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<document id="20266937D20B914FC8B7FF0C1AA439A6" ID-CLB-Dataset="272446" ID-DOI="10.1093/zoolinnean/zlad043" ID-GBIF-Dataset="81098796-8acf-4360-a625-ad76a281e99e" ID-ISSN="0024-4082" ID-Zenodo-Dep="8426263" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1696940821456" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan C. &amp; Cerdeño, Esperanza" docDate="2023" docId="03B17549E92047749D54FD3AFEE8FE06" docLanguage="en" docName="zlad043.pdf" docOrigin="Zoological Journal of the Linnean Society 199 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad043" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.1:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="1" docTitle="Protypotherium compressidens Ameghino 1891" docType="treatment" docVersion="6" lastPageNumber="432" masterDocId="FF880D31E92F47649E20FFB4FF9EFFC7" masterDocTitle="Systematic revision of the species of Protypotherium (Notoungulata: Interatheriidae) from the Santa Cruz Formation (Early-Middle Miocene), Argentinian Patagonia: a new phylogenetic hypothesis for the Interatheriidae" masterLastPageNumber="444" masterPageNumber="417" pageNumber="432" updateTime="1701999754155" updateUser="plazi">
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<mods:title id="02C32BE90C807DFE71759D16E90C7B9C">Systematic revision of the species of Protypotherium (Notoungulata: Interatheriidae) from the Santa Cruz Formation (Early-Middle Miocene), Argentinian Patagonia: a new phylogenetic hypothesis for the Interatheriidae</mods:title>
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<mods:namePart id="CE2A43AEA047AE918DB7FBD1D3E85DC3">Fernández, Mercedes</mods:namePart>
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<mods:namePart id="75B1FD0FCEDDCCF5B1E6440B83EC3A99">Fernicola, Juan C.</mods:namePart>
<mods:affiliation id="E928AC588D94E5BD9EB46C83CB3F7AA4">División Paleontología de Vertebrados, Museo Argentino de Ciencias Naturales Bernardino Rivadavia, Avenida Ángel Gallardo 470, C 1405 DJR Ciudad Autónoma de Buenos Aires, Argentina &amp; Laboratorio Anatomía y Evolución de los Vertebrados, Departamento de Ciencias Básicas, Universidad Nacional de Luján, Ruta 5 y Avenida Constitución 6700 Luján, Buenos Aires, Argentina &amp; Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Juan Domingo Perón 2158, C 1040 AAH, Buenos Aires, Argentina</mods:affiliation>
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<treatment id="03B17549E92047749D54FD3AFEE8FE06" ID-DOI="http://doi.org/10.5281/zenodo.10267836" ID-GBIF-Taxon="213996069" ID-Zenodo-Dep="10267836" LSID="urn:lsid:plazi:treatment:03B17549E92047749D54FD3AFEE8FE06" httpUri="http://treatment.plazi.org/id/03B17549E92047749D54FD3AFEE8FE06" lastPageId="16" lastPageNumber="432" pageId="15" pageNumber="432">
<subSubSection id="C30297D4E920476B9D54FD3AFAF7FD6E" box="[884,1385,654,681]" pageId="15" pageNumber="432" type="nomenclature">
<paragraph id="8BA7C45FE920476B9D54FD3AFAF7FD6E" blockId="15.[884,1385,654,681]" box="[884,1385,654,681]" pageId="15" pageNumber="432">
<heading id="D0EF7333E920476B9D54FD3AFAF7FD6E" bold="true" box="[884,1385,654,681]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="432" reason="2">
<taxonomicName id="4C18BFDCE920476B9D54FD3AFAF7FD6E" authority="Ameghino, 1891 b" authorityName="Ameghino" authorityYear="1891" box="[884,1385,654,681]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="compressidens">
<emphasis id="B96C184DE920476B9D54FD3AFAF7FD6E" bold="true" box="[884,1385,654,681]" pageId="15" pageNumber="432">
<emphasis id="B96C184DE920476B9D54FD3AFB38FD6E" bold="true" box="[884,1190,654,681]" italics="true" pageId="15" pageNumber="432">Protypotherium compressidens</emphasis>
<bibRefCitation id="EF89B9AEE920476B9A8AFD3BFAF7FD6E" author="Ameghino F" box="[1194,1385,655,681]" pageId="15" pageNumber="432" pagination="289 - 328" refId="ref21934" refString="Ameghino F. Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural 1891 b; 1: 289 - 328." type="journal article" year="1891">Ameghino, 1891b</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C30297D4E920476B9D0AFD01FC7BFCCB" pageId="15" pageNumber="432" type="description">
<paragraph id="8BA7C45FE920476B9D0AFD01FC7BFCCB" blockId="15.[810,1459,693,780]" pageId="15" pageNumber="432">
(
<figureCitation id="1323D8DAE920476B9D15FD01FCEFFD0A" box="[821,881,693,717]" captionStart="Figure 5" captionStartId="8.[129,194,1753,1777]" captionTargetBox="[257,1345,147,1723]" captionTargetId="figure-83@8.[254,1348,144,1726]" captionTargetPageId="8" captionText="Figure 5. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs.width of M1. B, Length vs. width of m1. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426276" httpUri="https://zenodo.org/record/8426276/files/figure.png" pageId="15" pageNumber="432">Figs 5</figureCitation>
,
<figureCitation id="1323D8DAE920476B9D5BFD01FC16FD0A" box="[891,904,693,717]" captionStart="Figure 6" captionStartId="9.[112,177,1757,1781]" captionTargetBox="[242,1330,147,1727]" captionTargetId="figure-87@9.[239,1333,144,1730]" captionTargetPageId="9" captionText="Figure 6. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs. width of M2. B, Length vs. width of m2. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426278" httpUri="https://zenodo.org/record/8426278/files/figure.png" pageId="15" pageNumber="432">6</figureCitation>
,
<figureCitation id="1323D8DAE920476B9DB3FD01FC51FD0A" box="[915,975,693,717]" captionStart="Figure 8" captionStartId="13.[113,178,1780,1804]" captionTargetBox="[188,1387,151,1749]" captionTargetId="figure-7@13.[186,1387,149,1752]" captionTargetPageId="13" captionText="Figure 8. Protypotherium praerutilum, in occlusal view.A, MACN-A 4044, incomplete palate. Protypotherium claudum, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)p3m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1dp1p2p4, and right i1 and dp1p2m3. Protypotherium compressidens, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)m3; (G) MACN-A 3867, right mandibular fragment with dp12p34 (trigonid); (H) MACN-A 97169732?g, right mandibular fragment with m1 (talonid)3; (I) MPM-PV 19761, right maxillary fragment with M12; (J) MPM-PV 19781, right mandibular fragment with p2m1; (K) MPM-PV 19782, less maxillary fragment with P2M3. Abbreviation: , indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426282" httpUri="https://zenodo.org/record/8426282/files/figure.png" pageId="15" pageNumber="432">8EK</figureCitation>
; Supporting Information, Table S2) 1891b
<taxonomicName id="4C18BFDCE920476B9DA7FD61FAB4FD2A" authority="Ameghino" authorityName="Ameghino" authorityYear="1891" box="[903,1322,725,749]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="compressidens">
<emphasis id="B96C184DE920476B9DA7FD61FB2DFD2A" box="[903,1203,725,749]" italics="true" pageId="15" pageNumber="432">Protypotherium compressidens</emphasis>
Ameghino
</taxonomicName>
, p. 292 (original description).
</paragraph>
</subSubSection>
<subSubSection id="C30297D4E920476B9D0AFC99FB4CFB10" pageId="15" pageNumber="432" type="materials_examined">
<paragraph id="8BA7C45FE920476B9D0AFC99FC35FC04" blockId="15.[810,1459,813,963]" pageId="15" pageNumber="432">
<materialsCitation id="3B70CE02E920476B9D0AFC99FC35FC04" ID-GBIF-Occurrence="4425895305" collectionCode="MACN-A" country="Argentina" location="Early - Middle Miocene" pageId="15" pageNumber="432" specimenCode="MACN-A 4029, MACN-A 4030" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<emphasis id="B96C184DE920476B9D0AFC99FC14FC82" box="[810,906,813,837]" italics="true" pageId="15" pageNumber="432">
<typeStatus id="54A37AFDE920476B9D0AFC99FC1BFC82" box="[810,901,813,837]" pageId="15" pageNumber="432" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DBBE6C24E920476B9DB3FC99FBACFC82" box="[915,1074,813,837]" collectionCode="MACN-A" pageId="15" pageNumber="432">MACN-A 4029</specimenCode>
(
<figureCitation id="1323D8DAE920476B9A61FC99FB18FC82" box="[1089,1158,813,837]" captionStart="Figure 8" captionStartId="13.[113,178,1780,1804]" captionTargetBox="[188,1387,151,1749]" captionTargetId="figure-7@13.[186,1387,149,1752]" captionTargetPageId="13" captionText="Figure 8. Protypotherium praerutilum, in occlusal view.A, MACN-A 4044, incomplete palate. Protypotherium claudum, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)p3m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1dp1p2p4, and right i1 and dp1p2m3. Protypotherium compressidens, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)m3; (G) MACN-A 3867, right mandibular fragment with dp12p34 (trigonid); (H) MACN-A 97169732?g, right mandibular fragment with m1 (talonid)3; (I) MPM-PV 19761, right maxillary fragment with M12; (J) MPM-PV 19781, right mandibular fragment with p2m1; (K) MPM-PV 19782, less maxillary fragment with P2M3. Abbreviation: , indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426282" httpUri="https://zenodo.org/record/8426282/files/figure.png" pageId="15" pageNumber="432">Fig. 8E</figureCitation>
), less maxillary fragment with P3M3, and
<specimenCode id="DBBE6C24E920476B9D8DFCF9FBD2FCA2" box="[941,1100,845,869]" collectionCode="MACN-A" pageId="15" pageNumber="432">MACN-A 4030</specimenCode>
(
<figureCitation id="1323D8DAE920476B9A7CFCF9FB3CFCA2" box="[1116,1186,845,869]" captionStart="Figure 8" captionStartId="13.[113,178,1780,1804]" captionTargetBox="[188,1387,151,1749]" captionTargetId="figure-7@13.[186,1387,149,1752]" captionTargetPageId="13" captionText="Figure 8. Protypotherium praerutilum, in occlusal view.A, MACN-A 4044, incomplete palate. Protypotherium claudum, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)p3m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1dp1p2p4, and right i1 and dp1p2m3. Protypotherium compressidens, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)m3; (G) MACN-A 3867, right mandibular fragment with dp12p34 (trigonid); (H) MACN-A 97169732?g, right mandibular fragment with m1 (talonid)3; (I) MPM-PV 19761, right maxillary fragment with M12; (J) MPM-PV 19781, right mandibular fragment with p2m1; (K) MPM-PV 19782, less maxillary fragment with P2M3. Abbreviation: , indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426282" httpUri="https://zenodo.org/record/8426282/files/figure.png" pageId="15" pageNumber="432">Fig. 8F</figureCitation>
), less mandibular fragment with p4 (talonid)m3, of the same individual (see
<bibRefCitation id="EF89B9AEE920476B9B67FCD8FC0AFC64" author="Fernandez M &amp; Fernicola JC &amp; Cerdeno E" pageId="15" pageNumber="432" pagination="473 - 98" refId="ref23394" refString="Fernandez M, Fernicola JC, Cerdeno E. Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino. Zootaxa 2018; 4387: 473 - 98. hups: // doi. org / 10.11646 / ZOOTAXA. 4387.3.4" type="journal article" year="2018">
Fernández
<emphasis id="B96C184DE920476B9D0AFC38FCC4FC64" box="[810,858,907,931]" italics="true" pageId="15" pageNumber="432">et al.</emphasis>
2018
</bibRefCitation>
),
<location id="8EC79284E920476B9D8CFC3FFB02FC64" LSID="urn:lsid:plazi:treatment:03B17549E92047749D54FD3AFEE8FE06:8EC79284E920476B9D8CFC3FFB02FC64" box="[940,1180,907,931]" country="Argentina" name="Early - Middle Miocene" pageId="15" pageNumber="432" stateProvince="Santa Cruz">EarlyMiddle Miocene</location>
, SCF,
<collectingRegion id="49DC0ABDE920476B9AFFFC3FFA2DFC64" box="[1247,1459,907,931]" country="Argentina" name="Santa Cruz" pageId="15" pageNumber="432">Santa Cruz Province</collectingRegion>
(
<collectingCountry id="F30F84CFE920476B9D11FC1FFC01FC04" box="[817,927,939,963]" name="Argentina" pageId="15" pageNumber="432">Argentina</collectingCountry>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA7C45FE920476B9D0AFC50FB4CFB10" blockId="15.[810,1460,996,1239]" pageId="15" pageNumber="432">
<emphasis id="B96C184DE920476B9D0AFC50FC75FC3B" box="[810,1003,996,1020]" italics="true" pageId="15" pageNumber="432">Referred materials:</emphasis>
MACN-A 41-55 lot b.f, less mandibular fragment with poorly preserved p34 (trigonid); MACN-A 3867 (
<figureCitation id="1323D8DAE920476B9D52FB97FC5EFBFC" box="[882,960,1059,1083]" captionStart="Figure 8" captionStartId="13.[113,178,1780,1804]" captionTargetBox="[188,1387,151,1749]" captionTargetId="figure-7@13.[186,1387,149,1752]" captionTargetPageId="13" captionText="Figure 8. Protypotherium praerutilum, in occlusal view.A, MACN-A 4044, incomplete palate. Protypotherium claudum, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)p3m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1dp1p2p4, and right i1 and dp1p2m3. Protypotherium compressidens, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)m3; (G) MACN-A 3867, right mandibular fragment with dp12p34 (trigonid); (H) MACN-A 97169732?g, right mandibular fragment with m1 (talonid)3; (I) MPM-PV 19761, right maxillary fragment with M12; (J) MPM-PV 19781, right mandibular fragment with p2m1; (K) MPM-PV 19782, less maxillary fragment with P2M3. Abbreviation: , indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426282" httpUri="https://zenodo.org/record/8426282/files/figure.png" pageId="15" pageNumber="432">Fig. 8G</figureCitation>
), right mandibular fragment with dp12p34 (trigonid); MACN-A 97169732?g (
<figureCitation id="1323D8DAE920476B9A85FBF6FB6FFB9D" box="[1189,1265,1090,1114]" captionStart="Figure 8" captionStartId="13.[113,178,1780,1804]" captionTargetBox="[188,1387,151,1749]" captionTargetId="figure-7@13.[186,1387,149,1752]" captionTargetPageId="13" captionText="Figure 8. Protypotherium praerutilum, in occlusal view.A, MACN-A 4044, incomplete palate. Protypotherium claudum, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)p3m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1dp1p2p4, and right i1 and dp1p2m3. Protypotherium compressidens, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)m3; (G) MACN-A 3867, right mandibular fragment with dp12p34 (trigonid); (H) MACN-A 97169732?g, right mandibular fragment with m1 (talonid)3; (I) MPM-PV 19761, right maxillary fragment with M12; (J) MPM-PV 19781, right mandibular fragment with p2m1; (K) MPM-PV 19782, less maxillary fragment with P2M3. Abbreviation: , indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426282" httpUri="https://zenodo.org/record/8426282/files/figure.png" pageId="15" pageNumber="432">Fig. 8H</figureCitation>
), right mandibular fragment with m1 (talonid)3; MPM-PV 19761 (
<figureCitation id="1323D8DAE920476B9B0AFBD5FAF5FBBE" box="[1322,1387,1121,1145]" captionStart="Figure 8" captionStartId="13.[113,178,1780,1804]" captionTargetBox="[188,1387,151,1749]" captionTargetId="figure-7@13.[186,1387,149,1752]" captionTargetPageId="13" captionText="Figure 8. Protypotherium praerutilum, in occlusal view.A, MACN-A 4044, incomplete palate. Protypotherium claudum, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)p3m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1dp1p2p4, and right i1 and dp1p2m3. Protypotherium compressidens, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)m3; (G) MACN-A 3867, right mandibular fragment with dp12p34 (trigonid); (H) MACN-A 97169732?g, right mandibular fragment with m1 (talonid)3; (I) MPM-PV 19761, right maxillary fragment with M12; (J) MPM-PV 19781, right mandibular fragment with p2m1; (K) MPM-PV 19782, less maxillary fragment with P2M3. Abbreviation: , indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426282" httpUri="https://zenodo.org/record/8426282/files/figure.png" pageId="15" pageNumber="432">Fig. 8I</figureCitation>
), right maxillary fragment with M12; MPM-PV 19781 (
<figureCitation id="1323D8DAE920476B9B0BFB35FAF2FB5E" box="[1323,1388,1153,1177]" captionStart="Figure 8" captionStartId="13.[113,178,1780,1804]" captionTargetBox="[188,1387,151,1749]" captionTargetId="figure-7@13.[186,1387,149,1752]" captionTargetPageId="13" captionText="Figure 8. Protypotherium praerutilum, in occlusal view.A, MACN-A 4044, incomplete palate. Protypotherium claudum, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)p3m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1dp1p2p4, and right i1 and dp1p2m3. Protypotherium compressidens, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)m3; (G) MACN-A 3867, right mandibular fragment with dp12p34 (trigonid); (H) MACN-A 97169732?g, right mandibular fragment with m1 (talonid)3; (I) MPM-PV 19761, right maxillary fragment with M12; (J) MPM-PV 19781, right mandibular fragment with p2m1; (K) MPM-PV 19782, less maxillary fragment with P2M3. Abbreviation: , indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426282" httpUri="https://zenodo.org/record/8426282/files/figure.png" pageId="15" pageNumber="432">Fig. 8J</figureCitation>
), right mandibular fragment with p2m1; and MPM-PV 19782 (
<figureCitation id="1323D8DAE920476B9BADFB14FCD7FB10" captionStart="Figure 8" captionStartId="13.[113,178,1780,1804]" captionTargetBox="[188,1387,151,1749]" captionTargetId="figure-7@13.[186,1387,149,1752]" captionTargetPageId="13" captionText="Figure 8. Protypotherium praerutilum, in occlusal view.A, MACN-A 4044, incomplete palate. Protypotherium claudum, in occlusal view. B, MACN-A 551, right mandibular fragment with p2 (alveolus)p3m2; (C) MLP 73-VII-6-4 d, less mandibular fragment with p2 (broken)m2 (trigonid); (D) MPEF-PV 1394, incomplete mandible with less i1dp1p2p4, and right i1 and dp1p2m3. Protypotherium compressidens, in occlusal view. E, MACN-A 4029, less maxillary fragment with P3M3; (F) MACN-A 4030, less mandibular fragment with p4 (talonid)m3; (G) MACN-A 3867, right mandibular fragment with dp12p34 (trigonid); (H) MACN-A 97169732?g, right mandibular fragment with m1 (talonid)3; (I) MPM-PV 19761, right maxillary fragment with M12; (J) MPM-PV 19781, right mandibular fragment with p2m1; (K) MPM-PV 19782, less maxillary fragment with P2M3. Abbreviation: , indicates broken tooth. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8426282" httpUri="https://zenodo.org/record/8426282/files/figure.png" pageId="15" pageNumber="432">Fig. 8K</figureCitation>
), less maxillary fragment with P2M3.
</paragraph>
</subSubSection>
<subSubSection id="C30297D4E920476B9D0AFB4DFA2DF90F" pageId="15" pageNumber="432" type="diagnosis">
<paragraph id="8BA7C45FE920476B9D0AFB4DFA2DF90F" blockId="15.[810,1462,1273,1736]" pageId="15" pageNumber="432">
<emphasis id="B96C184DE920476B9D0AFB4DFA87FAD6" box="[810,1305,1273,1297]" italics="true" pageId="15" pageNumber="432">
Extended diagnosis:
<taxonomicName id="4C18BFDCE920476B9DD3FB4DFA87FAD6" authorityName="Ameghino" authorityYear="1891" box="[1011,1305,1273,1297]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="compressidens">Protypotherium compressidens</taxonomicName>
</emphasis>
differs from
<taxonomicName id="4C18BFDCE920476B9BB8FB4DFC17FAF7" authorityName="Ameghino" authorityYear="1885" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="antiquum">
<emphasis id="B96C184DE920476B9BB8FB4DFC17FAF7" italics="true" pageId="15" pageNumber="432">Pr. antiquum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18BFDCE920476B9DB3FAADFC61FAF7" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[915,1023,1304,1328]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="australe">
<emphasis id="B96C184DE920476B9DB3FAADFC61FAF7" box="[915,1023,1304,1328]" italics="true" pageId="15" pageNumber="432">Pr. australe</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18BFDCE920476B9A2BFAADFB00FAF7" authorityName="Ameghino" authorityYear="1887" box="[1035,1182,1304,1328]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="praerutilum">
<emphasis id="B96C184DE920476B9A2BFAADFB00FAF7" box="[1035,1182,1304,1328]" italics="true" pageId="15" pageNumber="432">Pr. praerutilum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18BFDCE920476B9A89FAADFAB1FAF7" authorityName="Ameghino" authorityYear="1902" box="[1193,1327,1304,1328]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="columnifer">
<emphasis id="B96C184DE920476B9A89FAADFAB1FAF7" box="[1193,1327,1304,1328]" italics="true" pageId="15" pageNumber="432">Pr. columnifer</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18BFDCE920476B9B1AFAADFA33FAF7" authorityName="Ameghino" authorityYear="1889" box="[1338,1453,1304,1328]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="claudum">
<emphasis id="B96C184DE920476B9B1AFAADFA33FAF7" box="[1338,1453,1304,1328]" italics="true" pageId="15" pageNumber="432">Pr. claudum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18BFDCE920476B9D0AFA8CFC3AFA88" authorityName="Cabrera &amp; Kraglievich" authorityYear="1931" box="[810,932,1336,1359]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="minutum">
<emphasis id="B96C184DE920476B9D0AFA8CFC3AFA88" box="[810,932,1336,1359]" italics="true" pageId="15" pageNumber="432">Pr. minutum</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C18BFDCE920476B9DFAFA8CFBE0FA88" box="[986,1150,1335,1359]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="colloncurensis">
<emphasis id="B96C184DE920476B9DFAFA8CFBE0FA88" box="[986,1150,1335,1359]" italics="true" pageId="15" pageNumber="432">Pr. colloncurensis</emphasis>
</taxonomicName>
by narrower cheek teeth, even more noticeable in P2M3; from
<taxonomicName id="4C18BFDCE920476B9AB3FAE3FA88FAA9" authorityName="Ameghino" authorityYear="1885" box="[1171,1302,1367,1390]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="antiquum">
<emphasis id="B96C184DE920476B9AB3FAE3FA88FAA9" box="[1171,1302,1367,1390]" italics="true" pageId="15" pageNumber="432">Pr. antiquum</emphasis>
</taxonomicName>
by the talonid of m3 without a labial groove; from
<taxonomicName id="4C18BFDCE920476B9A96FAC3FAC0FA49" box="[1206,1374,1398,1422]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="colloncurensis">
<emphasis id="B96C184DE920476B9A96FAC3FAC0FA49" box="[1206,1374,1398,1422]" italics="true" pageId="15" pageNumber="432">Pr. colloncurensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C18BFDCE920476B9BB8FAC3FC0EFA6A" authorityName="Ameghino" authorityYear="1902" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="columnifer">
<emphasis id="B96C184DE920476B9BB8FAC3FC0EFA6A" italics="true" pageId="15" pageNumber="432">Pr. columnifer</emphasis>
</taxonomicName>
by the talonid of p24 sub-triangular in outline; from
<taxonomicName id="4C18BFDCE920476B9D0AFA01FC03FA0B" authorityName="Ameghino" authorityYear="1889" box="[810,925,1460,1484]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="claudum">
<emphasis id="B96C184DE920476B9D0AFA01FC03FA0B" box="[810,925,1460,1484]" italics="true" pageId="15" pageNumber="432">Pr. claudum</emphasis>
</taxonomicName>
by having p23 with a distinctive entoflexid; from
<taxonomicName id="4C18BFDCE920476B9BB8FA01FC30FA2B" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="colloncurensis">
<emphasis id="B96C184DE920476B9BB8FA01FC30FA2B" italics="true" pageId="15" pageNumber="432">Pr. colloncurensis</emphasis>
</taxonomicName>
by a descending process of the maxilla with a moderate development; from
<taxonomicName id="4C18BFDCE920476B9A0FFA40FB5AF9CC" authorityName="Ameghino" authorityYear="1887" box="[1071,1220,1523,1547]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="praerutilum">
<emphasis id="B96C184DE920476B9A0FFA40FB5AF9CC" box="[1071,1220,1523,1547]" italics="true" pageId="15" pageNumber="432">Pr. praerutilum</emphasis>
</taxonomicName>
by having a larger size; from
<taxonomicName id="4C18BFDCE920476B9D42F9A7FC42F9ED" authorityName="Cabrera &amp; Kraglievich" authorityYear="1931" box="[866,988,1555,1578]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="minutum">
<emphasis id="B96C184DE920476B9D42F9A7FC42F9ED" box="[866,988,1555,1578]" italics="true" pageId="15" pageNumber="432">Pr. minutum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C18BFDCE920476B9A2EF9A7FB0AF9ED" authorityName="Cabrera and Kraglievich" authorityYear="1931" box="[1038,1172,1554,1578]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="distinctum">
<emphasis id="B96C184DE920476B9A2EF9A7FB0AF9ED" box="[1038,1172,1554,1578]" italics="true" pageId="15" pageNumber="432">Pr. distinctum</emphasis>
</taxonomicName>
by less cement covering the cheek teeth, the P24 with an entoflexus located in a more distal position that is slightly less expanded, M13 with straighter ectolophs, and infraorbital foramen above premolars; from
<taxonomicName id="4C18BFDCE920476B9BB8F9C5FC0EF96F" authorityName="Ameghino" authorityYear="1902" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="columnifer">
<emphasis id="B96C184DE920476B9BB8F9C5FC0EF96F" italics="true" pageId="15" pageNumber="432">Pr. columnifer</emphasis>
</taxonomicName>
by the talonid of p4 narrower than the trigonid; and from
<taxonomicName id="4C18BFDCE920476B9D41F904FC7EF900" authorityName="Ameghino" authorityYear="1885" box="[865,992,1712,1735]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="antiquum">
<emphasis id="B96C184DE920476B9D41F904FC7EF900" box="[865,992,1712,1735]" italics="true" pageId="15" pageNumber="432">Pr. antiquum</emphasis>
</taxonomicName>
by the talonid of m3 without its labial groove.
</paragraph>
</subSubSection>
<subSubSection id="C30297D4E920476B9D0AF95CFA2AF8E7" pageId="15" pageNumber="432" type="distribution">
<paragraph id="8BA7C45FE920476B9D0AF95CFA2AF8E7" blockId="15.[810,1460,1768,1824]" pageId="15" pageNumber="432">
<emphasis id="B96C184DE920476B9D0AF95CFB5AF8C7" box="[810,1220,1768,1792]" italics="true" pageId="15" pageNumber="432">Geographic and stratigraphic provenance:</emphasis>
<collectingRegion id="49DC0ABDE920476B9AFAF95DFA2DF8C7" box="[1242,1459,1768,1793]" country="Argentina" name="Santa Cruz" pageId="15" pageNumber="432">Santa Cruz Province</collectingRegion>
(
<collectingCountry id="F30F84CFE920476B9D11F8BCFC02F8E7" box="[817,924,1800,1824]" name="Argentina" pageId="15" pageNumber="432">Argentina</collectingCountry>
), SCF,
<location id="8EC79284E920476B9DC1F8BCFB54F8E7" LSID="urn:lsid:plazi:treatment:03B17549E92047749D54FD3AFEE8FE06:8EC79284E920476B9DC1F8BCFB54F8E7" box="[993,1226,1800,1824]" country="Argentina" municipality="Santacrucian" name="Early - Middle Miocene" pageId="15" pageNumber="432" stateProvince="Santa Cruz">EarlyMiddle Miocene</location>
,
<collectingMunicipality id="6BC35E25E920476B9AF2F8BCFACDF8E7" box="[1234,1363,1800,1824]" pageId="15" pageNumber="432">Santacrucian</collectingMunicipality>
SALMA.
</paragraph>
</subSubSection>
<subSubSection id="C30297D4E920476B9D0AF8F6FABEF89E" box="[810,1312,1857,1881]" pageId="15" pageNumber="432" type="description">
<paragraph id="8BA7C45FE920476B9D0AF8F6FABEF89E" blockId="15.[810,1312,1857,1881]" box="[810,1312,1857,1881]" pageId="15" pageNumber="432">
<emphasis id="B96C184DE920476B9D0AF8F6FC22F89E" box="[810,956,1858,1881]" italics="true" pageId="15" pageNumber="432">Measurements:</emphasis>
Supporting Information, Table S2.
</paragraph>
</subSubSection>
<subSubSection id="C30297D4E92047749D0AF8CEFEE8FE06" lastPageId="16" lastPageNumber="433" pageId="15" pageNumber="432" type="discussion">
<paragraph id="8BA7C45FE92047749D0AF8CEFEE8FE06" blockId="15.[810,1459,1914,1976]" lastBlockId="16.[129,780,144,450]" lastPageId="16" lastPageNumber="433" pageId="15" pageNumber="432">
<emphasis id="B96C184DE920476B9D0AF8CEFC16F855" box="[810,904,1914,1938]" italics="true" pageId="15" pageNumber="432">Remarks:</emphasis>
The ratio between the width and length of each upper molar (R
<subScript id="179CC61AE920476B9DEAF81EFC76F87F" attach="left" box="[970,1000,1962,1976]" fontSize="6" pageId="15" pageNumber="432">W/L</subScript>
) of
<taxonomicName id="4C18BFDCE920476B9A34F82FFB23F875" authorityName="Ameghino" authorityYear="1891" box="[1044,1213,1946,1970]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="15" pageNumber="432" phylum="Chordata" rank="species" species="compressidens">
<emphasis id="B96C184DE920476B9A34F82FFB23F875" box="[1044,1213,1946,1970]" italics="true" pageId="15" pageNumber="432">Pr. compressidens</emphasis>
</taxonomicName>
(
<typeStatus id="54A37AFDE920476B9AEAF82EFAB5F875" box="[1226,1323,1946,1970]" pageId="15" pageNumber="432" type="holotype">holotype</typeStatus>
plus referred materials) for M1 and M2 is &lt;0.5, whereas it is ≥
<quantity id="4CE069BAE93F47749CBCFF24FD7BFF6F" box="[668,741,144,168]" metricMagnitude="-2" metricUnit="m" metricValue="1.397" pageId="16" pageNumber="433" unit="in" value="0.55">0.55 in</quantity>
<taxonomicName id="4C18BFDCE93F47749CCFFF25FF53FF00" baseAuthorityName="Ameghino" baseAuthorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="16" pageNumber="433" phylum="Chordata" rank="species" species="australe">
<emphasis id="B96C184DE93F47749CCFFF25FF53FF00" italics="true" pageId="16" pageNumber="433">Pr. australe</emphasis>
</taxonomicName>
and
<taxonomicName id="4C18BFDCE93F47749F25FF04FE03FF00" authorityName="Ameghino" authorityYear="1887" box="[261,413,175,199]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="16" pageNumber="433" phylum="Chordata" rank="species" species="praerutilum">
<emphasis id="B96C184DE93F47749F25FF04FE03FF00" box="[261,413,175,199]" italics="true" pageId="16" pageNumber="433">Pr. praerutilum</emphasis>
</taxonomicName>
, and for M3 (with enlarged distal lobe), R
<subScript id="179CC61AE93F47749EF7FF6BFF6BFF2A" attach="left" box="[215,245,223,237]" fontSize="6" pageId="16" pageNumber="433">W/L</subScript>
≤ 0.42, whereas it is ≥
<quantity id="4CE069BAE93F47749FCFFF7BFDA4FF20" box="[495,570,207,231]" metricMagnitude="-2" metricUnit="m" metricValue="1.1938" pageId="16" pageNumber="433" unit="in" value="0.47">0.47 in</quantity>
<taxonomicName id="4C18BFDCE93F47749C64FF7BFD2AFF20" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[580,692,207,231]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="16" pageNumber="433" phylum="Chordata" rank="species" species="australe">
<emphasis id="B96C184DE93F47749C64FF7BFD2AFF20" box="[580,692,207,231]" italics="true" pageId="16" pageNumber="433">Pr. australe</emphasis>
</taxonomicName>
and
<taxonomicName id="4C18BFDCE93F47749CCFFF7BFF6BFEC1" authorityName="Ameghino" authorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="16" pageNumber="433" phylum="Chordata" rank="species" species="praerutilum">
<emphasis id="B96C184DE93F47749CCFFF7BFF6BFEC1" italics="true" pageId="16" pageNumber="433">Pr. praerutilum</emphasis>
</taxonomicName>
. The scaưer diagrams for M1 (
<figureCitation id="1323D8DAE93F47749C0BFF5AFDEAFEC1" box="[555,628,238,262]" captionStart="Figure 5" captionStartId="8.[129,194,1753,1777]" captionTargetBox="[257,1345,147,1723]" captionTargetId="figure-83@8.[254,1348,144,1726]" captionTargetPageId="8" captionText="Figure 5. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs.width of M1. B, Length vs. width of m1. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426276" httpUri="https://zenodo.org/record/8426276/files/figure.png" pageId="16" pageNumber="433">Fig. 5A</figureCitation>
) and M2 (
<figureCitation id="1323D8DAE93F47749CC3FF5AFF00FEE2" captionStart="Figure 6" captionStartId="9.[112,177,1757,1781]" captionTargetBox="[242,1330,147,1727]" captionTargetId="figure-87@9.[239,1333,144,1730]" captionTargetPageId="9" captionText="Figure 6. Scaưer diagram of the Santacrucian species of Protypotherium (except for Pr.claudum). A, Length vs. width of M2. B, Length vs. width of m2. Abbreviations: L, length; W, width." figureDoi="http://doi.org/10.5281/zenodo.8426278" httpUri="https://zenodo.org/record/8426278/files/figure.png" pageId="16" pageNumber="433">Fig. 6A</figureCitation>
) show the separation of these species in terms of the length and width of M12. A similar condition is observed in m1 and m2: R
<subScript id="179CC61AE93F47749EE2FEE8FF41FEAD" attach="left" box="[194,223,348,362]" fontSize="6" pageId="16" pageNumber="433">L/W</subScript>
<quantity id="4CE069BAE93F47749EDCFEF8FEDAFEA3" box="[252,324,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.0413999999999999" pageId="16" pageNumber="433" unit="in" value="0.41">0.41 in</quantity>
<taxonomicName id="4C18BFDCE93F47749F6CFEF9FE68FEA3" authorityName="Ameghino" authorityYear="1891" box="[332,502,332,356]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="16" pageNumber="433" phylum="Chordata" rank="species" species="compressidens">
<emphasis id="B96C184DE93F47749F6CFEF9FE68FEA3" box="[332,502,332,356]" italics="true" pageId="16" pageNumber="433">Pr. compressidens</emphasis>
</taxonomicName>
, whereas it is ≥
<quantity id="4CE069BAE93F47749CBFFEF8FD79FEA3" box="[671,743,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.1938" pageId="16" pageNumber="433" unit="in" value="0.47">0.47 in</quantity>
<taxonomicName id="4C18BFDCE93F47749CCFFEF9FF53FE44" baseAuthorityName="Ameghino" baseAuthorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="16" pageNumber="433" phylum="Chordata" rank="species" species="australe">
<emphasis id="B96C184DE93F47749CCFFEF9FF53FE44" italics="true" pageId="16" pageNumber="433">Pr. australe</emphasis>
</taxonomicName>
and
<taxonomicName id="4C18BFDCE93F47749F21FED8FE08FE44" authorityName="Ameghino" authorityYear="1887" box="[257,406,363,387]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="16" pageNumber="433" phylum="Chordata" rank="species" species="praerutilum">
<emphasis id="B96C184DE93F47749F21FED8FE08FE44" box="[257,406,363,387]" italics="true" pageId="16" pageNumber="433">Pr. praerutilum</emphasis>
</taxonomicName>
, and in m3 (with enlarged talonid), R
<subScript id="179CC61AE93F47749EB0FE2FFF31FE6E" attach="left" box="[144,175,411,425]" fontSize="6" pageId="16" pageNumber="433">W/L</subScript>
≤ 0.33 for
<taxonomicName id="4C18BFDCE93F47749F00FE3FFE56FE64" authorityName="Ameghino" authorityYear="1891" box="[288,456,395,419]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="16" pageNumber="433" phylum="Chordata" rank="species" species="compressidens">
<emphasis id="B96C184DE93F47749F00FE3FFE56FE64" box="[288,456,395,419]" italics="true" pageId="16" pageNumber="433">Pr. compressidens</emphasis>
</taxonomicName>
, whereas it is ≥ 0.35 for
<taxonomicName id="4C18BFDCE93F47749CE1FE3FFF33FE05" baseAuthorityName="Ameghino" baseAuthorityYear="1887" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="16" pageNumber="433" phylum="Chordata" rank="species" species="australe">
<emphasis id="B96C184DE93F47749CE1FE3FFF33FE05" italics="true" pageId="16" pageNumber="433">Pr. australe</emphasis>
</taxonomicName>
and
<taxonomicName id="4C18BFDCE93F47749EFDFE1FFEEFFE05" authorityName="Ameghino" authorityYear="1887" box="[221,369,426,450]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="16" pageNumber="433" phylum="Chordata" rank="species" species="praerutilum">
<emphasis id="B96C184DE93F47749EFDFE1FFEEFFE05" box="[221,369,426,450]" italics="true" pageId="16" pageNumber="433">Pr. praerutilum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>