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16. 
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<figureCitation id="1320C6D8FFFEFFA2FD5DFAC3AFC4EFEF" box="[644,739,1404,1430]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[189,1398,788,1816]" captionTargetId="figure-11@5.[189,1398,788,1816]" captionTargetPageId="5" captionText="Fig. 1. Leaf margins of Peruvian Begonia. A. Entire, aciliate margins (B. stenotepala A.DC.). B. Crenate, aciliate margins (B. joshii Moonlight). C. Denticulate, ciliate margins (B. acerifolia Kunth). D. Irregularly double-dentate, ciliate margins (B. obtecticaulis Irmsch.). E. Serrulate, aciliate margins (B. glauca (Klotzsch) Ruiz &amp; Pav. exA.DC.). F. Serrate, ciliate margins (B. foliosa Kunth). Photographs taken by D.A. Purvis &amp; P.W. Moonlight in the living collections of the Royal Botanic Garden Edinburgh." figureDoi="http://doi.org/10.5281/zenodo.8178282" httpUri="https://zenodo.org/record/8178282/files/figure.png" pageId="91" pageNumber="92">Figs 1D</figureCitation>
, 
<figureCitation id="1320C6D8FFFEFFA2FD28FAC3AE32EFEF" box="[753,789,1404,1430]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[272,1315,265,1740]" captionTargetId="figure-13@6.[272,1315,265,1740]" captionTargetPageId="6" captionText="Fig. 2. Venation in Peruvian Begonia. A. Pinnate venation (B. yaracuyensis Moonlight). B. Pinnate venation, 3-veined from the base (B. glabra Aubl.). C. Palmate-pinnate venation (B. obtecticaulis Irmsch.). D. Palmate venation with one primary vein (B. stenotepalaL.B.Sm. &amp; B.G.Schub.).E. Palmate venation (B. lamolina Moonlight). F. Peltate venation with one primary vein (B. acerifolia Kunth.). G. Peltate venation (B. joshii Moonlight). Primary veins are shown in white, secondary veins from the petiole insertion in blue, and secondary veins from the primary veins in green. Photographs taken by D.A. Purvis &amp; P.W. Moonlight in the living collections of the Royal Botanic Garden Edinburgh." figureDoi="http://doi.org/10.5281/zenodo.8178284" httpUri="https://zenodo.org/record/8178284/files/figure.png" pageId="91" pageNumber="92">2C</figureCitation>
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<figureCitation id="1320C6D8FFFEFFA2FC88FAC3AEA1EFEF" box="[849,902,1404,1430]" captionStart="Fig" captionStartId="66.[189,232,1787,1813]" captionTargetBox="[194,1391,514,1732]" captionTargetId="figure-13@66.[189,1398,504,1740]" captionTargetPageId="66" captionText="Fig. 24. Distribution ofBegonia sect.Cyathocnemis (Klotzsch)A.DC. in Peru and surrounding countries. A. B. amoeboides Moonlight (red) andB. imbrexiformis Moonlight sp. nov. (blue).B. B. bracteosaA.DC. (red) and B. longinqua Moonlight (blue). C. B. cyathophora Poepp. &amp; Endl. (red), B. serratistipula Moonlight sp. nov. (blue) and C. vargasii Moonlight sp. nov. (green). D. B. lamolina Moonlight (red), B. lophoptera Rolfe (blue) and B. obtecticaulis Irmsch. (green). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178339" httpUri="https://zenodo.org/record/8178339/files/figure.png" pageId="91" pageNumber="92">24D</figureCitation>
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<figureCitation id="1320C6D8FFFEFFA2FC4BFAC3AE88EFEF" box="[914,943,1404,1430]" captionStart="Fig" captionStartId="92.[189,232,1787,1813]" captionTargetBox="[272,1315,265,1740]" captionTargetId="figure-13@92.[272,1315,265,1740]" captionTargetPageId="92" captionText="Fig. 33. Begonia obtecticaulis Irmsch. A. Habit. B. Leaf, abaxial surface. C. Leaf, adaxial surface. D. Stipule. E. Inflorescence. F. Bract. G. Pistillate flower, side view. H. Pistillate flower, front view. I. Cross section of ovary. J. Largest tepal of the pistillate flower. K. Smallest tepal of the pistillate flower. L. Pistils, side view. M. Pistils, front view. N. Staminate flower, side view. O. Staminate flower, front view. P. Androecium, side view. All photographs taken by D.A. Purvis in the living collections of the Royal Botanic Garden Edinburgh (Accession 20160137a, grown from seeds collected as part of P.W. Moonlight &amp; A. Daza 209)." figureDoi="http://doi.org/10.5281/zenodo.8178365" httpUri="https://zenodo.org/record/8178365/files/figure.png" pageId="91" pageNumber="92">33</figureCitation>
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Botanische Jahrbücher für 
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74 (4): 600 (
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<bibRefCitation id="EF8AA7ACFFFEFFA2FF64F991AF00EC31" author="Brako &amp; Zarucchi" box="[189,551,1582,1608]" firstAuthor="Brako" page="194" pageId="91" pageNumber="92" refId="ref225797" refString="Brako L. &amp; Zarucchi L. 1993. Catalogue of the Flowering Plants and Gymnosperms of Peru. Monographs in Systematic Botany from the Missouri Botanical Garden 45, Missouri Botanical Garden, St Louis." type="book" year="1993">Brako &amp; Zarucchi (1993: 194)</bibRefCitation>
; 
<bibRefCitation id="EF8AA7ACFFFEFFA2FDEAF991AEBAEC31" author="Leon &amp; Monsalve" box="[563,925,1582,1608]" firstAuthor="Leon" page="167" pageId="91" pageNumber="92" pagination="165 - 170" refId="ref229226" refString="Leon B. &amp; Monsalve C. 2006. Begoniaceae endemicas del Peru. Revista Peruana de Biologia 13 (2): 165 - 170. https: // doi. org / 10.15381 / rpb. v 13 i 2.1810" type="journal article" year="2006">
<collectingRegion id="49DF14BFFFFEFFA2FDEAF991AF48EC31" box="[563,623,1582,1608]" country="Nicaragua" name="Leon" pageId="91" pageNumber="92">León</collectingRegion>
&amp; Monsalve (2006: 167)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFFEFFA2FF64F9CDA972EC98" pageId="91" pageNumber="92" type="etymology">
<paragraph id="8BA4DA5DFFFEFFA2FF64F9CDAC62ECF5" blockId="91.[189,325,1650,1676]" box="[189,325,1650,1676]" pageId="91" pageNumber="92">
<heading id="D0EC6D31FFFEFFA2FF64F9CDAC62ECF5" bold="true" box="[189,325,1650,1676]" fontSize="11" level="3" pageId="91" pageNumber="92" reason="3">
