treatments-xml/data/03/B2/6B/03B26B4BFF79FF26FDAAFC2DAE09EF2B.xml
2024-06-21 12:22:17 +02:00

893 lines
93 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="38D1B516597A598FB3C4D01467915C4C" ID-DOI="https://doi.org/10.5852/ejt.2023.881.2175" ID-ISSN="2118-9773" ID-Zenodo-Dep="8178280" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690201937279" checkinUser="felipe" docAuthor="Moonlight, Peter. W., Jara-Muñoz, Orlando A., Purvis, David A., Delves, Jay, Allen, Josh P. &amp; Reynel, Carlos" docDate="2023" docId="03B26B4BFF79FF26FDAAFC2DAE09EF2B" docLanguage="en" docName="ejt-881_1-334.imf" docOrigin="European Journal of Taxonomy 881" docSource="http://dx.doi.org/10.5852/ejt.2023.881.2175" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia wollnyi Herzog" docType="treatment" docVersion="7" lastPageNumber="224" masterDocId="FF8B1333FFA5FFF9FFD9FFBFAD27EA79" masterDocTitle="The genus Begonia (Begoniaceae) in Peru" masterLastPageNumber="334" masterPageNumber="1" pageNumber="221" updateTime="1707123162844" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="8190125B93AC53B9A48A45930FC5ABFC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5551E1F3F013EF28D424AF93637A1BD2">
<mods:title id="4FD71290352336DB70D79B23BC5A5701">The genus Begonia (Begoniaceae) in Peru</mods:title>
</mods:titleInfo>
<mods:name id="DAA8208DC64AAE16120BD1A01351764D" type="personal">
<mods:role id="E1D2B38A492CEFD21051664032CC9C6F">
<mods:roleTerm id="FC1950B96854D2024BD49DB103CA521A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B8F768A7BAD6A8265526529A24FE0E3">Moonlight, Peter. W.</mods:namePart>
<mods:affiliation id="2BF6A1A0C975908C022DAD6B33EEBDBB">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK. &amp; Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland.</mods:affiliation>
<mods:nameIdentifier id="5C7A0E3504881B05901D458A5178CCC7" type="email">moonligp@tcd.ie</mods:nameIdentifier>
</mods:name>
<mods:name id="B28D8461A71A290D21A4C0713302E9BB" type="personal">
<mods:role id="0C0D20DBCB523794684B8785B3D5A907">
<mods:roleTerm id="26E1D7352D5CA3EA86377BE0B201B777">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BCCE5DA914DE80866CBD6EEE4396971E">Jara-Muñoz, Orlando A.</mods:namePart>
<mods:affiliation id="C2254DF5FDE44E7CC1E54A85B9C6B9AB">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá D. C., Colombia.</mods:affiliation>
<mods:nameIdentifier id="2E8889D5FBB9435FD314F7B3706F2173" type="email">oajaram@unal.edu.co</mods:nameIdentifier>
</mods:name>
<mods:name id="C6F75D25FA364089EF48F2C9F6D6C3E1" type="personal">
<mods:role id="38CC6AB9348485FB9FA50285348EE10E">
<mods:roleTerm id="75692C67725C785A992FB746CBCF65AC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A6B8B1E6DF3386C2EEFD2EAC20C0359B">Purvis, David A.</mods:namePart>
<mods:affiliation id="44C2556898F65A5A003E171E1B76EEB6">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="CCFC5026662EFEFBD9DD160FF0AE4CC8" type="email">dpurvis@rbge.org.uk</mods:nameIdentifier>
</mods:name>
<mods:name id="5C0B743C5DB4A14E479382FBFB72D8AA" type="personal">
<mods:role id="F328D47CAC5BD4299CC3E076F84CF5F5">
<mods:roleTerm id="825DCE370F290C22516B9BCBC0A2B3E6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E93FBC0A0B092586EBF462FBFB62FF33">Delves, Jay</mods:namePart>
<mods:affiliation id="FFF29430650E564954CFD028161EF410">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="6286D7F6E0BBE2899D0740AD403B189C" type="email">j.delves@westernsydney.edu.au</mods:nameIdentifier>
</mods:name>
<mods:name id="2900853FD3DD47B375446D7B0FD8396F" type="personal">
<mods:role id="EEA42E41B1E257830EF5C171EE7E7B00">
<mods:roleTerm id="8B4FF6E94ADAE94DE57E79D269E0F426">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A9F6FA8A1C6D5D55F9B6540B0B1DCEE2">Allen, Josh P.</mods:namePart>
<mods:affiliation id="F79987C6BD7D2B265E17EB6465537E38">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="FE38B3D41F4819328E53F060A27C10CA" type="email">myrmeciaman@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="DFAD0199193A891BF14B12D0F3E40969" type="personal">
<mods:role id="0C643C3076BB797C1087CEF1F1A86FEE">
<mods:roleTerm id="C2572FD788B87B1884B84D0E2310FDEC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="98DC9F640EABDCAF52E02A819FBF642A">Reynel, Carlos</mods:namePart>
<mods:affiliation id="9E08AFAC2561BDF47A225210F7F7A6E8">Facultad de Ciencias Forestales, Universidad Nacional Agraria-La Molina, Lima 12, Peru.</mods:affiliation>
<mods:nameIdentifier id="92D60589CBC48F63BE6400F6587A7854" type="email">reynel@lamolina.edu.pe</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="A344FE6A7860C1DD5038B51F2CCF49D1">text</mods:typeOfResource>
<mods:relatedItem id="9010BE912C800E64CEBC086858D585D7" type="host">
<mods:titleInfo id="9FD84B726BFACB5344636BE3810A0597">
<mods:title id="D5BF8BCF831C39DC2BE04D6CBEE25EB1">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="DBCFD28F8E5093C23E9471258D1216A3">
<mods:date id="C7CB8C67C963E68CCA4A2F8AADB922BB">2023</mods:date>
<mods:detail id="6129C3C7C68576A49016D8444AB35CB1" type="pubDate">
<mods:number id="D74E71BA4F6E7A23BAA9B0ED9697BD46">2023-07-18</mods:number>
</mods:detail>
<mods:detail id="E761FD9ADD4878414E68E65EC3233F72" type="volume">
<mods:number id="494DE0C5C6397FA2921E4E4738144810">881</mods:number>
</mods:detail>
<mods:extent id="DBA91EBF79BCB2FE5318062893B14F0E" unit="page">
<mods:start id="6EC4256D672160D693F8D3E42485508B">1</mods:start>
<mods:end id="3EF4B44A73D2F83F29A1AB853A153D74">334</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="26EA9CCA394FB1E605F5E4F7E0813C79">
<mods:url id="52BD894F9CF5B1E074517BC359690D7D">http://dx.doi.org/10.5852/ejt.2023.881.2175</mods:url>
</mods:location>
<mods:classification id="C4C27B5DB5F6BBF158062A16EA874AD4">journal article</mods:classification>
<mods:identifier id="4D9B5CF460275143912E4F9A68C394E7" type="DOI">https://doi.org/10.5852/ejt.2023.881.2175</mods:identifier>
<mods:identifier id="B0E7D43E4C8233CD089E5BEF3CCDEE1B" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="908F83F8DA80D8EA87E4E3D37C6FAE1F" type="Zenodo-Dep">8178280</mods:identifier>
</mods:mods>
<treatment id="03B26B4BFF79FF26FDAAFC2DAE09EF2B" LSID="urn:lsid:plazi:treatment:03B26B4BFF79FF26FDAAFC2DAE09EF2B" httpUri="http://treatment.plazi.org/id/03B26B4BFF79FF26FDAAFC2DAE09EF2B" lastPageId="223" lastPageNumber="224" pageId="220" pageNumber="221">
<subSubSection id="C30189D6FF79FF25FDAAFC2DAEE7E9D4" box="[627,960,914,941]" pageId="220" pageNumber="221" type="nomenclature">
<paragraph id="8BA4DA5DFF79FF25FDAAFC2DAEE7E9D4" blockId="220.[627,960,914,977]" box="[627,960,914,941]" pageId="220" pageNumber="221">
55.
<taxonomicName id="4C1BA1DEFF79FF25FD46FC2DAEE7E9D4" ID-CoL="87ZGH" ID-ENA="200057" authority="Herzog" authorityName="Herzog" box="[671,960,914,941]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="220" pageNumber="221" phylum="Tracheophyta" rank="species" species="wollnyi">
<emphasis id="B96F064FFF79FF25FD46FC2DAE44E9D4" bold="true" box="[671,867,914,941]" italics="true" pageId="220" pageNumber="221">Begonia wollnyi</emphasis>
Herzog
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF79FF25FD17FC09AE41E9A9" box="[718,870,950,977]" pageId="220" pageNumber="221" type="description">
<paragraph id="8BA4DA5DFF79FF25FD17FC09AE41E9A9" blockId="220.[627,960,914,977]" box="[718,870,950,977]" pageId="220" pageNumber="221">
<figureCitation id="1320C6D8FF79FF25FD17FC09AE1AE9A8" box="[718,829,950,977]" captionStart="Fig" captionStartId="204.[189,232,1822,1848]" captionTargetBox="[194,1392,558,1776]" captionTargetId="figure-14@204.[189,1398,548,1784]" captionTargetPageId="204" captionText="Fig. 65. Distribution of the acerifolia group of Begonia sect. Knesebeckia (Klotzsch) A.DC. in Peru and surrounding countries. A. B. acerifolia Kunth (red). B. B. arrogans Irmsch. (red) and B. ludwigii Irmsch. (blue). C. B. monadelpha (Ruiz ex Klotzsch) A.DC. (red) and B. serotina A.DC. (blue). D. B. huancabambae Moonlight sp. nov. (red) and B. wollnyi Herzog (blue). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178440" httpUri="https://zenodo.org/record/8178440/files/figure.png" pageId="220" pageNumber="221">Figs 65D</figureCitation>
,
<figureCitation id="1320C6D8FF79FF25FC90FC09AE41E9A9" box="[841,870,950,976]" captionStart="Fig" captionStartId="222.[189,232,1857,1883]" captionTargetBox="[246,1342,274,1819]" captionTargetId="figure-13@222.[246,1342,269,1819]" captionTargetPageId="222" captionText="Fig. 70. Begonia wollnyi Herzog. 1. Habit. 2. Leaf. 3. Staminate flower, front view. 4. Androecium, side view. 57. Stamens, side view. 8. Pistillate flower, side view. 9. Cross section of ovary, side view. 1011. Style, front and back view. 12. Cross section of ovary. Reproduction of an illustration of Begonia williamsii Rusby &amp; Nash by an unknown artist reproduced from Nash (1916: pl. 29) at the Royal Botanic Garden Edinburgh." figureDoi="http://doi.org/10.5281/zenodo.8178450" httpUri="https://zenodo.org/record/8178450/files/figure.png" pageId="220" pageNumber="221">70</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF79FF25FF64FC46A919EE6B" box="[189,1086,1016,1043]" pageId="220" pageNumber="221" type="reference_group">
<paragraph id="8BA4DA5DFF79FF25FF64FC46A919EE6B" blockId="220.[189,1399,1016,1725]" box="[189,1086,1016,1043]" pageId="220" pageNumber="221">
<emphasis id="B96F064FFF79FF25FF64FC46AE0DEE6B" box="[189,810,1016,1042]" italics="true" pageId="220" pageNumber="221">Repertorium specierum novarum regni vegetabilis</emphasis>
7: 63 (
<bibRefCitation id="EF8AA7ACFF79FF25FC54FC47A916EE6B" author="Herzog" box="[909,1073,1016,1042]" firstAuthor="Herzog" pageId="220" pageNumber="221" pagination="4 - 69" refId="ref227767" refString="Herzog T. C. J. 1909. Siphonogamae novae Bolivensis in itinere per Boliviam orientalem ab auctore letae. Repertorium specierum novarum regni vegetabilis 7: 4 - 69." type="journal article" year="1909">Herzog 1909</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF79FF25FB92FC46AEBEEE22" pageId="220" pageNumber="221" type="materials_examined">
<paragraph id="8BA4DA5DFF79FF25FB92FC46AEBEEE22" blockId="220.[189,1399,1016,1725]" pageId="220" pageNumber="221">
<emphasis id="B96F064FFF79FF25FBBEFC47A982EE6B" bold="true" box="[1127,1189,1016,1042]" pageId="220" pageNumber="221">Type</emphasis>
:
<materialsCitation id="3B73D000FF79FF25FB63FC47AEBEEE22" collectingDate="1907-10" collectionCode="Z" collectorName="T. C. J. Herzog" country="Bolivia" elevation="800" location="Im Bergwald der Quebrada de Cunucu" municipality="Cordillera de Sta. Cruz" pageId="220" pageNumber="221" specimenCode="Z-000001867" specimenCount="1" typeStatus="lectotype">
<collectingCountry id="F30C9ACDFF79FF25FB63FC47A816EE6B" box="[1210,1329,1016,1042]" name="Bolivia" pageId="220" pageNumber="221">BOLIVIA</collectingCountry>
<location id="8EC48C86FF79FF25FA8FFC46AFB2EE4F" LSID="urn:lsid:plazi:treatment:03B26B4BFF79FF26FDAAFC2DAE09EF2B:8EC48C86FF79FF25FA8FFC46AFB2EE4F" country="Bolivia" municipality="Cordillera de Sta. Cruz" name="Im Bergwald der Quebrada de Cunucu" pageId="220" pageNumber="221">Im Bergwald der Quebrada de Cuñucú</location>
(
<collectingMunicipality id="6BC04027FF79FF25FD71FBA3AEEFEE4F" box="[680,968,1052,1078]" pageId="220" pageNumber="221">Cordillera de Sta. Cruz</collectingMunicipality>
); ca
<elevation id="00363D6EFF79FF25FBDCFBA3A9B1EE4F" box="[1029,1174,1052,1079]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="220" pageNumber="221" unit="m" value="800.0">
<quantity id="4CE377B8FF79FF25FBDCFBA3A974EE4E" box="[1029,1107,1052,1079]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="220" pageNumber="221" unit="m" value="800.0">800 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF79FF25FB7EFBA3A806EE4F" box="[1191,1313,1052,1078]" pageId="220" pageNumber="221" value="1907-10">
<collectingDate id="EFE10575FF79FF25FB7EFBA3A806EE4F" box="[1191,1313,1052,1078]" pageId="220" pageNumber="221" value="1907-10">Oct. 1907</collectingDate>
</date>
;
<emphasis id="B96F064FFF79FF25FAE8FBA2AC41EE23" italics="true" pageId="220" pageNumber="221">
<collectorName id="26EEBF8BFF79FF25FAE8FBA2AC66EE23" pageId="220" pageNumber="221">T.C.J. Herzog</collectorName>
86
</emphasis>
;
<typeStatus id="54A064FFFF79FF25FEAFFBFFACC1EE23" box="[374,486,1088,1114]" pageId="220" pageNumber="221">lectotype</typeStatus>
:
<collectionCode id="ED0A4298FF79FF25FE28FBFEAF24EE22" box="[497,515,1089,1115]" country="Switzerland" name="Universität Zürich" pageId="220" pageNumber="221" type="Herbarium">Z</collectionCode>
[
<specimenCode id="DBBD7226FF79FF25FDC8FBFFAF9DEE23" box="[529,698,1088,1114]" pageId="220" pageNumber="221">Z-000001867</specimenCode>
],
<emphasis id="B96F064FFF79FF25FD15FBFFAEB5EE23" bold="true" box="[716,914,1088,1114]" pageId="220" pageNumber="221">designated here</emphasis>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF79FF25FF64FBDBA82CEEBF" pageId="220" pageNumber="221" type="reference_group">
<paragraph id="8BA4DA5DFF79FF25FF64FBDBACAEEEDB" blockId="220.[189,1399,1016,1725]" pageId="220" pageNumber="221">
<bibRefCitation id="EF8AA7ACFF79FF25FF64FBDBAF6DEE07" author="Smith &amp; Wasshausen" box="[189,586,1124,1150]" firstAuthor="Smith" page="68" pageId="220" pageNumber="221" pagination="5 - 78" refId="ref231911" refString="Smith L. B. &amp; Wasshausen D. C. 1989. Begoniaceae In: Lasser T. (ed.) Flora de Venezuela 4 (1): 5 - 78." type="journal article" year="1989">Smith &amp; Wasshausen (1989: 68)</bibRefCitation>
;
<bibRefCitation id="EF8AA7ACFF79FF25FD83FBDBAE60EE07" author="Golding" box="[602,839,1124,1150]" firstAuthor="Golding" page="49" pageId="220" pageNumber="221" pagination="49 - 53" refId="ref227451" refString="Golding J. 1999. Begonia wollnyi - emended. Begonian 66 (3): 49 - 53." type="journal article" year="1999">Golding (1999: 49)</bibRefCitation>
; Wasshausen
<emphasis id="B96F064FFF79FF25FC29FBDAA908EE07" box="[1008,1071,1124,1150]" italics="true" pageId="220" pageNumber="221">et al.</emphasis>
(2014: 386);
<bibRefCitation id="EF8AA7ACFF79FF25FB03FBDBACA2EEDB" author="Tebbitt" etAl="et al." firstAuthor="Tebbitt" page="218" pageId="220" pageNumber="221" pagination="215 - 226" refId="ref232737" refString="Tebbitt M. C., Andrada A. R., Kollmann J. C. &amp; Moonlight P. W. 2018 a. Taxonomy of Begonia wollnyi Herzog and Begonia arrogans Irmsch. Edinburgh Journal of Botany 75 (2): 215 - 226. https: // doi. org / 10.1017 / S 0960428618000069" type="journal article" year="2018" yearSuffix="a">
Tebbitt
<emphasis id="B96F064FFF79FF25FAEEFBDAA850EE07" box="[1335,1399,1124,1150]" italics="true" pageId="220" pageNumber="221">et al.</emphasis>
(2018a: 218)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BA4DA5DFF79FF25FF64FB13A82CEEBF" blockId="220.[189,1399,1016,1725]" box="[189,1291,1195,1222]" pageId="220" pageNumber="221">
<treatmentCitationGroup id="AB0BFD73FF79FF25FF64FB13A82CEEBF" box="[189,1291,1195,1222]" pageId="220" pageNumber="221">
<taxonomicName id="4C1BA1DEFF79FF25FF64FB13AE4EEEBC" ID-CoL="7SGV6" authority="L. B. Sm. &amp; B. G. Schub., Darwiniana" authorityName="L. B. Sm. &amp; B. G. Schub., Darwiniana" box="[189,873,1196,1222]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="220" pageNumber="221" phylum="Tracheophyta" rank="species" species="parodiana">
<emphasis id="B96F064FFF79FF25FF64FB13AC84EEBF" box="[189,419,1196,1222]" italics="true" pageId="220" pageNumber="221">Begonia parodiana</emphasis>
L.B.Sm. &amp; B.G.Schub.,
<treatmentCitation id="0ABAFC4CFF79FF25FD03FB13A820EEBF" author="Smith &amp; Schubert" box="[730,1287,1195,1222]" page="88" pageId="220" pageNumber="221" year="1941">
<emphasis id="B96F064FFF79FF25FD03FB13AE4EEEBC" box="[730,873,1196,1221]" italics="true" pageId="220" pageNumber="221">Darwiniana</emphasis>
5: 88 (
<bibRefCitation id="EF8AA7ACFF79FF25FC1EFB13A9D9EEBF" author="Smith &amp; Schubert" box="[967,1278,1196,1222]" firstAuthor="Smith" pageId="220" pageNumber="221" pagination="78 - 117" refId="ref231253" refString="Smith L. B. &amp; Schubert B. G. 1941 b. Revision de las especies Argentinas del genero Begonia. Darwiniana 5: 78 - 117. https: // www. jstor. org / stable / 23212166" type="journal article" year="1941" yearSuffix="b">Smith &amp; Schubert 1941b</bibRefCitation>
)
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF79FF25FACFFB13AEA6EF4B" pageId="220" pageNumber="221" type="materials_examined">
<paragraph id="8BA4DA5DFF79FF25FACFFB13AEA6EF4B" blockId="220.[189,1399,1016,1725]" pageId="220" pageNumber="221">
<emphasis id="B96F064FFF79FF25FAE9FB13A849EEBF" bold="true" box="[1328,1390,1196,1222]" pageId="220" pageNumber="221">Type</emphasis>
:
<materialsCitation id="3B73D000FF79FF25FF33FB6FAF38EF48" collectingDate="1939-10-29" collectionCode="GH, US" collectorName="W. J. Eyerdam &amp; A. A. Beetle" country="Argentina" county="Prov. Salta" elevation="800" latitude="-24.5" location="Cerros de rio Ytau" longLatPrecision="1251" longitude="-65.0" municipality="Dept. Oran" pageId="220" pageNumber="221" specimenCode="GH00068261" specimenCount="1">
<collectingCountry id="F30C9ACDFF79FF25FF33FB6FACB4EE93" box="[234,403,1232,1258]" name="Argentina" pageId="220" pageNumber="221">ARGENTINA</collectingCountry>
<emphasis id="B96F064FFF79FF25FE6FFB6FAF65EE90" bold="true" box="[438,578,1231,1258]" pageId="220" pageNumber="221">
<collectingCounty id="62C5A2D1FF79FF25FE6FFB6FAF65EE90" box="[438,578,1231,1258]" pageId="220" pageNumber="221">Prov. Salta</collectingCounty>
</emphasis>
<collectingMunicipality id="6BC04027FF79FF25FDB8FB6FAFCEEE93" box="[609,745,1232,1258]" pageId="220" pageNumber="221">Dept. Oran</collectingMunicipality>
,
<location id="8EC48C86FF79FF25FD2EFB6FAEFDEE93" LSID="urn:lsid:plazi:treatment:03B26B4BFF79FF26FDAAFC2DAE09EF2B:8EC48C86FF79FF25FD2EFB6FAEFDEE93" box="[759,986,1231,1258]" country="Argentina" county="Prov. Salta" latitude="-24.5" longLatPrecision="1251" longitude="-65.0" municipality="Dept. Oran" name="Cerros de rio Ytau" pageId="220" pageNumber="221">Cerros de río Ytau</location>
,
<locationDeviation id="9A16C37FFF79FF25FC31FB70A855EE90" box="[1000,1394,1231,1258]" pageId="220" pageNumber="221">
<quantity id="4CE377B8FF79FF25FC31FB70A913EE90" box="[1000,1076,1231,1258]" metricMagnitude="4" metricUnit="m" metricValue="5.4" pageId="220" pageNumber="221" unit="km" value="54.0">54 km</quantity>
west of Manuela Pedraza
</locationDeviation>
; [
<geoCoordinate id="EE2FBC9AFF79FF25FF28FB4DAC7FEF77" box="[241,344,1266,1294]" degrees="24" direction="south" minutes="30" orientation="latitude" pageId="220" pageNumber="221" precision="925" value="-24.5">24°30 S</geoCoordinate>
<geoCoordinate id="EE2FBC9AFF79FF25FEB9FB4CACF3EF77" box="[352,468,1266,1294]" degrees="65" direction="west" minutes="00" orientation="longitude" pageId="220" pageNumber="221" precision="925" value="-65.0">65°00 W</geoCoordinate>
];
<elevation id="00363D6EFF79FF25FE31FB4BAF55EF74" box="[488,626,1267,1294]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="220" pageNumber="221" unit="m" value="800.0">
<quantity id="4CE377B8FF79FF25FE31FB4BAF14EF77" box="[488,563,1268,1294]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="220" pageNumber="221" unit="m" value="800.0">800 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF79FF25FD59FB4BAE3CEF77" box="[640,795,1268,1294]" pageId="220" pageNumber="221" value="1939-10-29">
<collectingDate id="EFE10575FF79FF25FD59FB4BAE3CEF77" box="[640,795,1268,1294]" pageId="220" pageNumber="221" value="1939-10-29">29 Oct. 1939</collectingDate>
</date>
;
<emphasis id="B96F064FFF79FF25FCF1FB4BA9E1EF74" box="[808,1222,1267,1294]" italics="true" pageId="220" pageNumber="221">
<collectorName id="26EEBF8BFF79FF25FCF1FB4BAEECEF74" box="[808,971,1267,1293]" pageId="220" pageNumber="221">W.J. Eyerdam</collectorName>
&amp;
<collectorName id="26EEBF8BFF79FF25FC29FB4BA952EF74" box="[1008,1141,1267,1294]" pageId="220" pageNumber="221">A.A. Beetle</collectorName>
22726
</emphasis>
; holotype:
<collectionCode id="ED0A4298FF79FF25FA95FB4BA851EF77" box="[1356,1398,1268,1294]" country="USA" lsid="urn:lsid:biocol.org:col:15631" name="Harvard University - Gray Herbarium" pageId="220" pageNumber="221" type="Herbarium">GH</collectionCode>
[
<specimenCode id="DBBD7226FF79FF25FF28FAA8ACB0EF48" box="[241,407,1303,1329]" collectionCode="GH" country="USA" lsid="urn:lsid:biocol.org:col:15631" name="Harvard University - Gray Herbarium" pageId="220" pageNumber="221" type="Herbarium">GH00068261</specimenCode>
, photo
<collectionCode id="ED0A4298FF79FF25FE34FAA8AF3FEF48" box="[493,536,1303,1329]" pageId="220" pageNumber="221">US</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B73D000FF79FF25FDF2FAA8AEA6EF4B" box="[555,897,1303,1330]" collectingDate="1939-10-29" collectionCode="K, G" collectorName="W. J. Eyerdam &amp; A. A. Beetle" country="Argentina" county="Prov. Salta" elevation="800" latitude="-24.5" location="Cerros de rio Ytau" longLatPrecision="1251" longitude="-65.0" municipality="Dept. Oran" pageId="220" pageNumber="221" specimenCode="G00034152" specimenCount="1">
isotypes:
<collectionCode id="ED0A4298FF79FF25FD42FAA7AF94EF4B" box="[667,691,1304,1330]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="220" pageNumber="221" type="Herbarium">K</collectionCode>
,
<collectionCode id="ED0A4298FF79FF25FD66FAA8AFF3EF48" box="[703,724,1303,1329]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="220" pageNumber="221" type="Herbarium">G</collectionCode>
[
<specimenCode id="DBBD7226FF79FF25FD3BFAA8AE50EF4B" box="[738,887,1303,1330]" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="220" pageNumber="221" type="Herbarium">G00034152</specimenCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF79FF25FF64FA84AE23EFE4" pageId="220" pageNumber="221" type="reference_group">
<paragraph id="8BA4DA5DFF79FF25FF64FA84A919EF2C" blockId="220.[189,1399,1016,1725]" box="[189,1086,1339,1366]" pageId="220" pageNumber="221">
Wasshausen
<emphasis id="B96F064FFF79FF25FE8AFA83ACA9EF2C" box="[339,398,1339,1365]" italics="true" pageId="220" pageNumber="221">et al.</emphasis>
(2013: 385);
<bibRefCitation id="EF8AA7ACFF79FF25FDF6FA84AFD0EF2C" author="Delfini" box="[559,759,1339,1365]" firstAuthor="Delfini" page="8" pageId="220" pageNumber="221" pagination="1 - 16" refId="ref226758" refString="Delfini C. 2017. Begoniaceae. In: Zuloaga F. O. &amp; Belgrano J. (eds) Flora Vascular de la Republica Argentina, vol. 17: 1 - 16. Instituto de Botanica Darwinion, San Isidro, Argentina https: // doi. org / 10.2307 / j. ctt 20 p 56 nv. 4" type="journal article" year="2017">Delfini (2017: 8)</bibRefCitation>
;
<bibRefCitation id="EF8AA7ACFF79FF25FCDAFA84A91DEF2C" author="Tebbitt" box="[771,1082,1339,1365]" etAl="et al." firstAuthor="Tebbitt" page="218" pageId="220" pageNumber="221" pagination="215 - 226" refId="ref232737" refString="Tebbitt M. C., Andrada A. R., Kollmann J. C. &amp; Moonlight P. W. 2018 a. Taxonomy of Begonia wollnyi Herzog and Begonia arrogans Irmsch. Edinburgh Journal of Botany 75 (2): 215 - 226. https: // doi. org / 10.1017 / S 0960428618000069" type="journal article" year="2018" yearSuffix="a">
Tebbitt
<emphasis id="B96F064FFF79FF25FC84FA83AEBFEF2C" box="[861,920,1339,1365]" italics="true" pageId="220" pageNumber="221">et al.</emphasis>
(2018a: 218)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BA4DA5DFF79FF25FF64FADFAE23EFE4" blockId="220.[189,1399,1016,1725]" pageId="220" pageNumber="221">
<treatmentCitationGroup id="AB0BFD73FF79FF25FF64FADFAE23EFE4" pageId="220" pageNumber="221">
<treatmentCitation id="0ABAFC4CFF79FF25FF64FADFAE27EFE4" author="Irmsch., Botanische Jahrbucher" journal="Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie" page="605" pageId="220" pageNumber="221" score="478.66328617067893" volume="74" year="1949">
<taxonomicName id="4C1BA1DEFF79FF25FF64FADFAE72EF00" ID-CoL="87Y4Y" authority="Irmsch., Botanische Jahrbucher" authorityName="Irmsch., Botanische Jahrbucher" box="[189,853,1375,1401]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="220" pageNumber="221" phylum="Tracheophyta" rank="species" species="acrensis">
<emphasis id="B96F064FFF79FF25FF64FADFACBDEF00" box="[189,410,1376,1401]" italics="true" pageId="220" pageNumber="221">Begonia acrensis</emphasis>
Irmsch.,
<emphasis id="B96F064FFF79FF25FDF7FAE0AE72EF00" box="[558,853,1375,1401]" italics="true" pageId="220" pageNumber="221">Botanische Jahrbücher</emphasis>
</taxonomicName>
<emphasis id="B96F064FFF79FF25FCB6FAE0ACF0EFE4" italics="true" pageId="220" pageNumber="221">
für
<bibCitation id="5E31D231FF79FF25FC75FAE0ACF0EFE4" author="Systematik" journal="Pflanzengeschichte und Pflanzengeographie" page="605" pageId="220" pageNumber="221" score="109.65844492058366" volume="74" year="1949">Systematik, Pflanzengeschichte und Pflanzengeographie</bibCitation>
</emphasis>
74: 605 (
<bibRefCitation id="EF8AA7ACFF79FF25FD9FFA3CAFD0EFE4" author="Irmscher" box="[582,759,1411,1437]" firstAuthor="Irmscher" pageId="220" pageNumber="221" pagination="569 - 633" refId="ref228493" refString="Irmscher E. 1949. Begoniaceae, Beitrage zur Kenntnis der Begoniaceen Sudamerikas. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 74: 569 - 633." type="journal article" year="1949">Irmscher 1949</bibRefCitation>
)
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF79FF25FCD3FA3BACC2EF9C" pageId="220" pageNumber="221" type="materials_examined">
<paragraph id="8BA4DA5DFF79FF25FCD3FA3BACC2EF9C" blockId="220.[189,1399,1016,1725]" pageId="220" pageNumber="221">
<emphasis id="B96F064FFF79FF25FCF9FA3CAE79EFE4" bold="true" box="[800,862,1411,1437]" pageId="220" pageNumber="221">Type</emphasis>
:
<materialsCitation id="3B73D000FF79FF25FCB4FA3CA9E9EFB8" collectingDate="1911-06" collectionCode="US" collectorName="E. Ule" country="Brazil" location="rio Acre" pageId="220" pageNumber="221" specimenCode="US00115231" specimenCount="1" stateProvince="Acre State">
<collectingCountry id="F30C9ACDFF79FF25FCB4FA3CAEF1EFE4" box="[877,982,1411,1437]" name="Brazil" pageId="220" pageNumber="221">BRAZIL</collectingCountry>
<emphasis id="B96F064FFF79FF25FC2AFA3CA951EFE4" bold="true" box="[1011,1142,1411,1437]" pageId="220" pageNumber="221">
<collectingRegion id="49DF14BFFF79FF25FC2AFA3CA951EFE4" box="[1011,1142,1411,1437]" country="Brazil" name="Acre" pageId="220" pageNumber="221">Acre State</collectingRegion>
</emphasis>
<location id="8EC48C86FF79FF25FB56FA3CA9D5EFE4" LSID="urn:lsid:plazi:treatment:03B26B4BFF79FF26FDAAFC2DAE09EF2B:8EC48C86FF79FF25FB56FA3CA9D5EFE4" box="[1167,1266,1411,1437]" country="Brazil" name="rio Acre" pageId="220" pageNumber="221" stateProvince="Acre State">río Acre</location>
;
<date id="FFA5FC9DFF79FF25FB27FA3BA855EFE4" box="[1278,1394,1411,1438]" pageId="220" pageNumber="221" value="1911-06">
<collectingDate id="EFE10575FF79FF25FB27FA3BA855EFE4" box="[1278,1394,1411,1438]" pageId="220" pageNumber="221" value="1911-06">Jun. 1911</collectingDate>
</date>
;
<emphasis id="B96F064FFF79FF25FF33FA17AC5FEFB8" box="[234,376,1447,1473]" italics="true" pageId="220" pageNumber="221">
<collectorName id="26EEBF8BFF79FF25FF33FA17AC11EFB8" box="[234,310,1447,1473]" pageId="220" pageNumber="221">E. Ule</collectorName>
9649
</emphasis>
; lectotype:
<collectionCode id="ED0A4298FF79FF25FDDBFA18AF00EFB8" box="[514,551,1447,1473]" pageId="220" pageNumber="221">US</collectionCode>
[
<specimenCode id="DBBD7226FF79FF25FDE0FA18AFF4EFBB" box="[569,723,1447,1474]" pageId="220" pageNumber="221">US00115231</specimenCode>
], designated by
<bibRefCitation id="EF8AA7ACFF79FF25FC4AFA18A9EDEFB8" author="Tebbitt" box="[915,1226,1447,1473]" etAl="et al." firstAuthor="Tebbitt" page="218" pageId="220" pageNumber="221" pagination="215 - 226" refId="ref232737" refString="Tebbitt M. C., Andrada A. R., Kollmann J. C. &amp; Moonlight P. W. 2018 a. Taxonomy of Begonia wollnyi Herzog and Begonia arrogans Irmsch. Edinburgh Journal of Botany 75 (2): 215 - 226. https: // doi. org / 10.1017 / S 0960428618000069" type="journal article" year="2018" yearSuffix="a">
Tebbitt
<emphasis id="B96F064FFF79FF25FC34FA17A90FEFB8" box="[1005,1064,1447,1473]" italics="true" pageId="220" pageNumber="221">et al.</emphasis>
(2018a: 218)
</bibRefCitation>
;
</materialsCitation>
<materialsCitation id="3B73D000FF79FF25FB0CFA18ACC2EF9C" collectingDate="1911-06" collectionCode="G, K, U" collectorName="E. Ule" country="Brazil" location="rio Acre" pageId="220" pageNumber="221" specimenCode="U0000724" specimenCount="1" stateProvince="Acre State">
isolectotypes:
<collectionCode id="ED0A4298FF79FF25FF33FA74AC25EF9C" box="[234,258,1483,1509]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="220" pageNumber="221" type="Herbarium">G</collectionCode>
,
<collectionCode id="ED0A4298FF79FF25FED7FA74AC01EF9C" box="[270,294,1483,1509]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="220" pageNumber="221" type="Herbarium">K</collectionCode>
,
<collectionCode id="ED0A4298FF79FF25FEEBFA74AC60EF9C" box="[306,327,1483,1509]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15513" name="Nationaal Herbarium Nederland" pageId="220" pageNumber="221" type="Herbarium">U</collectionCode>
[
<specimenCode id="DBBD7226FF79FF25FE8CFA74ACFDEF9C" box="[341,474,1483,1509]" collectionCode="U" country="Netherlands" lsid="urn:lsid:biocol.org:col:15513" name="Nationaal Herbarium Nederland" pageId="220" pageNumber="221" type="Herbarium">U0000724</specimenCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF79FF25FF64FA50AC5EEC29" pageId="220" pageNumber="221" type="reference_group">
<paragraph id="8BA4DA5DFF79FF25FF64FA50AC80EC70" blockId="220.[189,1399,1016,1725]" box="[189,423,1519,1545]" pageId="220" pageNumber="221">
<bibRefCitation id="EF8AA7ACFF79FF25FF64FA50AC84EC70" author="Golding" box="[189,419,1519,1545]" firstAuthor="Golding" page="49" pageId="220" pageNumber="221" pagination="49 - 53" refId="ref227451" refString="Golding J. 1999. Begonia wollnyi - emended. Begonian 66 (3): 49 - 53." type="journal article" year="1999">Golding (1999: 49)</bibRefCitation>
.
</paragraph>
<paragraph id="8BA4DA5DFF79FF25FF64F9ACAC5EEC29" blockId="220.[189,1399,1016,1725]" pageId="220" pageNumber="221">
<treatmentCitationGroup id="AB0BFD73FF79FF25FF64F9ACAC5EEC29" pageId="220" pageNumber="221">
<taxonomicName id="4C1BA1DEFF79FF25FF64F9ACAF61EC55" ID-CoL="7SJ3P" authority="Rusby &amp; Nash" authorityName="Rusby &amp; Nash" box="[189,582,1554,1581]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="220" pageNumber="221" phylum="Tracheophyta" rank="species" species="williamsii">
<emphasis id="B96F064FFF79FF25FF64F9ACACBFEC55" box="[189,408,1554,1580]" italics="true" pageId="220" pageNumber="221">Begonia williamsii</emphasis>
Rusby &amp; Nash
</taxonomicName>
nom. illeg.; later homonym non R.S.
<location id="8EC48C86FF79FF25FC3EF9ADA97BEC55" LSID="urn:lsid:plazi:treatment:03B26B4BFF79FF26FDAAFC2DAE09EF2B:8EC48C86FF79FF25FC3EF9ADA97BEC55" box="[999,1116,1554,1580]" country="Bolivia" municipality="Type" name="Williams" pageId="220" pageNumber="221">Williams</location>
,
<treatmentCitation id="0ABAFC4CFF79FF25FBBFF9ACAC51EC29" author="Rusby &amp; Nash" page="47" pageId="220" pageNumber="221" year="1906">
<emphasis id="B96F064FFF79FF25FBBFF9ACA9E5EC55" box="[1126,1218,1555,1580]" italics="true" pageId="220" pageNumber="221">Torreya</emphasis>
6: 47 (
<bibRefCitation id="EF8AA7ACFF79FF25FAD7F9ADAC4BEC29" author="Rusby &amp; Nash" firstAuthor="Rusby" pageId="220" pageNumber="221" pagination="45 - 46" refId="ref230865" refString="Rusby H. H. &amp; Nash G. V. 1906. A new Begonia from Bolivia. Torreya 6: 45 - 46." type="journal article" year="1906">Rusby &amp; Nash 1906</bibRefCitation>
)
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF79FF25FE5BF988A972ECE0" pageId="220" pageNumber="221" type="materials_examined">
<paragraph id="8BA4DA5DFF79FF25FE5BF988A972ECE0" blockId="220.[189,1399,1016,1725]" pageId="220" pageNumber="221">
<emphasis id="B96F064FFF79FF25FE44F988ACFDEC28" bold="true" box="[413,474,1591,1617]" pageId="220" pageNumber="221">
<collectingMunicipality id="6BC04027FF79FF25FE44F988ACFDEC28" box="[413,474,1591,1617]" pageId="220" pageNumber="221">Type</collectingMunicipality>
</emphasis>
:
<materialsCitation id="3B73D000FF79FF25FE33F988A850EC0D" collectingDate="1901-11-14" collectionCode="NY" collectorName="R. S. Williams" country="Bolivia" elevation="420" latitude="-14.45" location="San Buena Ventura" longLatPrecision="1288" longitude="-67.55" municipality="San Buena Ventura" pageId="220" pageNumber="221" specimenCode="NY00118714" specimenCount="1">
<collectingCountry id="F30C9ACDFF79FF25FE33F988AF46EC28" box="[490,609,1591,1617]" name="Bolivia" pageId="220" pageNumber="221">BOLIVIA</collectingCountry>
<collectingMunicipality id="6BC04027FF79FF25FDA4F988AE43EC28" box="[637,868,1591,1617]" pageId="220" pageNumber="221">San Buena Ventura</collectingMunicipality>
; [
<geoCoordinate id="EE2FBC9AFF79FF25FCA0F98AAEC4EC28" box="[889,995,1589,1617]" degrees="14" direction="south" minutes="27" orientation="latitude" pageId="220" pageNumber="221" precision="925" value="-14.45">14°27 S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF79FF25FC28F98AA940EC28" box="[1009,1127,1589,1617]" degrees="67" direction="west" minutes="33" orientation="longitude" pageId="220" pageNumber="221" precision="925" value="-67.55">67°33 W</geoCoordinate>
];
<elevation id="00363D6EFF79FF25FBA5F988A82EEC29" box="[1148,1289,1590,1617]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="220" pageNumber="221" unit="m" value="420.0">
<quantity id="4CE377B8FF79FF25FBA5F988A9EFEC28" box="[1148,1224,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="220" pageNumber="221" unit="m" value="420.0">420 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF79FF25FAC1F988AC0FEC0D" pageId="220" pageNumber="221" value="1901-11-14">
<collectingDate id="EFE10575FF79FF25FAC1F988AC0FEC0D" pageId="220" pageNumber="221" value="1901-11-14">14 Nov. 1901</collectingDate>
</date>
;
<emphasis id="B96F064FFF79FF25FEEDF9E4AF23EC0D" box="[308,516,1626,1653]" italics="true" pageId="220" pageNumber="221">
<collectorName id="26EEBF8BFF79FF25FEEDF9E4ACF5EC0D" box="[308,466,1626,1653]" pageId="220" pageNumber="221">R.S. Williams</collectorName>
600
</emphasis>
; lectotype:
<collectionCode id="ED0A4298FF79FF25FD54F9E4AF91EC0C" box="[653,694,1627,1653]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="220" pageNumber="221" type="Herbarium">NY</collectionCode>
[
<specimenCode id="DBBD7226FF79FF25FD1EF9E5AE42EC0D" box="[711,869,1626,1652]" collectionCode="NY" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="220" pageNumber="221" type="Herbarium">NY00118714</specimenCode>
], designated by
<bibRefCitation id="EF8AA7ACFF79FF25FBFDF9E5A850EC0D" author="Smith &amp; Schubert" box="[1060,1399,1626,1652]" firstAuthor="Smith" page="84" pageId="220" pageNumber="221" pagination="71 - 87" refId="ref231295" refString="Smith L. B. &amp; Schubert B. G. 1944. Revision de las especies Bolivianas del genero Begonia. Revistas Cientificas de la Universidad Andina del Cusco 33 (87): 71 - 87." type="journal article" year="1944">Smith &amp; Schubert (1944: 84)</bibRefCitation>
</materialsCitation>
<materialsCitation id="3B73D000FF79FF25FF30F9C1A972ECE0" box="[233,1109,1662,1689]" collectingDate="1901-11-14" collectionCode="BM, K, US" collectorName="R. S. Williams" country="Bolivia" elevation="420" latitude="-14.45" location="San Buena Ventura" longLatPrecision="1288" longitude="-67.55" municipality="San Buena Ventura" pageId="220" pageNumber="221" specimenCode="BM000043984, K000536789, US00115497" specimenCount="1">
isolectotypes:
<collectionCode id="ED0A4298FF79FF25FE4BF9C0ACE7ECE0" box="[402,448,1663,1689]" country="United Kingdom" name="Bristol Museum" pageId="220" pageNumber="221">BM</collectionCode>
[
<specimenCode id="DBBD7226FF79FF25FE17F9C1AFAFECE1" box="[462,648,1662,1688]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="220" pageNumber="221">BM000043984</specimenCode>
],
<collectionCode id="ED0A4298FF79FF25FD42F9C0AF97ECE0" box="[667,688,1663,1689]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="220" pageNumber="221" type="Herbarium">K</collectionCode>
[
<specimenCode id="DBBD7226FF79FF25FD1BF9C1AE7BECE0" box="[706,860,1662,1689]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="220" pageNumber="221" type="Herbarium">K000536789</specimenCode>
],
<collectionCode id="ED0A4298FF79FF25FCADF9C1AEBEECE1" box="[884,921,1662,1688]" pageId="220" pageNumber="221">US</collectionCode>
[
<specimenCode id="DBBD7226FF79FF25FC73F9C1A963ECE0" box="[938,1092,1662,1689]" pageId="220" pageNumber="221">US00115497</specimenCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF79FF25FF64F91DAECFECC5" box="[189,1000,1698,1725]" pageId="220" pageNumber="221" type="reference_group">
<paragraph id="8BA4DA5DFF79FF25FF64F91DAECFECC5" blockId="220.[189,1399,1016,1725]" box="[189,1000,1698,1725]" pageId="220" pageNumber="221">
<bibRefCitation id="EF8AA7ACFF79FF25FF64F91DAC58ECC4" author="Nash" box="[189,383,1698,1725]" firstAuthor="Nash" page="57" pageId="220" pageNumber="221" pagination="355 - 360" refId="ref230318" refString="Nash G. V. 1916. Begonia williamsii, William's Begonia, native of Bolivia Addisonia 1: 37 - 38. Otto C. F. &amp; Dietrich A. G. 1836. Begonien. Allgemeine Gartenzeitung 4 (45): 355 - 360." type="journal article" year="1916">Nash (1916: 57)</bibRefCitation>
;
<bibRefCitation id="EF8AA7ACFF79FF25FE52F91DAFD5ECC5" author="Smith &amp; Schubert" box="[395,754,1698,1725]" firstAuthor="Smith" page="114" pageId="220" pageNumber="221" pagination="110 - 114" refId="ref231589" refString="Smith L. B. &amp; Schubert B. G. 1955. Studies in the Begoniaceae, IV. Journal of the Washington Academy of Sciences 45 (4): 110 - 114. https: // biostor. org / reference / 133744" type="journal article" year="1955">Smith &amp; Schubert (1955: 114)</bibRefCitation>
;
<bibRefCitation id="EF8AA7ACFF79FF25FD27F91DAEC3ECC5" author="Golding" box="[766,996,1698,1724]" firstAuthor="Golding" page="49" pageId="220" pageNumber="221" pagination="49 - 53" refId="ref227451" refString="Golding J. 1999. Begonia wollnyi - emended. Begonian 66 (3): 49 - 53." type="journal article" year="1999">Golding (1999: 49)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF79FF25FF64F959ACD2ED2A" pageId="220" pageNumber="221" type="etymology">
<paragraph id="8BA4DA5DFF79FF25FF64F959AC62ED79" blockId="220.[189,325,1766,1792]" box="[189,325,1766,1792]" pageId="220" pageNumber="221">
<heading id="D0EC6D31FF79FF25FF64F959AC62ED79" bold="true" box="[189,325,1766,1792]" fontSize="11" level="3" pageId="220" pageNumber="221" reason="3">
<emphasis id="B96F064FFF79FF25FF64F959AC62ED79" bold="true" box="[189,325,1766,1792]" pageId="220" pageNumber="221">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF79FF25FF64F8AAACD2ED2A" blockId="220.[187,1398,1813,1875]" pageId="220" pageNumber="221">Named for Walter Wollny, who was a German botanist and contemporary of Theodor Carl Julius Herzog, who described the species.</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF79FF24FF64F8C4A9D5EB15" lastPageId="221" lastPageNumber="222" pageId="220" pageNumber="221" type="materials_examined">
<paragraph id="8BA4DA5DFF79FF25FF64F8C4ACE3EDEC" blockId="220.[189,452,1915,1941]" box="[189,452,1915,1941]" pageId="220" pageNumber="221">
<heading id="D0EC6D31FF79FF25FF64F8C4ACE3EDEC" bold="true" box="[189,452,1915,1941]" fontSize="11" level="3" pageId="220" pageNumber="221" reason="3">
<emphasis id="B96F064FFF79FF25FF64F8C4ACE3EDEC" bold="true" box="[189,452,1915,1941]" pageId="220" pageNumber="221">Specimens examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF79FF24FF64F814A9D5EB15" blockId="220.[189,1399,1961,2025]" lastBlockId="221.[189,1399,265,364]" lastPageId="221" lastPageNumber="222" pageId="220" pageNumber="221">
<materialsCitation id="3B73D000FF79FF24FF64F814AE9BEB5A" collectingDate="1997-01-13" collectionCode="CUZ, MO, NY, US, USM" collectorName="P. Acevedo R." country="Peru" county="Prov. La Convencion" elevation="467" lastPageId="221" lastPageNumber="222" latitude="-11.785556" location="campamento San Martin-C" longLatPrecision="21" longitude="-72.699165" municipality="Dist. Camisea" pageId="220" pageNumber="221" specimenCode="MO-1835960, US00594261" specimenCount="1" stateProvince="Cusco Region">
<collectingCountry id="F30C9ACDFF79FF25FF64F814AC2FEDBC" box="[189,264,1963,1989]" name="Peru" pageId="220" pageNumber="221">PERU</collectingCountry>
<emphasis id="B96F064FFF79FF25FEF1F815AFD3EDBD" bold="true" box="[296,756,1961,1989]" pageId="220" pageNumber="221">
<collectingRegion id="49DF14BFFF79FF25FEF1F815ACFDEDBD" box="[296,474,1962,1988]" country="Peru" name="Cusco" pageId="220" pageNumber="221">Cusco Region</collectingRegion>
:
<collectingCounty id="62C5A2D1FF79FF25FE30F814AFD3EDBD" box="[489,756,1961,1989]" pageId="220" pageNumber="221">Prov. La Convención</collectingCounty>
</emphasis>
<collectingMunicipality id="6BC04027FF79FF25FCD7F815AE91EDBD" box="[782,950,1962,1988]" pageId="220" pageNumber="221">Dist. Camisea</collectingMunicipality>
,
<location id="8EC48C86FF79FF25FC1AF814A822EDBC" LSID="urn:lsid:plazi:treatment:03B26B4BFF79FF26FDAAFC2DAE09EF2B:8EC48C86FF79FF25FC1AF814A822EDBC" box="[963,1285,1963,1989]" country="Peru" county="Prov. La Convencion" latitude="-11.785556" longLatPrecision="21" longitude="-72.699165" municipality="Dist. Camisea" name="campamento San Martin-C" pageId="220" pageNumber="221" stateProvince="Cusco Region">campamento San Martín-C</location>
,
<location id="8EC48C86FF79FF25FAC8F815AC5EED91" LSID="urn:lsid:plazi:treatment:03B26B4BFF79FF26FDAAFC2DAE09EF2B:8EC48C86FF79FF25FAC8F815AC5EED91" country="Peru" county="Prov. La Convencion" latitude="-11.785556" longLatPrecision="21" longitude="-72.699165" municipality="Dist. Camisea" name="Camisea Production unit" pageId="220" pageNumber="221" stateProvince="Cusco Region">Camisea Production unit</location>
,
<locationDeviation id="9A16C37FFF79FF25FE5EF870AF0BED90" box="[391,556,1998,2025]" pageId="220" pageNumber="221">west of camp</locationDeviation>
;
<geoCoordinate id="EE2FBC9AFF79FF25FDE3F872AFEDED91" box="[570,714,1997,2025]" degrees="11" direction="south" minutes="47" orientation="latitude" pageId="220" pageNumber="221" precision="15" seconds="08" value="-11.785556">11°4708″ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF79FF25FD00F872AE50ED90" box="[729,887,1997,2025]" degrees="72" direction="west" minutes="41" orientation="longitude" pageId="220" pageNumber="221" precision="15" seconds="57" value="-72.699165">72°4157″ W</geoCoordinate>
;
<elevation id="00363D6EFF79FF25FC5CF871A933ED91" box="[901,1044,1998,2025]" metricMagnitude="2" metricUnit="m" metricValue="4.67" pageId="220" pageNumber="221" unit="m" value="467.0">
<quantity id="4CE377B8FF79FF25FC5CF871AEF5ED90" box="[901,978,1998,2025]" metricMagnitude="2" metricUnit="m" metricValue="4.67" pageId="220" pageNumber="221" unit="m" value="467.0">467 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF79FF25FBFDF871A9E6ED91" box="[1060,1217,1998,2025]" pageId="220" pageNumber="221" value="1997-01-13">
<collectingDate id="EFE10575FF79FF25FBFDF871A9E6ED91" box="[1060,1217,1998,2025]" pageId="220" pageNumber="221" value="1997-01-13">13 Jan. 1997</collectingDate>
</date>
;
<collectorName id="26EEBF8BFF79FF25FB09F870A850ED91" box="[1232,1399,1998,2024]" pageId="220" pageNumber="221">
<emphasis id="B96F064FFF79FF25FB09F870A855ED91" box="[1232,1394,1998,2024]" italics="true" pageId="220" pageNumber="221">P. Acevedo R</emphasis>
<emphasis id="B96F064FFF79FF25FAAAF870A850ED91" box="[1395,1399,1999,2024]" italics="true" pageId="220" pageNumber="221">.</emphasis>
</collectorName>
<emphasis id="B96F064FFF78FF24FF64FEB6ADDFEB5A" box="[189,248,265,291]" italics="true" pageId="221" pageNumber="222">8682</emphasis>
;
<collectionCode id="ED0A4298FF78FF24FED3FEB6AC6FEB5A" box="[266,328,265,291]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14364" lsid="urn:lsid:biocol.org:col:14364" name="Universidad Nacional San Antonio Abad del Cusco" pageId="221" pageNumber="222" type="Herbarium">CUZ</collectionCode>
,
<collectionCode id="ED0A4298FF78FF24FE8EFEB6ACA1EB5A" box="[343,390,265,291]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="221" pageNumber="222" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FF78FF24FE43FEB6AF1DEB5D" box="[410,570,265,292]" pageId="221" pageNumber="222">MO-1835960</specimenCode>
],
<collectionCode id="ED0A4298FF78FF24FD8DFEB5AF59EB5D" box="[596,638,266,292]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="221" pageNumber="222" type="Herbarium">NY</collectionCode>
,
<collectionCode id="ED0A4298FF78FF24FD55FEB6AF96EB5A" box="[652,689,265,291]" pageId="221" pageNumber="222">US</collectionCode>
[
<specimenCode id="DBBD7226FF78FF24FD1FFEB6AE46EB5D" box="[710,865,265,292]" pageId="221" pageNumber="222">US00594261</specimenCode>
],
<collectionCode id="ED0A4298FF78FF24FCA5FEB6AE9BEB5A" box="[892,956,265,291]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="221" pageNumber="222">USM</collectionCode>
</materialsCitation>
<materialsCitation id="3B73D000FF78FF24FC03FEB6A9D5EB15" collectingDate="1997-02-15" collectionCode="USM" collectorName="P. Nunez &amp; S. Baldeon M. &amp; A. Alonso &amp; J. Santisteban &amp; G. Valencia &amp; K. Anderson &amp; J. Luton &amp; J. Pacaya V. &amp; F. Ramos R." elevation="400" latitude="-11.7815" location="San Martin" longLatPrecision="13" longitude="-72.70167" municipality="Dist. Echarate" pageId="221" pageNumber="222" specimenCount="1">
<collectingMunicipality id="6BC04027FF78FF24FC03FEB6A9A1EB5A" box="[986,1158,265,291]" pageId="221" pageNumber="222">Dist. Echarate</collectingMunicipality>
,
<location id="8EC48C86FF78FF24FB4DFEB6A83AEB5A" LSID="urn:lsid:plazi:treatment:03B26B4BFF79FF26FDAAFC2DAE09EF2B:8EC48C86FF78FF24FB4DFEB6A83AEB5A" box="[1172,1309,265,291]" latitude="-11.7815" longLatPrecision="13" longitude="-72.70167" municipality="Dist. Echarate" name="San Martin" pageId="221" pageNumber="222">San Martín</location>
,
<locationDeviation id="9A16C37FFF78FF24FAF2FEB6ADCEEB3E" pageId="221" pageNumber="222">3 well site</locationDeviation>
;
<geoCoordinate id="EE2FBC9AFF78FF24FF20FE93ACA1EB31" box="[249,390,300,328]" degrees="11" direction="south" minutes="46.89" orientation="latitude" pageId="221" pageNumber="222" precision="9" value="-11.7815">11°46.89 S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF78FF24FE4EFE93AF16EB31" box="[407,561,300,328]" degrees="72" direction="west" minutes="42.10" orientation="longitude" pageId="221" pageNumber="222" precision="9" value="-72.70167">72°42.10 W</geoCoordinate>
;
<elevation id="00363D6EFF78FF24FD98FE91AFF4EB3E" box="[577,723,301,328]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="221" pageNumber="222" unit="m" value="400.0">
<quantity id="4CE377B8FF78FF24FD98FE91AFA8EB31" box="[577,655,302,328]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="221" pageNumber="222" unit="m" value="400.0">400 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF78FF24FD3DFE92AEAEEB31" box="[740,905,301,328]" pageId="221" pageNumber="222" value="1997-02-15">
<collectingDate id="EFE10575FF78FF24FD3DFE92AEAEEB31" box="[740,905,301,328]" pageId="221" pageNumber="222" value="1997-02-15">15 Feb. 1997</collectingDate>
</date>
;
<emphasis id="B96F064FFF78FF24FC40FE91A9BCEB15" italics="true" pageId="221" pageNumber="222">
<collectorName id="26EEBF8BFF78FF24FC40FE91A920EB3E" box="[921,1031,302,327]" pageId="221" pageNumber="222">P. Nuñez</collectorName>
,
<collectorName id="26EEBF8BFF78FF24FBCEFE91A9EEEB3E" box="[1047,1225,301,328]" pageId="221" pageNumber="222">S. Baldeón M.</collectorName>
,
<collectorName id="26EEBF8BFF78FF24FB00FE91A875EB3E" box="[1241,1362,301,328]" pageId="221" pageNumber="222">A. Alonso</collectorName>
,
<collectorName id="26EEBF8BFF78FF24FABBFE91AC6DEB15" pageId="221" pageNumber="222">J. Santisteban</collectorName>
,
<collectorName id="26EEBF8BFF78FF24FE8FFEEDACC3EB15" box="[342,484,338,364]" pageId="221" pageNumber="222">G. Valencia</collectorName>
,
<collectorName id="26EEBF8BFF78FF24FE29FEECAFA0EB15" box="[496,647,338,364]" pageId="221" pageNumber="222">K. Anderson</collectorName>
,
<collectorName id="26EEBF8BFF78FF24FD4AFEECAFD0EB15" box="[659,759,339,364]" pageId="221" pageNumber="222">J. Luton</collectorName>
,
<collectorName id="26EEBF8BFF78FF24FCDAFEECAEB1EB15" box="[771,918,339,364]" pageId="221" pageNumber="222">J. Pacaya V.</collectorName>
&amp;
<collectorName id="26EEBF8BFF78FF24FC62FEECA96DEB15" box="[955,1098,339,364]" pageId="221" pageNumber="222">F. Ramos R.</collectorName>
19107
</emphasis>
;
<collectionCode id="ED0A4298FF78FF24FB72FEEDA9C9EB15" box="[1195,1262,338,364]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="221" pageNumber="222">USM</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF78FF24FF64FE2BA963EFEB" pageId="221" pageNumber="222" type="description">
<paragraph id="8BA4DA5DFF78FF24FF64FE2BAC68EBD7" blockId="221.[189,335,404,430]" box="[189,335,404,430]" pageId="221" pageNumber="222">
<heading id="D0EC6D31FF78FF24FF64FE2BAC68EBD7" bold="true" box="[189,335,404,430]" fontSize="11" level="3" pageId="221" pageNumber="222" reason="3">
<emphasis id="B96F064FFF78FF24FF64FE2BAC68EBD7" bold="true" box="[189,335,404,430]" pageId="221" pageNumber="222">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF78FF24FF64FE7BA963EFEB" blockId="221.[187,1401,452,1427]" pageId="221" pageNumber="222">
Caulescent herb, to
<quantity id="4CE377B8FF78FF24FE70FE7BACF0EBA6" box="[425,471,452,479]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="221" pageNumber="222" unit="m" value="2.0">2 m</quantity>
high.
<emphasis id="B96F064FFF78FF24FDFDFE7AAF7AEBA6" box="[548,605,453,479]" italics="true" pageId="221" pageNumber="222">Stem</emphasis>
erect, becoming hardened at maturity and occasionally resembling a rhizome, rarely branching; internodes to
<quantity id="4CE377B8FF78FF24FD6AFE56AFC9E87A" box="[691,750,489,516]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="221" pageNumber="222" unit="cm" value="5.0">5 cm</quantity>
long, to
<quantity id="4CE377B8FF78FF24FC8EFE56AE8DE87A" box="[855,938,489,515]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="221" pageNumber="222" unit="mm" value="30.0">30 mm</quantity>
thick, succulent, pale brown, glabrous.
<emphasis id="B96F064FFF78FF24FF64FDB2AC3CE85E" box="[189,283,525,551]" italics="true" pageId="221" pageNumber="222">Stipules</emphasis>
persistent, oblong-ovate to triangular, 821 ×
<quantity id="4CE377B8FF78FF24FCE7FDB2AE84E851" box="[830,931,525,552]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="221" pageNumber="222" unit="mm" value="5.5" valueMax="8.0" valueMin="3.0">38 mm</quantity>
, apex acute, setose, opaque, pale green, glabrous, margin entire, aciliate.
<emphasis id="B96F064FFF78FF24FD8CFD8DAF80E832" box="[597,679,562,587]" italics="true" pageId="221" pageNumber="222">Leaves</emphasis>
38 per stem, alternate, basifixed or rarely peltate (never in
<collectingCountry id="F30C9ACDFF78FF24FF06FDE9AC3DE809" box="[223,282,598,624]" name="Peru" pageId="221" pageNumber="222">Peru</collectingCountry>
); petiole
<quantity id="4CE377B8FF78FF24FE57FDE9ACDFE808" box="[398,504,598,625]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.6" metricValueMin="0.5" pageId="221" pageNumber="222" unit="cm" value="10.5" valueMax="16.0" valueMin="5.0">516 cm</quantity>
long, green to red, glabrous; blade asymmetric, transversely broad-ovate to suborbicular, to 20 ×
<quantity id="4CE377B8FF78FF24FE29FDC5AF66E8EC" box="[496,577,634,661]" metricMagnitude="-1" metricUnit="m" metricValue="2.6" pageId="221" pageNumber="222" unit="cm" value="26.0">26 cm</quantity>
, succulent, apex acute to acuminate, base cordate, basal lobes not overlapping, sinus to
<quantity id="4CE377B8FF78FF24FE64FD20AF36E8C3" box="[445,529,671,698]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="221" pageNumber="222" unit="mm" value="35.0">35 mm</quantity>
deep, margin with 47 regular triangular lobes spaced evenly around the margin, serrate, ciliate, upper surface green, sometimes flushed silver between the veins, sparsely pilose, lower surface pale green to purple, glabrous to pilose, veins palmate-pinnate, 46 veined from the base, with 24 secondary veins on the larger side, 13 secondary veins on the smaller side.
<emphasis id="B96F064FFF78FF24FB43FCB3A865E95F" box="[1178,1346,780,806]" italics="true" pageId="221" pageNumber="222">Inflorescences</emphasis>
14 per stem, bisexual, axillary, erect, cymose, with up to 6 branches, bearing up to 16 staminate flowers and 16 pistillate flowers, protandrous; peduncle to
<quantity id="4CE377B8FF78FF24FCCCFCEAAE78E909" box="[789,863,853,880]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" pageId="221" pageNumber="222" unit="cm" value="19.0">19 cm</quantity>
long, pale green to red, glabrous or minutely glandular pubescent, bracts deciduous, elliptic to oblong, 59 ×
<quantity id="4CE377B8FF78FF24FC15FCC6A96DE9ED" box="[972,1098,889,916]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="5.0" metricValueMin="1.5" pageId="221" pageNumber="222" unit="mm" value="3.25" valueMax="5.0" valueMin="1.5">1.55 mm</quantity>
, opaque, green to white, minutely glandular pubescent, apex acute, margin entire, aciliate.
<emphasis id="B96F064FFF78FF24FC64FC21A9B7E9C1" box="[957,1168,926,952]" italics="true" pageId="221" pageNumber="222">Staminate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FF78FF24FAFCFC21A850E9C0" box="[1317,1399,926,953]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="221" pageNumber="222" unit="mm" value="14.0">14 mm</quantity>
long, glabrous; tepals 4, spreading, outer 2 broadly ovate, 513 ×
<quantity id="4CE377B8FF78FF24FC1EFC7CA91DE9A4" box="[967,1082,963,989]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="221" pageNumber="222" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">710 mm</quantity>
, apex rounded, pale green, glabrous to minutely glandular pubescent, margin entire, aciliate, inner 2 elliptic, 57 ×
<quantity id="4CE377B8FF78FF24FB16FC58A813EE7B" box="[1231,1332,999,1026]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="221" pageNumber="222" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
, apex rounded, pale green, glabrous, margin entire, aciliate; stamens 75100, spreading, yellow, filaments
<quantity id="4CE377B8FF78FF24FF64FB8FAC63EE33" box="[189,324,1072,1099]" metricMagnitude="-3" metricUnit="m" metricValue="1.625" metricValueMax="2.0" metricValueMin="1.25" pageId="221" pageNumber="222" unit="mm" value="1.625" valueMax="2.0" valueMin="1.25">1.252 mm</quantity>
long, free, anthers obovoid, 0.50.75 ×
<quantity id="4CE377B8FF78FF24FCF9FB8FAE59EE33" box="[800,894,1072,1098]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="221" pageNumber="222" unit="mm" value="0.3">0.3 mm</quantity>
, dehiscing via lateral slits, connectives not extended, symmetrically basifixed.
<emphasis id="B96F064FFF78FF24FDB9FBEBAE0EEE17" box="[608,809,1108,1134]" italics="true" pageId="221" pageNumber="222">Pistillate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FF78FF24FC1BFBEBA932EE16" box="[962,1045,1108,1135]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="221" pageNumber="222" unit="mm" value="15.0">15 mm</quantity>
long; bracteoles 2 or lacking, directly beneath the ovary, elliptic, ca 3 ×
<quantity id="4CE377B8FF78FF24FD64FBC6AE21EEEA" box="[701,774,1145,1171]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="221" pageNumber="222" unit="mm" value="1.0">1 mm</quantity>
, opaque, white, glabrous, apex acute, margin entire, aciliate; tepals 5, subequal, deciduous in fruit, spreading, elliptic to ovate, 59 ×
<quantity id="4CE377B8FF78FF24FB8FFB22A99FEEC1" box="[1110,1208,1181,1208]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="221" pageNumber="222" unit="mm" value="4.0" valueMax="6.0" valueMin="2.0">26 mm</quantity>
, apex acute, pale green, glabrous, margin entire, aciliate; ovary body ellipsoid, 410 ×
<quantity id="4CE377B8FF78FF24FC32FB7DA975EEA5" box="[1003,1106,1218,1244]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="221" pageNumber="222" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">37 mm</quantity>
, pale green flushed pink, glandular-pubescent, unequally 3-winged, wings semi-circular to triangular, largest 29 ×
<quantity id="4CE377B8FF78FF24FB24FB59A855EF78" box="[1277,1394,1254,1281]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="221" pageNumber="222" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">812 mm</quantity>
, smallest 17 ×
<quantity id="4CE377B8FF78FF24FEB6FAB4ACC5EF5C" box="[367,482,1291,1317]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="2.1" metricValueMin="0.8" pageId="221" pageNumber="222" unit="mm" value="14.5" valueMax="21.0" valueMin="8.0">821 mm</quantity>
; 3-locular, placentae branches divided, bearing ovules on both surfaces; styles 3, yellow, free,
<quantity id="4CE377B8FF78FF24FEA5FA90ACD0EF30" box="[380,503,1327,1354]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="221" pageNumber="222" unit="mm" value="3.25" valueMax="3.5" valueMin="3.0">33.5 mm</quantity>
long, once-divided, stigmatic papillae in a spirally twisted band.
<emphasis id="B96F064FFF78FF24FACDFA8FAC34EF14" italics="true" pageId="221" pageNumber="222">Fruiting pedicel</emphasis>
to
<quantity id="4CE377B8FF78FF24FEE3FAEBACAAEF17" box="[314,397,1364,1390]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="221" pageNumber="222" unit="mm" value="30.0">30 mm</quantity>
long.
<emphasis id="B96F064FFF78FF24FE00FAEBAF7FEF14" box="[473,600,1363,1389]" italics="true" pageId="221" pageNumber="222">Fruit body</emphasis>
ovoid, to 11 ×
<quantity id="4CE377B8FF78FF24FCD7FAEBAE71EF17" box="[782,854,1364,1390]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="221" pageNumber="222" unit="mm" value="8.0">8 mm</quantity>
, drying brown, wings same shape as in ovary, the largest expanding to 15 ×
<quantity id="4CE377B8FF78FF24FDC3FAC7AF57EFEB" box="[538,624,1400,1426]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="221" pageNumber="222" unit="mm" value="18.0">18 mm</quantity>
, the smallest expanding to 12 ×
<quantity id="4CE377B8FF78FF24FC33FAC7A967EFEB" box="[1002,1088,1400,1426]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="221" pageNumber="222" unit="mm" value="18.0">18 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF78FF24FF64FA04AE42EC7D" pageId="221" pageNumber="222" type="conservation">
<paragraph id="8BA4DA5DFF78FF24FF64FA04AF4AEFAC" blockId="221.[189,621,1466,1493]" box="[189,621,1466,1493]" pageId="221" pageNumber="222">
<heading id="D0EC6D31FF78FF24FF64FA04AF4AEFAC" bold="true" box="[189,621,1466,1493]" fontSize="11" level="3" pageId="221" pageNumber="222" reason="3">
<emphasis id="B96F064FFF78FF24FF64FA04AF4AEFAC" bold="true" box="[189,621,1466,1493]" pageId="221" pageNumber="222">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF78FF24FF64FA55AE42EC7D" blockId="221.[189,869,1514,1541]" box="[189,869,1514,1541]" pageId="221" pageNumber="222">
Assessed by
<bibRefCitation id="EF8AA7ACFF78FF24FE8DFA55AF6BEC7D" author="Tebbitt" box="[340,588,1514,1540]" etAl="et al." firstAuthor="Tebbitt" pageId="221" pageNumber="222" pagination="215 - 226" refId="ref232737" refString="Tebbitt M. C., Andrada A. R., Kollmann J. C. &amp; Moonlight P. W. 2018 a. Taxonomy of Begonia wollnyi Herzog and Begonia arrogans Irmsch. Edinburgh Journal of Botany 75 (2): 215 - 226. https: // doi. org / 10.1017 / S 0960428618000069" type="journal article" year="2018" yearSuffix="a">
Tebbitt
<emphasis id="B96F064FFF78FF24FE74FA54ACCEEC7D" box="[429,489,1514,1540]" italics="true" pageId="221" pageNumber="222">et al.</emphasis>
(2018a)
</bibRefCitation>
as Least Concern (LC).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF78FF26FF64F992A86BEB9A" lastPageId="223" lastPageNumber="224" pageId="221" pageNumber="222" type="discussion">
<paragraph id="8BA4DA5DFF78FF24FF64F992ACABEC3E" blockId="221.[189,396,1581,1607]" box="[189,396,1581,1607]" pageId="221" pageNumber="222">
<heading id="D0EC6D31FF78FF24FF64F992ACABEC3E" bold="true" box="[189,396,1581,1607]" fontSize="11" level="3" pageId="221" pageNumber="222" reason="3">
<emphasis id="B96F064FFF78FF24FF64F992ACABEC3E" bold="true" box="[189,396,1581,1607]" pageId="221" pageNumber="222">Synonymy notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF78FF24FF64F9E2AE54ECE2" blockId="221.[189,1399,1629,1692]" pageId="221" pageNumber="222">
The synonyms of
<taxonomicName id="4C1BA1DEFF78FF24FE4EF9E1AF36EC0E" box="[407,529,1629,1655]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="221" pageNumber="222" phylum="Tracheophyta" rank="species" species="wollnyi">
<emphasis id="B96F064FFF78FF24FE4EF9E1AF36EC0E" box="[407,529,1629,1655]" italics="true" pageId="221" pageNumber="222">B. wollnyi</emphasis>
</taxonomicName>
and the typification of
<taxonomicName id="4C1BA1DEFF78FF24FCF3F9E1A934EC0E" authority="Irmsch." authorityName="Irmsch." box="[810,1043,1629,1655]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="221" pageNumber="222" phylum="Tracheophyta" rank="species" species="acrensis">
<emphasis id="B96F064FFF78FF24FCF3F9E1AE88EC0E" box="[810,943,1630,1655]" italics="true" pageId="221" pageNumber="222">B. acrensis</emphasis>
Irmsch.
</taxonomicName>
and
<taxonomicName id="4C1BA1DEFF78FF24FB94F9E1AC60ECE2" authority="L. B. Sm. &amp; B. G. Schub." authorityName="L. B. Sm. &amp; B. G. Schub." class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="221" pageNumber="222" phylum="Tracheophyta" rank="species" species="parodiana">
<emphasis id="B96F064FFF78FF24FB94F9E1A9CDEC0E" box="[1101,1258,1629,1655]" italics="true" pageId="221" pageNumber="222">B. parodiana</emphasis>
L.B.Sm. &amp; B.G.Schub.
</taxonomicName>
are discussed in detail in
<bibRefCitation id="EF8AA7ACFF78FF24FDACF93EAE57ECE2" author="Tebbitt" box="[629,880,1665,1691]" etAl="et al." firstAuthor="Tebbitt" pageId="221" pageNumber="222" pagination="215 - 226" refId="ref232737" refString="Tebbitt M. C., Andrada A. R., Kollmann J. C. &amp; Moonlight P. W. 2018 a. Taxonomy of Begonia wollnyi Herzog and Begonia arrogans Irmsch. Edinburgh Journal of Botany 75 (2): 215 - 226. https: // doi. org / 10.1017 / S 0960428618000069" type="journal article" year="2018" yearSuffix="a">
Tebbitt
<emphasis id="B96F064FFF78FF24FD17F93DAE2EECE2" box="[718,777,1665,1691]" italics="true" pageId="221" pageNumber="222">et al.</emphasis>
(2018a)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BA4DA5DFF78FF24FF64F97BACBAECA7" blockId="221.[189,413,1732,1758]" box="[189,413,1732,1758]" pageId="221" pageNumber="222">
<heading id="D0EC6D31FF78FF24FF64F97BACBAECA7" bold="true" box="[189,413,1732,1758]" fontSize="11" level="3" pageId="221" pageNumber="222" reason="3">
<emphasis id="B96F064FFF78FF24FF64F97BACBAECA7" bold="true" box="[189,413,1732,1758]" pageId="221" pageNumber="222">Typification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF78FF24FF64F94BA9F1ED91" blockId="221.[189,1400,1780,2025]" pageId="221" pageNumber="222">
The protologue of
<taxonomicName id="4C1BA1DEFF78FF24FE42F94BAF33ED77" box="[411,532,1780,1806]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="221" pageNumber="222" phylum="Tracheophyta" rank="species" species="wollnyi">
<emphasis id="B96F064FFF78FF24FE42F94BAF33ED77" box="[411,532,1780,1806]" italics="true" pageId="221" pageNumber="222">B. wollnyi</emphasis>
</taxonomicName>
does not cite an herbarium thus it is appropriate to designate a
<typeStatus id="54A064FFFF78FF24FADCF94BA853ED77" box="[1285,1396,1780,1806]" pageId="221" pageNumber="222">lectotype</typeStatus>
.
<bibRefCitation id="EF8AA7ACFF78FF24FF67F8A7AC92ED4B" author="Tebbitt" box="[190,437,1816,1842]" etAl="et al." firstAuthor="Tebbitt" pageId="221" pageNumber="222" pagination="215 - 226" refId="ref232737" refString="Tebbitt M. C., Andrada A. R., Kollmann J. C. &amp; Moonlight P. W. 2018 a. Taxonomy of Begonia wollnyi Herzog and Begonia arrogans Irmsch. Edinburgh Journal of Botany 75 (2): 215 - 226. https: // doi. org / 10.1017 / S 0960428618000069" type="journal article" year="2018" yearSuffix="a">
Tebbitt
<emphasis id="B96F064FFF78FF24FECEF8A6AC75ED4B" box="[279,338,1816,1842]" italics="true" pageId="221" pageNumber="222">et al.</emphasis>
(2018a)
</bibRefCitation>
treated a sheet of the type collection held in Z as a
<typeStatus id="54A064FFFF78FF24FBC9F8A7A950ED4B" box="[1040,1143,1816,1842]" pageId="221" pageNumber="222">holotype</typeStatus>
and we are not aware of any other duplicates of this specimen. We designate this sheet as the
<typeStatus id="54A064FFFF78FF24FBE8F883A9BAED2F" box="[1073,1181,1852,1878]" pageId="221" pageNumber="222">lectotype</typeStatus>
collection herein.
<bibRefCitation id="EF8AA7ACFF78FF24FF64F8DEAC9AED02" author="Tebbitt" box="[189,445,1889,1915]" etAl="et al." firstAuthor="Tebbitt" pageId="221" pageNumber="222" pagination="215 - 226" refId="ref232737" refString="Tebbitt M. C., Andrada A. R., Kollmann J. C. &amp; Moonlight P. W. 2018 a. Taxonomy of Begonia wollnyi Herzog and Begonia arrogans Irmsch. Edinburgh Journal of Botany 75 (2): 215 - 226. https: // doi. org / 10.1017 / S 0960428618000069" type="journal article" year="2018" yearSuffix="a">
Tebbitt
<emphasis id="B96F064FFF78FF24FEC3F8DDAC7FED02" box="[282,344,1889,1915]" italics="true" pageId="221" pageNumber="222">et al.</emphasis>
(2018a)
</bibRefCitation>
also attempted to designate a
<typeStatus id="54A064FFFF78FF24FCE8F8DEAEBAED02" box="[817,925,1889,1915]" pageId="221" pageNumber="222">lectotype</typeStatus>
for
<taxonomicName id="4C1BA1DEFF78FF24FC0AF8DDA851ED02" authority="Rusby &amp; Nash. They" authorityName="Rusby &amp; Nash. They" box="[979,1398,1889,1915]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="221" pageNumber="222" phylum="Tracheophyta" rank="species" species="williamsii">
<emphasis id="B96F064FFF78FF24FC0AF8DDA94BED02" box="[979,1132,1889,1915]" italics="true" pageId="221" pageNumber="222">B. williamsii</emphasis>
Rusby &amp; Nash. They
</taxonomicName>
chose
<emphasis id="B96F064FFF78FF24FED1F839ACFBEDE6" box="[264,476,1925,1952]" italics="true" pageId="221" pageNumber="222">R.S. Williams 600</emphasis>
(NY), unaware that this sheet had already been designated as the
<typeStatus id="54A064FFFF78FF24FB33F83AA871EDE6" box="[1258,1366,1925,1951]" pageId="221" pageNumber="222">lectotype</typeStatus>
of this name.
<bibRefCitation id="EF8AA7ACFF78FF24FEE2F815AF46EDBD" author="Smith &amp; Schubert" box="[315,609,1962,1988]" firstAuthor="Smith" pageId="221" pageNumber="222" pagination="71 - 87" refId="ref231295" refString="Smith L. B. &amp; Schubert B. G. 1944. Revision de las especies Bolivianas del genero Begonia. Revistas Cientificas de la Universidad Andina del Cusco 33 (87): 71 - 87." type="journal article" year="1944">Smith &amp; Schubert (1944)</bibRefCitation>
cited this sheet as the “type”, which prior to 2001 this was sufficient to be considered lectotypification. We therefore consider this the date of lectotypification.
</paragraph>
<caption id="DF648AD5FF7BFF27FF64F8FEAF27ED91" ID-DOI="http://doi.org/10.5281/zenodo.8178450" ID-Zenodo-Dep="8178450" httpUri="https://zenodo.org/record/8178450/files/figure.png" pageId="222" pageNumber="223" startId="222.[189,232,1857,1883]" targetBox="[246,1342,274,1819]" targetPageId="222">
<paragraph id="8BA4DA5DFF7BFF27FF64F8FEAF27ED91" blockId="222.[189,1399,1857,2024]" pageId="222" pageNumber="223">
<emphasis id="B96F064FFF7BFF27FF64F8FEAC3AED22" bold="true" box="[189,285,1857,1883]" pageId="222" pageNumber="223">Fig. 70.</emphasis>
<taxonomicName id="4C1BA1DEFF7BFF27FEFEF8FDAF73ED25" authority="Herzog." authorityName="Herzog." box="[295,596,1857,1884]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="222" pageNumber="223" phylum="Tracheophyta" rank="species" species="wollnyi">
<emphasis id="B96F064FFF7BFF27FEFEF8FDACCCED22" box="[295,491,1857,1883]" italics="true" pageId="222" pageNumber="223">Begonia wollnyi</emphasis>
Herzog.
</taxonomicName>
<emphasis id="B96F064FFF7BFF27FD84F8FEAF4BED22" bold="true" box="[605,620,1857,1883]" pageId="222" pageNumber="223">1</emphasis>
. Habit.
<emphasis id="B96F064FFF7BFF27FD09F8FEAFF8ED22" bold="true" box="[720,735,1857,1883]" pageId="222" pageNumber="223">2</emphasis>
. Leaf.
<emphasis id="B96F064FFF7BFF27FCEEF8FEAE61ED22" bold="true" box="[823,838,1857,1883]" pageId="222" pageNumber="223">3</emphasis>
. Staminate flower, front view.
<emphasis id="B96F064FFF7BFF27FB67F8FEA9EAED22" bold="true" box="[1214,1229,1857,1883]" pageId="222" pageNumber="223">4</emphasis>
. Androecium, side view.
<emphasis id="B96F064FFF7BFF27FEE5F8DAAC40ED06" bold="true" box="[316,359,1893,1919]" pageId="222" pageNumber="223">57</emphasis>
. Stamens, side view.
<emphasis id="B96F064FFF7BFF27FDB1F8DAAF51ED06" bold="true" box="[616,630,1893,1919]" pageId="222" pageNumber="223">8</emphasis>
. Pistillate flower, side view.
<emphasis id="B96F064FFF7BFF27FC12F8DAAEFDED06" bold="true" box="[971,986,1893,1919]" pageId="222" pageNumber="223">9</emphasis>
. Cross section of ovary, side view.
<emphasis id="B96F064FFF7BFF27FF67F837AC22EDDB" bold="true" box="[190,261,1928,1954]" pageId="222" pageNumber="223">1011</emphasis>
. Style, front and back view.
<emphasis id="B96F064FFF7BFF27FD97F837AF4CEDDB" bold="true" box="[590,619,1928,1954]" pageId="222" pageNumber="223">12</emphasis>
. Cross section of ovary. Reproduction of an illustration of
<taxonomicName id="4C1BA1DEFF7BFF27FACAF836ACD4EDBC" authority="Rusby &amp; Nash" authorityName="Rusby &amp; Nash" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="222" pageNumber="223" phylum="Tracheophyta" rank="species" species="williamsii">
<emphasis id="B96F064FFF7BFF27FACAF836AC15EDBC" italics="true" pageId="222" pageNumber="223">Begonia williamsii</emphasis>
Rusby &amp; Nash
</taxonomicName>
by an unknown artist reproduced from
<bibRefCitation id="EF8AA7ACFF7BFF27FC3AF814A953EDBC" author="Nash" box="[995,1140,1963,1989]" firstAuthor="Nash" pageId="222" pageNumber="223" pagination="355 - 360" refId="ref230318" refString="Nash G. V. 1916. Begonia williamsii, William's Begonia, native of Bolivia Addisonia 1: 37 - 38. Otto C. F. &amp; Dietrich A. G. 1836. Begonien. Allgemeine Gartenzeitung 4 (45): 355 - 360." type="journal article" year="1916">Nash (1916</bibRefCitation>
: pl. 29) at the Royal Botanic Garden Edinburgh.
</paragraph>
</caption>
<paragraph id="8BA4DA5DFF7AFF26FF64FEB7AC97EB5A" blockId="223.[189,432,264,291]" box="[189,432,264,291]" pageId="223" pageNumber="224">
<heading id="D0EC6D31FF7AFF26FF64FEB7AC97EB5A" bold="true" box="[189,432,264,291]" fontSize="11" level="3" pageId="223" pageNumber="224" reason="3">
<emphasis id="B96F064FFF7AFF26FF64FEB7AC97EB5A" bold="true" box="[189,432,264,291]" pageId="223" pageNumber="224">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF7AFF26FF64FE86A86BEB9A" blockId="223.[189,1399,312,483]" pageId="223" pageNumber="224">
Most similar to
<taxonomicName id="4C1BA1DEFF7AFF26FE58FE86AF36EB2B" box="[385,529,313,338]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="arrogans">
<emphasis id="B96F064FFF7AFF26FE58FE86AF36EB2B" box="[385,529,313,338]" italics="true" pageId="223" pageNumber="224">B. arrogans</emphasis>
</taxonomicName>
(see Identification notes for
<taxonomicName id="4C1BA1DEFF7AFF26FCA9FE86A927EB2B" box="[880,1024,313,338]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="arrogans">
<emphasis id="B96F064FFF7AFF26FCA9FE86A927EB2B" box="[880,1024,313,338]" italics="true" pageId="223" pageNumber="224">B. arrogans</emphasis>
</taxonomicName>
). It is also possible to confuse
<taxonomicName id="4C1BA1DEFF7AFF26FF64FEE2AC12EB0F" box="[189,309,348,374]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="wollnyi">
<emphasis id="B96F064FFF7AFF26FF64FEE2AC12EB0F" box="[189,309,348,374]" italics="true" pageId="223" pageNumber="224">B. wollnyi</emphasis>
</taxonomicName>
with
<taxonomicName id="4C1BA1DEFF7AFF26FEA0FEE2AF2AEB0F" box="[377,525,348,374]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="acerifolia">
<emphasis id="B96F064FFF7AFF26FEA0FEE2AF2AEB0F" box="[377,525,348,374]" italics="true" pageId="223" pageNumber="224">B. acerifolia</emphasis>
</taxonomicName>
. Populations of
<taxonomicName id="4C1BA1DEFF7AFF26FD16FEE2AE43EB0F" box="[719,868,348,374]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="acerifolia">
<emphasis id="B96F064FFF7AFF26FD16FEE2AE43EB0F" box="[719,868,348,374]" italics="true" pageId="223" pageNumber="224">B. acerifolia</emphasis>
</taxonomicName>
overlap with those of
<taxonomicName id="4C1BA1DEFF7AFF26FBB4FEE2A9C2EB0F" box="[1133,1253,348,374]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="wollnyi">
<emphasis id="B96F064FFF7AFF26FBB4FEE2A9C2EB0F" box="[1133,1253,348,374]" italics="true" pageId="223" pageNumber="224">B. wollnyi</emphasis>
</taxonomicName>
in the south of
<collectingCountry id="F30C9ACDFF7AFF26FF07FE3EAC33EBE2" box="[222,276,385,411]" name="Peru" pageId="223" pageNumber="224">Peru</collectingCountry>
and
<collectingCountry id="F30C9ACDFF7AFF26FE97FE3FAC8FEBE3" box="[334,424,384,410]" name="Bolivia" pageId="223" pageNumber="224">Bolivia</collectingCountry>
. Individuals in these populations of
<taxonomicName id="4C1BA1DEFF7AFF26FC8DFE3EAECFEBE3" box="[852,1000,384,410]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="acerifolia">
<emphasis id="B96F064FFF7AFF26FC8DFE3EAECFEBE3" box="[852,1000,384,410]" italics="true" pageId="223" pageNumber="224">B. acerifolia</emphasis>
</taxonomicName>
almost always has peltate leaves, whereas those of
<taxonomicName id="4C1BA1DEFF7AFF26FE50FE1AAF26EBC7" box="[393,513,420,446]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="wollnyi">
<emphasis id="B96F064FFF7AFF26FE50FE1AAF26EBC7" box="[393,513,420,446]" italics="true" pageId="223" pageNumber="224">B. wollnyi</emphasis>
</taxonomicName>
are basifixed. Furthermore, two of the three wings on the ovaries and fruits of
<taxonomicName id="4C1BA1DEFF7AFF26FF04FE76AC56EB9A" box="[221,369,457,483]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="acerifolia">
<emphasis id="B96F064FFF7AFF26FF04FE76AC56EB9A" box="[221,369,457,483]" italics="true" pageId="223" pageNumber="224">B. acerifolia</emphasis>
</taxonomicName>
are reduced to ridges whereas those of
<taxonomicName id="4C1BA1DEFF7AFF26FC9BFE76AE9DEB9A" box="[834,954,457,483]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="wollnyi">
<emphasis id="B96F064FFF7AFF26FC9BFE76AE9DEB9A" box="[834,954,457,483]" italics="true" pageId="223" pageNumber="224">B. wollnyi</emphasis>
</taxonomicName>
have three well-developed wings.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF7AFF26FF64FDB4AE09EF2B" pageId="223" pageNumber="224" type="distribution">
<paragraph id="8BA4DA5DFF7AFF26FF64FDB4ACD2E85C" blockId="223.[189,501,523,549]" box="[189,501,523,549]" pageId="223" pageNumber="224">
<heading id="D0EC6D31FF7AFF26FF64FDB4ACD2E85C" bold="true" box="[189,501,523,549]" fontSize="11" level="3" pageId="223" pageNumber="224" reason="3">
<emphasis id="B96F064FFF7AFF26FF64FDB4ACD2E85C" bold="true" box="[189,501,523,549]" pageId="223" pageNumber="224">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF7AFF26FF64FD84A9C2E8E4" blockId="223.[189,1399,570,669]" pageId="223" pageNumber="224">
Known from
<collectingCountry id="F30C9ACDFF7AFF26FEBBFD84ACBCE82C" box="[354,411,571,597]" name="Peru" pageId="223" pageNumber="224">Peru</collectingCountry>
,
<collectingCountry id="F30C9ACDFF7AFF26FE73FD85AF01E82D" box="[426,550,570,596]" name="Venezuela" pageId="223" pageNumber="224">Venezuela</collectingCountry>
, and
<collectingCountry id="F30C9ACDFF7AFF26FDB3FD85AF92E82D" box="[618,693,570,596]" name="Brazil" pageId="223" pageNumber="224">Brazil</collectingCountry>
, where it has a peri-Amazonian distribution. Within
<collectingCountry id="F30C9ACDFF7AFF26FAE0FD84A855E82C" box="[1337,1394,571,597]" name="Peru" pageId="223" pageNumber="224">Peru</collectingCountry>
, known from a single population in
<collectingRegion id="49DF14BFFF7AFF26FD85FDE0AE25E800" box="[604,770,607,633]" country="Peru" name="Cusco" pageId="223" pageNumber="224">Cusco Region</collectingRegion>
(
<figureCitation id="1320C6D8FF7AFF26FCCBFDE0AEA7E803" box="[786,896,607,634]" captionStart="Fig" captionStartId="204.[189,232,1822,1848]" captionTargetBox="[194,1392,558,1776]" captionTargetId="figure-14@204.[189,1398,548,1784]" captionTargetPageId="204" captionText="Fig. 65. Distribution of the acerifolia group of Begonia sect. Knesebeckia (Klotzsch) A.DC. in Peru and surrounding countries. A. B. acerifolia Kunth (red). B. B. arrogans Irmsch. (red) and B. ludwigii Irmsch. (blue). C. B. monadelpha (Ruiz ex Klotzsch) A.DC. (red) and B. serotina A.DC. (blue). D. B. huancabambae Moonlight sp. nov. (red) and B. wollnyi Herzog (blue). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178440" httpUri="https://zenodo.org/record/8178440/files/figure.png" pageId="223" pageNumber="224">Fig. 65D</figureCitation>
). Found in Amazonian forest at an altitude of
<quantity id="4CE377B8FF7AFF26FF04FD3CAC45E8E4" box="[221,354,643,669]" metricMagnitude="2" metricUnit="m" metricValue="4.335" metricValueMax="4.67" metricValueMin="4.0" pageId="223" pageNumber="224" unit="m" value="433.5" valueMax="467.0" valueMin="400.0">400467 m</quantity>
a.s.l.
<taxonomicName id="4C1BA1DEFF7AFF26FE7EFD3BAF4FE8E4" box="[423,616,643,669]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="wollnyi">
<emphasis id="B96F064FFF7AFF26FE7EFD3BAF4FE8E4" box="[423,616,643,669]" italics="true" pageId="223" pageNumber="224">Begonia wollnyi</emphasis>
</taxonomicName>
drops its leaves and usually flowers in the dry season.
</paragraph>
<paragraph id="8BA4DA5DFF7AFF26FF64FD79AE4EE89A" blockId="223.[189,873,710,739]" box="[189,873,710,739]" pageId="223" pageNumber="224">
<emphasis id="B96F064FFF7AFF26FF64FD79AE4EE89A" bold="true" box="[189,873,710,739]" italics="true" pageId="223" pageNumber="224">
The
<taxonomicName id="4C1BA1DEFF7AFF26FF20FD78ACA2E89A" box="[249,389,711,739]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="maynensis">maynensis</taxonomicName>
group of
<taxonomicName id="4C1BA1DEFF7AFF26FDD1FD78AE4EE89A" box="[520,873,710,739]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="section" section="Knesebeckia">Begonia sect. Knesebeckia</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA4DA5DFF7AFF26FF64FD43AE88EE6A" blockId="223.[188,1400,764,1043]" pageId="223" pageNumber="224">
The maynensis group of
<emphasis id="B96F064FFF7AFF26FE33FD42AF23E96F" box="[490,516,765,790]" italics="true" pageId="223" pageNumber="224">B.</emphasis>
sect.
<taxonomicName id="4C1BA1DEFF7AFF26FD90FD43AFF8E96F" box="[585,735,764,790]" class="Magnoliopsida" family="Begoniaceae" genus="Knesebeckia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F064FFF7AFF26FD90FD43AFF8E96F" box="[585,735,764,790]" italics="true" pageId="223" pageNumber="224">Knesebeckia</emphasis>
</taxonomicName>
was revised and informally named by
<bibRefCitation id="EF8AA7ACFF7AFF26FB6DFD43AC39E943" author="Moonlight" etAl="et al." firstAuthor="Moonlight" pageId="223" pageNumber="224" pagination="179 - 198" refId="ref229967" refString="Moonlight P. W., Hughes M. &amp; Tebbitt M. C. 2017 b. Taxonomy of Begonia albomaculata and description of two new species endemic to Peru. Edinburgh Journal of Botany 74 (2): 179 - 198. https: // doi. org / 10.1017 / S 0960428617000075" type="journal article" year="2017" yearSuffix="b">
Moonlight
<emphasis id="B96F064FFF7AFF26FAE2FD42A85FE96F" box="[1339,1400,764,790]" italics="true" pageId="223" pageNumber="224">et al.</emphasis>
(2017b)
</bibRefCitation>
. It includes four species within
<emphasis id="B96F064FFF7AFF26FD7FFC9EAFE7E943" box="[678,704,801,826]" italics="true" pageId="223" pageNumber="224">B.</emphasis>
sect.
<taxonomicName id="4C1BA1DEFF7AFF26FCD2FC9FAE86E943" box="[779,929,800,826]" class="Magnoliopsida" family="Begoniaceae" genus="Knesebeckia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F064FFF7AFF26FCD2FC9FAE86E943" box="[779,929,800,826]" italics="true" pageId="223" pageNumber="224">Knesebeckia</emphasis>
</taxonomicName>
that are distinguished by their simple, unlobed leaves that are clustered towards the apex of the stem and their conspicuous, relatively persistent stipules. This concept of the group is mostly Peruvian but includes species also found in
<collectingCountry id="F30C9ACDFF7AFF26FF64FC33AC38E9DF" box="[189,287,908,934]" name="Ecuador" pageId="223" pageNumber="224">Ecuador</collectingCountry>
and
<collectingCountry id="F30C9ACDFF7AFF26FE80FC33AC83E9DF" box="[345,420,908,934]" name="Brazil" pageId="223" pageNumber="224">Brazil</collectingCountry>
. Recent phylogenetic evidence suggests that the maynensis group is monophyletic, though distantly related to the
<typeStatus id="54A064FFFF7AFF26FDFFFC0EAF7EE9B2" box="[550,601,945,971]" pageId="223" pageNumber="224">type</typeStatus>
species of
<emphasis id="B96F064FFF7AFF26FD04FC0EAFD1E9B3" box="[733,758,945,970]" italics="true" pageId="223" pageNumber="224">B.</emphasis>
sect.
<taxonomicName id="4C1BA1DEFF7AFF26FCE3FC0EA9D6E9B2" authority="(Moonlight et al. 2018)" baseAuthorityName="Moonlight" baseAuthorityYear="2018" box="[826,1265,945,971]" class="Magnoliopsida" family="Begoniaceae" genus="Knesebeckia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F064FFF7AFF26FCE3FC0EAEF7E9B2" box="[826,976,945,971]" italics="true" pageId="223" pageNumber="224">Knesebeckia</emphasis>
(
<bibRefCitation id="EF8AA7ACFF7AFF26FC39FC0EA9CDE9B2" author="Moonlight P. W. &amp; Reynel C." box="[992,1258,945,971]" journalOrPublisher="Phytotaxa" pageId="223" pageNumber="224" pagination="116 - 126" part="381" refId="ref229806" refString="Moonlight P. W. &amp; Reynel C. 2018. Two new species of Begonia from Peru. Phytotaxa 381 (1): 116 - 126. https: // doi. org / 10.11646 / phytotaxa. 381.1.15" title="Two new species of Begonia from Peru" type="journal article" year="2018">
Moonlight
<emphasis id="B96F064FFF7AFF26FBBFFC0EA985E9B2" box="[1126,1186,945,971]" italics="true" pageId="223" pageNumber="224">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
. The group should not however be elevated to section without a full taxonomic revision and phylogeny covering all members of
<emphasis id="B96F064FFF7AFF26FE89FC45AC4EEE6A" box="[336,361,1018,1043]" italics="true" pageId="223" pageNumber="224">B.</emphasis>
sect.
<taxonomicName id="4C1BA1DEFF7AFF26FE74FC46AE8CEE6A" authority="sensu Moonlight et al. (2018)" authorityName="Moonlight" authorityYear="2018" box="[429,939,1017,1043]" class="Magnoliopsida" family="Begoniaceae" genus="Knesebeckia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F064FFF7AFF26FE74FC46AF64EE6A" box="[429,579,1017,1043]" italics="true" pageId="223" pageNumber="224">Knesebeckia</emphasis>
sensu
<bibRefCitation id="EF8AA7ACFF7AFF26FD4AFC46AE8CEE6A" author="Moonlight P. W. &amp; Reynel C." box="[659,939,1017,1043]" journalOrPublisher="Phytotaxa" pageId="223" pageNumber="224" pagination="116 - 126" part="381" refId="ref229806" refString="Moonlight P. W. &amp; Reynel C. 2018. Two new species of Begonia from Peru. Phytotaxa 381 (1): 116 - 126. https: // doi. org / 10.11646 / phytotaxa. 381.1.15" title="Two new species of Begonia from Peru" type="journal article" year="2018">
Moonlight
<emphasis id="B96F064FFF7AFF26FCCFFC45AE76EE6A" box="[790,849,1017,1043]" italics="true" pageId="223" pageNumber="224">et al.</emphasis>
(2018)
</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA4DA5DFF7AFF26FF64FB84AE09EF2B" blockId="223.[188,1399,1083,1363]" pageId="223" pageNumber="224">
<bibRefCitation id="EF8AA7ACFF7AFF26FF64FB84ACF8EE2C" author="Moonlight" box="[189,479,1083,1109]" etAl="et al." firstAuthor="Moonlight" pageId="223" pageNumber="224" pagination="179 - 198" refId="ref229967" refString="Moonlight P. W., Hughes M. &amp; Tebbitt M. C. 2017 b. Taxonomy of Begonia albomaculata and description of two new species endemic to Peru. Edinburgh Journal of Botany 74 (2): 179 - 198. https: // doi. org / 10.1017 / S 0960428617000075" type="journal article" year="2017" yearSuffix="b">
Moonlight
<emphasis id="B96F064FFF7AFF26FE99FB83AC5BEE2C" box="[320,380,1083,1109]" italics="true" pageId="223" pageNumber="224">et al.</emphasis>
(2017b)
</bibRefCitation>
did not include three species then only known from
<collectingCountry id="F30C9ACDFF7AFF26FB91FB84A98DEE2C" box="[1096,1194,1083,1109]" name="Ecuador" pageId="223" pageNumber="224">Ecuador</collectingCountry>
in the maynensis group:
<taxonomicName id="4C1BA1DEFF7AFF26FEC8FBDFACFBEE00" box="[273,476,1119,1145]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="brandbygeana">
<emphasis id="B96F064FFF7AFF26FEC8FBDFACFBEE00" box="[273,476,1119,1145]" italics="true" pageId="223" pageNumber="224">B. brandbygeana</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1BA1DEFF7AFF26FE32FBDFAE54EE00" authority="L. B. Sm. &amp; Wassh." authorityName="L. B. Sm. &amp; Wassh." box="[491,883,1119,1146]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="oellgaardii">
<emphasis id="B96F064FFF7AFF26FE32FBDFAFA8EE00" box="[491,655,1119,1145]" italics="true" pageId="223" pageNumber="224">B. oellgaardii</emphasis>
L.B.Sm. &amp; Wassh.
</taxonomicName>
, and
<taxonomicName id="4C1BA1DEFF7AFF26FC76FBDFA80DEE00" authority="L. B. Sm. &amp; Wassh." authorityName="L. B. Sm. &amp; Wassh." box="[943,1322,1119,1146]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="sparreana">
<emphasis id="B96F064FFF7AFF26FC76FBDFA96EEE00" box="[943,1097,1120,1145]" italics="true" pageId="223" pageNumber="224">B. sparreana</emphasis>
L.B.Sm. &amp; Wassh.
</taxonomicName>
These species differ from the maynensis group sensu
<emphasis id="B96F064FFF7AFF26FD23FB3BAE61EEE4" box="[762,838,1156,1181]" italics="true" pageId="223" pageNumber="224">stricto</emphasis>
in their membranaceous leaves and in having non-glabrous leaves. In hindsight, these species should have been revised in
<bibRefCitation id="EF8AA7ACFF7AFF26FBEEFB17A870EEBB" author="Moonlight" box="[1079,1367,1192,1218]" etAl="et al." firstAuthor="Moonlight" pageId="223" pageNumber="224" pagination="179 - 198" refId="ref229967" refString="Moonlight P. W., Hughes M. &amp; Tebbitt M. C. 2017 b. Taxonomy of Begonia albomaculata and description of two new species endemic to Peru. Edinburgh Journal of Botany 74 (2): 179 - 198. https: // doi. org / 10.1017 / S 0960428617000075" type="journal article" year="2017" yearSuffix="b">
Moonlight
<emphasis id="B96F064FFF7AFF26FB61FB16A9D4EEBB" box="[1208,1267,1192,1218]" italics="true" pageId="223" pageNumber="224">et al.</emphasis>
(2017b)
</bibRefCitation>
as they are almost certainly the groups closest relatives. These species have not yet been included in any molecular phylogenies, so their placement remains uncertain. While
<taxonomicName id="4C1BA1DEFF7AFF26FC3FFB4EA994EF73" box="[998,1203,1264,1290]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="223" pageNumber="224" phylum="Tracheophyta" rank="species" species="brandbygeana">
<emphasis id="B96F064FFF7AFF26FC3FFB4EA994EF73" box="[998,1203,1264,1290]" italics="true" pageId="223" pageNumber="224">B. brandbygeana</emphasis>
</taxonomicName>
is now recorded from
<collectingCountry id="F30C9ACDFF7AFF26FF26FAAAAC1FEF56" box="[255,312,1301,1327]" name="Peru" pageId="223" pageNumber="224">Peru</collectingCountry>
, we refrain from including it or the two Ecuadorian endemics in the maynensis group here because this is a floristic and not a sectional revision.
</paragraph>
</subSubSection>
</treatment>
</document>