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<document id="2E3EFCB98297882C2560AAC8576B34CB" ID-DOI="10.11646/zootaxa.3426.1.2" ID-GBIF-Dataset="36838477-e1c9-49ef-a984-9e9a60869c13" ID-ISSN="1175-5326" ID-Zenodo-Dep="214004" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460509283957" checkinUser="plazi" docAuthor="Jaitrong, Weeyawat &amp; Hashimoto, Yoshiaki" docDate="2012" docId="03B1C028FF96231B699D241FD18D6276" docLanguage="en" docName="zt03426p044.pdf" docOrigin="Zootaxa 3426 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3426.1.2" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Aenictus minutulus" docType="treatment" docVersion="4" lastPageNumber="31" masterDocId="FF88B850FF972319690A2262D4116553" masterDocTitle="Revision of the Aenictus minutulus species group (Hymenoptera: Formicidae: Aenictinae) from Southeast Asia" masterLastPageNumber="44" masterPageNumber="29" pageNumber="30" updateTime="1698313181006" updateUser="plazi">
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<mods:title id="6813E42273FCD646E80B4A82B5015111">Revision of the Aenictus minutulus species group (Hymenoptera: Formicidae: Aenictinae) from Southeast Asia</mods:title>
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<mods:namePart id="EC6A70AFB80753F26C1DFC0610760A10">Jaitrong, Weeyawat</mods:namePart>
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<mods:namePart id="F52FBAF13EDEA747727FA84299877E25">Hashimoto, Yoshiaki</mods:namePart>
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<treatment id="03B1C028FF96231B699D241FD18D6276" ID-DOI="http://doi.org/10.5281/zenodo.5878879" ID-GBIF-Taxon="192432079" ID-Zenodo-Dep="5878879" LSID="urn:lsid:plazi:treatment:03B1C028FF96231B699D241FD18D6276" httpUri="http://treatment.plazi.org/id/03B1C028FF96231B699D241FD18D6276" lastPageId="2" lastPageNumber="31" pageId="1" pageNumber="30">
<subSubSection id="C30222B5FF962318699D241FD62363C4" box="[151,562,1661,1687]" pageId="1" pageNumber="30" type="nomenclature">
<paragraph id="8BA7713EFF962318699D241FD62363C4" blockId="1.[151,562,1661,1687]" box="[151,562,1661,1687]" pageId="1" pageNumber="30">
<heading id="D0EFC652FF962318699D241FD62363C4" bold="true" box="[151,562,1661,1687]" fontSize="11" level="1" pageId="1" pageNumber="30" reason="1">
<emphasis id="B96CAD2CFF962318699D241FD62363C4" bold="true" box="[151,562,1661,1687]" pageId="1" pageNumber="30">
<taxonomicName id="4C180ABDFF962318699D241FD59563C4" ID-CoL="64XW4" box="[151,388,1661,1687]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="30" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis id="B96CAD2CFF962318699D241FD59563C4" bold="true" box="[151,388,1661,1687]" italics="true" pageId="1" pageNumber="30">Aenictus minutulus</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30222B5FF962318699D24A6D6AD638A" box="[151,700,1731,1754]" pageId="1" pageNumber="30" type="reference_group">
<paragraph id="8BA7713EFF962318699D24A6D6AD638A" blockId="1.[151,700,1731,1754]" box="[151,700,1731,1754]" pageId="1" pageNumber="30">
<taxonomicName id="4C180ABDFF962318699D24A6D5286389" ID-CoL="64XW6" box="[151,313,1732,1754]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="30" phylum="Arthropoda" rank="species" species="piercei">
<emphasis id="B96CAD2CFF962318699D24A6D5286389" box="[151,313,1732,1754]" italics="true" pageId="1" pageNumber="30">Aenictus piercei</emphasis>
</taxonomicName>
group in Jaitrong et Yamane, 2011: 9.
</paragraph>
</subSubSection>
<subSubSection id="C30222B5FF96231B699D2564D76F646C" lastPageId="2" lastPageNumber="31" pageId="1" pageNumber="30" type="diagnosis">
<paragraph id="8BA7713EFF96231B699D2564D4F7644F" blockId="1.[151,1436,1798,2002]" lastBlockId="2.[151,1436,151,825]" lastPageId="2" lastPageNumber="31" pageId="1" pageNumber="30">
<emphasis id="B96CAD2CFF962318699D2564D500624C" bold="true" box="[151,273,1798,1823]" pageId="1" pageNumber="30">Diagnosis.</emphasis>
In an earlier paper (Jaitrong &amp; Yamane 2011) this species group was defined as follows: head in fullface view with occipital corner convex, and posterior margin almost straight to shallowly and broadly concave; occipital margin lacking collar; antenna 10-segmented; antennal scape short reaching only midlength of head; anterior clypeal margin roundly convex, lacking denticles; mandible subtriangular; its masticatory margin with a large apical tooth, medium-sized subapical and basal teeth, and 2-6 denticles between them; basal margin of mandible with conspicuous denticles; frontal carina short; parafrontal ridge absent; with mesosoma in profile promesonotum convex dorsally and sloping gradually to propodeum; metapleural groove present or absent (mesonotum and propodeum fused); propodeal junction angulated; subpetiolar process well developed, triangular or subrectangular. Head and first gastral segment entirely smooth and shiny. Body yellowish brown to reddish brown; typhlatta spot absent.
