543 lines
82 KiB
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543 lines
82 KiB
XML
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<mods:title id="028B5E5AA78936CD051830D78C03695E">Broeckinella Hensoni N. Sp., A New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran And A Revision Of The Genus Broeckinella Henson, 1948</mods:title>
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<treatment id="03B47D6A1C660136042EFD633F50FB0C" LSID="urn:lsid:plazi:treatment:03B47D6A1C660136042EFD633F50FB0C" httpUri="http://treatment.plazi.org/id/03B47D6A1C660136042EFD633F50FB0C" lastPageId="6" lastPageNumber="63" pageId="3" pageNumber="60">
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<subSubSection id="C3079FF71C660133042EFD633E3EFCD9" pageId="3" pageNumber="60" type="nomenclature">
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<paragraph id="8BA2CC7C1C660133042EFD633E3EFCD9" blockId="3.[816,1437,763,818]" pageId="3" pageNumber="60">
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<heading id="D0EA7B101C660133042EFD633E3EFCD9" centered="true" fontSize="36" level="2" pageId="3" pageNumber="60" reason="2">
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Family
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<taxonomicName id="4C1DB7FF1C6601330498FD633E3EFCD9" ID-CoL="7ZLC4" authority="(Munier-Chalmas, 1887) Maync, 1950" authorityName="Maync" authorityYear="1950" baseAuthorityName="Munier-Chalmas" baseAuthorityYear="1887" class="Globothalamea" family="Spirocyclinidae" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="60" phylum="Foraminifera" rank="family">Spirocyclinidae (Munier-Chalmas, 1887) Maync, 1950</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3079FF71C660134042EFCA13EE5FACF" lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60" type="discussion">
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<paragraph id="8BA2CC7C1C660133042EFCA13EE4FBAD" blockId="3.[816,1437,825,2045]" pageId="3" pageNumber="60">
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<emphasis id="B969106E1C660133042EFCA13EF9FCBA" bold="true" box="[816,929,825,849]" pageId="3" pageNumber="60">Remarks:</emphasis>
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In the definition of the families
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<taxonomicName id="4C1DB7FF1C6601330211FCA13E0DFC84" authorityName="Marie" authorityYear="1941" class="Globothalamea" family="Cyclamminidae" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="60" phylum="Foraminifera" rank="family">Cyclamminidae</taxonomicName>
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and
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<taxonomicName id="4C1DB7FF1C6601330490FCCF3821FC84" authority=", Albrich et al. (2015, p. 255)" authorityName="Albrich" authorityPageNumber="255" authorityYear="2015" box="[910,1401,855,879]" class="Globothalamea" family="Spirocyclinidae" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="60" phylum="Foraminifera" rank="family">
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Spirocyclinidae,
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<bibRefCitation id="EF8CB18D1C660133035BFCCF3846FC84" author="Albrich, S. & Boix, C. & Caus, E." box="[1093,1310,855,879]" pageId="3" pageNumber="60" pagination="245 - 267" refId="ref4304" refString="Albrich, S., Boix, C., Caus, E., 2015. Selected agglutinat- ed larger foraminifera from the Font de les Bagasses unit (Lower Campanian, southern Pyrenees). Carnets de Geologie 15 (18): 245 - 267." type="journal article" year="2015">Albrich et al. (2015</bibRefCitation>
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, p. 255)
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</taxonomicName>
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do not follow the classification of
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<bibRefCitation id="EF8CB18D1C660133039FFCEE3863FC65" author="Kaminski, M. A." box="[1153,1339,886,910]" pageId="3" pageNumber="60" pagination="89 - 108" refId="ref5055" refString="Kaminski, M. A., 2014. The year 2010 classification of the agglutinated Foraminifera. Micropaleontology, 60: 89 - 108." type="journal article" year="2014">Kaminski (2014)</bibRefCitation>
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. According to these authors “the genera with polygonal subepidermal patterns should be ascribed to the Family
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<taxonomicName id="4C1DB7FF1C6601330244FC2B3EC6FC01" class="Globothalamea" family="Spirocyclinidae" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="60" phylum="Foraminifera" rank="family">Spirocyclinidae</taxonomicName>
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instead of the
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<taxonomicName id="4C1DB7FF1C660133035DFC4A39AAFC01" authorityName="Marie" authorityYear="1941" box="[1091,1266,978,1002]" class="Globothalamea" family="Cyclamminidae" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="60" phylum="Foraminifera" rank="family">Cyclamminidae</taxonomicName>
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”. In this definition, the genus
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<taxonomicName id="4C1DB7FF1C66013304C4FC69393AFBE2" authorityName="HENSON" authorityYear="1948" box="[986,1122,1009,1033]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="60" phylum="Foraminifera" rank="genus">
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<emphasis id="B969106E1C66013304C4FC69393AFBE2" box="[986,1122,1009,1033]" italics="true" pageId="3" pageNumber="60">Broeckinella</emphasis>
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</taxonomicName>
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possessing a polygonal subepidermal pattern should therefore be included in the
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<taxonomicName id="4C1DB7FF1C660133026EFB973EE0FBAD" class="Globothalamea" family="Spirocyclinidae" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="60" phylum="Foraminifera" rank="family">Spirocyclinidae</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BA2CC7C1C660133042EFBD43E3EFB68" blockId="3.[816,1437,825,2045]" pageId="3" pageNumber="60">
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Subfamily
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<taxonomicName id="4C1DB7FF1C66013304ABFBD43E3EFB68" authority="Loeblich & Tappan, 1985" authorityName="Loeblich & Tappan" authorityYear="1985" class="Globothalamea" family="Spirocyclinidae" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="60" phylum="Foraminifera" rank="subFamily" subFamily="Pseudochoffatellinae">Pseudochoffatellinae Loeblich & Tappan, 1985</taxonomicName>
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</paragraph>
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<paragraph id="8BA2CC7C1C660133042EFB123852FB49" blockId="3.[816,1437,825,2045]" box="[816,1290,1162,1186]" pageId="3" pageNumber="60">
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Genus
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<taxonomicName id="4C1DB7FF1C6601330465FB1239FCFB49" authority="Henson, 1948" authorityName="HENSON" authorityYear="1948" box="[891,1188,1162,1186]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="60" phylum="Foraminifera" rank="genus">
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<emphasis id="B969106E1C6601330465FB12395FFB49" bold="true" box="[891,1031,1162,1186]" italics="true" pageId="3" pageNumber="60">Broeckinella</emphasis>
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<bibRefCitation id="EF8CB18D1C6601330311FB1339FCFB49" author="Henson, F. R. S." box="[1039,1188,1162,1186]" pageId="3" pageNumber="60" refId="ref4860" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History), 127 pp." type="book" year="1948">Henson, 1948</bibRefCitation>
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</taxonomicName>
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emended
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</paragraph>
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<paragraph id="8BA2CC7C1C660133042EFB313854FA18" blockId="3.[816,1437,825,2045]" pageId="3" pageNumber="60">
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<emphasis id="B969106E1C660133042EFB313EF9FB2A" bold="true" box="[816,929,1193,1217]" pageId="3" pageNumber="60">Remarks:</emphasis>
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The generic diagnosis of Loeblich and Tappan (1987, p. 104) only referred to the megalospheric form. It needs to be emended with respect to data obtained from a microspheric specimen described here. The diagnosis of Loeblich and Tappan (op. cit.) is emended as follows (added changes in bold letters)…….“chambers rapidly increasing in breadth to become arcuate but not completely cyclical, so that the test is successively flabelliform, complanate, and finally reniform
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<emphasis id="B969106E1C66013303B0FA0639A1FA1F" bold="true" pageId="3" pageNumber="60">in the macrospheric stage. Microspheric stage with a few annular chambers, so that the test is ovoid to discoidal</emphasis>
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”.
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</paragraph>
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<paragraph id="8BA2CC7C1C660133042EFA6239EAF9F9" blockId="3.[816,1437,825,2045]" box="[816,1202,1530,1554]" pageId="3" pageNumber="60">
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<taxonomicName id="4C1DB7FF1C660133042EFA6239EAF9F9" authority="Henson, 1948" authorityName="Henson" authorityYear="1948" box="[816,1202,1530,1554]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="60" phylum="Foraminifera" rank="species" species="arabica">
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<emphasis id="B969106E1C660133042EFA62394FF9F9" bold="true" box="[816,1047,1530,1554]" italics="true" pageId="3" pageNumber="60">Broeckinella arabica</emphasis>
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<bibRefCitation id="EF8CB18D1C6601330300FA6339EAF9F9" author="Henson, F. R. S." box="[1054,1202,1530,1554]" pageId="3" pageNumber="60" refId="ref4860" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History), 127 pp." type="book" year="1948">Henson, 1948</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="8BA2CC7C1C660133042EF9813E86F9DB" blockId="3.[816,1437,825,2045]" box="[816,990,1561,1585]" pageId="3" pageNumber="60">
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<figureCitation id="1326D0F91C660133042EF9813EF9F9DA" box="[816,929,1561,1585]" captionStart="Fig" captionStartId="4.[151,184,1322,1343]" captionTargetBox="[159,770,159,1285]" captionTargetId="figure-40@4.[151,770,151,1285]" captionTargetPageId="4" captionText="Fig. 4 Internal structure of Broeckinella arabica Henson. a Drawing of a slightly oblique equatorial section (slightly modified from Henson, 1947, fig. 13c; see also Cherchi and Schroeder 1978, fig. 1C). 1 = subepidermal layer (network; exoskeleton), 2 = zone with transverse partitions only, 3 = undivided chamber interior (lacking endoskeleton); septa with foramina. b Tangential equatorial section of Broeckinella arabica showing the three zones of Henson (1948). Thin-section Rt 85, Mandegan section. c Tangential equatorial section showing main partitions aligned between successive chambers (detail from Fig. 6d). Thin-section NG 42-1, Naghan section." figureDoi="http://doi.org/10.5281/zenodo.12583418" httpUri="https://zenodo.org/record/12583418/files/figure.png" pageId="3" pageNumber="60">Figs. 4b–d</figureCitation>
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,
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<figureCitation id="1326D0F91C66013304B1F9813E8EF9DA" box="[943,982,1561,1585]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="5.[225,258,1699,1720]" captionStartId-1="7.[188,223,1694,1715]" captionTargetBox-0="[225,1386,576,1662]" captionTargetBox-1="[188,1399,181,1657]" captionTargetId-0="figure-437@5.[225,1386,576,1662]" captionTargetId-1="figure-31@7.[188,1399,181,1657]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionText-0="Fig. 5 Broeckinella hensoni n. sp. (a) and Broeckinella arabica Henson (b–d) from the upper Maastrichtian of the Tarbur Formation, Naghan section. a Foraminiferal wackestone with Broeckinella hensoni n. sp., thin-section 2NG 197. b Foraminiferal wackestone with Broeckinella arabica Henson, 2NG 128-1. c–d Details from b, showing a microspheric specimen. Scale bars in c and d = 1 mm." captionText-1="Fig. 6 Broeckinella arabica Henson, upper Maastrichtian Tarbur Formation of the Mandegan (a, d) and Naghan sections (b– c, e–f), Zagros Zone, SW Iran. a, c–d Oblique equatorial sections, partly fragmentary, in some parts crossing the subepidermal network (e.g., a and left side of d). Note the undivided central part of the chambers in c. b Tangential section in the plane with only main partitions aligned between subsequent chambers (= zone 2 in Henson 1948; see Fig. 6a). e–f Oblique sections. Note initial planispiral part in the megalospheric specimen in e passing the proloculus (arrow). Note also the chambers undivided in the central part in e. Thin-sections: Rt 104 (a), NG 42-1 (b), NG 21 (c), Rt 85 (d), 2NG 49 (e), 2NG 112 (f). Scale bars 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12583420" figureDoi-1="http://doi.org/10.5281/zenodo.12583422" httpUri-0="https://zenodo.org/record/12583420/files/figure.png" httpUri-1="https://zenodo.org/record/12583422/files/figure.png" pageId="3" pageNumber="60">5–6</figureCitation>
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.
