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<document id="0A930360898702E81AA03678D4977FA4" ID-DOI="https://doi.org/10.5852/ejt.2023.881.2175" ID-ISSN="2118-9773" ID-Zenodo-Dep="8178280" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690201937279" checkinUser="felipe" docAuthor="Moonlight, Peter. W., Jara-Muñoz, Orlando A., Purvis, David A., Delves, Jay, Allen, Josh P. &amp; Reynel, Carlos" docDate="2023" docId="03B26B4BFE83FED0FDCFF9F0A9B0EB3E" docLanguage="en" docName="ejt-881_1-334.imf" docOrigin="European Journal of Taxonomy 881" docSource="http://dx.doi.org/10.5852/ejt.2023.881.2175" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia speculum Moonlight &amp; Tebbitt" docType="treatment" docVersion="7" lastPageNumber="296" masterDocId="FF8B1333FFA5FFF9FFD9FFBFAD27EA79" masterDocTitle="The genus Begonia (Begoniaceae) in Peru" masterLastPageNumber="334" masterPageNumber="1" pageNumber="295" updateTime="1707123162844" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3ED86A4ACB10E612B7AFA36387B4D84A">The genus Begonia (Begoniaceae) in Peru</mods:title>
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<mods:affiliation id="84C56DD0AA4204B3C85FBFBA6A111505">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK. &amp; Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland.</mods:affiliation>
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74. 
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<emphasis id="B96F064FFE83FEDFFD9BF9F0AE04EC10" bold="true" box="[578,803,1614,1641]" italics="true" pageId="294" pageNumber="295">Begonia speculum</emphasis>
Moonlight &amp; Tebbitt
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<figureCitation id="1320C6D8FE83FEDFFD17F9C9AE1CECE9" box="[718,827,1654,1680]" captionStart="Fig" captionStartId="288.[189,232,1857,1883]" captionTargetBox="[194,1390,1193,1812]" captionTargetId="figure-313@288.[189,1398,1183,1818]" captionTargetPageId="288" captionText="Fig. 90. Distribution of the Begonia sect. Ignota in Peru and surrounding countries. A. B. bifurcata L.B.Sm. &amp; B.G.Schub. (red), B. erythrothrix Tebbitt &amp; Moonlight (red), and B. occultata J.P.Allen &amp; Moonlight sp. nov. (green). B. B. speculum Moonlight &amp; Tebbitt (red), B. thyrsoidea Irmsch., and B. urubambensis Tebbitt (green). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178495" httpUri="https://zenodo.org/record/8178495/files/figure.png" pageId="294" pageNumber="295">Figs 90B</figureCitation>
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<emphasis id="B96F064FFE83FEDFFF64F904AF3AECAC" box="[189,541,1723,1749]" italics="true" pageId="294" pageNumber="295">Edinburgh Journal of Botany</emphasis>
74 (2): 112 (
<bibRefCitation id="EF8AA7ACFE83FEDFFD62F904AEDAECAC" author="Moonlight &amp; Tebbitt" box="[699,1021,1723,1749]" firstAuthor="Moonlight" pageId="294" pageNumber="295" pagination="111 - 122" refId="ref229847" refString="Moonlight P. W. &amp; Tebbitt M. C. 2016. Two new Peruvian species of Begonia (Begoniaceae) and an amended description of Begonia thyrsoidea. Edinburgh Journal of Botany 74 (2): 111 - 122. https: // doi. org / 10.1017 / S 0960428616000299" type="journal article" year="2016">Moonlight &amp; Tebbitt 2016</bibRefCitation>
).
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– 
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• 
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&amp; 
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158
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<emphasis id="B96F064FFE83FEDFFF64F8CAAC62EDF6" bold="true" box="[189,325,1909,1935]" pageId="294" pageNumber="295">Etymology</emphasis>
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The epithet is the Latin noun ‘ 
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<emphasis id="B96F064FFE83FEDFFDFDF818AFB3EDB8" box="[548,660,1959,1985]" italics="true" pageId="294" pageNumber="295">speculum</emphasis>
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’, which means ‘mirror’. It refers to the relatively symmetrical leaves of this species.
