322 lines
35 KiB
XML
322 lines
35 KiB
XML
<document id="161312860F27C062064D0229CD76A0BF" ID-DOI="10.11646/zootaxa.4034.2.8" ID-GBIF-Dataset="d3099fd2-efc2-45cd-a1f1-3d47c8312a4b" ID-ISSN="1175-5326" ID-Zenodo-Dep="239394" ID-ZooBank="5C08231F-2280-4FF8-A042-F94F04466AE3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461300330510" checkinUser="plazi" docAuthor="Cavalcanti, Thaynã, Santos, George Garcia & Pinheiro, Ulisses" docDate="2015" docId="03B2565BFD6EFFEB9EBBF88EFA2FFCE0" docLanguage="en" docName="zt04034p374.pdf" docOrigin="Zootaxa 4034 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Desmacella tylovariabilis Cavalcanti, Santos & Pinheiro, 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="371" masterDocId="FF8B2E23FD6BFFEC9E2CFFD1FF38FFD8" masterDocTitle="Desmacella Schmidt, 1870 from Brazil: Description of two new species and a review of records (Desmacellida: Demospongiae: Porifera)" masterLastPageNumber="374" masterPageNumber="364" pageNumber="369" updateTime="1698617292245" updateUser="plazi">
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<mods:mods id="2DED67F1DFE32A7575EF8BFAA1DF065C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2C1AEE4BAE7AD9E01B18D8270622B32E">
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<mods:title id="FB00CAB6B274DA33951BC7DC29E246F2">Desmacella Schmidt, 1870 from Brazil: Description of two new species and a review of records (Desmacellida: Demospongiae: Porifera)</mods:title>
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<mods:name id="45660616BF5AC92E62E64A069EA49B54" type="personal">
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<mods:roleTerm id="C867FAD8744B6D21A67C7808F0C760D3">Author</mods:roleTerm>
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<mods:namePart id="9225831140D4A94DFC38061B4C66EB07">Cavalcanti, Thaynã</mods:namePart>
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<mods:roleTerm id="CB8103FA6A837E9BA6B37099054DDED2">Author</mods:roleTerm>
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<mods:namePart id="7372EF561B0CEC7ACA6F5F534A28EB4C">Santos, George Garcia</mods:namePart>
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</mods:name>
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<mods:name id="18125EB0CE9D5EB1A0E3929ECB5D1E33" type="personal">
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<mods:roleTerm id="1A1368C48571B7211F1552F1A7BFA59C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2106B72246B4D8AEBCD17C79CC008ED1">Pinheiro, Ulisses</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="D9F85AC9544C496AE75E64797E61BD07">text</mods:typeOfResource>
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<mods:title id="E73CD795814B35D1879B7160F94E21C2">Zootaxa</mods:title>
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<mods:part id="1CAA3B2A9AC446B96FDCA73A35F7BA8F">
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<mods:date id="B340A5F64323A254CC10865CC5F62CE5">2015</mods:date>
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<mods:detail id="7E816DB0BC51F5469D0B79B3E169B108" type="volume">
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<mods:number id="A9D1BA300A7195740BFD1FB96BD6A541">4034</mods:number>
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<mods:detail id="87A4D81F728D9B767466CB8465BA67CD" type="issue">
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<mods:number id="D917A1DCE6DEF12727E4E572635BDC03">2</mods:number>
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<mods:classification id="24EFD2777883FC0D88495C86D4F3B011">journal article</mods:classification>
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<mods:identifier id="B9C7825AEA971A14FECE681D30F2FE60" type="DOI">10.11646/zootaxa.4034.2.8</mods:identifier>
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<treatment id="03B2565BFD6EFFEB9EBBF88EFA2FFCE0" ID-DOI="http://doi.org/10.5281/zenodo.6112719" ID-GBIF-Taxon="119643490" ID-Zenodo-Dep="6112719" LSID="urn:lsid:plazi:treatment:03B2565BFD6EFFEB9EBBF88EFA2FFCE0" httpUri="http://treatment.plazi.org/id/03B2565BFD6EFFEB9EBBF88EFA2FFCE0" lastPageId="7" lastPageNumber="371" pageId="5" pageNumber="369">
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<subSubSection id="C301B4C6FD6EFFE99EBBF88EFEB7F843" pageId="5" pageNumber="369" type="nomenclature">
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<paragraph id="8BA4E74DFD6EFFE99EBBF88EFD13F8A1" blockId="5.[151,555,1887,1947]" box="[151,555,1887,1913]" pageId="5" pageNumber="369">
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<heading id="D0EC5021FD6EFFE99EBBF88EFD13F8A1" bold="true" box="[151,555,1887,1913]" fontSize="11" level="1" pageId="5" pageNumber="369" reason="1">
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<emphasis id="B96F3B5FFD6EFFE99EBBF88EFD13F8A1" bold="true" box="[151,555,1887,1913]" pageId="5" pageNumber="369">
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||
<taxonomicName id="4C1B9CCEFD6EFFE99EBBF88EFEFFF8A1" box="[151,455,1887,1913]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="5" pageNumber="369" phylum="Porifera" rank="species" species="tylovariabilis" status="sp. nov.">
