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<emphasis id="B968870E0F11CB05FF341BACFE2EF825" bold="true" box="[151,404,1931,1956]" italics="true" pageId="6" pageNumber="77">Acanthopagrus sheim</emphasis>
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Iwatsuki, 2013: 85
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</taxonomicName>
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,
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<figureCitation id="132747990F11CB05FDD91BABFD06F824" box="[634,700,1932,1957]" captionStart="FIGURE 4" captionStartId="7.[151,250,1088,1110]" captionTargetBox="[329,1257,447,1066]" captionTargetId="figure@7.[329,1257,447,1066]" captionTargetPageId="7" captionText="FIGURE 4. Acanthopagrus sheim, PMNH 53008, 196 mm SL, WWFH, Karachi. Faint yellowish color in pelvic and anal fin and whitish lower margin of lower caudal lobe." httpUri="https://zenodo.org/record/228185/files/figure.png" pageId="6" pageNumber="77">Fig. 4</figureCitation>
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(c) (off Hawar Island, Oatar).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30608970F11CB04FF641B88FC19FE0A" lastPageId="7" lastPageNumber="78" pageId="6" pageNumber="77" type="diagnosis">
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<paragraph id="8BA35B1C0F11CB04FF641B88FC19FE0A" blockId="6.[151,1437,1931,2029]" lastBlockId="7.[151,1436,151,395]" lastPageId="7" lastPageNumber="78" pageId="6" pageNumber="77">
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<emphasis id="B968870E0F11CB05FF641B88FEFBF849" bold="true" box="[199,321,1967,1992]" pageId="6" pageNumber="77">Diagnosis.</emphasis>
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Distinguished from all congeners of
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<taxonomicName id="4C1C209F0F11CB05FD531B97FC21F849" box="[752,923,1968,1992]" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="77" phylum="Chordata" rank="genus">
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<emphasis id="B968870E0F11CB05FD531B97FC21F849" box="[752,923,1968,1992]" italics="true" pageId="6" pageNumber="77">Acanthopagrus</emphasis>
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</taxonomicName>
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by the following combination of characters: Body relatively deep and thicker, its depth 42–45% and width at pectoral base 16–20% of SL. Dorsal-fin rays XI–XII, 10–11 (mostly XI, 11); anal-fin rays III, 8; pored lateral line scales 42–44; scale rows above and below lateral line 4½–5½ and 11½–12½; scale rows between the fifth dorsal-fin spine base and lateral line 4½; no scales on preopercular flange; gill rakers 16–18 whereas 9–10 on the lower limb; 2nd anal-fin spine (2
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<collectingCountry id="F30B1B8C0F10CB04FB671CC4FB50FF7A" box="[1220,1258,227,251]" name="American Samoa" pageId="7" pageNumber="78">AS</collectingCountry>
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) longer than 3rd anal-fin spine, its ratio (2
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<collectingCountry id="F30B1B8C0F10CB04FE6A1D20FE57FE9E" box="[457,493,263,287]" name="American Samoa" pageId="7" pageNumber="78">AS</collectingCountry>
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/ 3
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<collectingCountry id="F30B1B8C0F10CB04FDB01D20FD83FE9E" box="[531,569,263,287]" name="American Samoa" pageId="7" pageNumber="78">AS</collectingCountry>
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) 1.2–1.6 (mean 1.4); least infraorbital depth 4.9–6.9% of SL; two to three black blotches just beneath inter-radial membranes between dorsal-fin rays, upper dorsal-fin membrane usually with darker margin; pelvic and anal-fins yellow; black streaks usually present on inter-radial membranes between yellow anal-fin rays; weak diffuse dark blotch at origin of lateral line.
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</paragraph>
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</subSubSection>
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<caption id="DF630B940F10CB04FF341867FDCDFBF4" httpUri="https://zenodo.org/record/228185/files/figure.png" pageId="7" pageNumber="78" targetBox="[329,1257,447,1066]" targetPageId="7">
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<paragraph id="8BA35B1C0F10CB04FF341867FDCDFBF4" blockId="7.[151,1435,1088,1141]" pageId="7" pageNumber="78">
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<emphasis id="B968870E0F10CB04FF341867FEA9FBD4" bold="true" box="[151,275,1088,1110]" pageId="7" pageNumber="78">FIGURE 4.</emphasis>
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<taxonomicName id="4C1C209F0F10CB04FEB91867FE4DFBD7" box="[282,503,1088,1110]" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="78" phylum="Chordata" rank="species" species="sheim">
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<emphasis id="B968870E0F10CB04FEB91867FE4DFBD7" box="[282,503,1088,1110]" italics="true" pageId="7" pageNumber="78">Acanthopagrus sheim</emphasis>
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</taxonomicName>
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, PMNH 53008, 196 mm SL, WWFH, Karachi. Faint yellowish color in pelvic and anal fin and whitish lower margin of lower caudal lobe.
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</paragraph>
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</caption>
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<subSubSection id="C30608970F10CB04FF6418BCFA21F9A2" pageId="7" pageNumber="78" type="description">
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<paragraph id="8BA35B1C0F10CB04FF6418BCFA21F9A2" blockId="7.[151,1437,1179,2003]" pageId="7" pageNumber="78">
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<emphasis id="B968870E0F10CB04FF6418BCFEE3FB35" bold="true" box="[199,345,1179,1204]" pageId="7" pageNumber="78">Description.</emphasis>
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Body deep, compressed; head 31–37% of SL, eye moderate in size, orbit diameter 7.9–9.4% of SL; dorsal profile of head steep and convex ascending sharply from just above eye; snout pointed; mouth slightly oblique, lips thick; maxilla reaching vertical at rear edge of pupil, longer than orbit diameter; upper-jaw length 12.8–14.6% of SL; lower jaw slightly included; four to six incisor like teeth (canines like in adults) in upper and lower jaws; three to five rows of well-developed molariform teeth in each side of both jaws, smaller anteriorly and larger posteriorly; four to five transverse rows of scales on cheek; preopercular flange lacking scales, posterior margin weakly serrated; low scaly sheath on bases of soft dorsal and anal-fin ray; dorsal-fin spines strong, first one is smaller (6.5–8% of SL), 4th one is longer (16–20.2% of SL); first anal-fin spine short (5.1–7.3% of SL), 2nd analfin spine heavy and long (17.1–22 % of SL), 3rd anal-fin spine thin and shorter (12.6–16.5 % of SL) than 2nd anal-fin spine; pectoral-fin tip nearly reaching level with first anal-fin spine base vertically, its length greater than head length; first pelvic-fin ray length clearly greater than third anal-fin spine; pelvic-fin spine longer than snout length.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30608970F10CB04FF641A09FC4AF89F" pageId="7" pageNumber="78" type="materials_examined">
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<paragraph id="8BA35B1C0F10CB04FF641A09FC4AF89F" blockId="7.[151,1437,1179,2003]" pageId="7" pageNumber="78">
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<emphasis id="B968870E0F10CB04FF641A09FE49F9C7" box="[199,499,1582,1606]" italics="true" pageId="7" pageNumber="78">Color of fresh specimens</emphasis>
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. Head and body silvery pale grey. Belly whitish silvery. Faint streaks along longitudinal rows of body scales in comparison to visible streaks in
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<taxonomicName id="4C1C209F0F10CB04FC301A75FBAEF9EB" box="[915,1044,1618,1642]" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="78" phylum="Chordata" rank="species" species="arabicus">
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<emphasis id="B968870E0F10CB04FC301A75FBAEF9EB" box="[915,1044,1618,1642]" italics="true" pageId="7" pageNumber="78">A. arabicus</emphasis>
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</taxonomicName>
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. Conspicuous diffused dark blotch at origin of lateral line. Dorsal-fin greyish to hyaline and upper margin of dorsal-fin membrane little blackish, dorsal-fin membrane between spines and soft rays with two to three black spots. Pelvic and anal-fins light yellow, but not bright yellow as in
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<taxonomicName id="4C1C209F0F10CB04FE741A99FDE6F957" box="[471,604,1726,1750]" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="78" phylum="Chordata" rank="species" species="arabicus">
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<emphasis id="B968870E0F10CB04FE741A99FDE6F957" box="[471,604,1726,1750]" italics="true" pageId="7" pageNumber="78">A. arabicus</emphasis>
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</taxonomicName>
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. Black streaks on inter-radial membranes between yellow anal-fin rays. Caudal fin blackish with some yellowish to greyish color, fin rays in terminal lower caudal fin lobe whitish. Pectoral-fins somewhat whitish hyaline, pale blotch at origin of pectoral-fin.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30608970F10CB04FF641B0EFE27F8E7" pageId="7" pageNumber="78" type="distribution">
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<paragraph id="8BA35B1C0F10CB04FF641B0EFE27F8E7" blockId="7.[151,1437,1179,2003]" pageId="7" pageNumber="78">
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<emphasis id="B968870E0F10CB04FF641B0EFEDAF8C3" bold="true" box="[199,352,1833,1858]" pageId="7" pageNumber="78">Distribution.</emphasis>
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<taxonomicName id="4C1C209F0F10CB04FECB1B0DFDE6F8C3" box="[360,604,1834,1858]" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="78" phylum="Chordata" rank="species" species="sheim">
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<emphasis id="B968870E0F10CB04FECB1B0DFDE6F8C3" box="[360,604,1834,1858]" italics="true" pageId="7" pageNumber="78">Acanthopagrus sheim</emphasis>
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</taxonomicName>
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is only known from The Gulf and
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<collectingCountry id="F30B1B8C0F10CB04FC491B0DFBF7F8C2" box="[1002,1101,1834,1859]" name="Pakistan" pageId="7" pageNumber="78">Pakistan</collectingCountry>
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. This species is very likely to occur in
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<collectingCountry id="F30B1B8C0F10CB04FF5A1B68FE9CF8E7" box="[249,294,1871,1894]" name="Iran" pageId="7" pageNumber="78">Iran</collectingCountry>
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and
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<collectingCountry id="F30B1B8C0F10CB04FEFF1B6AFE23F8E7" box="[348,409,1869,1894]" name="India" pageId="7" pageNumber="78">India</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30608970F10CB0BFF641B55FBB2FF79" lastPageId="8" lastPageNumber="79" pageId="7" pageNumber="78" type="discussion">
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<paragraph id="8BA35B1C0F10CB0BFF641B55FBB2FF79" blockId="7.[151,1437,1179,2003]" lastBlockId="8.[151,1436,151,248]" lastPageId="8" lastPageNumber="79" pageId="7" pageNumber="78">
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<emphasis id="B968870E0F10CB04FF641B55FE81F80B" bold="true" box="[199,315,1906,1930]" pageId="7" pageNumber="78">Remarks.</emphasis>
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One of the specimens (AMS B.
|
||
<quantity id="4CE4F6F90F10CB04FD631B55FCDDF80B" box="[704,871,1906,1930]" metricMagnitude="3" metricUnit="m" metricValue="8.280123999999999" pageId="7" pageNumber="78" unit="mm" value="8280124.0">8280, 124 mm</quantity>
|
||
SL, from Sind,
|
||
<collectingCountry id="F30B1B8C0F10CB04FB801B55FB3EF80A" box="[1059,1156,1906,1931]" name="Pakistan" pageId="7" pageNumber="78">Pakistan</collectingCountry>
|
||
) in Iwatsuki (2013) was identified as
|
||
<taxonomicName id="4C1C209F0F10CB04FE8B1BB1FE13F82F" box="[296,425,1942,1966]" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="78" phylum="Chordata" rank="species" species="arabicus">
|
||
<emphasis id="B968870E0F10CB04FE8B1BB1FE13F82F" box="[296,425,1942,1966]" italics="true" pageId="7" pageNumber="78">A. arabicus</emphasis>
|
||
</taxonomicName>
|
||
and erroneously listed in the description of
|
||
<taxonomicName id="4C1C209F0F10CB04FC3B1BB1FC43F82F" box="[920,1017,1942,1966]" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="78" phylum="Chordata" rank="species" species="sheim">
|
||
<emphasis id="B968870E0F10CB04FC3B1BB1FC43F82F" box="[920,1017,1942,1966]" italics="true" pageId="7" pageNumber="78">A. sheim</emphasis>
|
||
</taxonomicName>
|
||
in the “other specimens” in the same publication.
|
||
<taxonomicName id="4C1C209F0F10CB04FE8B1B9DFD9AF853" box="[296,544,1978,2002]" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="78" phylum="Chordata" rank="species" species="sheim">
|
||
<emphasis id="B968870E0F10CB04FE8B1B9DFD9AF853" box="[296,544,1978,2002]" italics="true" pageId="7" pageNumber="78">Acanthopagrus sheim</emphasis>
|
||
</taxonomicName>
|
||
can be easily mistaken for
|
||
<taxonomicName id="4C1C209F0F10CB04FCD21B9DFB30F853" box="[881,1162,1978,2002]" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="78" phylum="Chordata" rank="species" species="arabicus">
|
||
<emphasis id="B968870E0F10CB04FCD21B9DFB30F853" box="[881,1162,1978,2002]" italics="true" pageId="7" pageNumber="78">Acanthopagrus arabicus</emphasis>
|
||
</taxonomicName>
|
||
for its nearly identical appearance, scalation and morphometric proportions, but the local fishermen of Sindh coast consider them of different
|
||
<typeStatus id="54A7E5BE0F1FCB0BFEA31C9AFE84FF54" box="[256,318,189,213]" pageId="8" pageNumber="79">types</typeStatus>
|
||
.
|
||
<taxonomicName id="4C1C209F0F1FCB0BFEE81C9BFE17FF55" box="[331,429,188,212]" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="79" phylum="Chordata" rank="species" species="sheim">
|
||
<emphasis id="B968870E0F1FCB0BFEE81C9BFE17FF55" box="[331,429,188,212]" italics="true" pageId="8" pageNumber="79">A. sheim</emphasis>
|
||
</taxonomicName>
|
||
is locally known as Chaann and
|
||
<taxonomicName id="4C1C209F0F1FCB0BFC8F1C9BFC14FF55" box="[812,942,188,212]" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="79" phylum="Chordata" rank="species" species="arabicus">
|
||
<emphasis id="B968870E0F1FCB0BFC8F1C9BFC14FF55" box="[812,942,188,212]" italics="true" pageId="8" pageNumber="79">A. arabicus</emphasis>
|
||
</taxonomicName>
|
||
is called Nehra. There are also some color variations in this species compared to the specimens listed by Iwatsuki (2013).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |