treatments-xml/data/03/B2/6B/03B26B4BFFC8FF96FE24FCECAF1DED90.xml

<document id="33717AEC268EAFF732AF742F22F8BD7D" ID-DOI="https://doi.org/10.5852/ejt.2023.881.2175" ID-ISSN="2118-9773" ID-Zenodo-Dep="8178280" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690201937279" checkinUser="felipe" docAuthor="Moonlight, Peter. W., Jara-Muñoz, Orlando A., Purvis, David A., Delves, Jay, Allen, Josh P. &amp; Reynel, Carlos" docDate="2023" docId="03B26B4BFFC8FF96FE24FCECAF1DED90" docLanguage="en" docName="ejt-881_1-334.imf" docOrigin="European Journal of Taxonomy 881" docSource="http://dx.doi.org/10.5852/ejt.2023.881.2175" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia stenotepala L. B. Sm. &amp; B. G. Schub." docType="treatment" docVersion="7" lastPageNumber="112" masterDocId="FF8B1333FFA5FFF9FFD9FFBFAD27EA79" masterDocTitle="The genus Begonia (Begoniaceae) in Peru" masterLastPageNumber="334" masterPageNumber="1" pageNumber="110" updateTime="1707123162844" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="4053B5364CFA141B7A36A473A48CAA6F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="ADB537AD30B8640409E89AEA769ECFFD">
<mods:title id="EB7B0878870ABCAFC7910DB16C57030B">The genus Begonia (Begoniaceae) in Peru</mods:title>
</mods:titleInfo>
<mods:name id="15B2306E197128C16024DF7A4A40DED8" type="personal">
<mods:role id="AE77536A2E5014D2E85F993440D9CEA4">
<mods:roleTerm id="ABB7730F7108ACED417EFDFD2F166D04">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA2281AC357B5D7DCD12A8F3CAE33AC9">Moonlight, Peter. W.</mods:namePart>
<mods:affiliation id="46EFFB78EFB8F6C34464D69AABC4AFDA">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK. &amp; Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland.</mods:affiliation>
<mods:nameIdentifier id="B594441E0F96C9596C24824643F199AD" type="email">moonligp@tcd.ie</mods:nameIdentifier>
</mods:name>
<mods:name id="38889C3CB1370D52DEFEF5F0B35BAB26" type="personal">
<mods:role id="8F71A9CBB5EAE7AB2DCCD32530291FA2">
<mods:roleTerm id="5490E291AFED193A131705AE082345F6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6816CE1B39DB74F432D3A5C80D3FEC43">Jara-Muñoz, Orlando A.</mods:namePart>
<mods:affiliation id="DA89E8EB483835BA8F3EC108E765CDCA">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá D. C., Colombia.</mods:affiliation>
<mods:nameIdentifier id="D76C180A0980A49468C9211238B8BE06" type="email">oajaram@unal.edu.co</mods:nameIdentifier>
</mods:name>
<mods:name id="3671B4D4919AA499E494B90D0E3BD794" type="personal">
<mods:role id="721CC06A196518943F8CB919D4FC24CC">
<mods:roleTerm id="4768974A69CE4A9FC4829D60E4E44CCF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E7DC8B511EBD272F5837A069FB192E3B">Purvis, David A.</mods:namePart>
<mods:affiliation id="2D87F9BEE1634B76E80F65EAE6540769">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="5FF760F1631A3644C7A95A7EB39D4E39" type="email">dpurvis@rbge.org.uk</mods:nameIdentifier>
</mods:name>
<mods:name id="C6BE4CA455892148C40AEA2DBAF0618B" type="personal">
<mods:role id="D5B231892B158A6D9D1AF7B75CA01AD8">
<mods:roleTerm id="27EF5627E42A1E88DF46F733BC9713F8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3A56BC95D004C832B2261C01AD696223">Delves, Jay</mods:namePart>
<mods:affiliation id="27AB6BC66D90CC33F982AD344811BDB8">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="A23BA083108C9B50603D7C793C121709" type="email">j.delves@westernsydney.edu.au</mods:nameIdentifier>
</mods:name>
<mods:name id="1CE21F1CC01013F74D9F0318166A96BA" type="personal">
<mods:role id="DB7D5049D87C6429E5C0E427217B84C0">
<mods:roleTerm id="AFD0CE15DECEF17D27C3BEB0E3F6D737">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A7544C950141CB8780DB0FD63DC3E147">Allen, Josh P.</mods:namePart>
<mods:affiliation id="AA7B73FC94FE80A8E4DAC8E520CB18B7">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="BA075AC380F7D74A23AB0C71C9F76103" type="email">myrmeciaman@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="7CCA4592D6FCBD4E44388F7536E120D9" type="personal">
<mods:role id="36FE86DF4E4DA912268FC72F23C0807E">
<mods:roleTerm id="262C4796245CDC8563954B6BFE5C808C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="72DC51FBDBA72841723C9FA3981EEA7C">Reynel, Carlos</mods:namePart>
<mods:affiliation id="B307915984DCD6AAA8DCA2F6818612DE">Facultad de Ciencias Forestales, Universidad Nacional Agraria-La Molina, Lima 12, Peru.</mods:affiliation>
<mods:nameIdentifier id="DF133A8495AF09EA5DB8D9F3F94F7989" type="email">reynel@lamolina.edu.pe</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="4348424A34347AEAA7E741FEFD6C355E">text</mods:typeOfResource>
<mods:relatedItem id="2A3B9EA3005A59CE4C3B21936709A41E" type="host">
<mods:titleInfo id="81ED3FDDD71A59A0D6D2BD0E311D4D67">
<mods:title id="C10334340882A12178E4815F64D78A32">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="891600A3DF85B1903FB71EFCEBADF8A8">
<mods:date id="ECCEDEEAF9185065A03C4A4FB9F36C04">2023</mods:date>
<mods:detail id="5C1688B46F69F4B3AB1A9B7BF975165E" type="pubDate">
<mods:number id="25AD05D0DC4F7E060BB11E9BE0C09E35">2023-07-18</mods:number>
</mods:detail>
<mods:detail id="1716223B39BE255E6BB6F8F5C4469A3D" type="volume">
<mods:number id="B9E9A648F229B589F83305BDA6D2DB1D">881</mods:number>
</mods:detail>
<mods:extent id="DC0574AA8BB7191CE0F7430C8D8C2426" unit="page">
<mods:start id="63D281169B3A57BB28ECD4E1CA1AE53A">1</mods:start>
<mods:end id="B8641107C29803A62CA5DF9B6697504B">334</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="CEF2C68A6B5BD1DBC52181FAD20F0217">
<mods:url id="3297ADEA739C57388F5C1390F84DF406">http://dx.doi.org/10.5852/ejt.2023.881.2175</mods:url>
</mods:location>
<mods:classification id="1AC076463C8D3158CFDFA9390AE9934B">journal article</mods:classification>
<mods:identifier id="AFB3D45F1B4D84BEF870A7CDAFF3A4AB" type="DOI">https://doi.org/10.5852/ejt.2023.881.2175</mods:identifier>
<mods:identifier id="E964AC3E4EFB48863C7238658981EC25" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="EAEEC95DF0E96401CDA9B7A4C2FC099E" type="Zenodo-Dep">8178280</mods:identifier>
</mods:mods>
<treatment id="03B26B4BFFC8FF96FE24FCECAF1DED90" LSID="urn:lsid:plazi:treatment:03B26B4BFFC8FF96FE24FCECAF1DED90" httpUri="http://treatment.plazi.org/id/03B26B4BFFC8FF96FE24FCECAF1DED90" lastPageId="111" lastPageNumber="112" pageId="109" pageNumber="110">
<subSubSection id="C30189D6FFC8FF94FE24FCECA911E914" box="[509,1078,850,877]" pageId="109" pageNumber="110" type="nomenclature">
<paragraph id="8BA4DA5DFFC8FF94FE24FCECA911E914" blockId="109.[509,1078,850,913]" box="[509,1078,850,877]" pageId="109" pageNumber="110">
24. 
<taxonomicName id="4C1BA1DEFFC8FF94FDF0FCEDA911E914" ID-CoL="7SHLJ" authority="L. B. Sm. &amp; B. G. Schub." authorityName="L. B. Sm. &amp; B. G. Schub." box="[553,1078,850,877]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="stenotepala">
<emphasis id="B96F064FFFC8FF94FDF0FCEDAE38E914" bold="true" box="[553,799,850,877]" italics="true" pageId="109" pageNumber="110">Begonia stenotepala</emphasis>
L.B.Sm. &amp; B.G.Schub.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFC8FF94FD5AFCC9AE96E9E9" box="[643,945,886,913]" pageId="109" pageNumber="110" type="description">
<paragraph id="8BA4DA5DFFC8FF94FD5AFCC9AE96E9E9" blockId="109.[509,1078,850,913]" box="[643,945,886,913]" pageId="109" pageNumber="110">
<figureCitation id="1320C6D8FFC8FF94FD5AFCC9AFC5E9E9" box="[643,738,886,912]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[189,1398,788,1816]" captionTargetId="figure-11@5.[189,1398,788,1816]" captionTargetPageId="5" captionText="Fig. 1. Leaf margins of Peruvian Begonia. A. Entire, aciliate margins (B. stenotepala A.DC.). B. Crenate, aciliate margins (B. joshii Moonlight). C. Denticulate, ciliate margins (B. acerifolia Kunth). D. Irregularly double-dentate, ciliate margins (B. obtecticaulis Irmsch.). E. Serrulate, aciliate margins (B. glauca (Klotzsch) Ruiz &amp; Pav. exA.DC.). F. Serrate, ciliate margins (B. foliosa Kunth). Photographs taken by D.A. Purvis &amp; P.W. Moonlight in the living collections of the Royal Botanic Garden Edinburgh." figureDoi="http://doi.org/10.5281/zenodo.8178282" httpUri="https://zenodo.org/record/8178282/files/figure.png" pageId="109" pageNumber="110">Figs 1A</figureCitation>
, 
<figureCitation id="1320C6D8FFC8FF94FD36FCC9AE32E9E9" box="[751,789,886,912]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[272,1315,265,1740]" captionTargetId="figure-13@6.[272,1315,265,1740]" captionTargetPageId="6" captionText="Fig. 2. Venation in Peruvian Begonia. A. Pinnate venation (B. yaracuyensis Moonlight). B. Pinnate venation, 3-veined from the base (B. glabra Aubl.). C. Palmate-pinnate venation (B. obtecticaulis Irmsch.). D. Palmate venation with one primary vein (B. stenotepalaL.B.Sm. &amp; B.G.Schub.).E. Palmate venation (B. lamolina Moonlight). F. Peltate venation with one primary vein (B. acerifolia Kunth.). G. Peltate venation (B. joshii Moonlight). Primary veins are shown in white, secondary veins from the petiole insertion in blue, and secondary veins from the primary veins in green. Photographs taken by D.A. Purvis &amp; P.W. Moonlight in the living collections of the Royal Botanic Garden Edinburgh." figureDoi="http://doi.org/10.5281/zenodo.8178284" httpUri="https://zenodo.org/record/8178284/files/figure.png" pageId="109" pageNumber="110">2D</figureCitation>
, 
<figureCitation id="1320C6D8FFC8FF94FCF8FCC9AE61E9E8" box="[801,838,886,913]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[189,1398,1291,1854]" captionTargetId="figure-12@9.[189,1398,1291,1854]" captionTargetPageId="9" captionText="Fig. 5. Staminate tepal projection in Peruvian Begonia. A–B. Spreading tepals. C–D. Projecting tepals. A. B. glabra Aubl. B. B. bracteosa A.DC. C. B. stenotepala L.B.Sm. &amp; B.G.Schub. D. B. foliosa Kunth. Photographs taken by D.A. Purvis &amp; P.W. Moonlight in the living collections of the Royal Botanic Garden Edinburgh." figureDoi="http://doi.org/10.5281/zenodo.8178290" httpUri="https://zenodo.org/record/8178290/files/figure.png" pageId="109" pageNumber="110">5C</figureCitation>
, 
<figureCitation id="1320C6D8FFC8FF94FC8BFCC9AEAFE9E8" box="[850,904,886,913]" captionStart="Fig" captionStartId="99.[189,232,1822,1848]" captionTargetBox="[194,1391,556,1774]" captionTargetId="figure-78@99.[189,1398,546,1782]" captionTargetPageId="99" captionText="Fig. 35. Distribution of the stenotepala group of Begonia sect. Cyathocnemis (Klotzsch) A.DC. in Peru and surrounding countries. A. B. alto-peruviana A.DC. (red). B. B. brevicordata L.B.Sm. &amp; B.G.Schub. (red), B. deltoidesMoonlight sp. nov. (blue), and B. lucifuga Irmsch (green). C. B. longitepalaMoonlight sp. nov. (red) and B. subspinulosa Irmsch. (blue). D. B. stenotepala L.B.Sm. &amp; B.G.Schub. (red). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178371" httpUri="https://zenodo.org/record/8178371/files/figure.png" pageId="109" pageNumber="110">35D</figureCitation>
, 
<figureCitation id="1320C6D8FFC8FF94FC4DFCC9AE96E9E9" box="[916,945,886,912]" captionStart="Fig" captionStartId="110.[189,232,1787,1813]" captionTargetBox="[272,1315,265,1740]" captionTargetId="figure-13@110.[272,1315,265,1740]" captionTargetPageId="110" captionText="Fig. 38. Begonia stenotepala L.B.Sm. &amp; B.G.Schub. A. Habit. B. Leaf, adaxial surface. C. Leaf, abaxial surface. D. Stipule. E. Staminate flower, front view. F. Staminate flower, side view. G. Top tepal of the staminate flower. H. Bottom tepal of the staminate flower. I. Androecium, side view. J. Pistillate flower, front view. K. Pistillate flower, side view. L. Pistils, side view. M. Smallest tepal of the pistillate flower. N. Largest tepal of the pistillate flower. O. Bracteole. All photographs taken by D.A. Purvis &amp; P.W. Moonlight in the living collections of the Royal Botanic Garden Edinburgh (Accession 20141048, grown from seeds collected as part of P.W. Moonlight &amp; A. Daza 83)." figureDoi="http://doi.org/10.5281/zenodo.8178377" httpUri="https://zenodo.org/record/8178377/files/figure.png" pageId="109" pageNumber="110">38</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFC8FF94FF64FC07AC65E98C" pageId="109" pageNumber="110" type="reference_group">
<paragraph id="8BA4DA5DFFC8FF94FF64FC07AC65E98C" blockId="109.[189,1398,952,1085]" pageId="109" pageNumber="110">
<emphasis id="B96F064FFFC8FF94FF64FC07AEDFE9AB" box="[189,1016,952,978]" italics="true" pageId="109" pageNumber="110">Publications of the Field Museum of Natural History, Botanical Series</emphasis>
13 (4/1): 200 (
<bibRefCitation id="EF8AA7ACFFC8FF94FB7AFC07AC12E98C" author="Smith &amp; Schubert" firstAuthor="Smith" pageId="109" pageNumber="110" pagination="181 - 202" refId="ref231211" refString="Smith L. B. &amp; Schubert B. G. 1941 a. Flora of Peru, Begoniaceae. Publications of the Field Museum of Natural History, Botanical Series 13 (4 / 1): 181 - 202." type="journal article" year="1941" yearSuffix="a">Smith &amp; Schubert 1941a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFC8FF94FE9EFC63A871EE60" pageId="109" pageNumber="110" type="materials_examined">
<paragraph id="8BA4DA5DFFC8FF94FE9EFC63A871EE60" blockId="109.[189,1398,952,1085]" pageId="109" pageNumber="110">
– 
<emphasis id="B96F064FFFC8FF94FE80FC64ACB0E98C" bold="true" box="[345,407,987,1013]" pageId="109" pageNumber="110">Type</emphasis>
: 
<materialsCitation id="3B73D000FFC8FF94FE7EFC63A875EE60" collectingDate="1936-07-24" collectionCode="UC" collectorName="J. West" country="Peru" elevation="1150" latitude="-13.05" location="Cusco Region" longLatPrecision="1291" longitude="-71.53333" pageId="109" pageNumber="110" specimenCode="UC578669" specimenCount="1" stateProvince="Cusco" typeStatus="holotype">
<collectingCountry id="F30C9ACDFFC8FF94FE7EFC63ACD5E98F" box="[423,498,988,1014]" name="Peru" pageId="109" pageNumber="110">PERU</collectingCountry>
– 
<emphasis id="B96F064FFFC8FF94FDD1FC64AF94E98C" bold="true" box="[520,691,987,1013]" pageId="109" pageNumber="110">
<collectingRegion id="49DF14BFFFC8FF94FDD1FC64AF94E98C" box="[520,691,987,1013]" country="Peru" name="Cusco" pageId="109" pageNumber="110">Cusco Region</collectingRegion>
</emphasis>
• along 
<location id="8EC48C86FFC8FF94FCC9FC64AEEEE98C" LSID="urn:lsid:plazi:treatment:03B26B4BFFC8FF96FE24FCECAF1DED90:8EC48C86FFC8FF94FCC9FC64AEEEE98C" box="[784,969,987,1013]" country="Peru" latitude="-13.05" longLatPrecision="1291" longitude="-71.53333" name="rio Tambomayo" pageId="109" pageNumber="110" stateProvince="Cusco">río Tambomayo</location>
, 
<locationDeviation id="9A16C37FFFC8FF94FC0AFC64A855E98C" box="[979,1394,987,1013]" firstLocation="Tambomayo" pageId="109" pageNumber="110" secondLocation="Asuncion">between Tambomayo and Asunción</locationDeviation>
; [
<geoCoordinate id="EE2FBC9AFFC8FF94FF29FC40AC7DEE60" box="[240,346,1021,1049]" degrees="13" direction="south" minutes="03" orientation="latitude" pageId="109" pageNumber="110" precision="925" value="-13.05">13°03′ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFFC8FF94FEBEFC42ACFBEE60" box="[359,476,1021,1049]" degrees="71" direction="west" minutes="32" orientation="longitude" pageId="109" pageNumber="110" precision="925" value="-71.53333">71°32′ W</geoCoordinate>
]; 
<elevation id="00363D6EFFC8FF94FE29FC40AFE4EE60" box="[496,707,1023,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.4" metricValueMin="0.9" pageId="109" pageNumber="110" unit="m" value="1150.0" valueMax="1400.0" valueMin="900.0">
<quantity id="4CE377B8FFC8FF94FE29FC40AFA4EE60" box="[496,643,1023,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.4" metricValueMin="0.9" pageId="109" pageNumber="110" unit="m" value="1150.0" valueMax="1400.0" valueMin="900.0">900–1400 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFFC8FF94FD08FC40AE43EE60" box="[721,868,1023,1049]" pageId="109" pageNumber="110" value="1936-07-24">
<collectingDate id="EFE10575FFC8FF94FD08FC40AE43EE60" box="[721,868,1023,1049]" pageId="109" pageNumber="110" value="1936-07-24">24 Jul. 1936</collectingDate>
</date>
; 
<emphasis id="B96F064FFFC8FF94FCA9FC40A923EE60" box="[880,1028,1023,1049]" italics="true" pageId="109" pageNumber="110">J. West 7114</emphasis>
; 
<typeStatus id="54A064FFFFC8FF94FBCAFC40A959EE60" box="[1043,1150,1023,1049]" pageId="109" pageNumber="110">holotype</typeStatus>
: 
<collectionCode id="ED0A4298FFC8FF94FB53FC40A994EE60" box="[1162,1203,1023,1049]" country="USA" name="Upjohn Culture Collection" pageId="109" pageNumber="110">UC</collectionCode>
[
<specimenCode id="DBBD7226FFC8FF94FB18FC41A86CEE60" box="[1217,1355,1022,1049]" collectionCode="UC" country="USA" name="Upjohn Culture Collection" pageId="109" pageNumber="110">UC578669</specimenCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFC8FF94FF64FB9DAE86EE45" box="[189,929,1058,1085]" pageId="109" pageNumber="110" type="reference_group">
<paragraph id="8BA4DA5DFFC8FF94FF64FB9DAE86EE45" blockId="109.[189,1398,952,1085]" box="[189,929,1058,1085]" pageId="109" pageNumber="110">
<bibRefCitation id="EF8AA7ACFFC8FF94FF64FB9DAF01EE44" author="Brako &amp; Zarucchi" box="[189,550,1058,1085]" firstAuthor="Brako" page="195" pageId="109" pageNumber="110" refId="ref225797" refString="Brako L. &amp; Zarucchi L. 1993. Catalogue of the Flowering Plants and Gymnosperms of Peru. Monographs in Systematic Botany from the Missouri Botanical Garden 45, Missouri Botanical Garden, St Louis." type="book" year="1993">Brako &amp; Zarucchi (1993: 195)</bibRefCitation>
; 
<bibRefCitation id="EF8AA7ACFFC8FF94FDEAFB9DAEBAEE45" author="Leon &amp; Monsalve" box="[563,925,1058,1084]" firstAuthor="Leon" page="169" pageId="109" pageNumber="110" pagination="165 - 170" refId="ref229226" refString="Leon B. &amp; Monsalve C. 2006. Begoniaceae endemicas del Peru. Revista Peruana de Biologia 13 (2): 165 - 170. https: // doi. org / 10.15381 / rpb. v 13 i 2.1810" type="journal article" year="2006">
<collectingRegion id="49DF14BFFFC8FF94FDEAFB9DAF48EE45" box="[563,623,1058,1084]" country="Nicaragua" name="Leon" pageId="109" pageNumber="110">León</collectingRegion>
&amp; Monsalve (2006: 169)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFC8FF94FF64FBDBA839EEA9" pageId="109" pageNumber="110" type="etymology">
<paragraph id="8BA4DA5DFFC8FF94FF64FBDBAC62EE07" blockId="109.[189,325,1124,1150]" box="[189,325,1124,1150]" pageId="109" pageNumber="110">
<heading id="D0EC6D31FFC8FF94FF64FBDBAC62EE07" bold="true" box="[189,325,1124,1150]" fontSize="11" level="3" pageId="109" pageNumber="110" reason="3">
<emphasis id="B96F064FFFC8FF94FF64FBDBAC62EE07" bold="true" box="[189,325,1124,1150]" pageId="109" pageNumber="110">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFC8FF94FF64FB2CA839EEA9" blockId="109.[186,1399,1171,1233]" pageId="109" pageNumber="110">
The name derives from the Greek words, ‘ 
<emphasis id="B96F064FFFC8FF94FD40FB2CAFC4EED5" box="[665,739,1171,1196]" italics="true" pageId="109" pageNumber="110">stenos</emphasis>
’ and ‘ 
<emphasis id="B96F064FFFC8FF94FCF3FB2CAEA7EED4" box="[810,896,1171,1197]" italics="true" pageId="109" pageNumber="110">tepalus</emphasis>
’, meaning ‘narrow’ and ‘tepal’ respectively. This refers to the relatively narrow, triangular tepals of both the staminate and pistillate flowers.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFC8FF94FF64FB48AFFFED77" pageId="109" pageNumber="110" type="materials_examined">
<paragraph id="8BA4DA5DFFC8FF94FF64FB48AF09EF68" blockId="109.[189,558,1271,1297]" box="[189,558,1271,1297]" pageId="109" pageNumber="110">
<heading id="D0EC6D31FFC8FF94FF64FB48AF09EF68" bold="true" box="[189,558,1271,1297]" fontSize="11" level="3" pageId="109" pageNumber="110" reason="3">
<emphasis id="B96F064FFFC8FF94FF64FB48AF09EF68" bold="true" box="[189,558,1271,1297]" pageId="109" pageNumber="110">Selected specimens examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFC8FF94FF64FA98AFFFED77" blockId="109.[189,1399,1317,1806]" pageId="109" pageNumber="110">
<materialsCitation id="3B73D000FFC8FF94FF64FA98AF42EFF1" collectingDate="2004-04-25" collectingDateMax="2004-04-30" collectingDateMin="2004-04-25" collectionCode="US, USM" collectorName="N. Salinas &amp; H. Beltran &amp; R. B. Foster &amp; C. Vriesendorp" country="Peru" county="Dist. Echarate" elevation="750" latitude="-12.163333" location="Kapiromashi" longLatPrecision="21" longitude="-72.57528" municipality="Prov. La Convencion" pageId="109" pageNumber="110" specimenCode="US002222332" specimenCount="1" stateProvince="Cusco Region">
<collectingCountry id="F30C9ACDFFC8FF94FF64FA98AC2FEF38" box="[189,264,1319,1345]" name="Peru" pageId="109" pageNumber="110">PERU</collectingCountry>
– 
<emphasis id="B96F064FFFC8FF94FEE2FA99AE09EF38" bold="true" box="[315,814,1318,1345]" pageId="109" pageNumber="110">
<collectingRegion id="49DF14BFFFC8FF94FEE2FA99ACD0EF39" box="[315,503,1318,1344]" country="Peru" name="Cusco" pageId="109" pageNumber="110">Cusco Region</collectingRegion>
: 
<collectingMunicipality id="6BC04027FFC8FF94FDD6FA98AE09EF38" box="[527,814,1318,1345]" pageId="109" pageNumber="110">Prov. La Convención</collectingMunicipality>
</emphasis>
• 
<collectingCounty id="62C5A2D1FFC8FF94FC85FA99A937EF39" box="[860,1040,1318,1344]" pageId="109" pageNumber="110">Dist. Echarate</collectingCounty>
, 
<location id="8EC48C86FFC8FF94FBFFFA99A9E6EF39" LSID="urn:lsid:plazi:treatment:03B26B4BFFC8FF96FE24FCECAF1DED90:8EC48C86FFC8FF94FBFFFA99A9E6EF39" box="[1062,1217,1318,1344]" country="Peru" county="Dist. Echarate" latitude="-12.163333" longLatPrecision="21" longitude="-72.57528" municipality="Prov. La Convencion" name="Kapiromashi" pageId="109" pageNumber="110" stateProvince="Cusco Region">Kapiromashi</location>
, 
<geoCoordinate id="EE2FBC9AFFC8FF94FB0EFA98A856EF38" box="[1239,1393,1317,1345]" degrees="12" direction="south" minutes="09" orientation="latitude" pageId="109" pageNumber="110" precision="15" seconds="48" value="-12.163333">12°09′48″ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFFC8FF94FF64FAF7AC7CEF1C" box="[189,347,1352,1381]" degrees="72" direction="west" minutes="34" orientation="longitude" pageId="109" pageNumber="110" precision="15" seconds="31" value="-72.57528">72°34′31″ W</geoCoordinate>
; 
<elevation id="00363D6EFFC8FF94FEB3FAF5ACDDEF1D" box="[362,506,1354,1381]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="109" pageNumber="110" unit="m" value="750.0">
<quantity id="4CE377B8FFC8FF94FEB3FAF5AC9FEF1C" box="[362,440,1354,1381]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="109" pageNumber="110" unit="m" value="750.0">750 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFFC8FF94FDD2FAF5AFFEEF1D" box="[523,729,1354,1381]" pageId="109" pageNumber="110" value="2004-04-25" valueMax="2004-04-30" valueMin="2004-04-25">
<collectingDate id="EFE10575FFC8FF94FDD2FAF5AFFEEF1D" box="[523,729,1354,1381]" pageId="109" pageNumber="110" value="2004-04-25" valueMax="2004-04-30" valueMin="2004-04-25">25–30 Apr. 2004</collectingDate>
</date>
; 
<emphasis id="B96F064FFFC8FF94FD31FAF4ADDFEFFE" italics="true" pageId="109" pageNumber="110">
<collectorName id="26EEBF8BFFC8FF94FD31FAF4AE4FEF1D" box="[744,872,1354,1380]" pageId="109" pageNumber="110">N. Salinas</collectorName>
, 
<collectorName id="26EEBF8BFFC8FF94FCAEFAF4AEDDEF1D" box="[887,1018,1354,1380]" pageId="109" pageNumber="110">H. Beltrán</collectorName>
, 
<collectorName id="26EEBF8BFFC8FF94FBD0FAF4A9B3EF1D" box="[1033,1172,1355,1380]" pageId="109" pageNumber="110">R.B. Foster</collectorName>
&amp; 
<collectorName id="26EEBF8BFFC8FF94FB19FAF5A851EF1D" box="[1216,1398,1354,1380]" pageId="109" pageNumber="110">C. Vriesendorp</collectorName>
6691
</emphasis>
; 
<collectionCode id="ED0A4298FFC8FF94FECBFAD1AC10EFF1" box="[274,311,1390,1416]" pageId="109" pageNumber="110">US</collectionCode>
[
<specimenCode id="DBBD7226FFC8FF94FE8BFAD1ACDBEFF1" box="[338,508,1390,1416]" pageId="109" pageNumber="110">US002222332</specimenCode>
], 
<collectionCode id="ED0A4298FFC8FF94FDC7FAD1AF46EFF1" box="[542,609,1390,1416]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="109" pageNumber="110">USM</collectionCode>
.
</materialsCitation>
<materialsCitation id="3B73D000FFC8FF94FDAFFAD1A845EFD2" collectingDate="2007-08-05" collectingDateMax="2004-04-30" collectingDateMin="2004-04-25" collectionCode="MO" collectorName="I. Huamantupa" country="Peru" county="Dist. Kosnipata" elevation="700" latitude="-12.163333" location="pongo de Konec" longLatPrecision="21" longitude="-72.57528" municipality="Prov. Paucartambo" pageId="109" pageNumber="110" specimenCode="MO-2132027" specimenCount="1" stateProvince="Cusco">
– 
<emphasis id="B96F064FFFC8FF94FD4FFAD1AEB2EFF1" bold="true" box="[662,917,1390,1416]" pageId="109" pageNumber="110">
<collectingMunicipality id="6BC04027FFC8FF94FD4FFAD1AEB2EFF1" box="[662,917,1390,1416]" pageId="109" pageNumber="110">Prov. Paucartambo</collectingMunicipality>
</emphasis>
• 
<collectingCounty id="62C5A2D1FFC8FF94FC18FAD2A9A2EFFE" box="[961,1157,1389,1415]" pageId="109" pageNumber="110">Dist. Kosñipata</collectingCounty>
, 
<location id="8EC48C86FFC8FF94FB43FAD1A855EFF1" LSID="urn:lsid:plazi:treatment:03B26B4BFFC8FF96FE24FCECAF1DED90:8EC48C86FFC8FF94FB43FAD1A855EFF1" box="[1178,1394,1389,1416]" country="Peru" county="Dist. Kosnipata" latitude="-12.163333" longLatPrecision="21" longitude="-72.57528" municipality="Prov. Paucartambo" name="pongo de Konec" pageId="109" pageNumber="110" stateProvince="Cusco">pongo de Koñec</location>
; 
<geoCoordinate id="EE2FBC9AFFC8FF94FF64FA30AC77EFD2" box="[189,336,1423,1451]" degrees="12" direction="south" minutes="53" orientation="latitude" pageId="109" pageNumber="110" precision="15" seconds="53" value="-12.898055">12°53′53″ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFFC8FF94FEB9FA30ACD9EFD2" box="[352,510,1423,1451]" degrees="72" direction="west" minutes="22" orientation="longitude" pageId="109" pageNumber="110" precision="15" seconds="25" value="-72.37361">72°22′25″ W</geoCoordinate>
; 
<elevation id="00363D6EFFC8FF94FDD4FA2EAFB9EFD2" box="[525,670,1425,1451]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="109" pageNumber="110" unit="m" value="700.0">
<quantity id="4CE377B8FFC8FF94FDD4FA2EAF7CEFD2" box="[525,603,1425,1451]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="109" pageNumber="110" unit="m" value="700.0">700 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFFC8FF94FD77FA2EAE6FEFD2" box="[686,840,1425,1452]" pageId="109" pageNumber="110" value="2007-08-05">
<collectingDate id="EFE10575FFC8FF94FD77FA2EAE6FEFD2" box="[686,840,1425,1452]" pageId="109" pageNumber="110" value="2007-08-05">5 Aug. 2007</collectingDate>
</date>
; 
<emphasis id="B96F064FFFC8FF94FC8EFA2DA945EFD2" box="[855,1122,1425,1451]" italics="true" pageId="109" pageNumber="110">
<collectorName id="26EEBF8BFFC8FF94FC8EFA2DA92AEFD2" box="[855,1037,1426,1451]" pageId="109" pageNumber="110">I. Huamantupa</collectorName>
10294
</emphasis>
; 
<collectionCode id="ED0A4298FFC8FF94FBACFA2EA983EFD2" box="[1141,1188,1425,1451]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="109" pageNumber="110" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FFC8FF94FB61FA2EA87FEFD2" box="[1208,1368,1425,1451]" pageId="109" pageNumber="110">MO-2132027</specimenCode>
]
</materialsCitation>
• 
<materialsCitation id="3B73D000FFC8FF94FF64FA0BAEACEF8B" collectingDate="2014-08-04" collectionCode="E, MOL, USM" collectorName="P. W. Moonlight &amp; A. Daza" country="United States of America" elevation="1784" latitude="-13.073334" location="km 67 of road from Paucautambo to Pilcopata" longitude="-71.5558333" municipality="Moonlight" pageId="109" pageNumber="110" specimenCode="E00724451" specimenCount="1">
<location id="8EC48C86FFC8FF94FF64FA0BAFC3EFB7" LSID="urn:lsid:plazi:treatment:03B26B4BFFC8FF96FE24FCECAF1DED90:8EC48C86FFC8FF94FF64FA0BAFC3EFB7" box="[189,740,1460,1487]" country="United States of America" latitude="-13.073334" municipality="Moonlight" name="km 67 of road from Paucautambo to Pilcopata" pageId="109" pageNumber="110">km 67 of road from Paucautambo to Pilcopata</location>
; 
<geoCoordinate id="EE2FBC9AFFC8FF94FD29FA0BAEA7EFB7" box="[752,896,1459,1487]" degrees="13" direction="south" minutes="04" orientation="latitude" pageId="109" pageNumber="110" precision="15" seconds="24" value="-13.073334">13°04′24″ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFFC8FF94FC57FA0CA900EFB6" box="[910,1063,1459,1487]" pageId="109" pageNumber="110">71°33’21′ W</geoCoordinate>
; 
<elevation id="00363D6EFFC8FF94FBEDFA0BA9E9EFB7" box="[1076,1230,1460,1487]" metricMagnitude="3" metricUnit="m" metricValue="1.784" pageId="109" pageNumber="110" unit="m" value="1784.0">
<quantity id="4CE377B8FFC8FF94FBEDFA0BA9A9EFB6" box="[1076,1166,1460,1487]" metricMagnitude="3" metricUnit="m" metricValue="1.784" pageId="109" pageNumber="110" unit="m" value="1784.0">1784 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFFC8FF94FB04FA0BA855EFB7" box="[1245,1394,1460,1487]" pageId="109" pageNumber="110" value="2014-08-04">
<collectingDate id="EFE10575FFC8FF94FB04FA0BA855EFB7" box="[1245,1394,1460,1487]" pageId="109" pageNumber="110" value="2014-08-04">4 Aug. 2014</collectingDate>
</date>
; 
<emphasis id="B96F064FFFC8FF94FF64FA66AF04EF8B" box="[189,547,1496,1522]" italics="true" pageId="109" pageNumber="110">
<collectorName id="26EEBF8BFFC8FF94FF64FA66AC53EF8B" box="[189,372,1496,1522]" pageId="109" pageNumber="110">P.W. Moonlight</collectorName>
&amp; 
<collectorName id="26EEBF8BFFC8FF94FE45FA67ACDAEF8B" box="[412,509,1496,1522]" pageId="109" pageNumber="110">A. Daza</collectorName>
83
</emphasis>
; 
<collectionCode id="ED0A4298FFC8FF94FDECFA67AF60EF8B" box="[565,583,1496,1522]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="109" pageNumber="110" type="Herbarium">E</collectionCode>
[
<specimenCode id="DBBD7226FFC8FF94FD80FA67AFC6EF8A" box="[601,737,1496,1523]" collectionCode="E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="109" pageNumber="110" type="Herbarium">E00724451</specimenCode>
], 
<collectionCode id="ED0A4298FFC8FF94FD23FA67AE19EF8B" box="[762,830,1496,1522]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="109" pageNumber="110" type="Herbarium">MOL</collectionCode>
, 
<collectionCode id="ED0A4298FFC8FF94FC92FA67AEACEF8B" box="[843,907,1496,1522]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="109" pageNumber="110">USM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B73D000FFC8FF94FC7FFA67AC23EC40" collectingDate="1983-12-13" collectionCode="USM" collectorName="R. B. Foster &amp; T. Wachter" country="United States of America" county="Valle del Pilcopata" elevation="1250" latitude="-13.166667" location="Valle del Pilcopata" longLatPrecision="1291" longitude="-71.416664" municipality="Foster" pageId="109" pageNumber="110" specimenCount="1">
<collectingCounty id="62C5A2D1FFC8FF94FC7FFA67A9A0EF8B" box="[934,1159,1496,1522]" pageId="109" pageNumber="110">Valle del Pilcopata</collectingCounty>
, 
<locationDeviation id="9A16C37FFFC8FF94FB4DFA67AC10EC6C" pageId="109" pageNumber="110">road from Patria to Pillahuata</locationDeviation>
; 
<geoCoordinate id="EE2FBC9AFFC8FF94FE9FFA45AC89EC6C" box="[326,430,1530,1558]" degrees="13" direction="south" minutes="10" orientation="latitude" pageId="109" pageNumber="110" precision="925" value="-13.166667">13°10′ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFFC8FF94FE67FA45AF15EC6F" box="[446,562,1530,1558]" degrees="71" direction="west" minutes="25" orientation="longitude" pageId="109" pageNumber="110" precision="925" value="-71.416664">71°25′ W</geoCoordinate>
; 
<elevation id="00363D6EFFC8FF94FD9BFA44AE0DEC6C" box="[578,810,1531,1558]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="109" pageNumber="110" unit="m" value="1250.0" valueMax="1500.0" valueMin="1000.0">
<quantity id="4CE377B8FFC8FF94FD9BFA44AFC0EC6F" box="[578,743,1531,1558]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="109" pageNumber="110" unit="m" value="1250.0" valueMax="1500.0" valueMin="1000.0">1000–1500 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFFC8FF94FCE3FA44AEC6EC6C" box="[826,993,1531,1558]" pageId="109" pageNumber="110" value="1983-12-13">
<collectingDate id="EFE10575FFC8FF94FCE3FA44AEC6EC6C" box="[826,993,1531,1558]" pageId="109" pageNumber="110" value="1983-12-13">13 Dec. 1983</collectingDate>
</date>
; 
<emphasis id="B96F064FFFC8FF94FC29FA43A849EC6C" box="[1008,1390,1531,1558]" italics="true" pageId="109" pageNumber="110">
<collectorName id="26EEBF8BFFC8FF94FC29FA43A95DEC6C" box="[1008,1146,1532,1557]" pageId="109" pageNumber="110">R.B. Foster</collectorName>
&amp; 
<collectorName id="26EEBF8BFFC8FF94FB7EFA43A80EEC6C" box="[1191,1321,1531,1557]" pageId="109" pageNumber="110">T. Wachter</collectorName>
7465
</emphasis>
; 
<collectionCode id="ED0A4298FFC8FF94FF64F9A0AC27EC40" box="[189,256,1567,1593]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="109" pageNumber="110">USM</collectionCode>
.
</materialsCitation>
<materialsCitation id="3B73D000FFC8FF94FED4F9A0A9DDEC25" collectingDate="1991-08-10" collectingDateMax="2004-10-23" collectingDateMin="1945-07-15" collectionCode="MO, USM" collectorName="P. Nunez" country="United States of America" county="Valle del Pilcopata" elevation="643" latitude="-13.166667" location="Prov. Quispicanchis" longLatPrecision="1291" longitude="-71.416664" municipality="Prov. Quispicanchis" pageId="109" pageNumber="110" specimenCode="MO-2228053" specimenCount="1" stateProvince="Madre de Dios Region">
– 
<emphasis id="B96F064FFFC8FF94FEFFF9A0AF04EC41" bold="true" box="[294,547,1566,1593]" pageId="109" pageNumber="110">
<collectingMunicipality id="6BC04027FFC8FF94FEFFF9A0AF04EC41" box="[294,547,1566,1593]" pageId="109" pageNumber="110">Prov. Quispicanchis</collectingMunicipality>
</emphasis>
• 
<locationDeviation id="9A16C37FFFC8FF94FDE4F9A0A855EC40" box="[573,1394,1567,1593]" pageId="109" pageNumber="110">hills around río Araza between Pan de Azúcar and Quincemil airport</locationDeviation>
; 
<geoCoordinate id="EE2FBC9AFFC8FF94FF67F9FEAC04EC25" box="[190,291,1601,1629]" degrees="13" direction="south" minutes="13" orientation="latitude" pageId="109" pageNumber="110" precision="925" value="-13.216666">13°13′ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFFC8FF94FEE8F9FEAC84EC24" box="[305,419,1601,1629]" degrees="70" direction="west" minutes="45" orientation="longitude" pageId="109" pageNumber="110" precision="925" value="-70.75">70°45′ W</geoCoordinate>
; 
<elevation id="00363D6EFFC8FF94FE76F9FDAF1DEC25" box="[431,570,1602,1629]" metricMagnitude="2" metricUnit="m" metricValue="6.43" pageId="109" pageNumber="110" unit="m" value="643.0">
<quantity id="4CE377B8FFC8FF94FE76F9FDACDDEC24" box="[431,506,1602,1629]" metricMagnitude="2" metricUnit="m" metricValue="6.43" pageId="109" pageNumber="110" unit="m" value="643.0">643 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFFC8FF94FD91F9FDAFCDEC25" box="[584,746,1602,1628]" pageId="109" pageNumber="110" value="1991-08-10">
<collectingDate id="EFE10575FFC8FF94FD91F9FDAFCDEC25" box="[584,746,1602,1628]" pageId="109" pageNumber="110" value="1991-08-10">10 Aug. 1991</collectingDate>
</date>
; 
<emphasis id="B96F064FFFC8FF94FD2EF9FCAE79EC24" box="[759,862,1603,1629]" italics="true" pageId="109" pageNumber="110">
<collectorName id="26EEBF8BFFC8FF94FD2EF9FCAE79EC24" box="[759,862,1603,1629]" pageId="109" pageNumber="110">P. Núñez</collectorName>
</emphasis>
14129 (
<collectionCode id="ED0A4298FFC8FF94FC19F9FDAED6EC25" box="[960,1009,1602,1628]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="109" pageNumber="110" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FFC8FF94FBDBF9FDA985EC24" box="[1026,1186,1602,1629]" pageId="109" pageNumber="110">MO-2228053</specimenCode>
], 
<collectionCode id="ED0A4298FFC8FF94FB63F9FDA9DDEC25" box="[1210,1274,1602,1628]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="109" pageNumber="110">USM</collectionCode>
</materialsCitation>
<materialsCitation id="3B73D000FFC8FF94FAD8F9FCA802ECF9" collectingDate="1945-07-15" collectingDateMax="2004-10-23" collectingDateMin="1945-07-15" collectionCode="CUZ" collectorName="J. C. Vargas Calderon" country="United States of America" county="Valle del Pilcopata" elevation="1100" latitude="-13.166667" location="Mandor" longLatPrecision="1291" longitude="-71.416664" municipality="Foster" pageId="109" pageNumber="110" specimenCount="1" stateProvince="Madre de Dios Region">
• 
<location id="8EC48C86FFC8FF94FACAF9FDA855EC25" LSID="urn:lsid:plazi:treatment:03B26B4BFFC8FF96FE24FCECAF1DED90:8EC48C86FFC8FF94FACAF9FDA855EC25" box="[1299,1394,1602,1628]" country="United States of America" county="Valle del Pilcopata" latitude="-13.166667" longLatPrecision="1291" longitude="-71.416664" municipality="Foster" name="Mandor" pageId="109" pageNumber="110" stateProvince="Madre de Dios Region">Mandor</location>
, 
<location id="8EC48C86FFC8FF94FF64F9D9AC1AECF9" LSID="urn:lsid:plazi:treatment:03B26B4BFFC8FF96FE24FCECAF1DED90:8EC48C86FFC8FF94FF64F9D9AC1AECF9" box="[189,317,1638,1664]" country="United States of America" county="Valle del Pilcopata" latitude="-13.166667" longLatPrecision="1291" longitude="-71.416664" municipality="Foster" name="Marcapata" pageId="109" pageNumber="110" stateProvince="Madre de Dios Region">Marcapata</location>
; [
<geoCoordinate id="EE2FBC9AFFC8FF94FE88F9DBAC9EECF9" box="[337,441,1636,1664]" degrees="13" direction="south" minutes="21" orientation="latitude" pageId="109" pageNumber="110" precision="925" value="-13.35">13°21′ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFFC8FF94FE1FF9DBAF1CECF9" box="[454,571,1636,1664]" degrees="70" direction="west" minutes="52" orientation="longitude" pageId="109" pageNumber="110" precision="925" value="-70.86667">70°52′ W</geoCoordinate>
]; 
<elevation id="00363D6EFFC8FF94FD96F9D9AFCFECF9" box="[591,744,1638,1664]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="109" pageNumber="110" unit="m" value="1100.0">
<quantity id="4CE377B8FFC8FF94FD96F9D9AF8FECF9" box="[591,680,1638,1664]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="109" pageNumber="110" unit="m" value="1100.0">1100 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFFC8FF94FD2FF9DAAEADECF9" box="[758,906,1637,1664]" pageId="109" pageNumber="110" value="1945-07-15">
<collectingDate id="EFE10575FFC8FF94FD2FF9DAAEADECF9" box="[758,906,1637,1664]" pageId="109" pageNumber="110" value="1945-07-15">15 Jul. 1945</collectingDate>
</date>
; 
<emphasis id="B96F064FFFC8FF94FC4FF9D9A9FDECF9" box="[918,1242,1638,1664]" italics="true" pageId="109" pageNumber="110">
<collectorName id="26EEBF8BFFC8FF94FC4FF9D9A9B0ECF9" box="[918,1175,1638,1664]" pageId="109" pageNumber="110">J.C. Vargas Calderón</collectorName>
5242
</emphasis>
; 
<collectionCode id="ED0A4298FFC8FF94FB33F9D9A802ECF9" box="[1258,1317,1638,1664]" collectionName="CUZ" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14364" lsid="urn:lsid:biocol.org:col:14364" name="Universidad Nacional San Antonio Abad del Cusco" pageId="109" pageNumber="110" type="Herbarium">CUZ</collectionCode>
</materialsCitation>
<materialsCitation id="3B73D000FFC8FF94FAF5F9D9A91BECBE" collectingDate="2004-10-23" collectingDateMax="2004-10-23" collectingDateMin="1945-07-15" collectionCode="MO, US" collectorName="L. Valenzuela &amp; E. Suclli &amp; V. Chama &amp; J. La Torre" country="United States of America" county="Dist. Marcapata" elevation="615" latitude="-13.166667" location="Quincemil" longLatPrecision="1291" longitude="-71.416664" municipality="Quincemil" pageId="109" pageNumber="110" specimenCode="MO-1664172, US00900536" specimenCount="1" stateProvince="Madre de Dios Region">
• 
<collectingCounty id="62C5A2D1FFC8FF94FAE7F9D9AC1BECDD" pageId="109" pageNumber="110">Dist. Marcapata</collectingCounty>
, 
<collectingMunicipality id="6BC04027FFC8FF94FE93F936ACEEECDA" box="[330,457,1673,1699]" pageId="109" pageNumber="110">Quincemil</collectingMunicipality>
, 
<geoCoordinate id="EE2FBC9AFFC8FF94FE0EF937AF4FECDA" box="[471,616,1672,1700]" degrees="13" direction="south" minutes="42" orientation="latitude" pageId="109" pageNumber="110" precision="15" seconds="46" value="-13.712777">13°42′46″ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFFC8FF94FDAEF937AE32ECDD" box="[631,789,1672,1700]" degrees="70" direction="west" minutes="59" orientation="longitude" pageId="109" pageNumber="110" precision="15" seconds="40" value="-70.99444">70°59′40″ W</geoCoordinate>
; 
<elevation id="00363D6EFFC8FF94FCFAF936AECBECDA" box="[803,1004,1673,1700]" metricMagnitude="2" metricUnit="m" metricValue="6.15" metricValueMax="8.0" metricValueMin="4.3" pageId="109" pageNumber="110" unit="m" value="615.0" valueMax="800.0" valueMin="430.0">
<quantity id="4CE377B8FFC8FF94FCFAF936AE8DECDD" box="[803,938,1673,1700]" metricMagnitude="2" metricUnit="m" metricValue="6.15" metricValueMax="8.0" metricValueMin="4.3" pageId="109" pageNumber="110" unit="m" value="615.0" valueMax="800.0" valueMin="430.0">430–800 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFFC8FF94FC22F936A9BCECDA" box="[1019,1179,1673,1699]" pageId="109" pageNumber="110" value="2004-10-23">
<collectingDate id="EFE10575FFC8FF94FC22F936A9BCECDA" box="[1019,1179,1673,1699]" pageId="109" pageNumber="110" value="2004-10-23">23 Oct. 2004</collectingDate>
</date>
; 
<emphasis id="B96F064FFFC8FF94FB73F935AF70ECBF" italics="true" pageId="109" pageNumber="110">
<collectorName id="26EEBF8BFFC8FF94FB73F935A868ECDA" box="[1194,1359,1673,1699]" pageId="109" pageNumber="110">L. Valenzuela</collectorName>
, 
<collectorName id="26EEBF8BFFC8FF94FA84F935AC2EECBF" pageId="109" pageNumber="110">E. Suclli</collectorName>
,. 
<collectorName id="26EEBF8BFFC8FF94FEC1F912ACABECBF" box="[280,396,1708,1734]" pageId="109" pageNumber="110">V. Chama</collectorName>
, 
<collectorName id="26EEBF8BFFC8FF94FE4EF912AF3FECBF" box="[407,536,1709,1734]" pageId="109" pageNumber="110">J. La Torre</collectorName>
4267
</emphasis>
; 
<collectionCode id="ED0A4298FFC8FF94FDBCF912AFB3ECBE" box="[613,660,1709,1735]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="109" pageNumber="110" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FFC8FF94FD7DF913AE63ECBF" box="[676,836,1708,1734]" pageId="109" pageNumber="110">MO-1664172</specimenCode>
], 
<collectionCode id="ED0A4298FFC8FF94FC82F912AEA7ECBE" box="[859,896,1709,1735]" pageId="109" pageNumber="110">US</collectionCode>
[
<specimenCode id="DBBD7226FFC8FF94FC49F913A90CECBE" box="[912,1067,1708,1735]" pageId="109" pageNumber="110">US00900536</specimenCode>
].
</materialsCitation>
<materialsCitation id="3B73D000FFC8FF94FB9BF912AFFFED77" collectingDate="1964-06" collectingDateMax="2004-10-23" collectingDateMin="1945-07-15" collectionCode="MO" collectorName="J. C. Vargas Calderon" country="United States of America" county="Valle del Pilcopata" elevation="670" latitude="-13.166667" location="izquierda Madre de Dios" longLatPrecision="1291" longitude="-71.416664" municipality="Prov. Manu" pageId="109" pageNumber="110" specimenCode="MO-1835898" specimenCount="1" stateProvince="Madre de Dios Region">
– 
<emphasis id="B96F064FFFC8FF94FB8CF912AC76EC93" bold="true" pageId="109" pageNumber="110">
<collectingRegion id="49DF14BFFFC8FF94FB8CF912A848ECBF" box="[1109,1391,1708,1735]" country="United States of America" name="Madre de Dios Region" pageId="109" pageNumber="110">Madre de Dios Region</collectingRegion>
: 
<collectingMunicipality id="6BC04027FFC8FF94FF64F96FAC76EC93" box="[189,337,1744,1770]" pageId="109" pageNumber="110">Prov. Manu</collectingMunicipality>
</emphasis>
• 
<location id="8EC48C86FFC8FF94FEB1F96FAFADEC93" LSID="urn:lsid:plazi:treatment:03B26B4BFFC8FF96FE24FCECAF1DED90:8EC48C86FFC8FF94FEB1F96FAFADEC93" box="[360,650,1744,1770]" country="United States of America" county="Valle del Pilcopata" latitude="-13.166667" longLatPrecision="1291" longitude="-71.416664" municipality="Prov. Manu" name="izquierda Madre de Dios" pageId="109" pageNumber="110" stateProvince="Madre de Dios Region">izquierda Madre de Dios</location>
, 
<locationDeviation id="9A16C37FFFC8FF94FD4CF96EAE00EC92" box="[661,807,1745,1771]" pageId="109" pageNumber="110">cerca pongo</locationDeviation>
; [
<geoCoordinate id="EE2FBC9AFFC8FF94FCE0F971AE87EC93" box="[825,928,1742,1770]" pageId="109" pageNumber="110">12°62′ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFFC8FF94FC75F971A907EC92" box="[940,1056,1742,1771]" degrees="71" direction="west" minutes="22" orientation="longitude" pageId="109" pageNumber="110" precision="925" value="-71.36667">71°22′ W</geoCoordinate>
]; 
<elevation id="00363D6EFFC8FF94FBEAF96FA99CEC93" box="[1075,1211,1744,1771]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="109" pageNumber="110" unit="m" value="670.0">
<quantity id="4CE377B8FFC8FF94FBEAF96FA95BEC92" box="[1075,1148,1744,1771]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="109" pageNumber="110" unit="m" value="670.0">670 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFFC8FF94FB1EF96EA81BEC93" box="[1223,1340,1744,1771]" pageId="109" pageNumber="110" value="1964-06">
<collectingDate id="EFE10575FFC8FF94FB1EF96EA81BEC93" box="[1223,1340,1744,1771]" pageId="109" pageNumber="110" value="1964-06">Jun. 1964</collectingDate>
</date>
; 
<emphasis id="B96F064FFFC8FF94FA9EF96FACF0ED74" italics="true" pageId="109" pageNumber="110">
<collectorName id="26EEBF8BFFC8FF94FA9EF96FACA1ED74" pageId="109" pageNumber="110">J.C. Vargas Calderón</collectorName>
15526
</emphasis>
; 
<collectionCode id="ED0A4298FFC8FF94FE3EF94BAF31ED77" box="[487,534,1780,1806]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="109" pageNumber="110" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FFC8FF94FDFEF94CAFE0ED77" box="[551,711,1779,1806]" pageId="109" pageNumber="110">MO-1835898</specimenCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFC8FF96FF64F88AAC61E99F" lastPageId="111" lastPageNumber="112" pageId="109" pageNumber="110" type="description">
<paragraph id="8BA4DA5DFFC8FF94FF64F88AAC68ED36" blockId="109.[189,335,1845,1871]" box="[189,335,1845,1871]" pageId="109" pageNumber="110">
<heading id="D0EC6D31FFC8FF94FF64F88AAC68ED36" bold="true" box="[189,335,1845,1871]" fontSize="11" level="3" pageId="109" pageNumber="110" reason="3">
<emphasis id="B96F064FFFC8FF94FF64F88AAC68ED36" bold="true" box="[189,335,1845,1871]" pageId="109" pageNumber="110">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFC8FF96FF64F8DBAC61E99F" blockId="109.[187,1399,1892,2025]" lastBlockId="111.[187,1400,265,998]" lastPageId="111" lastPageNumber="112" pageId="109" pageNumber="110">
Caulescent herb, to 
<quantity id="4CE377B8FFC8FF94FE77F8DBACF9ED07" box="[430,478,1892,1918]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="109" pageNumber="110" unit="m" value="1.0">1 m</quantity>
high. 
<emphasis id="B96F064FFFC8FF94FDF4F8DBAF41ED07" box="[557,614,1892,1918]" italics="true" pageId="109" pageNumber="110">Stem</emphasis>
erect, rarely branching; internodes to 
<quantity id="4CE377B8FFC8FF94FBEEF8DBA953ED07" box="[1079,1140,1892,1918]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="109" pageNumber="110" unit="cm" value="7.0">7 cm</quantity>
long, to 
<quantity id="4CE377B8FFC8FF94FB3DF8DBA80CED07" box="[1252,1323,1892,1918]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="109" pageNumber="110" unit="mm" value="6.0">6 mm</quantity>
thick, succulent, pale green to pink, glabrous. 
<emphasis id="B96F064FFFC8FF94FD5BF838AFC7EDD8" box="[642,736,1927,1953]" italics="true" pageId="109" pageNumber="110">Stipules</emphasis>
late deciduous to persistent, lanceolate, 12–35 × 
<quantity id="4CE377B8FFC8FF94FAC9F838A855EDDB" box="[1296,1394,1927,1954]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="109" pageNumber="110" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
, apex acute, translucent, pale green, glabrous, margin entire, aciliate. 
<emphasis id="B96F064FFFC8FF94FC21F813A96DEDBC" box="[1016,1098,1964,1989]" italics="true" pageId="109" pageNumber="110">Leaves</emphasis>
4–8, alternate, basifixed; petiole 
<quantity id="4CE377B8FFC8FF94FECCF871AC49ED90" box="[277,366,1998,2025]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="9.0" metricValueMin="2.0" pageId="109" pageNumber="110" unit="cm" value="5.5" valueMax="9.0" valueMin="2.0">2–9 cm</quantity>
long, pale red, glabrous; blade strongly asymmetric, lanceolate to ovate, to 18 × 
<quantity id="4CE377B8FFC8FF94FAEDF871A855ED90" box="[1332,1394,1998,2025]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="109" pageNumber="110" unit="cm" value="9.0">9 cm</quantity>
, succulent, apex acuminate, base rounded, margin entire to serrulate, aciliate to ciliate, upper surface dark, glossy green, glabrous, lower surface green to deep red, glabrous, veins palmate but with one primary vein, 5–7 veined from the base, with 1–3 secondary veins on the larger side, 1–3 on the smaller side. 
<emphasis id="B96F064FFFCAFF96FF66FEC0AC40EBE0" box="[191,359,383,409]" italics="true" pageId="111" pageNumber="112">Inflorescences</emphasis>
1–3, bisexual, axillary, erect, cymose, branching up to 5 times, bearing up to 64 staminate flowers and 16 pistillate flowers, protandrous; peduncle to 
<quantity id="4CE377B8FFCAFF96FCB7FE19AE9FEBB9" box="[878,952,422,448]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="111" pageNumber="112" unit="cm" value="12.0">12 cm</quantity>
long, red, glabrous, bracts deciduous, ovate, 4–15 × 
<quantity id="4CE377B8FFCAFF96FEB5FE72ACC6EB91" box="[364,481,461,488]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="111" pageNumber="112" unit="mm" value="6.5" valueMax="10.0" valueMin="3.0">3–10 mm</quantity>
, opaque, brown, glabrous, apex rounded to acute, margin entire to serrulate, aciliate to ciliate. 
<emphasis id="B96F064FFFCAFF96FE4BFE4AAF42E877" box="[402,613,500,527]" italics="true" pageId="111" pageNumber="112">Staminate flowers</emphasis>
: pedicels to 
<quantity id="4CE377B8FFCAFF96FD26FE4BAE63E876" box="[767,836,500,527]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="111" pageNumber="112" unit="mm" value="5.0">5 mm</quantity>
long, glabrous; tepals 2, projecting, lanceolate, 5–18 × 
<quantity id="4CE377B8FFCAFF96FEC1FDA3AC5BE84F" box="[280,380,540,566]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="111" pageNumber="112" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 mm</quantity>
, apex acuminate, bright red or orange, glabrous, margin entire, aciliate; stamens ca 30, projecting, yellow, filaments ca 
<quantity id="4CE377B8FFCAFF96FDE2FDFCAFB1E824" box="[571,662,579,605]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="111" pageNumber="112" unit="mm" value="0.3">0.3 mm</quantity>
long, free, anthers linear, 1.5–2 × 
<quantity id="4CE377B8FFCAFF96FBEFFDFCA9B2E824" box="[1078,1173,579,606]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="111" pageNumber="112" unit="mm" value="0.5">0.5 mm</quantity>
, dehiscing through lateral slits, connectives extended, symmetrically basifixed. 
<emphasis id="B96F064FFFCAFF96FC59FDD5A96EE8FD" box="[896,1097,618,644]" italics="true" pageId="111" pageNumber="112">Pistillate flowers</emphasis>
: pedicels to 
<quantity id="4CE377B8FFCAFF96FB39FDD5A814E8FC" box="[1248,1331,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="111" pageNumber="112" unit="mm" value="15.0">15 mm</quantity>
long; bracteoles 3, positioned directly beneath the ovary, ovate, 5–5.5 × 
<quantity id="4CE377B8FFCAFF96FC16FD2EA947E8D5" box="[975,1120,657,684]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="111" pageNumber="112" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.5–5.5 mm</quantity>
, apex rounded, opaque, bright red or orange, glabrous, margin lacerate, ciliate; tepals 5, subequal, persistent in fruit, projecting, lanceolate, 5–18 × 
<quantity id="4CE377B8FFCAFF96FE7AFD5FAF2EE883" box="[419,521,736,762]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="9.0" metricValueMin="2.0" pageId="111" pageNumber="112" unit="mm" value="5.5" valueMax="9.0" valueMin="2.0">2–9 mm</quantity>
, apex acute, bright red or orange, glabrous, margin entire, aciliate; ovary body obovoid, 4–7 × 
<quantity id="4CE377B8FFCAFF96FE1CFCB8AF0BE95B" box="[453,556,775,802]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="111" pageNumber="112" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
, bright red or orange, glabrous, unequally 3-winged, wings triangular, largest 5–15 × 
<quantity id="4CE377B8FFCAFF96FEACFC91ACCDE930" box="[373,490,814,841]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="11.0" metricValueMin="4.0" pageId="111" pageNumber="112" unit="mm" value="7.5" valueMax="11.0" valueMin="4.0">4–11 mm</quantity>
, smallest two wings 3–8 × 
<quantity id="4CE377B8FFCAFF96FCE2FC91AE85E930" box="[827,930,814,841]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="111" pageNumber="112" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
; 3-locular, placentae branches divided, bearing ovules on both surfaces; styles 3, yellow, free, 
<quantity id="4CE377B8FFCAFF96FC94FCE9AE97E909" box="[845,944,854,880]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="111" pageNumber="112" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long, once-divided, stigmatic papillae in a spirally-twisted band. 
<emphasis id="B96F064FFFCAFF96FE26FCC1AFE5E9EE" box="[511,706,893,919]" italics="true" pageId="111" pageNumber="112">Fruiting pedicel</emphasis>
to 
<quantity id="4CE377B8FFCAFF96FD34FCC2AE65E9EE" box="[749,834,893,920]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="111" pageNumber="112" unit="mm" value="25.0">25 mm</quantity>
long. 
<emphasis id="B96F064FFFCAFF96FC48FCC1A936E9EE" box="[913,1041,893,919]" italics="true" pageId="111" pageNumber="112">Fruit body</emphasis>
ovoid, to 12 × 
<quantity id="4CE377B8FFCAFF96FB0BFCC2A83BE9EE" box="[1234,1308,893,919]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="111" pageNumber="112" unit="mm" value="6.0">6 mm</quantity>
, drying brown, wings same shape as in ovary, the largest expanding to 18 × 
<quantity id="4CE377B8FFCAFF96FC3FFC1BA91AE9C7" box="[998,1085,932,958]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="111" pageNumber="112" unit="mm" value="30.0">30 mm</quantity>
, the smallest expanding to 12 × 
<quantity id="4CE377B8FFCAFF96FF23FC74AC66E99F" box="[250,321,971,998]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="111" pageNumber="112" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF648AD5FFCBFF97FF64F944AECAED91" ID-DOI="http://doi.org/10.5281/zenodo.8178377" ID-Zenodo-Dep="8178377" httpUri="https://zenodo.org/record/8178377/files/figure.png" pageId="110" pageNumber="111" startId="110.[189,232,1787,1813]" targetBox="[272,1315,265,1740]" targetPageId="110">
<paragraph id="8BA4DA5DFFCBFF97FF64F944AECAED91" blockId="110.[188,1399,1787,2025]" pageId="110" pageNumber="111">
<emphasis id="B96F064FFFCBFF97FF64F944AC3EED6C" bold="true" box="[189,281,1787,1813]" pageId="110" pageNumber="111">Fig. 38.</emphasis>
<taxonomicName id="4C1BA1DEFFCBFF97FEC6F943AED2ED6C" authority="L. B. Sm. &amp; B. G. Schub. A. Habit. B. Leaf" box="[287,1013,1787,1814]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="110" pageNumber="111" phylum="Tracheophyta" rank="species" species="stenotepala">
<emphasis id="B96F064FFFCBFF97FEC6F943AF36ED6C" box="[287,529,1787,1813]" italics="true" pageId="110" pageNumber="111">Begonia stenotepala</emphasis>
L.B.Sm. &amp; B.G.Schub. 
<emphasis id="B96F064FFFCBFF97FCF0F944AE19ED6C" bold="true" box="[809,830,1787,1813]" pageId="110" pageNumber="111">A</emphasis>
. Habit. 
<emphasis id="B96F064FFFCBFF97FC43F944AE89ED6C" bold="true" box="[922,942,1787,1813]" pageId="110" pageNumber="111">B</emphasis>
. Leaf
</taxonomicName>
, adaxial surface. 
<emphasis id="B96F064FFFCBFF97FB62F944A9F7ED6C" bold="true" box="[1211,1232,1787,1813]" pageId="110" pageNumber="111">C</emphasis>
. Leaf, abaxial surface. 
<emphasis id="B96F064FFFCBFF97FEFAF8A1AC1FED41" bold="true" box="[291,312,1822,1848]" pageId="110" pageNumber="111">D</emphasis>
. Stipule. 
<emphasis id="B96F064FFFCBFF97FE76F8A1ACE4ED41" bold="true" box="[431,451,1822,1848]" pageId="110" pageNumber="111">E</emphasis>
. Staminate flower, front view. 
<emphasis id="B96F064FFFCBFF97FCE2F8A1AE6AED41" bold="true" box="[827,845,1822,1848]" pageId="110" pageNumber="111">F</emphasis>
. Staminate flower, side view. 
<emphasis id="B96F064FFFCBFF97FB62F8A1A9F5ED41" bold="true" box="[1211,1234,1822,1848]" pageId="110" pageNumber="111">G</emphasis>
. Top tepal of the staminate flower. 
<emphasis id="B96F064FFFCBFF97FE66F8FDACF1ED25" bold="true" box="[447,470,1858,1884]" pageId="110" pageNumber="111">H</emphasis>
. Bottom tepal of the staminate flower. 
<emphasis id="B96F064FFFCBFF97FC69F8FDAE9CED25" bold="true" box="[944,955,1858,1884]" pageId="110" pageNumber="111">I</emphasis>
. Androecium, side view. 
<emphasis id="B96F064FFFCBFF97FB2BF8FDA9D9ED25" bold="true" box="[1266,1278,1858,1884]" pageId="110" pageNumber="111">J</emphasis>
. Pistillate flower, front view. 
<emphasis id="B96F064FFFCBFF97FE43F8DAAC96ED06" bold="true" box="[410,433,1893,1919]" pageId="110" pageNumber="111">K</emphasis>
. Pistillate flower, side view. 
<emphasis id="B96F064FFFCBFF97FD26F8DAAE34ED06" bold="true" box="[767,787,1893,1919]" pageId="110" pageNumber="111">L</emphasis>
. Pistils, side view. 
<emphasis id="B96F064FFFCBFF97FC2BF8DAA929ED06" bold="true" box="[1010,1038,1893,1919]" pageId="110" pageNumber="111">M</emphasis>
. Smallest tepal of the pistillate flower. 
<emphasis id="B96F064FFFCBFF97FECFF837AC0CEDDB" bold="true" box="[278,299,1928,1954]" pageId="110" pageNumber="111">N</emphasis>
. Largest tepal of the pistillate flower. 
<emphasis id="B96F064FFFCBFF97FD29F837AE20EDDB" bold="true" box="[752,775,1928,1954]" pageId="110" pageNumber="111">O</emphasis>
. Bracteole. All photographs taken by D.A. Purvis &amp; P.W. Moonlight in the living collections of the Royal Botanic Garden Edinburgh (Accession 20141048, grown from seeds collected as part of 
<emphasis id="B96F064FFFCBFF97FDA5F870AEFBED91" box="[636,988,1998,2025]" italics="true" pageId="110" pageNumber="111">P.W. Moonlight &amp; A. Daza 83</emphasis>
).
</paragraph>
</caption>
<subSubSection id="C30189D6FFCAFF96FF64FBAEA84AEF30" pageId="111" pageNumber="112" type="conservation">
<paragraph id="8BA4DA5DFFCAFF96FF64FBAEAF4AEE52" blockId="111.[189,621,1040,1067]" box="[189,621,1040,1067]" pageId="111" pageNumber="112">
<heading id="D0EC6D31FFCAFF96FF64FBAEAF4AEE52" bold="true" box="[189,621,1040,1067]" fontSize="11" level="3" pageId="111" pageNumber="112" reason="3">
<emphasis id="B96F064FFFCAFF96FF64FBAEAF4AEE52" bold="true" box="[189,621,1040,1067]" pageId="111" pageNumber="112">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFCAFF96FF64FBFCA84AEF30" blockId="111.[189,1400,1091,1353]" pageId="111" pageNumber="112">
Previously assessed as Data Deficient (DD) by 
<bibRefCitation id="EF8AA7ACFFCAFF96FCDDFBFCA967EE24" author="Leon &amp; Monsalve" box="[772,1088,1091,1117]" firstAuthor="Leon" pageId="111" pageNumber="112" pagination="165 - 170" refId="ref229226" refString="Leon B. &amp; Monsalve C. 2006. Begoniaceae endemicas del Peru. Revista Peruana de Biologia 13 (2): 165 - 170. https: // doi. org / 10.15381 / rpb. v 13 i 2.1810" type="journal article" year="2006">
<collectingRegion id="49DF14BFFFCAFF96FCDDFBFCAE67EE24" box="[772,832,1091,1117]" country="Nicaragua" name="Leon" pageId="111" pageNumber="112">León</collectingRegion>
&amp; Monsalve (2006)
</bibRefCitation>
. The EOO of the species is &lt;
<quantity id="4CE377B8FFCAFF96FF2CFBD5AC79EEFD" box="[245,350,1130,1157]" metricMagnitude="6" metricUnit="m" metricValue="5.0" pageId="111" pageNumber="112" unit="km" value="5000.0">5000 km</quantity>
<superScript id="7C6E7715FFCAFF96FE86FBD5AC4FEE01" attach="right" box="[351,360,1130,1144]" fontSize="6" pageId="111" pageNumber="112">2</superScript>
, which is sufficiently small for the species to qualify as endangered. Within its range, however, 
<taxonomicName id="4C1BA1DEFFCAFF96FEE8FB2DACFBEED5" box="[305,476,1170,1196]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="111" pageNumber="112" phylum="Tracheophyta" rank="species" species="stenotepala">
<emphasis id="B96F064FFFCAFF96FEE8FB2DACFBEED5" box="[305,476,1170,1196]" italics="true" pageId="111" pageNumber="112">B. stenotepala</emphasis>
</taxonomicName>
is common and conspicuous, this species and frequently collected, especially on the borders of the road from Challabamba to Pilcopata, which forms one of the borders of Manu National Park. We estimate from fieldwork in 2014 that the population of 
<taxonomicName id="4C1BA1DEFFCAFF96FBF2FB5EA9F1EE83" box="[1067,1238,1248,1274]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="111" pageNumber="112" phylum="Tracheophyta" rank="species" species="stenotepala">
<emphasis id="B96F064FFFCAFF96FBF2FB5EA9F1EE83" box="[1067,1238,1248,1274]" italics="true" pageId="111" pageNumber="112">B. stenotepala</emphasis>
</taxonomicName>
here exceeds&gt; 50 k individuals and suspect similar populations are found within the species’ range, including within Manu National Park, and surrounding, intact forests. We assess 
<taxonomicName id="4C1BA1DEFFCAFF96FC70FA90A974EF30" box="[937,1107,1327,1353]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="111" pageNumber="112" phylum="Tracheophyta" rank="species" species="stenotepala">
<emphasis id="B96F064FFFCAFF96FC70FA90A974EF30" box="[937,1107,1327,1353]" italics="true" pageId="111" pageNumber="112">B. stenotepala</emphasis>
</taxonomicName>
as Least Concern (LC).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFCAFF96FF64FACBA9CDECAA" pageId="111" pageNumber="112" type="discussion">
<paragraph id="8BA4DA5DFFCAFF96FF64FACBAC97EFF7" blockId="111.[189,432,1396,1422]" box="[189,432,1396,1422]" pageId="111" pageNumber="112">
<heading id="D0EC6D31FFCAFF96FF64FACBAC97EFF7" bold="true" box="[189,432,1396,1422]" fontSize="11" level="3" pageId="111" pageNumber="112" reason="3">
<emphasis id="B96F064FFFCAFF96FF64FACBAC97EFF7" bold="true" box="[189,432,1396,1422]" pageId="111" pageNumber="112">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFCAFF96FF64FA18A9CDECAA" blockId="111.[188,1399,1447,1748]" pageId="111" pageNumber="112">
<taxonomicName id="4C1BA1DEFFCAFF96FF64FA18AC88EFB8" box="[189,431,1447,1473]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="111" pageNumber="112" phylum="Tracheophyta" rank="species" species="stenotepala">
<emphasis id="B96F064FFFCAFF96FF64FA18AC88EFB8" box="[189,431,1447,1473]" italics="true" pageId="111" pageNumber="112">Begonia stenotepala</emphasis>
</taxonomicName>
is trivial to identify when in flower on account of its staminate and pistillate flowers with vivid red tepals with acute apices. Similar coloured and shaped tepals are found only in distantly related species with completely different habits (e.g., 
<taxonomicName id="4C1BA1DEFFCAFF96FCE0FA49AEFFEC76" box="[825,984,1525,1551]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="111" pageNumber="112" phylum="Tracheophyta" rank="species" species="polypetala">
<emphasis id="B96F064FFFCAFF96FCE0FA49AEFFEC76" box="[825,984,1525,1551]" italics="true" pageId="111" pageNumber="112">B. polypetala</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C1BA1DEFFCAFF96FC31FA49A987EC76" box="[1000,1184,1526,1551]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="111" pageNumber="112" phylum="Tracheophyta" rank="species" species="rossmanniae">
<emphasis id="B96F064FFFCAFF96FC31FA49A987EC76" box="[1000,1184,1526,1551]" italics="true" pageId="111" pageNumber="112">B. rossmanniae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C1BA1DEFFCAFF96FB68FA49A842EC76" box="[1201,1381,1525,1551]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="111" pageNumber="112" phylum="Tracheophyta" rank="species" species="monadelpha">
<emphasis id="B96F064FFFCAFF96FB68FA49A842EC76" box="[1201,1381,1525,1551]" italics="true" pageId="111" pageNumber="112">B. monadelpha</emphasis>
</taxonomicName>
). When sterile, 
<taxonomicName id="4C1BA1DEFFCAFF96FEBFF9A2AF34EC4F" box="[358,531,1564,1590]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="111" pageNumber="112" phylum="Tracheophyta" rank="species" species="stenotepala">
<emphasis id="B96F064FFFCAFF96FEBFF9A2AF34EC4F" box="[358,531,1564,1590]" italics="true" pageId="111" pageNumber="112">B. stenotepala</emphasis>
</taxonomicName>
is recognised as an erect herb with transversely ovate leaves and large, persistent stipules. In this respect, it is very similar to 
<taxonomicName id="4C1BA1DEFFCAFF96FC84F9FBA93AEC27" box="[861,1053,1604,1630]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="111" pageNumber="112" phylum="Tracheophyta" rank="species" species="subspinulosa">
<emphasis id="B96F064FFFCAFF96FC84F9FBA93AEC27" box="[861,1053,1604,1630]" italics="true" pageId="111" pageNumber="112">B. subspinulosa</emphasis>
</taxonomicName>
, but the two species can be distinguished by their leaf margins. 
<taxonomicName id="4C1BA1DEFFCAFF96FD84F9D3AE68ECFC" box="[605,847,1643,1669]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="111" pageNumber="112" phylum="Tracheophyta" rank="species" species="stenotepala">
<emphasis id="B96F064FFFCAFF96FD84F9D3AE68ECFC" box="[605,847,1643,1669]" italics="true" pageId="111" pageNumber="112">Begonia stenotepala</emphasis>
</taxonomicName>
has entire to serrulate margins, which are often aciliate and do not look spinescent. In contrast, the leaf margins of 
<taxonomicName id="4C1BA1DEFFCAFF96FC2EF92CA990ECD5" box="[1015,1207,1682,1708]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="111" pageNumber="112" phylum="Tracheophyta" rank="species" species="subspinulosa">
<emphasis id="B96F064FFFCAFF96FC2EF92CA990ECD5" box="[1015,1207,1682,1708]" italics="true" pageId="111" pageNumber="112">B. subspinulosa</emphasis>
</taxonomicName>
are serrulate to serrate, always ciliate, with the serrations and ciliate margins appearing superficially sharp.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FFCAFF96FF64F940AF1DED90" pageId="111" pageNumber="112" type="distribution">
<paragraph id="8BA4DA5DFFCAFF96FF64F940ACD2ED60" blockId="111.[189,501,1791,1817]" box="[189,501,1791,1817]" pageId="111" pageNumber="112">
<heading id="D0EC6D31FFCAFF96FF64F940ACD2ED60" bold="true" box="[189,501,1791,1817]" fontSize="11" level="3" pageId="111" pageNumber="112" reason="3">
<emphasis id="B96F064FFFCAFF96FF64F940ACD2ED60" bold="true" box="[189,501,1791,1817]" pageId="111" pageNumber="112">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFFCAFF96FF64F88EAF1DED90" blockId="111.[188,1399,1841,2025]" pageId="111" pageNumber="112">
Endemic to 
<collectingCountry id="F30C9ACDFFCAFF96FE94F88DACA4ED35" box="[333,387,1842,1868]" name="Peru" pageId="111" pageNumber="112">Peru</collectingCountry>
and known from 
<collectingRegion id="49DF14BFFFCAFF96FD81F88EAF84ED32" box="[600,675,1841,1867]" country="Peru" name="Cusco" pageId="111" pageNumber="112">Cuzco</collectingRegion>
and 
<collectingRegion id="49DF14BFFFCAFF96FD06F88EAEA8ED32" box="[735,911,1841,1867]" country="Peru" name="Madre de Dios" pageId="111" pageNumber="112">Madre de Dios</collectingRegion>
Regions (
<figureCitation id="1320C6D8FFCAFF96FBD3F88EA95FED35" box="[1034,1144,1841,1868]" captionStart="Fig" captionStartId="99.[189,232,1822,1848]" captionTargetBox="[194,1391,556,1774]" captionTargetId="figure-78@99.[189,1398,546,1782]" captionTargetPageId="99" captionText="Fig. 35. Distribution of the stenotepala group of Begonia sect. Cyathocnemis (Klotzsch) A.DC. in Peru and surrounding countries. A. B. alto-peruviana A.DC. (red). B. B. brevicordata L.B.Sm. &amp; B.G.Schub. (red), B. deltoidesMoonlight sp. nov. (blue), and B. lucifuga Irmsch (green). C. B. longitepalaMoonlight sp. nov. (red) and B. subspinulosa Irmsch. (blue). D. B. stenotepala L.B.Sm. &amp; B.G.Schub. (red). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178371" httpUri="https://zenodo.org/record/8178371/files/figure.png" pageId="111" pageNumber="112">Fig. 35D</figureCitation>
). Found in lower and rarely middle montane Forest at an elevation of 
<quantity id="4CE377B8FFCAFF96FD2EF8E7AEADED0A" box="[759,906,1880,1907]" metricMagnitude="3" metricUnit="m" metricValue="1.275" metricValueMax="1.88" metricValueMin="0.67" pageId="111" pageNumber="112" unit="m" value="1275.0" valueMax="1880.0" valueMin="670.0">670–1880 m</quantity>
a.s.l. The vivid red flowers of the species and its projecting tepals suggest it may be hummingbird pollinated, but no pollinator records are known. 
<taxonomicName id="4C1BA1DEFFCAFF96FF65F817AC94EDB8" box="[188,435,1959,1985]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="111" pageNumber="112" phylum="Tracheophyta" rank="species" species="stenotepala">
<emphasis id="B96F064FFFCAFF96FF65F817AC94EDB8" box="[188,435,1959,1985]" italics="true" pageId="111" pageNumber="112">Begonia stenotepala</emphasis>
</taxonomicName>
has been collected in flower from April to December with a peak in July and August, which is the dry season.
</paragraph>
</subSubSection>
</treatment>
</document>