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<emphasis id="B96F3B5FFD6EFFE99EBBF88EFEFFF8A1" bold="true" box="[151,455,1887,1913]" italics="true" pageId="5" pageNumber="369">Desmacella tylovariabilis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25C8624FD6EFFE99FE2F88EFD13F8A1" box="[462,555,1887,1913]" pageId="5" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA4E74DFD6EFFE99EBBF853FEB7F843" blockId="5.[151,555,1887,1947]" box="[151,399,1922,1947]" pageId="5" pageNumber="369">
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||
(
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<figureCitation id="1320FBC8FD6EFFE99EB2F853FE31F843" box="[158,265,1922,1947]" captionStart="FIGURE 1" captionStartId="1.[151,250,1980,2002]" captionTargetBox="[154,1417,303,1955]" captionTargetId="figure@1.[154,1433,303,1955]" captionTargetPageId="1" captionText="FIGURE 1. Location of the collection sites of Desmacella species from Brazil. (1) Desmacella microsigmata sp. nov. (Rio Grande do Norte State); (2) Desmacella tylovariabilis sp. nov. (Rio de Janeiro State); (3) Desmacella annexa (São Paulo State)." httpUri="https://zenodo.org/record/239395/files/figure.png" pageId="5" pageNumber="369">Figures 1</figureCitation>
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,
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<figureCitation id="1320FBC8FD6EFFE99F3BF852FE1FF843" box="[279,295,1923,1947]" captionStart="FIGURE 3" captionStartId="6.[151,250,1663,1685]" captionTargetBox="[226,1359,215,1641]" captionTargetId="figure@6.[226,1360,193,1642]" captionTargetPageId="6" captionText="FIGURE 3. Desmacella tylovariabilis sp. nov. (MNRJ 7339) in SEM. A, preserved specimen; B, thick section showing the ectosome and choanosome; C, size variation of tylostyles; D, variation of tylostyle bases; E, sigmas; F, details of microspines on the tip of a sigma. Scale bars: A = 1 cm; B = 200 µm; C = 400 µm; D – E = 10 µm; F = 2 µm." httpUri="https://zenodo.org/record/239397/files/figure.png" pageId="5" pageNumber="369">3</figureCitation>
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, Table 1)
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</paragraph>
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</subSubSection>
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<subSubSection id="C301B4C6FD6EFFEA9EBBF81BFC0FF8EF" lastPageId="6" lastPageNumber="370" pageId="5" pageNumber="369" type="materials_examined">
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<paragraph id="8BA4E74DFD6EFFE99EBBF81BFC13F83B" blockId="5.[151,811,1994,2019]" box="[151,811,1994,2019]" pageId="5" pageNumber="369">
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<emphasis id="B96F3B5FFD6EFFE99EBBF81BFE01F83B" bold="true" box="[151,313,1994,2019]" pageId="5" pageNumber="369">
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<typeStatus id="54A059EFFD6EFFE99EBBF81BFFEAF83B" box="[151,210,1994,2019]" pageId="5" pageNumber="369">Type</typeStatus>
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locality:
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</emphasis>
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<collectingCountry id="F30CA7DDFD6EFFE99F6CF81BFEBFF83B" box="[320,391,1994,2019]" name="Brazil" pageId="5" pageNumber="369">Brazil</collectingCountry>
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, Rio de Janeiro State, Campos Basin.
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</paragraph>
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<caption id="DF64B7C5FD6DFFEA9EBBF9AEFB65F90A" httpUri="https://zenodo.org/record/239397/files/figure.png" pageId="6" pageNumber="370" targetBox="[226,1359,215,1641]" targetPageId="6">
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<paragraph id="8BA4E74DFD6DFFEA9EBBF9AEFB65F90A" blockId="6.[151,1436,1663,1746]" pageId="6" pageNumber="370">
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<emphasis id="B96F3B5FFD6DFFEA9EBBF9AEFE2CF94C" bold="true" box="[151,276,1663,1685]" pageId="6" pageNumber="370">FIGURE 3.</emphasis>
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<taxonomicName id="4C1B9CCEFD6DFFEA9F30F9AEFD1BF94D" box="[284,547,1663,1685]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="6" pageNumber="370" phylum="Porifera" rank="species" species="tylovariabilis" status="sp. nov.">
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<emphasis id="B96F3B5FFD6DFFEA9F30F9AEFD1BF94D" box="[284,547,1663,1685]" italics="true" pageId="6" pageNumber="370">Desmacella tylovariabilis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25C8624FD6DFFEA9C07F951FD45F94D" box="[555,637,1664,1685]" pageId="6" pageNumber="370" rank="species">
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<emphasis id="B96F3B5FFD6DFFEA9C07F951FD45F94D" bold="true" box="[555,637,1664,1685]" pageId="6" pageNumber="370">sp. nov.</emphasis>
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</taxonomicNameLabel>
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(MNRJ 7339) in SEM. A, preserved specimen; B, thick section showing the ectosome and choanosome; C, size variation of tylostyles; D, variation of tylostyle bases; E, sigmas; F, details of microspines on the tip of a sigma. Scale bars: A = 1 cm; B = 200 µm; C = 400 µm; D–E = 10 µm; F = 2 µm.
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</paragraph>
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</caption>
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<paragraph id="8BA4E74DFD6DFFEA9EEBF92BFC0FF8EF" blockId="6.[151,1436,1786,2027]" pageId="6" pageNumber="370">
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<emphasis id="B96F3B5FFD6DFFEA9EEBF92BFD31F8CB" bold="true" box="[199,521,1786,1811]" pageId="6" pageNumber="370">
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<typeStatus id="54A059EFFD6DFFEA9EEBF92BFE3AF8CB" box="[199,258,1786,1811]" pageId="6" pageNumber="370">Type</typeStatus>
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specimens:
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<typeStatus id="54A059EFFD6DFFEA9FB9F92BFD3CF8CB" box="[405,516,1786,1811]" pageId="6" pageNumber="370" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED0A7F88FD6DFFEA9C3EF92AFD65F8CA" box="[530,605,1787,1810]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="370">MNRJ</collectionCode>
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7339, Campos Basin (
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<geoCoordinate id="EE2F818AFD6DFFEA9D4EF92AFCEBF8CB" box="[866,979,1786,1811]" direction="south" orientation="latitude" pageId="6" pageNumber="370" precision="925" value="-22.966667">22° 58’ S</geoCoordinate>
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;
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<geoCoordinate id="EE2F818AFD6DFFEA9DCDF92AFB7CF8CB" box="[993,1092,1787,1811]" direction="west" orientation="longitude" pageId="6" pageNumber="370" precision="925" value="-42.016666">42° 1’W</geoCoordinate>
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), Rio de Janeiro State,
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<collectingCountry id="F30CA7DDFD6DFFEA9B7CF92BFAAFF8CB" box="[1360,1431,1786,1811]" name="Brazil" pageId="6" pageNumber="370">Brazil</collectingCountry>
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,
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<quantity id="4CE34AA8FD6DFFEA9EBBF8CEFFD3F8EE" box="[151,235,1823,1847]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="6" pageNumber="370" unit="m" value="1130.0">1130 m</quantity>
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depth, dragging, coll. CENPES/
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<collectionCode id="ED0A7F88FD6DFFEA9C75F8CEFDA4F8EF" box="[601,668,1823,1847]" pageId="6" pageNumber="370">UFRJ</collectionCode>
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, (
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<date id="FFA5C18DFD6DFFEA9C83F8CEFC14F8EF" box="[687,812,1823,1847]" pageId="6" pageNumber="370" value="2003-02-07">07/II/2003</date>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C301B4C6FD6DFFEB9EEBF893FC87FE98" lastPageId="7" lastPageNumber="371" pageId="6" pageNumber="370" type="diagnosis">
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<paragraph id="8BA4E74DFD6DFFEA9EEBF893FC3BF87B" blockId="6.[151,1436,1786,2027]" pageId="6" pageNumber="370">
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<emphasis id="B96F3B5FFD6DFFEA9EEBF893FE79F883" bold="true" box="[199,321,1858,1883]" pageId="6" pageNumber="370">Diagnosis.</emphasis>
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<taxonomicName id="4C1B9CCEFD6DFFEA9F64F893FD50F882" box="[328,616,1858,1882]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="6" pageNumber="370" phylum="Porifera" rank="species" species="tylovariabilis" status="sp. nov.">
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<emphasis id="B96F3B5FFD6DFFEA9F64F893FD50F882" box="[328,616,1858,1882]" italics="true" pageId="6" pageNumber="370">Desmacella tylovariabilis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25C8624FD6DFFEA9C43F893FDF0F883" box="[623,712,1858,1883]" pageId="6" pageNumber="370" rank="species">
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<emphasis id="B96F3B5FFD6DFFEA9C43F893FDF0F883" bold="true" box="[623,712,1858,1883]" pageId="6" pageNumber="370">sp. nov.</emphasis>
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</taxonomicNameLabel>
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is the only
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<taxonomicName id="4C1B9CCEFD6DFFEA9D65F893FCF5F882" box="[841,973,1858,1882]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="6" pageNumber="370" phylum="Porifera" rank="genus">
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<emphasis id="B96F3B5FFD6DFFEA9D65F893FCF5F882" box="[841,973,1858,1882]" italics="true" pageId="6" pageNumber="370">Desmacella</emphasis>
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</taxonomicName>
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in the Tropical Western Atlantic with the combination of large tylostyles (616.0
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<date id="FFA5C18DFD6DFFEA9C75F8B6FDB4F8A7" box="[601,652,1895,1919]" pageId="6" pageNumber="370" value="2011-10">x 11</date>
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µm, length x width) bearing rounded tyles in varied positions, and sigmas with microspined ends (34.2 µm, chord length).
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</paragraph>
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<paragraph id="8BA4E74DFD6DFFEA9EEBF87FFB2BF833" blockId="6.[151,1436,1786,2027]" pageId="6" pageNumber="370">
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<emphasis id="B96F3B5FFD6DFFEA9EEBF87FFD71F81F" bold="true" box="[199,585,1966,1991]" pageId="6" pageNumber="370">
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External morphology (
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<figureCitation id="1320FBC8FD6DFFEA9FF1F87FFD1EF81F" box="[477,550,1966,1991]" captionStart="FIGURE 3" captionStartId="6.[151,250,1663,1685]" captionTargetBox="[226,1359,215,1641]" captionTargetId="figure@6.[226,1360,193,1642]" captionTargetPageId="6" captionText="FIGURE 3. Desmacella tylovariabilis sp. nov. (MNRJ 7339) in SEM. A, preserved specimen; B, thick section showing the ectosome and choanosome; C, size variation of tylostyles; D, variation of tylostyle bases; E, sigmas; F, details of microspines on the tip of a sigma. Scale bars: A = 1 cm; B = 200 µm; C = 400 µm; D – E = 10 µm; F = 2 µm." httpUri="https://zenodo.org/record/239397/files/figure.png" pageId="6" pageNumber="370">Fig. 3</figureCitation>
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A).
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</emphasis>
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Massive sponge,
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<date id="FFA5C18DFD6DFFEA9D32F87FFC7CF81E" box="[798,836,1966,1991]" pageId="6" pageNumber="370">6 x</date>
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<quantity id="4CE34AA8FD6DFFEA9D63F87FFCA7F81E" box="[847,927,1966,1991]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="370" unit="cm" value="4.5">4.5 cm</quantity>
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(length x width) (
|
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<figureCitation id="1320FBC8FD6DFFEA9A55F87FFB87F81F" box="[1145,1215,1966,1991]" captionStart="FIGURE 3" captionStartId="6.[151,250,1663,1685]" captionTargetBox="[226,1359,215,1641]" captionTargetId="figure@6.[226,1360,193,1642]" captionTargetPageId="6" captionText="FIGURE 3. Desmacella tylovariabilis sp. nov. (MNRJ 7339) in SEM. A, preserved specimen; B, thick section showing the ectosome and choanosome; C, size variation of tylostyles; D, variation of tylostyle bases; E, sigmas; F, details of microspines on the tip of a sigma. Scale bars: A = 1 cm; B = 200 µm; C = 400 µm; D – E = 10 µm; F = 2 µm." httpUri="https://zenodo.org/record/239397/files/figure.png" pageId="6" pageNumber="370">Fig. 3</figureCitation>
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A). Hispid surface, without distinct oscules, fragile consistency and colour beige (fixed in ethanol).
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</paragraph>
|
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<paragraph id="8BA4E74DFD6CFFEB9EEBFF46FAA8FF0D" blockId="7.[151,1436,151,824]" pageId="7" pageNumber="371">
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||
<emphasis id="B96F3B5FFD6CFFEB9EEBFF46FE9DFF68" bold="true" box="[199,421,151,176]" pageId="7" pageNumber="371">
|
||
Skeleton (
|
||
<figureCitation id="1320FBC8FD6CFFEB9F11FF46FEBBFF68" box="[317,387,151,176]" captionStart="FIGURE 3" captionStartId="6.[151,250,1663,1685]" captionTargetBox="[226,1359,215,1641]" captionTargetId="figure@6.[226,1360,193,1642]" captionTargetPageId="6" captionText="FIGURE 3. Desmacella tylovariabilis sp. nov. (MNRJ 7339) in SEM. A, preserved specimen; B, thick section showing the ectosome and choanosome; C, size variation of tylostyles; D, variation of tylostyle bases; E, sigmas; F, details of microspines on the tip of a sigma. Scale bars: A = 1 cm; B = 200 µm; C = 400 µm; D – E = 10 µm; F = 2 µm." httpUri="https://zenodo.org/record/239397/files/figure.png" pageId="7" pageNumber="371">Fig. 3</figureCitation>
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B).
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</emphasis>
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Ectosomal skeleton with plumose bundles of tylostyles. Choanosomal skeleton composed of a vaguely halichondrioid skeleton; poorly developed spongin fibres. Sigmas appear to be distributed randomly.
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</paragraph>
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<paragraph id="8BA4E74DFD6CFFEB9EEBFF0EFC87FE98" blockId="7.[151,1436,151,824]" pageId="7" pageNumber="371">
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||
<emphasis id="B96F3B5FFD6CFFEB9EEBFF0EFEFAFF20" bold="true" box="[199,450,223,248]" pageId="7" pageNumber="371">
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Spicules (
|
||
<figureCitation id="1320FBC8FD6CFFEB9F15FF0EFE47FF20" box="[313,383,223,248]" captionStart="FIGURE 3" captionStartId="6.[151,250,1663,1685]" captionTargetBox="[226,1359,215,1641]" captionTargetId="figure@6.[226,1360,193,1642]" captionTargetPageId="6" captionText="FIGURE 3. Desmacella tylovariabilis sp. nov. (MNRJ 7339) in SEM. A, preserved specimen; B, thick section showing the ectosome and choanosome; C, size variation of tylostyles; D, variation of tylostyle bases; E, sigmas; F, details of microspines on the tip of a sigma. Scale bars: A = 1 cm; B = 200 µm; C = 400 µm; D – E = 10 µm; F = 2 µm." httpUri="https://zenodo.org/record/239397/files/figure.png" pageId="7" pageNumber="371">Fig. 3</figureCitation>
|
||
C–F).
|
||
</emphasis>
|
||
Tylostyles (315–
|
||
<emphasis id="B96F3B5FFD6CFFEB9CABFF31FDFEFF20" bold="true" box="[647,710,224,248]" italics="true" pageId="7" pageNumber="371">616.0</emphasis>
|
||
–
|
||
<date id="FFA5C18DFD6CFFEB9CF9FF0EFC00FF20" box="[725,824,223,248]" pageId="7" pageNumber="371" value="1050-10-06">1050 x 6</date>
|
||
–
|
||
<emphasis id="B96F3B5FFD6CFFEB9D6BFF31FC4DFF20" bold="true" box="[839,885,224,248]" italics="true" pageId="7" pageNumber="371">11.0</emphasis>
|
||
–16 µm, length x width): long, thin and smooth, straight to slightly curved, rounded tyles situated at varied positions at their bases (
|
||
<figureCitation id="1320FBC8FD6CFFEB9A13FED5FBB9FEC4" box="[1087,1153,260,285]" captionStart="FIGURE 3" captionStartId="6.[151,250,1663,1685]" captionTargetBox="[226,1359,215,1641]" captionTargetId="figure@6.[226,1360,193,1642]" captionTargetPageId="6" captionText="FIGURE 3. Desmacella tylovariabilis sp. nov. (MNRJ 7339) in SEM. A, preserved specimen; B, thick section showing the ectosome and choanosome; C, size variation of tylostyles; D, variation of tylostyle bases; E, sigmas; F, details of microspines on the tip of a sigma. Scale bars: A = 1 cm; B = 200 µm; C = 400 µm; D – E = 10 µm; F = 2 µm." httpUri="https://zenodo.org/record/239397/files/figure.png" pageId="7" pageNumber="371">Fig. 3</figureCitation>
|
||
C, D); Sigmas (25–
|
||
<emphasis id="B96F3B5FFD6CFFEB9B71FED5FAB6FEC4" bold="true" box="[1373,1422,260,284]" italics="true" pageId="7" pageNumber="371">34.2</emphasis>
|
||
– 48 µm, chord length): thin, abundant and microspined ends (
|
||
<figureCitation id="1320FBC8FD6CFFEB9D11FEF6FC47FE98" box="[829,895,295,320]" captionStart="FIGURE 3" captionStartId="6.[151,250,1663,1685]" captionTargetBox="[226,1359,215,1641]" captionTargetId="figure@6.[226,1360,193,1642]" captionTargetPageId="6" captionText="FIGURE 3. Desmacella tylovariabilis sp. nov. (MNRJ 7339) in SEM. A, preserved specimen; B, thick section showing the ectosome and choanosome; C, size variation of tylostyles; D, variation of tylostyle bases; E, sigmas; F, details of microspines on the tip of a sigma. Scale bars: A = 1 cm; B = 200 µm; C = 400 µm; D – E = 10 µm; F = 2 µm." httpUri="https://zenodo.org/record/239397/files/figure.png" pageId="7" pageNumber="371">Fig. 3</figureCitation>
|
||
E, F).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301B4C6FD6CFFEB9EEBFE9DFB9FFEBD" box="[199,1191,332,357]" pageId="7" pageNumber="371" type="distribution">
|
||
<paragraph id="8BA4E74DFD6CFFEB9EEBFE9DFB9FFEBD" blockId="7.[151,1436,151,824]" box="[199,1191,332,357]" pageId="7" pageNumber="371">
|
||
<emphasis id="B96F3B5FFD6CFFEB9EEBFE9DFE84FEBD" bold="true" box="[199,444,332,357]" pageId="7" pageNumber="371">
|
||
Distribution (
|
||
<figureCitation id="1320FBC8FD6CFFEB9F44FE9DFE94FEBD" box="[360,428,332,357]" captionStart="FIGURE 1" captionStartId="1.[151,250,1980,2002]" captionTargetBox="[154,1417,303,1955]" captionTargetId="figure@1.[154,1433,303,1955]" captionTargetPageId="1" captionText="FIGURE 1. Location of the collection sites of Desmacella species from Brazil. (1) Desmacella microsigmata sp. nov. (Rio Grande do Norte State); (2) Desmacella tylovariabilis sp. nov. (Rio de Janeiro State); (3) Desmacella annexa (São Paulo State)." httpUri="https://zenodo.org/record/239395/files/figure.png" pageId="7" pageNumber="371">Fig. 1</figureCitation>
|
||
).
|
||
</emphasis>
|
||
<collectingCountry id="F30CA7DDFD6CFFEB9FE8FE9DFD35FEBD" box="[452,525,332,357]" name="Brazil" pageId="7" pageNumber="371">Brazil</collectingCountry>
|
||
: Southeastern Region: Rio de Janeiro State: Campos Basin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301B4C6FD6CFFEB9EEBFEBEFA65FE75" pageId="7" pageNumber="371" type="etymology">
|
||
<paragraph id="8BA4E74DFD6CFFEB9EEBFEBEFA65FE75" blockId="7.[151,1436,151,824]" pageId="7" pageNumber="371">
|
||
<emphasis id="B96F3B5FFD6CFFEB9EEBFEBEFE76FE50" bold="true" box="[199,334,367,392]" pageId="7" pageNumber="371">Etymology.</emphasis>
|
||
The specific epithet
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9C6DFEA1FDEEFE50" box="[577,726,368,392]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="tylovariabilis">
|
||
<emphasis id="B96F3B5FFD6CFFEB9C6DFEA1FDEEFE50" box="[577,726,368,392]" italics="true" pageId="7" pageNumber="371">tylovariabilis</emphasis>
|
||
</taxonomicName>
|
||
refers to the variation observed on the tyles of the tylostyles,
|
||
<emphasis id="B96F3B5FFD6CFFEB9EBBFE45FFFAFE74" box="[151,194,404,428]" italics="true" pageId="7" pageNumber="371">tylo</emphasis>
|
||
= from the Greek
|
||
<emphasis id="B96F3B5FFD6CFFEB9FBDFE45FE81FE74" box="[401,441,404,428]" italics="true" pageId="7" pageNumber="371">tyle</emphasis>
|
||
, meaning knob or lump, and
|
||
<emphasis id="B96F3B5FFD6CFFEB9D2FFE44FC70FE74" box="[771,840,405,428]" italics="true" pageId="7" pageNumber="371">varius</emphasis>
|
||
= from the Latin, meaning different or various.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301B4C6FD6CFFEB9EEBFE66FA2FFCE0" pageId="7" pageNumber="371" type="discussion">
|
||
<paragraph id="8BA4E74DFD6CFFEB9EEBFE66FA2FFCE0" blockId="7.[151,1436,151,824]" pageId="7" pageNumber="371">
|
||
<emphasis id="B96F3B5FFD6CFFEB9EEBFE66FE03FE08" bold="true" box="[199,315,439,464]" pageId="7" pageNumber="371">Remarks.</emphasis>
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9F6AFE69FD53FE08" box="[326,619,440,464]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="tylovariabilis" status="sp. nov.">
|
||
<emphasis id="B96F3B5FFD6CFFEB9F6AFE69FD53FE08" box="[326,619,440,464]" italics="true" pageId="7" pageNumber="371">Desmacella tylovariabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25C8624FD6CFFEB9C5AFE66FDECFE08" box="[630,724,439,464]" pageId="7" pageNumber="371" rank="species">
|
||
<emphasis id="B96F3B5FFD6CFFEB9C5AFE66FDECFE08" bold="true" box="[630,724,439,464]" pageId="7" pageNumber="371">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is assigned to the genus because of its tylostyles and sigma spicules, and vaguely halichondrioid skeleton formed by large tylostyles. There are four species similar to
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9BADFE0CFE15FDC0" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="tylovariabilis" status="sp. nov.">
|
||
<emphasis id="B96F3B5FFD6CFFEB9BADFE0CFE15FDC0" italics="true" pageId="7" pageNumber="371">D. tylovariabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25C8624FD6CFFEB9F19FE2EFEA9FDC0" box="[309,401,511,536]" pageId="7" pageNumber="371" rank="species">
|
||
<emphasis id="B96F3B5FFD6CFFEB9F19FE2EFEA9FDC0" bold="true" box="[309,401,511,536]" pageId="7" pageNumber="371">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
:
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9F8FFDD0FD08FDC0" box="[419,560,512,536]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="grimaldii">
|
||
<emphasis id="B96F3B5FFD6CFFEB9F8FFDD0FD08FDC0" box="[419,560,512,536]" italics="true" pageId="7" pageNumber="371">D. grimaldii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9C6CFDD0FDFAFDC0" box="[576,706,512,536]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="informis">
|
||
<emphasis id="B96F3B5FFD6CFFEB9C6CFDD0FDFAFDC0" box="[576,706,512,536]" italics="true" pageId="7" pageNumber="371">D. informis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9CFFFDD0FC60FDCF" box="[723,856,512,536]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="inornata">
|
||
<emphasis id="B96F3B5FFD6CFFEB9CFFFDD0FC60FDCF" box="[723,856,512,536]" italics="true" pageId="7" pageNumber="371">D. inornata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9DBFFDD0FB29FDCF" box="[915,1041,512,536]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="topsenti">
|
||
<emphasis id="B96F3B5FFD6CFFEB9DBFFDD0FB29FDCF" box="[915,1041,512,536]" italics="true" pageId="7" pageNumber="371">D. topsenti</emphasis>
|
||
</taxonomicName>
|
||
. They occur in the North Atlantic Ocean, and have similarities regarding spicule dimensions (see Table 1). However, the above species differ from
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9BADFDF4FE15FDB8" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="tylovariabilis" status="sp. nov.">
|
||
<emphasis id="B96F3B5FFD6CFFEB9BADFDF4FE15FDB8" italics="true" pageId="7" pageNumber="371">D. tylovariabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25C8624FD6CFFEB9F18FD96FEB7FDB8" box="[308,399,583,608]" pageId="7" pageNumber="371" rank="species">
|
||
<emphasis id="B96F3B5FFD6CFFEB9F18FD96FEB7FDB8" bold="true" box="[308,399,583,608]" pageId="7" pageNumber="371">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
which has rounded tyles situated at varied positions at their bases. The new species differs from
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9EFAFDBCFE74FD5C" box="[214,332,621,644]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="annexa">
|
||
<emphasis id="B96F3B5FFD6CFFEB9EFAFDBCFE74FD5C" box="[214,332,621,644]" italics="true" pageId="7" pageNumber="371">D. annexa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9F70FDBCFEEFFD5C" box="[348,471,620,644]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="digitata">
|
||
<emphasis id="B96F3B5FFD6CFFEB9F70FDBCFEEFFD5C" box="[348,471,620,644]" italics="true" pageId="7" pageNumber="371">D. digitata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9FCBFDBCFDADFD5C" box="[487,661,620,644]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="polysigmata">
|
||
<emphasis id="B96F3B5FFD6CFFEB9FCBFDBCFDADFD5C" box="[487,661,620,644]" italics="true" pageId="7" pageNumber="371">D. polysigmata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9C89FDBCFC26FD5C" box="[677,798,620,644]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="pumilio">
|
||
<emphasis id="B96F3B5FFD6CFFEB9C89FDBCFC26FD5C" box="[677,798,620,644]" italics="true" pageId="7" pageNumber="371">D. pumilio</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9D75FDBCFCF9FD5C" box="[857,961,621,644]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="vicina">
|
||
<emphasis id="B96F3B5FFD6CFFEB9D75FDBCFCF9FD5C" box="[857,961,621,644]" italics="true" pageId="7" pageNumber="371">D. vicina</emphasis>
|
||
</taxonomicName>
|
||
, by the presence of only one category of sigmas.
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9ED8FD41FD2EFD70" box="[244,534,656,680]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="tylovariabilis" status="sp. nov.">
|
||
<emphasis id="B96F3B5FFD6CFFEB9ED8FD41FD2EFD70" box="[244,534,656,680]" italics="true" pageId="7" pageNumber="371">Desmacella tylovariabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25C8624FD6CFFEB9C33FD5EFD43FD70" box="[543,635,655,680]" pageId="7" pageNumber="371" rank="species">
|
||
<emphasis id="B96F3B5FFD6CFFEB9C33FD5EFD43FD70" bold="true" box="[543,635,655,680]" pageId="7" pageNumber="371">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is distinguished from
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9D50FD40FB5BFD70" box="[892,1123,656,680]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="infundibuliformis">
|
||
<emphasis id="B96F3B5FFD6CFFEB9D50FD40FCA8FD70" box="[892,912,657,680]" italics="true" pageId="7" pageNumber="371">D</emphasis>
|
||
.
|
||
<emphasis id="B96F3B5FFD6CFFEB9D8CFD41FB5BFD70" box="[928,1123,656,680]" italics="true" pageId="7" pageNumber="371">infundibuliformis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9A5FFD40FBE8FD7F" box="[1139,1232,656,680]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="jania">
|
||
<emphasis id="B96F3B5FFD6CFFEB9A5FFD40FBBFFD70" box="[1139,1159,657,680]" italics="true" pageId="7" pageNumber="371">D</emphasis>
|
||
.
|
||
<emphasis id="B96F3B5FFD6CFFEB9ABBFD41FBE8FD7F" box="[1175,1232,656,679]" italics="true" pageId="7" pageNumber="371">jania</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9ACCFD40FA49FD70" box="[1248,1393,656,680]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="meliorata">
|
||
<emphasis id="B96F3B5FFD6CFFEB9ACCFD40FBCCFD70" box="[1248,1268,657,680]" italics="true" pageId="7" pageNumber="371">D</emphasis>
|
||
.
|
||
<emphasis id="B96F3B5FFD6CFFEB9B28FD41FA49FD70" box="[1284,1393,656,680]" italics="true" pageId="7" pageNumber="371">meliorata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9BADFD40FE2CFD14" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="vagabunda">
|
||
<emphasis id="B96F3B5FFD6CFFEB9BADFD40FAADFD70" box="[1409,1429,657,680]" italics="true" pageId="7" pageNumber="371">D</emphasis>
|
||
.
|
||
<emphasis id="B96F3B5FFD6CFFEB9EBBFD65FE2CFD14" box="[151,276,692,716]" italics="true" pageId="7" pageNumber="371">vagabunda</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9F7AFD64FD39FD14" box="[342,513,692,716]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="vestibularis">
|
||
<emphasis id="B96F3B5FFD6CFFEB9F7AFD64FE52FD14" box="[342,362,693,716]" italics="true" pageId="7" pageNumber="371">D</emphasis>
|
||
.
|
||
<emphasis id="B96F3B5FFD6CFFEB9F52FD65FD39FD14" box="[382,513,692,716]" italics="true" pageId="7" pageNumber="371">vestibularis</emphasis>
|
||
</taxonomicName>
|
||
, by the new species’ larger tylostyles and sigmas (see Table 1).
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9B34FD65FE3FFD28" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="corrugata">
|
||
<emphasis id="B96F3B5FFD6CFFEB9B34FD65FE3FFD28" italics="true" pageId="7" pageNumber="371">Desmacella corrugata</emphasis>
|
||
</taxonomicName>
|
||
differs from the new species by the different morphology of its megascleres. Additionally, the new species differs from
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9F53FD2CFD13FCCC" box="[383,555,764,788]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="suberitoides">
|
||
<emphasis id="B96F3B5FFD6CFFEB9F53FD2CFD13FCCC" box="[383,555,764,788]" italics="true" pageId="7" pageNumber="371">D. suberitoides</emphasis>
|
||
</taxonomicName>
|
||
as that species presents two categories of tylostyles. Finally,
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9ACFFD2CFAA3FCCC" box="[1251,1435,764,788]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="tylovariabilis" status="sp. nov.">
|
||
<emphasis id="B96F3B5FFD6CFFEB9ACFFD2CFAA3FCCC" box="[1251,1435,764,788]" italics="true" pageId="7" pageNumber="371">D. tylovariabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25C8624FD6CFFEB9EBBFCCEFFC8FCE0" box="[151,240,799,824]" pageId="7" pageNumber="371" rank="species">
|
||
<emphasis id="B96F3B5FFD6CFFEB9EBBFCCEFFC8FCE0" bold="true" box="[151,240,799,824]" pageId="7" pageNumber="371">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
has massive growth form and larger spicules than those of
|
||
<taxonomicName id="4C1B9CCEFD6CFFEB9DA5FCF0FB7CFCEF" box="[905,1092,800,824]" class="Demospongiae" family="Desmacellidae" genus="Desmacella" kingdom="Animalia" order="Poecilosclerida" pageId="7" pageNumber="371" phylum="Porifera" rank="species" species="microsigmata" status="sp. nov.">
|
||
<emphasis id="B96F3B5FFD6CFFEB9DA5FCF0FB7CFCEF" box="[905,1092,800,824]" italics="true" pageId="7" pageNumber="371">D. microsigmata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25C8624FD6CFFEB9A67FCCEFB9CFCE0" box="[1099,1188,799,824]" pageId="7" pageNumber="371" rank="species">
|
||
<emphasis id="B96F3B5FFD6CFFEB9A67FCCEFB9CFCE0" bold="true" box="[1099,1188,799,824]" pageId="7" pageNumber="371">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(Table 1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |