treatments-xml/data/03/B2/6B/03B26B4BFFCEFF94FDB2F9E2AF4AE96C.xml
2024-06-21 12:22:17 +02:00

506 lines
53 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="AC30E311C9E39DD7851AD6000443BE08" ID-DOI="https://doi.org/10.5852/ejt.2023.881.2175" ID-ISSN="2118-9773" ID-Zenodo-Dep="8178280" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690201937279" checkinUser="felipe" docAuthor="Moonlight, Peter. W., Jara-Muñoz, Orlando A., Purvis, David A., Delves, Jay, Allen, Josh P. &amp; Reynel, Carlos" docDate="2023" docId="03B26B4BFFCEFF94FDB2F9E2AF4AE96C" docLanguage="en" docName="ejt-881_1-334.imf" docOrigin="European Journal of Taxonomy 881" docSource="http://dx.doi.org/10.5852/ejt.2023.881.2175" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia lucifuga Irmsch." docType="treatment" docVersion="7" lastPageNumber="110" masterDocId="FF8B1333FFA5FFF9FFD9FFBFAD27EA79" masterDocTitle="The genus Begonia (Begoniaceae) in Peru" masterLastPageNumber="334" masterPageNumber="1" pageNumber="108" updateTime="1707123162844" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="BAEA5188AED90C8B96DB8A8D1D4EB8F9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4F0177C13ABE94D9FB29F2B5095EEDF6">
<mods:title id="F26BF71B7256559EC0D31ACCD6E3854C">The genus Begonia (Begoniaceae) in Peru</mods:title>
</mods:titleInfo>
<mods:name id="BCF21C1CF1A9BADD4618F1361F9C9F15" type="personal">
<mods:role id="6906730C7BB0754BE6618B59116CEC47">
<mods:roleTerm id="DCB4DC3BD662FB9A9A13BC3E2DE00D68">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="642415915E8A70E00871062416363B5D">Moonlight, Peter. W.</mods:namePart>
<mods:affiliation id="64B4E0EFABE7EBF1FC59246364906E27">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK. &amp; Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland.</mods:affiliation>
<mods:nameIdentifier id="9AA790E1A9DDB93A9093132B4278D423" type="email">moonligp@tcd.ie</mods:nameIdentifier>
</mods:name>
<mods:name id="E03BE4E39B94EEF34875069EB695CE21" type="personal">
<mods:role id="9A8E6B66B4972EE139F82AAAD7EFAE2B">
<mods:roleTerm id="3203A5A4A496D4E498151D193C8115A2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6F9A52407BFC09196100F58ADA2B7573">Jara-Muñoz, Orlando A.</mods:namePart>
<mods:affiliation id="97B98DA68FE2BBFBABC1E274F8E8252D">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá D. C., Colombia.</mods:affiliation>
<mods:nameIdentifier id="85DFA6FD800AA506E3768242A63E0D2A" type="email">oajaram@unal.edu.co</mods:nameIdentifier>
</mods:name>
<mods:name id="44452F756049DED0ED99DB512830A1BA" type="personal">
<mods:role id="3AE4F031C2EDAD2A5FED6D3C4D621CFF">
<mods:roleTerm id="0F06C5EE283F148221545C586C6B0934">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A2FF9A89623D57025EC92A64B7DCA784">Purvis, David A.</mods:namePart>
<mods:affiliation id="6188A1C8FF708E1509CE1E3D479C62E5">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="AEA1BA1A330A4655555E70C356A38597" type="email">dpurvis@rbge.org.uk</mods:nameIdentifier>
</mods:name>
<mods:name id="75243A439DE5D847327426B90EBF55F7" type="personal">
<mods:role id="3D3B9D814E9331A8F44BD74F37848CF9">
<mods:roleTerm id="8C17B5585F364E07AE4D50860A74402A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6D61161DF085482D31F79E1814BBD189">Delves, Jay</mods:namePart>
<mods:affiliation id="B3148EC066E931E9DF521116349CCEA6">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="ED156DEAB3AF730781A0D6696AF31AEE" type="email">j.delves@westernsydney.edu.au</mods:nameIdentifier>
</mods:name>
<mods:name id="E36A1851727D0A231F2B7CB34FCE7D3D" type="personal">
<mods:role id="9167739115D28BF953B50E30D319FEC0">
<mods:roleTerm id="0E06F05BC42E3F18D4D50E2523201EB4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4624B23559C6B7447C0D50A448E1F7E1">Allen, Josh P.</mods:namePart>
<mods:affiliation id="AD4C2E25DCF0C0F33683F2B4488075EB">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="948BC963E269F71FD76D73831F27DE34" type="email">myrmeciaman@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="509113E5D8988882C042443F894FA549" type="personal">
<mods:role id="B82C0518B551A2397694A59790BDFB7D">
<mods:roleTerm id="A7AAA2FB992E7DB25796759C9A170F55">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC39362C634C02F36A6F8C9B7CEA2B10">Reynel, Carlos</mods:namePart>
<mods:affiliation id="B24CE43B8E7DADEAAE826456C89FCFBD">Facultad de Ciencias Forestales, Universidad Nacional Agraria-La Molina, Lima 12, Peru.</mods:affiliation>
<mods:nameIdentifier id="505CEAB620D5A40B60D1B79A4A53C95A" type="email">reynel@lamolina.edu.pe</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="D381854E54900B4DA439FE46F94EFBB2">text</mods:typeOfResource>
<mods:relatedItem id="A4D0DE17A5C87FE6D48AAD1F06A53252" type="host">
<mods:titleInfo id="D158108F30F6FD07B54A567C89B2F70D">
<mods:title id="BF86FE29BA4E4AB506979B25616C31DB">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="7F25030D73BE4A4553A850038E48ED51">
<mods:date id="5FD40054EA6180B4E132EDA3B33A231E">2023</mods:date>
<mods:detail id="A9BCA43D1920CDA12B88FB490DC546D6" type="pubDate">
<mods:number id="9B0F901672FC6D0C2B1D50D58418C7D5">2023-07-18</mods:number>
</mods:detail>
<mods:detail id="5779A0685DB4FB3D2B7DB05E4D31D990" type="volume">
<mods:number id="00CAD654F6F6B67451162EB46F4EFDA4">881</mods:number>
</mods:detail>
<mods:extent id="5C253268265E2AC69A42EB0A22B3E800" unit="page">
<mods:start id="71FC453C4CCF9781C61936DE974EF107">1</mods:start>
<mods:end id="64C1E9186F083DCB7A3A26A68786A416">334</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="2619C059BCAA878B4AA8E5B3078D14D2">
<mods:url id="8719D8BEF40B8F041FFA7442C204778D">http://dx.doi.org/10.5852/ejt.2023.881.2175</mods:url>
</mods:location>
<mods:classification id="46C2ADDAE2BCBA575D980111AEC2CA24">journal article</mods:classification>
<mods:identifier id="0113D35E30555F6503C382E84D401AAE" type="DOI">https://doi.org/10.5852/ejt.2023.881.2175</mods:identifier>
<mods:identifier id="38431AFAFD5A093F7D5ACA7E2331BA48" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="4E70751493EE4A231798C18E8AB0A129" type="Zenodo-Dep">8178280</mods:identifier>
</mods:mods>
<treatment id="03B26B4BFFCEFF94FDB2F9E2AF4AE96C" LSID="urn:lsid:plazi:treatment:03B26B4BFFCEFF94FDB2F9E2AF4AE96C" httpUri="http://treatment.plazi.org/id/03B26B4BFFCEFF94FDB2F9E2AF4AE96C" lastPageId="109" lastPageNumber="110" pageId="107" pageNumber="108">
<subSubSection id="C30189D6FFCEFF92FDB2F9E2AEEEEC0E" box="[619,969,1629,1656]" pageId="107" pageNumber="108" type="nomenclature">
<paragraph id="8BA4DA5DFFCEFF92FDB2F9E2AEEEEC0E" blockId="107.[619,969,1629,1693]" box="[619,969,1629,1656]" pageId="107" pageNumber="108">
23.
<taxonomicName id="4C1BA1DEFFCEFF92FD4EF9E2AEEEEC0E" ID-CoL="87YTL" authority="Irmsch." authorityName="Irmsch." box="[663,969,1629,1656]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="107" pageNumber="108" phylum="Tracheophyta" rank="species" species="lucifuga">
<emphasis id="B96F064FFFCEFF92FD4EF9E2AE4FEC01" bold="true" box="[663,872,1629,1656]" italics="true" pageId="107" pageNumber="108">Begonia lucifuga</emphasis>
Irmsch.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFCEFF92FD3FF93DAE6AECE4" box="[742,845,1666,1693]" pageId="107" pageNumber="108" type="description">
<paragraph id="8BA4DA5DFFCEFF92FD3FF93DAE6AECE4" blockId="107.[619,969,1629,1693]" box="[742,845,1666,1693]" pageId="107" pageNumber="108">
<figureCitation id="1320C6D8FFCEFF92FD3FF93DAE6AECE4" box="[742,845,1666,1693]" captionStart="Fig" captionStartId="99.[189,232,1822,1848]" captionTargetBox="[194,1391,556,1774]" captionTargetId="figure-78@99.[189,1398,546,1782]" captionTargetPageId="99" captionText="Fig. 35. Distribution of the stenotepala group of Begonia sect. Cyathocnemis (Klotzsch) A.DC. in Peru and surrounding countries. A. B. alto-peruviana A.DC. (red). B. B. brevicordata L.B.Sm. &amp; B.G.Schub. (red), B. deltoidesMoonlight sp. nov. (blue), and B. lucifuga Irmsch (green). C. B. longitepalaMoonlight sp. nov. (red) and B. subspinulosa Irmsch. (blue). D. B. stenotepala L.B.Sm. &amp; B.G.Schub. (red). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178371" httpUri="https://zenodo.org/record/8178371/files/figure.png" pageId="107" pageNumber="108">Fig. 35B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFCEFF92FF64F979AC11ED7C" pageId="107" pageNumber="108" type="reference_group">
<paragraph id="8BA4DA5DFFCEFF92FF64F979AC11ED7C" blockId="107.[189,1399,1734,1872]" pageId="107" pageNumber="108">
<bibCitation id="5E31D231FFCEFF92FF64F979AC14ED7C" journal="Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie" page="608" pageId="107" pageNumber="108" score="478.66328617067893" volume="74" year="1949">
<emphasis id="B96F064FFFCEFF92FF64F979A9BAEC99" box="[189,1181,1734,1760]" italics="true" pageId="107" pageNumber="108">
Botanische Jahrbücher für
<bibCitation id="5E31D231FFCEFF92FDDAF979A9BAEC99" author="Systematik" box="[515,1181,1734,1760]" journal="Pflanzengeschichte und Pflanzengeographie" page="608" pageId="107" pageNumber="108" score="109.65844492058366" volume="74" year="1949">Systematik, Pflanzengeschichte und Pflanzengeographie</bibCitation>
</emphasis>
<materialsCitation id="3B73D000FFCEFF92FB7DF979AC14ED7C" country="Peru" location="Junin Region" pageId="107" pageNumber="108" specimenCount="1" stateProvince="Junin">
74: 608 (
<bibRefCitation id="EF8AA7ACFFCEFF92FAD5F979AC0EED7C" author="Irmscher" firstAuthor="Irmscher" pageId="107" pageNumber="108" pagination="569 - 633" refId="ref228493" refString="Irmscher E. 1949. Begoniaceae, Beitrage zur Kenntnis der Begoniaceen Sudamerikas. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 74: 569 - 633." type="journal article" year="1949">Irmscher 1949</bibRefCitation>
)
</materialsCitation>
</bibCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFCEFF92FEE7F954A842ED52" pageId="107" pageNumber="108" type="materials_examined">
<paragraph id="8BA4DA5DFFCEFF92FEE7F954A842ED52" blockId="107.[189,1399,1734,1872]" pageId="107" pageNumber="108">
<emphasis id="B96F064FFFCEFF92FE8FF954ACB4ED7C" bold="true" box="[342,403,1771,1797]" pageId="107" pageNumber="108">Type</emphasis>
:
<materialsCitation id="3B73D000FFCEFF92FE7AF954A929ED52" collectingDate="1929-06-06" collectionCode="US" collectorName="E. P. Killip &amp; A. C. Smith" country="Peru" elevation="1800" latitude="-11.233334" location="Huacapistana" longLatPrecision="1295" longitude="-75.51667" municipality="Killip" pageId="107" pageNumber="108" specimenCode="US00955825" specimenCount="1" stateProvince="Junin Region" typeStatus="holotype">
<collectingCountry id="F30C9ACDFFCEFF92FE7AF954ACC9ED7C" box="[419,494,1771,1797]" name="Peru" pageId="107" pageNumber="108">PERU</collectingCountry>
<emphasis id="B96F064FFFCEFF92FDD7F955AF9FED7C" bold="true" box="[526,696,1770,1797]" pageId="107" pageNumber="108">
<collectingRegion id="49DF14BFFFCEFF92FDD7F955AF9FED7C" box="[526,696,1770,1797]" country="Peru" name="Junin" pageId="107" pageNumber="108">Junín Region</collectingRegion>
</emphasis>
<location id="8EC48C86FFCEFF92FD08F954AE54ED7C" LSID="urn:lsid:plazi:treatment:03B26B4BFFCEFF94FDB2F9E2AF4AE96C:8EC48C86FFCEFF92FD08F954AE54ED7C" box="[721,883,1771,1797]" country="Peru" latitude="-11.233334" longLatPrecision="1295" longitude="-75.51667" municipality="Killip" name="Huacapistana" pageId="107" pageNumber="108" stateProvince="Junin Region">Huacapistana</location>
; [
<geoCoordinate id="EE2FBC9AFFCEFF92FC5EF956AEC8ED7C" box="[903,1007,1769,1797]" degrees="11" direction="south" minutes="14" orientation="latitude" pageId="107" pageNumber="108" precision="925" value="-11.233334">11°14 S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFFCEFF92FC24F956A955ED7C" box="[1021,1138,1769,1797]" degrees="75" direction="west" minutes="31" orientation="longitude" pageId="107" pageNumber="108" precision="925" value="-75.51667">75°31 W</geoCoordinate>
];
<elevation id="00363D6EFFCEFF92FB5EF954A806ED7C" box="[1159,1313,1771,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="107" pageNumber="108" unit="m" value="1800.0">
<quantity id="4CE377B8FFCEFF92FB5EF954A9C6ED7C" box="[1159,1249,1771,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="107" pageNumber="108" unit="m" value="1800.0">1800 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFFCEFF92FAE9F954AC0FED53" pageId="107" pageNumber="108" value="1929-06-06">
<collectingDate id="EFE10575FFCEFF92FAE9F954AC0FED53" pageId="107" pageNumber="108" value="1929-06-06">6 Jun. 1929</collectingDate>
</date>
;
<emphasis id="B96F064FFFCEFF92FEEDF8AEAF82ED53" box="[308,677,1808,1835]" italics="true" pageId="107" pageNumber="108">
<collectorName id="26EEBF8BFFCEFF92FEEDF8AEAC88ED53" box="[308,431,1808,1834]" pageId="107" pageNumber="108">E.P. Killip</collectorName>
&amp;
<collectorName id="26EEBF8BFFCEFF92FE0DF8AFAF73ED53" box="[468,596,1808,1834]" pageId="107" pageNumber="108">A.C. Smith</collectorName>
27300
</emphasis>
;
<typeStatus id="54A064FFFFCEFF92FD6CF8AFAE07ED53" box="[693,800,1808,1834]" pageId="107" pageNumber="108">holotype</typeStatus>
:
<collectionCode id="ED0A4298FFCEFF92FCF2F8AFAE77ED53" box="[811,848,1808,1834]" pageId="107" pageNumber="108">US</collectionCode>
[
<specimenCode id="DBBD7226FFCEFF92FCBBF8AFAEDAED52" box="[866,1021,1808,1835]" pageId="107" pageNumber="108">US00955825</specimenCode>
];
</materialsCitation>
<materialsCitation id="3B73D000FFCEFF92FBCFF8AFA878ED52" box="[1046,1375,1808,1835]" collectingDate="1929-06-06" collectionCode="NY" collectorName="E. P. Killip &amp; A. C. Smith" country="United States of America" elevation="1800" latitude="-11.233334" location="Smith" longLatPrecision="1295" longitude="-75.51667" municipality="Killip" pageId="107" pageNumber="108" specimenCode="NY01085496" specimenCount="1" stateProvince="Junin Region" typeStatus="isotype">
<typeStatus id="54A064FFFFCEFF92FBCFF8AFA948ED53" box="[1046,1135,1808,1834]" pageId="107" pageNumber="108">isotype</typeStatus>
:
<collectionCode id="ED0A4298FFCEFF92FBA3F8AEA984ED52" box="[1146,1187,1809,1835]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="107" pageNumber="108" type="Herbarium">NY</collectionCode>
[
<specimenCode id="DBBD7226FFCEFF92FB6DF8AFA873ED52" box="[1204,1364,1808,1835]" collectionCode="NY" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="107" pageNumber="108" type="Herbarium">NY01085496</specimenCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFCEFF92FF64F88AA805ED36" box="[189,1314,1845,1872]" pageId="107" pageNumber="108" type="reference_group">
<paragraph id="8BA4DA5DFFCEFF92FF64F88AA805ED36" blockId="107.[189,1399,1734,1872]" box="[189,1314,1845,1872]" pageId="107" pageNumber="108">
<bibRefCitation id="EF8AA7ACFFCEFF92FF64F88AAF01ED36" author="Brako &amp; Zarucchi" box="[189,550,1845,1871]" firstAuthor="Brako" page="193" pageId="107" pageNumber="108" refId="ref225797" refString="Brako L. &amp; Zarucchi L. 1993. Catalogue of the Flowering Plants and Gymnosperms of Peru. Monographs in Systematic Botany from the Missouri Botanical Garden 45, Missouri Botanical Garden, St Louis." type="book" year="1993">Brako &amp; Zarucchi (1993: 193)</bibRefCitation>
;
<bibRefCitation id="EF8AA7ACFFCEFF92FDEBF88AAE8EED29" author="Vasquez" box="[562,937,1845,1872]" etAl="et al." firstAuthor="Vasquez" page="112 - 125" pageId="107" pageNumber="108" pagination="112 - 125" refId="ref233185" refString="Vasquez R., Rojas R. G,. Monteagudo A. M., Meza K. V., van der Werff H., Ortiz-Gentry R. &amp; Catchpole D. 2005. Flora vascular de la selva central del Peru: una aproximacion de la copmposicion floristica de tres areas naturales protegidas. Arnaldoa 12 (1 - 2): 112 - 125." type="journal article" year="2005">
Vásquez
<emphasis id="B96F064FFFCEFF92FD46F889AFFDED36" box="[671,730,1845,1871]" italics="true" pageId="107" pageNumber="108">et al.</emphasis>
(2005: 112125)
</bibRefCitation>
;
<bibRefCitation id="EF8AA7ACFFCEFF92FC6DF88AA839ED36" author="Leon &amp; Monsalve" box="[948,1310,1845,1871]" firstAuthor="Leon" page="167" pageId="107" pageNumber="108" pagination="165 - 170" refId="ref229226" refString="Leon B. &amp; Monsalve C. 2006. Begoniaceae endemicas del Peru. Revista Peruana de Biologia 13 (2): 165 - 170. https: // doi. org / 10.15381 / rpb. v 13 i 2.1810" type="journal article" year="2006">
<collectingRegion id="49DF14BFFFCEFF92FC6DF88AAED7ED36" box="[948,1008,1845,1871]" country="Nicaragua" name="Leon" pageId="107" pageNumber="108">León</collectingRegion>
&amp; Monsalve (2006: 167)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFCEFF92FF64F8C6AFF7ED91" pageId="107" pageNumber="108" type="etymology">
<paragraph id="8BA4DA5DFFCEFF92FF64F8C6AC62EDEA" blockId="107.[189,325,1913,1939]" box="[189,325,1913,1939]" pageId="107" pageNumber="108">
<heading id="D0EC6D31FFCEFF92FF64F8C6AC62EDEA" bold="true" box="[189,325,1913,1939]" fontSize="11" level="3" pageId="107" pageNumber="108" reason="3">
<emphasis id="B96F064FFFCEFF92FF64F8C6AC62EDEA" bold="true" box="[189,325,1913,1939]" pageId="107" pageNumber="108">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFCEFF92FF64F816AFF7ED91" blockId="107.[189,1399,1961,2024]" pageId="107" pageNumber="108">
The epithet derives from the Latin
<emphasis id="B96F064FFFCEFF92FDBBF816AFA1EDBA" box="[610,646,1961,1987]" italics="true" pageId="107" pageNumber="108">lux</emphasis>
and
<emphasis id="B96F064FFFCEFF92FD0DF816AE07EDBA" box="[724,800,1961,1987]" italics="true" pageId="107" pageNumber="108">fugure</emphasis>
meaning light and to escape. This refers to the species preferred habit, which is deep shade.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFC9FF95FF64FEB7AEE5E85F" pageId="108" pageNumber="109" type="materials_examined">
<paragraph id="8BA4DA5DFFC9FF95FF64FEB7ACE3EB5B" blockId="108.[189,452,264,290]" box="[189,452,264,290]" pageId="108" pageNumber="109">
<heading id="D0EC6D31FFC9FF95FF64FEB7ACE3EB5B" bold="true" box="[189,452,264,290]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
<emphasis id="B96F064FFFC9FF95FF64FEB7ACE3EB5B" bold="true" box="[189,452,264,290]" pageId="108" pageNumber="109">Specimens examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFC9FF95FF64FE87AEE5E85F" blockId="108.[189,1399,311,551]" pageId="108" pageNumber="109">
<materialsCitation id="3B73D000FFC9FF95FF64FE87A96EEBE0" collectingDate="2003-06-03" collectionCode="HOXA, MO" collectorName="A. Monteagudo &amp; G. Ortiz" country="Peru" county="Prov. Oxapampa" elevation="1385" latitude="-10.751111" location="Bosque de proteccion San Matias-San Carlos." longLatPrecision="21" longitude="-74.92972" pageId="108" pageNumber="109" specimenCode="MO-842669" specimenCount="1" stateProvince="Pasco Region">
<collectingCountry id="F30C9ACDFFC9FF95FF64FE87AC2FEB2B" box="[189,264,312,338]" name="Peru" pageId="108" pageNumber="109">PERU</collectingCountry>
<emphasis id="B96F064FFFC9FF95FEF6FE88AFE9EB28" bold="true" box="[303,718,311,337]" pageId="108" pageNumber="109">
<collectingRegion id="49DF14BFFFC9FF95FEF6FE88ACC6EB28" box="[303,481,311,337]" country="Peru" name="Pasco" pageId="108" pageNumber="109">Pasco Region</collectingRegion>
:
<collectingCounty id="62C5A2D1FFC9FF95FE2AFE88AFE9EB28" box="[499,718,311,337]" pageId="108" pageNumber="109">Prov. Oxapampa</collectingCounty>
</emphasis>
<collectingCounty id="62C5A2D1FFC9FF95FD28FE88AEB4EB28" box="[753,915,311,337]" pageId="108" pageNumber="109">Dist. Palcazú</collectingCounty>
,
<location id="8EC48C86FFC9FF95FC7DFE87AC36EB0C" LSID="urn:lsid:plazi:treatment:03B26B4BFFCEFF94FDB2F9E2AF4AE96C:8EC48C86FFC9FF95FC7DFE87AC36EB0C" country="Peru" county="Prov. Oxapampa" latitude="-10.751111" longLatPrecision="21" longitude="-74.92972" name="Bosque de proteccion San Matias-San Carlos." pageId="108" pageNumber="109" stateProvince="Pasco Region">Bosque de protección San Matias-San Carlos.</location>
<location id="8EC48C86FFC9FF95FEC5FEE4AE02EB0C" LSID="urn:lsid:plazi:treatment:03B26B4BFFCEFF94FDB2F9E2AF4AE96C:8EC48C86FFC9FF95FEC5FEE4AE02EB0C" box="[284,805,347,373]" country="Peru" county="Prov. Oxapampa" latitude="-10.751111" longLatPrecision="21" longitude="-74.92972" name="Bosque Secundario Sector Union-Shimakii" pageId="108" pageNumber="109" stateProvince="Pasco Region">Bosque Secundario Sector Unión-Shimakii</location>
;
<geoCoordinate id="EE2FBC9AFFC9FF95FCEDFEE6AEEFEB0C" box="[820,968,345,373]" degrees="10" direction="south" minutes="45" orientation="latitude" pageId="108" pageNumber="109" precision="15" seconds="04" value="-10.751111">10°4504″ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFFC9FF95FC01FEE6A950EB0C" box="[984,1143,345,373]" degrees="74" direction="west" minutes="55" orientation="longitude" pageId="108" pageNumber="109" precision="15" seconds="47" value="-74.92972">74°5547″ W</geoCoordinate>
;
<elevation id="00363D6EFFC9FF95FB5EFEE4A857EB0C" box="[1159,1392,347,374]" metricMagnitude="3" metricUnit="m" metricValue="1.385" metricValueMax="1.42" metricValueMin="1.35" pageId="108" pageNumber="109" unit="m" value="1385.0" valueMax="1420.0" valueMin="1350.0">
<quantity id="4CE377B8FFC9FF95FB5EFEE4A80AEB0C" box="[1159,1325,347,374]" metricMagnitude="3" metricUnit="m" metricValue="1.385" metricValueMax="1.42" metricValueMin="1.35" pageId="108" pageNumber="109" unit="m" value="1385.0" valueMax="1420.0" valueMin="1350.0">13501420 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFFC9FF95FF64FEC1AC68EBE1" box="[189,335,382,409]" pageId="108" pageNumber="109" value="2003-06-03">
<collectingDate id="EFE10575FFC9FF95FF64FEC1AC68EBE1" box="[189,335,382,409]" pageId="108" pageNumber="109" value="2003-06-03">3 Jun. 2003</collectingDate>
</date>
;
<emphasis id="B96F064FFFC9FF95FE86FEC0AFD7EBE1" box="[351,752,382,409]" italics="true" pageId="108" pageNumber="109">
<collectorName id="26EEBF8BFFC9FF95FE86FEC0AF3EEBE1" box="[351,537,382,409]" pageId="108" pageNumber="109">A. Monteagudo</collectorName>
&amp;
<collectorName id="26EEBF8BFFC9FF95FD9CFEC0AF8DEBE0" box="[581,682,383,409]" pageId="108" pageNumber="109">G. Ortiz</collectorName>
5685
</emphasis>
;
<collectionCode id="ED0A4298FFC9FF95FCDAFEC1AE7CEBE1" box="[771,859,382,408]" country="Peru" httpUri="http://grbio.org/cool/yriw-6hkr" name="Estación biológica del Jardin Botanico de Missouri" pageId="108" pageNumber="109" type="Herbarium">HOXA</collectionCode>
,
<collectionCode id="ED0A4298FFC9FF95FCB3FEC1AEBEEBE1" box="[874,921,382,408]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="108" pageNumber="109" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FFC9FF95FC74FEC1A919EBE1" box="[941,1086,382,408]" pageId="108" pageNumber="109">MO-842669</specimenCode>
]
</materialsCitation>
.
<materialsCitation id="3B73D000FFC9FF95FBADFEC2A838EB99" collectingDate="1985-03-07" collectionCode="MO" collectorName="B. A. Stein &amp; C. Todzia" country="Peru" county="Prov. Chanchamayo" elevation="1600" latitude="-11.0" location="Pampatigre" longLatPrecision="0" longitude="-75.166664" municipality="Romero" pageId="108" pageNumber="109" specimenCode="MO-1835893" specimenCount="1" stateProvince="Junin Region">
<emphasis id="B96F064FFFC9FF95FBADFEC2AC54EBC5" bold="true" pageId="108" pageNumber="109">
<collectingRegion id="49DF14BFFFC9FF95FBADFEC2A803EBE1" box="[1140,1316,381,408]" country="Peru" name="Junin" pageId="108" pageNumber="109">Junín Region</collectingRegion>
:
<collectingCounty id="62C5A2D1FFC9FF95FAEDFEC1AC54EBC5" pageId="108" pageNumber="109">Prov. Chanchamayo</collectingCounty>
</emphasis>
<collectingCounty id="62C5A2D1FFC9FF95FE48FE1DAF29EBC5" box="[401,526,418,444]" pageId="108" pageNumber="109">Santa Ana</collectingCounty>
, fondo Romero,
<location id="8EC48C86FFC9FF95FD00FE1DAE45EBC5" LSID="urn:lsid:plazi:treatment:03B26B4BFFCEFF94FDB2F9E2AF4AE96C:8EC48C86FFC9FF95FD00FE1DAE45EBC5" box="[729,866,418,444]" country="Peru" county="Prov. Chanchamayo" latitude="-11.0" longLatPrecision="0" longitude="-75.166664" municipality="Romero" name="Pampatigre" pageId="108" pageNumber="109" stateProvince="Junin Region">Pampatigre</location>
,
<locationDeviation id="9A16C37FFFC9FF95FCB6FE1DA80EEBC5" box="[879,1321,418,444]" pageId="108" pageNumber="109">above Santa Ana (SE of La Merced)</locationDeviation>
;
<geoCoordinate id="EE2FBC9AFFC9FF95FAEEFE1DA856EBC5" box="[1335,1393,418,444]" degrees="11" direction="south" orientation="latitude" pageId="108" pageNumber="109" precision="55555" value="-11.0">11°S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFFC9FF95FF66FE7BAC17EB99" box="[191,304,452,480]" degrees="75" direction="west" minutes="10" orientation="longitude" pageId="108" pageNumber="109" precision="925" value="-75.166664">75°10 W</geoCoordinate>
;
<elevation id="00363D6EFFC9FF95FEE4FE7AAF06EBA6" box="[317,545,453,480]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="108" pageNumber="109" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">
<quantity id="4CE377B8FFC9FF95FEE4FE7AACC7EB99" box="[317,480,453,480]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="108" pageNumber="109" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">15001700 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFFC9FF95FDF6FE79AFE4EB99" box="[559,707,453,480]" pageId="108" pageNumber="109" value="1985-03-07">
<collectingDate id="EFE10575FFC9FF95FDF6FE79AFE4EB99" box="[559,707,453,480]" pageId="108" pageNumber="109" value="1985-03-07">7 Mar. 1985</collectingDate>
</date>
;
<emphasis id="B96F064FFFC9FF95FD09FE79A903EBA6" box="[720,1060,453,480]" italics="true" pageId="108" pageNumber="109">
<collectorName id="26EEBF8BFFC9FF95FD09FE79AE61EB99" box="[720,838,454,480]" pageId="108" pageNumber="109">B.A. Stein</collectorName>
&amp;
<collectorName id="26EEBF8BFFC9FF95FCB6FE79AEC7EBA6" box="[879,992,453,480]" pageId="108" pageNumber="109">C. Todzia</collectorName>
2338
</emphasis>
;
<collectionCode id="ED0A4298FFC9FF95FBEDFE7AA944EBA6" box="[1076,1123,453,479]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="108" pageNumber="109" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FFC9FF95FBAFFE7AA832EB99" box="[1142,1301,453,480]" pageId="108" pageNumber="109">MO-1835893</specimenCode>
]
</materialsCitation>
<materialsCitation id="3B73D000FFC9FF95FAE3FE7AAE99E85F" collectingDate="2014-06-23" collectionCode="MOL" collectorName="P. W. Moonlight &amp; A. Daza" country="Peru" county="Prov. Chanchamayo" elevation="1523" latitude="-11.172501" location="Puente de Azucar" longLatPrecision="21" longitude="-75.453606" municipality="rio Tarma" pageId="108" pageNumber="109" specimenCount="1" stateProvince="Junin">
<locationDeviation id="9A16C37FFFC9FF95FAE3FE7AA908E87A" pageId="108" pageNumber="109">
<quantity id="4CE377B8FFC9FF95FAE3FE7AA851EBA6" box="[1338,1398,453,479]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="108" pageNumber="109" unit="km" value="1.0">1 km</quantity>
along path N along tributary of río Tarma from E side of Puente de Azúcar
</locationDeviation>
;
<geoCoordinate id="EE2FBC9AFFC9FF95FBE2FE58A9EEE87A" box="[1083,1225,487,515]" degrees="11" direction="south" minutes="10" orientation="latitude" pageId="108" pageNumber="109" precision="15" seconds="21" value="-11.172501">11°1021″ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFFC9FF95FB0FFE58A855E87A" box="[1238,1394,487,515]" degrees="75" direction="west" minutes="27" orientation="longitude" pageId="108" pageNumber="109" precision="15" seconds="13" value="-75.453606">75°2713″ W</geoCoordinate>
;
<elevation id="00363D6EFFC9FF95FF64FDB3AC71E85F" box="[189,342,524,551]" metricMagnitude="3" metricUnit="m" metricValue="1.523" pageId="108" pageNumber="109" unit="m" value="1523.0">
<quantity id="4CE377B8FFC9FF95FF64FDB3AC31E85E" box="[189,278,524,551]" metricMagnitude="3" metricUnit="m" metricValue="1.523" pageId="108" pageNumber="109" unit="m" value="1523.0">1523 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFFC9FF95FEBAFDB3ACDAE85F" box="[355,509,524,551]" pageId="108" pageNumber="109" value="2014-06-23">
<collectingDate id="EFE10575FFC9FF95FEBAFDB3ACDAE85F" box="[355,509,524,551]" pageId="108" pageNumber="109" value="2014-06-23">23 Jun. 2014</collectingDate>
</date>
;
<emphasis id="B96F064FFFC9FF95FDD0FDB2AE4EE85F" box="[521,873,524,551]" italics="true" pageId="108" pageNumber="109">
<collectorName id="26EEBF8BFFC9FF95FDD0FDB2AF98E85F" box="[521,703,524,550]" pageId="108" pageNumber="109">P.W. Moonlight</collectorName>
&amp;
<collectorName id="26EEBF8BFFC9FF95FD3CFDB2AE63E85F" box="[741,836,525,551]" pageId="108" pageNumber="109">A. Daza</collectorName>
46
</emphasis>
;
<collectionCode id="ED0A4298FFC9FF95FCA0FDB3AE99E85F" box="[889,958,524,550]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="108" pageNumber="109" type="Herbarium">MOL</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFC9FF95FF64FDF1AF68EFF9" pageId="108" pageNumber="109" type="description">
<paragraph id="8BA4DA5DFFC9FF95FF64FDF1AC68E811" blockId="108.[189,335,590,616]" box="[189,335,590,616]" pageId="108" pageNumber="109">
<heading id="D0EC6D31FFC9FF95FF64FDF1AC68E811" bold="true" box="[189,335,590,616]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
<emphasis id="B96F064FFFC9FF95FF64FDF1AC68E811" bold="true" box="[189,335,590,616]" pageId="108" pageNumber="109">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFC9FF95FF64FDC2AF68EFF9" blockId="108.[189,1400,637,1408]" pageId="108" pageNumber="109">
Caulescent herb, to
<quantity id="4CE377B8FFC9FF95FE7FFDC2ACC8E8EE" box="[422,495,637,663]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="108" pageNumber="109" unit="cm" value="40.0">40 cm</quantity>
high.
<emphasis id="B96F064FFFC9FF95FDE0FDC2AF55E8EE" box="[569,626,637,663]" italics="true" pageId="108" pageNumber="109">Stem</emphasis>
erect, rarely branching; internodes to
<quantity id="4CE377B8FFC9FF95FBE8FDC2A9A6E8EE" box="[1073,1153,637,664]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="108" pageNumber="109" unit="cm" value="2.5">2.5 cm</quantity>
long, to
<quantity id="4CE377B8FFC9FF95FB33FDC2A809E8EE" box="[1258,1326,637,663]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="108" pageNumber="109" unit="mm" value="6.0">6 mm</quantity>
thick, succulent, red, glabrous.
<emphasis id="B96F064FFFC9FF95FE30FD1FAF60E8C3" box="[489,583,672,698]" italics="true" pageId="108" pageNumber="109">Stipules</emphasis>
deciduous, ovate, 1525 ×
<quantity id="4CE377B8FFC9FF95FC4EFD1FA92BE8C2" box="[919,1036,672,699]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="108" pageNumber="109" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">510 mm</quantity>
, apex obtuse, translucent, red, glabrous, margin entire, aciliate.
<emphasis id="B96F064FFFC9FF95FD98FD7AAFB4E8A7" box="[577,659,709,734]" italics="true" pageId="108" pageNumber="109">Leaves</emphasis>
&gt; 3, alternate, basifixed; petiole
<quantity id="4CE377B8FFC9FF95FBCCFD7BA9ADE8A7" box="[1045,1162,708,735]" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="108" pageNumber="109" unit="cm" value="22.5" valueMax="30.0" valueMin="15.0">1530 cm</quantity>
long, red, glabrous; blade asymmetric, lanceolate, to 12 ×
<quantity id="4CE377B8FFC9FF95FD4EFD58AFFEE97B" box="[663,729,743,770]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="108" pageNumber="109" unit="cm" value="5.0">5 cm</quantity>
, succulent, apex acuminate, base obliquely truncate to obliquely cordate, basal lobes overlapping on the young leaves, sinus to
<quantity id="4CE377B8FFC9FF95FBD3FCB4A969E95F" box="[1034,1102,779,806]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="108" pageNumber="109" unit="mm" value="5.0">5 mm</quantity>
deep, margin serrulate to serrate, ciliate, upper surface dark green, glabrous, lower surface red, glabrous, veins palmate-pinnate, 68 veined from the base, with 57 secondary veins on the larger side, 25 secondary veins on the smaller side.
<emphasis id="B96F064FFFC9FF95FE86FCCAAF20E9F6" box="[351,519,885,911]" italics="true" pageId="108" pageNumber="109">Inflorescences</emphasis>
13, bisexual, axillary, erect, cymose, with 34 branches, bearing up to 8 staminate flowers and 8 pistillate flowers, protandrous; peduncle to
<quantity id="4CE377B8FFC9FF95FBC3FC25A97FE9CD" box="[1050,1112,922,948]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="108" pageNumber="109" unit="cm" value="7.0">7 cm</quantity>
long, colour unknown, glabrous, bracts deciduous, obovate, ca 23 ×
<quantity id="4CE377B8FFC9FF95FD27FC02AE6DE9AE" box="[766,842,957,983]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="108" pageNumber="109" unit="mm" value="1.0">1 mm</quantity>
, translucent, colour unknown, glabrous, apex truncate, margin entire, sparsely ciliate.
<emphasis id="B96F064FFFC9FF95FD4EFC5FAE4DE983" box="[663,874,992,1018]" italics="true" pageId="108" pageNumber="109">Staminate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FFC9FF95FBD8FC5FA973E982" box="[1025,1108,992,1019]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="108" pageNumber="109" unit="mm" value="24.0">24 mm</quantity>
long, glabrous; tepals 2, spreading, broadly ovate, 915 ×
<quantity id="4CE377B8FFC9FF95FD95FBBBAFE7EE67" box="[588,704,1028,1054]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="108" pageNumber="109" unit="mm" value="10.5" valueMax="13.0" valueMin="8.0">813 mm</quantity>
, apex rounded, pale pink, glabrous, margin entire, aciliate; stamens 2530, spreading, presumed yellow, filaments
<quantity id="4CE377B8FFC9FF95FC86FB98AEE4EE3B" box="[863,963,1063,1090]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="108" pageNumber="109" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
long, free, anthers ellipsoid, 12 ×
<quantity id="4CE377B8FFC9FF95FF64FBF5AC3CEE1C" box="[189,283,1098,1125]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="108" pageNumber="109" unit="mm" value="0.5">0.5 mm</quantity>
long, dehiscing via lateral slits, connectives extended to
<quantity id="4CE377B8FFC9FF95FC39FBF4A965EE1C" box="[992,1090,1099,1125]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="108" pageNumber="109" unit="mm" value="0.2">0.2 mm</quantity>
, symmetrically basifixed.
<emphasis id="B96F064FFFC9FF95FF64FBD1ACAFEEF1" box="[189,392,1134,1160]" italics="true" pageId="108" pageNumber="109">Pistillate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FFC9FF95FDFEFBD1AF5AEEF0" box="[551,637,1134,1161]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="108" pageNumber="109" unit="mm" value="15.0">15 mm</quantity>
long; bracteoles present, number unknown, positioned directly beneath the ovary, elliptic, ca 1.5 ×
<quantity id="4CE377B8FFC9FF95FDBAFB2DAF8DEED5" box="[611,682,1170,1196]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="108" pageNumber="109" unit="mm" value="1.0">1 mm</quantity>
, apex obtuse, translucent, colour unknown, glabrous, margin serrulate, ciliate; tepals 5, subequal, persistence in fruit unknown, projecting, ovate, 35 ×
<quantity id="4CE377B8FFC9FF95FAD5FB0AA854EEA9" box="[1292,1395,1205,1232]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="108" pageNumber="109" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
, apex obtuse to rounded, pink, glabrous, margin entire, aciliate; ovary body ovoid, ca 2.54 ×
<quantity id="4CE377B8FFC9FF95FAD4FB66A855EE8A" box="[1293,1394,1241,1267]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="108" pageNumber="109" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
, colour unknown, glabrous, unequally 3-winged, wings triangular, largest 35 ×
<quantity id="4CE377B8FFC9FF95FBA8FB43A9F1EF6E" box="[1137,1238,1276,1303]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="108" pageNumber="109" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
, smallest 34 ×
<quantity id="4CE377B8FFC9FF95FF01FAA0AC72EF43" box="[216,341,1311,1338]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="108" pageNumber="109" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 mm</quantity>
; 3-locular, placentae branches divided, bearing ovules on both surfaces; styles 3, colour unknown, free, ca
<quantity id="4CE377B8FFC9FF95FE78FAFCACDAEF27" box="[417,509,1347,1374]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="108" pageNumber="109" unit="mm" value="1.5">1.5 mm</quantity>
long, once-divided, stigmatic papillae in a spirally twisted band.
<emphasis id="B96F064FFFC9FF95FACDFAFBAC34EFF9" italics="true" pageId="108" pageNumber="109">Fruiting pedicel</emphasis>
unknown.