<emphasis id="B96F064FFFFEFFA2FF64F9CDAC62ECF5" bold="true" box="[189,325,1650,1676]" pageId="91" pageNumber="92">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFFEFFA2FF64F91DA972EC98" blockId="91.[189,1398,1698,1761]" pageId="91" pageNumber="92">
The epithet derives from the Latin words ‘ 
<emphasis id="B96F064FFFFEFFA2FD18F91DAE03ECC5" box="[705,804,1698,1724]" italics="true" pageId="91" pageNumber="92">obtectus</emphasis>
’ and ‘ 
<emphasis id="B96F064FFFFEFFA2FCAFF91DAE9AECC5" box="[886,957,1698,1724]" italics="true" pageId="91" pageNumber="92">caulis</emphasis>
’, meaning ‘hidden’ and ‘stem’. This refers to the large, persistent stipules that clasp the stem and hide it from view.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFFEFFA4FF64F8B5A9A9EB3E" lastPageId="93" lastPageNumber="94" pageId="91" pageNumber="92" type="materials_examined">
<paragraph id="8BA4DA5DFFFEFFA2FF64F8B5AF09ED5D" blockId="91.[189,558,1802,1828]" box="[189,558,1802,1828]" pageId="91" pageNumber="92">
<heading id="D0EC6D31FFFEFFA2FF64F8B5AF09ED5D" bold="true" box="[189,558,1802,1828]" fontSize="11" level="3" pageId="91" pageNumber="92" reason="3">
<emphasis id="B96F064FFFFEFFA2FF64F8B5AF09ED5D" bold="true" box="[189,558,1802,1828]" pageId="91" pageNumber="92">Selected specimens examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFFEFFA4FF64F884A9A9EB3E" blockId="91.[189,1400,1849,2025]" lastBlockId="93.[189,1399,263,327]" lastPageId="93" lastPageNumber="94" pageId="91" pageNumber="92">
<materialsCitation id="3B73D000FFFEFFA2FF64F884A9CFED03" collectingDate="1965-08-25" collectionCode="K, NY" collectorName="A. C. Hamilton &amp; P. M. Holligan" country="Peru" county="Prov. Huallaga" elevation="2600" latitude="-7.9166665" location="Valley of rio Apisoncho" longLatPrecision="1302" longitude="-77.166664" municipality="Valley" pageId="91" pageNumber="92" specimenCount="1" stateProvince="San Martin Region">
<collectingCountry id="F30C9ACDFFFEFFA2FF64F884AC2FED2C" box="[189,264,1851,1877]" name="Peru" pageId="91" pageNumber="92">PERU</collectingCountry>
– 
<emphasis id="B96F064FFFFEFFA2FEF2F885AFD7ED2C" bold="true" box="[299,752,1849,1877]" pageId="91" pageNumber="92">
<collectingRegion id="49DF14BFFFFEFFA2FEF2F885AF02ED2D" box="[299,549,1849,1876]" country="Peru" name="San Martin" pageId="91" pageNumber="92">San Martín Region</collectingRegion>
: 
<collectingCounty id="62C5A2D1FFFEFFA2FDEBF884AFD7ED2C" box="[562,752,1851,1877]" pageId="91" pageNumber="92">Prov. Huallaga</collectingCounty>
</emphasis>
• 
<location id="8EC48C86FFFEFFA2FCD6F885A908ED2D" LSID="urn:lsid:plazi:treatment:03B26B4BFFFEFFA7FD8BFAE8A850EB10:8EC48C86FFFEFFA2FCD6F885A908ED2D" box="[783,1071,1850,1877]" country="Peru" county="Prov. Huallaga" latitude="-7.9166665" longLatPrecision="1302" longitude="-77.166664" municipality="Valley" name="Valley of rio Apisoncho" pageId="91" pageNumber="92" stateProvince="San Martin Region">Valley of río Apisoncho</location>
, 
<locationDeviation id="9A16C37FFFFEFFA2FBE5F884A853ED2D" box="[1084,1396,1850,1877]" location="Jucusbamba" pageId="91" pageNumber="92">
<quantity id="4CE377B8FFFEFFA2FBE5F884A9AFED2D" box="[1084,1160,1850,1877]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="91" pageNumber="92" unit="km" value="30.0">30 km</quantity>
above Jucusbamba
</locationDeviation>
; 
<geoCoordinate id="EE2FBC9AFFFEFFA2FF66F8E1AC35ED00" box="[191,274,1886,1914]" degrees="7" direction="south" minutes="55" orientation="latitude" pageId="91" pageNumber="92" precision="925" value="-7.9166665">7°55′S</geoCoordinate>
, 
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; 
<elevation id="00363D6EFFFEFFA2FE4FF8E0AF0DED00" box="[406,554,1887,1914]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="91" pageNumber="92" unit="m" value="2600.0">
<quantity id="4CE377B8FFFEFFA2FE4FF8E0ACCAED03" box="[406,493,1887,1914]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="91" pageNumber="92" unit="m" value="2600.0">2600 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFFFEFFA2FDEFF8E0AFF4ED03" box="[566,723,1887,1914]" pageId="91" pageNumber="92" value="1965-08-25">
<collectingDate id="EFE10575FFFEFFA2FDEFF8E0AFF4ED03" box="[566,723,1887,1914]" pageId="91" pageNumber="92" value="1965-08-25">25Aug. 1965</collectingDate>
</date>
; 
<emphasis id="B96F064FFFFEFFA2FD04F8DFA9A9ED00" box="[733,1166,1887,1914]" italics="true" pageId="91" pageNumber="92">
<collectorName id="26EEBF8BFFFEFFA2FD04F8DFAEA0ED00" box="[733,903,1887,1914]" pageId="91" pageNumber="92">A.C. Hamilton</collectorName>
&amp; 
<collectorName id="26EEBF8BFFFEFFA2FC70F8DFA96AED00" box="[937,1101,1887,1913]" pageId="91" pageNumber="92">P.M. Holligan</collectorName>
1421
</emphasis>
; 
<collectionCode id="ED0A4298FFFEFFA2FB42F8DFA994ED03" box="[1179,1203,1888,1914]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="91" pageNumber="92" type="Herbarium">K</collectionCode>
, 
<collectionCode id="ED0A4298FFFEFFA2FB64F8DFA9CFED03" box="[1213,1256,1888,1914]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="91" pageNumber="92" type="Herbarium">NY</collectionCode>
</materialsCitation>
. – 
<materialsCitation id="3B73D000FFFEFFA2FAD8F8E1AEE9EDBA" collectingDate="2016-02-09" collectionCode="E, G, MO, MOL" collectorName="P. W. Moonlight &amp; A. Daza" country="Peru" county="Prov. Maranon" elevation="1464" latitude="-8.608056" location="route west of Uchiza" longLatPrecision="21" longitude="-76.67833" pageId="91" pageNumber="92" specimenCode="E00885461" specimenCount="1" stateProvince="Huanuco Region">
<emphasis id="B96F064FFFFEFFA2FAD8F8E1ACC0EDE7" bold="true" pageId="91" pageNumber="92">
<collectingRegion id="49DF14BFFFFEFFA2FAD8F8E1AC3DEDE7" country="Peru" name="Huanuco" pageId="91" pageNumber="92">Huánuco Region</collectingRegion>
: 
<collectingCounty id="62C5A2D1FFFEFFA2FEFEF83BACC0EDE7" box="[295,487,1923,1950]" pageId="91" pageNumber="92">Prov. Marañón</collectingCounty>
</emphasis>
• 
<location id="8EC48C86FFFEFFA2FDD8F83AAFDEEDE7" LSID="urn:lsid:plazi:treatment:03B26B4BFFFEFFA7FD8BFAE8A850EB10:8EC48C86FFFEFFA2FDD8F83AAFDEEDE7" box="[513,761,1924,1951]" country="Peru" county="Prov. Maranon" latitude="-8.608056" longLatPrecision="21" longitude="-76.67833" name="route west of Uchiza" pageId="91" pageNumber="92" stateProvince="Huanuco Region">route west of Uchiza</location>
