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<document id="6A8EB4B4EB3BA3CCF71D597F1ADA66AE" ID-DOI="https://doi.org/10.5852/ejt.2023.881.2175" ID-ISSN="2118-9773" ID-Zenodo-Dep="8178280" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690201937279" checkinUser="felipe" docAuthor="Moonlight, Peter. W., Jara-Muñoz, Orlando A., Purvis, David A., Delves, Jay, Allen, Josh P. &amp; Reynel, Carlos" docDate="2023" docId="03B26B4BFF0DFF50FE3EFEB6AE2BEFF8" docLanguage="en" docName="ejt-881_1-334.imf" docOrigin="European Journal of Taxonomy 881" docSource="http://dx.doi.org/10.5852/ejt.2023.881.2175" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia octopetala subsp. ovatiformis Irmsch." docType="treatment" docVersion="5" lastPageNumber="170" masterDocId="FF8B1333FFA5FFF9FFD9FFBFAD27EA79" masterDocTitle="The genus Begonia (Begoniaceae) in Peru" masterLastPageNumber="334" masterPageNumber="1" pageNumber="169" updateTime="1707123162844" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F2F11662BA7811F197075F0237A3F4B4">The genus Begonia (Begoniaceae) in Peru</mods:title>
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<mods:namePart id="DC247E9DFEFE42E83D6ABB5C5E239424">Moonlight, Peter. W.</mods:namePart>
<mods:affiliation id="4BC10A000B75CD4F0EF8257D4CAD9AC3">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK. &amp; Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland.</mods:affiliation>
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<mods:namePart id="DCF54FE0E31B6B49C15032B086D96588">Jara-Muñoz, Orlando A.</mods:namePart>
<mods:affiliation id="E0CD7E35920F8D53F1BE385B36E09F6A">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá D. C., Colombia.</mods:affiliation>
<mods:nameIdentifier id="BE83D94721983DC2CD64A0B1099DC438" type="email">oajaram@unal.edu.co</mods:nameIdentifier>
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<mods:namePart id="3598AC42502887D511E6ADD40F61E261">Purvis, David A.</mods:namePart>
<mods:affiliation id="2623A9653E99FE5C564CA844D07F8978">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="1DB29BF8DDDA1DE2BDAC3F0DB05C7227" type="email">dpurvis@rbge.org.uk</mods:nameIdentifier>
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<mods:namePart id="88F39D4083856AC435CEC6CA547A5E9E">Delves, Jay</mods:namePart>
<mods:affiliation id="F6FE9904CE03EE27CE9D493A4D756194">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="631D1BF4EE193D809CAA046AB30EE35E" type="email">j.delves@westernsydney.edu.au</mods:nameIdentifier>
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<mods:namePart id="D53019714EA2FCF0E52D3199C16556E8">Allen, Josh P.</mods:namePart>
<mods:affiliation id="F9894963FA8B9EE63A24A29781CA9B06">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
<mods:nameIdentifier id="EEEDCCE5DFAD86E2FEE19AAA825F62CF" type="email">myrmeciaman@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B960AEA32B3EFB73450F89077BD19775">Reynel, Carlos</mods:namePart>
<mods:affiliation id="90A3BFFEA79153B609B540DDC88D5949">Facultad de Ciencias Forestales, Universidad Nacional Agraria-La Molina, Lima 12, Peru.</mods:affiliation>
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<mods:title id="714511466C670CAF172A81E4B4D63D89">European Journal of Taxonomy</mods:title>
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<mods:date id="F6FF1024F31DFDC05458DFF9993557BB">2023</mods:date>
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<mods:number id="14D55DEA9FBFFDCAD8AEAF57262C106F">2023-07-18</mods:number>
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<mods:number id="BA47482E77423BD6C10BABEC39EA92B5">881</mods:number>
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<treatment id="03B26B4BFF0DFF50FE3EFEB6AE2BEFF8" LSID="urn:lsid:plazi:treatment:03B26B4BFF0DFF50FE3EFEB6AE2BEFF8" httpUri="http://treatment.plazi.org/id/03B26B4BFF0DFF50FE3EFEB6AE2BEFF8" lastPageId="169" lastPageNumber="170" pageId="168" pageNumber="169">
<subSubSection id="C30189D6FF0DFF51FE3EFEB6A96AEB5A" box="[487,1101,264,291]" pageId="168" pageNumber="169" type="nomenclature">
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36.2
<taxonomicName id="4C1BA1DEFF0DFF51FDFBFEB6A96AEB5A" ID-CoL="8CKQ4" authority="Irmsch." authorityName="Irmsch." box="[546,1101,264,291]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="168" pageNumber="169" phylum="Tracheophyta" rank="subSpecies" species="octopetala" subSpecies="ovatiformis">
<emphasis id="B96F064FFF0DFF51FDFBFEB6AE2DEB5A" bold="true" box="[546,778,264,291]" italics="true" pageId="168" pageNumber="169">Begonia octopetala</emphasis>
subsp.
<emphasis id="B96F064FFF0DFF51FCBAFEB7AECBEB5B" bold="true" box="[867,1004,264,290]" italics="true" pageId="168" pageNumber="169">ovatiformis</emphasis>
Irmsch.
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</paragraph>
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<subSubSection id="C30189D6FF0DFF51FD3FFE90AE69EB33" box="[742,846,303,330]" pageId="168" pageNumber="169" type="description">
<paragraph id="8BA4DA5DFF0DFF51FD3FFE90AE69EB33" blockId="168.[487,1101,264,330]" box="[742,846,303,330]" pageId="168" pageNumber="169">
<figureCitation id="1320C6D8FF0DFF51FD3FFE90AE69EB33" box="[742,846,303,330]" captionStart="Fig" captionStartId="160.[189,232,1822,1848]" captionTargetBox="[194,1391,558,1776]" captionTargetId="figure-14@160.[189,1398,548,1784]" captionTargetPageId="160" captionText="Fig. 52. Distribution of the octopetala group of Begonia sect. Eupetalum (Lindl.) A.DC. in Peru and surrounding countries. A. B. anemoniflora Irmsch. (red) and B. tumbezensis Irmsch. (blue). B. B. octopetala L.Hér. subsp. octopetala (red) and B. octopetala subsp. ovatiformis Irmsch. (blue). C. B. pleiopetala A.DC. (red) and B. pseudopleiopetala Tebbitt (blue). D. B. polypetala A.DC. (red). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178413" httpUri="https://zenodo.org/record/8178413/files/figure.png" pageId="168" pageNumber="169">Fig. 52B</figureCitation>
</paragraph>
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<subSubSection id="C30189D6FF0DFF51FF64FECBAC1FEBCC" pageId="168" pageNumber="169" type="reference_group">
<paragraph id="8BA4DA5DFF0DFF51FF64FECBAC1FEBCC" blockId="168.[189,1398,372,514]" pageId="168" pageNumber="169">
<emphasis id="B96F064FFF0DFF51FF64FECBA981EBF7" box="[189,1190,372,398]" italics="true" pageId="168" pageNumber="169">Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie</emphasis>
76: 75 (
<bibRefCitation id="EF8AA7ACFF0DFF51FAD5FECBAC0CEBCC" author="Irmscher E." firstAuthor="Irmscher" pageId="168" pageNumber="169" pagination="1 - 102" refId="ref228521" refString="Irmscher E. 1953. Systematische Studien uber Begoniaceen des tropischen Sudamerikas, besonders Brasiliens. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 76 (1): 1 - 102." type="journal article" year="1953">Irmscher 1953</bibRefCitation>
).
</paragraph>
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<subSubSection id="C30189D6FF0DFF51FEE5FE24A851EBA2" pageId="168" pageNumber="169" type="materials_examined">
<paragraph id="8BA4DA5DFF0DFF51FEE5FE24A851EBA2" blockId="168.[189,1398,372,514]" pageId="168" pageNumber="169">
<emphasis id="B96F064FFF0DFF51FEE5FE24AC6CEBCD" box="[316,331,411,436]" italics="true" pageId="168" pageNumber="169">–</emphasis>
<emphasis id="B96F064FFF0DFF51FE89FE24ACA9EBCC" bold="true" box="[336,398,411,437]" pageId="168" pageNumber="169">
<typeStatus id="54A064FFFF0DFF51FE89FE24ACA9EBCC" box="[336,398,411,437]" pageId="168" pageNumber="169">Type</typeStatus>
</emphasis>
:
<materialsCitation id="3B73D000FF0DFF51FE45FE24A851EBA2" collectingDate="1923-05-20" collectingDateMax="1923-06-01" collectingDateMin="1923-05-20" collectionCode="F, K" collectorName="J. E. Macbride" country="Peru" county="Prov. Huanuco" elevation="1980" latitude="-9.666667" location="Estacion near Muna" longLatPrecision="1299" longitude="-75.816666" municipality="Estacion" pageId="168" pageNumber="169" specimenCode="V0042326F" specimenCount="1" stateProvince="Huanuco Region" typeStatus="holotype">
<collectingCountry id="F30C9ACDFF0DFF51FE45FE24ACC0EBCC" box="[412,487,411,437]" name="Peru" pageId="168" pageNumber="169">PERU</collectingCountry>
–
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[
<collectingRegion id="49DF14BFFF0DFF51FDDFFE26AFF8EBCD" box="[518,735,409,436]" country="Peru" name="Huanuco" pageId="168" pageNumber="169">Huánuco Region</collectingRegion>
:
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]
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•
<location id="8EC48C86FF0DFF51FC1EFE25A996EBCC" LSID="urn:lsid:plazi:treatment:03B26B4BFF0DFF50FE3EFEB6AE2BEFF8:8EC48C86FF0DFF51FC1EFE25A996EBCC" box="[967,1201,410,437]" country="Peru" county="Prov. Huanuco" latitude="-9.666667" longLatPrecision="1299" longitude="-75.816666" municipality="Estacion" name="Estacion near Muna" pageId="168" pageNumber="169" stateProvince="Huanuco Region">Estacion near Muña</location>
; ca
<elevation id="00363D6EFF0DFF51FB02FE24A857EBCD" box="[1243,1392,410,437]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="168" pageNumber="169" unit="m" value="1980.0">
<quantity id="4CE377B8FF0DFF51FB02FE24A815EBCC" box="[1243,1330,411,437]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="168" pageNumber="169" unit="m" value="1980.0">1980 m</quantity>
a.s.l.
</elevation>
; [
<geoCoordinate id="EE2FBC9AFF0DFF51FF29FE00AC6CEBA2" box="[240,331,447,475]" degrees="9" direction="south" minutes="40" orientation="latitude" pageId="168" pageNumber="169" precision="925" value="-9.666667">9°40′ S</geoCoordinate>
,
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];
<date id="FFA5FC9DFF0DFF51FE06FE7EAFF5EBA2" box="[479,722,449,475]" pageId="168" pageNumber="169" value="1923-05-20" valueMax="1923-06-01" valueMin="1923-05-20">
<collectingDate id="EFE10575FF0DFF51FE06FE7EAFF5EBA2" box="[479,722,449,475]" pageId="168" pageNumber="169" value="1923-05-20" valueMax="1923-06-01" valueMin="1923-05-20">20 May–1 Jun. 1923</collectingDate>
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;
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<collectorName id="26EEBF8BFF0DFF51FD07FE7DAEA1EBA2" box="[734,902,449,475]" pageId="168" pageNumber="169">J.E. Macbride</collectorName>
4143
</emphasis>
;
<typeStatus id="54A064FFFF0DFF51FC01FE7EA964EBA2" box="[984,1091,449,475]" pageId="168" pageNumber="169">holotype</typeStatus>
:
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[
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, photo
<collectionCode id="ED0A4298FF0DFF51FA89FE7EA84DEBA2" box="[1360,1386,449,475]" pageId="168" pageNumber="169">K</collectionCode>
].
</materialsCitation>
</paragraph>
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<subSubSection id="C30189D6FF0DFF51FF64FE58A9B3E878" box="[189,1172,487,514]" pageId="168" pageNumber="169" type="reference_group">
<paragraph id="8BA4DA5DFF0DFF51FF64FE58A9B3E878" blockId="168.[189,1398,372,514]" box="[189,1172,487,514]" pageId="168" pageNumber="169">
<bibRefCitation id="EF8AA7ACFF0DFF51FF64FE58AF00E878" author="Brako &amp; Zarucchi" box="[189,551,487,513]" firstAuthor="Brako" page="194" pageId="168" pageNumber="169" refId="ref225797" refString="Brako L. &amp; Zarucchi L. 1993. Catalogue of the Flowering Plants and Gymnosperms of Peru. Monographs in Systematic Botany from the Missouri Botanical Garden 45, Missouri Botanical Garden, St Louis." type="book" year="1993">Brako &amp; Zarucchi (1993: 194)</bibRefCitation>
;
<bibRefCitation id="EF8AA7ACFF0DFF51FDEAFE58AEBBE878" author="Leon &amp; Monsalve" box="[563,924,487,513]" firstAuthor="Leon" page="167" pageId="168" pageNumber="169" pagination="165 - 170" refId="ref229226" refString="Leon B. &amp; Monsalve C. 2006. Begoniaceae endemicas del Peru. Revista Peruana de Biologia 13 (2): 165 - 170. https: // doi. org / 10.15381 / rpb. v 13 i 2.1810" type="journal article" year="2006">
<collectingRegion id="49DF14BFFF0DFF51FDEAFE58AF48E878" box="[563,623,487,513]" country="Nicaragua" name="Leon" pageId="168" pageNumber="169">León</collectingRegion>
&amp; Monsalve (2006: 167)
</bibRefCitation>
;
<bibRefCitation id="EF8AA7ACFF0DFF51FC71FE58A9B7E878" author="Tebbitt M. C." box="[936,1168,487,514]" journalOrPublisher="Novon" pageId="168" pageNumber="169" pagination="479 - 489" part="23" refId="ref232441" refString="Tebbitt M. C. 2015. Two new species of Andean tuberous Begonia in the B. octopetala group (Begoniaceae). Novon 23 (4): 479 - 489. https: // doi. org / 10.3417 / 2013027" title="Two new species of Andean tuberous Begonia in the B. octopetala group (Begoniaceae)" type="journal article" year="2015">Tebbitt (2015: 484)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF0DFF51FF64FD93AFC7E8E7" pageId="168" pageNumber="169" type="etymology">
<paragraph id="8BA4DA5DFF0DFF51FF64FD93AC62E83F" blockId="168.[189,325,556,582]" box="[189,325,556,582]" pageId="168" pageNumber="169">
<heading id="D0EC6D31FF0DFF51FF64FD93AC62E83F" bold="true" box="[189,325,556,582]" fontSize="11" level="3" pageId="168" pageNumber="169" reason="3">
<emphasis id="B96F064FFF0DFF51FF64FD93AC62E83F" bold="true" box="[189,325,556,582]" pageId="168" pageNumber="169">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF0DFF51FF64FDE1AFC7E8E7" blockId="168.[189,1399,605,671]" pageId="168" pageNumber="169">
<bibRefCitation id="EF8AA7ACFF0DFF51FF64FDE1AC5CE801" author="Irmscher" box="[189,379,605,632]" firstAuthor="Irmscher" pageId="168" pageNumber="169" pagination="1 - 102" refId="ref228521" refString="Irmscher E. 1953. Systematische Studien uber Begoniaceen des tropischen Sudamerikas, besonders Brasiliens. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 76 (1): 1 - 102." type="journal article" year="1953">Irmscher (1953)</bibRefCitation>
distinguished this
<taxonomicName id="4C1BA1DEFF0DFF51FD8FFDE1AFC0E801" box="[598,743,606,632]" pageId="168" pageNumber="169" rank="variety" variety="from">variety from</taxonomicName>
other members of the species primarily by its ovate leaf blades. The epithet refers to this characteristic.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF0DFF51FF64FD77AFD7E945" pageId="168" pageNumber="169" type="materials_examined">
<paragraph id="8BA4DA5DFF0DFF51FF64FD77AC9FE89B" blockId="168.[189,440,712,738]" box="[189,440,712,738]" pageId="168" pageNumber="169">
<heading id="D0EC6D31FF0DFF51FF64FD77AC9FE89B" bold="true" box="[189,440,712,738]" fontSize="11" level="3" pageId="168" pageNumber="169" reason="3">
<emphasis id="B96F064FFF0DFF51FF64FD77AC9FE89B" bold="true" box="[189,440,712,738]" pageId="168" pageNumber="169">Specimen examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF0DFF51FF64FD44AFD7E945" blockId="168.[189,1398,761,828]" pageId="168" pageNumber="169">
<materialsCitation id="3B73D000FF0DFF51FF64FD44AFCAE945" collectingDate="1863-05" collectionCode="K" country="Peru" county="Prov. Huanuco" elevation="2898" latitude="-9.666667" location="Clefts of rocks near Muna" longLatPrecision="1299" longitude="-75.816666" municipality="Clefts" pageId="168" pageNumber="169" specimenCount="1" stateProvince="Huanuco Region">
<collectingCountry id="F30C9ACDFF0DFF51FF64FD44AC2FE96C" box="[189,264,763,789]" name="Peru" pageId="168" pageNumber="169">PERU</collectingCountry>
–
<emphasis id="B96F064FFF0DFF51FEF0FD45AFE8E96C" bold="true" box="[297,719,762,789]" pageId="168" pageNumber="169">
<collectingRegion id="49DF14BFFF0DFF51FEF0FD45AF25E96D" box="[297,514,762,789]" country="Peru" name="Huanuco" pageId="168" pageNumber="169">Huánuco Region</collectingRegion>
:
<collectingCounty id="62C5A2D1FF0DFF51FDC8FD44AFE8E96C" box="[529,719,762,789]" pageId="168" pageNumber="169">Prov. Huánuco</collectingCounty>
</emphasis>
•
<location id="8EC48C86FF0DFF51FD33FD45A902E96C" LSID="urn:lsid:plazi:treatment:03B26B4BFF0DFF50FE3EFEB6AE2BEFF8:8EC48C86FF0DFF51FD33FD45A902E96C" box="[746,1061,762,789]" country="Peru" county="Prov. Huanuco" latitude="-9.666667" longLatPrecision="1299" longitude="-75.816666" municipality="Clefts" name="Clefts of rocks near Muna" pageId="168" pageNumber="169" stateProvince="Huanuco Region">Clefts of rocks near Muña</location>
; [