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<emphasis id="B96F064FFE82FEDEFF64FEB7ACE3EB5B" bold="true" box="[189,452,264,290]" pageId="295" pageNumber="296">Specimens examined</emphasis>
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</paragraph>
<paragraph id="8BA4DA5DFE82FEDEFF64FE86A850EBE2" blockId="295.[189,1399,310,411]" pageId="295" pageNumber="296">
<materialsCitation id="3B73D000FE82FEDEFF64FE86AED0EB0F" collectingDate="2018-07-04" collectionCode="USM" collectorName="P. W. Moonlight" country="Peru" county="Prov. Rioja" elevation="1297" latitude="-5.669722" location="Road from Pedro Ruiz to Rioja" longLatPrecision="21" longitude="-77.856384" pageId="295" pageNumber="296" specimenCount="1" stateProvince="San Martin Region">
<collectingCountry id="F30C9ACDFE82FEDEFF64FE86AC2FEB2A" box="[189,264,313,339]" name="Peru" pageId="295" pageNumber="296">PERU</collectingCountry>
– 
<emphasis id="B96F064FFE82FEDEFEF5FE87AFEBEB2B" bold="true" box="[300,716,311,338]" pageId="295" pageNumber="296">
<collectingRegion id="49DF14BFFE82FEDEFEF5FE87AF0EEB2B" box="[300,553,311,338]" country="Peru" name="San Martin" pageId="295" pageNumber="296">San Martín Region</collectingRegion>
: 
<collectingCounty id="62C5A2D1FE82FEDEFDE3FE87AFEBEB2B" box="[570,716,312,338]" pageId="295" pageNumber="296">Prov. Rioja</collectingCounty>
</emphasis>
• 
<location id="8EC48C86FE82FEDEFD34FE87A948EB2B" LSID="urn:lsid:plazi:treatment:03B26B4BFE83FED0FDCFF9F0A9B0EB3E:8EC48C86FE82FEDEFD34FE87A948EB2B" box="[749,1135,312,339]" country="Peru" county="Prov. Rioja" latitude="-5.669722" longLatPrecision="21" longitude="-77.856384" name="Road from Pedro Ruiz to Rioja" pageId="295" pageNumber="296" stateProvince="San Martin Region">Road from Pedro Ruiz to Rioja</location>
, 
<locationDeviation id="9A16C37FFE82FEDEFBA7FE87A9FAEB2B" box="[1150,1245,312,338]" pageId="295" pageNumber="296">km 393</locationDeviation>
; 
<geoCoordinate id="EE2FBC9AFE82FEDEFB34FE89A856EB2B" box="[1261,1393,310,338]" degrees="5" direction="south" minutes="40" orientation="latitude" pageId="295" pageNumber="296" precision="15" seconds="11" value="-5.669722">5°40′11″ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFE82FEDEFF64FEE4AC7DEB0E" box="[189,346,347,375]" degrees="77" direction="west" minutes="51" orientation="longitude" pageId="295" pageNumber="296" precision="15" seconds="23" value="-77.856384">77°51′23″ W</geoCoordinate>
; 
<elevation id="00363D6EFE82FEDEFEB1FEE3AF23EB0F" box="[360,516,348,375]" metricMagnitude="3" metricUnit="m" metricValue="1.297" pageId="295" pageNumber="296" unit="m" value="1297.0">
<quantity id="4CE377B8FE82FEDEFEB1FEE3ACE5EB0E" box="[360,450,348,375]" metricMagnitude="3" metricUnit="m" metricValue="1.297" pageId="295" pageNumber="296" unit="m" value="1297.0">1297 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFE82FEDEFDCAFEE3AFBCEB0F" box="[531,667,348,374]" pageId="295" pageNumber="296" value="2018-07-04">
<collectingDate id="EFE10575FE82FEDEFDCAFEE3AFBCEB0F" box="[531,667,348,374]" pageId="295" pageNumber="296" value="2018-07-04">4 Jul. 2018</collectingDate>
</date>
; 
<emphasis id="B96F064FFE82FEDEFD70FEE2AE82EB0F" box="[681,933,348,374]" italics="true" pageId="295" pageNumber="296">
<collectorName id="26EEBF8BFE82FEDEFD70FEE2AE46EB0F" box="[681,865,348,374]" pageId="295" pageNumber="296">P.W. Moonlight</collectorName>
1261
</emphasis>
; 
<collectionCode id="ED0A4298FE82FEDEFC6EFEE3AED0EB0F" box="[951,1015,348,374]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="295" pageNumber="296">USM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B73D000FE82FEDEFBCAFEE2A850EBE2" collectingDate="2010-07-09" collectionCode="USM" collectorName="M. M. Mora &amp; J. Perez" elevation="1750" latitude="-5.7" location="Bosque de Proteccion de Alto Mayo (BPAM)" longLatPrecision="1305" longitude="-77.73333" pageId="295" pageNumber="296" specimenCount="1">
<location id="8EC48C86FE82FEDEFBCAFEE2AC55EBE2" LSID="urn:lsid:plazi:treatment:03B26B4BFE83FED0FDCFF9F0A9B0EB3E:8EC48C86FE82FEDEFBCAFEE2AC55EBE2" latitude="-5.7" longLatPrecision="1305" longitude="-77.73333" name="Bosque de Proteccion de Alto Mayo (BPAM)" pageId="295" pageNumber="296">Bosque de Protección de Alto Mayo (BPAM)</location>
; 
<geoCoordinate id="EE2FBC9AFE82FEDEFEA7FEC0ACF3EBE2" box="[382,468,383,411]" degrees="5" direction="south" minutes="42" orientation="latitude" pageId="295" pageNumber="296" precision="925" value="-5.7">5°42′ S</geoCoordinate>
, 
<geoCoordinate id="EE2FBC9AFE82FEDEFE38FEC0AF75EBE2" box="[481,594,383,411]" degrees="77" direction="west" minutes="44" orientation="longitude" pageId="295" pageNumber="296" precision="925" value="-77.73333">77°44′ W</geoCoordinate>
; 
<elevation id="00363D6EFE82FEDEFD87FE3EAE67EBE3" box="[606,832,384,411]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="295" pageNumber="296" unit="m" value="1750.0" valueMax="1800.0" valueMin="1700.0">
<quantity id="4CE377B8FE82FEDEFD87FE3EAE27EBE2" box="[606,768,385,411]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="295" pageNumber="296" unit="m" value="1750.0" valueMax="1800.0" valueMin="1700.0">1700–1800 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFA5FC9DFE82FEDEFC97FE3EAEF5EBE2" box="[846,978,384,411]" pageId="295" pageNumber="296" value="2010-07-09">
<collectingDate id="EFE10575FE82FEDEFC97FE3EAEF5EBE2" box="[846,978,384,411]" pageId="295" pageNumber="296" value="2010-07-09">9 Jul. 2010</collectingDate>
</date>
; 
<emphasis id="B96F064FFE82FEDEFC07FE3EA807EBE2" box="[990,1312,385,411]" italics="true" pageId="295" pageNumber="296">
<collectorName id="26EEBF8BFE82FEDEFC07FE3EA94FEBE3" box="[990,1128,385,410]" pageId="295" pageNumber="296">M.M. Mora</collectorName>
&amp; 
<collectorName id="26EEBF8BFE82FEDEFB57FE3EA9CBEBE2" box="[1166,1260,385,411]" pageId="295" pageNumber="296">J. Pérez</collectorName>
883
</emphasis>
; 
<collectionCode id="ED0A4298FE82FEDEFAE9FE3EA854EBE2" box="[1328,1395,385,411]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="295" pageNumber="296">USM</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE82FEDEFF64FE7CA879EF51" pageId="295" pageNumber="296" type="description">
<paragraph id="8BA4DA5DFE82FEDEFF64FE7CAC68EBA4" blockId="295.[189,335,451,477]" box="[189,335,451,477]" pageId="295" pageNumber="296">
<heading id="D0EC6D31FE82FEDEFF64FE7CAC68EBA4" bold="true" box="[189,335,451,477]" fontSize="11" level="3" pageId="295" pageNumber="296" reason="3">
<emphasis id="B96F064FFE82FEDEFF64FE7CAC68EBA4" bold="true" box="[189,335,451,477]" pageId="295" pageNumber="296">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE82FEDEFF64FE4DA879EF51" blockId="295.[188,1400,498,1321]" pageId="295" pageNumber="296">
Caulescent, tuberous herb, to 
<quantity id="4CE377B8FE82FEDEFDC1FE4DAF46E874" box="[536,609,498,525]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="295" pageNumber="296" unit="cm" value="15.0">15 cm</quantity>
high. 
