365 lines
56 KiB
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365 lines
56 KiB
XML
<document id="05E05CF683C3C9A6E851F37C650D6EE5" ID-DOI="10.5852/ejt.2019.526" ID-GBIF-Dataset="2da65c0a-9c48-43f5-ada3-e088de28dec1" ID-ISSN="2118-9773" ID-Zenodo-Dep="3152217" ID-ZooBank="3FFC527D-E9CA-4D9F-BF59-076FA60171AC" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1558535825802" checkinUser="plazi" docAuthor="Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard" docDate="2019" docId="03B1DB4C8B0B4E314C53FEF7381EFDDC" docLanguage="en" docName="ej-526_nurinsiyah_neiber_hausdorf.pdf.imf" docOrigin="European Journal of Taxonomy 526" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Landouria abdidalem Nurinsiyah & Neiber & Hausdorf 2019, sp. nov." docType="treatment" docUuid="B91C66B0-56B9-40EB-800E-4C321959C0B2" docUuidSource="ZooBank" docVersion="15" lastPageNumber="41" masterDocId="FF88A3348B2D4E194E35FFFC3C0DFFDB" masterDocTitle="Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java" masterLastPageNumber="73" masterPageNumber="1" pageNumber="39" updateTime="1716316918745" updateUser="carolina">
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<mods:title id="6BA2AE58BA237901D1D0C46037C777D1">Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java</mods:title>
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<mods:namePart id="87492417A3DA3FC82DE7F8E1C3E1EA16">Nurinsiyah, Ayu Savitri</mods:namePart>
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<mods:affiliation id="2E2E4817972B3185A0ADB5AB6EED6CC7">Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia.</mods:affiliation>
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<mods:nameIdentifier id="0729BD56A37F54CB2A508B0A13540AD7" type="email">ayu_nurinsiyah@yahoo.com & ayus002@lipi.go.id</mods:nameIdentifier>
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<mods:namePart id="2C770ADB76D7394A7DFB43F19EA9D2AB">Neiber, Marco T.</mods:namePart>
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<mods:affiliation id="C722D6D65A91C953E449B39BB930BFB8">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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<mods:nameIdentifier id="C897F5A54AB85D6402126880D57C6615" type="email">mneiber@hotmail.de</mods:nameIdentifier>
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<mods:namePart id="59E5326E881E9A6FF2A5C49C22E0AE91">Hausdorf, Bernhard</mods:namePart>
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<mods:affiliation id="1501FE1017D65728D5D504ACFFD81ED8">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany.</mods:affiliation>
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<mods:nameIdentifier id="4B7AC31FB3F0AEF31751947075426DCA" type="email">hausdorf@zoologie.uni-hamburg.de</mods:nameIdentifier>
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<mods:title id="B6BE53DB1A7ED591FD3A623B24CF6C11">European Journal of Taxonomy</mods:title>
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<mods:date id="95704ABFDCC9B588213DDCB44387E671">2019</mods:date>
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<mods:number id="BA686A0B8D1D1D9BB385C4410AED8D56">2019-05-21</mods:number>
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<mods:number id="43B76968890065596F1773A7633064F8">526</mods:number>
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<treatment id="03B1DB4C8B0B4E314C53FEF7381EFDDC" ID-DOI="http://doi.org/10.5281/zenodo.5587348" ID-GBIF-Taxon="157239598" ID-Zenodo-Dep="5587348" LSID="urn:lsid:zoobank.org:act:B91C66B0-56B9-40EB-800E-4C321959C0B2" httpUri="http://treatment.plazi.org/id/03B1DB4C8B0B4E314C53FEF7381EFDDC" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
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<subSubSection id="C30239D18B0B4E3F4C53FEF738CCFE92" pageId="38" pageNumber="39" type="nomenclature">
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<paragraph id="8BA76A5A8B0B4E3F4C53FEF73FC3FEFD" blockId="38.[370,1217,267,365]" box="[614,974,267,294]" pageId="38" pageNumber="39">
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<heading id="D0EFDD368B0B4E3F4C53FEF73FC3FEFD" box="[614,974,267,294]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="39" reason="2">
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<taxonomicName id="4C1811D98B0B4E3F4C53FEF73F66FEFD" ID-CoL="7RTH6" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[614,875,267,294]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="abdidalem" status="sp. nov.">
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<emphasis id="B96CB6488B0B4E3F4C53FEF73F66FEFD" bold="true" box="[614,875,267,294]" italics="true" pageId="38" pageNumber="39">Landouria abdidalem</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25F0B338B0B4E3F4D46FEF03FC3FEFD" box="[883,974,268,294]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BA76A5A8B0B4E3F4F47FED238CCFE92" blockId="38.[370,1217,267,365]" box="[370,1217,302,329]" pageId="38" pageNumber="39">
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urn:lsid:zoobank.org:act:
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<uuid id="FFBE508F8B0B4E3F4CA6FED238CCFE92" box="[659,1217,302,329]" pageId="38" pageNumber="39">B91C66B0-56B9-40EB-800E-4C321959C0B2</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30239D18B0B4E3F4C50FEAE3FC2FEB7" box="[613,975,338,365]" pageId="38" pageNumber="39" type="description">
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<paragraph id="8BA76A5A8B0B4E3F4C50FEAE3FC2FEB7" blockId="38.[370,1217,267,365]" box="[613,975,338,365]" pageId="38" pageNumber="39">
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<figureCitation id="132376DF8B0B4E3F4C50FEAE3EB2FEB7" box="[613,703,338,364]" captionStart="Figs 15–24" captionStartId="21.[189,241,1646,1672]" captionTargetBox="[206,1377,265,1616]" captionTargetId="figure@21.[204,1383,242,1625]" captionTargetPageId="21" captionText="Figs 15–24. Genitalia of species of Landouria from Java. 15. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 16. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 17. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 18. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 19. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 20. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 21. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 22. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 23. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 24. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152243/files/figure.png" pageId="38" pageNumber="39">Figs 21</figureCitation>
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,
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<figureCitation id="132376DF8B0B4E3F4CFEFEAE3EE6FEB6" box="[715,747,338,365]" captionStart="Figs 29–47" captionStartId="29.[189,241,1366,1392]" captionTargetBox="[189,1399,649,1337]" captionTargetId="figure@29.[189,1399,649,1337]" captionTargetPageId="29" captionText="Figs 29–47. Flagella of species of Landouria from Java. 29. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 30. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 31. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 32. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 33. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 34. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 35. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 36. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 37. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 38. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). 39. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 40. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 41. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 42. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 43. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 44. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 45. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 46. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 47. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 1 mm." httpUri="https://zenodo.org/record/3152255/files/figure.png" pageId="38" pageNumber="39">35</figureCitation>
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,
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<figureCitation id="132376DF8B0B4E3F4CC2FEAE3F1AFEB6" box="[759,791,338,365]" captionStart="Figs 52–55" captionStartId="33.[189,241,1717,1743]" captionTargetBox="[189,1398,265,1993]" captionTargetId="figure@34.[189,1398,265,1993]" captionTargetPageId="34" captionText="Figs 52–55 (opposite page). Distribution of species of Landouria in Java (10′ grid). 52. L. winteriana (Pfeiffer, 1842), L. naggsi sp. nov., L. nusakambangensis sp. nov., L. petrukensis sp. nov., L. tholiformis sp. nov., L. tonywhitteni sp. nov., L. smimensis (Mousson, 1848). 53. L. conoidea (Leschke, 1914), L. abdidalem sp. nov., L. sewuensis sp. nov., L. sukoliloensis sp. nov., L. pacitanensis sp. nov., L. intumescens (Martens, 1867), L. madurensis sp. nov. 54. L. rotatoria (Pfeiffer, 1842), L. nodifera sp. nov., L. parahyangensis sp. nov., L. schepmani (Möllendorff, 1897), L. zonifera sp. nov., L. ciliocincta (Möllendorff, 1897), L. pakidulan sp. nov. 55. L. epiplatia (Möllendorff, 1897), L. monticola van Benthem Jutting, 1950, L. davini Dharma, 2015, L. menorehensis sp. nov., L. dharmai sp. nov., L. moussoniana (Martens, 1867)." httpUri="https://zenodo.org/record/3152265/files/figure.png" pageId="38" pageNumber="39">53</figureCitation>
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,
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<figureCitation id="132376DF8B0B4E3F4D16FEAE3F49FEB6" box="[803,836,338,365]" captionStart="Figs 56–59" captionStartId="40.[189,241,1859,1885]" captionTargetBox="[189,1399,265,2004]" captionTargetId="figure@39.[189,1399,265,2004]" captionTargetPageId="39" captionText="Figs 56–59 (opposite page). Shells of species of Landouria from Java. 56. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 57. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (holotype MZB 21432). 58. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 59. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152267/files/figure.png" pageId="38" pageNumber="39">56</figureCitation>
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;
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<tableCitation id="C69A5FE18B0B4E3F4D7AFEAE3FC2FEB7" box="[847,975,338,364]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="38" pageNumber="39" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30239D18B0B4E3F4E88FE6F3D6DFDF9" pageId="38" pageNumber="39" type="diagnosis">
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<paragraph id="8BA76A5A8B0B4E3F4E88FE6F3D3BFE76" blockId="38.[189,310,403,429]" box="[189,310,403,429]" pageId="38" pageNumber="39">
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<heading id="D0EFDD368B0B4E3F4E88FE6F3D3BFE76" bold="true" box="[189,310,403,429]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
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<emphasis id="B96CB6488B0B4E3F4E88FE6F3D3BFE76" bold="true" box="[189,310,403,429]" pageId="38" pageNumber="39">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA76A5A8B0B4E3F4E88FE3E3D6DFDF9" blockId="38.[189,1399,450,546]" pageId="38" pageNumber="39">
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<taxonomicName id="4C1811D98B0B4E3F4E88FE3E3DB3FE07" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[189,446,450,476]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="abdidalem" status="sp. nov.">
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<emphasis id="B96CB6488B0B4E3F4E88FE3E3DB3FE07" box="[189,446,450,476]" italics="true" pageId="38" pageNumber="39">Landouria abdidalem</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25F0B338B0B4E3F4FF3FE3E3E2FFE07" box="[454,546,450,476]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
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is characterized by a sharply keeled shell, a long and narrow penis and epiphallus, and a crenated flagellum, which is shorter than the epiphallus and abruptly tapering at its proximal end.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30239D18B0B4E3F4E88FDB63DC7FD03" pageId="38" pageNumber="39" type="etymology">
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<paragraph id="8BA76A5A8B0B4E3F4E88FDB63D49FDBF" blockId="38.[189,324,586,612]" box="[189,324,586,612]" pageId="38" pageNumber="39">
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<heading id="D0EFDD368B0B4E3F4E88FDB63D49FDBF" bold="true" box="[189,324,586,612]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
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<emphasis id="B96CB6488B0B4E3F4E88FDB63D49FDBF" bold="true" box="[189,324,586,612]" pageId="38" pageNumber="39">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA76A5A8B0B4E3F4E88FD843DC7FD03" blockId="38.[189,1399,632,729]" pageId="38" pageNumber="39">
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The name of the species is derived from the Javanese expression ʻAbdi Dalemʼ (used as a noun in apposition). Abdi Dalem is a person who devotes himself to serve the Keraton of
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<collectingRegion id="49DCA4B88B0B4E3F4AA7FD673915FD6E" box="[1170,1304,667,693]" country="Indonesia" name="Yogyakarta" pageId="38" pageNumber="39">Yogyakarta</collectingRegion>
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(Yogya Palace) and the Sultan.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30239D18B0B4E3F4E88FCFC38E1FB30" pageId="38" pageNumber="39" type="materials_examined">
|
||
<paragraph id="8BA76A5A8B0B4E3F4E88FCFC3DA7FCC1" blockId="38.[189,426,768,794]" box="[189,426,768,794]" pageId="38" pageNumber="39">
|
||
<heading id="D0EFDD368B0B4E3F4E88FCFC3DA7FCC1" bold="true" box="[189,426,768,794]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
|
||
<emphasis id="B96CB6488B0B4E3F4E88FCFC3DA7FCC1" bold="true" box="[189,426,768,794]" pageId="38" pageNumber="39">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="3B7060078B0B4E3F4EDFFCD33D58FC69" ID-GBIF-Occurrence="2243137485" collectingDate="2017-08-25" collectionCode="MZB" collectorName="B. Hausdorf" country="Indonesia" county="Kulonprogo" elevation="700" latitude="-7.748056" location="Kiskendo Cave" longLatPrecision="21" longitude="110.13139" municipality="Jatimulyo" pageId="38" pageNumber="39" specimenCode="21271" specimenCount="1" stateProvince="Yogyakarta" typeStatus="holotype">
|
||
<paragraph id="8BA76A5A8B0B4E3F4EDFFCD33D51FC92" blockId="38.[189,1399,815,947]" box="[234,348,815,841]" pageId="38" pageNumber="39">
|
||
<typeStatus id="54A3D4F88B0B4E3F4EDFFCD33D51FC92" box="[234,348,815,841]" pageId="38" pageNumber="39" type="holotype">
|
||
<emphasis id="B96CB6488B0B4E3F4EDFFCD33D51FC92" bold="true" box="[234,348,815,841]" pageId="38" pageNumber="39">Holotype</emphasis>
|
||
</typeStatus>
|
||
</paragraph>
|
||
<paragraph id="8BA76A5A8B0B4E3F4E88FCAE3D58FC69" blockId="38.[189,1399,815,947]" pageId="38" pageNumber="39">
|
||
<collectingCountry id="F30F2ACA8B0B4E3F4E88FCAE3D56FCB7" box="[189,347,850,876]" name="Indonesia" pageId="38" pageNumber="39">INDONESIA</collectingCountry>
|
||
• holotype (det. anat.);
|
||
<collectingRegion id="49DCA4B88B0B4E3F4CABFCAE3F2AFCB7" box="[670,807,850,876]" country="Indonesia" name="Yogyakarta" pageId="38" pageNumber="39">Yogyakarta</collectingRegion>
|
||
,
|
||
<collectingCounty id="62C612D68B0B4E3F4D09FCAE3FC0FCB7" box="[828,973,850,876]" pageId="38" pageNumber="39">Kulonprogo</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BC3F0208B0B4E3F4DD7FCAE3854FCB7" box="[994,1113,850,876]" pageId="38" pageNumber="39">Jatimulyo</collectingMunicipality>
|
||
, near
|
||
<location id="8EC73C818B0B4E3F4A84FCAE397FFCB7" LSID="urn:lsid:plazi:treatment:03B1DB4C8B0B4E314C53FEF7381EFDDC:8EC73C818B0B4E3F4A84FCAE397FFCB7" box="[1201,1394,850,876]" country="Indonesia" county="Kulonprogo" latitude="-7.748056" longLatPrecision="21" longitude="110.13139" municipality="Jatimulyo" name="Kiskendo Cave" pageId="38" pageNumber="39" stateProvince="Yogyakarta">Kiskendo Cave</location>
|
||
;
|
||
<geoCoordinate id="EE2C0C9D8B0B4E3F4E88FC8F3D4DFC54" box="[189,320,883,911]" degrees="7" direction="south" minutes="44" orientation="latitude" pageId="38" pageNumber="39" precision="15" seconds="53" value="-7.748056">7°44′53″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2C0C9D8B0B4E3F4F7AFC8F3DFCFC54" box="[335,497,883,911]" degrees="110" direction="east" minutes="07" orientation="longitude" pageId="38" pageNumber="39" precision="15" seconds="53" value="110.13139">110°07′53″ E</geoCoordinate>
|
||
;
|
||
<elevation id="00358D698B0B4E3F4FCAFC893E80FC54" box="[511,653,885,911]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="38" pageNumber="39" unit="m" value="700.0">
|
||
<quantity id="4CE0C7BF8B0B4E3F4FCAFC893E46FC54" box="[511,587,885,911]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="38" pageNumber="39" unit="m" value="700.0">700 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<collectingDate id="EFE2B5728B0B4E3F4CA9FC893F4EFC54" box="[668,835,885,912]" pageId="38" pageNumber="39" value="2017-08-25">
|
||
<date id="FFA64C9A8B0B4E3F4CA9FC893F4EFC54" box="[668,835,885,912]" pageId="38" pageNumber="39" value="2017-08-25">25 Aug. 2017</date>
|
||
</collectingDate>
|
||
;
|
||
<collectorName id="26ED0F8C8B0B4E3F4D64FC893FEFFC54" box="[849,994,885,911]" pageId="38" pageNumber="39">B. Hausdorf</collectorName>
|
||
leg.; D =
|
||
<quantity id="4CE0C7BF8B0B4E3F4A55FC8938C0FC54" box="[1120,1229,885,911]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="38" pageNumber="39" unit="mm" value="13.4">13.4 mm</quantity>
|
||
, H =
|
||
<quantity id="4CE0C7BF8B0B4E3F4B26FC89397FFC54" box="[1299,1394,885,911]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="38" pageNumber="39" unit="mm" value="6.8">6.8 mm</quantity>
|
||
;
|
||
<collectionCode id="ED09F29F8B0B4E3F4E88FC653CF0FC68" box="[189,253,921,947]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="38" pageNumber="39" type="Museum">MZB</collectionCode>
|
||
<specimenCode id="DBBEC2218B0B4E3F4F31FC643D5CFC69" box="[260,337,920,946]" pageId="38" pageNumber="39">21271</specimenCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B7060078B0B4E3F4EDFFC263821FBE1" ID-GBIF-Occurrence="2243137531" collectionCode="MZB" country="Indonesia" county="Purworejo" elevation="700" latitude="-7.7644444" location="Seplawan Cave" longLatPrecision="21" longitude="110.096115" municipality="Donorejo" pageId="38" pageNumber="39" specimenCode="17733" specimenCount="1" stateProvince="Central Java" typeStatus="paratype">
|
||
<paragraph id="8BA76A5A8B0B4E3F4EDFFC263D6AFC2F" blockId="38.[189,1399,986,1259]" box="[234,359,986,1012]" pageId="38" pageNumber="39">
|
||
<typeStatus id="54A3D4F88B0B4E3F4EDFFC263D6AFC2F" box="[234,359,986,1012]" pageId="38" pageNumber="39" type="paratype">
|
||
<emphasis id="B96CB6488B0B4E3F4EDFFC263D6AFC2F" bold="true" box="[234,359,986,1012]" pageId="38" pageNumber="39">Paratypes</emphasis>
|
||
</typeStatus>
|
||
</paragraph>
|
||
<paragraph id="8BA76A5A8B0B4E3F4E88FC0138E1FB30" blockId="38.[189,1399,986,1259]" pageId="38" pageNumber="39">
|
||
<collectingCountry id="F30F2ACA8B0B4E3F4E88FC013D56FBCC" box="[189,347,1021,1047]" name="Indonesia" pageId="38" pageNumber="39">INDONESIA</collectingCountry>
|
||
–
|
||
<collectingRegion id="49DCA4B88B0B4E3F4F4FFC003E10FBCC" box="[378,541,1020,1047]" country="Indonesia" name="Jawa Tengah" pageId="38" pageNumber="39">
|
||
<emphasis id="B96CB6488B0B4E3F4F4FFC003E10FBCC" bold="true" box="[378,541,1020,1047]" pageId="38" pageNumber="39">Central Java</emphasis>
|
||
</collectingRegion>
|
||
•
|
||
<specimenCount id="9D1EA1D38B0B4E3F4C0DFC013E87FBCC" box="[568,650,1021,1047]" count="1" pageId="38" pageNumber="39" type="generic">1 spec.</specimenCount>
|
||
(det. anat.);
|
||
<collectingCounty id="62C612D68B0B4E3F4D14FC013F93FBCC" box="[801,926,1021,1047]" pageId="38" pageNumber="39">Purworejo</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BC3F0208B0B4E3F4D9EFC013810FBCC" box="[939,1053,1021,1047]" pageId="38" pageNumber="39">Donorejo</collectingMunicipality>
|
||
, near
|
||
<location id="8EC73C818B0B4E3F4A51FC013910FBCC" LSID="urn:lsid:plazi:treatment:03B1DB4C8B0B4E314C53FEF7381EFDDC:8EC73C818B0B4E3F4A51FC013910FBCC" box="[1124,1309,1021,1047]" country="Indonesia" county="Purworejo" latitude="-7.7644444" longLatPrecision="21" longitude="110.096115" municipality="Donorejo" name="Seplawan Cave" pageId="38" pageNumber="39" stateProvince="Central Java">Seplawan Cave</location>
|
||
,
|
||
<originalDetermination id="A806D2A68B0B4E3F4B1FFC013DA8FBE1" pageId="38" pageNumber="39">shrubs and limestone rock</originalDetermination>
|
||
;
|
||
<geoCoordinate id="EE2C0C9D8B0B4E3F4F81FBE23E3AFBE1" box="[436,567,1054,1082]" degrees="7" direction="south" minutes="45" orientation="latitude" pageId="38" pageNumber="39" precision="15" seconds="52" value="-7.7644444">7°45′52″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2C0C9D8B0B4E3F4C72FBE23EE7FBE1" box="[583,746,1054,1082]" degrees="110" direction="east" minutes="05" orientation="longitude" pageId="38" pageNumber="39" precision="15" seconds="46" value="110.096115">110°05′46″ E</geoCoordinate>
|
||
;
|
||
<elevation id="00358D698B0B4E3F4CCCFBDC3F84FBE1" box="[761,905,1056,1082]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="38" pageNumber="39" unit="m" value="700.0">
|
||
<quantity id="4CE0C7BF8B0B4E3F4CCCFBDC3F4BFBE1" box="[761,838,1056,1082]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="38" pageNumber="39" unit="m" value="700.0">700 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<collectionCode id="ED09F29F8B0B4E3F4DACFBDC3FD4FBE1" box="[921,985,1056,1082]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="38" pageNumber="39" type="Museum">MZB</collectionCode>
|
||
<specimenCode id="DBBEC2218B0B4E3F4DD6FBDC3821FBE1" box="[995,1068,1056,1082]" pageId="38" pageNumber="39">17733</specimenCode>
|
||
•
|
||
<materialsCitation id="3B7060078B0B4E3F4A7FFBDC3DDEFB86" ID-GBIF-Occurrence="2243137346" collectionCode="MZB" country="Indonesia" county="Purworejo" elevation="700" latitude="-7.7644444" location="Seplawan Cave" longLatPrecision="21" longitude="110.096115" municipality="Donorejo" pageId="38" pageNumber="39" specimenCode="17732" specimenCount="3" stateProvince="Jawa Tengah" typeStatus="paratype">
|
||
<specimenCount id="9D1EA1D38B0B4E3F4A7FFBDC38AFFBE1" box="[1098,1186,1056,1082]" count="3" pageId="38" pageNumber="39" type="generic">3 spec.</specimenCount>
|
||
; same data as for preceding;
|
||
<collectionCode id="ED09F29F8B0B4E3F4F77FBB83D8FFB85" box="[322,386,1092,1118]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="38" pageNumber="39" type="Museum">MZB</collectionCode>
|
||
<specimenCode id="DBBEC2218B0B4E3F4FBFFBBF3DDEFB86" box="[394,467,1091,1117]" pageId="38" pageNumber="39">17732</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B7060078B0B4E3F4FDAFBBF3839FB86" ID-GBIF-Occurrence="2243137326" box="[495,1076,1091,1118]" collectionCode="MZB" country="Indonesia" county="Purworejo" elevation="700" latitude="-7.7644444" location="Seplawan Cave" longLatPrecision="21" longitude="110.096115" municipality="Donorejo" pageId="38" pageNumber="39" specimenCode="19314" specimenCount="5" stateProvince="Jawa Tengah" typeStatus="paratype">
|
||
<specimenCount id="9D1EA1D38B0B4E3F4FDAFBBF3E4BFB85" box="[495,582,1091,1118]" count="5" pageId="38" pageNumber="39" type="generic">5 spec.</specimenCount>
|
||
; same data as for preceding;
|
||
<collectionCode id="ED09F29F8B0B4E3F4D96FBB83FEEFB85" box="[931,995,1092,1118]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="38" pageNumber="39" type="Museum">MZB</collectionCode>
|
||
<specimenCode id="DBBEC2218B0B4E3F4DDEFBBF3839FB86" box="[1003,1076,1091,1117]" pageId="38" pageNumber="39">19314</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B7060078B0B4E3F4A65FBBF3DE9FB5B" ID-GBIF-Occurrence="2243137387" collectionCode="ZMH" country="Indonesia" county="Purworejo" elevation="700" latitude="-7.7644444" location="Seplawan Cave" longLatPrecision="21" longitude="110.096115" municipality="Donorejo" pageId="38" pageNumber="39" specimenCode="ZMH 133410" specimenCount="4" stateProvince="Jawa Tengah" typeStatus="paratype">
|
||
<specimenCount id="9D1EA1D38B0B4E3F4A65FBBF38AAFB85" box="[1104,1191,1091,1118]" count="4" pageId="38" pageNumber="39" type="generic">4 spec.</specimenCount>
|
||
; same data as for preceding;
|
||
<specimenCode id="DBBEC2218B0B4E3F4F77FB9B3DE9FB5B" box="[322,484,1126,1153]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="38" pageNumber="39" type="Museum">ZMH 133410</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B7060078B0B4E3F4C34FB9A3857FB5B" ID-GBIF-Occurrence="2243137442" box="[513,1114,1126,1153]" collectionCode="ZMH" country="Indonesia" county="Purworejo" elevation="700" latitude="-7.7644444" location="Seplawan Cave" longLatPrecision="21" longitude="110.096115" municipality="Donorejo" pageId="38" pageNumber="39" specimenCode="ZMH 133421" specimenCount="1" stateProvince="Jawa Tengah" typeStatus="paratype">
|
||
<specimenCount id="9D1EA1D38B0B4E3F4C34FB9A3E55FB5A" box="[513,600,1126,1153]" count="1" pageId="38" pageNumber="39" type="generic">1 spec.</specimenCount>
|
||
; same data as for preceding;
|
||
<specimenCode id="DBBEC2218B0B4E3F4D8DFB9B3857FB5B" box="[952,1114,1126,1153]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="38" pageNumber="39" type="Museum">ZMH 133421</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B7060078B0B4E3F4A42FB9A396EFB78" ID-GBIF-Occurrence="2243137490" collectionCode="ZMH" country="Indonesia" county="Kulonprogo" elevation="700" latitude="-7.748056" location="near Kiskendo Cave" longLatPrecision="21" longitude="110.13139" municipality="Jatimulyo" pageId="38" pageNumber="39" specimenCode="ZMH 140026" specimenCount="10" stateProvince="Yogyakarta" typeStatus="paratype">
|
||
<specimenCount id="9D1EA1D38B0B4E3F4A42FB9A38D1FB5A" box="[1143,1244,1126,1153]" count="10" pageId="38" pageNumber="39" type="generic">10 spec.</specimenCount>
|
||
;
|
||
<collectingRegion id="49DCA4B88B0B4E3F4ADCFB9A397FFB5B" box="[1257,1394,1126,1152]" country="Indonesia" name="Yogyakarta" pageId="38" pageNumber="39">Yogyakarta</collectingRegion>
|
||
,
|
||
<collectingCounty id="62C612D68B0B4E3F4E88FB753D43FB78" box="[189,334,1161,1187]" pageId="38" pageNumber="39">Kulonprogo</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BC3F0208B0B4E3F4F6EFB753DDFFB78" box="[347,466,1161,1187]" pageId="38" pageNumber="39">Jatimulyo</collectingMunicipality>
|
||
,
|
||
<location id="8EC73C818B0B4E3F4FEAFB763ED9FB7F" LSID="urn:lsid:plazi:treatment:03B1DB4C8B0B4E314C53FEF7381EFDDC:8EC73C818B0B4E3F4FEAFB763ED9FB7F" box="[479,724,1161,1188]" country="Indonesia" county="Kulonprogo" latitude="-7.748056" longLatPrecision="21" longitude="110.13139" municipality="Jatimulyo" name="near Kiskendo Cave" pageId="38" pageNumber="39" stateProvince="Yogyakarta">near Kiskendo Cave</location>
|
||
;
|
||
<geoCoordinate id="EE2C0C9D8B0B4E3F4CD7FB743F68FB7F" box="[738,869,1160,1188]" degrees="7" direction="south" minutes="44" orientation="latitude" pageId="38" pageNumber="39" precision="15" seconds="53" value="-7.748056">7°44′53″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2C0C9D8B0B4E3F4D41FB74381BFB7F" box="[884,1046,1160,1188]" degrees="110" direction="east" minutes="07" orientation="longitude" pageId="38" pageNumber="39" precision="15" seconds="53" value="110.13139">110°07′53″ E</geoCoordinate>
|
||
;
|
||
<elevation id="00358D698B0B4E3F4A11FB7638BFFB78" box="[1060,1202,1161,1188]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="38" pageNumber="39" unit="m" value="700.0">
|
||
<quantity id="4CE0C7BF8B0B4E3F4A11FB76387DFB7F" box="[1060,1136,1162,1188]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="38" pageNumber="39" unit="m" value="700.0">700 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<specimenCode id="DBBEC2218B0B4E3F4AF4FB76396EFB78" box="[1217,1379,1161,1188]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="38" pageNumber="39" type="Museum">ZMH 140026</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B7060078B0B4E3F4E88FB513E9CFB31" ID-GBIF-Occurrence="2243137457" collectedFrom="snakefruit plantation" collectionCode="MZB" country="Indonesia" elevation="730" latitude="-7.7438893" location="Sabrang Kidul" longLatPrecision="21" longitude="110.13194" municipality="Kulonprogo" pageId="38" pageNumber="39" specimenCode="17664" specimenCount="1" stateProvince="Yogyakarta" typeStatus="paratype">
|
||
<specimenCount id="9D1EA1D38B0B4E3F4E88FB513D15FB1C" box="[189,280,1197,1223]" count="1" pageId="38" pageNumber="39" type="generic">1 spec.</specimenCount>
|
||
(det. anat.);
|
||
<collectingRegion id="49DCA4B88B0B4E3F4FF3FB513E42FB1C" box="[454,591,1197,1223]" country="Indonesia" name="Yogyakarta" pageId="38" pageNumber="39">Yogyakarta</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BC3F0208B0B4E3F4C56FB513EF9FB1C" box="[611,756,1197,1223]" pageId="38" pageNumber="39">Kulonprogo</collectingMunicipality>
|
||
,
|
||
<location id="8EC73C818B0B4E3F4D3CFB513FB2FB1C" LSID="urn:lsid:plazi:treatment:03B1DB4C8B0B4E314C53FEF7381EFDDC:8EC73C818B0B4E3F4D3CFB513FB2FB1C" box="[777,959,1197,1223]" country="Indonesia" latitude="-7.7438893" longLatPrecision="21" longitude="110.13194" municipality="Kulonprogo" name="Sabrang Kidul" pageId="38" pageNumber="39" stateProvince="Yogyakarta">Sabrang Kidul</location>
|
||
,
|
||
<collectedFrom id="380429A78B0B4E3F4DE1FB5138DFFB1C" box="[980,1234,1197,1223]" pageId="38" pageNumber="39">snakefruit plantation</collectedFrom>
|
||
;
|
||
<geoCoordinate id="EE2C0C9D8B0B4E3F4AD2FB57397CFB1C" box="[1255,1393,1195,1223]" degrees="7" direction="south" minutes="44" orientation="latitude" pageId="38" pageNumber="39" precision="15" seconds="38" value="-7.7438893">7°44′38″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2C0C9D8B0B4E3F4E88FB323D50FB31" box="[189,349,1230,1258]" degrees="110" direction="east" minutes="07" orientation="longitude" pageId="38" pageNumber="39" precision="15" seconds="55" value="110.13194">110°07′55″ E</geoCoordinate>
|
||
;
|
||
<elevation id="00358D698B0B4E3F4F5FFB2C3DF9FB31" box="[362,500,1232,1258]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="38" pageNumber="39" unit="m" value="730.0">
|
||
<quantity id="4CE0C7BF8B0B4E3F4F5FFB2C3DB9FB31" box="[362,436,1232,1258]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="38" pageNumber="39" unit="m" value="730.0">730 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<collectionCode id="ED09F29F8B0B4E3F4C34FB2C3E4CFB31" box="[513,577,1232,1258]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="38" pageNumber="39" type="Museum">MZB</collectionCode>
|
||
<specimenCode id="DBBEC2218B0B4E3F4C7DFB2C3E9CFB31" box="[584,657,1232,1258]" pageId="38" pageNumber="39">17664</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B7060078B0B4E3F4C9FFB2C38E5FB30" ID-GBIF-Occurrence="2243137393" box="[682,1256,1232,1259]" collectionCode="MZB" country="Indonesia" elevation="730" latitude="-7.7438893" location="Sabrang Kidul" longLatPrecision="21" longitude="110.13194" municipality="Kulonprogo" pageId="38" pageNumber="39" specimenCode="19253" specimenCount="7" stateProvince="Jawa Tengah" typeStatus="paratype">
|
||
<specimenCount id="9D1EA1D38B0B4E3F4C9FFB2C3EF2FB31" box="[682,767,1232,1258]" count="7" pageId="38" pageNumber="39" type="generic">7 spec.</specimenCount>
|
||
; same data as for preceding;
|
||
<collectionCode id="ED09F29F8B0B4E3F4A61FB2C3899FB31" box="[1108,1172,1232,1258]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="38" pageNumber="39" type="Museum">MZB</collectionCode>
|
||
<specimenCode id="DBBEC2218B0B4E3F4AAEFB2C38E5FB30" box="[1179,1256,1232,1259]" pageId="38" pageNumber="39">19253</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
</subSubSection>
|
||
<subSubSection id="C30239D18B0B4E3F4E88FAED3E20F899" pageId="38" pageNumber="39" type="description">
|
||
<paragraph id="8BA76A5A8B0B4E3F4E88FAED3D43FAF0" blockId="38.[189,334,1297,1323]" box="[189,334,1297,1323]" pageId="38" pageNumber="39">
|
||
<heading id="D0EFDD368B0B4E3F4E88FAED3D43FAF0" bold="true" box="[189,334,1297,1323]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
|
||
<emphasis id="B96CB6488B0B4E3F4E88FAED3D43FAF0" bold="true" box="[189,334,1297,1323]" pageId="38" pageNumber="39">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA76A5A8B0B4E3F4E88FABC38FDF9D1" blockId="38.[189,1399,1344,1546]" pageId="38" pageNumber="39">
|
||
SHELL (
|
||
<figureCitation id="132376DF8B0B4E3F4F26FABC3D7DFA80" box="[275,368,1344,1371]" captionStart="Figs 56–59" captionStartId="40.[189,241,1859,1885]" captionTargetBox="[189,1399,265,2004]" captionTargetId="figure@39.[189,1399,265,2004]" captionTargetPageId="39" captionText="Figs 56–59 (opposite page). Shells of species of Landouria from Java. 56. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 57. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (holotype MZB 21432). 58. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 59. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152267/files/figure.png" pageId="38" pageNumber="39">Fig. 56;</figureCitation>
|
||
<tableCitation id="C69A5FE18B0B4E3F4F42FABC3DF6FA81" box="[375,507,1344,1370]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="38" pageNumber="39" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
|
||
). Depressed conical, with 5.75–6 hardly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes sparse, mainly at periphery and at umbilical side; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture rounded rhombic; upper insertion of peristome descending; peristome expanded, reflexed and thickened; umbilicus hardly eccentric, wide, comprising 33–48% of shell diameter, hardly obscured by columellar edge.
|
||
</paragraph>
|
||
<paragraph id="8BA76A5A8B0B4E3F4E88F9CD3E20F899" blockId="38.[189,1399,1585,1858]" pageId="38" pageNumber="39">
|
||
GENITALIA (
|
||
<figureCitation id="132376DF8B0B4E3F4F0BF9CD3D9AF990" box="[318,407,1585,1611]" captionStart="Figs 15–24" captionStartId="21.[189,241,1646,1672]" captionTargetBox="[206,1377,265,1616]" captionTargetId="figure@21.[204,1383,242,1625]" captionTargetPageId="21" captionText="Figs 15–24. Genitalia of species of Landouria from Java. 15. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 16. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 17. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 18. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 19. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 20. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 21. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 22. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 23. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 24. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152243/files/figure.png" pageId="38" pageNumber="39">Figs 21</figureCitation>
|
||
,
|
||
<figureCitation id="132376DF8B0B4E3F4F94F9CD3DCCF997" box="[417,449,1585,1612]" captionStart="Figs 29–47" captionStartId="29.[189,241,1366,1392]" captionTargetBox="[189,1399,649,1337]" captionTargetId="figure@29.[189,1399,649,1337]" captionTargetPageId="29" captionText="Figs 29–47. Flagella of species of Landouria from Java. 29. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 30. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 31. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 32. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 33. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 34. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 35. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 36. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 37. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 38. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). 39. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 40. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 41. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 42. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 43. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 44. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 45. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 46. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 47. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 1 mm." httpUri="https://zenodo.org/record/3152255/files/figure.png" pageId="38" pageNumber="39">35</figureCitation>
|
||
;
|
||
<tableCitation id="C69A5FE18B0B4E3F4FF9F9CD3E29F990" box="[460,548,1585,1611]" captionStart="Table 4" captionStartId="22.[188,257,551,579]" captionTargetBox="[205,1929,661,1732]" captionTargetPageId="22" captionText="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="38" pageNumber="39" tableUuid="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Table 4</tableCitation>
|
||
). Atrium short; penis long, becoming slightly broader proximally, without penis coecum; membranaceous sheath around distal part of penis connected with the epiphallus by tissue; penial retractor runs from diaphragm to proximal part of cylindrical epiphallus; flagellum shorter than epiphallus, broadest in its middle part, without node, abruptly tapering towards its blunt proximal end, crenated; vas deferens narrow and long; vagina long; oviduct shorter, broad, proximally tapering; peduncle of bursa copulatrix subdivided into a broader, spindle-shaped distal part and a narrower, longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30239D18B0B4E314E88F8953D60FE54" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39" type="discussion">
|
||
<paragraph id="8BA76A5A8B0B4E3F4E88F8953D22F858" blockId="38.[189,303,1897,1923]" box="[189,303,1897,1923]" pageId="38" pageNumber="39">
|
||
<heading id="D0EFDD368B0B4E3F4E88F8953D22F858" bold="true" box="[189,303,1897,1923]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
|
||
<emphasis id="B96CB6488B0B4E3F4E88F8953D22F858" bold="true" box="[189,303,1897,1923]" pageId="38" pageNumber="39">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA76A5A8B0B4E314E88F8643D60FE54" blockId="38.[189,1398,1943,2005]" lastBlockId="40.[189,1399,267,399]" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
|
||
With regard to the shell,
|
||
<taxonomicName id="4C1811D98B0B4E3F4FEAF8653E77F869" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[479,634,1944,1970]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="abdidalem" status="sp. nov.">
|
||
<emphasis id="B96CB6488B0B4E3F4FEAF8653E77F869" box="[479,634,1944,1970]" italics="true" pageId="38" pageNumber="39">L. abdidalem</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B0B4E3F4CB4F8643ED6F869" box="[641,731,1944,1970]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
resembles
|
||
<taxonomicName id="4C1811D98B0B4E3F4D6AF8653FF3F869" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[863,1022,1944,1970]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="tholiformis" status="sp. nov.">
|
||
<emphasis id="B96CB6488B0B4E3F4D6AF8653FF3F869" box="[863,1022,1944,1970]" italics="true" pageId="38" pageNumber="39">L. tholiformis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B0B4E3F4A30F864386DF869" box="[1029,1120,1944,1970]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="132376DF8B0B4E3F4A5AF86438CBF869" box="[1135,1222,1943,1970]" captionStart="Figs 48–51" captionStartId="31.[189,241,1824,1850]" captionTargetBox="[280,1308,265,2019]" captionTargetId="figure@32.[280,1308,265,2019]" captionTargetPageId="32" captionText="Figs 48–51 (opposite page). Shells of species of Landouria from Java. 48. L. parahyangensis sp. nov., West Java, Karangnunggal, surroundings of Pasir Taraje Cave (holotype, MZB 20429). 49. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 50. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 51. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152259/files/figure.png" pageId="38" pageNumber="39">Fig. 51</figureCitation>
|
||
), with which it clusters in the molecular phylogeny (
|
||
<figureCitation id="132376DF8B0B4E3F4C5AF8473EB9F80E" box="[623,692,1979,2005]" captionStart="Fig" captionStartId="7.[189,232,1718,1744]" captionTargetBox="[189,1398,270,1663]" captionTargetId="figure@7.[189,1398,270,1663]" captionTargetPageId="7" captionText="Fig. 1. Bayesian 50% majority rule consensus tree based on mitochondrial 16S rDNA sequences of species of Landouria Godwin-Austen, 1918 with Mandarina suenoae Minato, 1978 and Euhadra peliomphala (Pfeiffer, 1850) as outgroups. All species of Landouria in this tree are from Java except L. omphalostoma Páll-Gergely & Hunyadi, 2013 from China and L. timorensis Köhler et al. in press, L. montana Köhler et al. in press and Landouria sp. nov. (= L. winteriana (non Pfeiffer, 1842) in Köhler et al. (in press)) from Timor. Values at the nodes represent Bayesian posterior probabilities (left) and maximum likelihood bootstrap values (right). Numbers after taxon names correspond to MZB and ZMH catalogue numbers, AM catalogue numbers or GenBank accession numbers (see Table 1)." httpUri="https://zenodo.org/record/3152219/files/figure.png" pageId="38" pageNumber="39">Fig. 1</figureCitation>
|
||
). For differences, see remarks under that species. The range of
|
||
<taxonomicName id="4C1811D98B054E314EE9FEF03D78FEFE" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[220,373,267,293]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="abdidalem" status="sp. nov.">
|
||
<emphasis id="B96CB6488B054E314EE9FEF03D78FEFE" box="[220,373,267,293]" italics="true" pageId="40" pageNumber="41">L. abdidalem</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B054E314F4EFEF03DD8FEFD" box="[379,469,268,294]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
overlaps with that of
|
||
<taxonomicName id="4C1811D98B054E314CFAFEF03F40FEFD" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[719,845,268,294]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="zonifera" status="sp. nov.">
|
||
<emphasis id="B96CB6488B054E314CFAFEF03F40FEFD" box="[719,845,268,294]" italics="true" pageId="40" pageNumber="41">L. zonifera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B054E314D61FEF03FA0FEFD" box="[852,941,268,294]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
For differences, see remarks under that species. Furthermore, the range of
|
||
<taxonomicName id="4C1811D98B054E314C63FED33EFCFE92" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[598,753,303,329]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="abdidalem" status="sp. nov.">
|
||
<emphasis id="B96CB6488B054E314C63FED33EFCFE92" box="[598,753,303,329]" italics="true" pageId="40" pageNumber="41">L. abdidalem</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B054E314CCCFED33F59FE92" box="[761,852,303,329]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
overlaps with that of
|
||
<taxonomicName id="4C1811D98B054E314A60FED33919FE92" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[1109,1300,303,329]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="menorehensis" status="sp. nov.">
|
||
<emphasis id="B96CB6488B054E314A60FED33919FE92" box="[1109,1300,303,329]" italics="true" pageId="40" pageNumber="41">L. menorehensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B054E314B29FED3397AFE92" box="[1308,1399,303,329]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="132376DF8B054E314EF3FEAE3D16FEB7" box="[198,283,338,364]" captionStart="Figs 85–88" captionStartId="62.[189,241,1859,1885]" captionTargetBox="[232,1355,265,1830]" captionTargetId="figure@62.[232,1355,265,1830]" captionTargetPageId="62" captionText="Figs 85–88. Shells of species of Landouria from Java. 85. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 86. L. dharmai sp. nov., East Java, Tulungagung, Popoh (holotype, MZB 21272). 87. L. davini Dharma, 2015, Central Java, Baturraden, Pancuran Tujuh, Gunung Slamet (holotype, MZB 17063). 88. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152308/files/figure.png" pageId="40" pageNumber="41">Fig. 88</figureCitation>
|
||
) from which it can easily be distinguished by the sharply keeled, depressed conical, brownishcorneous shell.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30239D18B054E314E88FE42381EFDDC" box="[189,1043,446,519]" pageId="40" pageNumber="41" type="distribution">
|
||
<paragraph id="8BA76A5A8B054E314E88FE423D5BFE03" blockId="40.[189,342,446,472]" box="[189,342,446,472]" pageId="40" pageNumber="41">
|
||
<heading id="D0EFDD368B054E314E88FE423D5BFE03" bold="true" box="[189,342,446,472]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
|
||
<emphasis id="B96CB6488B054E314E88FE423D5BFE03" bold="true" box="[189,342,446,472]" pageId="40" pageNumber="41">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA76A5A8B054E314E88FE10381EFDDC" blockId="40.[189,1043,492,519]" box="[189,1043,492,519]" pageId="40" pageNumber="41">
|
||
<taxonomicName id="4C1811D98B054E314E88FE103DB1FDDD" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[189,444,492,518]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="abdidalem" status="sp. nov.">
|
||
<emphasis id="B96CB6488B054E314E88FE103DB1FDDD" box="[189,444,492,518]" italics="true" pageId="40" pageNumber="41">Landouria abdidalem</emphasis>
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</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B054E314FF6FE113E13FDDC" box="[451,542,493,519]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is endemic to the Menoreh Hills (
|
||
<figureCitation id="132376DF8B054E314D9BFE10380BFDDC" box="[942,1030,492,519]" captionStart="Figs 52–55" captionStartId="33.[189,241,1717,1743]" captionTargetBox="[189,1398,265,1993]" captionTargetId="figure@34.[189,1398,265,1993]" captionTargetPageId="34" captionText="Figs 52–55 (opposite page). Distribution of species of Landouria in Java (10′ grid). 52. L. winteriana (Pfeiffer, 1842), L. naggsi sp. nov., L. nusakambangensis sp. nov., L. petrukensis sp. nov., L. tholiformis sp. nov., L. tonywhitteni sp. nov., L. smimensis (Mousson, 1848). 53. L. conoidea (Leschke, 1914), L. abdidalem sp. nov., L. sewuensis sp. nov., L. sukoliloensis sp. nov., L. pacitanensis sp. nov., L. intumescens (Martens, 1867), L. madurensis sp. nov. 54. L. rotatoria (Pfeiffer, 1842), L. nodifera sp. nov., L. parahyangensis sp. nov., L. schepmani (Möllendorff, 1897), L. zonifera sp. nov., L. ciliocincta (Möllendorff, 1897), L. pakidulan sp. nov. 55. L. epiplatia (Möllendorff, 1897), L. monticola van Benthem Jutting, 1950, L. davini Dharma, 2015, L. menorehensis sp. nov., L. dharmai sp. nov., L. moussoniana (Martens, 1867)." httpUri="https://zenodo.org/record/3152265/files/figure.png" pageId="40" pageNumber="41">Fig. 53</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |