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<mods:title id="B7B9F8E02728635C8C313D825C5A91E2">The genus Begonia (Begoniaceae) in Peru</mods:title>
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<mods:affiliation id="6D120AC2E6542062A5FD5ABA3ADEE20E">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK. &amp; Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland.</mods:affiliation>
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48. 
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<emphasis id="B96F064FFF63FF3FFD93FDE1AFD9E801" bold="true" box="[586,766,605,632]" italics="true" pageId="198" pageNumber="199">Begonia velata</emphasis>
L.B.Sm. &amp; B.G.Schub.
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<emphasis id="B96F064FFF63FF3FFF64FD7DAEDFE8A4" box="[189,1016,706,733]" italics="true" pageId="198" pageNumber="199">Publications of the Field Museum of Natural History, Botanical Series</emphasis>
13 (4/1): 201 (
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– 
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• 
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&amp; Monsalve (2006: 170)
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.
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<emphasis id="B96F064FFF63FF3FFF64FC2EAC62E9D2" bold="true" box="[189,325,913,939]" pageId="198" pageNumber="199">Etymology</emphasis>
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From the Latin ‘ 
<emphasis id="B96F064FFF63FF3FFE51FC00ACFBE9A0" box="[392,476,959,985]" italics="true" pageId="198" pageNumber="199">velatus</emphasis>
’, meaning ‘partially hidden from view’. This most likely refers to the large bracts of the species, which obscure the young inflorescence.
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<emphasis id="B96F064FFF63FF3FFF64FB9BACE3EE47" bold="true" box="[189,452,1060,1086]" pageId="198" pageNumber="199">Specimens examined</emphasis>
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– 
<emphasis id="B96F064FFF63FF3FFEF5FBECAE14EE14" bold="true" box="[300,819,1106,1133]" pageId="198" pageNumber="199">
<collectingRegion id="49DF14BFFF63FF3FFEF5FBECAF10EE15" box="[300,567,1106,1133]" country="Peru" name="Lambayeque" pageId="198" pageNumber="199">Lambayeque Region</collectingRegion>
: 
<collectingCounty id="62C5A2D1FF63FF3FFD9DFBECAE14EE14" box="[580,819,1107,1133]" pageId="198" pageNumber="199">Prov. Lambayeque</collectingCounty>
</emphasis>
• 
<location id="8EC48C86FF63FF3FFC88FBECA907EE15" LSID="urn:lsid:plazi:treatment:03B26B4BFF63FF30FDC7FDE1AC0DEB31:8EC48C86FF63FF3FFC88FBECA907EE15" box="[849,1056,1106,1133]" country="Peru" county="Prov. Lambayeque" latitude="-6.1" longLatPrecision="1305" longitude="-79.433334" name="Bosque Chinama" pageId="198" pageNumber="199" stateProvince="Lambayeque Region">Bosque Chiñama</location>
; [
<geoCoordinate id="EE2FBC9AFF63FF3FFBEFFBEDA9B6EE14" box="[1078,1169,1105,1133]" degrees="6" direction="south" minutes="06" orientation="latitude" pageId="198" pageNumber="199" precision="925" value="-6.1">6°06′ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFF63FF3FFB79FBEEA830EE14" box="[1184,1303,1105,1133]" degrees="79" direction="west" minutes="26" orientation="longitude" pageId="198" pageNumber="199" precision="925" value="-79.433334">79°26′ W</geoCoordinate>
]; 
<elevation id="00363D6EFF63FF3FFAF4FBEDAC7AEEE9" metricMagnitude="3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="198" pageNumber="199" unit="m" value="2550.0" valueMax="2600.0" valueMin="2500.0">
<quantity id="4CE377B8FF63FF3FFAF4FBEDAC3DEEE9" metricMagnitude="3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="198" pageNumber="199" unit="m" value="2550.0" valueMax="2600.0" valueMin="2500.0">2500– 2600 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFF63FF3FFEB6FBC9AF3EEEE9" box="[367,537,1142,1168]" pageId="198" pageNumber="199" value="1988-08-23">
<collectingDate id="EFE10575FF63FF3FFEB6FBC9AF3EEEE9" box="[367,537,1142,1168]" pageId="198" pageNumber="199" value="1988-08-23">23 Aug. 1988</collectingDate>
</date>
; 
<emphasis id="B96F064FFF63FF3FFDF0FBC9AFF3EEE9" box="[553,724,1142,1168]" italics="true" pageId="198" pageNumber="199">
<collectorName id="26EEBF8BFF63FF3FFDF0FBC9AFA9EEE9" box="[553,654,1142,1168]" pageId="198" pageNumber="199">A. Cano</collectorName>
2087
</emphasis>
; 
<collectionCode id="ED0A4298FF63FF3FFD31FBC9AE2AEEE9" box="[744,781,1142,1168]" pageId="198" pageNumber="199">US</collectionCode>
[
<specimenCode id="DBBD7226FF63FF3FFCFBFBC9AE9AEEE9" box="[802,957,1142,1168]" pageId="198" pageNumber="199">US00222347</specimenCode>
].
</materialsCitation>
– 
<materialsCitation id="3B73D000FF63FF3FFC2AFBCAAE79EE83" collectingDate="2005-05-11" collectionCode="E, HUT, US" collectorName="E. Rodriguez R. &amp; E. Alvitez I. &amp; S. Orroyo A." county="Prov. San Miguel de Pallaques" elevation="3200" latitude="-7.0316668" location="Caserio Agua Blanca" longLatPrecision="21" longitude="-79.06944" municipality="Dist. Agua Blanca" pageId="198" pageNumber="199" specimenCode="E01007287, HUT41989, US00951216" specimenCount="1" stateProvince="Cajamarca Region">
<emphasis id="B96F064FFF63FF3FFC2AFBCAAC9CEECA" bold="true" pageId="198" pageNumber="199">
<collectingRegion id="49DF14BFFF63FF3FFC2AFBCAA9CFEEF6" box="[1011,1256,1141,1167]" inferredBySuffix="true" pageId="198" pageNumber="199">Cajamarca Region</collectingRegion>
: 
<collectingCounty id="62C5A2D1FF63FF3FFB20FBC9AC9CEECA" pageId="198" pageNumber="199">Prov. San Miguel de Pallaques</collectingCounty>
</emphasis>
• 
<collectingMunicipality id="6BC04027FF63FF3FFE08FB26AF8FEECA" box="[465,680,1177,1203]" pageId="198" pageNumber="199">Dist. Agua Blanca</collectingMunicipality>
, 
<location id="8EC48C86FF63FF3FFD6AFB26AE8BEECA" LSID="urn:lsid:plazi:treatment:03B26B4BFF63FF30FDC7FDE1AC0DEB31:8EC48C86FF63FF3FFD6AFB26AE8BEECA" box="[691,940,1177,1203]" county="Prov. San Miguel de Pallaques" latitude="-7.0316668" longLatPrecision="21" longitude="-79.06944" municipality="Dist. Agua Blanca" name="Caserio Agua Blanca" pageId="198" pageNumber="199" stateProvince="Cajamarca Region">Caserío Agua Blanca</location>
, 
<location id="8EC48C86FF63FF3FFC6EFB26A9C1EECA" LSID="urn:lsid:plazi:treatment:03B26B4BFF63FF30FDC7FDE1AC0DEB31:8EC48C86FF63FF3FFC6EFB26A9C1EECA" box="[951,1254,1177,1203]" country="United States of America" county="Prov. San Miguel de Pallaques" latitude="-7.0316668" longLatPrecision="21" longitude="-79.06944" municipality="Dist. Agua Blanca" name="Bosque del Cerro Quillon" pageId="198" pageNumber="199" stateProvince="Cajamarca Region">Bosque del Cerro Quillón</location>
; 
<geoCoordinate id="EE2FBC9AFF63FF3FFB28FB26A856EECA" box="[1265,1393,1175,1203]" degrees="7" direction="south" minutes="01" orientation="latitude" pageId="198" pageNumber="199" precision="15" seconds="54" value="-7.0316668">7°01′54″ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFF63FF3FFF64FB03AC7DEEAE" box="[189,346,1210,1239]" degrees="79" direction="west" minutes="04" orientation="longitude" pageId="198" pageNumber="199" precision="15" seconds="10" value="-79.06944">79°04′10″ W</geoCoordinate>
; 
<elevation id="00363D6EFF63FF3FFEBEFB03AF6AEEAF" box="[359,589,1212,1239]" metricMagnitude="3" metricUnit="m" metricValue="3.2" metricValueMax="3.3" metricValueMin="3.1" pageId="198" pageNumber="199" unit="m" value="3200.0" valueMax="3300.0" valueMin="3100.0">
<quantity id="4CE377B8FF63FF3FFEBEFB03AF2CEEAE" box="[359,523,1212,1239]" metricMagnitude="3" metricUnit="m" metricValue="3.2" metricValueMax="3.3" metricValueMin="3.1" pageId="198" pageNumber="199" unit="m" value="3200.0" valueMax="3300.0" valueMin="3100.0">3100–3300 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFF63FF3FFD85FB03AFD9EEAE" box="[604,766,1212,1239]" pageId="198" pageNumber="199" value="2005-05-11">
<collectingDate id="EFE10575FF63FF3FFD85FB03AFD9EEAE" box="[604,766,1212,1239]" pageId="198" pageNumber="199" value="2005-05-11">11 May 2005</collectingDate>
</date>
; 
<emphasis id="B96F064FFF63FF3FFCD5FB02A849EEAF" box="[780,1390,1212,1238]" italics="true" pageId="198" pageNumber="199">
<collectorName id="26EEBF8BFF63FF3FFCD5FB02AEEAEEAF" box="[780,973,1212,1238]" pageId="198" pageNumber="199">E. Rodríguez R.</collectorName>
, 
<collectorName id="26EEBF8BFF63FF3FFC05FB02A94EEEAF" box="[988,1129,1212,1238]" pageId="198" pageNumber="199">E. Alvítez I.</collectorName>
&amp; 
<collectorName id="26EEBF8BFF63FF3FFB4BFB03A80DEEAF" box="[1170,1322,1212,1238]" pageId="198" pageNumber="199">S. Orroyo A.</collectorName>
2720
</emphasis>
; 
<collectionCode id="ED0A4298FF63FF3FFF64FB5FADE8EE83" box="[189,207,1248,1274]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="198" pageNumber="199" type="Herbarium">E</collectionCode>
[
<specimenCode id="DBBD7226FF63FF3FFF3AFB60AC4CEE80" box="[227,363,1247,1273]" collectionCode="E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="198" pageNumber="199" type="Herbarium">E01007287</specimenCode>
], 
<collectionCode id="ED0A4298FF63FF3FFE5FFB5FACE5EE83" box="[390,450,1248,1274]" country="Japan" httpUri="http://grbio.org/cool/gtk4-xv5b" name="HUT Culture Collection" pageId="198" pageNumber="199">HUT</collectionCode>
[
<specimenCode id="DBBD7226FF63FF3FFE0AFB60AF45EE80" box="[467,610,1247,1273]" collectionCode="HUT" country="Japan" httpUri="http://grbio.org/cool/gtk4-xv5b" name="HUT Culture Collection" pageId="198" pageNumber="199">HUT41989</specimenCode>
], 
<collectionCode id="ED0A4298FF63FF3FFDA1FB60AFBAEE80" box="[632,669,1247,1273]" pageId="198" pageNumber="199">US</collectionCode>
[
<specimenCode id="DBBD7226FF63FF3FFD68FB60AE6BEE83" box="[689,844,1247,1274]" pageId="198" pageNumber="199">US00951216</specimenCode>
].
</materialsCitation>
– 
<materialsCitation id="3B73D000FF63FF3FFC58FB60AF15EF39" collectingDate="1988-04-21" collectionCode="F, MO" collectorName="C. Diaz &amp; S. Baldeon M." county="Prov. Cutervo" elevation="2750" latitude="-6.5166664" location="Carretera entre Llama y Huambos" longLatPrecision="1304" longitude="-79.083336" pageId="198" pageNumber="199" specimenCode="MO-098016" specimenCount="1">
<emphasis id="B96F064FFF63FF3FFC58FB60A911EE80" bold="true" box="[897,1078,1247,1273]" pageId="198" pageNumber="199">
<collectingCounty id="62C5A2D1FF63FF3FFC58FB60A911EE80" box="[897,1078,1247,1273]" pageId="198" pageNumber="199">Prov. Cutervo</collectingCounty>
</emphasis>
• 
<location id="8EC48C86FF63FF3FFB8CFB60AC17EF65" LSID="urn:lsid:plazi:treatment:03B26B4BFF63FF30FDC7FDE1AC0DEB31:8EC48C86FF63FF3FFB8CFB60AC17EF65" county="Prov. Cutervo" latitude="-6.5166664" longLatPrecision="1304" longitude="-79.083336" name="Carretera entre Llama y Huambos" pageId="198" pageNumber="199">Carretera entre Llama y Huambos</location>
, 
<location id="8EC48C86FF63FF3FFEE5FABCACF2EF64" LSID="urn:lsid:plazi:treatment:03B26B4BFF63FF30FDC7FDE1AC0DEB31:8EC48C86FF63FF3FFEE5FABCACF2EF64" box="[316,469,1283,1309]" country="United States of America" county="Prov. Cutervo" latitude="-6.5166664" longLatPrecision="1304" longitude="-79.083336" municipality="Llama" name="Tunaspampa" pageId="198" pageNumber="199">Tunaspampa</location>
; [
<geoCoordinate id="EE2FBC9AFF63FF3FFE31FABEAF64EF64" box="[488,579,1281,1309]" degrees="6" direction="south" minutes="31" orientation="latitude" pageId="198" pageNumber="199" precision="925" value="-6.5166664">6°31′ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFF63FF3FFD89FABCAFE2EF64" box="[592,709,1281,1309]" degrees="79" direction="west" minutes="05" orientation="longitude" pageId="198" pageNumber="199" precision="925" value="-79.083336">79°05′ W</geoCoordinate>
]; 
<elevation id="00363D6EFF63FF3FFD00FABDAE9BEF65" box="[729,956,1282,1309]" metricMagnitude="3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="198" pageNumber="199" unit="m" value="2750.0" valueMax="2900.0" valueMin="2600.0">
<quantity id="4CE377B8FF63FF3FFD00FABDAE5BEF64" box="[729,892,1282,1309]" metricMagnitude="3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="198" pageNumber="199" unit="m" value="2750.0" valueMax="2900.0" valueMin="2600.0">2600–2900 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFF63FF3FFC13FABCA940EF64" box="[970,1127,1283,1309]" pageId="198" pageNumber="199" value="1988-04-21">
<collectingDate id="EFE10575FF63FF3FFC13FABCA940EF64" box="[970,1127,1283,1309]" pageId="198" pageNumber="199" value="1988-04-21">21 Apr. 1988</collectingDate>
</date>
; 
<emphasis id="B96F064FFF63FF3FFBADFABCAC38EF39" italics="true" pageId="198" pageNumber="199">
<collectorName id="26EEBF8BFF63FF3FFBADFABCA9E8EF64" box="[1140,1231,1283,1309]" pageId="198" pageNumber="199">C. Díaz</collectorName>
&amp; 
<collectorName id="26EEBF8BFF63FF3FFB2FFABCADF9EF46" pageId="198" pageNumber="199">S. Baldeón M.</collectorName>
2880
</emphasis>
; 
<collectionCode id="ED0A4298FF63FF3FFEF5FA99AC1AEF39" box="[300,317,1318,1344]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="198" pageNumber="199" type="Herbarium">F</collectionCode>
, 
<collectionCode id="ED0A4298FF63FF3FFE9EFA99AC51EF39" box="[327,374,1318,1344]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="198" pageNumber="199" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FF63FF3FFE5FFA99AF30EF39" box="[390,535,1318,1344]" pageId="198" pageNumber="199">MO-098016</specimenCode>
]).
</materialsCitation>
– 
<materialsCitation id="3B73D000FF63FF3FFD93FA99A800EF1A" collectingDate="1986-03-05" collectionCode="HUT" collectorName="J. Mostacero L. &amp; S. Leiva G. &amp; F. Mejla C. &amp; F. Palaez P." county="Prov. San Miguel de Pallaques" elevation="3200" latitude="-7.016667" location="Cerro Quillon" longLatPrecision="1303" longitude="-79.066666" pageId="198" pageNumber="199" specimenCount="1">
<emphasis id="B96F064FFF63FF3FFD93FA99AEE3EF39" bold="true" box="[586,964,1317,1344]" pageId="198" pageNumber="199">
<collectingCounty id="62C5A2D1FF63FF3FFD93FA99AEE3EF39" box="[586,964,1317,1344]" pageId="198" pageNumber="199">Prov. San Miguel de Pallaques</collectingCounty>
</emphasis>
• 
<location id="8EC48C86FF63FF3FFC05FA99A9A6EF39" LSID="urn:lsid:plazi:treatment:03B26B4BFF63FF30FDC7FDE1AC0DEB31:8EC48C86FF63FF3FFC05FA99A9A6EF39" box="[988,1153,1318,1344]" county="Prov. San Miguel de Pallaques" latitude="-7.016667" longLatPrecision="1303" longitude="-79.066666" name="Cerro Quillon" pageId="198" pageNumber="199">Cerro Quillón</location>
; [
<geoCoordinate id="EE2FBC9AFF63FF3FFB4DFA99A9CCEF39" box="[1172,1259,1316,1344]" degrees="7" direction="south" minutes="01" orientation="latitude" pageId="198" pageNumber="199" precision="925" value="-7.016667">7°01′ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFF63FF3FFB2EFA99A84DEF39" box="[1271,1386,1316,1344]" degrees="79" direction="west" minutes="04" orientation="longitude" pageId="198" pageNumber="199" precision="925" value="-79.066666">79°04′ W</geoCoordinate>
]; 
<elevation id="00363D6EFF63FF3FFF62FAF6AC73EF1A" box="[187,340,1353,1379]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="198" pageNumber="199" unit="m" value="3200.0">
<quantity id="4CE377B8FF63FF3FFF62FAF6AC33EF1A" box="[187,276,1353,1379]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="198" pageNumber="199" unit="m" value="3200.0">3200 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFF63FF3FFEB8FAF6ACD4EF1A" box="[353,499,1353,1380]" pageId="198" pageNumber="199" value="1986-03-05">
<collectingDate id="EFE10575FF63FF3FFEB8FAF6ACD4EF1A" box="[353,499,1353,1380]" pageId="198" pageNumber="199" value="1986-03-05">5 Mar. 1986</collectingDate>
</date>
; 
<emphasis id="B96F064FFF63FF3FFDD8FAF5A9F2EF1A" box="[513,1237,1353,1379]" italics="true" pageId="198" pageNumber="199">
<collectorName id="26EEBF8BFF63FF3FFDD8FAF5AF9BEF1A" box="[513,700,1354,1379]" pageId="198" pageNumber="199">J. Mostacero L.</collectorName>
, 
<collectorName id="26EEBF8BFF63FF3FFD10FAF6AE68EF1A" box="[713,847,1353,1379]" pageId="198" pageNumber="199">S. Leiva G.</collectorName>
, 
<collectorName id="26EEBF8BFF63FF3FFC85FAF5AEC6EF1A" box="[860,993,1353,1379]" pageId="198" pageNumber="199">F. Mejla C.</collectorName>
&amp; 
<collectorName id="26EEBF8BFF63FF3FFBDEFAF5A9B4EF1A" box="[1031,1171,1353,1379]" pageId="198" pageNumber="199">F. Paláez P.</collectorName>
1258
</emphasis>
; 
<collectionCode id="ED0A4298FF63FF3FFB3CFAF6A804EF1A" box="[1253,1315,1353,1379]" collectionName="HUT" country="Japan" httpUri="http://grbio.org/cool/gtk4-xv5b" name="HUT Culture Collection" pageId="198" pageNumber="199">HUT</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF63FF31FF64FA35A97CEB15" lastPageId="200" lastPageNumber="201" pageId="198" pageNumber="199" type="description">
<paragraph id="8BA4DA5DFF63FF3FFF64FA35AC68EFDD" blockId="198.[189,335,1418,1444]" box="[189,335,1418,1444]" pageId="198" pageNumber="199">
<heading id="D0EC6D31FF63FF3FFF64FA35AC68EFDD" bold="true" box="[189,335,1418,1444]" fontSize="11" level="3" pageId="198" pageNumber="199" reason="3">
<emphasis id="B96F064FFF63FF3FFF64FA35AC68EFDD" bold="true" box="[189,335,1418,1444]" pageId="198" pageNumber="199">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF63FF31FF64FA06A97CEB15" blockId="198.[188,1401,1465,2020]" lastBlockId="200.[188,1399,265,364]" lastPageId="200" lastPageNumber="201" pageId="198" pageNumber="199">
Caulescent, tuberous herb, to 
<quantity id="4CE377B8FF63FF3FFDFEFA06AF70EFAA" box="[551,599,1465,1491]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="198" pageNumber="199" unit="m" value="1.0">1 m</quantity>
high. 
<emphasis id="B96F064FFF63FF3FFD7EFA06AFCBEFAA" box="[679,748,1465,1491]" italics="true" pageId="198" pageNumber="199">Tuber</emphasis>
ellipsoid, to 2 × 
<quantity id="4CE377B8FF63FF3FFC1DFA06A926EFAA" box="[964,1025,1465,1491]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="198" pageNumber="199" unit="cm" value="4.0">4 cm</quantity>
but likely much larger, with 1 growing point. 
<emphasis id="B96F064FFF63FF3FFEACFA63AC89EF8F" box="[373,430,1500,1526]" italics="true" pageId="198" pageNumber="199">Stem</emphasis>
erect, unbranched; internodes to 
<quantity id="4CE377B8FF63FF3FFCE2FA63AEA2EF8E" box="[827,901,1500,1527]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="198" pageNumber="199" unit="cm" value="14.0">14 cm</quantity>
long, to 
<quantity id="4CE377B8FF63FF3FFC29FA63A963EF8E" box="[1008,1092,1500,1527]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="198" pageNumber="199" unit="mm" value="10.0">10 mm</quantity>
thick, succulent, green to red, glabrous. 
<emphasis id="B96F064FFF63FF3FFEBEFA40ACE2EC60" box="[359,453,1535,1561]" italics="true" pageId="198" pageNumber="199">Stipules</emphasis>
deciduous, lanceolate to ovate, 4–12 × 
<quantity id="4CE377B8FF63FF3FFC44FA40A925EC63" box="[925,1026,1535,1562]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="198" pageNumber="199" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 mm</quantity>
, apex acute, translucent, colour unknown, glabrous, margin entire, aciliate. 
<emphasis id="B96F064FFF63FF3FFD67F99CAE37EC45" box="[702,784,1571,1596]" italics="true" pageId="198" pageNumber="199">Leaves</emphasis>
&gt; 5, alternate, basifixed; petiole 
<quantity id="4CE377B8FF63FF3FFB4DF99CA9DCEC44" box="[1172,1275,1571,1597]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="13.0" metricValueMin="2.0" pageId="198" pageNumber="199" unit="cm" value="7.5" valueMax="13.0" valueMin="2.0">2–13 cm</quantity>
long, pale brown to green, glabrous; blade transversely ovate, to 24 × 
<quantity id="4CE377B8FF63FF3FFC55F9F9AEFEEC19" box="[908,985,1606,1632]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="198" pageNumber="199" unit="cm" value="11.0">11 cm</quantity>
, membranaceous, apex acuminate, base cordate, basal lobes not overlapping, sinus to 
<quantity id="4CE377B8FF63FF3FFCDEF9D6AE7FECFA" box="[775,856,1641,1667]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="198" pageNumber="199" unit="mm" value="40.0">40 mm</quantity>
deep, margin serrate, with 3–5 triangular lobes on the broad side of the leaf and often with one on the narrow side of the leaf, aciliate, upper surface green, glabrous, lower surface green, glabrous, veins palmate-pinnate, 6–8 veined from the base, with 3–5 secondary veins on the larger side, 2–4 secondary veins on the smaller side. 
<emphasis id="B96F064FFF63FF3FFB41F96DA867EC95" box="[1176,1344,1746,1772]" italics="true" pageId="198" pageNumber="199">Inflorescences</emphasis>
2–4 per stem, bisexual, axillary, erect, cymose, with up to 3 branches, bearing up to 8 staminate flowers and 4 pistillate flowers, protandrous; peduncle to 
<quantity id="4CE377B8FF63FF3FFCDAF8A6AE6BED4A" box="[771,844,1817,1843]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" pageId="198" pageNumber="199" unit="cm" value="24.0">24 cm</quantity>
long, pale brown, glabrous to sparsely minute-glandular pilose at the apex, bracts deciduous, ovate, 15–28 × 
<quantity id="4CE377B8FF63FF3FFC76F883A902ED2E" box="[943,1061,1852,1879]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="2.2" metricValueMin="0.9" pageId="198" pageNumber="199" unit="mm" value="15.5" valueMax="22.0" valueMin="9.0">9–22 mm</quantity>
, translucent, white, sparsely minute-glandular pilose, apex rounded to obtuse, margin entire, aciliate. 
<emphasis id="B96F064FFF63FF3FFBCDF8DFA9C0ED00" box="[1044,1255,1887,1914]" italics="true" pageId="198" pageNumber="199">Staminate flowers</emphasis>
: pedicels to 
<quantity id="4CE377B8FF63FF3FFF64F83DAC36EDE4" box="[189,273,1922,1949]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="198" pageNumber="199" unit="mm" value="45.0">45 mm</quantity>
long, sparsely minute-glandular pilose; tepals 4, spreading, outer 2 ovate, 22–45 × 
<quantity id="4CE377B8FF63FF3FFB29F83DA854EDE4" box="[1264,1395,1922,1949]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="3.0" metricValueMin="1.6" pageId="198" pageNumber="199" unit="mm" value="23.0" valueMax="30.0" valueMin="16.0">16–30 mm</quantity>
, apex short-acuminate, white, sparsely minute-glandular pilose, margin entire to serrulate, aciliate, inner 2 obovate, 16–35 × 
<quantity id="4CE377B8FF63FF3FFE21F876AF5BED9A" box="[504,636,1993,2019]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.6" metricValueMin="1.2" pageId="198" pageNumber="199" unit="mm" value="19.0" valueMax="26.0" valueMin="12.0">12–26 mm</quantity>
, apex rounded, white, sparsely minute-glandular pilose, margin entire, aciliate; stamens 45–50, spreading, yellow, filaments 
<quantity id="4CE377B8FF62FF3EFC5CFEB6AEDAEB5A" box="[901,1021,265,292]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="199" pageNumber="200" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.5–4 mm</quantity>
long, free, anthers ovoid, 0.6–1 × 
<quantity id="4CE377B8FF62FF3EFF0EFE93AC15EB3E" box="[215,306,300,327]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="199" pageNumber="200" unit="mm" value="0.5">0.5 mm</quantity>
long, dehiscing via lateral slits, connectives extending to 
<quantity id="4CE377B8FF62FF3EFC31FE93A960EB3E" box="[1000,1095,300,327]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="199" pageNumber="200" unit="mm" value="0.2">0.2 mm</quantity>
, symmetrically basifixed. 
<emphasis id="B96F064FFF62FF3EFF64FEF0ACA0EB10" box="[189,391,335,361]" italics="true" pageId="199" pageNumber="200">Pistillate flowers</emphasis>
: pedicels to 
<quantity id="4CE377B8FF62FF3EFDFAFEF0AF50EB13" box="[547,631,335,362]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="199" pageNumber="200" unit="mm" value="40.0">40 mm</quantity>
long; bracteoles 2, positioned directly beneath the ovary, ovate, 18–20 × 
<quantity id="4CE377B8FF62FF3EFEFEFECCAC8FEBF4" box="[295,424,371,397]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="199" pageNumber="200" unit="mm" value="15.0" valueMax="18.0" valueMin="12.0">12–18 mm</quantity>
, apex rounded, translucent, white, glabrous, margin entire to serrate, aciliate; tepals 5, subequal, persistence unknown, projecting, the largest broadly-lanceolate to ovate, 18–30 × 
<quantity id="4CE377B8FF62FF3EFAD8FE29A855EBC9" box="[1281,1394,406,432]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.2" metricValueMin="0.8" pageId="199" pageNumber="200" unit="mm" value="15.0" valueMax="22.0" valueMin="8.0">8–22 mm</quantity>
, apex acuminate, white, sparsely minute-glandular pilose, margin entire, aciliate, the smallest lanceolate to ovate, 12–14 × 
<quantity id="4CE377B8FF62FF3EFE4AFE63AF20EB8E" box="[403,519,476,503]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="199" pageNumber="200" unit="mm" value="9.0" valueMax="12.0" valueMin="6.0">6–12 mm</quantity>
, apex obtuse, white, glabrous, margin entire, aciliate; ovary body obovoid, 9–14 × 
<quantity id="4CE377B8FF62FF3EFEC4FE40ACB4E863" box="[285,403,511,538]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="199" pageNumber="200" unit="mm" value="8.0" valueMax="10.0" valueMin="6.0">6–10 mm</quantity>
, colour unknown, glabrous, unequally 3-winged, wings triangular, largest 12–17 × 
<quantity id="4CE377B8FF62FF3EFF64FD9DAC14E844" box="[189,307,546,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="199" pageNumber="200" unit="mm" value="10.5" valueMax="15.0" valueMin="6.0">6–15 mm</quantity>
, smallest 8–15 × 
<quantity id="4CE377B8FF62FF3EFDD5FD9CAFA5E844" box="[524,642,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="12.0" metricValueMin="3.0" pageId="199" pageNumber="200" unit="mm" value="7.5" valueMax="12.0" valueMin="3.0">3–12 mm</quantity>
; 3-locular, placentae branches divided, bearing ovules on both surfaces; styles 3, yellow, free, 
<quantity id="4CE377B8FF6DFF31FDFEFEB6AFA0EB5A" box="[551,647,265,291]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="200" pageNumber="201" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
long, once-divided, stigmatic papillae in a spirally-twisted band. 
<emphasis id="B96F064FFF6DFF31FF65FE91AC5BEB3E" box="[188,380,301,327]" italics="true" pageId="200" pageNumber="201">Fruiting pedicel</emphasis>
to 
<quantity id="4CE377B8FF6DFF31FE7AFE92ACD1EB31" box="[419,502,301,328]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="200" pageNumber="201" unit="mm" value="50.0">50 mm</quantity>
long. 
<emphasis id="B96F064FFF6DFF31FD99FE91AF99EB3E" box="[576,702,301,327]" italics="true" pageId="200" pageNumber="201">Fruit body</emphasis>
ovoid, to 17 × 
<quantity id="4CE377B8FF6DFF31FCAAFE92AEEEEB31" box="[883,969,301,328]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="200" pageNumber="201" unit="mm" value="13.0">13 mm</quantity>
, drying brown, wings same shape as in ovary, largest expanding to 17 × 
<quantity id="4CE377B8FF6DFF31FD85FEEDAF95EB15" box="[604,690,338,364]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="200" pageNumber="201" unit="mm" value="17.0">17 mm</quantity>
, smallest expanding to 15 × 
<quantity id="4CE377B8FF6DFF31FBD8FEEDA971EB15" box="[1025,1110,338,364]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="200" pageNumber="201" unit="mm" value="13.0">13 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF648AD5FF62FF3EFF64F837AE11ED91" ID-DOI="http://doi.org/10.5281/zenodo.8178438" ID-Zenodo-Dep="8178438" httpUri="https://zenodo.org/record/8178438/files/figure.png" pageId="199" pageNumber="200" startId="199.[189,232,1928,1954]" targetBox="[215,1391,623,1852]" targetPageId="199">
<paragraph id="8BA4DA5DFF62FF3EFF64F837AE11ED91" blockId="199.[189,1399,1928,2025]" pageId="199" pageNumber="200">
<emphasis id="B96F064FFF62FF3EFF64F837AC3CEDDB" bold="true" box="[189,283,1928,1954]" pageId="199" pageNumber="200">Fig. 64.</emphasis>
<taxonomicName id="4C1BA1DEFF62FF3EFEFBF836AED6EDDB" authority="L. B. Sm. &amp; B. G. Schub. A. Habit. B. Pistillate" box="[290,1009,1928,1954]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="199" pageNumber="200" phylum="Tracheophyta" rank="species" species="velata">
<emphasis id="B96F064FFF62FF3EFEFBF836ACF3EDDB" box="[290,468,1928,1954]" italics="true" pageId="199" pageNumber="200">Begonia velata</emphasis>
L.B.Sm. &amp; B.G.Schub. 
<emphasis id="B96F064FFF62FF3EFD2AF837AE2FEDDB" bold="true" box="[755,776,1928,1954]" pageId="199" pageNumber="200">A</emphasis>
. Habit. 
<emphasis id="B96F064FFF62FF3EFCBEF837AE5CEDDB" bold="true" box="[871,891,1928,1954]" pageId="199" pageNumber="200">B</emphasis>
. Pistillate
</taxonomicName>
flower and bracteoles, side view. 
<emphasis id="B96F064FFF62FF3EFF64F814ADF5EDBC" bold="true" box="[189,210,1963,1989]" pageId="199" pageNumber="200">C</emphasis>
. Stigma and styles drawn from a desiccated fruit, side view. Illustration by Anna Dorward from 
<emphasis id="B96F064FFF62FF3EFA84F813AFC9ED91" italics="true" pageId="199" pageNumber="200">E. Rodríguez R., E. Alvítez I. &amp; S. Orroyo A. 2720</emphasis>
(US).
</paragraph>
</caption>
<subSubSection id="C30189D6FF6DFF31FF64FE2BAC57E8EA" pageId="200" pageNumber="201" type="conservation">
<paragraph id="8BA4DA5DFF6DFF31FF64FE2BAF4AEBD7" blockId="200.[189,621,404,430]" box="[189,621,404,430]" pageId="200" pageNumber="201">
<heading id="D0EC6D31FF6DFF31FF64FE2BAF4AEBD7" bold="true" box="[189,621,404,430]" fontSize="11" level="3" pageId="200" pageNumber="201" reason="3">
<emphasis id="B96F064FFF6DFF31FF64FE2BAF4AEBD7" bold="true" box="[189,621,404,430]" pageId="200" pageNumber="201">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF6DFF31FF64FE7BAC57E8EA" blockId="200.[188,1399,452,659]" pageId="200" pageNumber="201">
Assessed by 
<bibRefCitation id="EF8AA7ACFF6DFF31FE85FE7BAFB4EBA7" author="Leon &amp; Monsalve" box="[348,659,452,478]" firstAuthor="Leon" pageId="200" pageNumber="201" pagination="165 - 170" refId="ref229226" refString="Leon B. &amp; Monsalve C. 2006. Begoniaceae endemicas del Peru. Revista Peruana de Biologia 13 (2): 165 - 170. https: // doi. org / 10.15381 / rpb. v 13 i 2.1810" type="journal article" year="2006">
<collectingRegion id="49DF14BFFF6DFF31FE85FE7BACBFEBA7" box="[348,408,452,478]" country="Nicaragua" name="Leon" pageId="200" pageNumber="201">León</collectingRegion>
&amp; Monsalve (2006)
</bibRefCitation>
as Data Deficient (DD) and reassessed by 
<bibRefCitation id="EF8AA7ACFF6DFF31FB7EFE7BA874EBA7" author="Tebbitt" box="[1191,1363,452,478]" firstAuthor="Tebbitt" pageId="200" pageNumber="201" pagination="143 - 152" refId="ref232485" refString="Tebbitt M. C. 2016. Two new species of Andean Begonia (Begoniaceae). Edinburgh Journal of Botany 73 (1): 143 - 152. https: // doi. org / 10.1017 / S 0960428615000335" type="journal article" year="2016">Tebbitt (2016)</bibRefCitation>
as Vulnerable (VU D2). Known from five fragments of highly threatened northwest Peruvian relict montane forest on the western slopes of the Andes. The most distant sites are approximately 
<quantity id="4CE377B8FF6DFF31FAC2FDB3A851E85F" box="[1307,1398,524,550]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="200" pageNumber="201" unit="km" value="200.0">200 km</quantity>
apart and none of the five sites are protected. Given the species low EOO (ca 
<quantity id="4CE377B8FF6DFF31FB93FD8FA988E832" box="[1098,1199,560,587]" metricMagnitude="6" metricUnit="m" metricValue="2.5" pageId="200" pageNumber="201" unit="km" value="2500.0">2500 km</quantity>
<superScript id="7C6E7715FF6DFF31FB69FD8FA99EE847" attach="right" box="[1200,1209,560,574]" fontSize="6" pageId="200" pageNumber="201">2</superScript>
), its few known localities and fragmented distribution, and the threats to its habitat, we reassess 
<taxonomicName id="4C1BA1DEFF6DFF31FB82FDE9A9E4E816" box="[1115,1219,597,623]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="velata">
<emphasis id="B96F064FFF6DFF31FB82FDE9A9E4E816" box="[1115,1219,597,623]" italics="true" pageId="200" pageNumber="201">B. velata</emphasis>
</taxonomicName>
as Endangered (EN B1ab(iii)).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF6DFF31FF64FD03ACB5EFA2" pageId="200" pageNumber="201" type="discussion">
<paragraph id="8BA4DA5DFF6DFF31FF64FD03AC23E8AF" blockId="200.[189,260,700,726]" box="[189,260,700,726]" pageId="200" pageNumber="201">
<heading id="D0EC6D31FF6DFF31FF64FD03AC23E8AF" bold="true" box="[189,260,700,726]" fontSize="11" level="3" pageId="200" pageNumber="201" reason="3">
<emphasis id="B96F064FFF6DFF31FF64FD03AC23E8AF" bold="true" box="[189,260,700,726]" pageId="200" pageNumber="201">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF6DFF31FF64FD54AF57EE51" blockId="200.[188,1399,747,1064]" pageId="200" pageNumber="201">