<emphasis id="B96F064FFE82FEDEFD73FE4DAFC8E875" box="[682,751,498,524]" italics="true" pageId="295" pageNumber="296">Tuber</emphasis>
sub-globose, 1.5–2 × 
<quantity id="4CE377B8FE82FEDEFC2AFE4DA944E874" box="[1011,1123,498,525]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="295" pageNumber="296" unit="cm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 cm</quantity>
, with 1 growing points. 
<emphasis id="B96F064FFE82FEDEFF64FDA8AC25E848" box="[189,258,535,561]" italics="true" pageId="295" pageNumber="296">Stems</emphasis>
4, erect, unbranched; internodes to 
<quantity id="4CE377B8FE82FEDEFD61FDA9AE2CE848" box="[696,779,534,561]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="295" pageNumber="296" unit="cm" value="4.5">4.5 cm</quantity>
long, to 
<quantity id="4CE377B8FE82FEDEFCA5FDA9AEFFE848" box="[892,984,534,561]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="295" pageNumber="296" unit="mm" value="0.5">0.5 mm</quantity>
thick, wiry, pink to red, glabrous. 
<emphasis id="B96F064FFE82FEDEFF64FD84AC3CE82C" box="[189,283,571,597]" italics="true" pageId="295" pageNumber="296">Stipules</emphasis>
late deciduous, lanceolate, 1–4.5 × 
<quantity id="4CE377B8FE82FEDEFD60FD84AE3BE82C" box="[697,796,571,597]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="295" pageNumber="296" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
, apex acute, opaque, red to brown, glabrous, margin entire, aciliate. 
<emphasis id="B96F064FFE82FEDEFEACFDDFACE0E800" box="[373,455,608,633]" italics="true" pageId="295" pageNumber="296">Leaves</emphasis>
2–4 per stem, alternate, basifixed; petiole 
<quantity id="4CE377B8FE82FEDEFC67FDE0A90AE800" box="[958,1069,607,633]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="7.0" metricValueMin="0.8" pageId="295" pageNumber="296" unit="cm" value="3.9" valueMax="7.0" valueMin="0.8">0.8–7 cm</quantity>
long, pink to red, glabrous; blade subsymmetric, ovate, to 6.5 × 
<quantity id="4CE377B8FE82FEDEFDB5FD3CAF98E8E7" box="[620,703,643,670]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="295" pageNumber="296" unit="cm" value="3.5">3.5 cm</quantity>
, succulent, apex acute, base shallowly-cordate, basal lobes not overlapping, sinus to 
<quantity id="4CE377B8FE82FEDEFE37FD18AF14E8BB" box="[494,563,679,706]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="295" pageNumber="296" unit="mm" value="4.0">4 mm</quantity>
deep, margin irregularly crenate, ciliate, upper surface copper to dark green, glabrous, lower surface red, glabrous, veins palmate-pinnate, 5–7 veined from the base, 2–3 veins on the larger side, 2–3 veins on the smaller side. 
<emphasis id="B96F064FFE82FEDEFCBFFD4FA929E973" box="[870,1038,752,778]" italics="true" pageId="295" pageNumber="296">Inflorescences</emphasis>
1–3, bisexual, axillary, erect, cymose, with up to 4 branches, bearing up to 4 staminate flowers and 4 pistillate flowers, protandrous; peduncle to 
<quantity id="4CE377B8FE82FEDEFE93FC87ACBEE92A" box="[330,409,824,851]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="295" pageNumber="296" unit="cm" value="5.5">5.5 cm</quantity>
long, green to red, glabrous, bracts persistent, elliptic to lanceolate, 2–4 × 
<quantity id="4CE377B8FE82FEDEFB20FC87A855E92A" box="[1273,1394,824,851]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="295" pageNumber="296" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
, translucent, pale green, glabrous, apex obtuse, margin entire, glabrous. 
<emphasis id="B96F064FFE82FEDEFBCBFCE2A9C2E90F" box="[1042,1253,860,887]" italics="true" pageId="295" pageNumber="296">Staminate flowers</emphasis>
: pedicels to 
<quantity id="4CE377B8FE82FEDEFF66FC3FAC29E9E2" box="[191,270,896,923]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="295" pageNumber="296" unit="mm" value="15.0">15 mm</quantity>
long, glabrous; tepals 4, spreading, outer 2 elliptic to obovate, 6–7 × 
<quantity id="4CE377B8FE82FEDEFBF3FC3EA9AAE9E2" box="[1066,1165,897,923]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="295" pageNumber="296" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
, apex obtuse, white, glabrous, margin entire, aciliate, inner 2 elliptic to oblanceolate, 4–6 × 
<quantity id="4CE377B8FE82FEDEFC23FC1BA953E9C6" box="[1018,1140,932,959]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="295" pageNumber="296" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
, apex acute to obtuse, white, glabrous, margin entire, aciliate; stamens 15–20, spreading, yellow, filaments 
<quantity id="4CE377B8FE82FEDEFB49FC76A9D7E99A" box="[1168,1264,969,995]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="295" pageNumber="296" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
long, fused into a 
<quantity id="4CE377B8FE82FEDEFEDEFC52AC45EE71" box="[263,354,1005,1032]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="295" pageNumber="296" unit="mm" value="0.5">0.5 mm</quantity>
column, anthers cuboid, 0.5 × 
<quantity id="4CE377B8FE82FEDEFD08FC52AE49EE71" box="[721,878,1005,1032]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="295" pageNumber="296" unit="mm" value="0.625" valueMax="0.75" valueMin="0.5">0.5–0.75 mm</quantity>
long, dehiscing via lateral slits, connectives extended 
<quantity id="4CE377B8FE82FEDEFEEBFBAEACB5EE55" box="[306,402,1041,1068]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="295" pageNumber="296" unit="mm" value="0.1">0.1 mm</quantity>
, symmetrically basifixed. 
<emphasis id="B96F064FFE82FEDEFD17FBAEAEBEEE52" box="[718,921,1041,1067]" italics="true" pageId="295" pageNumber="296">Pistillate flowers</emphasis>
: pedicels to 
<quantity id="4CE377B8FE82FEDEFBE1FBAEA9A9EE55" box="[1080,1166,1041,1068]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="295" pageNumber="296" unit="mm" value="20.0">20 mm</quantity>
long; bracteoles 2, positioned directly beneath the ovary, elliptic to lanceolate, 2–4 × 
<quantity id="4CE377B8FE82FEDEFC15FB8AA960EE29" box="[972,1095,1077,1104]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="295" pageNumber="296" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
, apex obtuse, translucent, pale green, glabrous, margin entire, aciliate; tepals 5, subequal, persistent in fruit, spreading, the ovate to obovate, 3.5–6 × 
<quantity id="4CE377B8FE82FEDEFE5CFBC1ACD9EEE1" box="[389,510,1150,1177]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="295" pageNumber="296" unit="mm" value="4.25" valueMax="5.0" valueMin="3.5">3.5–5 mm</quantity>
, apex obtuse to rounded, white, glabrous, margin entire, aciliate; ovary body cuboid, 3.5–6 × 
<quantity id="4CE377B8FE82FEDEFEA3FB1DACD2EEC4" box="[378,501,1186,1213]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="6.0" metricValueMin="3.6" pageId="295" pageNumber="296" unit="mm" value="4.8" valueMax="6.0" valueMin="3.6">3.6–6 mm</quantity>
, pale green, glabrous, equally 3-winged, wings marginal, 0.5–1 × 
<quantity id="4CE377B8FE82FEDEFB21FB1DA855EEC4" box="[1272,1394,1186,1213]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="6.0" metricValueMin="3.5" pageId="295" pageNumber="296" unit="mm" value="4.75" valueMax="6.0" valueMin="3.5">3.5–6 mm</quantity>
; 2- or 3-locular, placentae branches entire, bearing ovules on both surfaces; styles 2 or 3, yellow, free to base, 
<quantity id="4CE377B8FE82FEDEFED8FB55AC44EF7C" box="[257,355,1258,1285]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="295" pageNumber="296" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long, 2-lobed, stigmatic papillae in a spiral band. 
<emphasis id="B96F064FFE82FEDEFC6DFB54A952EF7D" box="[948,1141,1258,1284]" italics="true" pageId="295" pageNumber="296">Fruiting pedicel</emphasis>
to 
<quantity id="4CE377B8FE82FEDEFB42FB55A9C8EF7C" box="[1179,1263,1258,1285]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="295" pageNumber="296" unit="mm" value="25.0">25 mm</quantity>
long. 
<emphasis id="B96F064FFE82FEDEFAE3FB54ADD1EF51" italics="true" pageId="295" pageNumber="296">Fruit body</emphasis>
globose, to 6 × 
<quantity id="4CE377B8FE82FEDEFE6DFAB1ACDCEF50" box="[436,507,1294,1321]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="295" pageNumber="296" unit="mm" value="6.0">6 mm</quantity>
, drying brown, wings same shape as in ovary, expanding to 
<quantity id="4CE377B8FE82FEDEFB65FAB1A831EF50" box="[1212,1302,1294,1321]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="295" pageNumber="296" unit="mm" value="2.5">2.5 mm</quantity>
wide.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE82FEDEFF64FAEEAC08EC52" pageId="295" pageNumber="296" type="conservation">
<paragraph id="8BA4DA5DFE82FEDEFF64FAEEAF4AEF12" blockId="295.[189,621,1361,1387]" box="[189,621,1361,1387]" pageId="295" pageNumber="296">
<heading id="D0EC6D31FE82FEDEFF64FAEEAF4AEF12" bold="true" box="[189,621,1361,1387]" fontSize="11" level="3" pageId="295" pageNumber="296" reason="3">
<emphasis id="B96F064FFE82FEDEFF64FAEEAF4AEF12" bold="true" box="[189,621,1361,1387]" pageId="295" pageNumber="296">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE82FEDEFF64FA3FAC08EC52" blockId="295.[189,1400,1408,1579]" pageId="295" pageNumber="296">
Assessed by 
<bibRefCitation id="EF8AA7ACFE82FEDEFE81FA3FAF80EFE3" author="Moonlight &amp; Tebbitt" box="[344,679,1408,1435]" firstAuthor="Moonlight" pageId="295" pageNumber="296" pagination="111 - 122" refId="ref229847" refString="Moonlight P. W. &amp; Tebbitt M. C. 2016. Two new Peruvian species of Begonia (Begoniaceae) and an amended description of Begonia thyrsoidea. Edinburgh Journal of Botany 74 (2): 111 - 122. https: // doi. org / 10.1017 / S 0960428616000299" type="journal article" year="2016">Moonlight &amp; Tebbitt (2016)</bibRefCitation>
as Vulnerable (VU D2), based upon its AOO of &lt;
<quantity id="4CE377B8FE82FEDEFAD7FA3EA86CEFE3" box="[1294,1355,1408,1435]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="295" pageNumber="296" unit="km" value="8.0">8 km</quantity>
<superScript id="7C6E7715FE82FEDEFA92FA3FA873EFF7" attach="left" box="[1355,1364,1408,1422]" fontSize="6" pageId="295" pageNumber="296">2</superScript>
. A further population has been identified 
<quantity id="4CE377B8FE82FEDEFDAEFA1BAF98EFC6" box="[631,703,1444,1471]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="295" pageNumber="296" unit="km" value="50.0">50 km</quantity>
to the SW of the 
<typeStatus id="54A064FFFE82FEDEFC5EFA1AAE9DEFC6" box="[903,954,1445,1471]" pageId="295" pageNumber="296">type</typeStatus>
locality and one further collection has been made at the 
<typeStatus id="54A064FFFE82FEDEFE52FA76AC99EF9A" box="[395,446,1481,1507]" pageId="295" pageNumber="296">type</typeStatus>
locality (
<emphasis id="B96F064FFE82FEDEFDF7FA75AE00EF9A" box="[558,807,1481,1507]" italics="true" pageId="295" pageNumber="296">P.W. Moonlight 1261</emphasis>
), where the population appeared stable. The AOO of the species is now 
<quantity id="4CE377B8FE82FEDEFE19FA52AF2CEC7E" box="[448,523,1517,1543]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="295" pageNumber="296" unit="km" value="12.0">12 km</quantity>
<superScript id="7C6E7715FE82FEDEFDD2FA53AF33EF83" attach="left" box="[523,532,1516,1530]" fontSize="6" pageId="295" pageNumber="296">2</superScript>
and remains within the criteria required for an assessment as Vulnerable (VU D2).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE82FEDEFF64F9ECAC3AEC9F" pageId="295" pageNumber="296" type="discussion">
<paragraph id="8BA4DA5DFE82FEDEFF64F9ECAC97EC17" blockId="295.[189,432,1619,1646]" box="[189,432,1619,1646]" pageId="295" pageNumber="296">
<heading id="D0EC6D31FE82FEDEFF64F9ECAC97EC17" bold="true" box="[189,432,1619,1646]" fontSize="11" level="3" pageId="295" pageNumber="296" reason="3">
<emphasis id="B96F064FFE82FEDEFF64F9ECAC97EC17" bold="true" box="[189,432,1619,1646]" pageId="295" pageNumber="296">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFE82FEDEFF64F93CAC3AEC9F" blockId="295.[189,1399,1667,1766]" pageId="295" pageNumber="296">
The only tuberous species of 
<taxonomicName id="4C1BA1DEFE82FEDEFDF2F93BAFA9ECE4" baseAuthorityName="Moonlight &amp; Fuentes" baseAuthorityYear="2022" box="[555,654,1668,1693]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="295" pageNumber="296" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F064FFE82FEDEFDF2F93BAFA9ECE4" box="[555,654,1668,1693]" italics="true" pageId="295" pageNumber="296">Begonia</emphasis>
</taxonomicName>
in northern 
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with subsymmetric, ovate leaves. It is also unusual among tuberous Peruvian begonias in its crenate leaf margins and stamens held on a short column.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FE82FED0FF64F8B1A9B0EB3E" lastPageId="297" lastPageNumber="298" pageId="295" pageNumber="296" type="distribution">
<paragraph id="8BA4DA5DFE82FEDEFF64F8B1ACD2ED51" blockId="295.[189,501,1806,1832]" box="[189,501,1806,1832]" pageId="295" pageNumber="296">
<heading id="D0EC6D31FE82FEDEFF64F8B1ACD2ED51" bold="true" box="[189,501,1806,1832]" fontSize="11" level="3" pageId="295" pageNumber="296" reason="3">
<emphasis id="B96F064FFE82FEDEFF64F8B1ACD2ED51" bold="true" box="[189,501,1806,1832]" pageId="295" pageNumber="296">Distribution and ecology</emphasis>
</heading>
</paragraph>
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Endemic to 
<collectingCountry id="F30C9ACDFE82FEDEFE95F881ACA5ED21" box="[332,386,1854,1880]" name="Peru" pageId="295" pageNumber="296">Peru</collectingCountry>
and Rioja Province, 
<collectingRegion id="49DF14BFFE82FEDEFDA2F881AE7BED2E" box="[635,860,1853,1880]" country="Peru" name="San Martin" pageId="295" pageNumber="296">San Martín Region</collectingRegion>
(
<figureCitation id="1320C6D8FE82FEDEFCB5F882AEFFED21" box="[876,984,1853,1880]" captionStart="Fig" captionStartId="288.[189,232,1857,1883]" captionTargetBox="[194,1390,1193,1812]" captionTargetId="figure-313@288.[189,1398,1183,1818]" captionTargetPageId="288" captionText="Fig. 90. Distribution of the Begonia sect. Ignota in Peru and surrounding countries. A. B. bifurcata L.B.Sm. &amp; B.G.Schub. (red), B. erythrothrix Tebbitt &amp; Moonlight (red), and B. occultata J.P.Allen &amp; Moonlight sp. nov. (green). B. B. speculum Moonlight &amp; Tebbitt (red), B. thyrsoidea Irmsch., and B. urubambensis Tebbitt (green). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178495" httpUri="https://zenodo.org/record/8178495/files/figure.png" pageId="295" pageNumber="296">Fig. 90B</figureCitation>
). Found in lower montane forest at an elevation of 
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a.s.l. where it grows directly on limestone in the shade of montane forest and with its tuber buried in alkaline soil. 
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<emphasis id="B96F064FFE82FEDEFD6DF838AEB6EDD9" box="[692,913,1926,1952]" italics="true" pageId="295" pageNumber="296">Begonia speculum</emphasis>
Moonlight &amp; Tebbitt
</taxonomicName>
has been collected from two populations in Bosque de Protección Alto Mayo and identified from photographs taken by Wenbo Chen of a third population ca 
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ESE of Nuevo 
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(
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, 
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; [
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a.s.l.]). As a tuberous herb, 
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<emphasis id="B96F064FFE8CFED0FD5BFEB5AE30EB5A" box="[642,791,265,291]" italics="true" pageId="297" pageNumber="298">B. speculum</emphasis>
</taxonomicName>
presumably has a dormant period but it has been collected in flower and fruit in February and July, so it remains unclear when this is.
</paragraph>
</subSubSection>
</treatment>
</document>