<emphasis id="B96F064FFFC9FF95FE4FFAD8ACF4EFF9" box="[406,467,1383,1408]" italics="true" pageId="108" pageNumber="109">Fruit</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFC9FF95FF64FA17A851ED2D" pageId="108" pageNumber="109" type="conservation">
<paragraph id="8BA4DA5DFFC9FF95FF64FA17AF4AEFBB" blockId="108.[189,621,1448,1474]" box="[189,621,1448,1474]" pageId="108" pageNumber="109">
<heading id="D0EC6D31FFC9FF95FF64FA17AF4AEFBB" bold="true" box="[189,621,1448,1474]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
<emphasis id="B96F064FFFC9FF95FF64FA17AF4AEFBB" bold="true" box="[189,621,1448,1474]" pageId="108" pageNumber="109">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFC9FF95FF64FA68A851ED2D" blockId="108.[188,1399,1495,1877]" pageId="108" pageNumber="109">
Previously assessed as Data Deficient (DD) by
<bibRefCitation id="EF8AA7ACFFC9FF95FD31FA68A930EF88" author="Leon &amp; Monsalve" box="[744,1047,1495,1521]" firstAuthor="Leon" pageId="108" pageNumber="109" pagination="165 - 170" refId="ref229226" refString="Leon B. &amp; Monsalve C. 2006. Begoniaceae endemicas del Peru. Revista Peruana de Biologia 13 (2): 165 - 170. https: // doi. org / 10.15381 / rpb. v 13 i 2.1810" type="journal article" year="2006">León &amp; Monsalve (2006)</bibRefCitation>
. Known from four collections across four populations with a combined EOO of
<quantity id="4CE377B8FFC9FF95FCD5FA44AE42EC6C" box="[780,869,1531,1557]" metricMagnitude="5" metricUnit="m" metricValue="2.1399999999999997" pageId="108" pageNumber="109" unit="km" value="214.0">214 km</quantity>
<superScript id="7C6E7715FFC9FF95FCBCFA45AE49EC71" attach="left" box="[869,878,1530,1544]" fontSize="6" pageId="108" pageNumber="109">2</superScript>
. Despite extensive fieldwork in the species range, we have only been able to locate a single individual (
<emphasis id="B96F064FFFC9FF95FC7CF9A0A830EC41" box="[933,1303,1566,1593]" italics="true" pageId="108" pageNumber="109">P.W. Moonlight &amp; A. Daza 36</emphasis>
), likely because the range of
<taxonomicName id="4C1BA1DEFFC9FF95FE12F9FDAF74EC25" box="[459,595,1602,1628]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="species" species="lucifuga">
<emphasis id="B96F064FFFC9FF95FE12F9FDAF74EC25" box="[459,595,1602,1628]" italics="true" pageId="108" pageNumber="109">B. lucifuga</emphasis>
</taxonomicName>
is centred on the valley of the río Chanchamayo. The Peruvian “Carretera Central” (National Route 22) descends from the Andes to the Amazon along this valley and is the only major road across the Andes for ca
<quantity id="4CE377B8FFC9FF95FD2FF936AE76ECDA" box="[758,849,1673,1699]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="108" pageNumber="109" unit="km" value="200.0">200 km</quantity>
in either direction. The Carretera Central has been a major route for more than 150 years and most of the Lower Montane Forest within the valley has been cleared for agriculture. We are also confident that
<taxonomicName id="4C1BA1DEFFC9FF95FCE4F96FAE99EC93" box="[829,958,1744,1770]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="species" species="lucifuga">
<emphasis id="B96F064FFFC9FF95FCE4F96FAE99EC93" box="[829,958,1744,1770]" italics="true" pageId="108" pageNumber="109">B. lucifuga</emphasis>
</taxonomicName>
does not grow in large numbers north of its known range in
<collectingRegion id="49DF14BFFFC9FF95FE18F94BAF45ED74" box="[449,610,1779,1806]" country="Peru" name="Pasco" pageId="108" pageNumber="109">Pasco Region</collectingRegion>
because this part of
<collectingCountry id="F30C9ACDFFC9FF95FC81F94BAEA9ED77" box="[856,910,1780,1806]" name="Peru" pageId="108" pageNumber="109">Peru</collectingCountry>
has for the past 20 years been one of the most intensively collected areas of tropical forest in the world. Considering its rarity, small range, and the habitat destruction across that range, we assess
<taxonomicName id="4C1BA1DEFFC9FF95FCD6F884AEB7ED2D" box="[783,912,1850,1876]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="species" species="lucifuga">
<emphasis id="B96F064FFFC9FF95FCD6F884AEB7ED2D" box="[783,912,1850,1876]" italics="true" pageId="108" pageNumber="109">B. lucifuga</emphasis>
</taxonomicName>
as Endangered (EN B1ab(iii)+B2ab(iii)).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFC9FF94FF64F8C3ACD6E824" lastPageId="109" lastPageNumber="110" pageId="108" pageNumber="109" type="discussion">
<paragraph id="8BA4DA5DFFC9FF95FF64F8C3AC97EDEF" blockId="108.[189,432,1916,1942]" box="[189,432,1916,1942]" pageId="108" pageNumber="109">
<heading id="D0EC6D31FFC9FF95FF64F8C3AC97EDEF" bold="true" box="[189,432,1916,1942]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
<emphasis id="B96F064FFFC9FF95FF64F8C3AC97EDEF" bold="true" box="[189,432,1916,1942]" pageId="108" pageNumber="109">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFC9FF94FF64F814AF6EEBC8" blockId="108.[188,1398,1963,2025]" lastBlockId="109.[189,1399,265,433]" lastPageId="109" lastPageNumber="110" pageId="108" pageNumber="109">
Within the stenotepala group,
<taxonomicName id="4C1BA1DEFFC9FF95FDC4F813AFB8EDBC" box="[541,671,1963,1989]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="species" species="lucifuga">
<emphasis id="B96F064FFFC9FF95FDC4F813AFB8EDBC" box="[541,671,1963,1989]" italics="true" pageId="108" pageNumber="109">B. lucifuga</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C1BA1DEFFC9FF95FCAFF813A909EDBC" box="[886,1070,1963,1989]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="species" species="brevicordata">
<emphasis id="B96F064FFFC9FF95FCAFF813A909EDBC" box="[886,1070,1963,1989]" italics="true" pageId="108" pageNumber="109">B. brevicordata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1BA1DEFFC9FF95FBBEF813A804EDBC" box="[1127,1315,1963,1989]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="species" species="subspinulosa">
<emphasis id="B96F064FFFC9FF95FBBEF813A804EDBC" box="[1127,1315,1963,1989]" italics="true" pageId="108" pageNumber="109">B. subspinulosa</emphasis>
</taxonomicName>
. These three species have white or pink tepals and the tepals on their staminate flowers have a rounded apex while the other species in the complex have bright red or orange tepals with an acute to narrowly acute apex.
<taxonomicName id="4C1BA1DEFFC8FF94FEDBFE92ACEAEB3F" box="[258,461,300,326]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="lucifuga">
<emphasis id="B96F064FFFC8FF94FEDBFE92ACEAEB3F" box="[258,461,300,326]" italics="true" pageId="109" pageNumber="110">Begonia lucifuga</emphasis>
</taxonomicName>
differs from these two species in its leaf margins, which are serrulate as opposed to serrate or dentate. Its leaves are also lanceolate in outline and have an acuminate apex, whereas those
<emphasis id="B96F064FFFC8FF94FF64FECCACB0EBF4" box="[189,407,371,397]" italics="true" pageId="109" pageNumber="110">
of
<taxonomicName id="4C1BA1DEFFC8FF94FF02FECBACB0EBF4" box="[219,407,371,397]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="subspinulosa">B. subspinulosa</taxonomicName>
</emphasis>
are ovate and those of
<taxonomicName id="4C1BA1DEFFC8FF94FD70FECBAE45EBF4" box="[681,866,371,397]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="brevicordata">
<emphasis id="B96F064FFFC8FF94FD70FECBAE45EBF4" box="[681,866,371,397]" italics="true" pageId="109" pageNumber="110">B. brevicordata</emphasis>
</taxonomicName>
are broadly ovate in outline and both species have short acuminate leaf apexes.
</paragraph>
<paragraph id="8BA4DA5DFFC8FF94FF64FE67ACD6E824" blockId="109.[189,1399,472,605]" pageId="109" pageNumber="110">
When sterile,
<taxonomicName id="4C1BA1DEFFC8FF94FEBFFE66ACCCEB8B" box="[358,491,472,498]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="lucifuga">
<emphasis id="B96F064FFFC8FF94FEBFFE66ACCCEB8B" box="[358,491,472,498]" italics="true" pageId="109" pageNumber="110">B. lucifuga</emphasis>
</taxonomicName>
is most likely to be confused with
<taxonomicName id="4C1BA1DEFFC8FF94FC45FE66A96EEB8B" box="[924,1097,472,498]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="stenotepala">
<emphasis id="B96F064FFFC8FF94FC45FE66A96EEB8B" box="[924,1097,472,498]" italics="true" pageId="109" pageNumber="110">B. stenotepala</emphasis>
</taxonomicName>
or
<emphasis id="B96F064FFFC8FF94FBACFE66A825EB8B" box="[1141,1282,472,498]" italics="true" pageId="109" pageNumber="110">
<taxonomicName id="4C1BA1DEFFC8FF94FBACFE66A9D9EB8B" box="[1141,1278,472,498]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="lucifuga">B. lucifuga</taxonomicName>
.
</emphasis>
It can be distinguished from
<taxonomicName id="4C1BA1DEFFC8FF94FE7BFE42AF6AE86F" box="[418,589,508,534]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="stenotepala">
<emphasis id="B96F064FFFC8FF94FE7BFE42AF6AE86F" box="[418,589,508,534]" italics="true" pageId="109" pageNumber="110">B. stenotepala</emphasis>
</taxonomicName>
by its stipules, which are deciduous whereas those of
<taxonomicName id="4C1BA1DEFFC8FF94FB14FE42A850E86F" box="[1229,1399,508,534]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="stenotepala">
<emphasis id="B96F064FFFC8FF94FB14FE42A850E86F" box="[1229,1399,508,534]" italics="true" pageId="109" pageNumber="110">B. stenotepala</emphasis>
</taxonomicName>
are persistent.
<taxonomicName id="4C1BA1DEFFC8FF94FEA9FD9FAF42E840" authority="Moonlight &amp; Jara-Muñoz &amp; Purvis &amp; Delves &amp; Allen &amp; Reynel, 2023" authorityName="Moonlight &amp; Jara-Muñoz &amp; Purvis &amp; Delves &amp; Allen &amp; Reynel" authorityYear="2023" box="[368,613,543,569]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="longitepala" status="sp. nov.">
<emphasis id="B96F064FFFC8FF94FEA9FD9FAF42E840" box="[368,613,543,569]" italics="true" pageId="109" pageNumber="110">Begonia longitepala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FFC8FF94FDA8FD9FAFF7E843" box="[625,720,544,570]" pageId="109" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
also has deciduous stipules but differs in its elliptic to oblanceolate leaf laminae.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFC8FF94FF64FD3BAF4AE96C" pageId="109" pageNumber="110" type="distribution">
<paragraph id="8BA4DA5DFFC8FF94FF64FD3BACD2E8E6" blockId="109.[189,501,644,671]" box="[189,501,644,671]" pageId="109" pageNumber="110">
<heading id="D0EC6D31FFC8FF94FF64FD3BACD2E8E6" bold="true" box="[189,501,644,671]" fontSize="11" level="3" pageId="109" pageNumber="110" reason="3">
<emphasis id="B96F064FFFC8FF94FF64FD3BACD2E8E6" bold="true" box="[189,501,644,671]" pageId="109" pageNumber="110">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFC8FF94FF64FD0CAF4AE96C" blockId="109.[188,1399,691,789]" pageId="109" pageNumber="110">
Endemic to
<collectingCountry id="F30C9ACDFFC8FF94FE91FD0BAC59E8B7" box="[328,382,692,718]" name="Peru" pageId="109" pageNumber="110">Peru</collectingCountry>
and known from
<collectingRegion id="49DF14BFFFC8FF94FD90FD0BAFAAE8B7" box="[585,653,692,718]" country="Peru" name="Pasco" pageId="109" pageNumber="110">Pasco</collectingRegion>
and
<collectingRegion id="49DF14BFFFC8FF94FD1DFD0CAE23E8B4" box="[708,772,691,717]" country="Peru" name="Junin" pageId="109" pageNumber="110">Junín</collectingRegion>
Regions (
<figureCitation id="1320C6D8FFC8FF94FCA0FD0CAEC3E8B7" box="[889,996,691,718]" captionStart="Fig" captionStartId="99.[189,232,1822,1848]" captionTargetBox="[194,1391,556,1774]" captionTargetId="figure-78@99.[189,1398,546,1782]" captionTargetPageId="99" captionText="Fig. 35. Distribution of the stenotepala group of Begonia sect. Cyathocnemis (Klotzsch) A.DC. in Peru and surrounding countries. A. B. alto-peruviana A.DC. (red). B. B. brevicordata L.B.Sm. &amp; B.G.Schub. (red), B. deltoidesMoonlight sp. nov. (blue), and B. lucifuga Irmsch (green). C. B. longitepalaMoonlight sp. nov. (red) and B. subspinulosa Irmsch. (blue). D. B. stenotepala L.B.Sm. &amp; B.G.Schub. (red). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178371" httpUri="https://zenodo.org/record/8178371/files/figure.png" pageId="109" pageNumber="110">Fig. 35B</figureCitation>
). Found in lower montane forest at an elevation of
<quantity id="4CE377B8FFC8FF94FEA8FD68AF34E888" box="[369,531,727,754]" metricMagnitude="3" metricUnit="m" metricValue="1.575" metricValueMax="1.8" metricValueMin="1.35" pageId="109" pageNumber="110" unit="m" value="1575.0" valueMax="1800.0" valueMin="1350.0">13501800 m</quantity>
a.s.l.
<taxonomicName id="4C1BA1DEFFC8FF94FD80FD67AE03E888" box="[601,804,727,753]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="lucifuga">
<emphasis id="B96F064FFFC8FF94FD80FD67AE03E888" box="[601,804,727,753]" italics="true" pageId="109" pageNumber="110">Begonia lucifuga</emphasis>
</taxonomicName>
is found in deep shade and humid areas, including in the shade of forests within gorges.
</paragraph>
</subSubSection>
</treatment>
</document>