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</paragraph>
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<paragraph id="8BA2CC7C1C660133042EF9AF3925F985" blockId="3.[816,1437,825,2045]" pageId="3" pageNumber="60">
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*1948
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<taxonomicName id="4C1DB7FF1C6601330460F9AF393BF9A4" authorityName="SCHLAGINTWEIT & RASHIDI" authorityYear="2020" box="[894,1123,1591,1615]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="60" phylum="Foraminifera" rank="species" species="arabica" status="n. gen., n. sp.">
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<emphasis id="B969106E1C6601330460F9AF393BF9A4" box="[894,1123,1591,1615]" italics="true" pageId="3" pageNumber="60">Broeckinella arabica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25AAD151C6601330372F9A0385DF9A4" box="[1132,1285,1592,1616]" pageId="3" pageNumber="60" rank="species">n. gen., n. sp.</taxonomicNameLabel>
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– Henson, p. 93, pl. 7, fig. 6, text-fig. 13a–c.
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</paragraph>
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<paragraph id="8BA2CC7C1C660133042EF9ED392AF940" blockId="3.[816,1437,825,2045]" pageId="3" pageNumber="60">
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1978
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<taxonomicName id="4C1DB7FF1C6601330462F9EC398CF967" authority="Henson" authorityName="Henson" box="[892,1236,1652,1676]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="60" phylum="Foraminifera" rank="species" species="arabica">
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<emphasis id="B969106E1C6601330462F9EC3934F967" box="[892,1132,1652,1676]" italics="true" pageId="3" pageNumber="60">Broeckinella arabica</emphasis>
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Henson
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</taxonomicName>
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– Cherchi and Schroeder, p. 514, fig. 1 A–C.
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</paragraph>
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<paragraph id="8BA2CC7C1C660133042EF92A3952F902" blockId="3.[816,1437,825,2045]" pageId="3" pageNumber="60">
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2004
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<taxonomicName id="4C1DB7FF1C660133046DF92A39D9F921" authorityName="Munier-Chalmas" authorityYear="1887" box="[883,1153,1714,1738]" class="Globothalamea" family="Dicyclinidae" genus="Dicyclina" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="60" phylum="Foraminifera" rank="species" species="schlumbergeri">
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<emphasis id="B969106E1C660133046DF92A39D9F921" box="[883,1153,1714,1738]" italics="true" pageId="3" pageNumber="60">Dicyclina schlumbergeri</emphasis>
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</taxonomicName>
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– Khosrow Tehrani and Afghah, pl. 2, fig. 4.
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</paragraph>
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<paragraph id="8BA2CC7C1C660133042EF9773EF4F8CD" blockId="3.[816,1437,825,2045]" pageId="3" pageNumber="60">
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2008
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<taxonomicName id="4C1DB7FF1C6601330471F9773922F8EC" authorityName="Munier-Chalmas" authorityYear="1887" box="[879,1146,1775,1799]" class="Globothalamea" family="Dicyclinidae" genus="Dicyclina" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="60" phylum="Foraminifera" rank="species" species="schlumbergeri">
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<emphasis id="B969106E1C6601330471F9773922F8EC" box="[879,1146,1775,1799]" italics="true" pageId="3" pageNumber="60">Dicyclina schlumbergeri</emphasis>
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</taxonomicName>
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– Khosrow Tehrani et al., pl. 1, fig. 6.
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</paragraph>
|
||
<paragraph id="8BA2CC7C1C660133042EF8B5396EF888" blockId="3.[816,1437,825,2045]" pageId="3" pageNumber="60">
|
||
2016
|
||
<taxonomicName id="4C1DB7FF1C660133046CF8B539ECF8AF" authority="Henson" authorityName="Henson" box="[882,1204,1837,1861]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="3" pageNumber="60" phylum="Foraminifera" rank="species" species="arabica">
|
||
<emphasis id="B969106E1C660133046CF8B53900F8AE" box="[882,1112,1837,1861]" italics="true" pageId="3" pageNumber="60">Broeckinella arabica</emphasis>
|
||
Henson
|
||
</taxonomicName>
|
||
– Schlagintweit and Rashidi, p. 57, fig. 9a–e.
|
||
</paragraph>
|
||
<paragraph id="8BA2CC7C1C660134042EF8F23834FD69" blockId="3.[816,1437,825,2045]" lastBlockId="4.[816,1437,158,1439]" lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60">
|
||
<emphasis id="B969106E1C660133042EF8F2396CF869" bold="true" box="[816,1076,1898,1922]" pageId="3" pageNumber="60">Emended description:</emphasis>
|
||
Test finely agglutinated, compressed, with numerous chambers rapidly increasing in width so that the
|
||
<emphasis id="B969106E1C66013304E0F83039FAF82B" bold="true" box="[1022,1186,1960,1984]" pageId="3" pageNumber="60">megalospheric</emphasis>
|
||
test becomes flabelliform. A tiny proloculus (diameter ~
|
||
<quantity id="4CE561991C66013303A3F85E3810F835" box="[1213,1352,1990,2014]" metricMagnitude="-5" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7" pageId="3" pageNumber="60" unit="mm" value="0.085" valueMax="0.1" valueMin="0.07">0.07-0.1 mm</quantity>
|
||
) is followed by about 8-10 planispiral chambers that increase rather rapidly in width (
|
||
<figureCitation id="1326D0F91C610134035FFF0639C8FF5D" box="[1089,1168,158,182]" captionStart="Fig" captionStartId="7.[188,223,1694,1715]" captionTargetBox="[188,1399,181,1657]" captionTargetId="figure-31@7.[188,1399,181,1657]" captionTargetPageId="7" captionText="Fig. 6 Broeckinella arabica Henson, upper Maastrichtian Tarbur Formation of the Mandegan (a, d) and Naghan sections (b– c, e–f), Zagros Zone, SW Iran. a, c–d Oblique equatorial sections, partly fragmentary, in some parts crossing the subepidermal network (e.g., a and left side of d). Note the undivided central part of the chambers in c. b Tangential section in the plane with only main partitions aligned between subsequent chambers (= zone 2 in Henson 1948; see Fig. 6a). e–f Oblique sections. Note initial planispiral part in the megalospheric specimen in e passing the proloculus (arrow). Note also the chambers undivided in the central part in e. Thin-sections: Rt 104 (a), NG 42-1 (b), NG 21 (c), Rt 85 (d), 2NG 49 (e), 2NG 112 (f). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12583422" httpUri="https://zenodo.org/record/12583422/files/figure.png" pageId="4" pageNumber="61">Fig. 6e</figureCitation>
|
||
). The chamber margins are subdivided by exoskeletal elements, both vertical and horizontal resulting in a meshwork (or pattern) of tiny polygonal cells displayed in shallow tangential sections (= zone
|
||
<quantity id="4CE561991C6101340493FE813EE1FEDA" box="[909,953,281,305]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="61" unit="in" value="1.0">1 in</quantity>
|
||
<bibRefCitation id="EF8CB18D1C61013404DCFE81390CFEDA" author="Henson, F. R. S." box="[962,1108,281,305]" pageId="4" pageNumber="61" refId="ref4860" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History), 127 pp." type="book" year="1948">Henson 1948</bibRefCitation>
|
||
; see
|
||
<figureCitation id="1326D0F91C6101340393FE813983FEDB" box="[1165,1243,281,305]" captionStart="Fig" captionStartId="4.[151,184,1322,1343]" captionTargetBox="[159,770,159,1285]" captionTargetId="figure-40@4.[151,770,151,1285]" captionTargetPageId="4" captionText="Fig. 4 Internal structure of Broeckinella arabica Henson. a Drawing of a slightly oblique equatorial section (slightly modified from Henson, 1947, fig. 13c; see also Cherchi and Schroeder 1978, fig. 1C). 1 = subepidermal layer (network; exoskeleton), 2 = zone with transverse partitions only, 3 = undivided chamber interior (lacking endoskeleton); septa with foramina. b Tangential equatorial section of Broeckinella arabica showing the three zones of Henson (1948). Thin-section Rt 85, Mandegan section. c Tangential equatorial section showing main partitions aligned between successive chambers (detail from Fig. 6d). Thin-section NG 42-1, Naghan section." figureDoi="http://doi.org/10.5281/zenodo.12583418" httpUri="https://zenodo.org/record/12583418/files/figure.png" pageId="4" pageNumber="61">Fig. 4a</figureCitation>
|
||
) (
|
||
<figureCitation id="1326D0F91C61013403EAFE813809FEDA" box="[1268,1361,281,305]" captionStart="Fig" captionStartId="4.[151,184,1322,1343]" captionTargetBox="[159,770,159,1285]" captionTargetId="figure-40@4.[151,770,151,1285]" captionTargetPageId="4" captionText="Fig. 4 Internal structure of Broeckinella arabica Henson. a Drawing of a slightly oblique equatorial section (slightly modified from Henson, 1947, fig. 13c; see also Cherchi and Schroeder 1978, fig. 1C). 1 = subepidermal layer (network; exoskeleton), 2 = zone with transverse partitions only, 3 = undivided chamber interior (lacking endoskeleton); septa with foramina. b Tangential equatorial section of Broeckinella arabica showing the three zones of Henson (1948). Thin-section Rt 85, Mandegan section. c Tangential equatorial section showing main partitions aligned between successive chambers (detail from Fig. 6d). Thin-section NG 42-1, Naghan section." figureDoi="http://doi.org/10.5281/zenodo.12583418" httpUri="https://zenodo.org/record/12583418/files/figure.png" pageId="4" pageNumber="61">Figs. 4b</figureCitation>
|
||
,
|
||
<figureCitation id="1326D0F91C6101340240FE813832FEDA" box="[1374,1386,281,305]" captionStart="Fig" captionStartId="7.[188,223,1694,1715]" captionTargetBox="[188,1399,181,1657]" captionTargetId="figure-31@7.[188,1399,181,1657]" captionTargetPageId="7" captionText="Fig. 6 Broeckinella arabica Henson, upper Maastrichtian Tarbur Formation of the Mandegan (a, d) and Naghan sections (b– c, e–f), Zagros Zone, SW Iran. a, c–d Oblique equatorial sections, partly fragmentary, in some parts crossing the subepidermal network (e.g., a and left side of d). Note the undivided central part of the chambers in c. b Tangential section in the plane with only main partitions aligned between subsequent chambers (= zone 2 in Henson 1948; see Fig. 6a). e–f Oblique sections. Note initial planispiral part in the megalospheric specimen in e passing the proloculus (arrow). Note also the chambers undivided in the central part in e. Thin-sections: Rt 104 (a), NG 42-1 (b), NG 21 (c), Rt 85 (d), 2NG 49 (e), 2NG 112 (f). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12583422" httpUri="https://zenodo.org/record/12583422/files/figure.png" pageId="4" pageNumber="61">6</figureCitation>
|
||
a-d). Toward the chamber interior a zone follows where only main partitions are present, aligned between successive chambers (= zone
|
||
<quantity id="4CE561991C61013404E0FEEC3972FE67" box="[1022,1066,372,396]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="61" unit="in" value="2.0">2 in</quantity>
|
||
<bibRefCitation id="EF8CB18D1C610134032DFEED399EFE67" author="Henson, F. R. S." box="[1075,1222,372,396]" pageId="4" pageNumber="61" refId="ref4860" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History), 127 pp." type="book" year="1948">Henson 1948</bibRefCitation>
|
||
; see
|
||
<figureCitation id="1326D0F91C610134021EFEEC3816FE67" box="[1280,1358,372,396]" captionStart="Fig" captionStartId="4.[151,184,1322,1343]" captionTargetBox="[159,770,159,1285]" captionTargetId="figure-40@4.[151,770,151,1285]" captionTargetPageId="4" captionText="Fig. 4 Internal structure of Broeckinella arabica Henson. a Drawing of a slightly oblique equatorial section (slightly modified from Henson, 1947, fig. 13c; see also Cherchi and Schroeder 1978, fig. 1C). 1 = subepidermal layer (network; exoskeleton), 2 = zone with transverse partitions only, 3 = undivided chamber interior (lacking endoskeleton); septa with foramina. b Tangential equatorial section of Broeckinella arabica showing the three zones of Henson (1948). Thin-section Rt 85, Mandegan section. c Tangential equatorial section showing main partitions aligned between successive chambers (detail from Fig. 6d). Thin-section NG 42-1, Naghan section." figureDoi="http://doi.org/10.5281/zenodo.12583418" httpUri="https://zenodo.org/record/12583418/files/figure.png" pageId="4" pageNumber="61">Fig. 4a</figureCitation>
|
||
) (
|
||
<figureCitation id="1326D0F91C6101340279FEEC3E14FE40" captionStart="Fig" captionStartId="4.[151,184,1322,1343]" captionTargetBox="[159,770,159,1285]" captionTargetId="figure-40@4.[151,770,151,1285]" captionTargetPageId="4" captionText="Fig. 4 Internal structure of Broeckinella arabica Henson. a Drawing of a slightly oblique equatorial section (slightly modified from Henson, 1947, fig. 13c; see also Cherchi and Schroeder 1978, fig. 1C). 1 = subepidermal layer (network; exoskeleton), 2 = zone with transverse partitions only, 3 = undivided chamber interior (lacking endoskeleton); septa with foramina. b Tangential equatorial section of Broeckinella arabica showing the three zones of Henson (1948). Thin-section Rt 85, Mandegan section. c Tangential equatorial section showing main partitions aligned between successive chambers (detail from Fig. 6d). Thin-section NG 42-1, Naghan section." figureDoi="http://doi.org/10.5281/zenodo.12583418" httpUri="https://zenodo.org/record/12583418/files/figure.png" pageId="4" pageNumber="61">Figs. 4c</figureCitation>
|
||
,
|
||
<figureCitation id="1326D0F91C6101340446FE0B3ED7FE40" box="[856,911,403,427]" captionStart="Fig" captionStartId="7.[188,223,1694,1715]" captionTargetBox="[188,1399,181,1657]" captionTargetId="figure-31@7.[188,1399,181,1657]" captionTargetPageId="7" captionText="Fig. 6 Broeckinella arabica Henson, upper Maastrichtian Tarbur Formation of the Mandegan (a, d) and Naghan sections (b– c, e–f), Zagros Zone, SW Iran. a, c–d Oblique equatorial sections, partly fragmentary, in some parts crossing the subepidermal network (e.g., a and left side of d). Note the undivided central part of the chambers in c. b Tangential section in the plane with only main partitions aligned between subsequent chambers (= zone 2 in Henson 1948; see Fig. 6a). e–f Oblique sections. Note initial planispiral part in the megalospheric specimen in e passing the proloculus (arrow). Note also the chambers undivided in the central part in e. Thin-sections: Rt 104 (a), NG 42-1 (b), NG 21 (c), Rt 85 (d), 2NG 49 (e), 2NG 112 (f). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12583422" httpUri="https://zenodo.org/record/12583422/files/figure.png" pageId="4" pageNumber="61">6b, d</figureCitation>
|
||
). The central part is undivided (
|
||
<figureCitation id="1326D0F91C61013403EFFE0B3815FE40" box="[1265,1357,403,427]" captionStart="Fig" captionStartId="4.[151,184,1322,1343]" captionTargetBox="[159,770,159,1285]" captionTargetId="figure-40@4.[151,770,151,1285]" captionTargetPageId="4" captionText="Fig. 4 Internal structure of Broeckinella arabica Henson. a Drawing of a slightly oblique equatorial section (slightly modified from Henson, 1947, fig. 13c; see also Cherchi and Schroeder 1978, fig. 1C). 1 = subepidermal layer (network; exoskeleton), 2 = zone with transverse partitions only, 3 = undivided chamber interior (lacking endoskeleton); septa with foramina. b Tangential equatorial section of Broeckinella arabica showing the three zones of Henson (1948). Thin-section Rt 85, Mandegan section. c Tangential equatorial section showing main partitions aligned between successive chambers (detail from Fig. 6d). Thin-section NG 42-1, Naghan section." figureDoi="http://doi.org/10.5281/zenodo.12583418" httpUri="https://zenodo.org/record/12583418/files/figure.png" pageId="4" pageNumber="61">Figs. 4b</figureCitation>
|
||
,
|
||
<figureCitation id="1326D0F91C6101340247FE0B38D5FE40" box="[1369,1421,403,427]" captionStart="Fig" captionStartId="7.[188,223,1694,1715]" captionTargetBox="[188,1399,181,1657]" captionTargetId="figure-31@7.[188,1399,181,1657]" captionTargetPageId="7" captionText="Fig. 6 Broeckinella arabica Henson, upper Maastrichtian Tarbur Formation of the Mandegan (a, d) and Naghan sections (b– c, e–f), Zagros Zone, SW Iran. a, c–d Oblique equatorial sections, partly fragmentary, in some parts crossing the subepidermal network (e.g., a and left side of d). Note the undivided central part of the chambers in c. b Tangential section in the plane with only main partitions aligned between subsequent chambers (= zone 2 in Henson 1948; see Fig. 6a). e–f Oblique sections. Note initial planispiral part in the megalospheric specimen in e passing the proloculus (arrow). Note also the chambers undivided in the central part in e. Thin-sections: Rt 104 (a), NG 42-1 (b), NG 21 (c), Rt 85 (d), 2NG 49 (e), 2NG 112 (f). Scale bars 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12583422" httpUri="https://zenodo.org/record/12583422/files/figure.png" pageId="4" pageNumber="61">6c, e</figureCitation>
|
||
). The multiple foramina are arranged in the median plane, not related to the main partitions (that can therefore not be termed septula). The
|
||
<emphasis id="B969106E1C6101340355FE7739B8FDEC" bold="true" box="[1099,1248,495,519]" pageId="4" pageNumber="61">microspheric</emphasis>
|
||
test is ovoid to discoidal due to final annular chambers (
|
||
<figureCitation id="1326D0F91C610134021CFD963836FDCE" box="[1282,1390,526,550]" captionStart="Fig" captionStartId="5.[225,258,1699,1720]" captionTargetBox="[225,1386,576,1662]" captionTargetId="figure-437@5.[225,1386,576,1662]" captionTargetPageId="5" captionText="Fig. 5 Broeckinella hensoni n. sp. (a) and Broeckinella arabica Henson (b–d) from the upper Maastrichtian of the Tarbur Formation, Naghan section. a Foraminiferal wackestone with Broeckinella hensoni n. sp., thin-section 2NG 197. b Foraminiferal wackestone with Broeckinella arabica Henson, 2NG 128-1. c–d Details from b, showing a microspheric specimen. Scale bars in c and d = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12583420" httpUri="https://zenodo.org/record/12583420/files/figure.png" pageId="4" pageNumber="61">Fig. 5b–c</figureCitation>
|
||
). A proloculus is not observable. The planispiral part, contrasting the A-form, consists of numerous (~ 26) chambers. The internal structure is identical to the A-form.
|
||
</paragraph>
|
||
<paragraph id="8BA2CC7C1C610134042EFD11398DFBFB" blockId="4.[816,1437,158,1439]" pageId="4" pageNumber="61">
|
||
<emphasis id="B969106E1C610134042EFD113EF9FD4A" bold="true" box="[816,929,649,673]" pageId="4" pageNumber="61">Remarks:</emphasis>
|
||
The material of
|
||
<taxonomicName id="4C1DB7FF1C6101340379FD123985FD4A" box="[1127,1245,649,673]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="61" phylum="Foraminifera" rank="species" species="arabica">
|
||
<emphasis id="B969106E1C6101340379FD123985FD4A" box="[1127,1245,649,673]" italics="true" pageId="4" pageNumber="61">B. arabica</emphasis>
|
||
</taxonomicName>
|
||
from the Tarbur Formation does not include centered sections. Data on the test dimensions therefore only comprise incomplete sections attaining a maximum size of
|
||
<quantity id="4CE561991C61013403A5FD7C3848FD17" box="[1211,1296,740,764]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="61" unit="mm" value="2.4">2.4 mm</quantity>
|
||
for megalospheric and up to
|
||
<quantity id="4CE561991C61013404E2FC9B390AFCF0" box="[1020,1106,771,795]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="4" pageNumber="61" unit="mm" value="4.6">4.6 mm</quantity>
|
||
for microspheric forms (
|
||
<tableCitation id="C69FF9C71C6101340275FC9B3E66FCD1" captionStart="Table 2" captionStartId="4.[267,332,1677,1701]" captionText="Table 2 Megalospheric versus microspheric generations of Broeckinella arabica Henson (A-form: data from Cherchi and Schroeder, 1978; B-form: this work, specimen shown in figure 5b–d)." pageId="4" pageNumber="61">Tab. 2</tableCitation>
|
||
).
