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<document id="DE44B2DA3FD076864ABB10C6574ED96C" ID-CLB-Dataset="25295" ID-DOI="10.5852/ejt.2019.558" ID-GBIF-Dataset="5eadba67-d4ee-4eb8-8ec9-bdb75cab1d59" ID-Zenodo-Dep="3474974" ID-ZooBank="5B9DC474-1F26-4732-B7B8-32C285FE0099" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570442388906" checkinUser="plazi" docAuthor="Fehér, Zoltán, Deli, Tamás, Erőss, Zoltán P. &amp; Lika, Romilda" docDate="2019" docId="03B17A5AF71CFC52FEF4F35A6BC5F97D" docLanguage="en" docName="ejt-558_feher_deli_eross.pdf.imf" docOrigin="European Journal of Taxonomy 558" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Virpazaria (Virpazaria) pageti subsp. kleteckii Stamol &amp; Subai in Reischutz 2012" docType="treatment" docVersion="13" lastPageNumber="16" masterDocId="FF880222F712FC5DFFCFF2516A65FFED" masterDocTitle="Taxonomic revision of the subterranean genus Virpazaria Gittenberger, 1969 (Gastropoda, Spelaeodiscidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="15" updateTime="1698757969035" updateUser="ExternalLinkService">
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<mods:title id="8EC8ABFD8F1E40B11904F87D8FD4E8A1">Taxonomic revision of the subterranean genus Virpazaria Gittenberger, 1969 (Gastropoda, Spelaeodiscidae)</mods:title>
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<mods:namePart id="01E32FCEE9E0251CC428A8A14DE0AE2A">Fehér, Zoltán</mods:namePart>
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<mods:namePart id="9F31F2B82233C6B27C52DDB25E1BF914">Deli, Tamás</mods:namePart>
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<mods:namePart id="67DD8DCBE9E6DE962FF0B432EE5C9E1F">Erőss, Zoltán P.</mods:namePart>
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<mods:namePart id="691D0535DC84B1C885ADAA877942DEE2">Lika, Romilda</mods:namePart>
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<mods:title id="37CBC4D1357E5D0137A594A8750251CE">European Journal of Taxonomy</mods:title>
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<mods:date id="A930380630A9A6F81B2C22FB500AEE94">2019</mods:date>
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<treatment id="03B17A5AF71CFC52FEF4F35A6BC5F97D" ID-DOI="http://doi.org/10.5281/zenodo.3475257" ID-GBIF-Taxon="159654643" ID-Zenodo-Dep="3475257" LSID="urn:lsid:plazi:treatment:03B17A5AF71CFC52FEF4F35A6BC5F97D" httpUri="http://treatment.plazi.org/id/03B17A5AF71CFC52FEF4F35A6BC5F97D" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<subSubSection id="C30298C7F71CFC53FEF4F35A6E9CFEC8" box="[315,1273,266,294]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BA7CB4CF71CFC53FEF4F35A6E9CFEC8" blockId="14.[315,1273,266,294]" box="[315,1273,266,294]" pageId="14" pageNumber="15">
<heading id="D0EF7C20F71CFC53FEF4F35A6E9CFEC8" box="[315,1273,266,294]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
<taxonomicName id="4C18B0CFF71CFC53FEF4F35A6E9CFEC8" ID-CoL="7J5NH" authority="Stamol &amp; Subai in Reischutz et al., 2012" authorityName="Stamol &amp; Subai in Reischutz" authorityYear="2012" box="[315,1273,266,294]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="subSpecies" species="pageti" subGenus="Virpazaria" subSpecies="kleteckii">
<emphasis id="B96C175EF71CFC53FEF4F35A6974FECB" bold="true" box="[315,785,267,294]" italics="true" pageId="14" pageNumber="15">Virpazaria (Virpazaria) pageti kleteckii</emphasis>
Štamol &amp; Subai in
<bibRefCitation id="EF89B6BDF71CFC53FC36F35A6E9CFEC8" author="Reischutz A. &amp; Reischutz P. L." box="[1017,1273,267,294]" etAl="et al." firstAuthor="Reischutz" journalOrPublisher="Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft" pageId="14" pageNumber="15" pagination="17 - 18" part="19" refId="ref13210" refString="Reischutz A. &amp; Reischutz P. L. 2012. Virpazaria nicoleae nov. spec. (Spelaeodiscidae: Pulmonata: Gastropoda) aus Montenegro. Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft 19: 17 - 18." title="Virpazaria nicoleae nov. spec. (Spelaeodiscidae: Pulmonata: Gastropoda) aus Montenegro" type="journal article" year="2012">
Reischütz
<emphasis id="B96C175EF71CFC53FBBBF35D6ECAFEC8" box="[1140,1199,267,293]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
, 2012
</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30298C7F71CFC53FF72F31F6E3FFE8A" box="[189,1114,331,359]" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8BA7CB4CF71CFC53FF72F31F6E3FFE8A" blockId="14.[189,1114,331,359]" box="[189,1114,331,359]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="AB08EC62F71CFC53FF72F31F6E3FFE8A" box="[189,1114,331,359]" pageId="14" pageNumber="15">
<taxonomicName id="4C18B0CFF71CFC53FF72F31F6E6FFE8A" ID-CoL="7J5NH" authority="Stamol &amp; Subai in Reischutz et al., 2012: 32" authorityName="Stamol &amp; Subai in Reischutz" authorityPageNumber="32" authorityYear="2012" box="[189,1034,331,359]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="subSpecies" species="pageti" subSpecies="kleteckii">
<emphasis id="B96C175EF71CFC53FF72F31F6B91FE8A" box="[189,500,333,359]" italics="true" pageId="14" pageNumber="15">Virpazaria pageti kleteckii</emphasis>
Štamol &amp; Subai in
<treatmentCitation id="0AB9ED5DF71CFC53FD13F31C6E6FFE8A" author="Reischutz A. &amp; Reischutz P. L." box="[732,1034,333,359]" page="32" pageId="14" pageNumber="15" year="2012">
<bibRefCitation id="EF89B6BDF71CFC53FD13F31C6E6FFE8A" author="Reischutz A. &amp; Reischutz P. L." box="[732,1034,333,359]" etAl="et al." firstAuthor="Reischutz" journalOrPublisher="Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft" pageId="14" pageNumber="15" pagination="17 - 18" part="19" refId="ref13210" refString="Reischutz A. &amp; Reischutz P. L. 2012. Virpazaria nicoleae nov. spec. (Spelaeodiscidae: Pulmonata: Gastropoda) aus Montenegro. Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft 19: 17 - 18." title="Virpazaria nicoleae nov. spec. (Spelaeodiscidae: Pulmonata: Gastropoda) aus Montenegro" type="journal article" year="2012">
Reischütz
<emphasis id="B96C175EF71CFC53FC98F31F69F7FE8A" box="[855,914,333,359]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
, 2012: 32
</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, fig. 2.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30298C7F71CFC53FF72F3DF6BC6FE17" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BA7CB4CF71CFC53FF72F3DF6B53FE45" blockId="14.[189,310,398,424]" box="[189,310,398,424]" pageId="14" pageNumber="15">
<heading id="D0EF7C20F71CFC53FF72F3DF6B53FE45" bold="true" box="[189,310,398,424]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B96C175EF71CFC53FF72F3DF6B53FE45" bold="true" box="[189,310,398,424]" pageId="14" pageNumber="15">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA7CB4CF71CFC53FF72F3EC6BC6FE17" blockId="14.[189,1398,445,506]" pageId="14" pageNumber="15">Distinguished from nominate form by smaller, non-detached aperture, which is less covered by parietal flap in frontal view.</paragraph>
</subSubSection>
<caption id="DF679BC4F71CFC53FF72F536680AF806" ID-DOI="http://doi.org/10.5281/zenodo.3474984" ID-Zenodo-Dep="3474984" httpUri="https://zenodo.org/record/3474984/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1895,1921]" targetBox="[189,1399,582,1858]" targetPageId="14">
<paragraph id="8BA7CB4CF71CFC53FF72F536680AF806" blockId="14.[189,1398,1895,2027]" pageId="14" pageNumber="15">
<emphasis id="B96C175EF71CFC53FF72F5366B75F86C" bold="true" box="[189,272,1895,1921]" pageId="14" pageNumber="15">Fig. 5.</emphasis>
Shells of species of
<taxonomicName id="4C18B0CFF71CFC53FDD0F53969EBF86C" authority="Gittenberger, 1969" authorityName="Gittenberger" authorityYear="1969" box="[543,910,1895,1921]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
<emphasis id="B96C175EF71CFC53FDD0F53968F9F86C" box="[543,668,1896,1921]" italics="true" pageId="14" pageNumber="15">Virpazaria</emphasis>
<bibRefCitation id="EF89B6BDF71CFC53FD66F53669EBF86C" author="Gittenberger E." box="[681,910,1895,1921]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="14" pageNumber="15" pagination="287 - 306" part="43" refId="ref12709" refString="Gittenberger E. 1969. Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306." title="Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae" type="journal article" year="1969">Gittenberger, 1969</bibRefCitation>
</taxonomicName>
.
<emphasis id="B96C175EF71CFC53FC50F53669B3F86C" bold="true" box="[927,982,1895,1921]" pageId="14" pageNumber="15">A–E</emphasis>
.
<taxonomicName id="4C18B0CFF71CFC53FC24F5396855F849" authority="Reischutz &amp; Reischutz, 2012" authorityName="A. Reischutz &amp; P. L. Reischutz" authorityYear="2012" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="nicoleae" subGenus="Virpazaria">
<emphasis id="B96C175EF71CFC53FC24F5396F13F86C" box="[1003,1398,1895,1921]" italics="true" pageId="14" pageNumber="15">Virpazaria (Virpazaria) nicoleae</emphasis>
<bibRefCitation id="EF89B6BDF71CFC53FF72F5DB6855F849" author="Reischutz A. &amp; Reischutz P. L." box="[189,560,1930,1956]" firstAuthor="Reischutz" journalOrPublisher="Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft" pageId="14" pageNumber="15" pagination="17 - 18" part="19" refId="ref13210" refString="Reischutz A. &amp; Reischutz P. L. 2012. Virpazaria nicoleae nov. spec. (Spelaeodiscidae: Pulmonata: Gastropoda) aus Montenegro. Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft 19: 17 - 18." title="Virpazaria nicoleae nov. spec. (Spelaeodiscidae: Pulmonata: Gastropoda) aus Montenegro" type="journal article" year="2012">Reischütz &amp; Reischütz, 2012</bibRefCitation>
</taxonomicName>
, holotype (NHMW 108113).
<emphasis id="B96C175EF71CFC53FC63F5DB698DF849" bold="true" box="[940,1000,1930,1956]" pageId="14" pageNumber="15">F–M</emphasis>
.
<taxonomicName id="4C18B0CFF71CFC53FBCFF5DA6BFAF82A" authority="Gittenberger, 1969" authorityName="Gittenberger" authorityYear="1969" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="pageti" subGenus="Virpazaria">
<emphasis id="B96C175EF71CFC53FBCFF5DA6F13F849" box="[1024,1398,1931,1957]" italics="true" pageId="14" pageNumber="15">Virpazaria (Virpazaria) pageti</emphasis>
<bibRefCitation id="EF89B6BDF71CFC53FF72F5FC6BFAF82A" author="Gittenberger E." box="[189,415,1965,1991]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="14" pageNumber="15" pagination="287 - 306" part="43" refId="ref12709" refString="Gittenberger E. 1969. Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306." title="Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae" type="journal article" year="1969">Gittenberger, 1969</bibRefCitation>
</taxonomicName>
. F–I. Between Fort Prevlaka and the border station (HNHM 104391). J–M. Igalo (HNHM 102250). Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection id="C30298C7F71DFC52FF72F35A68EDFDA8" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BA7CB4CF71DFC52FF72F35A6BCFFEC8" blockId="15.[189,426,267,293]" box="[189,426,267,293]" pageId="15" pageNumber="16">
<heading id="D0EF7C20F71DFC52FF72F35A6BCFFEC8" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B96C175EF71DFC52FF72F35A6BCFFEC8" bold="true" box="[189,426,267,293]" pageId="15" pageNumber="16">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA7CB4CF71DFC52FF25F36B6B39FEB9" blockId="15.[189,1398,314,410]" box="[234,348,314,340]" pageId="15" pageNumber="16">
<emphasis id="B96C175EF71DFC52FF25F36B6B39FEB9" bold="true" box="[234,348,314,340]" pageId="15" pageNumber="16">
<typeStatus id="54A375EEF71DFC52FF25F36B6B39FEB9" box="[234,348,314,340]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA7CB4CF71DFC52FF72F30C6B6BFE77" blockId="15.[189,1398,314,410]" pageId="15" pageNumber="16">
<materialsCitation id="3B70C111F71DFC52FF72F30C6B6BFE77" ID-GBIF-Occurrence="2424151048" collectingDate="2010-04-06" collectionCode="CNHM" collectorName="R. Ozimec" country="Croatia" county="Vilina Cave" location="Konavle Region" municipality="Gruda" pageId="15" pageNumber="16" specimenCode="CNHM 10393" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F30F8BDCF71DFC52FF72F30C6B5FFE9A" box="[189,314,349,375]" name="Croatia" pageId="15" pageNumber="16">CROATIA</collectingCountry>
•
<location id="8EC79D97F71DFC52FE9AF30D687CFE9B" LSID="urn:lsid:plazi:treatment:03B17A5AF71CFC52FEF4F35A6BC5F97D:8EC79D97F71DFC52FE9AF30D687CFE9B" box="[341,537,348,374]" country="Croatia" county="Vilina Cave" municipality="Gruda" name="Konavle Region" pageId="15" pageNumber="16">Konavle Region</location>
,
<collectingMunicipality id="6BC35136F71DFC52FDEAF30D6814FE9B" box="[549,625,348,374]" pageId="15" pageNumber="16">Gruda</collectingMunicipality>
,
<collectingCounty id="62C6B3C0F71DFC52FDB2F30D6968FE9A" box="[637,781,348,375]" pageId="15" pageNumber="16">Vilina Cave</collectingCounty>
; UTM: BN81;
<date id="FFA6ED8CF71DFC52FC0EF30D6E35FE9A" box="[961,1104,348,375]" pageId="15" pageNumber="16" value="2010-04-06">
<collectingDate id="EFE21464F71DFC52FC0EF30D6E35FE9A" box="[961,1104,348,375]" pageId="15" pageNumber="16" value="2010-04-06">6 Apr. 2010</collectingDate>
</date>
;
<collectorName id="26EDAE9AF71DFC52FB92F30C6EBEFE9B" box="[1117,1243,348,375]" pageId="15" pageNumber="16">R. Ozimec</collectorName>
leg.;
<specimenCode id="DBBE6337F71DFC52FAD1F30C6B6FFE77" collectionCode="CNHM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" lsid="urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="15" pageNumber="16" type="Museum">CNHM 10393</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA7CB4CF71DFC52FF25F3906B02FE36" blockId="15.[189,1399,449,581]" box="[234,359,449,475]" pageId="15" pageNumber="16">
<emphasis id="B96C175EF71DFC52FF25F3906B02FE36" bold="true" box="[234,359,449,475]" pageId="15" pageNumber="16">
<typeStatus id="54A375EEF71DFC52FF25F3906B02FE36" box="[234,359,449,475]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA7CB4CF71DFC52FF72F3B468EDFDA8" blockId="15.[189,1399,449,581]" pageId="15" pageNumber="16">
<materialsCitation id="3B70C111F71DFC52FF72F3B46B57FDCF" ID-GBIF-Occurrence="2424151003" collectingDate="2010-04-06" collectionCode="CNHM" collectorName="Croatian Biospeleological Society" country="Croatia" county="Vilina Cave" location="Konavle Region" municipality="Gruda" pageId="15" pageNumber="16" specimenCount="2" typeStatus="paratype">
<collectingCountry id="F30F8BDCF71DFC52FF72F3B46B5FFE12" box="[189,314,485,511]" name="Croatia" pageId="15" pageNumber="16">CROATIA</collectingCountry>
•
<specimenCount id="9D1E00C5F71DFC52FE9CF3B46BCDFE12" box="[339,424,485,511]" pageId="15" pageNumber="16" type="generic">2 spec.</specimenCount>
; same collection data as for holotype;
<collectorName id="26EDAE9AF71DFC52FCA4F3B56E9EFE13" box="[875,1275,484,510]" pageId="15" pageNumber="16">Croatian Biospeleological Society</collectorName>
collection [not seen]
</materialsCitation>
•
<materialsCitation id="3B70C111F71DFC52FE83F05968E1FDA8" ID-GBIF-Occurrence="2424146955" collectingDate="2011-03" collectorName="A. and P. L. Reischutz &amp; Reischutz" country="Croatia" location="Dubravka village" pageId="15" pageNumber="16" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D1E00C5F71DFC52FE83F0596BC4FDCF" box="[332,417,520,546]" pageId="15" pageNumber="16" type="generic">1 spec.</specimenCount>
; in
<location id="8EC79D97F71DFC52FE02F05968FEFDCF" LSID="urn:lsid:plazi:treatment:03B17A5AF71CFC52FEF4F35A6BC5F97D:8EC79D97F71DFC52FE02F05968FEFDCF" box="[461,667,520,546]" country="Croatia" name="Dubravka village" pageId="15" pageNumber="16">Dubravka village</location>
, ca
<locationDeviation id="9A15D26EF71DFC52FD06F05669CCFDCF" box="[713,937,519,546]" location="Gruda" pageId="15" pageNumber="16">
<quantity id="4CE066A9F71DFC52FD06F0566961FDCF" box="[713,772,519,546]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="16" unit="km" value="5.0">5 km</quantity>
NE of Gruda
</locationDeviation>
;
<date id="FFA6ED8CF71DFC52FC7AF0596E54FDCF" box="[949,1073,520,546]" pageId="15" pageNumber="16" value="2011-03">
<collectingDate id="EFE21464F71DFC52FC7AF0596E54FDCF" box="[949,1073,520,546]" pageId="15" pageNumber="16" value="2011-03">Mar. 2011</collectingDate>
</date>
;
<collectorName id="26EDAE9AF71DFC52FBF3F0596F59FDCF" box="[1084,1340,520,546]" pageId="15" pageNumber="16">A. and P.L. Reischütz</collectorName>
leg.;
<collectorName id="26EDAE9AF71DFC52FF72F07A6B54FDA8" box="[189,305,555,581]" pageId="15" pageNumber="16">Reischütz</collectorName>
private collection [not seen]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30298C7F71DFC52FF72F03D682FFD58" box="[189,586,620,694]" pageId="15" pageNumber="16" type="description">
<paragraph id="8BA7CB4CF71DFC52FF72F03D6B2BFD6B" blockId="15.[189,334,620,646]" box="[189,334,620,646]" pageId="15" pageNumber="16">
<heading id="D0EF7C20F71DFC52FF72F03D6B2BFD6B" bold="true" box="[189,334,620,646]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B96C175EF71DFC52FF72F03D6B2BFD6B" bold="true" box="[189,334,620,646]" pageId="15" pageNumber="16">Dimensions</emphasis>
</heading>
</paragraph>
<paragraph id="8BA7CB4CF71DFC52FF72F0CA682FFD58" blockId="15.[189,586,667,694]" box="[189,586,667,694]" pageId="15" pageNumber="16">
H
<quantity id="4CE066A9F71DFC52FF16F0CA6B1DFD58" box="[217,376,667,694]" metricMagnitude="-3" metricUnit="m" metricValue="1.575" metricValueMax="1.65" metricValueMin="1.5" pageId="15" pageNumber="16" unit="mm" value="1.575" valueMax="1.65" valueMin="1.5">1.5–1.65 mm</quantity>
; W 2.75–3.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="C30298C7F71DFC52FF72F08D6BC5F97D" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8BA7CB4CF71DFC52FF72F08D6B98FD1B" blockId="15.[189,509,732,758]" box="[189,509,732,758]" pageId="15" pageNumber="16">
<heading id="D0EF7C20F71DFC52FF72F08D6B98FD1B" bold="true" box="[189,509,732,758]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B96C175EF71DFC52FF72F08D6B98FD1B" bold="true" box="[189,509,732,758]" pageId="15" pageNumber="16">Distribution and remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA7CB4CF71DFC52FF72F15A69B9FA2C" blockId="15.[189,1399,779,1474]" pageId="15" pageNumber="16">
This species (including its subspecies) has a restricted range north of Kotor Bay in
<collectingCountry id="F30F8BDCF71DFC52FB91F15A6ED1FCC8" box="[1118,1204,779,805]" name="Croatia" pageId="15" pageNumber="16">Croatia</collectingCountry>
and
<collectingCountry id="F30F8BDCF71DFC52FB28F15A6F13FCC8" box="[1255,1398,779,805]" name="Serbia" pageId="15" pageNumber="16">Montenegro</collectingCountry>
(
<bibRefCitation id="EF89B6BDF71DFC52FF0AF17F6BFAFCA5" author="Gittenberger E." box="[197,415,814,840]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="15" pageNumber="16" pagination="287 - 306" part="43" refId="ref12709" refString="Gittenberger E. 1969. Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306." title="Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae" type="journal article" year="1969">Gittenberger 1969</bibRefCitation>
;
<bibRefCitation id="EF89B6BDF71DFC52FE64F17F68C3FCA5" author="Reischutz A. &amp; Reischutz P. L." box="[427,678,814,840]" etAl="et al." firstAuthor="Reischutz" journalOrPublisher="Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft" pageId="15" pageNumber="16" pagination="17 - 18" part="19" refId="ref13210" refString="Reischutz A. &amp; Reischutz P. L. 2012. Virpazaria nicoleae nov. spec. (Spelaeodiscidae: Pulmonata: Gastropoda) aus Montenegro. Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft 19: 17 - 18." title="Virpazaria nicoleae nov. spec. (Spelaeodiscidae: Pulmonata: Gastropoda) aus Montenegro" type="journal article" year="2012">
Reischütz
<emphasis id="B96C175EF71DFC52FDEAF17E6805FCA5" box="[549,608,814,840]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2012
</bibRefCitation>
). The
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lot of the nominate form was collected from debris at the mouth of the Sutorina River. Later, this species was found in rock crevices at the eastern side of Prevlaka Peninsula, between Njivice and Igalo very near the
<typeStatus id="54A375EEF71DFC52FC5AF12469ADFC62" box="[917,968,885,911]" pageId="15" pageNumber="16">type</typeStatus>
locality (see
<bibRefCitation id="EF89B6BDF71DFC52FBA5F1256F20FC63" author="Gittenberger E." box="[1130,1349,884,910]" firstAuthor="Gittenberger" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien" pageId="15" pageNumber="16" pagination="423 - 426" part="90" refId="ref12766" refString="Gittenberger E. 1988. Uber Virpazaria (V.) pageti Gittenberger 1969, Gyralina (G.) pageti n. sp. und deren Sekundarfundorte (Mollusca: Pulmonata). Annalen des Naturhistorischen Museums in Wien 90 B: 423 - 426." title="Uber Virpazaria (V.) pageti Gittenberger 1969, Gyralina (G.) pageti n. sp. und deren Sekundarfundorte (Mollusca: Pulmonata)" type="journal article" year="1988">Gittenberger 1988</bibRefCitation>
). At another point of the Prevlaka, ca
<quantity id="4CE066A9F71DFC52FD88F1C668FCFC5C" box="[583,665,919,946]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="15" pageNumber="16" unit="km" value="3.5">3.5 km</quantity>
as the crow flies from the former site,
<bibRefCitation id="EF89B6BDF71DFC52FBA7F1C66F13FC5F" author="Reischutz A. &amp; Reischutz P. L." box="[1128,1398,919,946]" etAl="et al." firstAuthor="Reischutz" journalOrPublisher="Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft" pageId="15" pageNumber="16" pagination="17 - 18" part="19" refId="ref13210" refString="Reischutz A. &amp; Reischutz P. L. 2012. Virpazaria nicoleae nov. spec. (Spelaeodiscidae: Pulmonata: Gastropoda) aus Montenegro. Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft 19: 17 - 18." title="Virpazaria nicoleae nov. spec. (Spelaeodiscidae: Pulmonata: Gastropoda) aus Montenegro" type="journal article" year="2012">
Reischütz
<emphasis id="B96C175EF71DFC52FB2BF1C96F45FC5F" box="[1252,1312,920,946]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
(2012)
</bibRefCitation>
reported further shells of this species (four adults and
<specimenCount id="9D1E00C5F71DFC52FCF1F1EA69B1FC3B" box="[830,980,955,982]" pageId="15" pageNumber="16" type="juvenile">15 juveniles</specimenCount>
). They were distinguished from the nominate form by their smaller shell, finer and denser ribbing, smaller parietal flap, as well as the nondetached aperture, and were described as
<taxonomicName id="4C18B0CFF71DFC52FD6BF6536993FBF6" box="[676,1014,1025,1051]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="subSpecies" species="pageti" subSpecies="alexanderi">
<emphasis id="B96C175EF71DFC52FD6BF6536993FBF6" box="[676,1014,1025,1051]" italics="true" pageId="15" pageNumber="16">Virpazaria pageti alexanderi</emphasis>
</taxonomicName>
. In 2018, we were able to obtain a larger sample of
<emphasis id="B96C175EF71DFC52FE58F674682BFBD3" box="[407,590,1060,1086]" italics="true" pageId="15" pageNumber="16">
V. p.
<taxonomicName id="4C18B0CFF71DFC52FE1EF675682BFBD3" authorityName="Reischutz &amp; Subai" authorityYear="2012" box="[465,590,1060,1086]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="alexanderi">alexanderi</taxonomicName>
</emphasis>
from the
<typeStatus id="54A375EEF71DFC52FD0EF6746891FBD2" box="[705,756,1061,1087]" pageId="15" pageNumber="16">type</typeStatus>
locality that provided a good opportunity to study size extremes and the variability of character states within this population. In our sample (HNHM 104391), shell width ranged between 2.1 and 3.0 mm, shell height between 1.3 and
<quantity id="4CE066A9F71DFC52FBF8F63A6EF3FB68" box="[1079,1174,1131,1157]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="15" pageNumber="16" unit="mm" value="1.9">1.9 mm</quantity>
, number of whorls between 3¾ and 4¼, number of ribs on the last whorl between 48 and 83 and we found specimens with detached and non-detached apertures as well. The values and character states given by
<bibRefCitation id="EF89B6BDF71DFC52FB2CF6E06B69FB03" author="Gittenberger E." firstAuthor="Gittenberger" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien" pageId="15" pageNumber="16" pagination="423 - 426" part="90" refId="ref12766" refString="Gittenberger E. 1988. Uber Virpazaria (V.) pageti Gittenberger 1969, Gyralina (G.) pageti n. sp. und deren Sekundarfundorte (Mollusca: Pulmonata). Annalen des Naturhistorischen Museums in Wien 90 B: 423 - 426." title="Uber Virpazaria (V.) pageti Gittenberger 1969, Gyralina (G.) pageti n. sp. und deren Sekundarfundorte (Mollusca: Pulmonata)" type="journal article" year="1988">Gittenberger (1988)</bibRefCitation>
for the nominate form and those given by
<bibRefCitation id="EF89B6BDF71DFC52FD3FF6856990FB03" author="Reischutz A. &amp; Reischutz P. L." box="[752,1013,1236,1262]" etAl="et al." firstAuthor="Reischutz" journalOrPublisher="Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft" pageId="15" pageNumber="16" pagination="17 - 18" part="19" refId="ref13210" refString="Reischutz A. &amp; Reischutz P. L. 2012. Virpazaria nicoleae nov. spec. (Spelaeodiscidae: Pulmonata: Gastropoda) aus Montenegro. Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft 19: 17 - 18." title="Virpazaria nicoleae nov. spec. (Spelaeodiscidae: Pulmonata: Gastropoda) aus Montenegro" type="journal article" year="2012">
Reischütz
<emphasis id="B96C175EF71DFC52FCA6F68469C7FB03" box="[873,930,1236,1262]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
(2012)
</bibRefCitation>
for
<emphasis id="B96C175EF71DFC52FBECF6846EB3FB03" box="[1059,1238,1236,1262]" italics="true" pageId="15" pageNumber="16">
V. p.
<taxonomicName id="4C18B0CFF71DFC52FB96F6856EB3FB03" authorityName="Reischutz &amp; Subai" authorityYear="2012" box="[1113,1238,1236,1262]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="alexanderi">alexanderi</taxonomicName>
</emphasis>
all fall within these ranges. This alone might be a good reason for treating them as synonyms, which is only reinforced once their geographical distribution is also taken into consideration. The two
<typeStatus id="54A375EEF71DFC52FB8DF74A6E10FAD8" box="[1090,1141,1307,1333]" pageId="15" pageNumber="16">type</typeStatus>
localities are not only close to each other, but are located along the foot of the same limestone hill, which is composed of the same Upper Cretaceous geological formation. Having no geographical or geological barrier in between, it is likely that the two sites contain the same subterranean population and an extensive sampling along the Prevlaka Peninsula might detect this species at further locations.
</paragraph>
<paragraph id="8BA7CB4CF71DFC52FF72F7BB6BC5F97D" blockId="15.[189,1399,1513,1680]" pageId="15" pageNumber="16">
<taxonomicName id="4C18B0CFF71DFC52FF72F7BB6B97F9EE" box="[189,498,1513,1539]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="subSpecies" species="pageti" subSpecies="kleteckii">
<emphasis id="B96C175EF71DFC52FF72F7BB6B97F9EE" box="[189,498,1513,1539]" italics="true" pageId="15" pageNumber="16">Virpazaria pageti kleteckii</emphasis>
</taxonomicName>
was found at two locations somewhat north of the Prevlaka; namely at Vilina Spilja near Gruda and at Dubravka ca
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NE of Gruda. The original description was based on four specimens only (three from Vilina Spilja and one from Dubravka). Until further available material makes it possible to test the stability of distinguishing features,
<emphasis id="B96C175EF71DFC52FC0DF4026E01F980" box="[962,1124,1619,1645]" italics="true" pageId="15" pageNumber="16">V. p. kleteckii</emphasis>
must be accepted as a distinct subspecies.
</paragraph>
</subSubSection>
</treatment>
</document>
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