; 
<geoCoordinate id="EE2FBC9AFFFEFFA2FCDCF83CAEA1EDE7" box="[773,902,1923,1951]" degrees="8" direction="south" minutes="36" orientation="latitude" pageId="91" pageNumber="92" precision="15" seconds="29" value="-8.608056">8°36′29″ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFFFEFFA2FC4DF83CA908EDE6" box="[916,1071,1923,1951]" degrees="76" direction="west" minutes="40" orientation="longitude" pageId="91" pageNumber="92" precision="15" seconds="42" value="-76.67833">76°40′42″ W</geoCoordinate>
; 
<elevation id="00363D6EFFFEFFA2FBE2F83BA9F2EDE7" box="[1083,1237,1924,1951]" metricMagnitude="3" metricUnit="m" metricValue="1.464" pageId="91" pageNumber="92" unit="m" value="1464.0">
<quantity id="4CE377B8FFFEFFA2FBE2F83BA9B2EDE6" box="[1083,1173,1924,1951]" metricMagnitude="3" metricUnit="m" metricValue="1.464" pageId="91" pageNumber="92" unit="m" value="1464.0">1464 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFFFEFFA2FB3BF83BA855EDE7" box="[1250,1394,1924,1950]" pageId="91" pageNumber="92" value="2016-02-09">
<collectingDate id="EFE10575FFFEFFA2FB3BF83BA855EDE7" box="[1250,1394,1924,1950]" pageId="91" pageNumber="92" value="2016-02-09">9 Feb. 2016</collectingDate>
</date>
; 
<emphasis id="B96F064FFFFEFFA2FF64F815AF65EDBA" box="[189,578,1961,1988]" italics="true" pageId="91" pageNumber="92">
<collectorName id="26EEBF8BFFFEFFA2FF64F815AC50EDBA" box="[189,375,1961,1987]" pageId="91" pageNumber="92">P.W. Moonlight</collectorName>
&amp; 
<collectorName id="26EEBF8BFFFEFFA2FE7FF815AF2DEDBA" box="[422,522,1962,1988]" pageId="91" pageNumber="92">A. Daza</collectorName>
208
</emphasis>
; 
<collectionCode id="ED0A4298FFFEFFA2FD8EF815AF4EEDBD" box="[599,617,1962,1988]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="91" pageNumber="92" type="Herbarium">E</collectionCode>
[
<specimenCode id="DBBD7226FFFEFFA2FDA6F816AE21EDBD" box="[639,774,1961,1988]" collectionCode="E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="91" pageNumber="92" type="Herbarium">E00885461</specimenCode>
], 
<collectionCode id="ED0A4298FFFEFFA2FCFAF816AE1CEDBA" box="[803,827,1961,1987]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="91" pageNumber="92" type="Herbarium">G</collectionCode>
, 
<collectionCode id="ED0A4298FFFEFFA2FC92F816AE5AEDBA" box="[843,893,1961,1987]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="91" pageNumber="92" type="Herbarium">MO</collectionCode>
, 
<collectionCode id="ED0A4298FFFEFFA2FC57F816AEE9EDBA" box="[910,974,1961,1987]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="91" pageNumber="92" type="Herbarium">MOL</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B73D000FFFEFFA2FC29F816A9B6ED90" collectingDate="2016-02-09" collectionCode="E, MOL, P" collectorName="P. W. Moonlight &amp; A. Daza" country="Peru" county="Prov. Maranon" elevation="2086" latitude="-8.645278" location="Uchiza" longLatPrecision="21" longitude="-76.81528" pageId="91" pageNumber="92" specimenCode="E00885463" specimenCount="1" stateProvince="Huanuco">
ibid.; 
<geoCoordinate id="EE2FBC9AFFFEFFA2FBE0F817A998EDBA" box="[1081,1215,1960,1988]" degrees="8" direction="south" minutes="38" orientation="latitude" pageId="91" pageNumber="92" precision="15" seconds="43" value="-8.645278">8°38′43″ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFFFEFFA2FB09F817A855EDBD" box="[1232,1394,1960,1988]" degrees="76" direction="west" minutes="48" orientation="longitude" pageId="91" pageNumber="92" precision="15" seconds="55" value="-76.81528">76°48′55″ W</geoCoordinate>
; 
<elevation id="00363D6EFFFEFFA2FF64F871AC73ED91" box="[189,340,1998,2025]" metricMagnitude="3" metricUnit="m" metricValue="2.086" pageId="91" pageNumber="92" unit="m" value="2086.0">
<quantity id="4CE377B8FFFEFFA2FF64F871AC32ED90" box="[189,277,1998,2025]" metricMagnitude="3" metricUnit="m" metricValue="2.086" pageId="91" pageNumber="92" unit="m" value="2086.0">2086 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFFFEFFA2FEB9F871ACCBED91" box="[352,492,1998,2024]" pageId="91" pageNumber="92" value="2016-02-09">
<collectingDate id="EFE10575FFFEFFA2FEB9F871ACCBED91" box="[352,492,1998,2024]" pageId="91" pageNumber="92" value="2016-02-09">9 Feb. 2016</collectingDate>
</date>
; 
<emphasis id="B96F064FFFFEFFA2FE2EF870AE09ED91" box="[503,814,1998,2025]" italics="true" pageId="91" pageNumber="92">
<collectorName id="26EEBF8BFFFEFFA2FE2EF870AF8BED91" box="[503,684,1998,2024]" pageId="91" pageNumber="92">P.W. Moonlight</collectorName>
&amp; 
<collectorName id="26EEBF8BFFFEFFA2FD09F870AE09ED91" box="[720,814,1999,2025]" pageId="91" pageNumber="92">A. Daza</collectorName>
</emphasis>
212 (
<collectionCode id="ED0A4298FFFEFFA2FCA8F871AEA3ED91" box="[881,900,1998,2024]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="91" pageNumber="92" type="Herbarium">E</collectionCode>
[
<specimenCode id="DBBD7226FFFEFFA2FC4DF871A93BED90" box="[916,1052,1998,2025]" collectionCode="E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="91" pageNumber="92" type="Herbarium">E00885463</specimenCode>
], 
<collectionCode id="ED0A4298FFFEFFA2FBEAF871A950ED91" box="[1075,1143,1998,2024]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="91" pageNumber="92" type="Herbarium">MOL</collectionCode>
, 
<collectionCode id="ED0A4298FFFEFFA2FB5BF870A9B6ED90" box="[1154,1169,1999,2025]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="91" pageNumber="92" type="Herbarium">P</collectionCode>
</materialsCitation>
<materialsCitation id="3B73D000FFFEFFA4FB4EF870A9A9EB3E" collectingDate="2021-07-21" collectionCode="USM" collectorName="P. Gonzalez &amp; P. Arista" country="Peru" county="Cholon" elevation="3200" lastPageId="93" lastPageNumber="94" latitude="-8.645278" location="San Pedro de Chonta" longLatPrecision="21" longitude="-76.81528" pageId="91" pageNumber="92" specimenCode="E00885463" specimenCount="1" stateProvince="Huanuco">
• Dist. 
<collectingCounty id="62C5A2D1FFFEFFA2FB3FF871A818ED91" box="[1254,1343,1998,2024]" pageId="91" pageNumber="92">Cholón</collectingCounty>
, 
<location id="8EC48C86FFFEFFA4FA93F871ACAFEB5A" LSID="urn:lsid:plazi:treatment:03B26B4BFFFEFFA7FD8BFAE8A850EB10:8EC48C86FFFEFFA4FA93F871ACAFEB5A" country="Peru" county="Cholon" lastPageId="93" lastPageNumber="94" latitude="-8.645278" longLatPrecision="21" longitude="-76.81528" name="San Pedro de Chonta" pageId="91" pageNumber="92" stateProvince="Huanuco">San Pedro de Chonta</location>
, 
<locationDeviation id="9A16C37FFFF8FFA4FE4CFEB6A99BEB5A" box="[405,1212,265,291]" pageId="93" pageNumber="94">retorno desde el campamento hacia el pueblo San Pedro de Chonta</locationDeviation>
; 
<geoCoordinate id="EE2FBC9AFFF8FFA4FB10FEB8A856EB5A" box="[1225,1393,263,291]" degrees="8" direction="south" minutes="38" orientation="latitude" pageId="93" pageNumber="94" precision="1" seconds="13.42" value="-8.637061">8°38′13.42″ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFFF8FFA4FF64FE94AC5AEB3E" box="[189,381,299,327]" degrees="76" direction="west" minutes="53" orientation="longitude" pageId="93" pageNumber="94" precision="1" seconds="08.57" value="-76.88571">76°53′08.57″ W</geoCoordinate>
; 
<quantity id="4CE377B8FFF8FFA4FE50FE92ACC5EB3E" box="[393,482,301,327]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="93" pageNumber="94" unit="m" value="3200.0">
<elevation id="00363D6EFFF8FFA4FE50FE92ACC5EB3E" box="[393,482,301,327]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="93" pageNumber="94" unit="m" value="3200.0">3200 m</elevation>
</quantity>
a.s.l.; 
<date id="FFA5FC9DFFF8FFA4FDF6FE92AFE4EB3E" box="[559,707,300,327]" pageId="93" pageNumber="94" value="2021-07-21">
<collectingDate id="EFE10575FFF8FFA4FDF6FE92AFE4EB3E" box="[559,707,300,327]" pageId="93" pageNumber="94" value="2021-07-21">21 Jul. 2021</collectingDate>
</date>
; 
<emphasis id="B96F064FFFF8FFA4FD16FE92A910EB3F" box="[719,1079,300,327]" italics="true" pageId="93" pageNumber="94">
<collectorName id="26EEBF8BFFF8FFA4FD16FE92AE7DEB3F" box="[719,858,300,326]" pageId="93" pageNumber="94">P. González</collectorName>
&amp; 
<collectorName id="26EEBF8BFFF8FFA4FC59FE92AEC2EB3E" box="[896,997,301,327]" pageId="93" pageNumber="94">P. Arista</collectorName>
10016
</emphasis>
; 
<collectionCode id="ED0A4298FFF8FFA4FB9EFE92A9ADEB3E" box="[1095,1162,301,327]" pageId="93" pageNumber="94">USM</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF648AD5FFF9FFA5FF64F944AF25ED91" ID-DOI="http://doi.org/10.5281/zenodo.8178365" ID-Zenodo-Dep="8178365" httpUri="https://zenodo.org/record/8178365/files/figure.png" pageId="92" pageNumber="93" startId="92.[189,232,1787,1813]" targetBox="[272,1315,265,1740]" targetPageId="92">
<paragraph id="8BA4DA5DFFF9FFA5FF64F944AF25ED91" blockId="92.[189,1401,1787,2025]" pageId="92" pageNumber="93">
<emphasis id="B96F064FFFF9FFA5FF64F944AC39ED6C" bold="true" box="[189,286,1787,1813]" pageId="92" pageNumber="93">Fig. 33.</emphasis>
<taxonomicName id="4C1BA1DEFFF9FFA5FEF0F943AE51ED6C" authority="Irmsch. A. Habit. B. Leaf" box="[297,886,1787,1814]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="92" pageNumber="93" phylum="Tracheophyta" rank="species" species="obtecticaulis">
<emphasis id="B96F064FFFF9FFA5FEF0F943AF09ED6C" box="[297,558,1787,1813]" italics="true" pageId="92" pageNumber="93">Begonia obtecticaulis</emphasis>
Irmsch. 
<emphasis id="B96F064FFFF9FFA5FD44F944AF95ED6C" bold="true" box="[669,690,1787,1813]" pageId="92" pageNumber="93">A</emphasis>
. Habit. 
<emphasis id="B96F064FFFF9FFA5FCCEF944AE0CED6C" bold="true" box="[791,811,1787,1813]" pageId="92" pageNumber="93">B</emphasis>
. Leaf
</taxonomicName>
, abaxial surface. 
<emphasis id="B96F064FFFF9FFA5FB95F944A946ED6C" bold="true" box="[1100,1121,1787,1813]" pageId="92" pageNumber="93">C</emphasis>
. Leaf, adaxial surface. 
<emphasis id="B96F064FFFF9FFA5FF64F8A1ADF5ED41" bold="true" box="[189,210,1822,1848]" pageId="92" pageNumber="93">D</emphasis>
. Stipule. 
<emphasis id="B96F064FFFF9FFA5FE93F8A1AC7CED41" bold="true" box="[330,347,1822,1848]" pageId="92" pageNumber="93">E</emphasis>
. Inflorescence. 
<emphasis id="B96F064FFFF9FFA5FDC3F8A1AF0BED41" bold="true" box="[538,556,1822,1848]" pageId="92" pageNumber="93">F</emphasis>
. Bract. 
<emphasis id="B96F064FFFF9FFA5FD49F8A1AF82ED41" bold="true" box="[656,677,1822,1848]" pageId="92" pageNumber="93">G</emphasis>
. Pistillate flower, side view. 
<emphasis id="B96F064FFFF9FFA5FBDBF8A1A93EED41" bold="true" box="[1026,1049,1822,1848]" pageId="92" pageNumber="93">H</emphasis>
. Pistillate flower, front view. 
<emphasis id="B96F064FFFF9FFA5FF64F8FDADEFED25" bold="true" box="[189,200,1858,1884]" pageId="92" pageNumber="93">I</emphasis>
. Cross section of ovary. 
<emphasis id="B96F064FFFF9FFA5FE24F8FDAF2BED25" bold="true" box="[509,524,1858,1884]" pageId="92" pageNumber="93">J</emphasis>
. Largest tepal of the pistillate flower. 
<emphasis id="B96F064FFFF9FFA5FC3BF8FDAEDEED25" bold="true" box="[994,1017,1858,1884]" pageId="92" pageNumber="93">K</emphasis>
. Smallest tepal of the pistillate flower. 
<emphasis id="B96F064FFFF9FFA5FECCF8DAAC0EED06" bold="true" box="[277,297,1893,1919]" pageId="92" pageNumber="93">L</emphasis>
. Pistils, side view. 
<emphasis id="B96F064FFFF9FFA5FDD0F8DAAF02ED06" bold="true" box="[521,549,1893,1919]" pageId="92" pageNumber="93">M</emphasis>
. Pistils, front view. 
<emphasis id="B96F064FFFF9FFA5FCD6F8DAAE03ED06" bold="true" box="[783,804,1893,1919]" pageId="92" pageNumber="93">N</emphasis>
. Staminate flower, side view. 
<emphasis id="B96F064FFFF9FFA5FB5AF8DBA9BDED07" bold="true" box="[1155,1178,1892,1918]" pageId="92" pageNumber="93">O</emphasis>
. Staminate flower, front view. 
<emphasis id="B96F064FFFF9FFA5FE9AF837AC72EDDB" bold="true" box="[323,341,1928,1954]" pageId="92" pageNumber="93">P</emphasis>
. Androecium, side view. All photographs taken by D.A. Purvis in the living collections of the Royal Botanic Garden Edinburgh (Accession 20160137a, grown from seeds collected as part of 
<emphasis id="B96F064FFFF9FFA5FA9BF813ACD6ED91" italics="true" pageId="92" pageNumber="93">P.W. Moonlight &amp; A. Daza 209</emphasis>
).
</paragraph>
</caption>
<subSubSection id="C30189D6FFF8FFA4FF64FED1AF6BEEBD" pageId="93" pageNumber="94" type="description">
<paragraph id="8BA4DA5DFFF8FFA4FF64FED1AC68EBF1" blockId="93.[189,335,366,392]" box="[189,335,366,392]" pageId="93" pageNumber="94">
<heading id="D0EC6D31FFF8FFA4FF64FED1AC68EBF1" bold="true" box="[189,335,366,392]" fontSize="11" level="3" pageId="93" pageNumber="94" reason="3">
<emphasis id="B96F064FFFF8FFA4FF64FED1AC68EBF1" bold="true" box="[189,335,366,392]" pageId="93" pageNumber="94">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFF8FFA4FF64FE22AF6BEEBD" blockId="93.[187,1401,413,1220]" pageId="93" pageNumber="94">
Caulescent herb, to 
<quantity id="4CE377B8FFF8FFA4FE6DFE22ACDBEBCE" box="[436,508,413,440]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="93" pageNumber="94" unit="m" value="1.5">1.5 m</quantity>
high. 
<emphasis id="B96F064FFFF8FFA4FD89FE22AFAEEBCE" box="[592,649,413,439]" italics="true" pageId="93" pageNumber="94">Stem</emphasis>
erect, branching; internodes to 
<quantity id="4CE377B8FFF8FFA4FBCCFE22A94EEBCE" box="[1045,1129,413,440]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="93" pageNumber="94" unit="cm" value="9.5">9.5 cm</quantity>
long, to 
<quantity id="4CE377B8FFF8FFA4FB39FE22A80EEBCE" box="[1248,1321,413,439]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="93" pageNumber="94" unit="mm" value="8.0">8 mm</quantity>
thick, succulent, pale green to dark red, glabrous. 
<emphasis id="B96F064FFFF8FFA4FD1EFE7FAE02EBA3" box="[711,805,448,474]" italics="true" pageId="93" pageNumber="94">Stipules</emphasis>
in unequal pairs, persistent, lanceolate to broadly ovate, clasping the stem, 9–35 × 
<quantity id="4CE377B8FFF8FFA4FD9DFE5BAF9FEB87" box="[580,696,484,510]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.8" metricValueMin="0.4" pageId="93" pageNumber="94" unit="mm" value="11.0" valueMax="18.0" valueMin="4.0">4–18 mm</quantity>
, apex acute, base cuneate, translucent, pale green to brown, glabrous, margin entire, aciliate. 
<emphasis id="B96F064FFFF8FFA4FD93FDB7AFBBE858" box="[586,668,520,545]" italics="true" pageId="93" pageNumber="94">Leaves</emphasis>
&gt; 5, alternate, basifixed; petiole 
<quantity id="4CE377B8FFF8FFA4FBFEFDB8A98AE85B" box="[1063,1197,519,546]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="4.5" metricValueMin="0.8" pageId="93" pageNumber="94" unit="cm" value="2.65" valueMax="4.5" valueMin="0.8">0.8–4.5 cm</quantity>
long, pale green to red, glabrous; blade asymmetric, lanceolate to ovate, to 9 × 
<quantity id="4CE377B8FFF8FFA4FC47FD94AED5E83C" box="[926,1010,555,582]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="93" pageNumber="94" unit="cm" value="3.5">3.5 cm</quantity>
, succulent, apex acuminate, base transversely cordate, basal lobes rounded, sinus to 
<quantity id="4CE377B8FFF8FFA4FCDFFDF1AE6EE810" box="[774,841,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="93" pageNumber="94" unit="mm" value="6.0">6 mm</quantity>
deep, margin irregularly double-dentate, ciliate, upper surface bright green, glabrous, lower surface pale grey-green, glabrous, veins palmate-pinnate, 4–6 veined from the base, with 2–4 secondary veins on the larger side, 1–3 secondary veins on the smaller side. 
<emphasis id="B96F064FFFF8FFA4FE81FD06AF27E8AA" box="[344,512,697,723]" italics="true" pageId="93" pageNumber="94">Inflorescences</emphasis>
1–3 per stem, bisexual, axillary, erect, cymose, with 2 branches, bearing up to 4 staminate flowers and 8 pistillate flowers, protandrous; peduncle to 
<quantity id="4CE377B8FFF8FFA4FBDFFD63A967E88E" box="[1030,1088,732,759]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="93" pageNumber="94" unit="cm" value="8.0">8 cm</quantity>
long, red, glabrous, bracts persistent, lanceolate, 7–10 × 
<quantity id="4CE377B8FFF8FFA4FDFAFCBFAFAEE963" box="[547,649,768,794]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="93" pageNumber="94" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
, translucent, white, glabrous, apex acuminate, margin lacerate, ciliate. 
<emphasis id="B96F064FFFF8FFA4FECDFC9BACC0E944" box="[276,487,803,830]" italics="true" pageId="93" pageNumber="94">Staminate flowers</emphasis>
: pedicels to 
<quantity id="4CE377B8FFF8FFA4FDA6FC9CAFF5E947" box="[639,722,803,830]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="93" pageNumber="94" unit="mm" value="12.0">12 mm</quantity>
long, glabrous; tepals 2, spreading, broadly ovate, 12–14 × 
<quantity id="4CE377B8FFF8FFA4FF0EFCF9AC7DE918" box="[215,346,838,865]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="93" pageNumber="94" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 mm</quantity>
, apex obtuse, white to pink, glabrous, margin entire, aciliate or rarely ciliate at the base; stamens 25–30, spreading, yellow, filaments 
<quantity id="4CE377B8FFF8FFA4FD08FCD5AE6FE9FC" box="[721,840,874,901]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="93" pageNumber="94" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 mm</quantity>
long, free, anthers ellipsoid, 1.5–2 × 
<quantity id="4CE377B8FFF8FFA4FADFFCD5A855E9FC" box="[1286,1394,874,901]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="93" pageNumber="94" unit="mm" value="0.75">0.75 mm</quantity>
, dehiscing via lateral slits, connectives extending to 
<quantity id="4CE377B8FFF8FFA4FCCEFC31AE53E9D1" box="[791,884,910,936]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="93" pageNumber="94" unit="mm" value="0.4">0.4 mm</quantity>
, symmetrically basifixed. 
<emphasis id="B96F064FFFF8FFA4FB7FFC31A849E9D1" box="[1190,1390,910,936]" italics="true" pageId="93" pageNumber="94">Pistillate flowers</emphasis>
: pedicels to 
<quantity id="4CE377B8FFF8FFA4FE9AFC0EACA0E9B5" box="[323,391,945,972]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="93" pageNumber="94" unit="mm" value="8.0">8 mm</quantity>
long; bracteoles 2, lanceolate to ovate, ca 1 × 
<quantity id="4CE377B8FFF8FFA4FC70FC0EA921E9B5" box="[937,1030,945,972]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="93" pageNumber="94" unit="mm" value="0.5">0.5 mm</quantity>
, translucent, white tinged pink; tepals 5, subequal, deciduous in fruit, spreading, elliptic to ovate, 7–15 × 
<quantity id="4CE377B8FFF8FFA4FBDFFC6BA94EE996" box="[1030,1129,980,1007]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="93" pageNumber="94" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
, apex rounded to acute, white, glabrous, margin entire, aciliate; ovary body ovoid, ca 7 × 
<quantity id="4CE377B8FFF8FFA4FC07FC47A90EEE6A" box="[990,1065,1016,1043]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="93" pageNumber="94" unit="mm" value="6.0">6 mm</quantity>
, white, glabrous, unequally 3-winged, wings triangular, largest ca 10 × 
<quantity id="4CE377B8FFF8FFA4FD6BFBA3AE20EE4F" box="[690,775,1052,1078]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="93" pageNumber="94" unit="mm" value="10.0">10 mm</quantity>
, smallest ca 10 × 
<quantity id="4CE377B8FFF8FFA4FC0BFBA3A93FEE4F" box="[978,1048,1052,1078]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="93" pageNumber="94" unit="mm" value="3.0">3 mm</quantity>
; 3-locular, placentae branches divided, bearing ovules on both surfaces; styles 3, yellow, free, ca 
<quantity id="4CE377B8FFF8FFA4FC07FB80A902EE23" box="[990,1061,1087,1114]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="93" pageNumber="94" unit="mm" value="5.0">5 mm</quantity>
long, irregularly 2–5 times-divided, stigmatic papillae in a spirally twisted band. 
<emphasis id="B96F064FFFF8FFA4FCF3FBDCAECDEE05" box="[810,1002,1122,1148]" italics="true" pageId="93" pageNumber="94">Fruiting pedicel</emphasis>
to 
<quantity id="4CE377B8FFF8FFA4FBD7FBDCA947EE04" box="[1038,1120,1123,1149]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="93" pageNumber="94" unit="mm" value="18.0">18 mm</quantity>
long. 