|
||
<taxonomicName id="4C1DB7FF1C610134044BFCBB3E9DFCD1" box="[853,965,802,826]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="61" phylum="Foraminifera" rank="species" species="arabica">
|
||
<emphasis id="B969106E1C610134044BFCBB3E9DFCD1" box="[853,965,802,826]" italics="true" pageId="4" pageNumber="61">B. arabica</emphasis>
|
||
</taxonomicName>
|
||
is intermediate in size between the distinctly smaller
|
||
<taxonomicName id="4C1DB7FF1C61013404BBFCD93967FCB3" box="[933,1087,833,856]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="61" phylum="Foraminifera" rank="species" species="neumannae">
|
||
<emphasis id="B969106E1C61013404BBFCD93967FCB3" box="[933,1087,833,856]" italics="true" pageId="4" pageNumber="61">B. neumannae</emphasis>
|
||
</taxonomicName>
|
||
and the larger
|
||
<taxonomicName id="4C1DB7FF1C61013403F9FCD93801FCB3" authorityName="SCHLAGINTWEIT & RASHIDI" authorityYear="2020" box="[1255,1369,832,856]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="61" phylum="Foraminifera" rank="species" species="hensoni" status="n. sp.">
|
||
<emphasis id="B969106E1C61013403F9FCD93801FCB3" box="[1255,1369,832,856]" italics="true" pageId="4" pageNumber="61">B. hensoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25AAD151C610134027CFCD938C5FCB3" box="[1378,1437,833,856]" pageId="4" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
|
||
(
|
||
<tableCitation id="C69FF9C71C6101340426FCC73E26FC9C" box="[824,894,863,887]" captionStart="Table 3" captionStartId="5.[251,316,158,182]" captionText="Table 3 Summary table of the biometric parameters of Broeckinella species. Data from Henson (1948), Gendrot (1968), Cherchi and Schroeder (1978), and this work. * measured from the original papers." pageId="4" pageNumber="61">Tab. 3</tableCitation>
|
||
). The description of
|
||
<bibRefCitation id="EF8CB18D1C610134037AFCF8385CFC9C" author="Henson, F. R. S." box="[1124,1284,863,887]" pageId="4" pageNumber="61" refId="ref4860" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History), 127 pp." type="book" year="1948">Henson (1948)</bibRefCitation>
|
||
was based on a single specimen, a megalospheric form (see
|
||
<bibRefCitation id="EF8CB18D1C6101340256FCE63947FC5F" author="Cherchi, A. & Schroeder, R." pageId="4" pageNumber="61" pagination="513 - 520" refId="ref4416" refString="Cherchi, A., Schroeder, R., 1978. Revision of the type of Broeckinella arabica Henson (Foram.) from Qatar Peninsula (Arabia). Neues Jahrbuch fur Geologie und Palaontologie Monatshefte, 9: 513 - 520." type="journal article" year="1978">Cherchi and Schroeder, 1978</bibRefCitation>
|
||
). The specimen from the Tarbur Formation illustrated in
|
||
<figureCitation id="1326D0F91C610134035CFC2339CBFC38" box="[1090,1171,955,979]" captionStart="Fig" captionStartId="5.[225,258,1699,1720]" captionTargetBox="[225,1386,576,1662]" captionTargetId="figure-437@5.[225,1386,576,1662]" captionTargetPageId="5" captionText="Fig. 5 Broeckinella hensoni n. sp. (a) and Broeckinella arabica Henson (b–d) from the upper Maastrichtian of the Tarbur Formation, Naghan section. a Foraminiferal wackestone with Broeckinella hensoni n. sp., thin-section 2NG 197. b Foraminiferal wackestone with Broeckinella arabica Henson, 2NG 128-1. c–d Details from b, showing a microspheric specimen. Scale bars in c and d = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12583420" httpUri="https://zenodo.org/record/12583420/files/figure.png" pageId="4" pageNumber="61">Fig. 5b</figureCitation>
|
||
is the first record of a microspheric specimen. The differences between the two generations are summarized in
|
||
<tableCitation id="C69FF9C71C6101340360FC60398DFBFB" box="[1150,1237,1016,1040]" captionStart="Table 2" captionStartId="4.[267,332,1677,1701]" captionText="Table 2 Megalospheric versus microspheric generations of Broeckinella arabica Henson (A-form: data from Cherchi and Schroeder, 1978; B-form: this work, specimen shown in figure 5b–d)." pageId="4" pageNumber="61">Table 2.</tableCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA2CC7C1C610134042EFB8038CAFB0C" blockId="4.[816,1437,158,1439]" pageId="4" pageNumber="61">
|
||
<taxonomicName id="4C1DB7FF1C610134042EFB803EFCFBC4" box="[816,932,1047,1071]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="61" phylum="Foraminifera" rank="species" species="arabica">
|
||
<emphasis id="B969106E1C610134042EFB803EFCFBC4" box="[816,932,1047,1071]" italics="true" pageId="4" pageNumber="61">B. arabica</emphasis>
|
||
</taxonomicName>
|
||
was figured as
|
||
<taxonomicName id="4C1DB7FF1C610134034BFB8F3838FBC4" authorityName="Munier-Chalmas" authorityYear="1887" box="[1109,1376,1047,1071]" class="Globothalamea" family="Dicyclinidae" genus="Dicyclina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="61" phylum="Foraminifera" rank="species" species="schlumbergeri">
|
||
<emphasis id="B969106E1C610134034BFB8F3838FBC4" box="[1109,1376,1047,1071]" italics="true" pageId="4" pageNumber="61">Dicyclina schlumbergeri</emphasis>
|
||
</taxonomicName>
|
||
from the Maastrichtian of the Tarbur Formation by Khosrow Tehrani and Afghah (2004) and the Maastrichtian of the Amiran Formation (SW
|
||
<collectingCountry id="F30A8CEC1C6101340354FBEC392FFB60" box="[1098,1143,1140,1163]" name="Iran" pageId="4" pageNumber="61">Iran</collectingCountry>
|
||
, Zagros Zone) where the authors distinguished a
|
||
<emphasis id="B969106E1C6101340357FB0A3849FB41" box="[1097,1297,1170,1194]" italics="true" pageId="4" pageNumber="61">Loftusia-Dicyclina</emphasis>
|
||
assemblage zone.
|
||
<taxonomicName id="4C1DB7FF1C610134046BFB2839DAFB23" authorityName="Munier-Chalmas" authorityYear="1887" box="[885,1154,1200,1224]" class="Globothalamea" family="Dicyclinidae" genus="Dicyclina" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="61" phylum="Foraminifera" rank="species" species="schlumbergeri">
|
||
<emphasis id="B969106E1C610134046BFB2839DAFB23" box="[885,1154,1200,1224]" italics="true" pageId="4" pageNumber="61">Dicyclina schlumbergeri</emphasis>
|
||
</taxonomicName>
|
||
is rather common in the Tarbur Formation (see
|
||
<bibRefCitation id="EF8CB18D1C6101340339FB5738DFFB0C" author="Schlagintweit, F. & Rashidi, K." box="[1063,1415,1231,1255]" pageId="4" pageNumber="61" pagination="21 - 29" refId="ref5870" refString="Schlagintweit, F., Rashidi, K., 2018. Suraqalatia brasieri Gormus, Lawa & Nuaimy, 2017 (Larger benthic foraminifera; Suraqalatiidae n. fam.) from the Late Maastrichtian of the Tarbur Formation and remarks on Dicyclina Munier-Chalmas. Acta Palaeontologica Romaniae, 14 (1): 21 - 29." type="journal article" year="2018">Schlagintweit and Rashidi, 2018</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA2CC7C1C610134042EFB763EE5FACF" blockId="4.[816,1437,158,1439]" pageId="4" pageNumber="61">
|
||
<taxonomicName id="4C1DB7FF1C610134042EFB763862FAED" authority="Schlagintweit and Rashidi" authorityName="Schlagintweit and Rashidi" authorityYear="2020" box="[816,1338,1262,1286]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="61" phylum="Foraminifera" rank="species" species="hensoni" status="sp. nov.">
|
||
<emphasis id="B969106E1C610134042EFB763943FAED" bold="true" box="[816,1051,1262,1286]" italics="true" pageId="4" pageNumber="61">Broeckinella hensoni</emphasis>
|
||
Schlagintweit and Rashidi
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25AAD151C610134025FFB7738CDFAED" box="[1345,1429,1263,1286]" pageId="4" pageNumber="61" rank="species">nov. sp.</taxonomicNameLabel>
|
||
<figureCitation id="1326D0F91C610134042EFA953EDFFACE" box="[816,903,1293,1317]" captionStart="Fig" captionStartId="5.[225,258,1699,1720]" captionTargetBox="[225,1386,576,1662]" captionTargetId="figure-437@5.[225,1386,576,1662]" captionTargetPageId="5" captionText="Fig. 5 Broeckinella hensoni n. sp. (a) and Broeckinella arabica Henson (b–d) from the upper Maastrichtian of the Tarbur Formation, Naghan section. a Foraminiferal wackestone with Broeckinella hensoni n. sp., thin-section 2NG 197. b Foraminiferal wackestone with Broeckinella arabica Henson, 2NG 128-1. c–d Details from b, showing a microspheric specimen. Scale bars in c and d = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12583420" httpUri="https://zenodo.org/record/12583420/files/figure.png" pageId="4" pageNumber="61">Figs. 5a</figureCitation>
|
||
,
|
||
<figureCitation id="1326D0F91C610134048CFA953EEEFACE" box="[914,950,1293,1317]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[203,236,1747,1768]" captionStartId-1="9.[193,226,1666,1687]" captionTargetBox-0="[203,1406,252,1710]" captionTargetBox-1="[193,1394,268,1629]" captionTargetId-0="figure-60@8.[203,1406,252,1710]" captionTargetId-1="figure-37@9.[193,1394,268,1629]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 7 Broeckinella hensoni n. sp., upper Maastrichtian Tarbur Formation of the Naghan section, Zagros Zone, SW Iran. a, c Subaxial sections. Specimen shown in A displays more than 40 chambers in the uncoiled adult part. d Detail from c showing initial coiled part. e Oblique equatorial section. f Slightly oblique equatorial section showing flabelliform test morphology (test diameter: ~6.6 mm). Thin-sections: 2NG 153 (a), 2NGN (b, f), 2NG 38 (c–d), NG 197-1 (e). Scale bars 1.0 mm." captionText-1="Fig. 8 Broeckinella hensoni n. sp., upper Maastrichtian Tarbur Formation of the Naghan (a, d–e) and Mandegan (b–c, f) sections, Zagros Zone, SW Iran. a Oblique equatorial section. Note porcelaneous test of Cuvillerinella sp. below. b Detail showing fine subepidermal network (sn, below), followed by zone with radial partitions (rp) only and finally the undivided central chamber part with foramina (f) piercing the septa (s). c Oblique section cutting the initial planispiral part. d–e Oblique equatorial sections showing zones of different internal structure (see Text-Figure 3). f Detail from c showing initial planispirally coiled part with numerous chambers. Note also the fine subepidermal network and main partitions (right side). Thin-sections: 2NG 166 (a), Rt 67-1 (b), RT 72 (c, f), 2NG 118 (d), 2NG 172 (e). Scale bars 1.0 mm for a, c–e, 0.2 mm for b and f." figureDoi-0="http://doi.org/10.5281/zenodo.12583424" figureDoi-1="http://doi.org/10.5281/zenodo.12583426" httpUri-0="https://zenodo.org/record/12583424/files/figure.png" httpUri-1="https://zenodo.org/record/12583426/files/figure.png" pageId="4" pageNumber="61">7-8</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF629CF41C6101340789FAB23FF8F9BF" ID-DOI="http://doi.org/10.5281/zenodo.12583418" ID-Zenodo-Dep="12583418" httpUri="https://zenodo.org/record/12583418/files/figure.png" pageId="4" pageNumber="61" startId="4.[151,184,1322,1343]" targetBox="[159,770,159,1285]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BA2CC7C1C6101340789FAB23FF8F9BF" blockId="4.[151,771,1322,1620]" pageId="4" pageNumber="61">
|
||
<emphasis id="B969106E1C6101340789FAB23D8FFAD4" bold="true" box="[151,215,1322,1343]" pageId="4" pageNumber="61">Fig. 4</emphasis>
|
||
Internal structure of
|
||
<taxonomicName id="4C1DB7FF1C61013406A3FAB23FB4FAAB" authority="Henson." authorityName="Henson." box="[445,748,1322,1344]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="61" phylum="Foraminifera" rank="species" species="arabica">
|
||
<emphasis id="B969106E1C61013406A3FAB23FD7FAAB" box="[445,655,1322,1344]" italics="true" pageId="4" pageNumber="61">Broeckinella arabica</emphasis>
|
||
Henson.