<geoCoordinate id="EE2FBC9AFF0DFF51FBE3FD46A9B3E96C" box="[1082,1172,761,789]" degrees="9" direction="south" minutes="40" orientation="latitude" pageId="168" pageNumber="169" precision="925" value="-9.666667">9°40′ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFF0DFF51FB7BFD46A83FE96C" box="[1186,1304,761,789]" degrees="75" direction="west" minutes="49" orientation="longitude" pageId="168" pageNumber="169" precision="925" value="-75.816666">75°49′ W</geoCoordinate>
];
<elevation id="00363D6EFF0DFF51FAF4FD45AC71E942" metricMagnitude="3" metricUnit="m" metricValue="2.8975" metricValueMax="3.05" metricValueMin="2.745" pageId="168" pageNumber="169" unit="m" value="2897.5" valueMax="3050.0" valueMin="2745.0">
<quantity id="4CE377B8FF0DFF51FAF4FD45AC31E945" metricMagnitude="3" metricUnit="m" metricValue="2.8975" metricValueMax="3.05" metricValueMin="2.745" pageId="168" pageNumber="169" unit="m" value="2897.5" valueMax="3050.0" valueMin="2745.0">2745– 3050 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFF0DFF51FEBAFC9DACFAE942" box="[355,477,801,828]" pageId="168" pageNumber="169" value="1863-05">
<collectingDate id="EFE10575FF0DFF51FEBAFC9DACFAE942" box="[355,477,801,828]" pageId="168" pageNumber="169" value="1863-05">May 1863</collectingDate>
</date>
,
<emphasis id="B96F064FFF0DFF51FE33FC9EAFE3E942" box="[490,708,801,827]" italics="true" pageId="168" pageNumber="169">coll. unknown 185</emphasis>
;
<collectionCode id="ED0A4298FF0DFF51FD0DFC9DAFCAE945" box="[724,749,802,828]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="168" pageNumber="169" type="Herbarium">K</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF0DFF51FF64FCDAAF46ED5C" pageId="168" pageNumber="169" type="description">
<paragraph id="8BA4DA5DFF0DFF51FF64FCDAAC68E906" blockId="168.[189,335,869,895]" box="[189,335,869,895]" pageId="168" pageNumber="169">
<heading id="D0EC6D31FF0DFF51FF64FCDAAC68E906" bold="true" box="[189,335,869,895]" fontSize="11" level="3" pageId="168" pageNumber="169" reason="3">
<emphasis id="B96F064FFF0DFF51FF64FCDAAC68E906" bold="true" box="[189,335,869,895]" pageId="168" pageNumber="169">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF0DFF51FF64FC28AF46ED5C" blockId="168.[189,1401,919,1830]" pageId="168" pageNumber="169">
Acaulescent, rhizomatous herb, to
<quantity id="4CE377B8FF0DFF51FD8CFC28AFB9E9CB" box="[597,670,919,946]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" pageId="168" pageNumber="169" unit="cm" value="45.0">45 cm</quantity>
high.
<emphasis id="B96F064FFF0DFF51FD31FC28AE69E9C8" box="[744,846,919,945]" italics="true" pageId="168" pageNumber="169">Rhizome</emphasis>
ellipsoid, ca 3.5 ×
<quantity id="4CE377B8FF0DFF51FBF6FC27A94BE9CB" box="[1071,1132,920,946]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="168" pageNumber="169" unit="cm" value="2.0">2 cm</quantity>
, with 1 growing point.
<emphasis id="B96F064FFF0DFF51FF64FC01AC3CE9A1" box="[189,283,958,984]" italics="true" pageId="168" pageNumber="169">Stipules</emphasis>
persistent, triangular, ca 9 ×
<quantity id="4CE377B8FF0DFF51FD57FC01AFFDE9A1" box="[654,730,958,984]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="168" pageNumber="169" unit="mm" value="9.0">9 mm</quantity>
, apex obtuse, opaque, brown, glabrous, margin entire, aciliate to sparsely ciliate.
<emphasis id="B96F064FFF0DFF51FDD9FC5AAF75E987" box="[512,594,997,1022]" italics="true" pageId="168" pageNumber="169">Leaves</emphasis>
1–3, alternate, basifixed; petiole
<quantity id="4CE377B8FF0DFF51FC3FFC5BA97AE986" box="[998,1117,996,1023]" metricMagnitude="-1" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="168" pageNumber="169" unit="cm" value="15.5" valueMax="17.0" valueMin="14.0">14–17 cm</quantity>
long, colour unknown, lanate; blade subsymmetric, triangular in outline, to 21 ×
<quantity id="4CE377B8FF0DFF51FC82FBB5AE9BEE5C" box="[859,956,1034,1061]" metricMagnitude="-1" metricUnit="m" metricValue="1.55" pageId="168" pageNumber="169" unit="cm" value="15.5">15.5 cm</quantity>
, succulent, apex obtuse, base cordate, basal lobes not overlapping or overlapping, sinus to
<quantity id="4CE377B8FF0DFF51FCFFFB8EAE5EEE35" box="[806,889,1073,1100]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="168" pageNumber="169" unit="mm" value="50.0">50 mm</quantity>
deep, margin with 0–5 short, crenate lobes on both sides of the lamina, dentate, ciliate, upper surface green, pilose, lower surface pale green, glabrous to sparsely pilose, veins palmate but with 1 primary vein, 5–6 veined from the base, with 1–3 secondary veins on the larger side, 1–3 on the smaller side.
<emphasis id="B96F064FFF0DFF51FCACFB1BA93AEEC7" box="[885,1053,1188,1214]" italics="true" pageId="168" pageNumber="169">Inflorescences</emphasis>
1–3, bisexual, axillary, erect, a dichasial or monochasial cyme, with 1–3 branches, bearing up to 3 staminate flowers and 2 pistillate flowers, protandrous; peduncle to
<quantity id="4CE377B8FF0DFF51FD8CFB4EAFB8EF75" box="[597,671,1265,1292]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" pageId="168" pageNumber="169" unit="cm" value="45.0">45 cm</quantity>
long, red, sparsely to densely lanate, bracts persistent, ovate to lanceolate, 5–11 ×
<quantity id="4CE377B8FF0DFF51FE18FAA7AF0FEF4B" box="[449,552,1304,1330]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="168" pageNumber="169" unit="mm" value="5.0" valueMax="8.0" valueMin="2.0">2–8 mm</quantity>
, opaque, white, green, or red, glabrous to lanate, apex acute to obtuse, margin entire, aciliate.
<emphasis id="B96F064FFF0DFF51FE0DFA81AF80EF21" box="[468,679,1342,1368]" italics="true" pageId="168" pageNumber="169">Staminate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FF0DFF51FC9AFA81AEA7EF20" box="[835,896,1342,1369]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="168" pageNumber="169" unit="cm" value="3.0">3 cm</quantity>
long, sparsely to densely lanate; tepals 6, spreading, oblanceolate to obovate, 17–22 ×
<quantity id="4CE377B8FF0DFF51FD0DFADAAE6FEF06" box="[724,840,1381,1407]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.7" metricValueMin="0.7" pageId="168" pageNumber="169" unit="mm" value="12.0" valueMax="17.0" valueMin="7.0">7–17 mm</quantity>
, apex rounded, white to pink, glabrous, margin entire, aciliate; stamens 50–75, spreading, yellow, filaments
<quantity id="4CE377B8FF0DFF51FC59FA34AEC7EFDC" box="[896,992,1419,1445]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="168" pageNumber="169" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long, free, anthers obovoid, ca 1 ×
<quantity id="4CE377B8FF0DFF51FF64FA0EAC3FEFB5" box="[189,280,1457,1484]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="168" pageNumber="169" unit="mm" value="0.5">0.5 mm</quantity>
, dehiscing via lateral slits, connectives extended to ca
<quantity id="4CE377B8FF0DFF51FC5DFA0EAEF8EFB5" box="[900,991,1457,1484]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="168" pageNumber="169" unit="mm" value="0.2">0.2 mm</quantity>
, symmetrically basifixed.
<emphasis id="B96F064FFF0DFF51FAD2FA0EAC35EF8B" italics="true" pageId="168" pageNumber="169">Pistillate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FF0DFF51FE70FA67ACC3EF8B" box="[425,484,1496,1522]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="168" pageNumber="169" unit="cm" value="9.0">9 cm</quantity>
long; bracteoles unknown or lacking; tepals ca 7, subequal, late deciduous in fruit, spreading, obovate, 7–17 ×
<quantity id="4CE377B8FF0DFF51FD89FA41AFE2EC60" box="[592,709,1534,1561]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="168" pageNumber="169" unit="mm" value="10.0" valueMax="13.0" valueMin="7.0">7–13 mm</quantity>
, apex rounded to truncate, white to pink, glabrous on the inner surface, sparsely to densely lanate on the outer surface, margin entire, aciliate; ovary body ovoid to broadly obovoid, 6–13 ×
<quantity id="4CE377B8FF0DFF51FDC8F9F4AFA1EC1C" box="[529,646,1611,1638]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="13.0" metricValueMin="5.0" pageId="168" pageNumber="169" unit="mm" value="9.0" valueMax="13.0" valueMin="5.0">5–13 mm</quantity>
, white to pink, densely lanate, unequally 3-winged, the largest rectangular, ascending, 7–13 ×
<quantity id="4CE377B8FF0DFF51FDF6F9CEAF85ECF5" box="[559,674,1649,1676]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="2.5" metricValueMin="0.7" pageId="168" pageNumber="169" unit="mm" value="16.0" valueMax="25.0" valueMin="7.0">7–25 mm</quantity>
, the smallest a semi-circular ridge,
<quantity id="4CE377B8FF0DFF51FB9BF9CEA99EECF5" box="[1090,1209,1649,1676]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="168" pageNumber="169" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
wide; 3-locular, placentae branches divided, bearing ovules on both surfaces; styles 3, yellow, free,
<quantity id="4CE377B8FF0DFF51FB46F927A825ECCB" box="[1183,1282,1688,1714]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="168" pageNumber="169" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long,&gt; 5 times-divided, stigmatic papillae in a spirally twisted band.
<emphasis id="B96F064FFF0DFF51FC5FF900A96FECA1" box="[902,1096,1726,1752]" italics="true" pageId="168" pageNumber="169">Fruiting pedicel</emphasis>
to
<quantity id="4CE377B8FF0DFF51FBB6F901A98DECA0" box="[1135,1194,1726,1753]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="168" pageNumber="169" unit="cm" value="9.0">9 cm</quantity>
long.
<emphasis id="B96F064FFF0DFF51FB21F900A850ECA1" box="[1272,1399,1726,1752]" italics="true" pageId="168" pageNumber="169">Fruit body</emphasis>
obovoid, to 13 ×
<quantity id="4CE377B8FF0DFF51FE57F95AACC1EC86" box="[398,486,1765,1791]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="168" pageNumber="169" unit="mm" value="13.0">13 mm</quantity>
, drying brown, wings same shape as in ovary, the largest expanding to 13 ×
<quantity id="4CE377B8FF0DFF51FF66F8B4AC32ED5F" box="[191,277,1803,1830]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="168" pageNumber="169" unit="mm" value="50.0">50 mm</quantity>
, the smallest not expanding.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF0DFF51FF64F8EFA92CED91" pageId="168" pageNumber="169" type="conservation">
<paragraph id="8BA4DA5DFF0DFF51FF64F8EFAF4AED13" blockId="168.[189,621,1871,1898]" box="[189,621,1871,1898]" pageId="168" pageNumber="169">
<heading id="D0EC6D31FF0DFF51FF64F8EFAF4AED13" bold="true" box="[189,621,1871,1898]" fontSize="11" level="3" pageId="168" pageNumber="169" reason="3">
<emphasis id="B96F064FFF0DFF51FF64F8EFAF4AED13" bold="true" box="[189,621,1871,1898]" pageId="168" pageNumber="169">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF0DFF51FF64F83EA92CED91" blockId="168.[189,1398,1921,2025]" pageId="168" pageNumber="169">
Previously assessed by
<bibRefCitation id="EF8AA7ACFF0DFF51FE10F83DAFCBEDE2" author="Leon &amp; Monsalve" box="[457,748,1921,1948]" firstAuthor="Leon" pageId="168" pageNumber="169" pagination="165 - 170" refId="ref229226" refString="Leon B. &amp; Monsalve C. 2006. Begoniaceae endemicas del Peru. Revista Peruana de Biologia 13 (2): 165 - 170. https: // doi. org / 10.15381 / rpb. v 13 i 2.1810" type="journal article" year="2006">
<collectingRegion id="49DF14BFFF0DFF51FE10F83DAF22EDE5" box="[457,517,1922,1948]" country="Nicaragua" name="Leon" pageId="168" pageNumber="169">León</collectingRegion>
&amp; Monsalve (2006)
</bibRefCitation>
as Data Deficient (DD). Known from a single population but not collected for &lt;100 years, though this area is poorly collected. We have been unable to visit this locality so assess
<taxonomicName id="4C1BA1DEFF0DFF51FE54F870AFCBED91" box="[397,748,1998,2025]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="168" pageNumber="169" phylum="Tracheophyta" rank="variety" species="octopetala" variety="ovatiformis">
<emphasis id="B96F064FFF0DFF51FE54F870AF0CED91" box="[397,555,1998,2024]" italics="true" pageId="168" pageNumber="169">B. octopetala</emphasis>
var.