</paragraph>
<paragraph id="8BA7713EFF95231B69C32345D76F646C" blockId="2.[151,1436,151,825]" box="[201,894,295,320]" pageId="2" pageNumber="31">Size variation occurs among individuals from single colonies.</paragraph>
</subSubSection>
<subSubSection id="C30222B5FF95231B69C3232ED103666A" pageId="2" pageNumber="31" type="discussion">
<paragraph id="8BA7713EFF95231B69C3232ED6C664FF" blockId="2.[151,1436,151,825]" pageId="2" pageNumber="31">
<emphasis id="B96CAD2CFF95231B69C3232ED52C6437" bold="true" box="[201,317,332,356]" pageId="2" pageNumber="31">Remarks.</emphasis>
This is a group of rather small species, measuring
<quantity id="4CE0DCDBFF95231B6A78232ED0016437" box="[882,1040,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.2" metricValueMin="1.8" pageId="2" pageNumber="31" unit="mm" value="2.5" valueMax="3.2" valueMin="1.8">1.80-3.20 mm</quantity>
in total body length. It is most similar to the
<taxonomicName id="4C180ABDFF95231B68022312D56F64DB" box="[264,382,368,392]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B96CAD2CFF95231B68022312D56F64DB" box="[264,382,368,392]" italics="true" pageId="2" pageNumber="31">A. javanus</emphasis>
</taxonomicName>
group in terms of body size and coloration, but in the former the anterior clypeal margin lacks denticles, while it has several denticles in the latter.
</paragraph>
<paragraph id="8BA7713EFF95231B69C323D5D0FC67D6" blockId="2.[151,1436,151,825]" pageId="2" pageNumber="31">
The
<taxonomicName id="4C180ABDFF95231B69F523DAD5A66483" box="[255,439,440,464]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="piercei">
<emphasis id="B96CAD2CFF95231B69F523DAD5A66483" box="[255,439,440,464]" italics="true" pageId="2" pageNumber="31">Aenictus piercei</emphasis>
</taxonomicName>
group was established by Jaitrong and Yamane (2011) to include five named species occurring in Southeast Asia:
<taxonomicName id="4C180ABDFF95231B68E323BED02664A7" authority="Terayama et Kubota, 1993" authorityName="Terayama et Kubota" authorityYear="1993" box="[489,1079,476,501]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="changmaianus">
<emphasis id="B96CAD2CFF95231B68E323BED6E664A7" box="[489,759,476,500]" italics="true" pageId="2" pageNumber="31">Aenictus changmaianus</emphasis>
<bibRefCitation id="EF890CCFFF95231B6A0823BFD02664A7" author="Terayama et Kubota" box="[770,1079,476,501]" firstAuthor="Terayama" pageId="2" pageNumber="31" pagination="68 - 72" refId="ref8681" refString="Terayama, M. &amp; Kubota, A. (1993) The army ant genus Aenictus (Hymenoptera: Formicidae) from Thailand and Vietnam, with descriptions of three new species. Bulletin of the Biogeographical Society of Japan, (2) 48, 68 - 72." type="journal article" year="1993">Terayama et Kubota, 1993</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C180ABDFF95231B6D4C23BED16464A7" authority="Terayama, 1984" authorityName="Terayama" authorityYear="1984" box="[1094,1397,476,501]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="lifuiae">
<emphasis id="B96CAD2CFF95231B6D4C23BED0A164A7" box="[1094,1200,476,500]" italics="true" pageId="2" pageNumber="31">A. lifuiae</emphasis>
Terayama, 1984
</taxonomicName>
;
<taxonomicName id="4C180ABDFF95231B6C8E23BED62E6744" authority="Terayama et Yamane, 1989" authorityName="Terayama et Yamane" authorityYear="1989" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis id="B96CAD2CFF95231B6C8E23BED517674B" italics="true" pageId="2" pageNumber="31">A. minutulus</emphasis>
<bibRefCitation id="EF890CCFFF95231B68062062D62E6744" author="Terayama et Yamane" box="[268,575,511,536]" firstAuthor="Terayama" pageId="2" pageNumber="31" pagination="597" refId="ref8732" refString="Terayama, M. &amp; Yamane, S. (1989) The army ant genus Aenictus (Hymenoptera: Formicinae) from Sumatra, with descriptions of three new species. Japanese Journal of Entomology, (3) 57, 597 - 603." type="journal article" year="1989">Terayama et Yamane, 1989</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C180ABDFF95231B6B402062D7616744" authority="Emery, 1894" authorityName="Emery" authorityYear="1894" box="[586,880,511,536]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="peguensis">
<emphasis id="B96CAD2CFF95231B6B402062D6C6674B" box="[586,727,512,536]" italics="true" pageId="2" pageNumber="31">A. peguensis</emphasis>
Emery, 1894
</taxonomicName>
; and
<taxonomicName id="4C180ABDFF95231B6AA12062D1406744" authority="Wheeler et Chapman, 1930" authorityName="Wheeler et Chapman" authorityYear="1930" box="[939,1361,511,536]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="piercei">
<emphasis id="B96CAD2CFF95231B6AA12062D007674B" box="[939,1046,512,536]" italics="true" pageId="2" pageNumber="31">A. piercei</emphasis>
Wheeler et Chapman, 1930
</taxonomicName>
. However, after carefully examining the
<typeStatus id="54A3CF9CFF95231B6B2D2047D646676E" box="[551,599,549,573]" pageId="2" pageNumber="31">type</typeStatus>
material and an associated specimen of
<taxonomicName id="4C180ABDFF95231B6D2C2046D085676F" box="[1062,1172,548,572]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="piercei">
<emphasis id="B96CAD2CFF95231B6D2C2046D085676F" box="[1062,1172,548,572]" italics="true" pageId="2" pageNumber="31">A. piercei</emphasis>
</taxonomicName>
we concluded that this species should be removed from this group and that it is a member of the
<taxonomicName id="4C180ABDFF95231B6AC6202AD0536733" box="[972,1090,584,608]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="B96CAD2CFF95231B6AC6202AD0536733" box="[972,1090,584,608]" italics="true" pageId="2" pageNumber="31">A. javanus</emphasis>
</taxonomicName>
group (
<taxonomicName id="4C180ABDFF95231B6D90202AD1146733" box="[1178,1285,584,608]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="piercei">
<emphasis id="B96CAD2CFF95231B6D90202AD1146733" box="[1178,1285,584,608]" italics="true" pageId="2" pageNumber="31">A. piercei</emphasis>
</taxonomicName>
has denticles on the anterior margin of clypeus). The
<taxonomicName id="4C180ABDFF95231B6B5E200ED6D067D7" box="[596,705,620,644]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="piercei">
<emphasis id="B96CAD2CFF95231B6B5E200ED6D067D7" box="[596,705,620,644]" italics="true" pageId="2" pageNumber="31">A. piercei</emphasis>
</taxonomicName>
group is renamed here as the
<taxonomicName id="4C180ABDFF95231B6D05200ED08F67D7" box="[1039,1182,620,644]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis id="B96CAD2CFF95231B6D05200ED08F67D7" box="[1039,1182,620,644]" italics="true" pageId="2" pageNumber="31">A. minutulus</emphasis>
</taxonomicName>
group.
</paragraph>
<paragraph id="8BA7713EFF95231B69C320EDD103666A" blockId="2.[151,1436,151,825]" pageId="2" pageNumber="31">
After a careful examination of the smaller specimens of
<taxonomicName id="4C180ABDFF95231B6A4820F2D7B867FB" box="[834,937,656,680]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="lifuiae">
<emphasis id="B96CAD2CFF95231B6A4820F2D7B867FB" box="[834,937,656,680]" italics="true" pageId="2" pageNumber="31">A. lifuiae</emphasis>
</taxonomicName>
collected from Iriomote-jima and Okinawajima, southern
<collectingCountry id="F30F31AEFF95231B683520D7D593679F" box="[319,386,693,716]" name="Japan" pageId="2" pageNumber="31">Japan</collectingCountry>
, we found that the mandible is almost linear, the anterior clypeal margin is straight or weakly concave, and that a gap occurs between the mandibles and anterior clypeal margin. These characteristics are used to separate the
<taxonomicName id="4C180ABDFF95231B684B209ED5C66647" box="[321,471,764,788]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B96CAD2CFF95231B684B209ED5C66647" box="[321,471,764,788]" italics="true" pageId="2" pageNumber="31">A. ceylonicus</emphasis>
</taxonomicName>
group from the other groups of the genus
<taxonomicName id="4C180ABDFF95231B6ABC209ED0076647" box="[950,1046,764,788]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B96CAD2CFF95231B6ABC209ED0076647" box="[950,1046,764,788]" italics="true" pageId="2" pageNumber="31">Aenictus</emphasis>
</taxonomicName>
, and also these specimens possess other conditions shared with the
<taxonomicName id="4C180ABDFF95231B6B0F2142D68B666B" box="[517,666,800,824]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B96CAD2CFF95231B6B0F2142D68B666B" box="[517,666,800,824]" italics="true" pageId="2" pageNumber="31">A. ceylonicus</emphasis>
</taxonomicName>
group. We transfer
<taxonomicName id="4C180ABDFF95231B6A732142D7F1666B" box="[889,992,800,824]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="lifuiae">
<emphasis id="B96CAD2CFF95231B6A732142D7F1666B" box="[889,992,800,824]" italics="true" pageId="2" pageNumber="31">A. lifuiae</emphasis>
</taxonomicName>
to the
<taxonomicName id="4C180ABDFF95231B6D262142D0D3666B" box="[1068,1218,800,824]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B96CAD2CFF95231B6D262142D0D3666B" box="[1068,1218,800,824]" italics="true" pageId="2" pageNumber="31">A. ceylonicus</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
<subSection id="E29768D5FF95231B699D21E9D58761CD" pageId="2" pageNumber="31" type="multiple">
<subSubSection id="C30222B5FF95231B699D21E9D59866F6" box="[151,393,907,933]" pageId="2" pageNumber="31" type="multiple">
<paragraph id="8BA7713EFF95231B699D21E9D59866F6" blockId="2.[151,393,907,933]" box="[151,393,907,933]" pageId="2" pageNumber="31">
<heading id="D0EFC652FF95231B699D21E9D59866F6" bold="true" box="[151,393,907,933]" fontSize="11" level="1" pageId="2" pageNumber="31" reason="1">
<emphasis id="B96CAD2CFF95231B699D21E9D59866F6" bold="true" box="[151,393,907,933]" pageId="2" pageNumber="31">Checklist of species</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30222B5FF95231B699D21B1D58761CD" pageId="2" pageNumber="31" type="synonymic_list">
<paragraph id="8BA7713EFF95231B699D21B1D6C066B9" blockId="2.[151,721,978,1183]" box="[151,721,978,1003]" pageId="2" pageNumber="31">
<taxonomicName id="4C180ABDFF95231B699D21B1D6C066B9" authority="Terayama et Kubota, 1993" authorityName="Terayama et Kubota" authorityYear="1993" box="[151,721,978,1003]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="changmaianus">
<emphasis id="B96CAD2CFF95231B699D21B1D5B366B8" box="[151,418,979,1003]" italics="true" pageId="2" pageNumber="31">Aenictus changmaianus</emphasis>
<bibRefCitation id="EF890CCFFF95231B68A321B1D6C066B9" author="Terayama et Kubota" box="[425,721,978,1003]" firstAuthor="Terayama" pageId="2" pageNumber="31" pagination="68 - 72" refId="ref8681" refString="Terayama, M. &amp; Kubota, A. (1993) The army ant genus Aenictus (Hymenoptera: Formicidae) from Thailand and Vietnam, with descriptions of three new species. Bulletin of the Biogeographical Society of Japan, (2) 48, 68 - 72." type="journal article" year="1993">Terayama et Kubota, 1993</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA7713EFF95231B699D2194D5AF615C" blockId="2.[151,721,978,1183]" box="[151,446,1014,1039]" pageId="2" pageNumber="31">
<taxonomicName id="4C180ABDFF95231B699D2194D54C615D" box="[151,349,1014,1038]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B96CAD2CFF95231B699D2194D54C615D" box="[151,349,1014,1038]" italics="true" pageId="2" pageNumber="31">Aenictus minimus</emphasis>
</taxonomicName>
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<emphasis id="B96CAD2CFF95231B686E2195D5AF615C" bold="true" box="[356,446,1015,1039]" pageId="2" pageNumber="31">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BA7713EFF95231B699D2679D6B56161" blockId="2.[151,721,978,1183]" box="[151,676,1050,1075]" pageId="2" pageNumber="31">
<taxonomicName id="4C180ABDFF95231B699D2679D6B56161" authority="Terayama et Yamane, 1989" authorityName="Terayama et Yamane" authorityYear="1989" box="[151,676,1050,1075]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis id="B96CAD2CFF95231B699D2679D57D6160" box="[151,364,1051,1075]" italics="true" pageId="2" pageNumber="31">Aenictus minutulus</emphasis>
<bibRefCitation id="EF890CCFFF95231B68792679D6B56161" author="Terayama et Yamane" box="[371,676,1050,1075]" firstAuthor="Terayama" pageId="2" pageNumber="31" pagination="597" refId="ref8732" refString="Terayama, M. &amp; Yamane, S. (1989) The army ant genus Aenictus (Hymenoptera: Formicinae) from Sumatra, with descriptions of three new species. Japanese Journal of Entomology, (3) 57, 597 - 603." type="journal article" year="1989">Terayama et Yamane, 1989</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA7713EFF95231B699D265CD6136105" blockId="2.[151,721,978,1183]" box="[151,514,1086,1111]" pageId="2" pageNumber="31">
<taxonomicName id="4C180ABDFF95231B699D265CD6136105" authority="Emery, 1895" authorityName="Emery" authorityYear="1895" box="[151,514,1086,1111]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="peguensis">
<emphasis id="B96CAD2CFF95231B699D265CD57D6105" box="[151,364,1086,1110]" italics="true" pageId="2" pageNumber="31">Aenictus peguensis</emphasis>
<bibRefCitation id="EF890CCFFF95231B687E265DD6136105" author="Emery" box="[372,514,1086,1111]" firstAuthor="Emery" pageId="2" pageNumber="31" pagination="450" refId="ref8238" refString="Emery, C. (1895) Viaggio di Leonardo Fea in Birmania e regioni vicine. 63. Formiche di Birmania, del Tenasserim e dei Monti Carin, raccolte da L. Fea. Annali del Museo Civico di Storia Naturale di Genova, (2) 14, 450 - 483." type="journal article" year="1895">Emery, 1895</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BA7713EFF95231B699D2601D5E06128" blockId="2.[151,721,978,1183]" box="[151,497,1123,1147]" pageId="2" pageNumber="31">
<taxonomicName id="4C180ABDFF95231B699D2601D5806128" box="[151,401,1123,1147]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="subterraneus" status="sp. nov.">
<emphasis id="B96CAD2CFF95231B699D2601D5806128" box="[151,401,1123,1147]" italics="true" pageId="2" pageNumber="31">Aenictus subterraneus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F1057FF95231B689D2601D5E06128" box="[407,497,1123,1147]" pageId="2" pageNumber="31" rank="species">
<emphasis id="B96CAD2CFF95231B689D2601D5E06128" bold="true" box="[407,497,1123,1147]" pageId="2" pageNumber="31">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BA7713EFF95231B699D26E4D58761CD" blockId="2.[151,721,978,1183]" box="[151,406,1158,1183]" pageId="2" pageNumber="31">
<taxonomicName id="4C180ABDFF95231B699D26E4D4E961CD" box="[151,248,1158,1182]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B96CAD2CFF95231B699D26E4D4E961CD" box="[151,248,1158,1182]" italics="true" pageId="2" pageNumber="31">Aenictus</emphasis>
</taxonomicName>
sp.56 of WJT
</paragraph>
</subSubSection>
</subSection>
<subSection id="E29768D5FF95231B699D2691D18D6276" pageId="2" pageNumber="31" type="key">
<subSubSection id="C30222B5FF95231B699D2691D66A6058" box="[151,635,1266,1291]" pageId="2" pageNumber="31" type="multiple">
<paragraph id="8BA7713EFF95231B699D2691D66A6058" blockId="2.[151,635,1266,1291]" box="[151,635,1266,1291]" pageId="2" pageNumber="31">
<heading id="D0EFC652FF95231B699D2691D66A6058" bold="true" box="[151,635,1266,1291]" fontSize="10" level="2" pageId="2" pageNumber="31" reason="2">
<emphasis id="B96CAD2CFF95231B699D2691D66A6058" bold="true" box="[151,635,1266,1291]" pageId="2" pageNumber="31">Key to species based on the worker caste</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30222B5FF95231B699D2758D18D6276" pageId="2" pageNumber="31" type="key">
<key id="FF89260FFF95231B699D2758D18D6276" pageId="2" pageNumber="31">
<keyStep id="30EC699BFF95231B699D2758D18D60D9" pageId="2" pageNumber="31">
<paragraph id="8BA7713EFF95231B699D2758D18D603F" blockId="2.[151,1437,1338,1829]" pageId="2" pageNumber="31">
<keyLead id="30E9D20BFF95231B699D2758D18D603F" pageId="2" pageNumber="31">
1. Promesonotum in profile with clearly convex dorsal outline; propodeum dorsally extremely lower than promesonotum (
<figureCitation id="13236DBBFF95231B6C7B2758D4FF603F" captionStart="FIGURE 2" captionStartId="5.[151,250,1545,1568]" captionTargetBox="[188,1398,193,1524]" captionTargetId="figure@5.[188,1398,193,1524]" captionTargetPageId="5" captionText="FIGURE 2. Aenictus sp. 56 of WJT (specimen from Borneo, FI 92 - 337). A, Head in full-face view; B, body in dorsal view; C, body in profile; D, declivity of propodeum, obliquely posterior view; E, propodeal junction, petiole and postpetiole in profile; F, mandible." httpUri="https://zenodo.org/record/214006/files/figure.png" pageId="2" pageNumber="31">Figs 2</figureCitation>
C, 5C, 6C).......................................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BA7713EFF95231B699D2717D18D60D9" blockId="2.[151,1437,1338,1829]" box="[151,1436,1397,1418]" pageId="2" pageNumber="31">
<keyLead id="30E9D20BFF95231B699D2717D18D60D9" box="[151,1436,1397,1418]" pageId="2" pageNumber="31">
- Mesosoma dorsally almost flat or feebly convex (
<figureCitation id="13236DBBFF95231B6BB62717D6E460D9" box="[700,757,1397,1418]" captionStart="FIGURE 1" captionStartId="4.[151,250,1795,1818]" captionTargetBox="[153,1433,193,1773]" captionTargetId="figure@4.[153,1434,193,1774]" captionTargetPageId="4" captionText="FIGURE 1. Aenictus changmaianus (A, B, C, paratype; D, E, non-type from Chiang Mai Province, N. Thailand). A, Head in full-face view; B, body in dorsal view; C, body in profile; D, propodeal junction, petiole and postpetiole in profile; E, mandible." httpUri="https://zenodo.org/record/214005/files/figure.png" pageId="2" pageNumber="31">Figs 1</figureCitation>
C, 3C, 4C).............................................. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EC699BFF95231B699D27F0D18D6097" pageId="2" pageNumber="31">
<paragraph id="8BA7713EFF95231B699D27F0D18D60F4" blockId="2.[151,1437,1338,1829]" box="[151,1436,1426,1447]" pageId="2" pageNumber="31">
<keyLead id="30E9D20BFF95231B699D27F0D18D60F4" box="[151,1436,1426,1447]" pageId="2" pageNumber="31">
2 Declivity of propodeum shallowly concave, encircled with a distinct rim (
<figureCitation id="13236DBBFF95231B6AAF27F0D7CC60F4" box="[933,989,1426,1447]" captionStart="FIGURE 2" captionStartId="5.[151,250,1545,1568]" captionTargetBox="[188,1398,193,1524]" captionTargetId="figure@5.[188,1398,193,1524]" captionTargetPageId="5" captionText="FIGURE 2. Aenictus sp. 56 of WJT (specimen from Borneo, FI 92 - 337). A, Head in full-face view; B, body in dorsal view; C, body in profile; D, declivity of propodeum, obliquely posterior view; E, propodeal junction, petiole and postpetiole in profile; F, mandible." httpUri="https://zenodo.org/record/214006/files/figure.png" pageId="2" pageNumber="31">Fig. 2</figureCitation>
CE) (Borneo and Java)...
<emphasis id="B96CAD2CFF95231B6DF327F0D11E60F4" box="[1273,1295,1426,1447]" italics="true" pageId="2" pageNumber="31">A.</emphasis>
sp. 56 of WJT
</keyLead>
</paragraph>
<paragraph id="8BA7713EFF95231B699D27CDD18D6097" blockId="2.[151,1437,1338,1829]" box="[151,1436,1455,1476]" pageId="2" pageNumber="31">
<keyLead id="30E9D20BFF95231B699D27CDD18D6097" box="[151,1436,1455,1476]" pageId="2" pageNumber="31">
<emphasis id="B96CAD2CFF95231B699D27CDD48E6097" bold="true" box="[151,159,1455,1476]" pageId="2" pageNumber="31">-</emphasis>
Declivity of propodeum with lateral carinae, but not demarcated basally by a transverse carina (
<figureCitation id="13236DBBFF95231B6D7C27CDD0A06097" box="[1142,1201,1455,1476]" captionStart="FIGURE 5" captionStartId="10.[151,250,1774,1797]" captionTargetBox="[153,1434,196,1749]" captionTargetId="figure@10.[153,1434,196,1753]" captionTargetPageId="10" captionText="FIGURE 5. Aenictus peguensis (A, C, lectotype; B, D, E, non-type from S. Vietnam). A, Head in full-face view; B, body in dorsal view; C, body in profile; D, declivity of propodeum, obliquely posterior view; E, propodeal junction, petiole and postpetiole in profile; F, mandible." httpUri="https://zenodo.org/record/214009/files/figure.png" pageId="2" pageNumber="31">Figs 5</figureCitation>
DE, 6BD)....... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EC699BFF95231B699D27AFD18A6369" pageId="2" pageNumber="31">
<paragraph id="8BA7713EFF95231B699D27AFD18A60AC" blockId="2.[151,1437,1338,1829]" pageId="2" pageNumber="31">
<keyLead id="30E9D20BFF95231B699D27AFD18A60AC" pageId="2" pageNumber="31">
3 In full-face view posterior margin of head convex (
<figureCitation id="13236DBBFF95231B6BDA27AFD71860B1" box="[720,777,1485,1506]" captionStart="FIGURE 5" captionStartId="10.[151,250,1774,1797]" captionTargetBox="[153,1434,196,1749]" captionTargetId="figure@10.[153,1434,196,1753]" captionTargetPageId="10" captionText="FIGURE 5. Aenictus peguensis (A, C, lectotype; B, D, E, non-type from S. Vietnam). A, Head in full-face view; B, body in dorsal view; C, body in profile; D, declivity of propodeum, obliquely posterior view; E, propodeal junction, petiole and postpetiole in profile; F, mandible." httpUri="https://zenodo.org/record/214009/files/figure.png" pageId="2" pageNumber="31">Fig. 5</figureCitation>
A); propodeal junction angulated, right-angled (
<figureCitation id="13236DBBFF95231B6DD327AFD10360B1" box="[1241,1298,1485,1506]" captionStart="FIGURE 5" captionStartId="10.[151,250,1774,1797]" captionTargetBox="[153,1434,196,1749]" captionTargetId="figure@10.[153,1434,196,1753]" captionTargetPageId="10" captionText="FIGURE 5. Aenictus peguensis (A, C, lectotype; B, D, E, non-type from S. Vietnam). A, Head in full-face view; B, body in dorsal view; C, body in profile; D, declivity of propodeum, obliquely posterior view; E, propodeal junction, petiole and postpetiole in profile; F, mandible." httpUri="https://zenodo.org/record/214009/files/figure.png" pageId="2" pageNumber="31">Fig. 5</figureCitation>
E); larger species (TL
<quantity id="4CE0DCDBFF95231B683D2788D5D460AC" box="[311,453,1514,1535]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.15" metricValueMin="2.95" pageId="2" pageNumber="31" unit="mm" value="3.05" valueMax="3.15" valueMin="2.95">2.953.15 mm</quantity>
; HW
<quantity id="4CE0DCDBFF95231B68F72788D69B60AC" box="[509,650,1514,1535]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="5.8" metricValueMin="5.2" pageId="2" pageNumber="31" unit="mm" value="0.55" valueMax="0.58" valueMin="0.52">0.520.58 mm</quantity>
) (
<collectingCountry id="F30F31AEFF95231B6B962788D6FE60AC" box="[668,751,1514,1535]" name="Vietnam" pageId="2" pageNumber="31">Vietnam</collectingCountry>
and
<collectingCountry id="F30F31AEFF95231B6A142788D76C60AC" box="[798,893,1514,1535]" name="Myanmar" pageId="2" pageNumber="31">Myanmar</collectingCountry>
)..................................
<taxonomicName id="4C180ABDFF95231B6C292788D18A60AC" ID-CoL="64Y7R" box="[1315,1435,1514,1535]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="peguensis">
<emphasis id="B96CAD2CFF95231B6C292788D18A60AC" box="[1315,1435,1514,1535]" italics="true" pageId="2" pageNumber="31">A. peguensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA7713EFF95231B699D2465D18A6369" blockId="2.[151,1437,1338,1829]" pageId="2" pageNumber="31">
<keyLead id="30E9D20BFF95231B699D2465D18A6369" pageId="2" pageNumber="31">
<emphasis id="B96CAD2CFF95231B699D2465D48E634F" bold="true" box="[151,159,1543,1564]" pageId="2" pageNumber="31">-</emphasis>
In full-face view posterior margin of head almost straight or feebly concave (
<figureCitation id="13236DBBFF95231B6ADA2465D01B634F" box="[976,1034,1543,1564]" captionStart="FIGURE 6" captionStartId="12.[151,250,1980,2003]" captionTargetBox="[190,1409,193,1944]" captionTargetId="figure@12.[190,1409,193,1946]" captionTargetPageId="12" captionText="FIGURE 6. Aenictus subterraneus sp. nov. (holotype). A, Head in full-face view; B, body in dorsal view; C, body in profile; D, propodeal junction, petiole and postpetiole in profile; E, mandible." httpUri="https://zenodo.org/record/214010/files/figure.png" pageId="2" pageNumber="31">Fig. 6</figureCitation>
A); propodeal junction roundly angulated (
<figureCitation id="13236DBBFF95231B69E22447D5336369" box="[232,290,1573,1594]" captionStart="FIGURE 6" captionStartId="12.[151,250,1980,2003]" captionTargetBox="[190,1409,193,1944]" captionTargetId="figure@12.[190,1409,193,1946]" captionTargetPageId="12" captionText="FIGURE 6. Aenictus subterraneus sp. nov. (holotype). A, Head in full-face view; B, body in dorsal view; C, body in profile; D, propodeal junction, petiole and postpetiole in profile; E, mandible." httpUri="https://zenodo.org/record/214010/files/figure.png" pageId="2" pageNumber="31">Fig. 6</figureCitation>
C); smaller species (TL
<quantity id="4CE0DCDBFF95231B6B032447D6876369" box="[521,662,1573,1594]" metricMagnitude="-3" metricUnit="m" metricValue="2.125" metricValueMax="2.15" metricValueMin="2.1" pageId="2" pageNumber="31" unit="mm" value="2.125" valueMax="2.15" valueMin="2.1">2.102.15 mm</quantity>
; HW
<quantity id="4CE0DCDBFF95231B6BC42447D74A6369" box="[718,859,1573,1594]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="2" pageNumber="31" unit="mm" value="0.49" valueMax="0.5" valueMin="0.48">0.480.50 mm</quantity>
) (Borneo)....................
<taxonomicName id="4C180ABDFF95231B6DBB2447D1586369" box="[1201,1353,1573,1594]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="subterraneus" status="sp. nov.">
<emphasis id="B96CAD2CFF95231B6DBB2447D1586369" box="[1201,1353,1573,1594]" italics="true" pageId="2" pageNumber="31">A. subterraneus</emphasis>
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<emphasis id="B96CAD2CFF95231B6C452447D18A6369" bold="true" box="[1359,1435,1573,1594]" pageId="2" pageNumber="31">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EC699BFF95231B699D2420D18D63FC" pageId="2" pageNumber="31">
<paragraph id="8BA7713EFF95231B699D2420D18A6327" blockId="2.[151,1437,1338,1829]" pageId="2" pageNumber="31">
<keyLead id="30E9D20BFF95231B699D2420D18A6327" pageId="2" pageNumber="31">
4 Basal margin of mandible with 12 small denticles next to large basal tooth and followed by a larger denticle (
<figureCitation id="13236DBBFF95231B6C0E2420D12C6304" box="[1284,1341,1602,1623]" captionStart="FIGURE 3" captionStartId="6.[151,250,1810,1833]" captionTargetBox="[185,1401,193,1788]" captionTargetId="figure@6.[185,1401,193,1789]" captionTargetPageId="6" captionText="FIGURE 3. Aenictus minimus sp. nov. (holotype). A, Head in full-face view; B, body in dorsal view; C, body in profile; D, propodeal junction, petiole and postpetiole in profile; E, mandible." httpUri="https://zenodo.org/record/214007/files/figure.png" pageId="2" pageNumber="31">Fig. 3</figureCitation>
E); antennal scape relatively short (SI 6364) (N.
<collectingCountry id="F30F31AEFF95231B6B62243DD6AA6327" box="[616,699,1631,1652]" name="Vietnam" pageId="2" pageNumber="31">Vietnam</collectingCountry>
).............................................
<taxonomicName id="4C180ABDFF95231B6DD72402D1586326" box="[1245,1353,1632,1653]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
<emphasis id="B96CAD2CFF95231B6DD72402D1586326" box="[1245,1353,1632,1653]" italics="true" pageId="2" pageNumber="31">A. minimus</emphasis>
</taxonomicName>
<emphasis id="B96CAD2CFF95231B6C45243DD18A6327" bold="true" box="[1359,1435,1631,1652]" pageId="2" pageNumber="31">
<taxonomicNameLabel id="A25F1057FF95231B6C45243DD18A6327" box="[1359,1435,1631,1652]" pageId="2" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BA7713EFF95231B699D241FD18D63FC" blockId="2.[151,1437,1338,1829]" pageId="2" pageNumber="31">
<keyLead id="30E9D20BFF95231B699D241FD18D63FC" pageId="2" pageNumber="31">
<emphasis id="B96CAD2CFF95231B699D241FD48E63C1" bold="true" box="[151,159,1661,1682]" pageId="2" pageNumber="31">-</emphasis>
Basal margin of mandible with 35 denticles that gradually reduce in size toward base of mandible (
<figureCitation id="13236DBBFF95231B6DB4241FD0EB63C1" box="[1214,1274,1661,1682]" captionStart="FIGURE 1" captionStartId="4.[151,250,1795,1818]" captionTargetBox="[153,1433,193,1773]" captionTargetId="figure@4.[153,1434,193,1774]" captionTargetPageId="4" captionText="FIGURE 1. Aenictus changmaianus (A, B, C, paratype; D, E, non-type from Chiang Mai Province, N. Thailand). A, Head in full-face view; B, body in dorsal view; C, body in profile; D, propodeal junction, petiole and postpetiole in profile; E, mandible." httpUri="https://zenodo.org/record/214005/files/figure.png" pageId="2" pageNumber="31">Figs 1</figureCitation>
E, 4E); antennal scape relatively long (SI 6772).......................................................................... 5
</keyLead>
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<keyStep id="30EC699BFF95231B699D24D5D18D6276" pageId="2" pageNumber="31">
<paragraph id="8BA7713EFF95231B699D24D5D18763B9" blockId="2.[151,1437,1338,1829]" pageId="2" pageNumber="31">
<keyLead id="30E9D20BFF95231B699D24D5D18763B9" pageId="2" pageNumber="31">
5 Metapleural gland bulla relatively large; distance between propodeal spiracle and metapleural gland bulla shorter than or almost as long as spiracular diameter (
<figureCitation id="13236DBBFF95231B6B5B24B7D69763B9" box="[593,646,1749,1770]" captionStart="FIGURE 1" captionStartId="4.[151,250,1795,1818]" captionTargetBox="[153,1433,193,1773]" captionTargetId="figure@4.[153,1434,193,1774]" captionTargetPageId="4" captionText="FIGURE 1. Aenictus changmaianus (A, B, C, paratype; D, E, non-type from Chiang Mai Province, N. Thailand). A, Head in full-face view; B, body in dorsal view; C, body in profile; D, propodeal junction, petiole and postpetiole in profile; E, mandible." httpUri="https://zenodo.org/record/214005/files/figure.png" pageId="2" pageNumber="31">Fig. 1</figureCitation>
D) (
<collectingCountry id="F30F31AEFF95231B6BA524B7D71363B9" box="[687,770,1749,1770]" name="Vietnam" pageId="2" pageNumber="31">Vietnam</collectingCountry>
and
<collectingCountry id="F30F31AEFF95231B6A3A24B7D79663B9" box="[816,903,1749,1770]" name="Thailand" pageId="2" pageNumber="31">Thailand</collectingCountry>
).............................
<taxonomicName id="4C180ABDFF95231B6DE524B7D18763B9" box="[1263,1430,1749,1770]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="changmaianus">
<emphasis id="B96CAD2CFF95231B6DE524B7D18763B9" box="[1263,1430,1749,1770]" italics="true" pageId="2" pageNumber="31">A. changmaianus</emphasis>
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<paragraph id="8BA7713EFF95231B699D2490D18D6276" blockId="2.[151,1437,1338,1829]" pageId="2" pageNumber="31">
<keyLead id="30E9D20BFF95231B699D2490D18D6276" pageId="2" pageNumber="31">
<emphasis id="B96CAD2CFF95231B699D2490D48E6254" bold="true" box="[151,159,1778,1799]" pageId="2" pageNumber="31">-</emphasis>
Metapleural gland bulla relatively small; distance between propodeal spiracle and metapleural gland bulla clearly longer than spiracular diameter (
<figureCitation id="13236DBBFF95231B68AC256DD5CE6277" box="[422,479,1807,1828]" captionStart="FIGURE 4" captionStartId="8.[151,250,1967,1990]" captionTargetBox="[184,1403,193,1912]" captionTargetId="figure@8.[184,1403,193,1914]" captionTargetPageId="8" captionText="FIGURE 4. Aenictus minutulus (A, B, C, E, paratype; D, non-type from W. Java). A, Head in full-face view; B, body in dorsal view; C, body in profile; D, propodeal junction, petiole and postpetiole in profile; E, mandible." httpUri="https://zenodo.org/record/214008/files/figure.png" pageId="2" pageNumber="31">Fig. 4</figureCitation>
D) (Sumatra and Java)..................................................
<taxonomicName id="4C180ABDFF95231B6C292572D18D6276" ID-CoL="64XW4" box="[1315,1436,1808,1829]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis id="B96CAD2CFF95231B6C292572D18D6276" box="[1315,1436,1808,1829]" italics="true" pageId="2" pageNumber="31">A. minutulus</emphasis>
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