|
||
</taxonomicName>
|
||
<emphasis id="B969106E1C61013405E9FAB23E5BFAD4" bold="true" box="[759,771,1322,1343]" pageId="4" pageNumber="61">a</emphasis>
|
||
Drawing of a slightly oblique equatorial section (slightly modified from Henson, 1947, fig. 13c; see also
|
||
<bibRefCitation id="EF8CB18D1C610134059AFAF93C6FFA78" author="Cherchi, A. & Schroeder, R." pageId="4" pageNumber="61" pagination="513 - 520" refId="ref4416" refString="Cherchi, A., Schroeder, R., 1978. Revision of the type of Broeckinella arabica Henson (Foram.) from Qatar Peninsula (Arabia). Neues Jahrbuch fur Geologie und Palaontologie Monatshefte, 9: 513 - 520." type="journal article" year="1978">Cherchi and Schroeder 1978</bibRefCitation>
|
||
, fig. 1C). 1 = subepidermal layer (network; exoskeleton), 2 = zone with transverse partitions only, 3 = undivided chamber interior (lacking endoskeleton); septa with foramina.
|
||
<emphasis id="B969106E1C61013407F9FA483DACFA0E" bold="true" box="[231,244,1488,1509]" pageId="4" pageNumber="61">b</emphasis>
|
||
Tangential equatorial section of
|
||
<taxonomicName id="4C1DB7FF1C6101340529FA483E5BFA0D" box="[567,771,1488,1510]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="61" phylum="Foraminifera" rank="species" species="arabica">
|
||
<emphasis id="B969106E1C6101340529FA483E5BFA0D" box="[567,771,1488,1510]" italics="true" pageId="4" pageNumber="61">Broeckinella arabica</emphasis>
|
||
</taxonomicName>
|
||
showing the three zones of
|
||
<bibRefCitation id="EF8CB18D1C61013406B9FA743F63F9EA" author="Henson, F. R. S." box="[423,571,1515,1537]" pageId="4" pageNumber="61" refId="ref4860" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History), 127 pp." type="book" year="1948">Henson (1948)</bibRefCitation>
|
||
. Thin-section Rt 85, Mandegan section.
|
||
<emphasis id="B969106E1C610134067DF99F3C36F9F7" bold="true" box="[355,366,1543,1564]" pageId="4" pageNumber="61">c</emphasis>
|
||
Tangential equatorial section showing main partitions aligned between successive chambers (detail from Fig. 6d). Thin-section NG 42-1, Naghan section.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3079FF71C610134042EFAB43971FA74" pageId="4" pageNumber="61" type="materials_examined">
|
||
<paragraph id="8BA2CC7C1C610134042EFAB43971FA74" blockId="4.[816,1437,158,1439]" pageId="4" pageNumber="61">
|
||
<emphasis id="B969106E1C610134042EFAB43EF9FAAF" bold="true" box="[816,929,1324,1348]" pageId="4" pageNumber="61">
|
||
<typeStatus id="54A672DE1C610134042EFAB43EC4FAAF" box="[816,924,1324,1348]" pageId="4" pageNumber="61" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Equatorial section of an assumed macrospheric specimen showing flabelliform test and illustrated in
|
||
<figureCitation id="1326D0F91C610134042EFAF03E22FA6B" box="[816,890,1384,1408]" captionStart="Fig" captionStartId="8.[203,236,1747,1768]" captionTargetBox="[203,1406,252,1710]" captionTargetId="figure-60@8.[203,1406,252,1710]" captionTargetPageId="8" captionText="Fig. 7 Broeckinella hensoni n. sp., upper Maastrichtian Tarbur Formation of the Naghan section, Zagros Zone, SW Iran. a, c Subaxial sections. Specimen shown in A displays more than 40 chambers in the uncoiled adult part. d Detail from c showing initial coiled part. e Oblique equatorial section. f Slightly oblique equatorial section showing flabelliform test morphology (test diameter: ~6.6 mm). Thin-sections: 2NG 153 (a), 2NGN (b, f), 2NG 38 (c–d), NG 197-1 (e). Scale bars 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12583424" httpUri="https://zenodo.org/record/12583424/files/figure.png" pageId="4" pageNumber="61">Fig. 7f</figureCitation>
|
||
, thin-section 2NG N. The diameter of the
|
||
<typeStatus id="54A672DE1C610134025EFAF038C5FA6B" box="[1344,1437,1384,1408]" pageId="4" pageNumber="61" type="holotype">holotype</typeStatus>
|
||
specimen is ~
|
||
<quantity id="4CE561991C61013404DCFA1F3979FA74" box="[962,1057,1415,1439]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" pageId="4" pageNumber="61" unit="mm" value="3.65">3.65 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF629CF41C6101340615F9153852F928" pageId="4" pageNumber="61" startId="4.[267,332,1677,1701]" targetType="table">
|
||
<paragraph id="8BA2CC7C1C6101340615F9153852F928" blockId="4.[267,1322,1676,1731]" pageId="4" pageNumber="61">
|
||
<emphasis id="B969106E1C6101340615F9153C38F94E" bold="true" box="[267,352,1677,1701]" pageId="4" pageNumber="61">Table 2</emphasis>
|
||
Megalospheric
|
||
<emphasis id="B969106E1C6101340513F9153F09F94F" box="[525,593,1677,1700]" italics="true" pageId="4" pageNumber="61">versus</emphasis>
|
||
microspheric generations of
|
||
<taxonomicName id="4C1DB7FF1C6101340496F914399AF94F" authority="Henson" authorityName="Henson" box="[904,1218,1676,1700]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="4" pageNumber="61" phylum="Foraminifera" rank="species" species="arabica">
|
||
<emphasis id="B969106E1C6101340496F9143931F94F" box="[904,1129,1676,1700]" italics="true" pageId="4" pageNumber="61">Broeckinella arabica</emphasis>
|
||
Henson
|
||
</taxonomicName>
|
||
(A-form: data from
|
||
<bibRefCitation id="EF8CB18D1C6101340686F9333F8AF928" author="Cherchi, A. & Schroeder, R." box="[408,722,1707,1731]" pageId="4" pageNumber="61" pagination="513 - 520" refId="ref4416" refString="Cherchi, A., Schroeder, R., 1978. Revision of the type of Broeckinella arabica Henson (Foram.) from Qatar Peninsula (Arabia). Neues Jahrbuch fur Geologie und Palaontologie Monatshefte, 9: 513 - 520." type="journal article" year="1978">Cherchi and Schroeder, 1978</bibRefCitation>
|
||
; B-form: this work, specimen shown in figure 5b–d).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA2CC7C1C6101340619F972385AF82A" pageId="4" pageNumber="61">
|
||
<table id="F91D3EDC1C61FECF0619F9723849F82A" box="[263,1297,1770,1985]" gridcols="3" gridrows="6" pageId="4" pageNumber="61">
|
||
<tr id="352DCE3E1C61FECF0619F9723849F8E9" box="[263,1297,1770,1794]" gridrow="0" pageId="4" pageNumber="61">
|
||
<th id="76FCA7421C61FECF0619F9723F35F8E9" box="[263,621,1770,1794]" gridcol="0" gridrow="0" pageId="4" pageNumber="61">generation</th>
|
||
<th id="76FCA7421C61FECF05DCF9723E9CF8E9" box="[706,964,1770,1794]" gridcol="1" gridrow="0" pageId="4" pageNumber="61">megalospheric (A) form</th>
|
||
<th id="76FCA7421C61FECF033FF9723849F8E9" box="[1057,1297,1770,1794]" gridcol="2" gridrow="0" pageId="4" pageNumber="61">microspheric (B) form</th>
|
||
</tr>
|
||
<tr id="352DCE3E1C61FECF0619F8923849F8C9" box="[263,1297,1802,1826]" gridrow="1" pageId="4" pageNumber="61">
|
||
<th id="76FCA7421C61FECF0619F8923F35F8C9" box="[263,621,1802,1826]" gridcol="0" gridrow="1" pageId="4" pageNumber="61">number chambers planispiral part</th>
|
||
<td id="76FCA7421C61FECF05DCF8923E9CF8C9" box="[706,964,1802,1826]" gridcol="1" gridrow="1" pageId="4" pageNumber="61">~10</td>
|
||
<td id="76FCA7421C61FECF033FF8923849F8C9" box="[1057,1297,1802,1826]" gridcol="2" gridrow="1" pageId="4" pageNumber="61">~26</td>
|
||
</tr>
|
||
<tr id="352DCE3E1C61FECF0619F8B23849F8A9" box="[263,1297,1834,1858]" gridrow="2" pageId="4" pageNumber="61">
|
||
<th id="76FCA7421C61FECF0619F8B23F35F8A9" box="[263,621,1834,1858]" gridcol="0" gridrow="2" pageId="4" pageNumber="61">diameter proloculus</th>
|
||
<td id="76FCA7421C61FECF05DCF8B23E9CF8A9" box="[706,964,1834,1858]" gridcol="1" gridrow="2" pageId="4" pageNumber="61">0.07-0.1 mm</td>
|
||
<td id="76FCA7421C61FECF033FF8B23849F8A9" box="[1057,1297,1834,1858]" gridcol="2" gridrow="2" pageId="4" pageNumber="61">not discernible</td>
|
||
</tr>
|
||
<tr id="352DCE3E1C61FECF0619F8D23849F86A" box="[263,1297,1866,1921]" gridrow="3" pageId="4" pageNumber="61">
|
||
<th id="76FCA7421C61FECF0619F8D23F35F86A" box="[263,621,1866,1921]" gridcol="0" gridrow="3" pageId="4" pageNumber="61">test diameter</th>
|
||
<td id="76FCA7421C61FECF05DCF8D23E9CF86A" box="[706,964,1866,1921]" gridcol="1" gridrow="3" pageId="4" pageNumber="61">3.1 mm (holotype)</td>
|
||
<td id="76FCA7421C61FECF033FF8D23849F86A" box="[1057,1297,1866,1921]" gridcol="2" gridrow="3" pageId="4" pageNumber="61">~4.6 mm (~4.2 preserved part)</td>
|
||
</tr>
|
||
<tr id="352DCE3E1C61FECF0619F8113849F84A" box="[263,1297,1929,1953]" gridrow="4" pageId="4" pageNumber="61">
|
||
<th id="76FCA7421C61FECF0619F8113F35F84A" box="[263,621,1929,1953]" gridcol="0" gridrow="4" pageId="4" pageNumber="61">adult final chambers</th>
|
||
<td id="76FCA7421C61FECF05DCF8113E9CF84A" box="[706,964,1929,1953]" gridcol="1" gridrow="4" pageId="4" pageNumber="61">flabelliform</td>
|
||
<td id="76FCA7421C61FECF033FF8113849F84A" box="[1057,1297,1929,1953]" gridcol="2" gridrow="4" pageId="4" pageNumber="61">annular</td>
|
||
</tr>
|
||
<tr id="352DCE3E1C61FECF0619F8313849F82A" box="[263,1297,1961,1985]" gridrow="5" pageId="4" pageNumber="61">
|
||
<th id="76FCA7421C61FECF0619F8313F35F82A" box="[263,621,1961,1985]" gridcol="0" gridrow="5" pageId="4" pageNumber="61">test morphology</th>