<emphasis id="B96F064FFFF8FFA4FB71FBDCA803EE05" box="[1192,1316,1122,1148]" italics="true" pageId="93" pageNumber="94">Fruit body</emphasis>
ovoid, to 9 × 
<quantity id="4CE377B8FFF8FFA4FECBFB39AC7BEED9" box="[274,348,1158,1184]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="93" pageNumber="94" unit="mm" value="6.0">6 mm</quantity>
, drying brown, wings same shape as in ovary, the largest expanding 12 × 
<quantity id="4CE377B8FFF8FFA4FB33FB39A864EED9" box="[1258,1347,1158,1184]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="93" pageNumber="94" unit="mm" value="18.0">18 mm</quantity>
, the smallest expanding to 12 × 
<quantity id="4CE377B8FFF8FFA4FDD8FB16AF6FEEBD" box="[513,584,1193,1220]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="93" pageNumber="94" unit="mm" value="6.0">6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFF8FFA4FF64FB54A902EFBB" pageId="93" pageNumber="94" type="conservation">
<paragraph id="8BA4DA5DFFF8FFA4FF64FB54AF4AEF7C" blockId="93.[189,621,1259,1285]" box="[189,621,1259,1285]" pageId="93" pageNumber="94">
<heading id="D0EC6D31FFF8FFA4FF64FB54AF4AEF7C" bold="true" box="[189,621,1259,1285]" fontSize="11" level="3" pageId="93" pageNumber="94" reason="3">
<emphasis id="B96F064FFFF8FFA4FF64FB54AF4AEF7C" bold="true" box="[189,621,1259,1285]" pageId="93" pageNumber="94">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFF8FFA4FF64FAA5A902EFBB" blockId="93.[189,1399,1306,1474]" pageId="93" pageNumber="94">
Previously assessed by 
<bibRefCitation id="EF8AA7ACFFF8FFA4FE01FAA5AE2DEF4D" author="Leon &amp; Monsalve" box="[472,778,1306,1332]" firstAuthor="Leon" pageId="93" pageNumber="94" pagination="165 - 170" refId="ref229226" refString="Leon B. &amp; Monsalve C. 2006. Begoniaceae endemicas del Peru. Revista Peruana de Biologia 13 (2): 165 - 170. https: // doi. org / 10.15381 / rpb. v 13 i 2.1810" type="journal article" year="2006">
<collectingRegion id="49DF14BFFFF8FFA4FE01FAA5AF33EF4D" box="[472,532,1306,1332]" country="Nicaragua" name="Leon" pageId="93" pageNumber="94">León</collectingRegion>
&amp; Monsalve (2006)
</bibRefCitation>
as Endangered (EN); however, their concept of the species included specimens of 
<taxonomicName id="4C1BA1DEFFF8FFA4FDECFA81AE93EF21" authority="C. DC. Known" authorityName="C. DC. Known" box="[565,948,1341,1368]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="93" pageNumber="94" phylum="Tracheophyta" rank="species" species="albomaculata">
<emphasis id="B96F064FFFF8FFA4FDECFA81AFDAEF2E" box="[565,765,1341,1367]" italics="true" pageId="93" pageNumber="94">B. albomaculata</emphasis>
C.DC. Known
</taxonomicName>
from three populations, but the 
<typeStatus id="54A064FFFFF8FFA4FA9DFA81A850EF21" box="[1348,1399,1342,1368]" pageId="93" pageNumber="94">type</typeStatus>
locality has undergone severe deforestation in the last century and it was last collected there in 1923. Given its EOO of &lt;
<quantity id="4CE377B8FFF8FFA4FE69FA3BAF31EFE7" box="[432,534,1412,1439]" metricMagnitude="6" metricUnit="m" metricValue="5.0" pageId="93" pageNumber="94" unit="km" value="5000.0">5000 km</quantity>
<superScript id="7C6E7715FFF8FFA4FDCFFA3BAF38EFEB" attach="left" box="[534,543,1412,1426]" fontSize="6" pageId="93" pageNumber="94">2</superScript>
, that is known from three populations, and its probable extirpation from the 
<typeStatus id="54A064FFFFF8FFA4FF30FA17AC3BEFBB" box="[233,284,1448,1474]" pageId="93" pageNumber="94">type</typeStatus>
locality, we assess 
<taxonomicName id="4C1BA1DEFFF8FFA4FE26FA16AF9FEFBB" box="[511,696,1448,1474]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="93" pageNumber="94" phylum="Tracheophyta" rank="species" species="obtecticaulis">
<emphasis id="B96F064FFFF8FFA4FE26FA16AF9FEFBB" box="[511,696,1448,1474]" italics="true" pageId="93" pageNumber="94">B. obtecticaulis</emphasis>
</taxonomicName>
as Endangered (EN B1ab(iv)).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFF8FFA4FF64FA55AFFFED2D" pageId="93" pageNumber="94" type="discussion">
<paragraph id="8BA4DA5DFFF8FFA4FF64FA55AC23EC7D" blockId="93.[189,260,1514,1540]" box="[189,260,1514,1540]" pageId="93" pageNumber="94">
<heading id="D0EC6D31FFF8FFA4FF64FA55AC23EC7D" bold="true" box="[189,260,1514,1540]" fontSize="11" level="3" pageId="93" pageNumber="94" reason="3">
<emphasis id="B96F064FFFF8FFA4FF64FA55AC23EC7D" bold="true" box="[189,260,1514,1540]" pageId="93" pageNumber="94">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFF8FFA4FF64F9A7AC67EC2F" blockId="93.[189,1398,1560,1622]" pageId="93" pageNumber="94">
Description of the pistillate flowers made using living accession 20160137a at the Royal Botanic Garden 
<collectingRegion id="49DF14BFFFF8FFA4FF64F983AC1BEC2F" box="[189,316,1596,1622]" country="United Kingdom" name="City of Edinburgh" pageId="93" pageNumber="94">Edinburgh</collectingRegion>
.
</paragraph>
<paragraph id="8BA4DA5DFFF8FFA4FF64F9C2AC97ECE1" blockId="93.[189,432,1661,1688]" box="[189,432,1661,1688]" pageId="93" pageNumber="94">
<heading id="D0EC6D31FFF8FFA4FF64F9C2AC97ECE1" bold="true" box="[189,432,1661,1688]" fontSize="11" level="3" pageId="93" pageNumber="94" reason="3">
<emphasis id="B96F064FFFF8FFA4FF64F9C2AC97ECE1" bold="true" box="[189,432,1661,1688]" pageId="93" pageNumber="94">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFF8FFA4FF64F912AFFFED2D" blockId="93.[187,1400,1708,1877]" pageId="93" pageNumber="94">
<taxonomicName id="4C1BA1DEFFF8FFA4FF64F912ACE6ECBF" box="[189,449,1708,1734]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="93" pageNumber="94" phylum="Tracheophyta" rank="species" species="obtecticaulis">
<emphasis id="B96F064FFFF8FFA4FF64F912ACE6ECBF" box="[189,449,1708,1734]" italics="true" pageId="93" pageNumber="94">Begonia obtecticaulis</emphasis>
</taxonomicName>
is one of three Peruvian caulescent begonias with large, persistent stipules that clasp the internodes and are longer than the internodes. This character is shared with 
<taxonomicName id="4C1BA1DEFFF8FFA4FB6EF96EA85FEC93" box="[1207,1400,1744,1770]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="93" pageNumber="94" phylum="Tracheophyta" rank="species" species="inbrexiformis">
