treatments-xml/data/03/B1/9C/03B19C3FFFFA32491081FD00FECC8DB4.xml
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<document id="1A4DCAAC6FD1008AAAF175FC524FD5EF" ID-DOI="10.1649/0010-065x(2002)056[0123:paoang]2.0.co;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4891486" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622574605291" checkinUser="carolina" docAuthor="Ocampo, Federico C." docDate="2002" docId="03B19C3FFFFA32491081FD00FECC8DB4" docLanguage="en" docName="TheColeopBull.56.1.123-126.pdf.imf" docOrigin="The Coleopterists Bulletin 56 (1)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Procoilodes adrastus Ocampo 2002, new species" docType="treatment" docVersion="3" lastPageNumber="126" masterDocId="FF88E447FFF8324A112BFFDDFF9B8C12" masterDocTitle="Procoilodes adrastus Ocampo, A New Genus And Species Of Miocene Hybosorid In Amber From The Dominican Republic (Coleoptera: Scarabaeoidea: Hybosoridae)" masterLastPageNumber="126" masterPageNumber="123" pageNumber="125" updateTime="1698952667531" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C23FCDBD4C1EB8FE49C419E9DDA46B47">Procoilodes adrastus Ocampo, A New Genus And Species Of Miocene Hybosorid In Amber From The Dominican Republic (Coleoptera: Scarabaeoidea: Hybosoridae)</mods:title>
</mods:titleInfo>
<mods:name id="E2A34EAD967D52B15CEB8D641FBFD65B" type="personal">
<mods:role id="4D55FC7E2A17E0645B36FD6E1B505A8F">
<mods:roleTerm id="6FF1E5009297657A7E0D87B67B8C90FD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5E9202330273DDDD07FF8CE5043B4E03">Ocampo, Federico C.</mods:namePart>
<mods:affiliation id="BB47C7F2145A30FC7B3FCBB2C631E2AB">Systematics Research Collections, W 436 Nebraska Hall University of Nebraska State Museum Lincoln, NE 68588, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="7981D85000EA12720C0ECF6F08D4DDAA" type="email">focampo@unlserve.unl.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="202EAAB1F17B91321473FC17051707DE">text</mods:typeOfResource>
<mods:relatedItem id="FCA3AFD69A249E41D1F7E45CE3BA247B" type="host">
<mods:titleInfo id="2F7DC366AE4E24596146003B2DBA1165">
<mods:title id="9C2C89B3146E6205BEE14D12032F4060">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="8ACDAEBEF6C917FF8F4DDA44FA112607">
<mods:date id="3B71D9BBF04BC5F09467C7DE7BA327C5">2002</mods:date>
<mods:detail id="C11D8E989DDF9FDA00CAC6D0D47AC637" type="pubDate">
<mods:number id="3A01254BCDD11EC14D6BBCA9F0533DAE">2002-03-01</mods:number>
</mods:detail>
<mods:detail id="E1B374DF60230F65D0DEC6F3E8479AC8" type="volume">
<mods:number id="8982CE93ABDFCA221AE37C8B1EF94D47">56</mods:number>
</mods:detail>
<mods:detail id="BA206B7CE6CECF3F022FF5A4A3D797A2" type="issue">
<mods:number id="21327D7A9E275B496593D549FB91D600">1</mods:number>
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<mods:start id="A429D732570A70DBF9C1B9B48938D12A">123</mods:start>
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</mods:extent>
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<mods:classification id="4F45756322490D14C1D7FFE9DD115A1C">journal article</mods:classification>
<mods:identifier id="45D117CAAB17B1F9F4A34F862F28FAD8" type="DOI">10.1649/0010-065x(2002)056[0123:paoang]2.0.co;2</mods:identifier>
<mods:identifier id="83D24E0A2D888937B501767CF852E1D1" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="37021DB5DB8AC2AC292A4B01FCCB11AF" type="Zenodo-Dep">4891486</mods:identifier>
</mods:mods>
<treatment id="03B19C3FFFFA32491081FD00FECC8DB4" LSID="urn:lsid:plazi:treatment:03B19C3FFFFA32491081FD00FECC8DB4" httpUri="http://treatment.plazi.org/id/03B19C3FFFFA32491081FD00FECC8DB4" lastPageId="3" lastPageNumber="126" pageId="2" pageNumber="125">
<subSubSection id="C3027EA2FFFA32481081FD00FCE48EE0" box="[426,895,733,754]" pageId="2" pageNumber="125" type="nomenclature">
<paragraph id="8BA72D29FFFA32481081FD00FCE48EE0" blockId="2.[426,895,733,781]" box="[426,895,733,754]" pageId="2" pageNumber="125">
<heading id="D0EF9A45FFFA32481081FD00FCE48EE0" box="[426,895,733,754]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="125" reason="2">
<taxonomicName id="4C1856AAFFFA32481081FD00FD748EE0" ID-CoL="4MNXS" authority="Ocampo" authorityName="Ocampo" authorityYear="2002" box="[426,751,733,754]" class="Insecta" family="Hybosoridae" genus="Procoilodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="125" phylum="Arthropoda" rank="species" species="adrastus" status="sp. nov.">
<emphasis id="B96CF13BFFFA32481081FD00FD108EE0" box="[426,651,733,754]" italics="true" pageId="2" pageNumber="125">Procoilodes adrastus</emphasis>
Ocampo
</taxonomicName>
,
<emphasis id="B96CF13BFFFA324813D7FD00FCE48EE0" bold="true" box="[764,895,733,754]" pageId="2" pageNumber="125">
<taxonomicNameLabel id="A25F4C40FFFA324813D7FD00FCE48EE0" box="[764,895,733,754]" pageId="2" pageNumber="125" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3027EA2FFFA32481375FD25FD4F8F1F" box="[606,724,760,781]" pageId="2" pageNumber="125" type="description">
<paragraph id="8BA72D29FFFA32481375FD25FD4F8F1F" blockId="2.[426,895,733,781]" box="[606,724,760,781]" pageId="2" pageNumber="125">
(
<figureCitation id="132331ACFFFA3248134DFD25FD378F1F" box="[614,684,760,781]" captionStart="Fig" captionStartId="1.[269,303,1214,1233]" captionTargetBox="[283,1040,169,1192]" captionTargetId="figure-12@1.[283,1040,169,1193]" captionTargetPageId="1" captionText="Fig. 1. Holotype of Procoilodes adrastus embedded in amber, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4891488" httpUri="https://zenodo.org/record/4891488/files/figure.png" pageId="2" pageNumber="125">Figs. 1</figureCitation>
4)
</paragraph>
</subSubSection>
<subSubSection id="C3027EA2FFFA32481038FCFAFE568F9F" pageId="2" pageNumber="125" type="materials_examined">
<paragraph id="8BA72D29FFFA32481038FCFAFE568F9F" blockId="2.[249,1081,807,909]" pageId="2" pageNumber="125">
<emphasis id="B96CF13BFFFA32481038FCFAFE268F2E" bold="true" box="[275,445,807,828]" pageId="2" pageNumber="125">Type Material.</emphasis>
<materialsCitation id="3B702774FFFA324810EEFCFAFC818F45" collectionCode="AMNH" country="Dominican Republic" location="Republic" pageId="2" pageNumber="125" specimenCode="DR-10-136" specimenCount="1" typeStatus="holotype">
<typeStatus id="54A3938BFFFA324810EEFCFAFDBD8F2E" box="[453,550,807,828]" pageId="2" pageNumber="125" type="holotype">Holotype</typeStatus>
from
<collectingCountry id="F30F6DB9FFFA32481341FCFAFC9C8F2E" box="[618,775,807,828]" name="Dominican Republic" pageId="2" pageNumber="125">the Dominican</collectingCountry>
<location id="8EC77BF2FFFA32481224FCFAFCF68F2E" LSID="urn:lsid:plazi:treatment:03B19C3FFFFA32491081FD00FECC8DB4:8EC77BF2FFFA32481224FCFAFCF68F2E" box="[783,877,807,828]" country="Dominican Republic" name="Republic" pageId="2" pageNumber="125">Republic</location>
(specimen number
<collectionCode id="ED09B5ECFFFA324811D2FC9FFED28F45" box="[249,329,834,855]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="125">AMNH</collectionCode>
<specimenCode id="DBBE8552FFFA32481079FC9FFE4B8F45" box="[338,464,834,855]" collectionCode="DR-" pageId="2" pageNumber="125">DR­10136</specimenCode>
) embedded in Miocene amber
</materialsCitation>
.
<materialsCitation id="3B702774FFFA3248120EFC9FFE528F9F" country="Dominican Republic" county="New York" location="Amber Fossil Collection" municipality="American Museum of Natural History" pageId="2" pageNumber="125" specimenCount="1" typeStatus="holotype">
<typeStatus id="54A3938BFFFA3248120EFC9FFC1D8F45" box="[805,902,834,855]" pageId="2" pageNumber="125" type="holotype">Holotype</typeStatus>
deposited in the
<location id="8EC77BF2FFFA324811D2FC80FD9A8F60" LSID="urn:lsid:plazi:treatment:03B19C3FFFFA32491081FD00FECC8DB4:8EC77BF2FFFA324811D2FC80FD9A8F60" box="[249,513,861,882]" country="Dominican Republic" county="New York" municipality="American Museum of Natural History" name="Amber Fossil Collection" pageId="2" pageNumber="125">Amber Fossil Collection</location>
,
<location id="8EC77BF2FFFA32481327FC80FD748F60" LSID="urn:lsid:plazi:treatment:03B19C3FFFFA32491081FD00FECC8DB4:8EC77BF2FFFA32481327FC80FD748F60" box="[524,751,861,882]" country="Dominican Republic" county="New York" municipality="American Museum of Natural History" name="Dept. of Entomology" pageId="2" pageNumber="125">Dept. of Entomology</location>
,
<collectingMunicipality id="6BC3B753FFFA324813D0FC80FED08F9F" pageId="2" pageNumber="125">American Museum of Natural History</collectingMunicipality>
,
<collectingCounty id="62C655A5FFFA3248107CFCA5FE528F9F" box="[343,457,888,909]" pageId="2" pageNumber="125">New York</collectingCounty>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3027EA2FFFA3248103BFC7BFD13892F" pageId="2" pageNumber="125" type="description">
<paragraph id="8BA72D29FFFA3248103BFC7BFD13892F" blockId="2.[249,1083,934,1341]" pageId="2" pageNumber="125">
<emphasis id="B96CF13BFFFA3248103BFC7BFE6B8FAB" bold="true" box="[272,496,934,953]" pageId="2" pageNumber="125">
Description.
<typeStatus id="54A3938BFFFA324810BBFC7BFE778FAB" box="[400,492,934,953]" pageId="2" pageNumber="125" type="holotype">Holotype</typeStatus>
.
</emphasis>
Length
<quantity id="4CE080CCFFFA3248136BFC7BFD0C8FAB" box="[576,663,934,953]" metricMagnitude="-3" metricUnit="m" metricValue="4.73" pageId="2" pageNumber="125" unit="mm" value="4.73">4.73 mm</quantity>
; greatest width
<quantity id="4CE080CCFFFA32481207FC7BFC188FAB" box="[812,899,934,953]" metricMagnitude="-3" metricUnit="m" metricValue="2.76" pageId="2" pageNumber="125" unit="mm" value="2.76">2.76 mm</quantity>
. Color dark brown.
<emphasis id="B96CF13BFFFA324811D2FC63FEB08FC3" box="[249,299,958,977]" italics="true" pageId="2" pageNumber="125">Head</emphasis>
. Frons with surface heavily and finely tuberculate, vertex with 2 conical tubercles. Eye canthus elongate, rectangular, apex truncate. Frontoclypeal suture not visible. Clypeus with surface similar to that of frons; anterior margin slightly rounded, lateral margins arcuate and narrowly reflexed. Eyes subglobose. Mandibles large; external edge broadly rounded; dorsal surface rugose, concave; subapex with well­developed, dorsal tooth. Maxilla and galea with dense setae; maxillary palpus with 4 segments. Labium with sparse, long setae.
<emphasis id="B96CF13BFFFA3248109EFB8DFD898871" box="[437,530,1104,1123]" italics="true" pageId="2" pageNumber="125">Pronotum</emphasis>
. Surface weakly convex, heavily and finely tuberculate, slightly depressed on middle in anterior half. Edges of anterior angles, lateral margin, and sides of posterior margin smooth.
<taxonomicName id="4C1856AAFFFA32481374FB5DFD218881" box="[607,698,1152,1171]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="CoL" kingdom="Fungi" order="Corynexochida" pageId="2" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="B96CF13BFFFA32481374FB5DFD218881" box="[607,698,1152,1171]" italics="true" pageId="2" pageNumber="125">Scutellum</emphasis>
</taxonomicName>
. Shape subtriangular; apex acute.
<emphasis id="B96CF13BFFFA324812D1FB5DFBAF8881" box="[1018,1076,1152,1171]" italics="true" pageId="2" pageNumber="125">Elytra</emphasis>
. Surface shagreened, with 1 stria next to elytral suture. Humeral umbone elevated.
<emphasis id="B96CF13BFFFA32481523FB44FBAF88BE" box="[1032,1076,1177,1196]" italics="true" pageId="2" pageNumber="125">Legs</emphasis>
. Protibiae (Fig. 3) on external margin with 3 long, acute teeth and with 2 denticles between both apical and median and between median and basal teeth, and with 5 denticles behind basal tooth; apical spur not reaching apex of second tarsal segment. Protarsus with segments 14 subequal in length and each shorter than fifth, each with slender distal setae. Meso­ and metatarsi with segments 1 and 5 longer than any of segments 2 4, segments 14 with slender distal setae.
</paragraph>
</subSubSection>
<subSubSection id="C3027EA2FFFA32481038FA86FC4689B5" pageId="2" pageNumber="125" type="discussion">
<paragraph id="8BA72D29FFFA32481038FA86FC4689B5" blockId="2.[249,1081,1371,1556]" pageId="2" pageNumber="125">
<emphasis id="B96CF13BFFFA32481038FA86FE1B8962" bold="true" box="[275,384,1371,1392]" pageId="2" pageNumber="125">Remarks.</emphasis>
The tree that produced the resin that trapped this specimen is
<taxonomicName id="4C1856AAFFFA324811D2FAAAFD80899E" authority="Poinar" authorityName="Poinar" authorityYear="1991" box="[249,539,1399,1420]" class="Magnoliopsida" family="Fabaceae" genus="Hymenaea" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="125" phylum="Tracheophyta" rank="species" species="protera">
<emphasis id="B96CF13BFFFA324811D2FAAAFE5C899E" box="[249,455,1399,1420]" italics="true" pageId="2" pageNumber="125">Hymenaea protera</emphasis>
Poinar
</taxonomicName>
(Leguminosae, Caesalpinioidea). The amber in which it is embedded was dated as Miocene in age (Grimaldi 1995).
</paragraph>
</subSubSection>
<subSubSection id="C3027EA2FFFA32491038FA70FECC8DB4" lastPageId="3" lastPageNumber="126" pageId="2" pageNumber="125" type="etymology">
<paragraph id="8BA72D29FFFA32481038FA70FDD08A06" blockId="2.[249,1081,1371,1556]" pageId="2" pageNumber="125">
<emphasis id="B96CF13BFFFA32481038FA70FE0889D0" bold="true" box="[275,403,1453,1474]" pageId="2" pageNumber="125">Etymology.</emphasis>
From the classical mythological name
<emphasis id="B96CF13BFFFA3248125DFA70FC4889D0" box="[886,979,1453,1474]" italics="true" pageId="2" pageNumber="125">Adrastus</emphasis>
, which means he who holds his position or the unescapable, a name perfectly applied to this specimen which has been embedded and immovable in amber for more than 15 million years.
</paragraph>
<paragraph id="8BA72D29FFFB324911DDFFB6FC198C6C" blockId="3.[246,898,107,126]" box="[246,898,107,126]" pageId="3" pageNumber="126">126 THE COLEOPTERISTS BULLETIN 56(1), 2002</paragraph>
<paragraph id="8BA72D29FFFB3249103BFF0AFBAD8CFE" blockId="3.[272,1078,215,236]" box="[272,1078,215,236]" pageId="3" pageNumber="126">
<materialsCitation id="3B702774FFFB3249103BFF0AFBAD8CFE" box="[272,1078,215,236]" collectorName="Natural History, NY" location="American Museum of Natural History" pageId="3" pageNumber="126" specimenCount="1" stateProvince="Saint David">
I thank
<collectingRegion id="49DCE3CBFFFB32491041FF0AFE318CFE" box="[362,426,215,236]" country="Saint Vincent and the Grenadines" name="Saint David" pageId="3" pageNumber="126">David</collectingRegion>
Grimaldi (
<location id="8EC77BF2FFFB3249130DFF0AFC548CFE" LSID="urn:lsid:plazi:treatment:03B19C3FFFFA32491081FD00FECC8DB4:8EC77BF2FFFB3249130DFF0AFC548CFE" box="[550,975,215,236]" name="American Museum of Natural History" pageId="3" pageNumber="126" stateProvince="Saint David">
American Museum of
<collectorName id="26ED48FFFFFB32491208FF0AFC548CFE" box="[803,975,215,236]" pageId="3" pageNumber="126">Natural History</collectorName>
</location>
, NY) for
</materialsCitation>
</paragraph>
<paragraph id="8BA72D29FFFB324911DDFF2CFECC8DB4" blockId="3.[246,1078,241,422]" pageId="3" pageNumber="126">
making the specimen available to me for study. I am grateful to Brett Ratcliffe and Mary Liz Jameson (University of
<collectingRegion id="49DCE3CBFFFB324913BDFED1FD608D33" box="[662,763,268,289]" country="United States of America" name="Nebraska" pageId="3" pageNumber="126">Nebraska</collectingRegion>
, Lincoln, NE) for their guidance in preparing the manuscript. Two anonymous reviewers are thanked for their helpful remarks to improve the manuscript. This project was supported by an NSF/ Biotic Survey and Inventory Grant (DEB 9870202) to B. C. Ratcliffe and an NSF/ PEET grant (DEB 9712447) to B. C. Ratcliffe and M. L. Jameson.
</paragraph>
</subSubSection>
</treatment>
</document>