|
||
<td id="76FCA7421C61FECF05DCF8313E9CF82A" box="[706,964,1961,1985]" gridcol="1" gridrow="5" pageId="4" pageNumber="61">fan-shaped</td>
|
||
<td id="76FCA7421C61FECF033FF8313849F82A" box="[1057,1297,1961,1985]" gridcol="2" gridrow="5" pageId="4" pageNumber="61">ovoidal to discoidal</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF629CF41C60013507E5FF063861FF3F" pageId="5" pageNumber="62" startId="5.[251,316,158,182]" targetType="table">
|
||
<paragraph id="8BA2CC7C1C60013507E5FF063861FF3F" blockId="5.[251,1356,158,212]" pageId="5" pageNumber="62">
|
||
<emphasis id="B969106E1C60013507E5FF063C08FF5D" bold="true" box="[251,336,158,182]" pageId="5" pageNumber="62">Table 3</emphasis>
|
||
Summary table of the biometric parameters of
|
||
<taxonomicName id="4C1DB7FF1C6001350454FF063975FF5D" box="[842,1069,158,182]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="62" phylum="Foraminifera" rank="species" species="undetermined">
|
||
<emphasis id="B969106E1C6001350454FF063E8AFF5D" box="[842,978,158,182]" italics="true" pageId="5" pageNumber="62">Broeckinella</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
Data from
|
||
<bibRefCitation id="EF8CB18D1C60013503B8FF063810FF5D" author="Henson, F. R. S." box="[1190,1352,158,182]" pageId="5" pageNumber="62" refId="ref4860" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History), 127 pp." type="book" year="1948">Henson (1948)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8CB18D1C6001350611FF243CEFFF3F" author="Gendrot, C." box="[271,439,188,212]" pageId="5" pageNumber="62" pagination="657 - 694" refId="ref4824" refString="Gendrot, C., 1968. Stratigraphie et micropaleontologie du Senonien de la region des Martigues pres Marseille (Bouches-du-Rhone). Eclogae Geologicae Helvetiae, 61 (2): 657 - 694." type="journal article" year="1968">Gendrot (1968)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8CB18D1C60013506DCFF243E5EFF3F" author="Cherchi, A. & Schroeder, R." box="[450,774,188,212]" pageId="5" pageNumber="62" pagination="513 - 520" refId="ref4416" refString="Cherchi, A., Schroeder, R., 1978. Revision of the type of Broeckinella arabica Henson (Foram.) from Qatar Peninsula (Arabia). Neues Jahrbuch fur Geologie und Palaontologie Monatshefte, 9: 513 - 520." type="journal article" year="1978">Cherchi and Schroeder (1978)</bibRefCitation>
|
||
, and this work. * measured from the original papers.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA2CC7C1C60013506C7FF643987FDFB" pageId="5" pageNumber="62">
|
||
<table id="F91D3EDC1C60FECF07E9FF643814FDFA" box="[247,1356,252,529]" gridcols="4" gridrows="7" pageId="5" pageNumber="62">
|
||
<tr id="352DCE3E1C60FECF07E9FF643814FEFF" box="[247,1356,252,276]" gridrow="0" pageId="5" pageNumber="62">
|
||
<th id="76FCA7421C60FECF07E9FF643F7FFEFF" box="[247,551,252,276]" gridcol="0" gridrow="0" pageId="5" pageNumber="62">
|
||
<emphasis id="B969106E1C60013506C7FF643F7FFEFF" bold="true" box="[473,551,252,276]" pageId="5" pageNumber="62">species</emphasis>
|
||
</th>
|
||
<th id="76FCA7421C60FECF0554FF643E63FEFF" box="[586,827,252,276]" gridcol="1" gridrow="0" pageId="5" pageNumber="62">
|
||
<taxonomicName id="4C1DB7FF1C6001350594FF643FA4FEFF" box="[650,764,252,276]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="62" phylum="Foraminifera" rank="species" species="arabica">
|
||
<emphasis id="B969106E1C6001350594FF643FA4FEFF" bold="true" box="[650,764,252,276]" italics="true" pageId="5" pageNumber="62">B. arabica</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76FCA7421C60FECF0477FF643967FEFF" box="[873,1087,252,276]" gridcol="2" gridrow="0" pageId="5" pageNumber="62">
|
||
<taxonomicName id="4C1DB7FF1C600135049DFF64397CFEFF" box="[899,1060,252,276]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="62" phylum="Foraminifera" rank="species" species="neumannae">
|
||
<emphasis id="B969106E1C600135049DFF64397CFEFF" bold="true" box="[899,1060,252,276]" italics="true" pageId="5" pageNumber="62">B. neumannae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76FCA7421C60FECF0374FF643814FEFF" box="[1130,1356,252,276]" gridcol="3" gridrow="0" pageId="5" pageNumber="62">
|
||
<emphasis id="B969106E1C600135039FFF64386DFEFF" bold="true" box="[1153,1333,252,276]" pageId="5" pageNumber="62">
|
||
<taxonomicName id="4C1DB7FF1C600135039FFF6439ABFEFF" authorityName="SCHLAGINTWEIT & RASHIDI" authorityYear="2020" box="[1153,1267,252,276]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="62" phylum="Foraminifera" rank="species" species="grandis" status="n. sp.">
|
||
<emphasis id="B969106E1C600135039FFF6439ABFEFF" bold="true" box="[1153,1267,252,276]" italics="true" pageId="5" pageNumber="62">B. grandis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25AAD151C60013503E7FF64386DFEFF" box="[1273,1333,252,276]" pageId="5" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="352DCE3E1C60FECF07E9FE833814FEB9" box="[247,1356,283,338]" gridrow="1" pageId="5" pageNumber="62">
|
||
<th id="76FCA7421C60FECF07E9FE833F7FFEB9" box="[247,551,283,338]" gridcol="0" gridrow="1" pageId="5" pageNumber="62">Type-locality</th>
|
||
<td id="76FCA7421C60FECF0554FE833E63FEB9" box="[586,827,283,338]" gridcol="1" gridrow="1" pageId="5" pageNumber="62">
|
||
Maastrichtian of Qatar (
|
||
<bibRefCitation id="EF8CB18D1C6001350569FEA23E56FEB9" author="Henson, F. R. S." box="[631,782,314,338]" pageId="5" pageNumber="62" refId="ref4860" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History), 127 pp." type="book" year="1948">Henson, 1948</bibRefCitation>
|
||
)
|
||
</td>
|
||
<td id="76FCA7421C60FECF0477FE833967FEB9" box="[873,1087,283,338]" gridcol="2" gridrow="1" pageId="5" pageNumber="62">
|
||
Santonian of France
|
||
<bibRefCitation id="EF8CB18D1C600135049FFEA2397FFEB9" author="Gendrot, C." box="[897,1063,314,338]" pageId="5" pageNumber="62" pagination="657 - 694" refId="ref4824" refString="Gendrot, C., 1968. Stratigraphie et micropaleontologie du Senonien de la region des Martigues pres Marseille (Bouches-du-Rhone). Eclogae Geologicae Helvetiae, 61 (2): 657 - 694." type="journal article" year="1968">Gendrot (1968)</bibRefCitation>
|
||
</td>
|
||
<td id="76FCA7421C60FECF0374FE833814FEB9" box="[1130,1356,283,338]" gridcol="3" gridrow="1" pageId="5" pageNumber="62">Maastrichtian of Iran</td>
|
||
</tr>
|
||
<tr id="352DCE3E1C60FECF07E9FEC23814FE7A" box="[247,1356,346,401]" gridrow="2" pageId="5" pageNumber="62">
|
||
<th id="76FCA7421C60FECF07E9FEC23F7FFE7A" box="[247,551,346,401]" gridcol="0" gridrow="2" pageId="5" pageNumber="62">Test diameter (mm)</th>
|
||
<td id="76FCA7421C60FECF0554FEC23E63FE7A" box="[586,827,346,401]" gridcol="1" gridrow="2" pageId="5" pageNumber="62">up to 3.1 (A-form) up to 4.6 (B-form)</td>
|
||
<td id="76FCA7421C60FECF0477FEC23967FE7A" box="[873,1087,346,401]" gridcol="2" gridrow="2" pageId="5" pageNumber="62">up to 1.5 mm (A-form)</td>
|
||
<td id="76FCA7421C60FECF0374FEC23814FE7A" box="[1130,1356,346,401]" gridcol="3" gridrow="2" pageId="5" pageNumber="62">up to 7.5 (?B-form)</td>
|
||
</tr>
|
||
<tr id="352DCE3E1C60FECF07E9FE013814FE5A" box="[247,1356,409,433]" gridrow="3" pageId="5" pageNumber="62">
|
||
<th id="76FCA7421C60FECF07E9FE013F7FFE5A" box="[247,551,409,433]" gridcol="0" gridrow="3" pageId="5" pageNumber="62">Test thickness (mm)</th>
|
||
<td id="76FCA7421C60FECF0554FE013E63FE5A" box="[586,827,409,433]" gridcol="1" gridrow="3" pageId="5" pageNumber="62">~0.3</td>
|
||
<td id="76FCA7421C60FECF0477FE013967FE5A" box="[873,1087,409,433]" gridcol="2" gridrow="3" pageId="5" pageNumber="62">up to 0.22</td>
|
||
<td id="76FCA7421C60FECF0374FE013814FE5A" box="[1130,1356,409,433]" gridcol="3" gridrow="3" pageId="5" pageNumber="62">up to 0.4</td>
|
||
</tr>
|
||
<tr id="352DCE3E1C60FECF07E9FE213814FE3A" box="[247,1356,441,465]" gridrow="4" pageId="5" pageNumber="62">
|
||
<th id="76FCA7421C60FECF07E9FE213F7FFE3A" box="[247,551,441,465]" gridcol="0" gridrow="4" pageId="5" pageNumber="62">Number chambers last mm</th>
|
||
<td id="76FCA7421C60FECF0554FE213E63FE3A" box="[586,827,441,465]" gridcol="1" gridrow="4" pageId="5" pageNumber="62">8-14 *</td>
|
||
<td id="76FCA7421C60FECF0477FE213967FE3A" box="[873,1087,441,465]" gridcol="2" gridrow="4" pageId="5" pageNumber="62">6-8 *</td>
|
||
<td id="76FCA7421C60FECF0374FE213814FE3A" box="[1130,1356,441,465]" gridcol="3" gridrow="4" pageId="5" pageNumber="62">5-7</td>
|
||
</tr>
|
||
<tr id="352DCE3E1C60FECF07E9FE413814FE1A" box="[247,1356,473,497]" gridrow="5" pageId="5" pageNumber="62">
|
||
<th id="76FCA7421C60FECF07E9FE413F7FFE1A" box="[247,551,473,497]" gridcol="0" gridrow="5" pageId="5" pageNumber="62">Diameter foramina (mm)</th>
|
||
<td id="76FCA7421C60FECF0554FE413E63FE1A" box="[586,827,473,497]" gridcol="1" gridrow="5" pageId="5" pageNumber="62">0.02-0.04</td>
|
||
<td id="76FCA7421C60FECF0477FE413967FE1A" box="[873,1087,473,497]" gridcol="2" gridrow="5" pageId="5" pageNumber="62">0.06</td>
|
||
<td id="76FCA7421C60FECF0374FE413814FE1A" box="[1130,1356,473,497]" gridcol="3" gridrow="5" pageId="5" pageNumber="62">0.05-0.07</td>
|
||
</tr>
|
||
<tr id="352DCE3E1C60FECF07E9FE613814FDFA" box="[247,1356,505,529]" gridrow="6" pageId="5" pageNumber="62">
|
||
<th id="76FCA7421C60FECF07E9FE613F7FFDFA" box="[247,551,505,529]" gridcol="0" gridrow="6" pageId="5" pageNumber="62">Proloculus diameter (mm)</th>
|
||
<td id="76FCA7421C60FECF0554FE613E63FDFA" box="[586,827,505,529]" gridcol="1" gridrow="6" pageId="5" pageNumber="62">~0.1</td>
|
||
<td id="76FCA7421C60FECF0477FE613967FDFA" box="[873,1087,505,529]" gridcol="2" gridrow="6" pageId="5" pageNumber="62">0.07-0.09 *</td>
|
||
<td id="76FCA7421C60FECF0374FE613814FDFA" box="[1130,1356,505,529]" gridcol="3" gridrow="6" pageId="5" pageNumber="62">-</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF629CF41C60013507FFF93B3F89F8E7" ID-DOI="http://doi.org/10.5281/zenodo.12583420" ID-Zenodo-Dep="12583420" httpUri="https://zenodo.org/record/12583420/files/figure.png" pageId="5" pageNumber="62" startId="5.[225,258,1699,1720]" targetBox="[225,1386,576,1662]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BA2CC7C1C60013507FFF93B3F89F8E7" blockId="5.[225,1363,1699,1804]" pageId="5" pageNumber="62">
|
||
<emphasis id="B969106E1C60013507FFF93B3C44F953" bold="true" box="[225,284,1699,1720]" pageId="5" pageNumber="62">Fig. 5</emphasis>
|
||
<taxonomicName id="4C1DB7FF1C6001350630F93B3CA3F952" authorityName="SCHLAGINTWEIT & RASHIDI" authorityYear="2020" box="[302,507,1699,1721]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="62" phylum="Foraminifera" rank="species" species="hensoni" status="sp. nov.">
|
||
<emphasis id="B969106E1C6001350630F93B3CA3F952" box="[302,507,1699,1721]" italics="true" pageId="5" pageNumber="62">Broeckinella hensoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25AAD151C600135051DF93C3F61F952" box="[515,569,1700,1721]" pageId="5" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
|
||
(
|
||
<emphasis id="B969106E1C6001350556F93B3F0CF953" bold="true" box="[584,596,1699,1720]" pageId="5" pageNumber="62">a</emphasis>
|
||
) and
|
||
<taxonomicName id="4C1DB7FF1C6001350591F93B3EF6F952" authority="Henson" authorityName="Henson" box="[655,942,1699,1721]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="62" phylum="Foraminifera" rank="species" species="arabica">
|
||
<emphasis id="B969106E1C6001350591F93B3E04F952" box="[655,860,1699,1721]" italics="true" pageId="5" pageNumber="62">Broeckinella arabica</emphasis>
|
||
Henson
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B969106E1C60013504A3F93B3EBCF953" bold="true" box="[957,996,1699,1720]" pageId="5" pageNumber="62">b–d</emphasis>
|
||
) from the upper Maastrichtian of the Tarbur Formation, Naghan section.
|
||
<emphasis id="B969106E1C600135055DF9273F17F93F" bold="true" box="[579,591,1727,1748]" pageId="5" pageNumber="62">a</emphasis>
|
||
Foraminiferal wackestone with
|
||
<taxonomicName id="4C1DB7FF1C600135048EF9273905F93E" authorityName="SCHLAGINTWEIT & RASHIDI" authorityYear="2020" box="[912,1117,1727,1749]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="62" phylum="Foraminifera" rank="species" species="hensoni" status="sp. nov.">
|
||
<emphasis id="B969106E1C600135048EF9273905F93E" box="[912,1117,1727,1749]" italics="true" pageId="5" pageNumber="62">Broeckinella hensoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25AAD151C6001350379F95839C6F93E" box="[1127,1182,1728,1749]" pageId="5" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
|
||
, thin-section 2NG 197.
|
||
<emphasis id="B969106E1C600135060CF9433C47F91B" bold="true" box="[274,287,1755,1776]" pageId="5" pageNumber="62">b</emphasis>
|
||
Foraminiferal wackestone with
|
||
<taxonomicName id="4C1DB7FF1C6001350545F9423E22F91B" authority="Henson" authorityName="Henson" box="[603,890,1754,1776]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="5" pageNumber="62" phylum="Foraminifera" rank="species" species="arabica">
|
||
<emphasis id="B969106E1C6001350545F9423E70F91B" box="[603,808,1754,1776]" italics="true" pageId="5" pageNumber="62">Broeckinella arabica</emphasis>
|
||
Henson
|
||
</taxonomicName>
|
||
, 2NG 128-1.
|
||
<emphasis id="B969106E1C600135031FF943397DF91B" bold="true" box="[1025,1061,1755,1776]" pageId="5" pageNumber="62">c–d</emphasis>
|
||
Details from
|
||
<emphasis id="B969106E1C60013503B3F94339E2F91B" bold="true" box="[1197,1210,1755,1776]" pageId="5" pageNumber="62">b</emphasis>
|
||
, showing a microspheric specimen. Scale bars in
|
||
<emphasis id="B969106E1C600135052CF96E3F65F8E0" bold="true" box="[562,573,1782,1803]" pageId="5" pageNumber="62">c</emphasis>
|
||
and
|
||
<emphasis id="B969106E1C6001350572F96E3F21F8E0" bold="true" box="[620,633,1782,1803]" pageId="5" pageNumber="62">d</emphasis>
|
||
= 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3079FF71C6301360789FF063FAFFE85" pageId="6" pageNumber="63" type="etymology">
|
||
<paragraph id="8BA2CC7C1C6301360789FF063FFCFEDA" blockId="6.[151,772,158,1256]" pageId="6" pageNumber="63">
|
||
<emphasis id="B969106E1C6301360789FF063C43FF5D" bold="true" box="[151,283,158,182]" pageId="6" pageNumber="63">Etymology:</emphasis>
|
||
The species name is dedicated to Francis Roger Spencer Henson (1901-1967) in honour for his fundamental works on larger imperforate foraminifera from the Middle East area. For further information on F.R.S. Henson and his merits see
|
||
<bibRefCitation id="EF8CB18D1C63013606E0FE813FF9FEDA" author="Hughes, G. W." box="[510,673,281,305]" pageId="6" pageNumber="63" pagination="145 - 157" refId="ref4898" refString="Hughes, G. W., 2013. Micropalaeontologists of Saudi Arabia and other Gulf countries. In: Bowden, A. J., Gregory, F. J., Henderson, A. S. (eds): Landmarks in Foraminiferal Micropalaeontology: History and Development. The Micropaleontological Society, Spec. Pubs.: 145 - 157." type="book chapter" year="2013">Hughes (2013)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA2CC7C1C6301360789FEAF3F3CFEA4" blockId="6.[151,772,158,1256]" box="[151,612,311,335]" pageId="6" pageNumber="63">
|
||
<emphasis id="B969106E1C6301360789FEAF3C6EFEA4" bold="true" box="[151,310,311,335]" pageId="6" pageNumber="63">Type-locality:</emphasis>
|
||
Naghan section (
|
||
<figureCitation id="1326D0F91C63013606EEFEAF3F0EFEA4" box="[496,598,311,335]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[425,460,1385,1406]" captionStartId-1="3.[285,320,677,698]" captionTargetBox-0="[243,1415,157,1348]" captionTargetBox-1="[182,1432,149,640]" captionTargetId-0="figure-639@2.[243,1415,157,1348]" captionTargetId-1="figure-1214@3.[182,1432,149,640]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 2 Location of the Naghan and Mandegan sections in the Zagros Zone, SW Iran." captionText-1="Fig. 3 Naghan section, along the road from Naghan to Izeh with the type-level of Broeckinella hensoni n. sp. marked by an asterisk (left side)." figureDoi-0="http://doi.org/10.5281/zenodo.12583412" figureDoi-1="http://doi.org/10.5281/zenodo.12583414" httpUri-0="https://zenodo.org/record/12583412/files/figure.png" httpUri-1="https://zenodo.org/record/12583414/files/figure.png" pageId="6" pageNumber="63">Figs. 2–3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA2CC7C1C6301360789FECE3FAFFE85" blockId="6.[151,772,158,1256]" box="[151,759,342,366]" pageId="6" pageNumber="63">
|
||
<emphasis id="B969106E1C6301360789FECE3C4EFE85" bold="true" box="[151,278,342,366]" pageId="6" pageNumber="63">Type-level:</emphasis>
|
||
Late Maastrichtian of the Tarbur Formation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3079FF71C6301360789FEED3F50FB0C" pageId="6" pageNumber="63" type="description">
|
||
<paragraph id="8BA2CC7C1C6301360789FEED3F9FFC38" blockId="6.[151,772,158,1256]" pageId="6" pageNumber="63">
|
||
<emphasis id="B969106E1C6301360789FEED3C7CFE66" bold="true" box="[151,292,373,397]" pageId="6" pageNumber="63">Description:</emphasis>
|
||
Test finely, agglutinating, large (up to almost
|
||
<quantity id="4CE561991C63013607C8FE0B3C43FE40" box="[214,283,403,427]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="63" unit="mm" value="8.0">8 mm</quantity>
|
||
), flattened with an early evolute planispiral stage followed by a flabelli- to reniform adult growth stage (
|
||
<figureCitation id="1326D0F91C63013607FDFE493C6AFE02" box="[227,306,465,489]" captionStart="Fig" captionStartId="8.[203,236,1747,1768]" captionTargetBox="[203,1406,252,1710]" captionTargetId="figure-60@8.[203,1406,252,1710]" captionTargetPageId="8" captionText="Fig. 7 Broeckinella hensoni n. sp., upper Maastrichtian Tarbur Formation of the Naghan section, Zagros Zone, SW Iran. a, c Subaxial sections. Specimen shown in A displays more than 40 chambers in the uncoiled adult part. d Detail from c showing initial coiled part. e Oblique equatorial section. f Slightly oblique equatorial section showing flabelliform test morphology (test diameter: ~6.6 mm). Thin-sections: 2NG 153 (a), 2NGN (b, f), 2NG 38 (c–d), NG 197-1 (e). Scale bars 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12583424" httpUri="https://zenodo.org/record/12583424/files/figure.png" pageId="6" pageNumber="63">Fig. 7f</figureCitation>
|
||
). Axial sections display almost constant chamber width (
|
||
<figureCitation id="1326D0F91C630136065BFE773CC8FDEC" box="[325,400,495,519]" captionStart="Fig" captionStartId="8.[203,236,1747,1768]" captionTargetBox="[203,1406,252,1710]" captionTargetId="figure-60@8.[203,1406,252,1710]" captionTargetPageId="8" captionText="Fig. 7 Broeckinella hensoni n. sp., upper Maastrichtian Tarbur Formation of the Naghan section, Zagros Zone, SW Iran. a, c Subaxial sections. Specimen shown in A displays more than 40 chambers in the uncoiled adult part. d Detail from c showing initial coiled part. e Oblique equatorial section. f Slightly oblique equatorial section showing flabelliform test morphology (test diameter: ~6.6 mm). Thin-sections: 2NG 153 (a), 2NGN (b, f), 2NG 38 (c–d), NG 197-1 (e). Scale bars 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12583424" httpUri="https://zenodo.org/record/12583424/files/figure.png" pageId="6" pageNumber="63">Fig. 7a</figureCitation>
|
||
). Further details on the early stage are limited by the available sections. The adult part consists of up to 50 chambers. There are five to seven chambers in the last mm of the test. Wall finely agglutinated. Exoskeleton consisting of one short horizontal and two orders (short, and longer) of subepidermal vertical (or radial) partitions (beams and rafters) together forming a subepidermal network (
|
||
<figureCitation id="1326D0F91C6301360684FD5E3CB0FD35" box="[410,488,710,734]" captionStart="Fig" captionStartId="9.[193,226,1666,1687]" captionTargetBox="[193,1394,268,1629]" captionTargetId="figure-37@9.[193,1394,268,1629]" captionTargetPageId="9" captionText="Fig. 8 Broeckinella hensoni n. sp., upper Maastrichtian Tarbur Formation of the Naghan (a, d–e) and Mandegan (b–c, f) sections, Zagros Zone, SW Iran. a Oblique equatorial section. Note porcelaneous test of Cuvillerinella sp. below. b Detail showing fine subepidermal network (sn, below), followed by zone with radial partitions (rp) only and finally the undivided central chamber part with foramina (f) piercing the septa (s). c Oblique section cutting the initial planispiral part. d–e Oblique equatorial sections showing zones of different internal structure (see Text-Figure 3). f Detail from c showing initial planispirally coiled part with numerous chambers. Note also the fine subepidermal network and main partitions (right side). Thin-sections: 2NG 166 (a), Rt 67-1 (b), RT 72 (c, f), 2NG 118 (d), 2NG 172 (e). Scale bars 1.0 mm for a, c–e, 0.2 mm for b and f." figureDoi="http://doi.org/10.5281/zenodo.12583426" httpUri="https://zenodo.org/record/12583426/files/figure.png" pageId="6" pageNumber="63">Fig. 8e</figureCitation>
|
||
). Due to the shortness of the secondary partitions this network of polygonal cells is only visible in shallow tangential sections. Longer vertical partitions (beams) are aligned between successive chambers, and reach slightly deeper into the chamber lumen. The central part of the chamber lumen is undivid- ed (e.g.,
|
||
<figureCitation id="1326D0F91C630136061EFCE63C08FC7D" box="[256,336,894,918]" captionStart="Fig" captionStartId="9.[193,226,1666,1687]" captionTargetBox="[193,1394,268,1629]" captionTargetId="figure-37@9.[193,1394,268,1629]" captionTargetPageId="9" captionText="Fig. 8 Broeckinella hensoni n. sp., upper Maastrichtian Tarbur Formation of the Naghan (a, d–e) and Mandegan (b–c, f) sections, Zagros Zone, SW Iran. a Oblique equatorial section. Note porcelaneous test of Cuvillerinella sp. below. b Detail showing fine subepidermal network (sn, below), followed by zone with radial partitions (rp) only and finally the undivided central chamber part with foramina (f) piercing the septa (s). c Oblique section cutting the initial planispiral part. d–e Oblique equatorial sections showing zones of different internal structure (see Text-Figure 3). f Detail from c showing initial planispirally coiled part with numerous chambers. Note also the fine subepidermal network and main partitions (right side). Thin-sections: 2NG 166 (a), Rt 67-1 (b), RT 72 (c, f), 2NG 118 (d), 2NG 172 (e). Scale bars 1.0 mm for a, c–e, 0.2 mm for b and f." figureDoi="http://doi.org/10.5281/zenodo.12583426" httpUri="https://zenodo.org/record/12583426/files/figure.png" pageId="6" pageNumber="63">Fig. 8e</figureCitation>
|
||
in the lower part; see
|
||
<bibRefCitation id="EF8CB18D1C630136057CFCE63FA7FC7D" author="Henson, F. R. S." box="[610,767,894,918]" pageId="6" pageNumber="63" refId="ref4860" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History), 127 pp." type="book" year="1948">Henson, 1948</bibRefCitation>
|
||
: fig.13c). Multiple foramina (one row) arranged in the median plane. Microspheric specimens not reported.
|
||
</paragraph>
|
||
<paragraph id="8BA2CC7C1C6301360789FC423CFEFC19" blockId="6.[151,772,158,1256]" box="[151,422,986,1010]" pageId="6" pageNumber="63">
|
||
<emphasis id="B969106E1C6301360789FC423C7CFC19" bold="true" box="[151,292,986,1010]" pageId="6" pageNumber="63">Dimensions:</emphasis>
|
||
See table 3.
|
||
</paragraph>
|
||
<paragraph id="8BA2CC7C1C6301360789FC613F50FB0C" blockId="6.[151,772,158,1256]" pageId="6" pageNumber="63">
|
||
<emphasis id="B969106E1C6301360789FC613C6FFBFA" bold="true" box="[151,311,1017,1041]" pageId="6" pageNumber="63">Comparisons:</emphasis>
|
||
Amongst the three Late Cretaceous species of the genus,
|
||
<taxonomicName id="4C1DB7FF1C6301360647FB8F3F63FBC4" authorityName="SCHLAGINTWEIT & RASHIDI" authorityYear="2020" box="[345,571,1047,1071]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="6" pageNumber="63" phylum="Foraminifera" rank="species" species="hensoni">
|
||
<emphasis id="B969106E1C6301360647FB8F3F63FBC4" box="[345,571,1047,1071]" italics="true" pageId="6" pageNumber="63">Broeckinella hensoni</emphasis>
|
||
</taxonomicName>
|
||
is distinctly larger then
|
||
<taxonomicName id="4C1DB7FF1C63013607CFFBAF3CFCFBA6" authority="Henson" authorityName="Henson" box="[209,420,1078,1102]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="6" pageNumber="63" phylum="Foraminifera" rank="species" species="arabica">
|
||
<emphasis id="B969106E1C63013607CFFBAF3C1EFBA5" box="[209,326,1078,1102]" italics="true" pageId="6" pageNumber="63">B. arabica</emphasis>
|
||
Henson
|
||
</taxonomicName>
|
||
(double the size!) and
|
||
<taxonomicName id="4C1DB7FF1C63013605AFFBAF3C15FB87" authority="Gendrot" authorityName="Gendrot" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="6" pageNumber="63" phylum="Foraminifera" rank="species" species="neumannae">
|
||
<emphasis id="B969106E1C63013605AFFBAF3DB4FB87" italics="true" pageId="6" pageNumber="63">B. neumannae</emphasis>
|
||
Gendrot
|
||
</taxonomicName>
|
||
(five times larger!) (see table 3). With respect to the time-equivalent
|
||
<taxonomicName id="4C1DB7FF1C63013606C4FBEC3F13FB60" box="[474,587,1139,1163]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="6" pageNumber="63" phylum="Foraminifera" rank="species" species="arabica">
|
||
<emphasis id="B969106E1C63013606C4FBEC3F13FB60" box="[474,587,1139,1163]" italics="true" pageId="6" pageNumber="63">B. arabica</emphasis>
|
||
</taxonomicName>
|
||
, the distinctively lower number of chambers per last mm test length (
|
||
<quantity id="4CE561991C63013605C9FB0A3D89FB23" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.556" metricValueMin="2.032" pageId="6" pageNumber="63" unit="in" value="11.0" valueMax="14.0" valueMin="8.0">8 to 14 in</quantity>
|
||
<taxonomicName id="4C1DB7FF1C63013607C5FB293C16FB23" box="[219,334,1200,1224]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="6" pageNumber="63" phylum="Foraminifera" rank="species" species="arabica">
|
||
<emphasis id="B969106E1C63013607C5FB293C16FB23" box="[219,334,1200,1224]" italics="true" pageId="6" pageNumber="63">B. arabica</emphasis>
|
||
</taxonomicName>
|
||
against
|
||
<quantity id="4CE561991C63013606B0FB283F48FB23" box="[430,528,1200,1224]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" metricValueMax="1.778" metricValueMin="1.27" pageId="6" pageNumber="63" unit="in" value="6.0" valueMax="7.0" valueMin="5.0">5 to 7 in</quantity>
|
||
<taxonomicName id="4C1DB7FF1C6301360504FB293FD6FB23" authorityName="SCHLAGINTWEIT & RASHIDI" authorityYear="2020" box="[538,654,1200,1224]" class="Globothalamea" family="Cyclamminidae" genus="Broeckinella" kingdom="Chromista" order="Loftusiida" pageId="6" pageNumber="63" phylum="Foraminifera" rank="species" species="hensoni">
|
||
<emphasis id="B969106E1C6301360504FB293FD6FB23" box="[538,654,1200,1224]" italics="true" pageId="6" pageNumber="63">B. hensoni</emphasis>
|
||
</taxonomicName>
|
||
), due to a greater chamber height, is notable.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |