497 lines
69 KiB
XML
497 lines
69 KiB
XML
<document id="D5D4167D09D93A1DAABAB799F4CEE51A" ID-CLB-Dataset="21443" ID-DOI="10.11646/zootaxa.4816.3.4" ID-GBIF-Dataset="8de569a5-ae44-4994-8313-57163e9a4514" ID-ISSN="1175-5326" ID-Zenodo-Dep="3954447" ID-ZooBank="FA112B5F-E75E-4E6F-8E4C-E0D87674FCCA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1595339868132" checkinUser="plazi" docAuthor="Ahyong, Shane T. & Ng, Peter K. L." docDate="2020" docId="03B087FADC14FFC3C798FD69FB8E2BB6" docLanguage="en" docName="zootaxa.4816.3.4.pdf" docOrigin="Zootaxa 4816 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nepinnotheres fulvia Ahyong & Ng 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="346" masterDocId="FF89FF82DC1EFFCEC70FFFABFFCA2C29" masterDocTitle="New species of pinnotherid crabs from Southeast Asia and Papua New Guinea (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="349" masterPageNumber="333" pageNumber="343" updateTime="1698843768168" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="6ACCA46ED97B0FAE068CA82457413A52" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CD0AC681844AC4A83300C874D0281A14">
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<mods:title id="F5FFFC250344C88BA034CB8FDF5BF2F9">New species of pinnotherid crabs from Southeast Asia and Papua New Guinea (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:name id="B8CCF833D79055C1D662076EA3A7FD76" type="personal">
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<mods:roleTerm id="DDFA03BE031B1B844DDFD68DF10AB349">Author</mods:roleTerm>
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<mods:namePart id="4B8EF4D5364B4866681D34F8C5F9AC15">Ahyong, Shane T.</mods:namePart>
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<mods:affiliation id="7432376FEDFE15B40A2826FCCA405EDF">Australian Museum Research Institute, Australian Museum, 1 William Street, Sydney NSW 2010, Australia, and School of Biological, Earth & Environmental Sciences, University of New South Wales, NSW 2052, Australia.</mods:affiliation>
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</mods:name>
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<mods:name id="18C235B9FD6ADC09C5334B4B18560EBE" type="personal">
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<mods:roleTerm id="3D4825D654AC334688430CA32EB60F1F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="91D4F46345CB1959BE1D8E6EA788AC6F">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="8ED2EB872E6F0B6238A4417127043DFE">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore. peterng @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 5946 - 0608</mods:affiliation>
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<mods:typeOfResource id="6AE1226AC34E9D9DD69B767B48D4BE51">text</mods:typeOfResource>
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<mods:title id="F086678CE2F23A520799C6D3805B0F4D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="C11A541BE5E42E34A72F8412C353C688">
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<mods:date id="A797D641967B78A2233C4A6FCAE0F337">2020</mods:date>
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<mods:number id="371E38732B3A73DB79570ED843E9A19B">2020-07-17</mods:number>
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<mods:detail id="BB6C679CA494A8343BFEEBE56700CB0E" type="volume">
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<mods:number id="3334899248FECF69C2C336777DDA6833">4816</mods:number>
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<mods:number id="2BC5C7E3ED39E0D2A718222CAE07C09C">3</mods:number>
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<mods:identifier id="CC9ACB2E6279E3F4269B057DA38CB118" type="DOI">10.11646/zootaxa.4816.3.4</mods:identifier>
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<treatment id="03B087FADC14FFC3C798FD69FB8E2BB6" ID-DOI="http://doi.org/10.5281/zenodo.4330024" ID-GBIF-Taxon="165857118" ID-Zenodo-Dep="4330024" LSID="urn:lsid:plazi:treatment:03B087FADC14FFC3C798FD69FB8E2BB6" httpUri="http://treatment.plazi.org/id/03B087FADC14FFC3C798FD69FB8E2BB6" lastPageId="13" lastPageNumber="346" pageId="10" pageNumber="343">
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<subSubSection id="C3036567DC14FFC4C798FD69FE312EF4" box="[151,507,706,733]" pageId="10" pageNumber="343" type="nomenclature">
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<paragraph id="8BA636ECDC14FFC4C798FD69FE312EF4" blockId="10.[151,507,706,769]" box="[151,507,706,733]" pageId="10" pageNumber="343">
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<heading id="D0EE8180DC14FFC4C798FD69FE312EF4" bold="true" box="[151,507,706,733]" fontSize="11" level="1" pageId="10" pageNumber="343" reason="1">
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<emphasis id="B96DEAFEDC14FFC4C798FD69FE312EF4" bold="true" box="[151,507,706,733]" pageId="10" pageNumber="343">
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<taxonomicName id="4C194D6FDC14FFC4C798FD69FE5C2EF4" ID-CoL="84H22" authority="Ahyong & Ng, 2020" authorityName="Ahyong & Ng" authorityYear="2020" box="[151,406,706,733]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="343" phylum="Arthropoda" rank="species" species="fulvia" status="sp. nov.">
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<emphasis id="B96DEAFEDC14FFC4C798FD69FE5C2EF4" bold="true" box="[151,406,706,733]" italics="true" pageId="10" pageNumber="343">Nepinnotheres fulvia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25E5785DC14FFC4C691FD68FE312EF4" box="[414,507,707,733]" pageId="10" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3036567DC14FFC4C798FD4CFED82F28" box="[151,274,743,769]" pageId="10" pageNumber="343" type="description">
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<paragraph id="8BA636ECDC14FFC4C798FD4CFED82F28" blockId="10.[151,507,706,769]" box="[151,274,743,769]" pageId="10" pageNumber="343">
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(
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<figureCitation id="13222A69DC14FFC4C790FD4CFF252F28" box="[159,239,743,769]" captionStart="FIGURE 6" captionStartId="12.[151,250,1811,1836]" captionTargetBox="[162,1429,245,1773]" captionTargetId="figure@12.[151,1436,237,1786]" captionTargetPageId="12" captionText="FIGURE 6. Nepinnotheres fulvia sp. nov., male holotype, cl 2.8 mm, cw 2.9 mm, ZRC 2018.1391. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum and anterior pleon; E, right chela; F–I, right P2–5; J, right P2 proximal articles, ventral view; K, pleon; L, left G1, pleonal view; M, left G2, pleonal view. Scales: A = 1.0 mm; B, L, M = 0.25 mm; C–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954459" httpUri="https://zenodo.org/record/3954459/files/figure.png" pageId="10" pageNumber="343">Figs. 6</figureCitation>
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,
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<figureCitation id="13222A69DC14FFC4C7F4FD4CFEC02F28" box="[251,266,743,769]" captionStart="FIGURE 7" captionStartId="13.[151,250,1530,1555]" captionTargetBox="[155,1435,190,1494]" captionTargetId="figure@13.[151,1436,181,1505]" captionTargetPageId="13" captionText="FIGURE 7. Nepinnotheres fulvia sp. nov., female paratype, cl 3.0 mm, cw 3.1 mm, ZRC 2018.1392. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, right P2–5; J, right P2 proximal articles, ventral view; K, pleon. Scales: A, K = 1.0 mm; B = 0.25 mm; C–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954461" httpUri="https://zenodo.org/record/3954461/files/figure.png" pageId="10" pageNumber="343">7</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3036567DC14FFC4C798FC84FDC22F9C" pageId="10" pageNumber="343" type="materials_examined">
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<paragraph id="8BA636ECDC14FFC4C798FC84FDC22F9C" blockId="10.[151,1437,814,2029]" pageId="10" pageNumber="343">
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<emphasis id="B96DEAFEDC14FFC4C798FC84FEF52F60" bold="true" box="[151,319,814,841]" pageId="10" pageNumber="343">Type material</emphasis>
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||
.
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<materialsCitation id="3B713CB1DC14FFC4C640FC84FD0E2FB8" ID-GBIF-Occurrence="2816749305" collectingDate="2012-12-07" collectionCode="PAPUA, NIUGINI" collectorName="Fulvia & G. B. Sowerby & A. Anker" country="Papua New Guinea" elevation="10" latitude="-5.137667" location="Tabad Island" longLatPrecision="1" longitude="145.812" pageId="10" pageNumber="343" specimenCode="ZRC 2018.1391" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="54A2884EDC14FFC4C640FC84FE2D2F60" box="[335,487,815,841]" pageId="10" pageNumber="343" type="holotype">HOLOTYPE</typeStatus>
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||
:
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<specimenCode id="DBBF9E97DC14FFC4C6FCFC84FD662F60" box="[499,684,815,841]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="343">ZRC 2018.1391</specimenCode>
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, male (cl
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||
<quantity id="4CE19B09DC14FFC4C411FC84FCB22F60" box="[798,888,815,841]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="10" pageNumber="343" unit="mm" value="2.8">2.8 mm</quantity>
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, cw
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<quantity id="4CE19B09DC14FFC4C4A1FC84FBC02F60" box="[942,1034,815,841]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="343" unit="mm" value="2.9">2.9 mm</quantity>
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), west of
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<location id="8EC66037DC14FFC4C375FC84FAC72F60" LSID="urn:lsid:plazi:treatment:03B087FADC14FFC3C798FD69FB8E2BB6:8EC66037DC14FFC4C375FC84FAC72F60" box="[1146,1293,815,841]" country="Papua New Guinea" latitude="-5.137667" longLatPrecision="1" longitude="145.812" name="Tabad Island" pageId="10" pageNumber="343">Tabad Island</location>
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,
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||
<collectingCountry id="F30E767CDC14FFC4C215FC84FF212F44" name="Papua New Guinea" pageId="10" pageNumber="343">Papua New Guinea</collectingCountry>
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,
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<geoCoordinate id="EE2D502BDC14FFC4C7F9FCF9FE592F44" box="[246,403,850,877]" degrees="5" direction="south" minutes="08.26002" orientation="latitude" pageId="10" pageNumber="343" precision="1" value="-5.137667">5°08.26002’S</geoCoordinate>
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,
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<geoCoordinate id="EE2D502BDC14FFC4C691FCF9FD902F44" box="[414,602,850,877]" degrees="145" direction="east" minutes="48.71976" orientation="longitude" pageId="10" pageNumber="343" precision="1" value="145.812">145°48.71976’E</geoCoordinate>
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,
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<quantity id="4CE19B09DC14FFC4C56AFCF8FD552F44" box="[613,671,851,877]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="343" unit="m" value="10.0">
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<elevation id="0034D1DFDC14FFC4C56AFCF8FD552F44" box="[613,671,851,877]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="343" unit="m" value="10.0">10 m</elevation>
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</quantity>
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, from
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||
<emphasis id="B96DEAFEDC14FFC4C5E7FCF8FC532F44" box="[744,921,851,877]" italics="true" pageId="10" pageNumber="343">
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||
<collectorName id="26EC533ADC14FFC4C5E7FCF8FCFB2F44" box="[744,817,851,877]" pageId="10" pageNumber="343">Fulvia</collectorName>
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australis
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</emphasis>
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||
(
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<collectorName id="26EC533ADC14FFC4C4A6FCF8FB832F44" box="[937,1097,851,877]" pageId="10" pageNumber="343">G.B. Sowerby</collectorName>
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II, 1834) (
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<taxonomicName id="4C194D6FDC14FFC4C3B0FCF8FAF82F47" authorityName="Lamarck" authorityYear="1809" box="[1215,1330,851,878]" class="Bivalvia" family="Cardiidae" kingdom="Animalia" order="Veneroida" pageId="10" pageNumber="343" phylum="Mollusca" rank="family">Cardiidae</taxonomicName>
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),
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<collectionCode id="ED08AE29DC14FFC4C24AFCF8FECE2FB8" pageId="10" pageNumber="343">PAPUA NIUGINI</collectionCode>
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stn PR198, coll.
