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<document id="824B3EB47FDFF46C2883F8F7B3539AE5" ID-DOI="https://doi.org/10.5852/ejt.2023.881.2175" ID-ISSN="2118-9773" ID-Zenodo-Dep="8178280" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690201937279" checkinUser="felipe" docAuthor="Moonlight, Peter. W., Jara-Muñoz, Orlando A., Purvis, David A., Delves, Jay, Allen, Josh P. &amp; Reynel, Carlos" docDate="2023" docId="03B26B4BFF61FF3FFDD3F90FA982E859" docLanguage="en" docName="ejt-881_1-334.imf" docOrigin="European Journal of Taxonomy 881" docSource="http://dx.doi.org/10.5852/ejt.2023.881.2175" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia piurensis L. B. Sm. &amp; B. G. Schub." docType="treatment" docVersion="8" lastPageNumber="199" masterDocId="FF8B1333FFA5FFF9FFD9FFBFAD27EA79" masterDocTitle="The genus Begonia (Begoniaceae) in Peru" masterLastPageNumber="334" masterPageNumber="1" pageNumber="197" updateTime="1707123162844" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="4B5A5E851BB101562527B1D668FA5FCF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FF4A1E24B35E73C6A1A8F70F30D3B5F8">
<mods:title id="997D86DB3BDDC5C9547137E717B277F6">The genus Begonia (Begoniaceae) in Peru</mods:title>
</mods:titleInfo>
<mods:name id="F302DA48E3421FC11A6B029C0CE0DE7F" type="personal">
<mods:role id="23070E739DA12241B8ACBC8C8C46B2E7">
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<mods:namePart id="157F304E1EE62DFAED60340AB7D56C8F">Moonlight, Peter. W.</mods:namePart>
<mods:affiliation id="A6628EEC22C046C03A5AC86A124EF784">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK. &amp; Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland.</mods:affiliation>
<mods:nameIdentifier id="EC5FEC5F399D37BC8D1AA24C137D97CA" type="email">moonligp@tcd.ie</mods:nameIdentifier>
</mods:name>
<mods:name id="52C4075E224818E7918D195C4528745A" type="personal">
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<mods:roleTerm id="02D6E6D8D2ECDF991B2563DA7DDF8751">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B897EB7DF26574CE7A70151C0F71AC02">Jara-Muñoz, Orlando A.</mods:namePart>
<mods:affiliation id="72F1180A63C78281572915FD471EFA24">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá D. C., Colombia.</mods:affiliation>
<mods:nameIdentifier id="4034E39ED513AEE6C1806DDB859605B8" type="email">oajaram@unal.edu.co</mods:nameIdentifier>
</mods:name>
<mods:name id="11034E90D64CA6944CFBFD36D0DF0153" type="personal">
<mods:role id="1EDD338EDDFBA8D96909F9BF51D64353">
<mods:roleTerm id="ACBF6CF927D10EA30400BDE80D90FE3D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="07A754C9E4761173A9922F0361DC0F4B">Purvis, David A.</mods:namePart>
<mods:affiliation id="D518C8CE33FD8D07B9D19427B2674F9E">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="ADE9A80E6E8A53D20A4CF77F2242459B" type="email">dpurvis@rbge.org.uk</mods:nameIdentifier>
</mods:name>
<mods:name id="BBF2B186511BF35B3EE712B72BD46D56" type="personal">
<mods:role id="9171C7AFBE9073126C64569D6772901C">
<mods:roleTerm id="B925DF6CE753AD5222D6D53BD71CCC88">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8E894C56D4415F81D25FB4FA9C3401A7">Delves, Jay</mods:namePart>
<mods:affiliation id="C5B7AF8383C2F39999045A9112124910">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="AAB6893C021C5278EC6FA6CBE6D90D14" type="email">j.delves@westernsydney.edu.au</mods:nameIdentifier>
</mods:name>
<mods:name id="8867514E287BC3D53EE67C71F1F61145" type="personal">
<mods:role id="D73D04D1F83DFDB5B2D941E63611750D">
<mods:roleTerm id="CC714BA0DA6C0AAD2CC43931B27E5116">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A271B2E0658E9B5D9754588D665D11B0">Allen, Josh P.</mods:namePart>
<mods:affiliation id="7B9FFD4BAE1FACEFF9FCC4C52DFB197B">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="10BF351D90642B8116742DE1F12C8909" type="email">myrmeciaman@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="7493FED5D46E5CC39DCD15F3EFA44135" type="personal">
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<mods:roleTerm id="D6628A17680767F655A264B7B1083BE4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6B9CBB38CC4D59749DBECC37B2C0C457">Reynel, Carlos</mods:namePart>
<mods:affiliation id="9A694CCFFCEAEC59F9EA4EDE53A22477">Facultad de Ciencias Forestales, Universidad Nacional Agraria-La Molina, Lima 12, Peru.</mods:affiliation>
<mods:nameIdentifier id="B1C82BA497EF410CA633B3F42FE5A1E1" type="email">reynel@lamolina.edu.pe</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="956A4B2E32B81F306A6697D5A78E77E3">text</mods:typeOfResource>
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<mods:titleInfo id="38EFB755CAF394BBFB06B8AB906C3D3D">
<mods:title id="5B279F215AE102F3D0030A96F8FFD093">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="ADEAE2C0041BB4CD0A47B5475958981E">
<mods:date id="A71712B58A6FADE059EC115186F43D66">2023</mods:date>
<mods:detail id="21A2AEB63AB0C97D2A15EBD24912025B" type="pubDate">
<mods:number id="55A4CDC103877D084DBEF85F366AA4D7">2023-07-18</mods:number>
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<mods:detail id="5DDE2096111B54E4842DD357C748AA19" type="volume">
<mods:number id="C002EE703E2BE6F60D57093436BF42C0">881</mods:number>
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<mods:start id="D173E16E68BBB60C4FC7840642FDBD69">1</mods:start>
<mods:end id="0D5F9E006C4B8C6A0858F7FD521D4AD2">334</mods:end>
</mods:extent>
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<mods:location id="AD06B600817598E1A1E31848F85B954E">
<mods:url id="4DF83B612F047FB739D44C002632F4B7">http://dx.doi.org/10.5852/ejt.2023.881.2175</mods:url>
</mods:location>
<mods:classification id="6E09EF91125EAF2169D3BDE3116546B4">journal article</mods:classification>
<mods:identifier id="40F396BD9D84652B2DD25668EA364858" type="DOI">https://doi.org/10.5852/ejt.2023.881.2175</mods:identifier>
<mods:identifier id="3C5019BAF088600B36B2841BF6C33A0C" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="497CA3A5C5766309C0346806579721B9" type="Zenodo-Dep">8178280</mods:identifier>
</mods:mods>
<treatment id="03B26B4BFF61FF3FFDD3F90FA982E859" LSID="urn:lsid:plazi:treatment:03B26B4BFF61FF3FFDD3F90FA982E859" httpUri="http://treatment.plazi.org/id/03B26B4BFF61FF3FFDD3F90FA982E859" lastPageId="198" lastPageNumber="199" pageId="196" pageNumber="197">
<subSubSection id="C30189D6FF61FF3DFDD3F90FA90EECB3" box="[522,1065,1712,1738]" pageId="196" pageNumber="197" type="nomenclature">
<paragraph id="8BA4DA5DFF61FF3DFDD3F90FA90EECB3" blockId="196.[522,1065,1712,1738]" box="[522,1065,1712,1738]" pageId="196" pageNumber="197">
47.
<taxonomicName id="4C1BA1DEFF61FF3DFDEFF90FA90EECB3" ID-CoL="7SGZ7" authority="L. B. Sm. &amp; B. G. Schub." box="[566,1065,1712,1738]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="196" pageNumber="197" phylum="Tracheophyta" rank="species" species="piurensis">
<emphasis id="B96F064FFF61FF3DFDEFF90FAE35ECB3" bold="true" box="[566,786,1712,1738]" italics="true" pageId="196" pageNumber="197">Begonia piurensis</emphasis>
L.B.Sm. &amp; B.G.Schub.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF61FF3DFF64F94CAC65ED48" pageId="196" pageNumber="197" type="reference_group">
<paragraph id="8BA4DA5DFF61FF3DFF64F94CAC65ED48" blockId="196.[189,1399,1779,2024]" pageId="196" pageNumber="197">
<emphasis id="B96F064FFF61FF3DFF64F94CAEDFED74" box="[189,1016,1779,1805]" italics="true" pageId="196" pageNumber="197">Publications of the Field Museum of Natural History, Botanical Series</emphasis>
13 (4/1): 197 (
<bibRefCitation id="EF8AA7ACFF61FF3DFB7AF94CAC12ED48" author="Smith &amp; Schubert" firstAuthor="Smith" pageId="196" pageNumber="197" pagination="181 - 202" refId="ref231211" refString="Smith L. B. &amp; Schubert B. G. 1941 a. Flora of Peru, Begoniaceae. Publications of the Field Museum of Natural History, Botanical Series 13 (4 / 1): 181 - 202." type="journal article" year="1941" yearSuffix="a">Smith &amp; Schubert 1941a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF61FF3DFE92F8A7ACDBEDD9" pageId="196" pageNumber="197" type="materials_examined">
<paragraph id="8BA4DA5DFF61FF3DFE92F8A7ACDBEDD9" blockId="196.[189,1399,1779,2024]" pageId="196" pageNumber="197">
–
<emphasis id="B96F064FFF61FF3DFEBEF8A8AC84ED48" bold="true" box="[359,419,1815,1841]" pageId="196" pageNumber="197">Type</emphasis>
:
<materialsCitation id="3B73D000FF61FF3DFE6DF8A7A9C6ED02" collectingDate="1939-04-04" collectionCode="UC" collectorName="H. E. Stork" country="Peru" county="Prov. Huancabamba" elevation="1200" latitude="-5.366667" location="Quebradas immediately NW of Canchaque" longLatPrecision="1305" longitude="-79.6" pageId="196" pageNumber="197" specimenCode="UC640259" specimenCount="1" stateProvince="Piura Region" typeStatus="holotype">
<collectingCountry id="F30C9ACDFF61FF3DFE6DF8A7ACD8ED4B" box="[436,511,1816,1842]" name="Peru" pageId="196" pageNumber="197">PERU</collectingCountry>
–
<emphasis id="B96F064FFF61FF3DFDFAF8A8AEC1ED48" bold="true" box="[547,998,1815,1841]" pageId="196" pageNumber="197">
<collectingRegion id="49DF14BFFF61FF3DFDFAF8A8AFF6ED48" box="[547,721,1815,1841]" country="Peru" name="Piura" pageId="196" pageNumber="197">Piura Region</collectingRegion>
:
<collectingCounty id="62C5A2D1FF61FF3DFD39F8A8AEC1ED48" box="[736,998,1815,1841]" pageId="196" pageNumber="197">Prov. Huancabamba</collectingCounty>
</emphasis>
•
<location id="8EC48C86FF61FF3DFBD8F8A8AC54ED2F" LSID="urn:lsid:plazi:treatment:03B26B4BFF61FF3FFDD3F90FA982E859:8EC48C86FF61FF3DFBD8F8A8AC54ED2F" country="Peru" county="Prov. Huancabamba" latitude="-5.366667" longLatPrecision="1305" longitude="-79.6" name="Quebradas immediately NW of Canchaque" pageId="196" pageNumber="197" stateProvince="Piura Region">Quebradas immediately NW of Canchaque</location>
;
<locationDeviation id="9A16C37FFF61FF3DFE5AF883A9DAED2F" box="[387,1277,1852,1878]" pageId="196" pageNumber="197">on clay banks beside trail, also commonly along road below Canchaque</locationDeviation>
; [
<geoCoordinate id="EE2FBC9AFF61FF3DFACCF885A856ED2F" box="[1301,1393,1850,1878]" degrees="5" direction="south" minutes="22" orientation="latitude" pageId="196" pageNumber="197" precision="925" value="-5.366667">5°22′ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF61FF3DFF33F8E0AC47ED02" box="[234,352,1887,1915]" degrees="79" direction="west" minutes="36" orientation="longitude" pageId="196" pageNumber="197" precision="925" value="-79.6">79°36′ W</geoCoordinate>
];
<elevation id="00363D6EFF61FF3DFEACF8DEAF36ED03" box="[373,529,1888,1915]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="196" pageNumber="197" unit="m" value="1200.0">
<quantity id="4CE377B8FF61FF3DFEACF8DEACF7ED02" box="[373,464,1889,1915]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="196" pageNumber="197" unit="m" value="1200.0">1200 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF61FF3DFDC6F8DEAF88ED02" box="[543,687,1889,1915]" pageId="196" pageNumber="197" value="1939-04-04">
<collectingDate id="EFE10575FF61FF3DFDC6F8DEAF88ED02" box="[543,687,1889,1915]" pageId="196" pageNumber="197" value="1939-04-04">4 Apr. 1939</collectingDate>
</date>
;
<emphasis id="B96F064FFF61FF3DFD65F8DEAEACED02" box="[700,907,1888,1915]" italics="true" pageId="196" pageNumber="197">
<collectorName id="26EEBF8BFF61FF3DFD65F8DEAE1DED03" box="[700,826,1888,1914]" pageId="196" pageNumber="197">H.E. Stork</collectorName>
11393
</emphasis>
;
<typeStatus id="54A064FFFF61FF3DFC42F8DFA921ED03" box="[923,1030,1888,1914]" pageId="196" pageNumber="197">holotype</typeStatus>
:
<collectionCode id="ED0A4298FF61FF3DFBCAF8DEA91BED02" box="[1043,1084,1889,1915]" country="USA" name="Upjohn Culture Collection" pageId="196" pageNumber="197">UC</collectionCode>
[
<specimenCode id="DBBD7226FF61FF3DFB92F8DFA9F2ED02" box="[1099,1237,1888,1915]" collectionCode="UC" country="USA" name="Upjohn Culture Collection" pageId="196" pageNumber="197">UC640259</specimenCode>
];
</materialsCitation>
<materialsCitation id="3B73D000FF61FF3DFB30F8DFACDBEDD9" collectingDate="1939-04-04" collectionCode="G, GH, K" collectorName="H. E. Stork" country="Peru" county="Prov. Huancabamba" elevation="1200" latitude="-5.366667" location="Quebradas immediately NW of Canchaque" longLatPrecision="1305" longitude="-79.6" pageId="196" pageNumber="197" specimenCode="GH00068265" specimenCount="1" stateProvince="Piura Region" typeStatus="isotypes">
<typeStatus id="54A064FFFF61FF3DFB30F8DFA869ED03" box="[1257,1358,1888,1914]" pageId="196" pageNumber="197">isotypes</typeStatus>
:
<collectionCode id="ED0A4298FF61FF3DFA83F8DEA855ED02" box="[1370,1394,1889,1915]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="196" pageNumber="197" type="Herbarium">G</collectionCode>
,
<collectionCode id="ED0A4298FF61FF3DFF35F83AAC31EDE6" box="[236,278,1925,1951]" country="USA" lsid="urn:lsid:biocol.org:col:15631" name="Harvard University - Gray Herbarium" pageId="196" pageNumber="197" type="Herbarium">GH</collectionCode>
[
<specimenCode id="DBBD7226FF61FF3DFEFDF83AACEAEDD9" box="[292,461,1925,1952]" collectionCode="GH" country="USA" lsid="urn:lsid:biocol.org:col:15631" name="Harvard University - Gray Herbarium" pageId="196" pageNumber="197" type="Herbarium">GH00068265</specimenCode>
],
<collectionCode id="ED0A4298FF61FF3DFE06F839ACDFEDD9" box="[479,504,1926,1952]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="196" pageNumber="197" type="Herbarium">K</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF61FF3DFF64F815AC5DED91" pageId="196" pageNumber="197" type="reference_group">
<paragraph id="8BA4DA5DFF61FF3DFF64F815AC5DED91" blockId="196.[189,1399,1779,2024]" pageId="196" pageNumber="197">
<bibRefCitation id="EF8AA7ACFF61FF3DFF64F815AF37EDBD" author="Smith &amp; Schubert" box="[189,528,1961,1988]" firstAuthor="Smith" page="39" pageId="196" pageNumber="197" pagination="36 - 40" refId="ref231547" refString="Smith L. B. &amp; Schubert B. G. 1952. Plants collected in Ecuador by W. H. Camp, Begoniaceae. Memoirs of the New York Botanical Garden 8 (1): 36 - 40." type="journal article" year="1952">Smith &amp; Schubert (1952: 39)</bibRefCitation>
;
<bibRefCitation id="EF8AA7ACFF61FF3DFDC5F815AEBDEDBD" author="Smith &amp; Wasshausen" box="[540,922,1962,1988]" firstAuthor="Smith" page="247" pageId="196" pageNumber="197" pagination="233 - 256" refId="ref231729" refString="Smith L. B. &amp; Wasshausen D. C. 1979. Begonia of Ecuador. Phytologia 44 (4): 233 - 256. https: // biostor. org / reference / 63675" type="journal article" year="1979">Smith &amp; Wasshausen (1979: 247</bibRefCitation>
,
<bibRefCitation id="EF8AA7ACFF61FF3DFC7DF815A93BEDBD" author="Smith &amp; Wasshausen" box="[932,1052,1962,1988]" firstAuthor="Smith" page="298" pageId="196" pageNumber="197" pagination="441 - 451" refId="ref231766" refString="Smith L. B. &amp; Wasshausen D. C. 1983. Notes on Begoniaceae, I. Phytologia 52 (7): 441 - 451. https: // biostor. org / reference / 208201" type="journal article" year="1983">1983: 298</bibRefCitation>
,
<bibRefCitation id="EF8AA7ACFF61FF3DFBFFF815A9B7EDBD" author="Smith &amp; Wasshausen" box="[1062,1168,1961,1988]" firstAuthor="Smith" page="52" pageId="196" pageNumber="197" pagination="4 - 65" refId="ref231872" refString="Smith L. B. &amp; Wasshausen D. C. 1986. Begoniaceae. In: Harling G. &amp; Andersson L. (eds) Flora of Ecuador 25 (133): 4 - 65." type="journal article" year="1986">1986: 52</bibRefCitation>
);
<bibRefCitation id="EF8AA7ACFF61FF3DFB7DF815AC51ED91" author="Brako &amp; Zarucchi" firstAuthor="Brako" page="194" pageId="196" pageNumber="197" refId="ref225797" refString="Brako L. &amp; Zarucchi L. 1993. Catalogue of the Flowering Plants and Gymnosperms of Peru. Monographs in Systematic Botany from the Missouri Botanical Garden 45, Missouri Botanical Garden, St Louis." type="book" year="1993">Brako &amp; Zarucchi (1993: 194)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF60FF3CFF64FEB6AC3CEB0F" pageId="197" pageNumber="198" type="etymology">
<paragraph id="8BA4DA5DFF60FF3CFF64FEB6AC62EB5A" blockId="197.[189,325,265,291]" box="[189,325,265,291]" pageId="197" pageNumber="198">
<heading id="D0EC6D31FF60FF3CFF64FEB6AC62EB5A" bold="true" box="[189,325,265,291]" fontSize="11" level="3" pageId="197" pageNumber="198" reason="3">
<emphasis id="B96F064FFF60FF3CFF64FEB6AC62EB5A" bold="true" box="[189,325,265,291]" pageId="197" pageNumber="198">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF60FF3CFF64FE87AC3CEB0F" blockId="197.[189,1398,312,374]" pageId="197" pageNumber="198">
The
<typeStatus id="54A064FFFF60FF3CFF28FE86AC03EB2A" box="[241,292,313,339]" pageId="197" pageNumber="198">type</typeStatus>
specimen was collected in Peru’s
<collectingRegion id="49DF14BFFF60FF3CFD6DFE87AE69EB2B" box="[692,846,312,338]" country="Peru" name="Piura" pageId="197" pageNumber="198">Piura Region</collectingRegion>
by H.E. Stork and the species is named for this locality.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF60FF3CFF64FE21A855E8B8" pageId="197" pageNumber="198" type="materials_examined">
<paragraph id="8BA4DA5DFF60FF3CFF64FE21ACE3EBC1" blockId="197.[189,452,414,440]" box="[189,452,414,440]" pageId="197" pageNumber="198">
<heading id="D0EC6D31FF60FF3CFF64FE21ACE3EBC1" bold="true" box="[189,452,414,440]" fontSize="11" level="3" pageId="197" pageNumber="198" reason="3">
<emphasis id="B96F064FFF60FF3CFF64FE21ACE3EBC1" bold="true" box="[189,452,414,440]" pageId="197" pageNumber="198">Specimens examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF60FF3CFF64FE70A855E8B8" blockId="197.[189,1399,461,706]" pageId="197" pageNumber="198">
<materialsCitation id="3B73D000FF60FF3CFF64FE70AC54E848" collectingDate="2004-06-22" collectionCode="US" collectorName="A. Sagastegui A. &amp; C. Tellez &amp; F. Torres &amp; M. Berru &amp; W. Garcia" country="Peru" county="Prov. Morropon" elevation="1320" latitude="-5.048611" location="El Palto-Santo Domingo" longLatPrecision="21" longitude="-79.851944" pageId="197" pageNumber="198" specimenCode="US01913139" specimenCount="1" stateProvince="Piura Region">
<collectingCountry id="F30C9ACDFF60FF3CFF64FE70AC2FEB90" box="[189,264,463,489]" name="Peru" pageId="197" pageNumber="198">PERU</collectingCountry>
–
<emphasis id="B96F064FFF60FF3CFEEAFE71AFEBEB91" bold="true" box="[307,716,461,488]" pageId="197" pageNumber="198">
<collectingRegion id="49DF14BFFF60FF3CFEEAFE71ACC3EB91" box="[307,484,462,488]" country="Peru" name="Piura" pageId="197" pageNumber="198">Piura Region</collectingRegion>
:
<collectingCounty id="62C5A2D1FF60FF3CFE21FE71AFEBEB91" box="[504,716,461,488]" pageId="197" pageNumber="198">Prov. Morropón</collectingCounty>
</emphasis>
•
<location id="8EC48C86FF60FF3CFD2BFE71A904EB91" LSID="urn:lsid:plazi:treatment:03B26B4BFF61FF3FFDD3F90FA982E859:8EC48C86FF60FF3CFD2BFE71A904EB91" box="[754,1059,462,488]" country="Peru" county="Prov. Morropon" latitude="-5.048611" longLatPrecision="21" longitude="-79.851944" name="El Palto-Santo Domingo" pageId="197" pageNumber="198" stateProvince="Piura Region">El Palto-Santo Domingo</location>
,
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,
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;
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<quantity id="4CE377B8FF60FF3CFF64FE4DAC30E874" box="[189,279,498,525]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="197" pageNumber="198" unit="m" value="1320.0">1320 m</quantity>
a.s.l.
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;
<date id="FFA5FC9DFF60FF3CFEBEFE4DAF23E875" box="[359,516,498,525]" pageId="197" pageNumber="198" value="2004-06-22">
<collectingDate id="EFE10575FF60FF3CFEBEFE4DAF23E875" box="[359,516,498,525]" pageId="197" pageNumber="198" value="2004-06-22">22 Jun. 2004</collectingDate>
</date>
;
<emphasis id="B96F064FFF60FF3CFDCAFE4CA867E875" box="[531,1344,498,525]" italics="true" pageId="197" pageNumber="198">
<collectorName id="26EEBF8BFF60FF3CFDCAFE4CAFFCE874" box="[531,731,499,525]" pageId="197" pageNumber="198">A. Sagástegui A.</collectorName>
,
<collectorName id="26EEBF8BFF60FF3CFD30FE4CAE70E875" box="[745,855,498,525]" pageId="197" pageNumber="198">C. Téllez</collectorName>
,
<collectorName id="26EEBF8BFF60FF3CFCBCFE4CAEF6E875" box="[869,977,499,524]" pageId="197" pageNumber="198">F. Torres</collectorName>
,
<collectorName id="26EEBF8BFF60FF3CFC06FE4CA96AE874" box="[991,1101,499,525]" pageId="197" pageNumber="198">M. Berrú</collectorName>
&amp;
<collectorName id="26EEBF8BFF60FF3CFBACFE4CA9C8E874" box="[1141,1263,499,525]" pageId="197" pageNumber="198">W. García</collectorName>
17485
</emphasis>
;
<collectionCode id="ED0A4298FF60FF3CFA88FE4DA851E875" box="[1361,1398,498,524]" pageId="197" pageNumber="198">US</collectionCode>
[
<specimenCode id="DBBD7226FF60FF3CFF1EFDA8AC45E848" box="[199,354,535,561]" pageId="197" pageNumber="198">US01913139</specimenCode>
].
</materialsCitation>
<materialsCitation id="3B73D000FF60FF3CFEA2FDA8A803E82C" collectingDate="1987-06-18" collectionCode="F, MO, USM" collectorName="C. Diaz &amp; S. Baldeon M." county="Prov. Huancabamba" elevation="1700" latitude="-5.350278" location="Chorro Blanco" longLatPrecision="21" longitude="-79.60361" municipality="Dist. Canchaque" pageId="197" pageNumber="198" specimenCode="MO-098011" specimenCount="1">
–
<emphasis id="B96F064FFF60FF3CFE4DFDA8AFBEE848" bold="true" box="[404,665,535,561]" pageId="197" pageNumber="198">
<collectingCounty id="62C5A2D1FF60FF3CFE4DFDA8AFBEE848" box="[404,665,535,561]" pageId="197" pageNumber="198">Prov. Huancabamba</collectingCounty>
</emphasis>
•
<collectingMunicipality id="6BC04027FF60FF3CFD6BFDA9AE5FE849" box="[690,888,534,560]" pageId="197" pageNumber="198">Dist. Canchaque</collectingMunicipality>
,
<location id="8EC48C86FF60FF3CFC5CFDA9A91FE849" LSID="urn:lsid:plazi:treatment:03B26B4BFF61FF3FFDD3F90FA982E859:8EC48C86FF60FF3CFC5CFDA9A91FE849" box="[901,1080,534,560]" country="United States of America" county="Prov. Huancabamba" latitude="-5.350278" longLatPrecision="21" longitude="-79.60361" municipality="Dist. Canchaque" name="Chorro Blanco" pageId="197" pageNumber="198">Chorro Blanco</location>
;
<geoCoordinate id="EE2FBC9AFF60FF3CFB9CFDAAA9E0E848" box="[1093,1223,533,561]" degrees="5" direction="south" minutes="21" orientation="latitude" pageId="197" pageNumber="198" precision="15" seconds="01" value="-5.350278">5°21′01″ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF60FF3CFB0CFDAAA855E848" box="[1237,1394,533,561]" degrees="79" direction="west" minutes="36" orientation="longitude" pageId="197" pageNumber="198" precision="15" seconds="13" value="-79.60361">79°36′13″ W</geoCoordinate>
;
<elevation id="00363D6EFF60FF3CFF64FD84AC86E82C" box="[189,417,571,598]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="197" pageNumber="198" unit="m" value="1700.0" valueMax="1900.0" valueMin="1500.0">
<quantity id="4CE377B8FF60FF3CFF64FD84AC47E82C" box="[189,352,571,598]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="197" pageNumber="198" unit="m" value="1700.0" valueMax="1900.0" valueMin="1500.0">1500–1900 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF60FF3CFE69FD84AF6BE82C" box="[432,588,571,597]" pageId="197" pageNumber="198" value="1987-06-18">
<collectingDate id="EFE10575FF60FF3CFE69FD84AF6BE82C" box="[432,588,571,597]" pageId="197" pageNumber="198" value="1987-06-18">18 Jun. 1987</collectingDate>
</date>
;
<emphasis id="B96F064FFF60FF3CFD83FD84AEEDE82C" box="[602,970,571,597]" italics="true" pageId="197" pageNumber="198">
<collectorName id="26EEBF8BFF60FF3CFD83FD84AF91E82C" box="[602,694,571,597]" pageId="197" pageNumber="198">C. Díaz</collectorName>
&amp;
<collectorName id="26EEBF8BFF60FF3CFD07FD84AEAFE82C" box="[734,904,571,597]" pageId="197" pageNumber="198">S. Baldeón M.</collectorName>
2465
</emphasis>
;
<collectionCode id="ED0A4298FF60FF3CFC02FD84AECBE82C" box="[987,1004,571,597]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="197" pageNumber="198" type="Herbarium">F</collectionCode>
,
<collectionCode id="ED0A4298FF60FF3CFC20FD84A90FE82C" box="[1017,1064,571,597]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="197" pageNumber="198" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FF60FF3CFBE3FD84A9EDE82C" box="[1082,1226,571,597]" pageId="197" pageNumber="198">MO-098011</specimenCode>
],
<collectionCode id="ED0A4298FF60FF3CFB3DFD84A803E82C" box="[1252,1316,571,597]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="197" pageNumber="198">USM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B73D000FF60FF3CFAE7FD84A806E800" collectingDate="1948-05-22" collectionCode="USM" collectorName="R. Ferreyra" elevation="1250" latitude="-5.366667" location="Dist. Canchaque" longLatPrecision="1305" longitude="-79.6" municipality="Dist. Canchaque" pageId="197" pageNumber="198" specimenCount="1">
<collectingMunicipality id="6BC04027FF60FF3CFAE7FD84AC63E800" pageId="197" pageNumber="198">Dist. Canchaque</collectingMunicipality>
; [
<geoCoordinate id="EE2FBC9AFF60FF3CFE83FDE2AC92E800" box="[346,437,605,633]" degrees="5" direction="south" minutes="22" orientation="latitude" pageId="197" pageNumber="198" precision="925" value="-5.366667">5°22′ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF60FF3CFE1DFDE2AF1CE800" box="[452,571,605,633]" degrees="79" direction="west" minutes="36" orientation="longitude" pageId="197" pageNumber="198" precision="925" value="-79.6">79°36′ W</geoCoordinate>
];
<elevation id="00363D6EFF60FF3CFD8BFDE0AE1EE800" box="[594,825,607,633]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="197" pageNumber="198" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">
<quantity id="4CE377B8FF60FF3CFD8BFDE0AFD0E800" box="[594,759,607,633]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="197" pageNumber="198" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">1200–1300 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF60FF3CFC90FDE0AEC9E800" box="[841,1006,607,633]" pageId="197" pageNumber="198" value="1948-05-22">
<collectingDate id="EFE10575FF60FF3CFC90FDE0AEC9E800" box="[841,1006,607,633]" pageId="197" pageNumber="198" value="1948-05-22">22 May 1948</collectingDate>
</date>
;
<emphasis id="B96F064FFF60FF3CFC25FDDFA9E8E800" box="[1020,1231,607,633]" italics="true" pageId="197" pageNumber="198">
<collectorName id="26EEBF8BFF60FF3CFC25FDDFA9ADE800" box="[1020,1162,608,633]" pageId="197" pageNumber="198">R. Ferreyra</collectorName>
3092
</emphasis>
;
<collectionCode id="ED0A4298FF60FF3CFB38FDE0A806E800" box="[1249,1313,607,633]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="197" pageNumber="198">USM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B73D000FF60FF3CFAE7FDE0A855E8B8" collectingDate="2016-01-27" collectionCode="E, MO, MOL" collectorName="P. W. Moonlight &amp; A. Daza" elevation="1199" latitude="-5.375" location="Ravine 100 m from centre of Canchaque" longLatPrecision="21" longitude="-79.606384" municipality="Dist. Canchaque" pageId="197" pageNumber="198" specimenCode="E00885468" specimenCount="1">
<collectingMunicipality id="6BC04027FF60FF3CFAE7FDE0AC65E8E4" pageId="197" pageNumber="198">Dist. Canchaque</collectingMunicipality>
,
<location id="8EC48C86FF60FF3CFE89FD3CAE67E8E4" LSID="urn:lsid:plazi:treatment:03B26B4BFF61FF3FFDD3F90FA982E859:8EC48C86FF60FF3CFE89FD3CAE67E8E4" box="[336,832,643,670]" latitude="-5.375" longLatPrecision="21" longitude="-79.606384" municipality="Dist. Canchaque" name="Ravine 100 m from centre of Canchaque" pageId="197" pageNumber="198">
Ravine
<quantity id="4CE377B8FF60FF3CFE74FD3CACDDE8E7" box="[429,506,643,670]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="197" pageNumber="198" unit="m" value="100.0">100 m</quantity>
from centre of Canchaque
</location>
,
<locationDeviation id="9A16C37FFF60FF3CFC97FD3BA9F8E8E4" box="[846,1247,643,670]" pageId="197" pageNumber="198">over metal suspension footbridge</locationDeviation>
,
<geoCoordinate id="EE2FBC9AFF60FF3CFB34FD3DA856E8E4" box="[1261,1393,642,670]" degrees="5" direction="south" minutes="22" orientation="latitude" pageId="197" pageNumber="198" precision="15" seconds="30" value="-5.375">5°22′30″ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF60FF3CFF64FD19AC7FE8BB" box="[189,344,678,706]" degrees="79" direction="west" minutes="36" orientation="longitude" pageId="197" pageNumber="198" precision="15" seconds="23" value="-79.606384">79°36′23″ W</geoCoordinate>
;
<elevation id="00363D6EFF60FF3CFEBDFD18ACDBE8B8" box="[356,508,679,706]" metricMagnitude="3" metricUnit="m" metricValue="1.199" pageId="197" pageNumber="198" unit="m" value="1199.0">
<quantity id="4CE377B8FF60FF3CFEBDFD18AC9BE8BB" box="[356,444,679,706]" metricMagnitude="3" metricUnit="m" metricValue="1.199" pageId="197" pageNumber="198" unit="m" value="1199.0">1199 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF60FF3CFDD0FD18AF85E8B8" box="[521,674,679,706]" pageId="197" pageNumber="198" value="2016-01-27">
<collectingDate id="EFE10575FF60FF3CFDD0FD18AF85E8B8" box="[521,674,679,706]" pageId="197" pageNumber="198" value="2016-01-27">27 Jan. 2016</collectingDate>
</date>
;
<emphasis id="B96F064FFF60FF3CFD77FD17A93EE8B8" box="[686,1049,679,706]" italics="true" pageId="197" pageNumber="198">
<collectorName id="26EEBF8BFF60FF3CFD77FD17AE43E8B8" box="[686,868,679,705]" pageId="197" pageNumber="198">P.W. Moonlight</collectorName>
&amp;
<collectorName id="26EEBF8BFF60FF3CFC50FD17AECEE8B8" box="[905,1001,680,706]" pageId="197" pageNumber="198">A. Daza</collectorName>
111
</emphasis>
;
<collectionCode id="ED0A4298FF60FF3CFBF1FD17A91DE8BB" box="[1064,1082,680,706]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="197" pageNumber="198" type="Herbarium">E</collectionCode>
[
<specimenCode id="DBBD7226FF60FF3CFB92FD18A9F4E8BB" box="[1099,1235,679,706]" collectionCode="E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="197" pageNumber="198" type="Herbarium">E00885468</specimenCode>
],
<collectionCode id="ED0A4298FF60FF3CFB32FD18A83AE8B8" box="[1259,1309,679,705]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="197" pageNumber="198" type="Herbarium">MO</collectionCode>
,
<collectionCode id="ED0A4298FF60FF3CFAF0FD18A849E8B8" box="[1321,1390,679,705]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="197" pageNumber="198" type="Herbarium">MOL</collectionCode>
.
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</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF60FF3CFF64FD56A81EEC71" pageId="197" pageNumber="198" type="description">
<paragraph id="8BA4DA5DFF60FF3CFF64FD56AC68E97A" blockId="197.[189,335,745,771]" box="[189,335,745,771]" pageId="197" pageNumber="198">
<heading id="D0EC6D31FF60FF3CFF64FD56AC68E97A" bold="true" box="[189,335,745,771]" fontSize="11" level="3" pageId="197" pageNumber="198" reason="3">
<emphasis id="B96F064FFF60FF3CFF64FD56AC68E97A" bold="true" box="[189,335,745,771]" pageId="197" pageNumber="198">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF60FF3CFF64FCA6A81EEC71" blockId="197.[188,1400,793,1544]" pageId="197" pageNumber="198">
Caulescent, tuberous herb, to
<quantity id="4CE377B8FF60FF3CFDC2FCA6AF6FE94D" box="[539,584,793,820]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="197" pageNumber="198" unit="m" value="1.0">1 m</quantity>
high.
<emphasis id="B96F064FFF60FF3CFD4AFCA6AFFFE94A" box="[659,728,793,819]" italics="true" pageId="197" pageNumber="198">Tuber</emphasis>
spheroid, ca 4 ×
<quantity id="4CE377B8FF60FF3CFC7DFCA6AEC5E94D" box="[932,994,793,820]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="197" pageNumber="198" unit="cm" value="4.0">4 cm</quantity>
, with 1 growing point.
<emphasis id="B96F064FFF60FF3CFB2CFCA5A809E94D" box="[1269,1326,794,820]" italics="true" pageId="197" pageNumber="198">Stem</emphasis>
erect, unbranched; internodes to
<quantity id="4CE377B8FF60FF3CFE23FC81AF63E921" box="[506,580,830,856]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="197" pageNumber="198" unit="cm" value="20.0">20 cm</quantity>
long, to
<quantity id="4CE377B8FF60FF3CFD6BFC81AFD0E921" box="[690,759,830,856]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="197" pageNumber="198" unit="mm" value="7.0">7 mm</quantity>
thick, succulent, green, glabrous.
<emphasis id="B96F064FFF60FF3CFB48FC82A9C8E92E" box="[1169,1263,829,855]" italics="true" pageId="197" pageNumber="198">Stipules</emphasis>
deciduous, ovate, ca 12 ×
<quantity id="4CE377B8FF60FF3CFEB2FCDDAC94E905" box="[363,435,866,892]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="197" pageNumber="198" unit="mm" value="6.0">6 mm</quantity>
, apex acute, translucent, pale green, glabrous, margin entire, aciliate.
<emphasis id="B96F064FFF60FF3CFB30FCDDA81CE902" box="[1257,1339,866,891]" italics="true" pageId="197" pageNumber="198">Leaves</emphasis>
3–5, alternate, basifixed; petiole
<quantity id="4CE377B8FF60FF3CFDD6FC39AFA7E9D9" box="[527,640,902,929]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="8.5" metricValueMin="3.0" pageId="197" pageNumber="198" unit="cm" value="5.75" valueMax="8.5" valueMin="3.0">3–8.5 cm</quantity>
long, light green, glabrous; blade asymmetric, ovate, to 16 ×
<quantity id="4CE377B8FF60FF3CFF64FC15ADDEE9BC" box="[189,249,938,965]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="197" pageNumber="198" unit="cm" value="8.0">8 cm</quantity>
, succulent, apex acuminate, base cordate, basal lobes not overlapping, sinus to
<quantity id="4CE377B8FF60FF3CFB4CFC15A9FFE9BC" box="[1173,1240,938,965]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="197" pageNumber="198" unit="mm" value="2.0">2 mm</quantity>
deep, margin dentate, ciliate, upper surface dark green, glabrous, lower surface pale green, glabrous, veins palmate-pinnate, 6–8 veined from the base, with 2–4 secondary veins on the larger side, 1–3 secondary veins on the smaller side.
<emphasis id="B96F064FFF60FF3CFE6FFBA8AF79EE48" box="[438,606,1047,1073]" italics="true" pageId="197" pageNumber="198">Inflorescences</emphasis>
1–3 per stem, bisexual, axillary, erect, cymose, with 6 branches, bearing up to 32 staminate flowers and 32 pistillate flowers, protandrous; peduncle to
<quantity id="4CE377B8FF60FF3CFB6BFB84A9DCEE2C" box="[1202,1275,1083,1109]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="197" pageNumber="198" unit="cm" value="11.0">11 cm</quantity>
long, pale green, glabrous, bracts deciduous, elliptic to ovate, 4–6 ×
<quantity id="4CE377B8FF60FF3CFC59FBE0AEDAEE03" box="[896,1021,1119,1146]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="197" pageNumber="198" unit="mm" value="2.75" valueMax="4.0" valueMin="1.5">1.5–4 mm</quantity>
, opaque, pale brown, glabrous, apex truncate, margin undulate, ciliate.
<emphasis id="B96F064FFF60FF3CFD4AFB3BAE41EEE4" box="[659,870,1155,1182]" italics="true" pageId="197" pageNumber="198">Staminate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FF60FF3CFC26FB3CA975EEE7" box="[1023,1106,1155,1182]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="197" pageNumber="198" unit="mm" value="10.0">10 mm</quantity>
long, glabrous; tepals 4, spreading, outer 2 ovate, 6–8 ×
<quantity id="4CE377B8FF60FF3CFDE1FB18AFB9EEBB" box="[568,670,1191,1218]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="197" pageNumber="198" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
, apex rounded, white, glabrous, margin entire, ciliate, inner 2 oblanceolate, 7–8 ×
<quantity id="4CE377B8FF60FF3CFE74FB73AF36EE9F" box="[429,529,1228,1254]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="197" pageNumber="198" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
, apex rounded, white to pink, glabrous, margin entire, aciliate; stamens ca 25, spreading, free, filaments
<quantity id="4CE377B8FF60FF3CFDCCFB4FAFACEF73" box="[533,651,1264,1291]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="197" pageNumber="198" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 mm</quantity>
long, free, anthers elliptic, 0.5–1 ×
<quantity id="4CE377B8FF60FF3CFBFFFB4FA9A5EF73" box="[1062,1154,1264,1290]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="197" pageNumber="198" unit="mm" value="0.4">0.4 mm</quantity>
, dehiscing via lateral slits, connectives extending to
<quantity id="4CE377B8FF60FF3CFDF2FAABAFAEEF57" box="[555,649,1300,1326]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="197" pageNumber="198" unit="mm" value="0.2">0.2 mm</quantity>
, symmetrically basifixed.
<emphasis id="B96F064FFF60FF3CFC19FAABA9AEEF57" box="[960,1161,1300,1326]" italics="true" pageId="197" pageNumber="198">Pistillate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FF60FF3CFAFAFAABA850EF57" box="[1315,1399,1300,1326]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="197" pageNumber="198" unit="mm" value="27.0">27 mm</quantity>
long; bracteoles lacking; tepals 5, subequal, deciduous in fruit, spreading, obovate to broadly obovate, 4.5–9 ×
<quantity id="4CE377B8FF60FF3CFEC4FAE3ACB1EF0E" box="[285,406,1372,1399]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="7.5" metricValueMin="2.0" pageId="197" pageNumber="198" unit="mm" value="4.75" valueMax="7.5" valueMin="2.0">2–7.5 mm</quantity>
, apex obtuse, white to pink, glabrous, margin entire, aciliate; ovary body ovoid, 4.5–7 ×
<quantity id="4CE377B8FF60FF3CFF0DFA3FAC68EFE2" box="[212,335,1408,1435]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="6.5" metricValueMin="4.0" pageId="197" pageNumber="198" unit="mm" value="5.25" valueMax="6.5" valueMin="4.0">4–6.5 mm</quantity>
, white to pink, glabrous, sub-equally 3-winged, wings triangular, 5.5–10.5 ×
<quantity id="4CE377B8FF60FF3CFB06FA3FA857EFE2" box="[1247,1392,1408,1435]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.5" metricValueMin="3.5" pageId="197" pageNumber="198" unit="mm" value="6.0" valueMax="8.5" valueMin="3.5">3.5–8.5 mm</quantity>
; 3-locular, placentae branches divided, bearing ovules on both surfaces; styles 3, yellow, free, ca
<quantity id="4CE377B8FF60FF3CFAEAFA1AA85FEFC6" box="[1331,1400,1445,1471]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="197" pageNumber="198" unit="mm" value="2.0">2 mm</quantity>
long, once-divided, stigmatic papillae in a spirally twisted band.
<emphasis id="B96F064FFF60FF3CFC6FFA75A950EF9A" box="[950,1143,1481,1507]" italics="true" pageId="197" pageNumber="198">Fruiting pedicel</emphasis>
to
<quantity id="4CE377B8FF60FF3CFB45FA76A9C8EF9A" box="[1180,1263,1481,1507]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="197" pageNumber="198" unit="mm" value="32.0">32 mm</quantity>
long.
<emphasis id="B96F064FFF60FF3CFAE3FA75ADD1EC7E" italics="true" pageId="197" pageNumber="198">Fruit body</emphasis>
ovoid, to 9 ×
<quantity id="4CE377B8FF60FF3CFE42FA52ACC5EC71" box="[411,482,1517,1544]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="197" pageNumber="198" unit="mm" value="8.0">8 mm</quantity>
, drying brown, wings same shape as in ovary, expanding to 11 ×
<quantity id="4CE377B8FF60FF3CFB06FA52A812EC71" box="[1247,1333,1517,1544]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="197" pageNumber="198" unit="mm" value="10.0">10 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF60FF3CFF64F98FAFD9ED73" pageId="197" pageNumber="198" type="conservation">
<paragraph id="8BA4DA5DFF60FF3CFF64F98FAF4AEC33" blockId="197.[189,621,1583,1610]" box="[189,621,1583,1610]" pageId="197" pageNumber="198">
<heading id="D0EC6D31FF60FF3CFF64F98FAF4AEC33" bold="true" box="[189,621,1583,1610]" fontSize="11" level="3" pageId="197" pageNumber="198" reason="3">
<emphasis id="B96F064FFF60FF3CFF64F98FAF4AEC33" bold="true" box="[189,621,1583,1610]" pageId="197" pageNumber="198">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF60FF3CFF64F9DFAFD9ED73" blockId="197.[189,1399,1631,1802]" pageId="197" pageNumber="198">
Known from two populations within
<collectingCountry id="F30C9ACDFF60FF3CFD51F9DFAFE6EC03" box="[648,705,1632,1658]" name="Peru" pageId="197" pageNumber="198">Peru</collectingCountry>
, which are both close to rapidly expanding villages and vulnerable to agricultural and urban expansion. It is relatively common and frequently collected in
<collectingCountry id="F30C9ACDFF60FF3CFF64F918AC38ECB8" box="[189,287,1703,1729]" name="Ecuador" pageId="197" pageNumber="198">Ecuador</collectingCountry>
where it is known from ca 5 populations, and the total EOO of the species is ca 18
<quantity id="4CE377B8FF60FF3CFAD5F917A841ECB8" box="[1292,1382,1703,1730]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="197" pageNumber="198" unit="km" value="400.0">400 km</quantity>
<superScript id="7C6E7715FF60FF3CFABEF918A857ECCC" attach="right" box="[1383,1392,1703,1717]" fontSize="6" pageId="197" pageNumber="198">2</superScript>
. Given the species’ EOO, relatively few populations, and the threats to its Peruvian populations, we assess
<taxonomicName id="4C1BA1DEFF60FF3CFED5F94EACBEED73" box="[268,409,1777,1802]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="197" pageNumber="198" phylum="Tracheophyta" rank="species" species="piurensis">
<emphasis id="B96F064FFF60FF3CFED5F94EACBEED73" box="[268,409,1777,1802]" italics="true" pageId="197" pageNumber="198">B. piurensis</emphasis>
</taxonomicName>
as Vulnerable (VU B1ab(iii)).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF60FF3FFF64F88DA948EB3F" lastPageId="198" lastPageNumber="199" pageId="197" pageNumber="198" type="discussion">
<paragraph id="8BA4DA5DFF60FF3CFF64F88DAC97ED35" blockId="197.[189,432,1842,1868]" box="[189,432,1842,1868]" pageId="197" pageNumber="198">
<heading id="D0EC6D31FF60FF3CFF64F88DAC97ED35" bold="true" box="[189,432,1842,1868]" fontSize="11" level="3" pageId="197" pageNumber="198" reason="3">
<emphasis id="B96F064FFF60FF3CFF64F88DAC97ED35" bold="true" box="[189,432,1842,1868]" pageId="197" pageNumber="198">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF60FF3FFF64F8DCA948EB3F" blockId="197.[189,1400,1890,2025]" lastBlockId="198.[189,1401,265,327]" lastPageId="198" lastPageNumber="199" pageId="197" pageNumber="198">
<taxonomicName id="4C1BA1DEFF60FF3CFF64F8DCACB1ED05" box="[189,406,1891,1916]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="197" pageNumber="198" phylum="Tracheophyta" rank="species" species="piurensis">
<emphasis id="B96F064FFF60FF3CFF64F8DCACB1ED05" box="[189,406,1891,1916]" italics="true" pageId="197" pageNumber="198">Begonia piurensis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C1BA1DEFF60FF3CFD59F8DCAE7DED05" authority="L. B. Sm." authorityName="L. B. Sm." box="[640,858,1890,1916]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="197" pageNumber="198" phylum="Tracheophyta" rank="species" species="velata">
<emphasis id="B96F064FFF60FF3CFD59F8DCAFCBED05" box="[640,748,1890,1916]" italics="true" pageId="197" pageNumber="198">B. velata</emphasis>
L.B.Sm.
</taxonomicName>
&amp;. B.G.Schub. (see below) but superficially resembles several species of
<emphasis id="B96F064FFF60FF3CFDC2F838AF12EDD9" box="[539,565,1927,1952]" italics="true" pageId="197" pageNumber="198">B.</emphasis>
sect.
<taxonomicName id="4C1BA1DEFF60FF3CFDA2F839AE04EDD9" authorityName="Klotzsch, Gattungen und Arten 1854: 247 (Klotzsch" authorityYear="1855" baseAuthorityName="Klotzsch" box="[635,803,1926,1952]" class="Magnoliopsida" family="Begoniaceae" genus="Cyathocnemis" kingdom="Plantae" order="Cucurbitales" pageId="197" pageNumber="198" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F064FFF60FF3CFDA2F839AE04EDD9" box="[635,803,1926,1952]" italics="true" pageId="197" pageNumber="198">Cyathocnemis</emphasis>
</taxonomicName>
in that it is a large, upright herb with stems with elongate internodes and transversely ovate leaves. It differs from all species in this section in its staminate flowers with four tepals (vs two) and in having a tuber, but this is rarely recorded on herbarium labels. Note that while
<taxonomicName id="4C1BA1DEFF63FF3FFE59FEB5AF28EB5A" box="[384,527,266,291]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="198" pageNumber="199" phylum="Tracheophyta" rank="species" species="piurensis">
<emphasis id="B96F064FFF63FF3FFE59FEB5AF28EB5A" box="[384,527,266,291]" italics="true" pageId="198" pageNumber="199">B. piurensis</emphasis>
</taxonomicName>
is tuberous we include it in the key as both a tuberous and non-tuberous species because this character is only noted on a single Peruvian specimen label.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF63FF3FFF64FED2A982E859" pageId="198" pageNumber="199" type="distribution">
<paragraph id="8BA4DA5DFF63FF3FFF64FED2ACD2EBF1" blockId="198.[189,501,365,392]" box="[189,501,365,392]" pageId="198" pageNumber="199">
<heading id="D0EC6D31FF63FF3FFF64FED2ACD2EBF1" bold="true" box="[189,501,365,392]" fontSize="11" level="3" pageId="198" pageNumber="199" reason="3">
<emphasis id="B96F064FFF63FF3FFF64FED2ACD2EBF1" bold="true" box="[189,501,365,392]" pageId="198" pageNumber="199">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF63FF3FFF64FE22A982E859" blockId="198.[188,1398,412,544]" pageId="198" pageNumber="199">
Known from
<collectingCountry id="F30C9ACDFF63FF3FFE82FE23AC9AEBCF" box="[347,445,412,438]" name="Ecuador" pageId="198" pageNumber="199">Ecuador</collectingCountry>
and
<collectingCountry id="F30C9ACDFF63FF3FFE2CFE22AF09EBCE" box="[501,558,413,439]" name="Peru" pageId="198" pageNumber="199">Peru</collectingCountry>
, including
<collectingRegion id="49DF14BFFF63FF3FFD76FE23AE6EEBCF" box="[687,841,412,438]" country="Peru" name="Piura" pageId="198" pageNumber="199">Piura Region</collectingRegion>
in
<collectingCountry id="F30C9ACDFF63FF3FFCB7FE22AE8FEBCE" box="[878,936,413,439]" name="Peru" pageId="198" pageNumber="199">Peru</collectingCountry>
. Known from northwest Peruvian relict montane forest at an elevation of
<quantity id="4CE377B8FF63FF3FFDE7FE00AFF8EBA3" box="[574,735,447,474]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.9" metricValueMin="1.2" pageId="198" pageNumber="199" unit="m" value="1550.0" valueMax="1900.0" valueMin="1200.0">1200–1900 m</quantity>
a.s.l. (
<figureCitation id="1320C6D8FF63FF3FFCF3FE00AEB3EBA0" box="[810,916,447,473]" captionStart="Fig" captionStartId="193.[189,232,1892,1918]" captionTargetBox="[194,1392,1227,1846]" captionTargetId="figure-268@193.[189,1398,1217,1852]" captionTargetPageId="193" captionText="Fig. 62. Distribution of Begonia sect. Knesebeckia in Peru. A. B. brandbygeana L.B.Sm. &amp; Wassh. (red) and B. parcifolia (blue). B. B. piurensis L.B.Sm. &amp; B.G.Schub. (red) and B. velata L.B.Sm. &amp; B.G.Schub. (blue). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178434" httpUri="https://zenodo.org/record/8178434/files/figure.png" pageId="198" pageNumber="199">Fig. 62B</figureCitation>
).
<taxonomicName id="4C1BA1DEFF63FF3FFC7FFE7FA95CEBA0" box="[934,1147,448,473]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="198" pageNumber="199" phylum="Tracheophyta" rank="species" species="piurensis">
<emphasis id="B96F064FFF63FF3FFC7FFE7FA95CEBA0" box="[934,1147,448,473]" italics="true" pageId="198" pageNumber="199">Begonia piurensis</emphasis>
</taxonomicName>
is a tuberous species, and it appears to die back to its tuber in the dry season before beginning to grow at the start of the wet season (January) before flowering and fruiting later in the wet season (April to June).
</paragraph>
</subSubSection>
</treatment>
</document>
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