<bibRefCitation id="EF8AA7ACFF6DFF31FF64FD54AC4EE97F" author="Tebbitt M. C." box="[189,361,747,774]" journalOrPublisher="Edinburgh Journal of Botany" pageId="200" pageNumber="201" pagination="217 - 228" part="74" refId="ref232526" refString="Tebbitt M. C. 2017. Recircumscription and new synonyms of Begonia acerifolia (Begoniaceae) and amended descriptions of the poorly known Begonia hydrophylloides and Begonia velata. Edinburgh Journal of Botany 74 (2): 217 - 228. https: // doi. org / 10.1017 / S 0960428617000105" title="Recircumscription and new synonyms of Begonia acerifolia (Begoniaceae) and amended descriptions of the poorly known Begonia hydrophylloides and Begonia velata" type="journal article" year="2017">Tebbitt (2017)</bibRefCitation>
speculated on a possible hybrid origin of 
<taxonomicName id="4C1BA1DEFF6DFF31FCABFD53AEF9E97C" box="[882,990,747,773]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="velata">
<emphasis id="B96F064FFF6DFF31FCABFD53AEF9E97C" box="[882,990,747,773]" italics="true" pageId="200" pageNumber="201">B. velata</emphasis>
</taxonomicName>
, highlighting the possible role of 
<taxonomicName id="4C1BA1DEFF6DFF31FF64FCAFAC77E953" box="[189,336,784,810]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="acerifolia">
<emphasis id="B96F064FFF6DFF31FF64FCAFAC77E953" box="[189,336,784,810]" italics="true" pageId="200" pageNumber="201">B. acerifolia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1BA1DEFF6DFF31FE5FFCAFAF2FE953" box="[390,520,784,810]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="ludwigii">
<emphasis id="B96F064FFF6DFF31FE5FFCAFAF2FE953" box="[390,520,784,810]" italics="true" pageId="200" pageNumber="201">B. ludwigii</emphasis>
</taxonomicName>
as parents. Tebbitt’s concept of 
<taxonomicName id="4C1BA1DEFF6DFF31FCA0FCAFAEC5E953" box="[889,994,784,810]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="velata">
<emphasis id="B96F064FFF6DFF31FCA0FCAFAEC5E953" box="[889,994,784,810]" italics="true" pageId="200" pageNumber="201">B. velata</emphasis>
</taxonomicName>
was a misapplied name (see Notes under 
<taxonomicName id="4C1BA1DEFF6DFF31FEDFFC8AAF7BE937" authority="Moonlight" authorityName="Moonlight &amp; Jara-Muñoz &amp; Purvis &amp; Delves &amp; Allen &amp; Reynel" authorityYear="2023" box="[262,604,820,846]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="huancabambae" status="sp. nov.">
<emphasis id="B96F064FFF6DFF31FEDFFC8AACFDE937" box="[262,474,820,846]" italics="true" pageId="200" pageNumber="201">B. huancabambae</emphasis>
Moonlight
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FF6DFF31FDBBFC8AAF9AE937" box="[610,701,820,847]" pageId="200" pageNumber="201" rank="species">sp. nov.</taxonomicNameLabel>
) and 
<taxonomicName id="4C1BA1DEFF6DFF31FD23FC8AAE45E937" box="[762,866,820,846]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="velata">
<emphasis id="B96F064FFF6DFF31FD23FC8AAE45E937" box="[762,866,820,846]" italics="true" pageId="200" pageNumber="201">B. velata</emphasis>
</taxonomicName>
is clearly most closely related to 
<taxonomicName id="4C1BA1DEFF6DFF31FB3BFC8AA849E937" box="[1250,1390,821,846]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="piurensis">
<emphasis id="B96F064FFF6DFF31FB3BFC8AA849E937" box="[1250,1390,821,846]" italics="true" pageId="200" pageNumber="201">B. piurensis</emphasis>
</taxonomicName>
. These two species are the only tuberous species of Andean 
<taxonomicName id="4C1BA1DEFF6DFF31FCAAFCE6AEF1E90B" box="[883,982,857,882]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F064FFF6DFF31FCAAFCE6AEF1E90B" box="[883,982,857,882]" italics="true" pageId="200" pageNumber="201">Begonia</emphasis>
</taxonomicName>
with&gt; 
<quantity id="4CE377B8FF6DFF31FBE9FCE7A95EE90A" box="[1072,1145,856,883]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="200" pageNumber="201" unit="cm" value="50.0">50 cm</quantity>
tall stems. These two species are both found in the Amotape-Huancabamba Region of northern 
<collectingCountry id="F30C9ACDFF6DFF31FBF6FCC2A942E9EE" box="[1071,1125,893,919]" name="Peru" pageId="200" pageNumber="201">Peru</collectingCountry>
and southern 
<collectingCountry id="F30C9ACDFF6DFF31FAD6FCC2A854E9EE" box="[1295,1395,893,919]" name="Ecuador" pageId="200" pageNumber="201">Ecuador</collectingCountry>
. The new species is found further south than 
<taxonomicName id="4C1BA1DEFF6DFF31FD0AFC1DAE78E9C2" box="[723,863,930,955]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="piurensis">
<emphasis id="B96F064FFF6DFF31FD0AFC1DAE78E9C2" box="[723,863,930,955]" italics="true" pageId="200" pageNumber="201">B. piurensis</emphasis>
</taxonomicName>
, which is only found within 
<collectingRegion id="49DF14BFFF6DFF31FB62FC1EA870E9C2" box="[1211,1367,929,955]" country="Peru" name="Piura" pageId="200" pageNumber="201">Piura Region</collectingRegion>
in 
<collectingCountry id="F30C9ACDFF6DFF31FF66FC79ADDFE999" box="[191,248,966,992]" name="Peru" pageId="200" pageNumber="201">Peru</collectingCountry>
, and the two species appear to be ecologically distinct. 
<taxonomicName id="4C1BA1DEFF6DFF31FC5DFC79A97CE9A6" box="[900,1115,966,991]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="piurensis">
<emphasis id="B96F064FFF6DFF31FC5DFC79A97CE9A6" box="[900,1115,966,991]" italics="true" pageId="200" pageNumber="201">Begonia piurensis</emphasis>
</taxonomicName>
is found to an elevation of 
<quantity id="4CE377B8FF6DFF31FF06FC55AC1EEE7D" box="[223,313,1002,1028]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="200" pageNumber="201" unit="m" value="2000.0">2000 m</quantity>
a.s.l. whereas 
<taxonomicName id="4C1BA1DEFF6DFF31FE35FC55AF70EE7A" box="[492,599,1001,1027]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="velata">
<emphasis id="B96F064FFF6DFF31FE35FC55AF70EE7A" box="[492,599,1001,1027]" italics="true" pageId="200" pageNumber="201">B. velata</emphasis>
</taxonomicName>
has only been collected above 
<quantity id="4CE377B8FF6DFF31FC09FC56A90EEE7D" box="[976,1065,1001,1028]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="200" pageNumber="201" unit="m" value="2600.0">2600 m</quantity>
a.s.l. in cooler forest where most of the precipitation is from fog.
</paragraph>
<paragraph id="8BA4DA5DFF6DFF31FF64FBEFAC97EE13" blockId="200.[189,432,1104,1130]" box="[189,432,1104,1130]" pageId="200" pageNumber="201">
<heading id="D0EC6D31FF6DFF31FF64FBEFAC97EE13" bold="true" box="[189,432,1104,1130]" fontSize="11" level="3" pageId="200" pageNumber="201" reason="3">
<emphasis id="B96F064FFF6DFF31FF64FBEFAC97EE13" bold="true" box="[189,432,1104,1130]" pageId="200" pageNumber="201">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF6DFF31FF64FB3EA92CEF55" blockId="200.[189,1399,1152,1324]" pageId="200" pageNumber="201">
<taxonomicName id="4C1BA1DEFF6DFF31FF64FB3EAC57EEE3" box="[189,368,1152,1178]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="velata">
<emphasis id="B96F064FFF6DFF31FF64FB3EAC57EEE3" box="[189,368,1152,1178]" italics="true" pageId="200" pageNumber="201">Begonia velata</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4C1BA1DEFF6DFF31FD8BFB3EAFF8EEE3" box="[594,735,1153,1178]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="piurensis">
<emphasis id="B96F064FFF6DFF31FD8BFB3EAFF8EEE3" box="[594,735,1153,1178]" italics="true" pageId="200" pageNumber="201">B. piurensis</emphasis>
</taxonomicName>
, but is easy to distinguish when in flower or fruit by its larger bracts (15–28 × 
<quantity id="4CE377B8FF6DFF31FE12FB1BAF1BEEC6" box="[459,572,1188,1215]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="2.2" metricValueMin="0.9" pageId="200" pageNumber="201" unit="mm" value="15.5" valueMax="22.0" valueMin="9.0">9–22 mm</quantity>
vs 4–6 × 
<quantity id="4CE377B8FF6DFF31FD6BFB1BAE08EEC7" box="[690,815,1188,1215]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="200" pageNumber="201" unit="mm" value="2.75" valueMax="4.0" valueMin="1.5">1.5–4 mm</quantity>
); its larger staminate flowers (
<quantity id="4CE377B8FF6DFF31FB4BFB1BA833EEC6" box="[1170,1300,1188,1215]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" metricValueMax="9.0" metricValueMin="4.4" pageId="200" pageNumber="201" unit="mm" value="67.0" valueMax="90.0" valueMin="44.0">44–90 mm</quantity>
wide vs 
<quantity id="4CE377B8FF6DFF31FF64FB77AC1BEE9A" box="[189,316,1224,1251]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="200" pageNumber="201" unit="mm" value="14.0" valueMax="16.0" valueMin="12.0">12–16 mm</quantity>
wide); its bracteolate pistillate flowers (vs ebracteolate); and its larger fruits bodies (to 17 × 
<quantity id="4CE377B8FF6DFF31FF64FB52AC36EF7E" box="[189,273,1261,1287]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="200" pageNumber="201" unit="mm" value="12.0">12 mm</quantity>
vs 9 × 
<quantity id="4CE377B8FF6DFF31FEB2FB52AC92EF7E" box="[363,437,1261,1287]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="200" pageNumber="201" unit="mm" value="8.0">8 mm</quantity>
). When sterile, 
<taxonomicName id="4C1BA1DEFF6DFF31FDA9FB51AFFEEF7E" box="[624,729,1261,1287]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="velata">
<emphasis id="B96F064FFF6DFF31FDA9FB51AFFEEF7E" box="[624,729,1261,1287]" italics="true" pageId="200" pageNumber="201">B. velata</emphasis>
</taxonomicName>
may be distinguished from 
<taxonomicName id="4C1BA1DEFF6DFF31FBF1FB51A992EF7E" box="[1064,1205,1262,1287]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="piurensis">
<emphasis id="B96F064FFF6DFF31FBF1FB51A992EF7E" box="[1064,1205,1262,1287]" italics="true" pageId="200" pageNumber="201">B. piurensis</emphasis>
</taxonomicName>
by the cusps on the longest edge of its leaf, which give the blade an angular appearance.
</paragraph>
<paragraph id="8BA4DA5DFF6DFF31FF64FAEBACB5EFA2" blockId="200.[188,1400,1364,1499]" pageId="200" pageNumber="201">
Many specimens of 
<taxonomicName id="4C1BA1DEFF6DFF31FE69FAEBAF3CEF17" box="[432,539,1364,1390]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="velata">
<emphasis id="B96F064FFF6DFF31FE69FAEBAF3CEF17" box="[432,539,1364,1390]" italics="true" pageId="200" pageNumber="201">B. velata</emphasis>
</taxonomicName>
have also been confused with the superficially similar 
<taxonomicName id="4C1BA1DEFF6DFF31FB74FAEBA865EF17" box="[1197,1346,1364,1390]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="acerifolia">
<emphasis id="B96F064FFF6DFF31FB74FAEBA865EF17" box="[1197,1346,1364,1390]" italics="true" pageId="200" pageNumber="201">B. acerifolia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1BA1DEFF6DFF31FF64FAC6ACB3EFEB" authority="Moonlight &amp; Jara-Muñoz &amp; Purvis &amp; Delves &amp; Allen &amp; Reynel, 2023" authorityName="Moonlight &amp; Jara-Muñoz &amp; Purvis &amp; Delves &amp; Allen &amp; Reynel" authorityYear="2023" box="[189,404,1400,1426]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="huancabambae" status="sp. nov.">
<emphasis id="B96F064FFF6DFF31FF64FAC6ACB3EFEB" box="[189,404,1400,1426]" italics="true" pageId="200" pageNumber="201">B. huancabambae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FF6DFF31FE47FAC6ACDAEFEB" box="[414,509,1400,1427]" pageId="200" pageNumber="201" rank="species">sp. nov.</taxonomicNameLabel>
), which are both rhizomatous rather than tuberous herbs but have similarly shaped leaves. Within the range of 
<taxonomicName id="4C1BA1DEFF6DFF31FD89FA22AF9FEFCF" box="[592,696,1436,1462]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="velata">
<emphasis id="B96F064FFF6DFF31FD89FA22AF9FEFCF" box="[592,696,1436,1462]" italics="true" pageId="200" pageNumber="201">B. velata</emphasis>
</taxonomicName>
, both species have a conspicuous indumentum on both sides of the leaf lamina.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF6DFF30FF64F9BCAC0DEB31" lastPageId="201" lastPageNumber="202" pageId="200" pageNumber="201" type="distribution">
<paragraph id="8BA4DA5DFF6DFF31FF64F9BCACD2EC64" blockId="200.[189,501,1539,1565]" box="[189,501,1539,1565]" pageId="200" pageNumber="201">
<heading id="D0EC6D31FF6DFF31FF64F9BCACD2EC64" bold="true" box="[189,501,1539,1565]" fontSize="11" level="3" pageId="200" pageNumber="201" reason="3">
<emphasis id="B96F064FFF6DFF31FF64F9BCACD2EC64" bold="true" box="[189,501,1539,1565]" pageId="200" pageNumber="201">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF6DFF31FF64F98CA922ECEC" blockId="200.[189,1399,1587,1686]" pageId="200" pageNumber="201">
Endemic to 
<collectingCountry id="F30C9ACDFF6DFF31FE93F98CACA7EC34" box="[330,384,1587,1613]" name="Peru" pageId="200" pageNumber="201">Peru</collectingCountry>
and known from 
<collectingRegion id="49DF14BFFF6DFF31FD96F98CAFC0EC34" box="[591,743,1587,1613]" country="Peru" name="Lambayeque" pageId="200" pageNumber="201">Lambayeque</collectingRegion>
and 
<collectingRegion id="49DF14BFFF6DFF31FCC6F98CAEBBEC34" box="[799,924,1587,1613]" country="Peru" name="Cajamarca" pageId="200" pageNumber="201">Cajamarca</collectingRegion>
Regions (
<figureCitation id="1320C6D8FF6DFF31FBCDF98CA958EC34" box="[1044,1151,1587,1613]" captionStart="Fig" captionStartId="193.[189,232,1892,1918]" captionTargetBox="[194,1392,1227,1846]" captionTargetId="figure-268@193.[189,1398,1217,1852]" captionTargetPageId="193" captionText="Fig. 62. Distribution of Begonia sect. Knesebeckia in Peru. A. B. brandbygeana L.B.Sm. &amp; Wassh. (red) and B. parcifolia (blue). B. B. piurensis L.B.Sm. &amp; B.G.Schub. (red) and B. velata L.B.Sm. &amp; B.G.Schub. (blue). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178434" httpUri="https://zenodo.org/record/8178434/files/figure.png" pageId="200" pageNumber="201">Fig. 62B</figureCitation>
), where it is found in northwest Peruvian montane forests at an elevation of 
<quantity id="4CE377B8FF6DFF31FCE7F9E8AEC6EC0B" box="[830,993,1623,1650]" metricMagnitude="3" metricUnit="m" metricValue="2.9" metricValueMax="3.3" metricValueMin="2.5" pageId="200" pageNumber="201" unit="m" value="2900.0" valueMax="3300.0" valueMin="2500.0">2500–3300 m</quantity>
a.s.l. 
<taxonomicName id="4C1BA1DEFF6DFF31FBFEF9E7A9FEEC08" box="[1063,1241,1623,1649]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="velata">
<emphasis id="B96F064FFF6DFF31FBFEF9E7A9FEEC08" box="[1063,1241,1623,1649]" italics="true" pageId="200" pageNumber="201">Begonia velata</emphasis>
</taxonomicName>
is a tuberous species and dies back to its tuber in the dry season (June to November).
</paragraph>
<paragraph id="8BA4DA5DFF6DFF31FF64F900AE7AECA5" blockId="200.[189,861,1727,1756]" box="[189,861,1727,1756]" pageId="200" pageNumber="201">
<emphasis id="B96F064FFF6DFF31FF64F900AE7AECA5" bold="true" box="[189,861,1727,1756]" italics="true" pageId="200" pageNumber="201">
The 
<taxonomicName id="4C1BA1DEFF6DFF31FF20F900AC5EECA5" box="[249,377,1727,1756]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="acerifolia">acerifolia</taxonomicName>
group of 
<taxonomicName id="4C1BA1DEFF6DFF31FE22F900AE7AECA5" box="[507,861,1727,1756]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="section" section="Knesebeckia">Begonia sect. Knesebeckia</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA4DA5DFF6DFF31FF64F94BAE9BED90" blockId="200.[187,1399,1780,2025]" pageId="200" pageNumber="201">
The acerifolia group of 
<taxonomicName id="4C1BA1DEFF6DFF31FE08F94AAE2AED77" box="[465,781,1780,1807]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="section" section="Knesebeckia">
<emphasis id="B96F064FFF6DFF31FE08F94AAF13ED77" box="[465,564,1781,1806]" italics="true" pageId="200" pageNumber="201">Begonia</emphasis>
sect. 
<emphasis id="B96F064FFF6DFF31FDAEF94BAE2AED77" box="[631,781,1780,1806]" italics="true" pageId="200" pageNumber="201">Knesebeckia</emphasis>
</taxonomicName>
was defined by 
<bibRefCitation id="EF8AA7ACFF6DFF31FC10F94BA957ED77" author="Tebbitt" box="[969,1136,1780,1806]" firstAuthor="Tebbitt" pageId="200" pageNumber="201" pagination="143 - 152" refId="ref232485" refString="Tebbitt M. C. 2016. Two new species of Andean Begonia (Begoniaceae). Edinburgh Journal of Botany 73 (1): 143 - 152. https: // doi. org / 10.1017 / S 0960428615000335" type="journal article" year="2016">Tebbitt (2016)</bibRefCitation>
to include ten Andean species of 
<taxonomicName id="4C1BA1DEFF6DFF31FEE4F8A5ACB9ED4A" box="[317,414,1818,1843]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F064FFF6DFF31FEE4F8A5ACB9ED4A" box="[317,414,1818,1843]" italics="true" pageId="200" pageNumber="201">Begonia</emphasis>
</taxonomicName>
, though two of the original members are now considered synonyms (
<bibRefCitation id="EF8AA7ACFF6DFF31FB05F8A6ADDAED2E" author="Tebbitt" etAl="et al." firstAuthor="Tebbitt" pageId="200" pageNumber="201" pagination="199 - 215" refId="ref232682" refString="Tebbitt M. C., Hughes M. &amp; Moonlight P. W. 2017. Taxonomy of the Begonia tiliifolia group, including descriptions of two new species. Edinburgh Journal of Botany 74 (2): 199 - 215. https: // doi. org / 10.1017 / S 0960428617000087" type="journal article" year="2017">
Tebbitt 
<emphasis id="B96F064FFF6DFF31FAE2F8A5A851ED4A" box="[1339,1398,1817,1843]" italics="true" pageId="200" pageNumber="201">et al.</emphasis>
2017
</bibRefCitation>
). It includes relatively seasonally dry adapted members of the section that die back to above-ground stems in the dry season. Recent phylogenetic evidence demonstrates that this group is polyphyletic (
<bibRefCitation id="EF8AA7ACFF6DFF31FF1CF839ACFFEDD9" author="Moonlight P. W. &amp; Reynel C." box="[197,472,1926,1952]" journalOrPublisher="Phytotaxa" pageId="200" pageNumber="201" pagination="116 - 126" part="381" refId="ref229806" refString="Moonlight P. W. &amp; Reynel C. 2018. Two new species of Begonia from Peru. Phytotaxa 381 (1): 116 - 126. https: // doi. org / 10.11646 / phytotaxa. 381.1.15" title="Two new species of Begonia from Peru" type="journal article" year="2018">
Moonlight 
<emphasis id="B96F064FFF6DFF31FE96F838ACA8EDD9" box="[335,399,1926,1952]" italics="true" pageId="200" pageNumber="201">et al.</emphasis>
2018
</bibRefCitation>
) but we retain it here as it is a useful morphological grouping. Following 
<bibRefCitation id="EF8AA7ACFF6DFF31FF64F815ACF6EDBD" author="Moonlight P. W. &amp; Reynel C." box="[189,465,1962,1988]" journalOrPublisher="Phytotaxa" pageId="200" pageNumber="201" pagination="116 - 126" part="381" refId="ref229806" refString="Moonlight P. W. &amp; Reynel C. 2018. Two new species of Begonia from Peru. Phytotaxa 381 (1): 116 - 126. https: // doi. org / 10.11646 / phytotaxa. 381.1.15" title="Two new species of Begonia from Peru" type="journal article" year="2018">
Moonlight 
<emphasis id="B96F064FFF6DFF31FE99F814AC5CEDBD" box="[320,379,1962,1988]" italics="true" pageId="200" pageNumber="201">et al.</emphasis>
(2018)
</bibRefCitation>
we exclude 
<taxonomicName id="4C1BA1DEFF6DFF31FDBDF814AE0CEDBD" box="[612,811,1962,1988]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="urubambensis">
<emphasis id="B96F064FFF6DFF31FDBDF814AE0CEDBD" box="[612,811,1962,1988]" italics="true" pageId="200" pageNumber="201">B. urubambensis</emphasis>
</taxonomicName>
from the group, which differs in lacking an above ground-stem and is distantly related to other species in the group.
</paragraph>
<paragraph id="8BA4DA5DFF6CFF30FF64FEB6AC0DEB31" blockId="201.[189,1399,265,328]" pageId="201" pageNumber="202">
Of the seven currently recognised members of the acerifolia group, six are found in 
<collectingCountry id="F30C9ACDFF6CFF30FB75FEB6A9C5EB5A" box="[1196,1250,265,291]" name="Peru" pageId="201" pageNumber="202">Peru</collectingCountry>
and two are endemic.
</paragraph>
</subSubSection>
</treatment>
</document>