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<document id="F74D31AB60ACC4A82E0BAFA85236D39E" ID-DOI="10.5252/zoosystema2024v46a7" ID-ISSN="1638-9387" ID-Zenodo-Dep="10863451" ID-ZooBank="A7CA7D85-2633-4930-BA12-ACFCB3D0DE21" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1711190289521" checkinUser="plazi" docAuthor="Boyko, Christopher B. &amp; Williams, Jason D." docDate="2024" docId="03B0992AFFACAB46FEB7F9FAE4CBFD65" docLanguage="en" docName="zoosystema.2024.46.7.pdf" docOrigin="Zoosystema 46 (7)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a7.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Rhopalione rusa Boyko &amp; Williams 2024, n. sp." docType="treatment" docUuid="C780B367-B63A-495A-9951-717DBE2F0549" docUuidSource="ZooBank" docVersion="2" lastPageNumber="172" masterDocId="FF89E152FFBAAB5FFFFAFF88E641FF35" masterDocTitle="New records of bopyrid parasites (Crustacea, Isopoda, Epicaridea) of pea crabs (Crustacea, Decapoda, Pinnotheridae) with descriptions of two new species of Rhopalione Pérez, 1921 and a review of the genus" masterLastPageNumber="175" masterPageNumber="149" pageNumber="169" updateTime="1711190782758" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="10DE54DC1598F78F2BB4694B9BF01B86">New records of bopyrid parasites (Crustacea, Isopoda, Epicaridea) of pea crabs (Crustacea, Decapoda, Pinnotheridae) with descriptions of two new species of Rhopalione Pérez, 1921 and a review of the genus</mods:title>
</mods:titleInfo>
<mods:name id="4AD26A7BD6C0CD74BE87A8371FD18046" type="personal">
<mods:role id="4AE44C13A109FFC8558FA431D4D7BC56">
<mods:roleTerm id="DF195A9D6E1B41895E89958A21E1F5DB">Author</mods:roleTerm>
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<mods:namePart id="26F28E5812DB46850E993489F05AEB6C">Boyko, Christopher B.</mods:namePart>
<mods:affiliation id="81AD9F0556B25C45E7FEC3A467C716F0">Department of Biology, Hofstra University, 1000 Hempstead Turnpike, Hempstead, NY 11549 (United States); and Division of Invertebrate Zoology, American Museum of Natural History, 200 Central Park West, New York, NY, 10024 (United States) cboyko @ amnh. org (corresponding author)</mods:affiliation>
<mods:nameIdentifier id="79E882A98E344147FCDF149FB65C1AFC" type="email">cboyko@amnh.org</mods:nameIdentifier>
</mods:name>
<mods:name id="D811547D2CF2D025FDA6E8CD94678FE5" type="personal">
<mods:role id="014F9CB6B86F6D94A1DC27CC4ABFF302">
<mods:roleTerm id="0CEA81450478693B4282AB79268E8FDE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="864A9ED4A774AAD48D39CAF46F2BB842">Williams, Jason D.</mods:namePart>
<mods:affiliation id="A614287D2354161E0FDC61E924CA856C">Department of Biology, Hofstra University, 1000 Hempstead Turnpike, Hempstead, NY 11549 (United States) jason. d. williams @ hofstra. edu</mods:affiliation>
<mods:nameIdentifier id="E910742056380987FC4BCC83093FA9D3" type="email">williams@hofstra.edu</mods:nameIdentifier>
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<mods:typeOfResource id="4FBF483F64013CA1EC8533B9B6289044">text</mods:typeOfResource>
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<mods:title id="64C13EC885DF9B207DB768123466895F">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="42BC18453CB4194BEF2ABDCB27BE2044">
<mods:date id="C406B84AF82F1BD131725A8A0D8A635C">2024</mods:date>
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<mods:number id="1059BAF2310D7D37F408F075678C0AD8">2024-03-22</mods:number>
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<mods:number id="498892DBF4CF21B334A7BA309823E240">46</mods:number>
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<mods:url id="31978BA8819EF6723D7646FDC5FFC48E">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a7.pdf</mods:url>
</mods:location>
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<mods:identifier id="3C85F54F27F9E4076D6D16BD56BB2B59" type="DOI">10.5252/zoosystema2024v46a7</mods:identifier>
<mods:identifier id="DE0596C667FDD606F9564987A884EE4E" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="CDA68F8461490950C32A902573CF97F1" type="Zenodo-Dep">10863451</mods:identifier>
<mods:identifier id="4BAEEBF7B14C613886B6EF74DE18B068" type="ZooBank">A7CA7D85-2633-4930-BA12-ACFCB3D0DE21</mods:identifier>
</mods:mods>
<treatment id="03B0992AFFACAB46FEB7F9FAE4CBFD65" LSID="urn:lsid:zoobank.org:act:C780B367-B63A-495A-9951-717DBE2F0549" httpUri="http://treatment.plazi.org/id/03B0992AFFACAB46FEB7F9FAE4CBFD65" lastPageId="25" lastPageNumber="172" pageId="22" pageNumber="169">
<subSubSection id="C3037BB7FFACAB49FEB7F9FAE47FF9B8" box="[333,574,1650,1677]" pageId="22" pageNumber="169" type="nomenclature">
<paragraph id="8BA6283CFFACAB49FEB7F9FAE47FF9B8" blockId="22.[333,574,1650,1708]" box="[333,574,1650,1677]" pageId="22" pageNumber="169">
<heading id="D0EE9F50FFACAB49FEB7F9FAE47FF9B8" box="[333,574,1650,1677]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="169" reason="2">
<taxonomicName id="4C1953BFFFACAB49FEB7F9FAE7BDF9B8" authority="Boyko &amp; Williams, 2024" authorityName="Boyko &amp; Williams" authorityYear="2024" box="[333,508,1650,1677]" class="Malacostraca" family="Bopyridae" genus="Rhopalione" kingdom="Animalia" order="Isopoda" pageId="22" pageNumber="169" phylum="Arthropoda" rank="species" species="rusa" status="sp. nov.">
<emphasis id="B96DF42EFFACAB49FEB7F9FAE7BDF9B8" bold="true" box="[333,508,1650,1677]" italics="true" pageId="22" pageNumber="169">Rhopalione rusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4955FFACAB49FDF9F9FBE47FF9B8" box="[515,574,1651,1677]" pageId="22" pageNumber="169" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3037BB7FFACAB49FE7BF91AE6A4F8B0" pageId="22" pageNumber="169" type="description">
<paragraph id="8BA6283CFFACAB49FE7BF91AE44BF999" blockId="22.[333,574,1650,1708]" box="[385,522,1682,1708]" pageId="22" pageNumber="169">
(
<figureCitation id="132234B9FFACAB49FE72F91AE798F999" box="[392,473,1682,1708]" captionStart="FIG" captionStartId="23.[132,143,1779,1797]" captionTargetBox="[132,1455,95,1732]" captionTargetId="figure-18@23.[132,1455,215,1732]" captionTargetPageId="23" captionText="FIG. 10. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259) and allotype (ZRC 2023.0260) specimens: A, ventral view of host pinnotherid Arcotheres similis (Bürger, 1895) with Rhopalione sp. in situ on left side of pleon (posterior end of male shown by arrow);B, holotype female, dorsal view; C, holotype female,ventral view; D, allotype male, dorsal view. Scale bars: A, 2 mm; B-D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10863477" httpUri="https://zenodo.org/record/10863477/files/figure.png" pageId="22" pageNumber="169">Figs 10</figureCitation>
;
<figureCitation id="132234B9FFACAB49FE1FF91AE440F999" box="[485,513,1682,1708]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="22" pageNumber="169">11</figureCitation>
)
</paragraph>
<paragraph id="8BA6283CFFACAB49FF5BF95EE4ABF9DD" blockId="22.[161,746,1750,1768]" box="[161,746,1750,1768]" pageId="22" pageNumber="169">
<uri id="FF88243EFFACAB49FF5BF95EE4ABF9DD" box="[161,746,1750,1768]" pageId="22" pageNumber="169">
urn:lsid:zoobank.org:act:
<uuid id="FFBF12E9FFACAB49FE9DF95EE4ABF9DD" box="[359,746,1750,1768]" pageId="22" pageNumber="169">C780B367-B63A-495A-9951-717DBE2F0549</uuid>
</uri>
</paragraph>
<paragraph id="8BA6283CFFACAB49FF7EF881E458F80E" blockId="22.[132,775,1801,1851]" pageId="22" pageNumber="169">
“bopyrids under pleon”
<bibRefCitation id="EF8855CDFFACAB49FE62F881E4E5F814" author="NG P. K. L. &amp; AHYONG S. T." box="[408,676,1801,1825]" pageId="22" pageNumber="169" pagination="134 - 248" refId="ref22405" refString="NG P. K. L. &amp; AHYONG S. T. 2022. - The pea crab genus Arcotheres Manning, 1993 (Crustacea: Brachyura: Pinnotheridae) from Singapore and Peninsular Malaysia, with a reappraisal of diagnostic characters and descriptions of two new genera. Raffles Bulletin of Zoology 70: 134 - 248. https: // doi. org / 10.26107 / RBZ- 2022 - 0009" type="journal article" year="2022">Ng &amp; Ahyong 2022: 173</bibRefCitation>
[ex ZRC 2019.0534; specimens examined herein].
</paragraph>
<paragraph id="8BA6283CFFACAB49FF7EF8DBE6A4F8B0" blockId="22.[132,775,1875,1925]" pageId="22" pageNumber="169">
“unidentified bopyrid” [of
<taxonomicName id="4C1953BFFFACAB49FE7FF8DBE463F85E" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[389,546,1875,1899]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="169" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B96DF42EFFACAB49FE7FF8DBE463F85E" box="[389,546,1875,1899]" italics="true" pageId="22" pageNumber="169">Arcotheres similis</emphasis>
</taxonomicName>
]
<bibRefCitation id="EF8855CDFFACAB49FDBAF8DBE6EBF8B0" author="WILLIAMS J. D. &amp; BOYKO C. B. &amp; TRI N. V." pageId="22" pageNumber="169" pagination="531 - 541" refId="ref24339" refString="WILLIAMS J. D., BOYKO C. B. &amp; TRI N. V. 2023. - Description of a new genus and species of bopyrid (Isopoda: Epicaridea: Bopyridae) from the pinnotherid crab Plenotheres coarctatus (Burger, 1895) associated with mangrove clams from Vietnam. Raffles Bulletin of Zoology 71: 531 - 541. https: // doi. org / 10.26107 / RBZ- 2023 - 0040" type="journal article" year="2023">
Williams
<emphasis id="B96DF42EFFACAB49FD67F8DCE48BF85E" box="[669,714,1875,1899]" italics="true" pageId="22" pageNumber="169">et al.</emphasis>
2023: 533
</bibRefCitation>
[list].
</paragraph>
</subSubSection>
<subSubSection id="C3037BB7FFACAB49FF7EF816E3E5FE46" pageId="22" pageNumber="169" type="materials_examined">
<paragraph id="8BA6283CFFACAB49FF7EF816E359FE11" blockId="22.[132,776,1949,2026]" lastBlockId="22.[811,1458,215,504]" pageId="22" pageNumber="169">
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<smallCapsWord id="8D40BEE0FFACAB49FF47F82BE75AF881" baselines="1968" box="[189,283,1955,1972]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="22" pageNumber="169">MATERIAL</smallCapsWord>
. —
<emphasis id="B96DF42EFFACAB49FEB0F815E7E8F880" bold="true" box="[330,425,1949,1973]" pageId="22" pageNumber="169">
<typeStatus id="54A2969EFFACAB49FEB0F815E7E8F880" box="[330,425,1949,1973]" pageId="22" pageNumber="169" type="holotype">Holotype</typeStatus>
</emphasis>
.
<emphasis id="B96DF42EFFACAB49FE4CF817E458F882" bold="true" box="[438,537,1951,1975]" pageId="22" pageNumber="169">
<collectingCountry id="F30E68ACFFACAB49FE4CF817E458F882" box="[438,537,1951,1975]" name="Singapore" pageId="22" pageNumber="169">Singapore</collectingCountry>
</emphasis>
• ovigerous
<specimenCount id="9D1FE3B5FFACAB49FD6DF828E4E9F881" box="[663,680,1952,1972]" count="1" pageId="22" pageNumber="169" type="female"></specimenCount>
(
<quantity id="4CE185D9FFACAB49FD4DF815E546F880" box="[695,775,1949,1973]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="22" pageNumber="169" unit="mm" value="6.8">6.8 mm</quantity>
TL), from left side of pleon of female
<taxonomicName id="4C1953BFFFACAB49FDF6F830E680F8DF" authority="(Burger, 1895)" baseAuthorityName="Burger" baseAuthorityYear="1895" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="169" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B96DF42EFFACAB49FDF6F830E4EEF8E5" box="[524,687,1976,2000]" italics="true" pageId="22" pageNumber="169">Arcotheres similis</emphasis>
(Bürger, 1895)
</taxonomicName>
(
<collectionCode id="ED08B0F9FFACAB49FF34F85AE740F8DF" box="[206,257,2002,2026]" country="Singapore" name="Zoological Reference Collection, National University of Singapore" pageId="22" pageNumber="169">ZRC</collectionCode>
2019.0534; 6.0 mm CL,
<quantity id="4CE185D9FFACAB49FDF8F85AE411F8DF" box="[514,592,2002,2026]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="22" pageNumber="169" unit="mm" value="7.6">7.6 mm</quantity>
CW), ex
<taxonomicName id="4C1953BFFFACAB49FD4AF85AE20EFFDA" authority="Hanley, 1843" authorityName="Hanley" authorityYear="1843" box="[688,1103,215,2026]" class="Bivalvia" family="Mytilidae" genus="Modiolus" kingdom="Animalia" order="Mytilida" pageId="22" pageNumber="169" phylum="Mollusca" rank="species" species="philippinarum">
<emphasis id="B96DF42EFFACAB49FD4AF85AE5FDFFDA" box="[688,956,215,2026]" italics="true" pageId="22" pageNumber="169">Modiolus philippinarum</emphasis>
Hanley, 1843
</taxonomicName>
(
<taxonomicName id="4C1953BFFFACAB49FBA4FF5FE33EFFDA" authority="Rafinesque, 1815" authorityName="Rafinesque" authorityYear="1815" box="[1118,1407,215,239]" class="Bivalvia" family="Mytilidae" kingdom="Animalia" order="Mytilida" pageId="22" pageNumber="169" phylum="Mollusca" rank="family">
Mytilidae
<bibRefCitation id="EF8855CDFFACAB49FB30FF5FE33EFFDA" author="RAFINESQUE C. S." box="[1226,1407,215,239]" pageId="22" pageNumber="169" refId="ref23178" refString="RAFINESQUE C. S. 1815. - Analyse de la nature ou Tableau de l'univers et des corps organises. Palerme, 224 p. https: // doi. org / 10.5962 / bhl. title. 106607" type="book" year="1815">Rafinesque, 1815</bibRefCitation>
</taxonomicName>
); off Changi Beach, intertidal area next to Carpark 7, near ferry terminal;
<date id="FFA70EFCFFACAB49FCD1FE84E5DCFE11" box="[811,925,268,292]" pageId="22" pageNumber="169" value="2019-04-22">22.IV.2019</date>
; S. K. Tan
<emphasis id="B96DF42EFFACAB49FBF0FE85E278FE11" box="[1034,1081,268,292]" italics="true" pageId="22" pageNumber="169">et al.</emphasis>
leg.; ZRC 2023.0259.
</paragraph>
<paragraph id="8BA6283CFFACAB49FCD7FEA0E3E5FE46" blockId="22.[811,1458,215,504]" pageId="22" pageNumber="169">
<materialsCitation id="3B712261FFACAB49FCD7FEA0E3E5FE46" collectingDate="2019-04-22" collectorName="S. K. Tan" country="Singapore" location="Mature" pageId="22" pageNumber="169" specimenCount="1" specimenCount-male="1" typeStatus="allotype">
<emphasis id="B96DF42EFFACAB49FCD7FEA0E5C4FE75" bold="true" box="[813,901,296,320]" pageId="22" pageNumber="169">
<typeStatus id="54A2969EFFACAB49FCD7FEA0E5C4FE75" box="[813,901,296,320]" pageId="22" pageNumber="169" type="allotype">Allotype</typeStatus>
</emphasis>
.
<emphasis id="B96DF42EFFACAB49FC6EFEA0E5BAFE75" bold="true" box="[916,1019,296,320]" pageId="22" pageNumber="169">
<collectingCountry id="F30E68ACFFACAB49FC6EFEA0E5BAFE75" box="[916,1019,296,320]" name="Singapore" pageId="22" pageNumber="169">Singapore</collectingCountry>
</emphasis>
<location id="8EC67EE7FFACAB49FBECFEAFE220FE0B" LSID="urn:lsid:plazi:treatment:03B0992AFFACAB46FEB7F9FAE4CBFD65:8EC67EE7FFACAB49FBECFEAFE220FE0B" box="[1046,1121,295,318]" country="Singapore" name="Mature" pageId="22" pageNumber="169">Mature</location>
<specimenCount id="9D1FE3B5FFACAB49FB93FEA1E23CFE08" box="[1129,1149,297,317]" count="1" pageId="22" pageNumber="169" type="male"></specimenCount>
(
<quantity id="4CE185D9FFACAB49FB74FEAEE2A0FE0B" box="[1166,1249,294,318]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="22" pageNumber="169" unit="mm" value="4.1">4.1 mm</quantity>
TL), collected with holotype; off Changi Beach, intertidal area next to Carpark 7, near ferry terminal;
<date id="FFA70EFCFFACAB49FC3AFED3E273FE46" box="[960,1074,347,371]" pageId="22" pageNumber="169" value="2019-04-22">
<collectingDate id="EFE3F714FFACAB49FC3AFED3E273FE46" box="[960,1074,347,371]" pageId="22" pageNumber="169" value="2019-04-22">22.IV.2019</collectingDate>
</date>
;
<collectorName id="26EC4DEAFFACAB49FBC6FED3E2D7FE46" box="[1084,1174,347,371]" pageId="22" pageNumber="169">S. K. Tan</collectorName>
<emphasis id="B96DF42EFFACAB49FB67FED4E28DFE46" box="[1181,1228,347,371]" italics="true" pageId="22" pageNumber="169">et al.</emphasis>
leg.; ZRC 2023.0260
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3037BB7FFACAB49FCD7FEFEE3E8FEC2" pageId="22" pageNumber="169" type="description">
<paragraph id="8BA6283CFFACAB49FCD7FEFEE3E8FEC2" blockId="22.[811,1458,215,504]" pageId="22" pageNumber="169">
<emphasis id="B96DF42EFFACAB49FCD7FEFEE5C5FEBB" bold="true" box="[813,900,374,398]" pageId="22" pageNumber="169">
<typeStatus id="54A2969EFFACAB49FCD7FEFEE5C5FEBB" box="[813,900,374,398]" pageId="22" pageNumber="169" type="paratype">Paratype</typeStatus>
</emphasis>
.
<emphasis id="B96DF42EFFACAB49FC6BFEFFE5B5FEBA" bold="true" box="[913,1012,375,399]" pageId="22" pageNumber="169">
<collectingCountry id="F30E68ACFFACAB49FC6BFEFFE5B5FEBA" box="[913,1012,375,399]" name="Singapore" pageId="22" pageNumber="169">Singapore</collectingCountry>
</emphasis>
• 1 ovigerous
<specimenCount id="9D1FE3B5FFACAB49FB7CFEF1E2D6FEB8" box="[1158,1175,377,397]" count="1" pageId="22" pageNumber="169" type="female"></specimenCount>
(
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TL; male not present), from right side of pleon of female
<taxonomicName id="4C1953BFFFACAB49FB5DFE18E2BDFE9D" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1191,1276,400,424]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="169" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B96DF42EFFACAB49FB5DFE18E2F7FE92" box="[1191,1206,400,423]" italics="true" pageId="22" pageNumber="169">A</emphasis>
.
<emphasis id="B96DF42EFFACAB49FB3BFE18E2BDFE9D" box="[1217,1276,400,424]" italics="true" pageId="22" pageNumber="169">similis</emphasis>
</taxonomicName>
(ZRC 2019.0534;
<quantity id="4CE185D9FFACAB49FCD1FE22E537FEF7" box="[811,886,426,450]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="22" pageNumber="169" unit="mm" value="5.6">5.6 mm</quantity>
CL,
<quantity id="4CE185D9FFACAB49FC5EFE22E5AEFEF7" box="[932,1007,426,450]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="22" pageNumber="169" unit="mm" value="7.5">7.5 mm</quantity>
CW), ex
<taxonomicName id="4C1953BFFFACAB49FBB1FE22E3EEFEF6" authority="Hanley, 1843" authorityName="Hanley" authorityYear="1843" box="[1099,1455,426,451]" class="Bivalvia" family="Mytilidae" genus="Modiolus" kingdom="Animalia" order="Mytilida" pageId="22" pageNumber="169" phylum="Mollusca" rank="species" species="philippinarum">
<emphasis id="B96DF42EFFACAB49FBB1FE22E36AFEF7" box="[1099,1323,426,450]" italics="true" pageId="22" pageNumber="169">Modiolus philippinarum</emphasis>
Hanley, 1843
</taxonomicName>
(
<taxonomicName id="4C1953BFFFACAB49FCCBFE4DE5D3FEE8" authorityName="Rafinesque" authorityYear="1815" box="[817,914,453,477]" class="Bivalvia" family="Mytilidae" kingdom="Animalia" order="Mytilida" pageId="22" pageNumber="169" phylum="Mollusca" rank="family">Mytilidae</taxonomicName>
); off Changi Beach, intertidal area next to Carpark 7, near ferry terminal;
<date id="FFA70EFCFFACAB49FC44FE68E271FECD" box="[958,1072,480,504]" pageId="22" pageNumber="169" value="2019-04-22">22.IV.2019</date>
; S. K. Tan
<emphasis id="B96DF42EFFACAB49FB61FE68E28AFEC2" box="[1179,1227,479,503]" italics="true" pageId="22" pageNumber="169">et al.</emphasis>
leg.; ZRC 2023.0261.
</paragraph>
</subSubSection>
<subSubSection id="C3037BB7FFACAB49FCD7FD98E3DFFDCE" pageId="22" pageNumber="169" type="etymology">
<paragraph id="8BA6283CFFACAB49FCD7FD98E3DFFDCE" blockId="22.[811,1457,527,763]" pageId="22" pageNumber="169">
<smallCapsWord id="8D40BEE0FFACAB49FCD7FD98E5E8FD13" baselines="545,546" box="[813,937,528,551]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="22" pageNumber="169">ETYMOLOGY</smallCapsWord>
. — The species is named after the
<typeStatus id="54A2969EFFACAB49FAECFD98E300FD1D" box="[1302,1345,528,552]" pageId="22" pageNumber="169">type</typeStatus>
locality of Changi Beach which, in 1604, was originally called “Tanjong Rusa” (tanjong meaning cape and rusa being Malay for deer which could refer to the extinct Indian muntjac (
<taxonomicName id="4C1953BFFFACAB49FB74FDD7E5E4FDA4" authority="(Zimmerman, 1780))" baseAuthorityName="Zimmerman" baseAuthorityYear="1780" class="Mammalia" family="Cervidae" genus="Muntiacus" kingdom="Animalia" order="Artiodactyla" pageId="22" pageNumber="169" phylum="Chordata" rank="species" species="muntjak">
<emphasis id="B96DF42EFFACAB49FB74FDD7E309FD43" box="[1166,1352,606,630]" italics="true" pageId="22" pageNumber="169">Muntiacus muntjak</emphasis>
(Zimmerman, 1780))
</taxonomicName>
or the once thought extinct Sambar deer (
<emphasis id="B96DF42EFFACAB49FAC9FDF2E3F1FDA4" box="[1331,1456,633,657]" italics="true" pageId="22" pageNumber="169">Rusa unicolor</emphasis>
Kerr, 1792). The change of name from Tanjong Rusa to Tanjong Changi occurred between 1604 and 1828 (
<bibRefCitation id="EF8855CDFFACAB49FB0BFD26E3E8FDF3" author="GIBSON-HILL C. A." box="[1265,1449,686,710]" pageId="22" pageNumber="169" pagination="163 - 214" refId="ref20688" refString="GIBSON-HILL C. A. 1954. - Singapore: Notes on the History of the Old Strait, 1580 - 1850. Journal of the Malayan Branch of the Royal Asiatic Society 27 (1): 163 - 214. https: // www. jstor. org / stable / 41486179" type="journal article" year="1954">Gibson-Hill 1954</bibRefCitation>
;
<bibRefCitation id="EF8855CDFFACAB49FCD7FD40E597FDD5" author="BORSCHBERG P." box="[813,982,712,736]" pageId="22" pageNumber="169" pagination="373 - 390" refId="ref19545" refString="BORSCHBERG P. 2017. - Singapore and its Straits, c. 1500 - 1800. Indonesia and the Malay World 45 (133): 373 - 390. https: // doi. org / 10.1080 / 13639811.2017.1340493" type="journal article" year="2017">Borschberg 2017</bibRefCitation>
; https://eresources.nlb.gov.sg/infopedia/articles/ SIP_245_
<date id="FFA70EFCFFACAB49FC6EFD6BE248FDCE" box="[916,1033,739,763]" pageId="22" pageNumber="169" value="2004-12-15">2004-12-15.</date>
html). The name is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3037BB7FFACAB47FCD7FCBEE36FF979" lastPageId="24" lastPageNumber="171" pageId="22" pageNumber="169" type="description">
<paragraph id="8BA6283CFFACAB49FCD7FCBEE3F0F8DE" blockId="22.[810,1457,822,2028]" pageId="22" pageNumber="169">
<smallCapsWord id="8D40BEE0FFACAB49FCD7FCBEE5FCFC7B" baselines="841,841" box="[813,957,822,848]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="22" pageNumber="169">DESCRIPTION</smallCapsWord>
<emphasis id="B96DF42EFFACAB49FCD7FCDDE537FC5A" box="[813,886,853,879]" italics="true" pageId="22" pageNumber="169">Female</emphasis>
<typeStatus id="54A2969EFFACAB49FCD7FCFDE5D2FCBA" box="[813,915,885,911]" pageId="22" pageNumber="169" type="holotype">Holotype</typeStatus>
(
<figureCitation id="132234B9FFACAB49FC59FCFDE5B3FCBA" box="[931,1010,885,911]" captionStart="FIG" captionStartId="23.[132,143,1779,1797]" captionTargetBox="[132,1455,95,1732]" captionTargetId="figure-18@23.[132,1455,215,1732]" captionTargetPageId="23" captionText="FIG. 10. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259) and allotype (ZRC 2023.0260) specimens: A, ventral view of host pinnotherid Arcotheres similis (Bürger, 1895) with Rhopalione sp. in situ on left side of pleon (posterior end of male shown by arrow);B, holotype female, dorsal view; C, holotype female,ventral view; D, allotype male, dorsal view. Scale bars: A, 2 mm; B-D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10863477" httpUri="https://zenodo.org/record/10863477/files/figure.png" pageId="22" pageNumber="169">Figs 10</figureCitation>
A-C; 11A-G). Length
<quantity id="4CE185D9FFACAB49FB11FCFDE307FCBA" box="[1259,1350,885,911]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="22" pageNumber="169" unit="mm" value="6.8">6.8 mm</quantity>
; maximal width at posterior margin of pereomere 4 5.0 mm (not including extended coxal plates); head length
<quantity id="4CE185D9FFACAB49FB29FC3DE36DFCFA" box="[1235,1324,949,975]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="22" pageNumber="169" unit="mm" value="0.9">0.9 mm</quantity>
; head width
<quantity id="4CE185D9FFACAB49FCD7FC5DE5C9FCDA" box="[813,904,981,1007]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="22" pageNumber="169" unit="mm" value="1.4">1.4 mm</quantity>
. Body nearly straight (
<figureCitation id="132234B9FFACAB49FB7BFC5CE344FCDA" box="[1153,1285,980,1007]" captionStart="FIG" captionStartId="23.[132,143,1779,1797]" captionTargetBox="[132,1455,95,1732]" captionTargetId="figure-18@23.[132,1455,215,1732]" captionTargetPageId="23" captionText="FIG. 10. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259) and allotype (ZRC 2023.0260) specimens: A, ventral view of host pinnotherid Arcotheres similis (Bürger, 1895) with Rhopalione sp. in situ on left side of pleon (posterior end of male shown by arrow);B, holotype female, dorsal view; C, holotype female,ventral view; D, allotype male, dorsal view. Scale bars: A, 2 mm; B-D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10863477" httpUri="https://zenodo.org/record/10863477/files/figure.png" pageId="22" pageNumber="169">Fig. 10A, B</figureCitation>
), subcircular in outline but approximately as wide as long when including coxal plates, not distorted; all somites distinct. Head ovate, separated from pereon, 1.6 × wider than long, anterior and posterior margins convex, lateral margins rounded; wide frontal lamina. Eyes absent. Antennules (
<figureCitation id="132234B9FFACAB49FB6DFBFCE2B6FBBB" box="[1175,1271,1140,1166]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="22" pageNumber="169">Fig. 11A</figureCitation>
) of three articles each, antennae (
<figureCitation id="132234B9FFACAB49FC24FB1CE201FB9B" box="[990,1088,1172,1198]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="22" pageNumber="169">Fig. 11A</figureCitation>
) of five articles each, both visible dorsally, few setae distally. Maxilliped (
<figureCitation id="132234B9FFACAB49FB3FFB3CE366FBFB" box="[1221,1319,1204,1230]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="22" pageNumber="169">Fig. 11D</figureCitation>
) longer than wide; broad non-articulated palp with blunt apex; spur elongate, recurved, distally triangular (
<figureCitation id="132234B9FFACAB49FB68FB7BE2B2FA3B" box="[1170,1267,1267,1294]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="22" pageNumber="169">Fig. 11D</figureCitation>
inset), covered by large round accessory lobe. Barbula (
<figureCitation id="132234B9FFACAB49FB4EFA9BE353FA1B" box="[1204,1298,1299,1326]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="22" pageNumber="169">Fig. 11B</figureCitation>
) laterally with pair of elongate, smooth, broad falcate projections, distally tapering; shallowly convex medially. Pereon nearly straight (
<figureCitation id="132234B9FFACAB49FCCCFAFBE5D8FAB8" box="[822,921,1395,1421]" captionStart="FIG" captionStartId="23.[132,143,1779,1797]" captionTargetBox="[132,1455,95,1732]" captionTargetId="figure-18@23.[132,1455,215,1732]" captionTargetPageId="23" captionText="FIG. 10. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259) and allotype (ZRC 2023.0260) specimens: A, ventral view of host pinnotherid Arcotheres similis (Bürger, 1895) with Rhopalione sp. in situ on left side of pleon (posterior end of male shown by arrow);B, holotype female, dorsal view; C, holotype female,ventral view; D, allotype male, dorsal view. Scale bars: A, 2 mm; B-D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10863477" httpUri="https://zenodo.org/record/10863477/files/figure.png" pageId="22" pageNumber="169">Fig. 10B</figureCitation>
). Pereomeres dorsally distinct, produced laterally into blunt, rounded lobes; widest at pereomere 4; margins weakly curved, mid-dorsal bosses or projections absent; ovoid dorsolateral bosses on pereomeres 1-6, round on pereomere 7, all dorsolateral bosses distinctly separated from lateral portion of pereomeres; ovoid coxal plates on all seven pereomeres, shorter than dorsolateral bosses on pereomeres 1-4, longer than dorsolateral bosses on pereomeres 5-7, dorsolateral bosses and coxal plates in posterior pereomeres not fused. Oostegites completely enclosing brood pouch (
<figureCitation id="132234B9FFACAB49FB63F91AE2B6F999" box="[1177,1271,1682,1708]" captionStart="FIG" captionStartId="23.[132,143,1779,1797]" captionTargetBox="[132,1455,95,1732]" captionTargetId="figure-18@23.[132,1455,215,1732]" captionTargetPageId="23" captionText="FIG. 10. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259) and allotype (ZRC 2023.0260) specimens: A, ventral view of host pinnotherid Arcotheres similis (Bürger, 1895) with Rhopalione sp. in situ on left side of pleon (posterior end of male shown by arrow);B, holotype female, dorsal view; C, holotype female,ventral view; D, allotype male, dorsal view. Scale bars: A, 2 mm; B-D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10863477" httpUri="https://zenodo.org/record/10863477/files/figure.png" pageId="22" pageNumber="169">Fig. 10C</figureCitation>
), strongly vaulted ventrally, not protruding beyond anterior margins of body, not visible dorsally. Oostegite 1 (
<figureCitation id="132234B9FFACAB49FB64F95AE35EF9D9" box="[1182,1311,1746,1772]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="22" pageNumber="169">Fig. 11E, F</figureCitation>
) longer than wide; anterior lobe rounded, larger than distal lobe; internal ridge with few irregular low digitations (
<figureCitation id="132234B9FFACAB49FB09F89AE314F819" box="[1267,1365,1810,1836]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="22" pageNumber="169">Fig. 11F</figureCitation>
); posterior lobe with acute, triangular distal projection on margin. Oostegite 5 (
<figureCitation id="132234B9FFACAB49FC40F8D9E25AF859" box="[954,1051,1873,1900]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="22" pageNumber="169">Fig. 11C</figureCitation>
) posterior margin fringed with setae. Pereopods isomorphic, subchelate (
<figureCitation id="132234B9FFACAB49FB67F8F9E35BF8BE" box="[1181,1306,1905,1931]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="22" pageNumber="169">Fig. 11E, G</figureCitation>
), small curved dactylus, propodus ovate with scales around region that meets tip of dactylus, carpus and merus triangular, ischium and basis tubular, carpus with distal setae and scales. Pleon short
</paragraph>
<caption id="DF6678B4FFADAB48FF7EF97BE483F806" ID-DOI="http://doi.org/10.5281/zenodo.10863477" ID-Zenodo-Dep="10863477" httpUri="https://zenodo.org/record/10863477/files/figure.png" pageId="23" pageNumber="170" startId="23.[132,143,1779,1797]" targetBox="[132,1455,95,1732]" targetPageId="23" targetType="figure">
<paragraph id="8BA6283CFFADAB48FF7EF97BE483F806" blockId="23.[132,1455,1779,1843]" pageId="23" pageNumber="170">
<smallCapsWord id="8D40BEE0FFADAB48FF7EF97BE6DCF831" baselines="1792,1793" box="[132,157,1779,1797]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="23" pageNumber="170">FIG</smallCapsWord>
. 10.
<emphasis id="B96DF42EFFADAB48FF32F97BE69AF830" bold="true" box="[200,219,1779,1797]" pageId="23" pageNumber="170"></emphasis>
<taxonomicName id="4C1953BFFFADAB48FF1BF97BE728F830" authority="Boyko &amp; Williams, 2024" authorityName="Boyko &amp; Williams" authorityYear="2024" box="[225,361,1779,1797]" class="Malacostraca" family="Bopyridae" genus="Rhopalione" kingdom="Animalia" order="Isopoda" pageId="23" pageNumber="170" phylum="Arthropoda" rank="species" species="rusa" status="sp. nov.">
<emphasis id="B96DF42EFFADAB48FF1BF97BE728F830" box="[225,361,1779,1797]" italics="true" pageId="23" pageNumber="170">Rhopalione rusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4955FFADAB48FE8AF97BE7E0F830" box="[368,417,1779,1797]" pageId="23" pageNumber="170" rank="species">n. sp.</taxonomicNameLabel>
holotype (ZRC 2023.0259) and allotype (ZRC 2023.0260) specimens:
<emphasis id="B96DF42EFFADAB48FC08F97BE5BEF830" bold="true" box="[1010,1023,1779,1797]" pageId="23" pageNumber="170">A</emphasis>
, ventral view of host pinnotherid
<taxonomicName id="4C1953BFFFADAB48FAE6F97BE6B6F829" authority="(Burger, 1895)" baseAuthorityName="Burger" baseAuthorityYear="1895" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="170" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B96DF42EFFADAB48FAE6F97BE3EEF830" box="[1308,1455,1779,1797]" italics="true" pageId="23" pageNumber="170">Arcotheres similis</emphasis>
(Bürger, 1895)
</taxonomicName>
with
<taxonomicName id="4C1953BFFFADAB48FED8F882E7E1F829" box="[290,416,1802,1820]" class="Malacostraca" family="Bopyridae" genus="Rhopalione" kingdom="Animalia" order="Isopoda" pageId="23" pageNumber="170" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B96DF42EFFADAB48FED8F882E7C1F829" box="[290,384,1802,1820]" italics="true" pageId="23" pageNumber="170">Rhopalione</emphasis>
sp.
</taxonomicName>
<emphasis id="B96DF42EFFADAB48FE59F882E793F829" box="[419,466,1802,1820]" italics="true" pageId="23" pageNumber="170">in situ</emphasis>
on left side of pleon (posterior end of male shown by arrow);
<emphasis id="B96DF42EFFADAB48FC3CF882E592F829" bold="true" box="[966,979,1802,1820]" pageId="23" pageNumber="170">B</emphasis>
, holotype female, dorsal view;
<emphasis id="B96DF42EFFADAB48FB28F882E2A1F829" bold="true" box="[1234,1248,1802,1820]" pageId="23" pageNumber="170">C</emphasis>
, holotype female,ventral view;
<emphasis id="B96DF42EFFADAB48FF4EF8A9E683F806" bold="true" box="[180,194,1825,1843]" pageId="23" pageNumber="170">D</emphasis>
, allotype male, dorsal view. Scale bars: A, 2 mm; B-D, 1 mm.
</paragraph>
</caption>
<paragraph id="8BA6283CFFADAB48FF7EF8F9E24EF8D9" blockId="23.[132,777,1905,2027]" lastBlockId="23.[811,1455,1905,2028]" pageId="23" pageNumber="170">
(
<figureCitation id="132234B9FFADAB48FF76F8F9E6B0F8BE" box="[140,241,1905,1931]" captionStart="FIG" captionStartId="23.[132,143,1779,1797]" captionTargetBox="[132,1455,95,1732]" captionTargetId="figure-18@23.[132,1455,215,1732]" captionTargetPageId="23" captionText="FIG. 10. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259) and allotype (ZRC 2023.0260) specimens: A, ventral view of host pinnotherid Arcotheres similis (Bürger, 1895) with Rhopalione sp. in situ on left side of pleon (posterior end of male shown by arrow);B, holotype female, dorsal view; C, holotype female,ventral view; D, allotype male, dorsal view. Scale bars: A, 2 mm; B-D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10863477" httpUri="https://zenodo.org/record/10863477/files/figure.png" pageId="23" pageNumber="170">Fig. 10B</figureCitation>
), five pleomeres, all dorsally distinct, each with lateral plates produced into long, slender, distally rounded lobes, similar in size and shape to corresponding five pairs of biramous pleopods (
<figureCitation id="132234B9FFADAB48FEA7F859E7FEF8DE" box="[349,447,2001,2027]" captionStart="FIG" captionStartId="23.[132,143,1779,1797]" captionTargetBox="[132,1455,95,1732]" captionTargetId="figure-18@23.[132,1455,215,1732]" captionTargetPageId="23" captionText="FIG. 10. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259) and allotype (ZRC 2023.0260) specimens: A, ventral view of host pinnotherid Arcotheres similis (Bürger, 1895) with Rhopalione sp. in situ on left side of pleon (posterior end of male shown by arrow);B, holotype female, dorsal view; C, holotype female,ventral view; D, allotype male, dorsal view. Scale bars: A, 2 mm; B-D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10863477" httpUri="https://zenodo.org/record/10863477/files/figure.png" pageId="23" pageNumber="170">Figs 10B</figureCitation>
;
<figureCitation id="132234B9FFADAB48FE33F859E7BAF8DE" box="[457,507,2001,2027]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="23" pageNumber="170">11C</figureCitation>
); endopodites with small basal swelling (observable in lateral view), decreasing in size on posterior pleomeres. Uniramous uropods (
<figureCitation id="132234B9FFADAB48FAF4F819E32CF89E" box="[1294,1389,1937,1963]" captionStart="FIG" captionStartId="23.[132,143,1779,1797]" captionTargetBox="[132,1455,95,1732]" captionTargetId="figure-18@23.[132,1455,215,1732]" captionTargetPageId="23" captionText="FIG. 10. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259) and allotype (ZRC 2023.0260) specimens: A, ventral view of host pinnotherid Arcotheres similis (Bürger, 1895) with Rhopalione sp. in situ on left side of pleon (posterior end of male shown by arrow);B, holotype female, dorsal view; C, holotype female,ventral view; D, allotype male, dorsal view. Scale bars: A, 2 mm; B-D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10863477" httpUri="https://zenodo.org/record/10863477/files/figure.png" pageId="23" pageNumber="170">Fig. 10B</figureCitation>
;
<figureCitation id="132234B9FFADAB48FA8DF819E3E7F89E" box="[1399,1446,1937,1963]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="23" pageNumber="170">11C</figureCitation>
) similar in size and shape to pleopods and adjacent lateral plates of pleomere 5.
</paragraph>
<caption id="DF6678B4FFA2AB47FF7CFB25E4B0FA05" ID-DOI="http://doi.org/10.5281/zenodo.10863481" ID-Zenodo-Dep="10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="24" pageNumber="171" startId="24.[134,143,1197,1215]" targetBox="[132,1455,216,1147]" targetPageId="24" targetType="figure">
<paragraph id="8BA6283CFFA2AB47FF7CFB25E4B0FA05" blockId="24.[133,1457,1196,1328]" pageId="24" pageNumber="171">
<smallCapsWord id="8D40BEE0FFA2AB47FF7CFB25E6DCFB88" baselines="1210,1210" box="[134,157,1197,1215]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="24" pageNumber="171">FIG</smallCapsWord>
. 11. —
<taxonomicName id="4C1953BFFFA2AB47FF24FB25E725FB8A" authority="Boyko &amp; Williams, 2024" authorityName="Boyko &amp; Williams" authorityYear="2024" box="[222,356,1197,1215]" class="Malacostraca" family="Bopyridae" genus="Rhopalione" kingdom="Animalia" order="Isopoda" pageId="24" pageNumber="171" phylum="Arthropoda" rank="species" species="rusa" status="sp. nov.">
<emphasis id="B96DF42EFFA2AB47FF24FB25E725FB8A" box="[222,356,1197,1215]" italics="true" pageId="24" pageNumber="171">Rhopalione rusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4955FFA2AB47FE93FB25E7D8FB8A" box="[361,409,1197,1215]" pageId="24" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
holotype (ZRC 2023.0259,
<emphasis id="B96DF42EFFA2AB47FD81FB25E4DCFB8B" bold="true" box="[635,669,1196,1215]" pageId="24" pageNumber="171">A -G</emphasis>
) and allotype (ZRC 2023.0260,
<emphasis id="B96DF42EFFA2AB47FC58FB25E588FB8A" bold="true" box="[930,969,1197,1215]" pageId="24" pageNumber="171">H -M</emphasis>
) specimens (ZRC 2019.0534):
<emphasis id="B96DF42EFFA2AB47FB32FB25E294FB8A" bold="true" box="[1224,1237,1197,1215]" pageId="24" pageNumber="171">A</emphasis>
, left antennae;
<emphasis id="B96DF42EFFA2AB47FAAEFB25E320FB8A" bold="true" box="[1364,1377,1197,1215]" pageId="24" pageNumber="171">B</emphasis>
, barbula, left side;
<emphasis id="B96DF42EFFA2AB47FF34FB4BE69DFBE0" bold="true" box="[206,220,1219,1237]" pageId="24" pageNumber="171">C</emphasis>
, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods;
<emphasis id="B96DF42EFFA2AB47FF7FFB52E6D2FBD9" bold="true" box="[133,147,1242,1260]" pageId="24" pageNumber="171">D</emphasis>
, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (
<emphasis id="B96DF42EFFA2AB47FBF1FB52E252FBD9" bold="true" box="[1035,1043,1242,1260]" pageId="24" pageNumber="171">*</emphasis>
);
<emphasis id="B96DF42EFFA2AB47FBDBFB52E26CFBD9" bold="true" box="[1057,1069,1242,1260]" pageId="24" pageNumber="171">E</emphasis>
, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view;
<emphasis id="B96DF42EFFA2AB47FE9CFB78E735FA37" bold="true" box="[358,372,1264,1282]" pageId="24" pageNumber="171">G</emphasis>
, left pereopod 7;
<emphasis id="B96DF42EFFA2AB47FDFFFB79E452FA36" bold="true" box="[517,531,1265,1283]" pageId="24" pageNumber="171">H</emphasis>
, ventral view;
<emphasis id="B96DF42EFFA2AB47FD72FB79E4CFFA36" bold="true" box="[648,654,1265,1283]" pageId="24" pageNumber="171">I</emphasis>
, left antennae;
<emphasis id="B96DF42EFFA2AB47FCF7FB79E556FA36" bold="true" box="[781,791,1265,1283]" pageId="24" pageNumber="171">J</emphasis>
, right pereopod 1;
<emphasis id="B96DF42EFFA2AB47FC4EFB79E580FA36" bold="true" box="[948,961,1265,1283]" pageId="24" pageNumber="171">K</emphasis>
, right pereopod 1, close-up of dactyl and propodus;
<emphasis id="B96DF42EFFA2AB47FA8FFB79E3C1FA36" bold="true" box="[1397,1408,1265,1283]" pageId="24" pageNumber="171">L</emphasis>
, right pereopod 7;
<emphasis id="B96DF42EFFA2AB47FF15FA8FE741FA2C" bold="true" box="[239,256,1287,1305]" pageId="24" pageNumber="171">M</emphasis>
, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations:
<emphasis id="B96DF42EFFA2AB47FC37FA8FE5A5FA2C" bold="true" box="[973,996,1287,1305]" pageId="24" pageNumber="171">En</emphasis>
, endopod;
<emphasis id="B96DF42EFFA2AB47FBBEFA8FE21BFA2C" bold="true" box="[1092,1114,1287,1305]" pageId="24" pageNumber="171">Ex</emphasis>
, exopod;
<emphasis id="B96DF42EFFA2AB47FB54FA8FE285FA2C" bold="true" box="[1198,1220,1287,1305]" pageId="24" pageNumber="171">Lp</emphasis>
, lateral plate;
<emphasis id="B96DF42EFFA2AB47FAC0FA8FE31BFA2C" bold="true" box="[1338,1370,1287,1305]" pageId="24" pageNumber="171">Uro</emphasis>
, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm.
</paragraph>
</caption>
<paragraph id="8BA6283CFFA2AB47FF7EFAFBE6F8FAB8" blockId="24.[130,777,1395,2028]" box="[132,185,1395,1421]" pageId="24" pageNumber="171">
<emphasis id="B96DF42EFFA2AB47FF7EFAFBE6F8FAB8" box="[132,185,1395,1421]" italics="true" pageId="24" pageNumber="171">Male</emphasis>
</paragraph>
<paragraph id="8BA6283CFFA2AB47FF7EFA1BE36FF979" blockId="24.[130,777,1395,2028]" lastBlockId="24.[811,1457,1395,1612]" pageId="24" pageNumber="171">
<typeStatus id="54A2969EFFA2AB47FF7EFA1BE69CFA98" box="[132,221,1427,1453]" pageId="24" pageNumber="171" type="allotype">Allotype</typeStatus>
(
<figureCitation id="132234B9FFA2AB47FF12FA1BE70AFA98" box="[232,331,1427,1453]" captionStart="FIG" captionStartId="23.[132,143,1779,1797]" captionTargetBox="[132,1455,95,1732]" captionTargetId="figure-18@23.[132,1455,215,1732]" captionTargetPageId="23" captionText="FIG. 10. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259) and allotype (ZRC 2023.0260) specimens: A, ventral view of host pinnotherid Arcotheres similis (Bürger, 1895) with Rhopalione sp. in situ on left side of pleon (posterior end of male shown by arrow);B, holotype female, dorsal view; C, holotype female,ventral view; D, allotype male, dorsal view. Scale bars: A, 2 mm; B-D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10863477" httpUri="https://zenodo.org/record/10863477/files/figure.png" pageId="24" pageNumber="171">Figs 10D</figureCitation>
;
<figureCitation id="132234B9FFA2AB47FEAFFA1BE72FFA98" box="[341,366,1427,1453]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="24" pageNumber="171">11</figureCitation>
H-M). Length
<quantity id="4CE185D9FFA2AB47FDF6FA1BE420FA9B" box="[524,609,1427,1454]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="24" pageNumber="171" unit="mm" value="4.1">4.1 mm</quantity>
; maximal width at pereomeres 2-5
<quantity id="4CE185D9FFA2AB47FEBCFA3BE7DFFAF8" box="[326,414,1459,1485]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="24" pageNumber="171" unit="mm" value="1.3">1.3 mm</quantity>
; head length
<quantity id="4CE185D9FFA2AB47FDD1FA3BE4C2FAF8" box="[555,643,1459,1485]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="24" pageNumber="171" unit="mm" value="0.4">0.4 mm</quantity>
; head width
<quantity id="4CE185D9FFA2AB47FF79FA5BE698FAD8" box="[131,217,1491,1517]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="24" pageNumber="171" unit="mm" value="0.8">0.8 mm</quantity>
. Body elongate, fusiform, straight, length 3.2 × width; all somites dorsally and ventrally distinct (
<figureCitation id="132234B9FFA2AB47FDB7FA7BE4F7F938" box="[589,694,1523,1549]" captionStart="FIG" captionStartId="23.[132,143,1779,1797]" captionTargetBox="[132,1455,95,1732]" captionTargetId="figure-18@23.[132,1455,215,1732]" captionTargetPageId="23" captionText="FIG. 10. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259) and allotype (ZRC 2023.0260) specimens: A, ventral view of host pinnotherid Arcotheres similis (Bürger, 1895) with Rhopalione sp. in situ on left side of pleon (posterior end of male shown by arrow);B, holotype female, dorsal view; C, holotype female,ventral view; D, allotype male, dorsal view. Scale bars: A, 2 mm; B-D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10863477" httpUri="https://zenodo.org/record/10863477/files/figure.png" pageId="24" pageNumber="171">Figs 10D</figureCitation>
;
<figureCitation id="132234B9FFA2AB47FD38FA7BE4B9F938" box="[706,760,1523,1549]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="24" pageNumber="171">11H</figureCitation>
). Head transversely ovate in dorsal view, about half as long as pereomere 1 (along midline) (
<figureCitation id="132234B9FFA2AB47FE4FF9BAE454F978" box="[437,533,1586,1613]" captionStart="FIG" captionStartId="23.[132,143,1779,1797]" captionTargetBox="[132,1455,95,1732]" captionTargetId="figure-18@23.[132,1455,215,1732]" captionTargetPageId="23" captionText="FIG. 10. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259) and allotype (ZRC 2023.0260) specimens: A, ventral view of host pinnotherid Arcotheres similis (Bürger, 1895) with Rhopalione sp. in situ on left side of pleon (posterior end of male shown by arrow);B, holotype female, dorsal view; C, holotype female,ventral view; D, allotype male, dorsal view. Scale bars: A, 2 mm; B-D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10863477" httpUri="https://zenodo.org/record/10863477/files/figure.png" pageId="24" pageNumber="171">Fig. 10D</figureCitation>
); anterior and posterior margins weakly convex; irregularly shaped eyes. Antennules (
<figureCitation id="132234B9FFA2AB47FF76F9FAE69EF9B9" box="[140,223,1650,1676]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="24" pageNumber="171">Fig. 11I</figureCitation>
) of three articles each, antennae (
<figureCitation id="132234B9FFA2AB47FDCCF9FAE4C8F9B9" box="[566,649,1650,1676]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="24" pageNumber="171">Fig. 11I</figureCitation>
) of five articles each, both with terminal setae. Pereomeres 1-7 (
<figureCitation id="132234B9FFA2AB47FD64F91AE543F999" box="[670,770,1682,1708]" captionStart="FIG" captionStartId="23.[132,143,1779,1797]" captionTargetBox="[132,1455,95,1732]" captionTargetId="figure-18@23.[132,1455,215,1732]" captionTargetPageId="23" captionText="FIG. 10. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259) and allotype (ZRC 2023.0260) specimens: A, ventral view of host pinnotherid Arcotheres similis (Bürger, 1895) with Rhopalione sp. in situ on left side of pleon (posterior end of male shown by arrow);B, holotype female, dorsal view; C, holotype female,ventral view; D, allotype male, dorsal view. Scale bars: A, 2 mm; B-D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10863477" httpUri="https://zenodo.org/record/10863477/files/figure.png" pageId="24" pageNumber="171">Figs 10D</figureCitation>
;
<figureCitation id="132234B9FFA2AB47FF78F93AE6F4F9F9" box="[130,181,1714,1740]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="24" pageNumber="171">11H</figureCitation>
) subequal in length, lateral margins rounded, posterior margins straight or at most slightly concave; pereomeres 2-5 subequal in width; pereomeres 1, 6, 7 narrower, subequal in width; midventral projections absent (
<figureCitation id="132234B9FFA2AB47FDE5F89AE4C4F819" box="[543,645,1810,1836]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="24" pageNumber="171">Fig. 11H</figureCitation>
). Pereopods isomorphic in size and shape, subchelate, dactyli of anterior two pairs longer than others, all dactyli with setae, propodus ovate, with setae along edge corresponding to region that meets distal end of dactylus, carpus and merus triangular, end of carpus with small setae and scales, ischium and basis tubular (
<figureCitation id="132234B9FFA2AB47FF18F859E7C5F8DE" box="[226,388,2001,2028]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="24" pageNumber="171">Fig. 11H, J, K</figureCitation>
). Pleon (
<figureCitation id="132234B9FFA2AB47FE19F859E47AF8DE" box="[483,571,2001,2027]" captionStart="FIG" captionStartId="20.[132,143,867,885]" captionTargetBox="[132,1455,214,825]" captionTargetId="figure-559@20.[132,1455,214,825]" captionTargetPageId="20" captionText="FIG. 8. — Rhopalione racemus n. sp. holotype and allotype specimens (SAM C16386): A, holotype female, dorsal view; B, holotype female, ventral view; C, allotype male, dorsal view. Scale bars: A, B, 2 mm; C, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10863471" httpUri="https://zenodo.org/record/10863471/files/figure.png" pageId="24" pageNumber="171">Figs 8C</figureCitation>
;
<figureCitation id="132234B9FFA2AB47FDBDF859E41EF8DE" box="[583,607,2001,2027]" captionStart="FIG" captionStartId="21.[132,143,1246,1264]" captionTargetBox="[132,1455,216,1187]" captionTargetId="graphics-18@21.[148,1455,216,1187]" captionTargetPageId="21" captionText="FIG. 9. — Rhopalione racemus n. sp. holotype (A-H) and allotype (I-L) specimens (SAM C16386): A, left antennae; B, barbula; C, left maxilliped, with base of barbular lobes shown; D, left oostegite 1, outer view; E, left oostegite 1, inner view; F, pereomere 7 and pleon, ventral view; G, left pereopod 1 and coxal plate; H, left pereopod 7; I, ventral view; J, right antennae, oral cone and maxilliped; K, right pereopod 1; L, left pereopod 7. Abbreviations: CP, coxal plate; IL, inner barbular lobe; Max, maxilliped; OL, outer barbular lobe; Or, oral cone. Scale bars: A, G, H, 100 µm; B, D, E, I, 500 µm; C, 250 µm; F, 1 mm; J-L, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863475" httpUri="https://zenodo.org/record/10863475/files/figure.png" pageId="24" pageNumber="171">9I</figureCitation>
) broadly tapering with triangular general outline; pleomeres dorsally and ventrally distinct, distal margins splayed, posteriorly slightly recurved, lateral margins rounded; pleomeres 1-4 with medioventral tubercles (
<figureCitation id="132234B9FFA2AB47FBF6FA5BE231FAD8" box="[1036,1136,1491,1517]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="24" pageNumber="171">Fig. 11H</figureCitation>
); pleopods 1-5 (
<figureCitation id="132234B9FFA2AB47FAE7FA5BE3E9FAD8" box="[1309,1448,1491,1517]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="24" pageNumber="171">Fig. 11H, M</figureCitation>
) reniform, uniramous, with minute setae in concave region (
<figureCitation id="132234B9FFA2AB47FCCCF99AE5E3F918" box="[822,930,1554,1581]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="24" pageNumber="171">Fig. 11M</figureCitation>
). Pleotelson (
<figureCitation id="132234B9FFA2AB47FBCFF99AE2DFF918" box="[1077,1182,1554,1581]" captionStart="FIG" captionStartId="23.[132,143,1779,1797]" captionTargetBox="[132,1455,95,1732]" captionTargetId="figure-18@23.[132,1455,215,1732]" captionTargetPageId="23" captionText="FIG. 10. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259) and allotype (ZRC 2023.0260) specimens: A, ventral view of host pinnotherid Arcotheres similis (Bürger, 1895) with Rhopalione sp. in situ on left side of pleon (posterior end of male shown by arrow);B, holotype female, dorsal view; C, holotype female,ventral view; D, allotype male, dorsal view. Scale bars: A, 2 mm; B-D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10863477" httpUri="https://zenodo.org/record/10863477/files/figure.png" pageId="24" pageNumber="171">Figs 10D</figureCitation>
;
<figureCitation id="132234B9FFA2AB47FB51F99BE29FF918" box="[1195,1246,1555,1581]" captionStart="FIG" captionStartId="24.[134,143,1197,1215]" captionTargetBox="[132,1455,216,1147]" captionTargetId="graphics-16@24.[132,1455,216,1147]" captionTargetPageId="24" captionText="FIG. 11. — Rhopalione rusa n. sp. holotype (ZRC 2023.0259, A-G) and allotype (ZRC 2023.0260,H-M) specimens (ZRC 2019.0534): A, left antennae; B, barbula, left side;C, dorsal view of terminus of pleon showing fifth pleomere with lateral plates on both sides and biramous pleopod shown only on right side and uropods; D, left maxilliped,outer view;inset shows inner view of anterior lobe and spur obscured by lobe of maxilliped (*); E, left oostegite 1 and pereopod 1, outer view; F, left oostegite 1, inner view; G, left pereopod 7; H, ventral view;I, left antennae; J, right pereopod 1; K, right pereopod 1, close-up of dactyl and propodus; L, right pereopod 7; M, pleomere showing pleopod 4 on right side and mid-ventral tubercule. Abbreviations: En, endopod; Ex, exopod; Lp, lateral plate; Uro, uropods. Scale bars: A, G, L, M, 100 µm; B, D-F, 250 µm; C, H, 500 µm; J-K, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10863481" httpUri="https://zenodo.org/record/10863481/files/figure.png" pageId="24" pageNumber="171">11H</figureCitation>
) narrow anteriorly, widening posteriorly, indented posteromedially.
</paragraph>
</subSubSection>
<subSubSection id="C3037BB7FFA2AB46FCD7F9FBE4CBFD65" lastPageId="25" lastPageNumber="172" pageId="24" pageNumber="171" type="discussion">
<paragraph id="8BA6283CFFA2AB47FCD7F9FBE5D0F9BE" blockId="24.[811,1456,1651,2028]" box="[813,913,1651,1677]" pageId="24" pageNumber="171">
<smallCapsWord id="8D40BEE0FFA2AB47FCD7F9FBE5D0F9BE" baselines="1670,1670" box="[813,913,1651,1677]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="24" pageNumber="171">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="8BA6283CFFA2AB46FCD7F91AE451FEA4" blockId="24.[811,1456,1651,2028]" lastBlockId="25.[132,777,215,592]" lastPageId="25" lastPageNumber="172" pageId="24" pageNumber="171">
Females of
<taxonomicName id="4C1953BFFFA2AB47FC5FF91AE20BF999" authority="Boyko &amp; Williams, 2024" authorityName="Boyko &amp; Williams" authorityYear="2024" box="[933,1098,1682,1708]" class="Malacostraca" family="Bopyridae" genus="Rhopalione" kingdom="Animalia" order="Isopoda" pageId="24" pageNumber="171" phylum="Arthropoda" rank="species" species="rusa" status="sp. nov.">
<emphasis id="B96DF42EFFA2AB47FC5FF91AE20BF999" box="[933,1098,1682,1708]" italics="true" pageId="24" pageNumber="171">Rhopalione rusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4955FFA2AB47FBABF91BE2CDF998" box="[1105,1164,1683,1709]" pageId="24" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
are most similar to
<taxonomicName id="4C1953BFFFA2AB47FA9AF91BE516F9F9" authority="Boyko &amp; Williams, 2024" authorityName="Boyko &amp; Williams" authorityYear="2024" class="Malacostraca" family="Bopyridae" genus="Rhopalione" kingdom="Animalia" order="Isopoda" pageId="24" pageNumber="171" phylum="Arthropoda" rank="species" species="racemus" status="n. sp.">
<emphasis id="B96DF42EFFA2AB47FA9AF91BE331F999" box="[1376,1392,1683,1708]" italics="true" pageId="24" pageNumber="171">R</emphasis>
.
<emphasis id="B96DF42EFFA2AB47FA84F91BE516F9F9" italics="true" pageId="24" pageNumber="171">racemus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4955FFA2AB47FCA4F93BE5D8F9F8" box="[862,921,1715,1741]" pageId="24" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
in the shape of the pereomeres, which are greatly inflated laterally, forming a kind of barbell shape; however, the females differ in the form of the pleopods and lateral plates which are long and slender in
<taxonomicName id="4C1953BFFFA2AB47FB42F89AE343F81E" authority="Boyko &amp; Williams, 2024" authorityName="Boyko &amp; Williams" authorityYear="2024" box="[1208,1282,1810,1836]" class="Malacostraca" family="Bopyridae" genus="Rhopalione" kingdom="Animalia" order="Isopoda" pageId="24" pageNumber="171" phylum="Arthropoda" rank="species" species="rusa" status="n. sp.">
<emphasis id="B96DF42EFFA2AB47FB42F89AE289F81E" box="[1208,1224,1810,1835]" italics="true" pageId="24" pageNumber="171">R</emphasis>
.
<emphasis id="B96DF42EFFA2AB47FB2DF89AE343F81E" box="[1239,1282,1810,1835]" italics="true" pageId="24" pageNumber="171">rusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4955FFA2AB47FAF0F89AE304F819" box="[1290,1349,1810,1836]" pageId="24" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
but short and broad in
<taxonomicName id="4C1953BFFFA2AB47FC39F8BAE275F87E" authority="Boyko &amp; Williams, 2024" authorityName="Boyko &amp; Williams" authorityYear="2024" box="[963,1076,1842,1868]" class="Malacostraca" family="Bopyridae" genus="Rhopalione" kingdom="Animalia" order="Isopoda" pageId="24" pageNumber="171" phylum="Arthropoda" rank="species" species="racemus" status="n. sp.">
<emphasis id="B96DF42EFFA2AB47FC39F8BAE592F87E" box="[963,979,1842,1867]" italics="true" pageId="24" pageNumber="171">R</emphasis>
.
<emphasis id="B96DF42EFFA2AB47FC18F8BAE275F87E" box="[994,1076,1842,1867]" italics="true" pageId="24" pageNumber="171">racemus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4955FFA2AB47FBC7F8BAE238F879" box="[1085,1145,1842,1868]" pageId="24" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
Additionally, the barbula of females of
<taxonomicName id="4C1953BFFFA2AB47FC5CF8DAE5B0F85E" authority="Boyko &amp; Williams, 2024" authorityName="Boyko &amp; Williams" authorityYear="2024" box="[934,1009,1874,1900]" class="Malacostraca" family="Bopyridae" genus="Rhopalione" kingdom="Animalia" order="Isopoda" pageId="24" pageNumber="171" phylum="Arthropoda" rank="species" species="rusa" status="n. sp.">
<emphasis id="B96DF42EFFA2AB47FC5CF8DAE5F4F85E" box="[934,949,1874,1899]" italics="true" pageId="24" pageNumber="171">R</emphasis>
.
<emphasis id="B96DF42EFFA2AB47FC3FF8DAE5B0F85E" box="[965,1009,1874,1899]" italics="true" pageId="24" pageNumber="171">rusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4955FFA2AB47FC03F8DAE276F859" box="[1017,1079,1874,1900]" pageId="24" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
bears two subequal lobes whereas that of females of
<taxonomicName id="4C1953BFFFA2AB47FC17F8FAE21BF8BE" authority="Boyko &amp; Williams, 2024" authorityName="Boyko &amp; Williams" authorityYear="2024" box="[1005,1114,1906,1932]" class="Malacostraca" family="Bopyridae" genus="Rhopalione" kingdom="Animalia" order="Isopoda" pageId="24" pageNumber="171" phylum="Arthropoda" rank="species" species="racemus" status="n. sp.">
<emphasis id="B96DF42EFFA2AB47FC17F8FAE5BCF8BE" box="[1005,1021,1906,1931]" italics="true" pageId="24" pageNumber="171">R</emphasis>
.
<emphasis id="B96DF42EFFA2AB47FBF2F8FAE21BF8BE" box="[1032,1114,1906,1931]" italics="true" pageId="24" pageNumber="171">racemus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4955FFA2AB47FB9AF8FAE2DBF8B9" box="[1120,1178,1906,1932]" pageId="24" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
have the outer lobe much longer than the inner. Males of the two species are also similar with both having five pairs of reniform pleopods but
<taxonomicName id="4C1953BFFFA2AB47FA9EF83AE3EEF8FE" authority="Boyko &amp; Williams, 2024" authorityName="Boyko &amp; Williams" authorityYear="2024" box="[1380,1455,1970,1996]" class="Malacostraca" family="Bopyridae" genus="Rhopalione" kingdom="Animalia" order="Isopoda" pageId="24" pageNumber="171" phylum="Arthropoda" rank="species" species="rusa" status="n. sp.">
<emphasis id="B96DF42EFFA2AB47FA9EF83AE335F8FE" box="[1380,1396,1970,1995]" italics="true" pageId="24" pageNumber="171">R</emphasis>
.
<emphasis id="B96DF42EFFA2AB47FA7EF83AE3EEF8FE" box="[1412,1455,1970,1995]" italics="true" pageId="24" pageNumber="171">rusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4955FFA2AB47FCD7F85AE52AF8D9" box="[813,875,2002,2028]" pageId="24" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
has midventral tubercles on pleomeres 1-4 whereas
<taxonomicName id="4C1953BFFFA3AB46FF7EFF50E6B1FFC4" authority="Boyko &amp; Williams, 2024" authorityName="Boyko &amp; Williams" authorityYear="2024" box="[132,240,216,242]" class="Malacostraca" family="Bopyridae" genus="Rhopalione" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="172" phylum="Arthropoda" rank="species" species="racemus" status="n. sp.">
<emphasis id="B96DF42EFFA3AB46FF7EFF50E6D5FFC4" box="[132,148,216,241]" italics="true" pageId="25" pageNumber="172">R</emphasis>
.
<emphasis id="B96DF42EFFA3AB46FF5AFF50E6B1FFC4" box="[160,240,216,241]" italics="true" pageId="25" pageNumber="172">racemus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4955FFA3AB46FF0FFF50E76CFFC7" box="[245,301,216,242]" pageId="25" pageNumber="172" rank="species">n. sp.</taxonomicNameLabel>
lack midventral tubercles.
<taxonomicName id="4C1953BFFFA3AB46FDBAFF5FE546FFC4" authority="Boyko &amp; Williams, 2024" authorityName="Boyko &amp; Williams" authorityYear="2024" box="[576,775,215,241]" class="Malacostraca" family="Bopyridae" genus="Rhopalione" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="172" phylum="Arthropoda" rank="species" species="racemus" status="sp. nov.">
<emphasis id="B96DF42EFFA3AB46FDBAFF5FE546FFC4" box="[576,775,215,241]" italics="true" pageId="25" pageNumber="172">Rhopalione racemus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4955FFA3AB46FF7EFF70E6FFFE27" box="[132,190,248,274]" pageId="25" pageNumber="172" rank="species">n. sp.</taxonomicNameLabel>
is known only from
<taxonomicName id="4C1953BFFFA3AB46FE60FF7FE4E3FE24" baseAuthorityName="Baker" baseAuthorityYear="1907" box="[410,674,247,273]" class="Malacostraca" family="Pinnotheridae" genus="Discorsotheres" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="172" phylum="Arthropoda" rank="species" species="subglobosus">
<emphasis id="B96DF42EFFA3AB46FE60FF7FE4E3FE24" box="[410,674,247,273]" italics="true" pageId="25" pageNumber="172">Discorsotheres subglobosus</emphasis>
</taxonomicName>
collected in South Australia (Temperate Australasia) vs
<taxonomicName id="4C1953BFFFA3AB46FD99FE9FE546FE04" authority="Boyko &amp; Williams, 2024" authorityName="Boyko &amp; Williams" authorityYear="2024" box="[611,775,279,305]" class="Malacostraca" family="Bopyridae" genus="Rhopalione" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="172" phylum="Arthropoda" rank="species" species="rusa" status="sp. nov.">
<emphasis id="B96DF42EFFA3AB46FD99FE9FE546FE04" box="[611,775,279,305]" italics="true" pageId="25" pageNumber="172">Rhopalione rusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E4955FFA3AB46FF7EFEBFE680FE64" box="[132,193,311,337]" pageId="25" pageNumber="172" rank="species">n. sp.</taxonomicNameLabel>
, which is only known from
<taxonomicName id="4C1953BFFFA3AB46FE0BFEBFE4E0FE64" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[497,673,311,337]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="172" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B96DF42EFFA3AB46FE0BFEBFE4E0FE64" box="[497,673,311,337]" italics="true" pageId="25" pageNumber="172">Arcotheres similis</emphasis>
</taxonomicName>
collected in
<collectingCountry id="F30E68ACFFA3AB46FF5AFEDFE74AFE44" box="[160,267,343,369]" name="Singapore" pageId="25" pageNumber="172">Singapore</collectingCountry>
(Central Indo-Pacific) (see
<bibRefCitation id="EF8855CDFFA3AB46FDC8FEDFE546FE44" author="SPALDING M. D. &amp; FOX H. E. &amp; ALLEN G. R. &amp; DAVIDSON N. &amp; FERDANA Z. A. &amp; FINLAYSON M. &amp; HALPERN B. S." box="[562,775,343,369]" pageId="25" pageNumber="172" pagination="573 - 583" refId="ref23998" refString="SPALDING M. D., FOX H. E., ALLEN G. R., DAVIDSON N., FERDANA Z. A., FINLAYSON M., HALPERN B. S., ET AL. 2007. - Marine ecoregions of the world: a bioregionalization of coastal and shelf areas. BioScience 57 (7): 573 - 583. https: // doi. org / 10.1641 / B 570707" type="journal article" year="2007">
Spalding
<emphasis id="B96DF42EFFA3AB46FD6CFED0E488FE44" box="[662,713,343,369]" italics="true" pageId="25" pageNumber="172">et al.</emphasis>
2007
</bibRefCitation>
for discussion of marine ecoregions).
</paragraph>
<paragraph id="8BA6283CFFA3AB46FF61FE1EE4CBFD65" blockId="25.[132,777,215,592]" pageId="25" pageNumber="172">
The material of
<taxonomicName id="4C1953BFFFA3AB46FEBFFE1EE7F9FE85" authorityName="Perez" authorityYear="1920" box="[325,440,406,432]" class="Malacostraca" family="Bopyridae" genus="Rhopalione" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="B96DF42EFFA3AB46FEBFFE1EE7F9FE85" box="[325,440,406,432]" italics="true" pageId="25" pageNumber="172">Rhopalione</emphasis>
</taxonomicName>
cited by
<bibRefCitation id="EF8855CDFFA3AB46FDE1FE1FE546FE84" author="NG P. K. L. &amp; AHYONG S. T." box="[539,775,406,433]" pageId="25" pageNumber="172" pagination="134 - 248" refId="ref22405" refString="NG P. K. L. &amp; AHYONG S. T. 2022. - The pea crab genus Arcotheres Manning, 1993 (Crustacea: Brachyura: Pinnotheridae) from Singapore and Peninsular Malaysia, with a reappraisal of diagnostic characters and descriptions of two new genera. Raffles Bulletin of Zoology 70: 134 - 248. https: // doi. org / 10.26107 / RBZ- 2022 - 0009" type="journal article" year="2022">Ng &amp; Ahyong (2022)</bibRefCitation>
comprises the type series as designated herein. The
<typeStatus id="54A2969EFFA3AB46FD52FE3EE546FEE5" box="[680,775,438,464]" pageId="25" pageNumber="172" type="holotype">holotype</typeStatus>
female left approximately four puncture holes from the mouthparts in ventral pleomeres 1 and 2 (toward midline) of the host; the
<typeStatus id="54A2969EFFA3AB46FED3FD9FE7C4FD04" box="[297,389,535,561]" pageId="25" pageNumber="172" type="paratype">paratype</typeStatus>
female left puncture holes in ventral pleomeres 1 and 2 (toward midline) of the host.
</paragraph>
</subSubSection>
</treatment>
</document>