<emphasis id="B96F064FFF0DFF51FDBCF871AFCBED91" box="[613,748,1998,2024]" italics="true" pageId="168" pageNumber="169">ovatiformis</emphasis>
</taxonomicName>
as Data Deficient (DD).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF0CFF50FF64FEB6A842EEB0" pageId="169" pageNumber="170" type="discussion">
<paragraph id="8BA4DA5DFF0CFF50FF64FEB6AC23EB5A" blockId="169.[189,260,265,291]" box="[189,260,265,291]" pageId="169" pageNumber="170">
<heading id="D0EC6D31FF0CFF50FF64FEB6AC23EB5A" bold="true" box="[189,260,265,291]" fontSize="11" level="3" pageId="169" pageNumber="170" reason="3">
<emphasis id="B96F064FFF0CFF50FF64FEB6AC23EB5A" bold="true" box="[189,260,265,291]" pageId="169" pageNumber="170">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF0CFF50FF64FE88ACDFE91F" blockId="169.[188,1399,311,870]" pageId="169" pageNumber="170">
The taxonomic status of this subspecies is unclear. It is known from
<specimenCount id="9D1D11D4FF0CFF50FC3FFE87A9B0EB28" box="[998,1175,311,338]" pageId="169" pageNumber="170" type="generic">two specimens</specimenCount>
, which share some characteristics with
<taxonomicName id="4C1BA1DEFF0CFF50FE69FEE3AF74EB0C" baseAuthorityName="Knowles &amp; Wescott" baseAuthorityYear="1837" box="[432,595,347,373]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="species" species="octopetala">
<emphasis id="B96F064FFF0CFF50FE69FEE3AF74EB0C" box="[432,595,347,373]" italics="true" pageId="169" pageNumber="170">B. octopetala</emphasis>
</taxonomicName>
and others with
<taxonomicName id="4C1BA1DEFF0CFF50FCF1FEE3A976EB0C" authority="Irmsch." authorityName="Irmsch." box="[808,1105,347,373]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="species" species="aequatorialis">
<emphasis id="B96F064FFF0CFF50FCF1FEE3AECCEB0C" box="[808,1003,347,373]" italics="true" pageId="169" pageNumber="170">B. aequatorialis</emphasis>
Irmsch.
</taxonomicName>
The
<typeStatus id="54A064FFFF0CFF50FB4CFEE4A9F4EB0C" box="[1173,1235,347,373]" pageId="169" pageNumber="170">types</typeStatus>
of these two species differ substantially: the
<typeStatus id="54A064FFFF0CFF50FDE9FEC0AF44EBE0" box="[560,611,383,409]" pageId="169" pageNumber="170">type</typeStatus>
of
<taxonomicName id="4C1BA1DEFF0CFF50FD53FEC0AE0FEBE1" baseAuthorityName="Knowles &amp; Wescott" baseAuthorityYear="1837" box="[650,808,382,408]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="species" species="octopetala">
<emphasis id="B96F064FFF0CFF50FD53FEC0AE0FEBE1" box="[650,808,382,408]" italics="true" pageId="169" pageNumber="170">B. octopetala</emphasis>
</taxonomicName>
has broadly ovate leaves that lack a distinct apex; subsymmetric inflorescences; staminate flowers with&gt; 6 tepals;&gt;
<quantity id="4CE377B8FF0CFF50FC10FE1DA93EEBC5" box="[969,1049,418,445]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="169" pageNumber="170" unit="cm" value="1.5">1.5 cm</quantity>
broad tepals on the staminate flowers; and ascending largest fruit wings. In contrast, the
<typeStatus id="54A064FFFF0CFF50FCA0FE79AE8BEB99" box="[889,940,454,480]" pageId="169" pageNumber="170">type</typeStatus>
of
<taxonomicName id="4C1BA1DEFF0CFF50FC0FFE79A9B0EBA6" box="[982,1175,453,479]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="species" species="aequatorialis">
<emphasis id="B96F064FFF0CFF50FC0FFE79A9B0EBA6" box="[982,1175,453,479]" italics="true" pageId="169" pageNumber="170">B. aequatorialis</emphasis>
</taxonomicName>
has much smaller, deltoid leaves with a distinct apex; asymmetric inflorescences; staminate flowers with 6 tepals; &lt;
<quantity id="4CE377B8FF0CFF50FAFEFE56A850E87A" box="[1319,1399,489,516]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="169" pageNumber="170" unit="cm" value="1.5">1.5 cm</quantity>
broad tepals on the staminate flowers; and non-ascending largest fruit wings. Irmscher’s concept of
<taxonomicName id="4C1BA1DEFF0CFF50FF64FD8EAF61E833" box="[189,582,560,586]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="subSpecies" species="octopetala" subSpecies="ovatiformis">
<emphasis id="B96F064FFF0CFF50FF64FD8EAC78E833" box="[189,351,560,586]" italics="true" pageId="169" pageNumber="170">B. octopetala</emphasis>
subsp.
<emphasis id="B96F064FFF0CFF50FE66FD8FAF61E833" box="[447,582,560,586]" italics="true" pageId="169" pageNumber="170">ovatiformis</emphasis>
</taxonomicName>
is intermediate between these two species. It differs from the
<typeStatus id="54A064FFFF0CFF50FA9DFD8FA850E833" box="[1348,1399,560,586]" pageId="169" pageNumber="170">type</typeStatus>
of
<taxonomicName id="4C1BA1DEFF0CFF50FF06FDEBAC58E814" baseAuthorityName="Knowles &amp; Wescott" baseAuthorityYear="1837" box="[223,383,595,621]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="species" species="octopetala">
<emphasis id="B96F064FFF0CFF50FF06FDEBAC58E814" box="[223,383,595,621]" italics="true" pageId="169" pageNumber="170">B. octopetala</emphasis>
</taxonomicName>
in having deltoid leaf laminae with a clear apex; six tepals on the staminate flower; and ascending largest fruit wings but differs from the
<typeStatus id="54A064FFFF0CFF50FCE1FDC8AE4CE8E8" box="[824,875,631,657]" pageId="169" pageNumber="170">type</typeStatus>
of
<taxonomicName id="4C1BA1DEFF0CFF50FC4AFDC8A974E8E8" box="[915,1107,631,657]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="species" species="aequatorialis">
<emphasis id="B96F064FFF0CFF50FC4AFDC8A974E8E8" box="[915,1107,631,657]" italics="true" pageId="169" pageNumber="170">B. aequatorialis</emphasis>
</taxonomicName>
in having subsymmetric inflorescences and broader tepals on the staminate flowers. The boundaries between the three taxa have become blurred due to the recent collection of several collections of
<taxonomicName id="4C1BA1DEFF0CFF50FBD9FD01A986E8A1" baseAuthorityName="Knowles &amp; Wescott" baseAuthorityYear="1837" box="[1024,1185,702,728]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="species" species="octopetala">
<emphasis id="B96F064FFF0CFF50FBD9FD01A986E8A1" box="[1024,1185,702,728]" italics="true" pageId="169" pageNumber="170">B. octopetala</emphasis>
</taxonomicName>
from the Andean cordillera. These specimens tend to have smaller, more triangular shaped leaves (e.g.,
<emphasis id="B96F064FFF0CFF50FB7AFD5DAC75E967" italics="true" pageId="169" pageNumber="170">P.W. Moonlight &amp; A. Daza 106</emphasis>
) and narrower tepals on the staminate flower (e.g.,
<emphasis id="B96F064FFF0CFF50FC65FCBAA816E966" box="[956,1329,773,799]" italics="true" pageId="169" pageNumber="170">P.W. Moonlight &amp; A. Daza 229</emphasis>
) than the
<typeStatus id="54A064FFFF0CFF50FF33FC96AC3AE93A" box="[234,285,809,835]" pageId="169" pageNumber="170">type</typeStatus>
of
<taxonomicName id="4C1BA1DEFF0CFF50FE9CFC96ACC3E93B" baseAuthorityName="Knowles &amp; Wescott" baseAuthorityYear="1837" box="[325,484,808,834]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="species" species="octopetala">
<emphasis id="B96F064FFF0CFF50FE9CFC96ACC3E93B" box="[325,484,808,834]" italics="true" pageId="169" pageNumber="170">B. octopetala</emphasis>
</taxonomicName>
but remain distinct from the
<typeStatus id="54A064FFFF0CFF50FCE7FC96AE56E93A" box="[830,881,809,835]" pageId="169" pageNumber="170">type</typeStatus>
of
<taxonomicName id="4C1BA1DEFF0CFF50FC43FC96A83EE93B" box="[922,1305,808,834]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="subSpecies" species="octopetala" subSpecies="ovatiformis">
<emphasis id="B96F064FFF0CFF50FC43FC96A91FE93B" box="[922,1080,808,834]" italics="true" pageId="169" pageNumber="170">B. octopetala</emphasis>
subsp.
<emphasis id="B96F064FFF0CFF50FB4BFC97A83EE93B" box="[1170,1305,808,834]" italics="true" pageId="169" pageNumber="170">ovatiformis</emphasis>
</taxonomicName>
in their non-ascending fruit wings.
</paragraph>
<paragraph id="8BA4DA5DFF0CFF50FF64FC32AFCAEE6B" blockId="169.[189,1401,909,1042]" pageId="169" pageNumber="170">
Mark Tebbitt is currently undertaking a full taxonomic revision of
<emphasis id="B96F064FFF0CFF50FC13FC31AEC3E9DE" box="[970,996,910,935]" italics="true" pageId="169" pageNumber="170">B.</emphasis>
sect.
<taxonomicName id="4C1BA1DEFF0CFF50FBFEFC32A980E9DE" box="[1063,1191,909,935]" class="Magnoliopsida" family="Begoniaceae" genus="Eupetalum" kingdom="Plantae" order="Cucurbitales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F064FFF0CFF50FBFEFC32A980E9DE" box="[1063,1191,909,935]" italics="true" pageId="169" pageNumber="170">Eupetalum</emphasis>
</taxonomicName>
(Tebbitt in prep.) following his recent revision of the closely related
<emphasis id="B96F064FFF0CFF50FCF1FC0DAE66E9B2" box="[808,833,946,971]" italics="true" pageId="169" pageNumber="170">B.</emphasis>
sect.
<taxonomicName id="4C1BA1DEFF0CFF50FC52FC0EAEDCE9B2" box="[907,1019,945,971]" pageId="169" pageNumber="323" rank="section" section="Australes">
<emphasis id="B96F064FFF0CFF50FC52FC0EAEDCE9B2" box="[907,1019,945,971]" italics="true" pageId="169" pageNumber="170">Australes</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8AA7ACFF0CFF50FBD4FC0EA98BE9B2" author="Tebbitt M. C." box="[1037,1196,945,971]" journalOrPublisher="American Begonia Society, Sacramento" pageId="169" pageNumber="170" refId="ref232580" refString="Tebbitt M. C. 2020. Tuberous Begonias: a Monograph of Begonia Section Australes. American Begonia Society, Sacramento." title="Tuberous Begonias: a Monograph of Begonia Section Australes" type="book" year="2020">Tebbitt 2020</bibRefCitation>
). We suspect the boundaries of these species may change as a result of this revision but maintain
<taxonomicName id="4C1BA1DEFF0CFF50FB5CFC6AAC63EE6B" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="subSpecies" species="octopetala" subSpecies="ovatiformis">
<emphasis id="B96F064FFF0CFF50FB5CFC6AA803E997" box="[1157,1316,980,1006]" italics="true" pageId="169" pageNumber="170">B. octopetala</emphasis>
subsp.
<emphasis id="B96F064FFF0CFF50FF64FC47AC63EE6B" box="[189,324,1016,1042]" italics="true" pageId="169" pageNumber="170">ovatiformis</emphasis>
</taxonomicName>
as a distinct taxon in this treatment.
</paragraph>
<paragraph id="8BA4DA5DFF0CFF50FF64FB86AC97EE2D" blockId="169.[189,432,1081,1108]" box="[189,432,1081,1108]" pageId="169" pageNumber="170">
<heading id="D0EC6D31FF0CFF50FF64FB86AC97EE2D" bold="true" box="[189,432,1081,1108]" fontSize="11" level="3" pageId="169" pageNumber="170" reason="3">
<emphasis id="B96F064FFF0CFF50FF64FB86AC97EE2D" bold="true" box="[189,432,1081,1108]" pageId="169" pageNumber="170">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF0CFF50FF64FBD7A842EEB0" blockId="169.[189,1398,1128,1226]" pageId="169" pageNumber="170">
Distinct from most specimens of
<taxonomicName id="4C1BA1DEFF0CFF50FD8EFBD6AFDEEEFB" baseAuthorityName="Knowles &amp; Wescott" baseAuthorityYear="1837" box="[599,761,1128,1154]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="species" species="octopetala">
<emphasis id="B96F064FFF0CFF50FD8EFBD6AFDEEEFB" box="[599,761,1128,1154]" italics="true" pageId="169" pageNumber="170">B. octopetala</emphasis>
</taxonomicName>
in its leaf laminae, which are deltoid in outline (vs circular to rarely deltoid), and its staminate flowers, which have 6 (vs 6–11 tepals). The only character that can distinguish this subspecies reliably is its ascending (vs non-ascending) fruit and ovary wings.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF0CFF50FF64FB4EAE2BEFF8" pageId="169" pageNumber="170" type="distribution">
<paragraph id="8BA4DA5DFF0CFF50FF64FB4EACD2EF72" blockId="169.[189,501,1265,1291]" box="[189,501,1265,1291]" pageId="169" pageNumber="170">
<heading id="D0EC6D31FF0CFF50FF64FB4EACD2EF72" bold="true" box="[189,501,1265,1291]" fontSize="11" level="3" pageId="169" pageNumber="170" reason="3">
<emphasis id="B96F064FFF0CFF50FF64FB4EACD2EF72" bold="true" box="[189,501,1265,1291]" pageId="169" pageNumber="170">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF0CFF50FF64FA9FAE2BEFF8" blockId="169.[189,1399,1312,1409]" pageId="169" pageNumber="170">
Endemic to
<collectingCountry id="F30C9ACDFF0CFF50FE8BFA9FACAFEF43" box="[338,392,1312,1338]" name="Peru" pageId="169" pageNumber="170">Peru</collectingCountry>
and
<collectingRegion id="49DF14BFFF0CFF50FE10FA9FAFB4EF43" box="[457,659,1312,1338]" country="Peru" name="Huanuco" pageId="169" pageNumber="170">Huánuco Region</collectingRegion>
(
<figureCitation id="1320C6D8FF0CFF50FD7EFA9FAE31EF42" box="[679,790,1312,1339]" captionStart="Fig" captionStartId="160.[189,232,1822,1848]" captionTargetBox="[194,1391,558,1776]" captionTargetId="figure-14@160.[189,1398,548,1784]" captionTargetPageId="160" captionText="Fig. 52. Distribution of the octopetala group of Begonia sect. Eupetalum (Lindl.) A.DC. in Peru and surrounding countries. A. B. anemoniflora Irmsch. (red) and B. tumbezensis Irmsch. (blue). B. B. octopetala L.Hér. subsp. octopetala (red) and B. octopetala subsp. ovatiformis Irmsch. (blue). C. B. pleiopetala A.DC. (red) and B. pseudopleiopetala Tebbitt (blue). D. B. polypetala A.DC. (red). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178413" httpUri="https://zenodo.org/record/8178413/files/figure.png" pageId="169" pageNumber="170">Fig. 52B</figureCitation>
). Known from
<quantity id="4CE377B8FF0CFF50FC0DFA9FA95DEF43" box="[980,1146,1312,1339]" metricMagnitude="3" metricUnit="m" metricValue="2.5149999999999997" metricValueMax="3.05" metricValueMin="1.98" pageId="169" pageNumber="170" unit="m" value="2515.0" valueMax="3050.0" valueMin="1980.0">1980–3050 m</quantity>
a.s.l. in elevation in middle and upper montane forest. Like the
<typeStatus id="54A064FFFF0CFF50FD18FAFBAFD3EF27" box="[705,756,1348,1374]" pageId="169" pageNumber="170">type</typeStatus>
subspecies,
<taxonomicName id="4C1BA1DEFF0CFF50FC53FAFBA9CBEF24" box="[906,1260,1347,1374]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="169" pageNumber="170" phylum="Tracheophyta" rank="variety" species="octopetala" variety="ovatiformis">
<emphasis id="B96F064FFF0CFF50FC53FAFBA90DEF24" box="[906,1066,1347,1373]" italics="true" pageId="169" pageNumber="170">B. octopetala</emphasis>
var.
<emphasis id="B96F064FFF0CFF50FBBCFAFCA9CBEF24" box="[1125,1260,1347,1373]" italics="true" pageId="169" pageNumber="170">ovatiformis</emphasis>
</taxonomicName>
is tuberous and likely dies down to its tuber in the dry season.
</paragraph>
</subSubSection>
</treatment>
</document>
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