<emphasis id="B96F064FFFF8FFA4FB6EF96EA85FEC93" box="[1207,1400,1744,1770]" italics="true" pageId="93" pageNumber="94">B. inbrexiformis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1BA1DEFFF8FFA4FF37F94BAC89ED74" authority="Moonlight &amp; Jara-Muñoz &amp; Purvis &amp; Delves &amp; Allen &amp; Reynel, 2023" authorityName="Moonlight &amp; Jara-Muñoz &amp; Purvis &amp; Delves &amp; Allen &amp; Reynel" authorityYear="2023" box="[238,430,1779,1805]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="93" pageNumber="94" phylum="Tracheophyta" rank="species" species="serratistipula" status="sp. nov.">
<emphasis id="B96F064FFFF8FFA4FF37F94BAC89ED74" box="[238,430,1779,1805]" italics="true" pageId="93" pageNumber="94">B. serratistipula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FFF8FFA4FE6DF94BAF28ED77" box="[436,527,1780,1806]" pageId="93" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName id="4C1BA1DEFFF8FFA4FDCDF94BAE31ED74" box="[532,790,1779,1805]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="93" pageNumber="94" phylum="Tracheophyta" rank="species" species="obtecticaulis">
<emphasis id="B96F064FFFF8FFA4FDCDF94BAE31ED74" box="[532,790,1779,1805]" italics="true" pageId="93" pageNumber="94">Begonia obtecticaulis</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="4C1BA1DEFFF8FFA4FB89F94BA832ED74" authority="Moonlight &amp; Jara-Muñoz &amp; Purvis &amp; Delves &amp; Allen &amp; Reynel, 2023" authorityName="Moonlight &amp; Jara-Muñoz &amp; Purvis &amp; Delves &amp; Allen &amp; Reynel" authorityYear="2023" box="[1104,1301,1779,1805]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="93" pageNumber="94" phylum="Tracheophyta" rank="species" species="imbrexiformis" status="sp. nov.">
<emphasis id="B96F064FFFF8FFA4FB89F94BA832ED74" box="[1104,1301,1779,1805]" italics="true" pageId="93" pageNumber="94">B. imbrexiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FFF8FFA4FAC4F94BA850ED77" box="[1309,1399,1780,1806]" pageId="93" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
by its palmate-pinnate (vs pinnate) leaf venation and from 
<taxonomicName id="4C1BA1DEFFF8FFA4FC87F8A7A93AED48" authority="Moonlight &amp; Jara-Muñoz &amp; Purvis &amp; Delves &amp; Allen &amp; Reynel, 2023" authorityName="Moonlight &amp; Jara-Muñoz &amp; Purvis &amp; Delves &amp; Allen &amp; Reynel" authorityYear="2023" box="[862,1053,1815,1841]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="93" pageNumber="94" phylum="Tracheophyta" rank="species" species="serratistipula" status="sp. nov.">
<emphasis id="B96F064FFFF8FFA4FC87F8A7A93AED48" box="[862,1053,1815,1841]" italics="true" pageId="93" pageNumber="94">B. serratistipula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FFF8FFA4FBFAF8A8A95BED48" box="[1059,1148,1815,1841]" pageId="93" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
by the entire margins to its stipules (vs lacerate in the bottom third).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFF8FFA7FF64F8C3A850EB10" lastPageId="94" lastPageNumber="95" pageId="93" pageNumber="94" type="distribution">
<paragraph id="8BA4DA5DFFF8FFA4FF64F8C3ACD2EDEF" blockId="93.[189,501,1916,1942]" box="[189,501,1916,1942]" pageId="93" pageNumber="94">
<heading id="D0EC6D31FFF8FFA4FF64F8C3ACD2EDEF" bold="true" box="[189,501,1916,1942]" fontSize="11" level="3" pageId="93" pageNumber="94" reason="3">
<emphasis id="B96F064FFFF8FFA4FF64F8C3ACD2EDEF" bold="true" box="[189,501,1916,1942]" pageId="93" pageNumber="94">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFF8FFA7FF64F814A850EB10" blockId="93.[188,1398,1963,2025]" lastBlockId="94.[189,1399,265,362]" lastPageId="94" lastPageNumber="95" pageId="93" pageNumber="94">
Endemic to 
<collectingCountry id="F30C9ACDFFF8FFA4FE91F814AC59EDBC" box="[328,382,1963,1989]" name="Peru" pageId="93" pageNumber="94">Peru</collectingCountry>
and known from 
<collectingRegion id="49DF14BFFFF8FFA4FD93F814AFEBEDBC" box="[586,716,1963,1989]" country="Peru" name="San Martin" pageId="93" pageNumber="94">San Martín</collectingRegion>
and 
<collectingRegion id="49DF14BFFFF8FFA4FCDBF814AE4BEDBC" box="[770,876,1963,1989]" country="Peru" name="Huanuco" pageId="93" pageNumber="94">Huánuco</collectingRegion>
Regions (
<figureCitation id="1320C6D8FFF8FFA4FC38F814A969EDBC" box="[993,1102,1963,1989]" captionStart="Fig" captionStartId="66.[189,232,1787,1813]" captionTargetBox="[194,1391,514,1732]" captionTargetId="figure-13@66.[189,1398,504,1740]" captionTargetPageId="66" captionText="Fig. 24. Distribution ofBegonia sect.Cyathocnemis (Klotzsch)A.DC. in Peru and surrounding countries. A. B. amoeboides Moonlight (red) andB. imbrexiformis Moonlight sp. nov. (blue).B. B. bracteosaA.DC. (red) and B. longinqua Moonlight (blue). C. B. cyathophora Poepp. &amp; Endl. (red), B. serratistipula Moonlight sp. nov. (blue) and C. vargasii Moonlight sp. nov. (green). D. B. lamolina Moonlight (red), B. lophoptera Rolfe (blue) and B. obtecticaulis Irmsch. (green). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178339" httpUri="https://zenodo.org/record/8178339/files/figure.png" pageId="93" pageNumber="94">Fig. 24D</figureCitation>
). Found in lower, middle, and upper montane forest at an elevation of 
<quantity id="4CE377B8FFF8FFA4FD15F871AE48ED90" box="[716,879,1998,2025]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="3.2" metricValueMin="1.4" pageId="93" pageNumber="94" unit="m" value="2300.0" valueMax="3200.0" valueMin="1400.0">1400–3200 m</quantity>
a.s.l. The species has most frequently been collected on the margins of humid forests, particularly in areas of transition to drier Andean shrublands. There are only a few natural areas of dry shrubland in the Andes of 
<collectingRegion id="49DF14BFFFFBFFA7FC3DFE93A94EEB3F" box="[996,1129,300,326]" country="Peru" name="San Martin" pageId="94" pageNumber="95">San Martín</collectingRegion>
and 
<collectingRegion id="49DF14BFFFFBFFA7FB7DFE93A829EB3F" box="[1188,1294,300,326]" country="Peru" name="Huanuco" pageId="94" pageNumber="95">Huánuco</collectingRegion>
Regions at the elevation favoured by 
<taxonomicName id="4C1BA1DEFFFBFFA7FDD0FEEFAFE6EB10" box="[521,705,335,361]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="94" pageNumber="95" phylum="Tracheophyta" rank="species" species="obtecticaulis">
<emphasis id="B96F064FFFFBFFA7FDD0FEEFAFE6EB10" box="[521,705,335,361]" italics="true" pageId="94" pageNumber="95">B. obtecticaulis</emphasis>
</taxonomicName>
but some, such as Ongón District, remain poorly collected.
</paragraph>
</subSubSection>
</treatment>
</document>