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<collectorName id="26EC533ADC14FFC4C6C8FCDCFDFB2FB8" box="[455,561,887,913]" pageId="10" pageNumber="343">A. Anker</collectorName>
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,
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||
<date id="FFA7102CDC14FFC4C532FCDCFD0E2FB8" box="[573,708,887,913]" pageId="10" pageNumber="343" value="2012-12-07">
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<collectingDate id="EFE3E9C4DC14FFC4C532FCDCFD0E2FB8" box="[573,708,887,913]" pageId="10" pageNumber="343" value="2012-12-07">7 Dec 2012</collectingDate>
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</date>
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</materialsCitation>
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.
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<materialsCitation id="3B713CB1DC14FFC4C5C0FCDCFDCF2F9C" ID-GBIF-Occurrence="2816749303" collectingDate="2012-12-07" collectionCode="ZRC" collectorName="Fulvia & G. B. Sowerby & A. Anker" country="Papua New Guinea" elevation="10" latitude="-5.137667" location="Tabad Island" longLatPrecision="1" longitude="145.812" pageId="10" pageNumber="343" specimenCode="ZRC 2018.1392" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54A2884EDC14FFC4C5C0FCDCFC952FB8" box="[719,863,887,913]" pageId="10" pageNumber="343" type="paratype">PARATYPE</typeStatus>
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:
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<specimenCode id="DBBF9E97DC14FFC4C466FCDCFBE82FB8" box="[873,1058,887,913]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="343">ZRC 2018.1392</specimenCode>
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, female (cl 3.0 mm, cw
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<quantity id="4CE19B09DC14FFC4C23AFCDCFA5A2FB8" box="[1333,1424,887,913]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="10" pageNumber="343" unit="mm" value="3.1">3.1 mm</quantity>
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), collected together with holotype
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</materialsCitation>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3036567DC14FFC5C7C8FC15FE6A2F76" lastPageId="11" lastPageNumber="344" pageId="10" pageNumber="343" type="description">
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<paragraph id="8BA636ECDC14FFC4C7C8FC15FB7E286C" blockId="10.[151,1437,814,2029]" pageId="10" pageNumber="343">
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<emphasis id="B96DEAFEDC14FFC4C7C8FC15FE9B2FF0" bold="true" box="[199,337,958,985]" pageId="10" pageNumber="343">Description</emphasis>
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. MALE: Carapace subcircular. Front slightly produced, anterior margin weakly concave in dorsal view. Dorsum smooth, glabrous. Epistome with broad, triangular interantennular septum; median buccal margin with narrow, acute, median point. Antennular sinus of similar size to orbit; antennules folded obliquely. Antennal articles 1 and 2 fused to epistome. Eyes visible in dorsal view, filling orbit, cornea pigmented.
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</paragraph>
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<paragraph id="8BA636ECDC14FFC4C7C8FBE4FAB128D0" blockId="10.[151,1437,814,2029]" pageId="10" pageNumber="343">Maxilliped 3 ischiomerus length about twice width; inner distal margin with prominent, obtuse angle, margin distal and proximal to angle straight; outer margin strongly convex. Carpus shorter than propodus. Propodus spatulate, tapering in distal half, apex rounded, dorsally and distally setose, length 2.8× dactylus length. Dactylus digitiform, distally setose, inserted at propodal midlength, apex reaching to midway between insertion and end of propodus. Exopod inner margin straight, outer margin broadly convex; flagellum with 2 articles, distally setose.</paragraph>
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<paragraph id="8BA636ECDC14FFC4C7C8FAA8FEB02A30" blockId="10.[151,1437,814,2029]" pageId="10" pageNumber="343">Chelipeds stout, inflated, symmetrical from left to right. Dactylus and pollex crossing distally, with distinct gape. Dactylus arcuate, dorsal margin arcuate, sparsely setose; occlusal margin setose, with blunt triangular tooth proximal to midlength, concave in distal half, with rounded dentate subdistal lobe near base of simple, falcate dactylar apex. Pollex relatively straight, apex simple, with fringe of short setae on inner ventral margin extending onto proximal half of palm; occlusal margin setose, irregularly and minutely dentate, with short spinules on distal half, with low subdistal lobe occluding with corresponding dactylar lobe when fingers closed. Propodus palm dorsal margin 1.3× height, 1.2× length of dactylus; scattered dorsal setae; ventral margin weakly concave. Carpus unarmed, inner margin setose.</paragraph>
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<paragraph id="8BA636ECDC14FFC4C7C8F988FBB92A80" blockId="10.[151,1437,814,2029]" pageId="10" pageNumber="343">
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Pereopods 2–5 (walking legs 1–4) symmetrical from left to right, unarmed; extensor and flexor margins setose, P3 and 4 also with natatory setae; relative lengths in decreasing order: P3
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<emphasis id="B96DEAFEDC14FFC4C4C3F9E3FC152A48" box="[972,991,1608,1633]" italics="true" pageId="10" pageNumber="343">></emphasis>
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P4
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<emphasis id="B96DEAFEDC14FFC4C305F9E3FBD72A48" box="[1034,1053,1608,1633]" italics="true" pageId="10" pageNumber="343">></emphasis>
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P2
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<emphasis id="B96DEAFEDC14FFC4C348F9E3FB902A48" box="[1095,1114,1608,1633]" italics="true" pageId="10" pageNumber="343">></emphasis>
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P5. P2 basis anterior surface granulate. P3 merus 0.6× pcl. P2–5 dactyli similar, evenly arcuate, distally spiniform, P2–4 dactylus 0.6× propodus length, P5 dactylus 0.7× propodus length. Relative dactylus lengths: P2 = P3 = P4
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<emphasis id="B96DEAFEDC14FFC4C33AF93BFB822A80" box="[1077,1096,1680,1705]" italics="true" pageId="10" pageNumber="343">></emphasis>
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P5.
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</paragraph>
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<paragraph id="8BA636ECDC14FFC4C7C8F918FAA12AD8" blockId="10.[151,1437,814,2029]" pageId="10" pageNumber="343">Thoracic sternum anterior margin distinctly concave medially; sternites 1–3 fused; demarcation between sternites 2 and 3 partially indicated by transverse median groove; sterno-pleonal cavity reaching sternite ¾ suture.</paragraph>
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<paragraph id="8BA636ECDC14FFC4C7C8F950FD912B10" blockId="10.[151,1437,814,2029]" pageId="10" pageNumber="343">Pleon slender, margins evenly converging distally; comprising 6 free somites and telson, widest at somites 2 and 3; telson wider than long, apex rounded.</paragraph>
|
||
<paragraph id="8BA636ECDC14FFC4C7C8F8E8FC982B8C" blockId="10.[151,1437,814,2029]" pageId="10" pageNumber="343">Gonopod 1 elongate, slender, arcuate, dorsoventrally compressed, about 2/3 length of pleon, tapering to simple apex; margins setose, longest on outer margin. Gonopod 2 endopod simple, about ¼ length of gonopod 1; exopod short, slender, about as long as endopod. Pleopods 3–5 absent.</paragraph>
|
||
<paragraph id="8BA636ECDC14FFC5C7C8F804FAE02CFE" blockId="10.[151,1437,814,2029]" lastBlockId="11.[151,1437,153,2015]" lastPageId="11" lastPageNumber="344" pageId="10" pageNumber="343">FEMALE: Carapace subcircular. Front not produced, anterior margin weakly concave in dorsal view. Dorsum smooth, very few, minute, scattered setae. Epistome with broad, triangular interantennular septum; median buccal margin with narrow, acute, median point. Antennular sinus of similar size to orbit; antennules folded obliquely. Antennal articles 1 and 2 fused to epistome. Eyes not visible in dorsal view, filling orbit, cornea pigmented.</paragraph>
|
||
<paragraph id="8BA636ECDC15FFC5C7C8FF4AFAE92DA2" blockId="11.[151,1437,153,2015]" pageId="11" pageNumber="344">Maxilliped 3 ischiomerus length about twice width; inner distal margin with prominent, obtuse angle, margin distal and proximal to angle straight; outer margin strongly convex. Carpus shorter than propodus. Propodus spatulate, tapering in distal half, apex rounded, dorsally and distally setose, length 3.2× dactylus length. Dactylus digitiform, distally setose, inserted at propodal midlength, apex reaching to midway between insertion and end of propodus. Exopod inner margin straight, outer margin sinuous; flagellum with 2 articles, distally setose.</paragraph>
|
||
<paragraph id="8BA636ECDC15FFC5C7C8FE3EFBD22E16" blockId="11.[151,1437,153,2015]" pageId="11" pageNumber="344">Chelipeds symmetrical from left to right, surfaces sparsely setose. Dactylus and pollex relatively straight, crossing distally, without gape. Dactylus occlusal margin with blunt triangular tooth proximal to midlength, straight in distal half, sparsely setose, minutely denticulate. Pollex occlusal margin irregular, sparsely setose, minutely denticulate; with fringe of short setae on inner ventral margin. Propodus palm dorsal margin 1.5× height, 1.2× length of dactylus; ventral margin weakly concave. Carpus unarmed, inner margin setose.</paragraph>
|
||
<paragraph id="8BA636ECDC15FFC5C7C8FDE2FF302EDA" blockId="11.[151,1437,153,2015]" pageId="11" pageNumber="344">
|
||
Pereopods 2–5 (walking legs 1–4) symmetrical from left to right, unarmed, setose on flexor and extensor margins of merus, carpus, propodus, and dactylus, P3–4 also with natatory setae; relative lengths in decreasing order: P3
|
||
<emphasis id="B96DEAFEDC15FFC5C7B3FD39FF052E82" box="[188,207,658,683]" italics="true" pageId="11" pageNumber="344">></emphasis>
|
||
P4
|
||
<emphasis id="B96DEAFEDC15FFC5C7F5FD39FEC72E82" box="[250,269,658,683]" italics="true" pageId="11" pageNumber="344">></emphasis>
|
||
P2
|
||
<emphasis id="B96DEAFEDC15FFC5C637FD39FE812E82" box="[312,331,658,683]" italics="true" pageId="11" pageNumber="344">></emphasis>
|
||
P5. P3 merus 0.5× pcl. P2 basis anterior surface granulate. Dactyli similar, evenly arcuate, distally spiniform; P2–4 dactyli 0.5× propodus length, of P5, 0.6× propodus length. Relative dactylus lengths: P3> P2 = P4
|
||
<emphasis id="B96DEAFEDC15FFC5C7B3FD71FF052EDA" box="[188,207,730,755]" italics="true" pageId="11" pageNumber="344">></emphasis>
|
||
P5.
|
||
</paragraph>
|
||
<paragraph id="8BA636ECDC15FFC5C7C8FD56FAFA2F3E" blockId="11.[151,1437,153,2015]" box="[199,1328,765,791]" pageId="11" pageNumber="344">Thoracic sternum anterior margin shallowly concave medially; sternites 1–3 indistinguishably fused.</paragraph>
|
||
<paragraph id="8BA636ECDC15FFC5C7C8FC8AFE6A2F76" blockId="11.[151,1437,153,2015]" pageId="11" pageNumber="344">
|
||
Pleon of
|
||
<typeStatus id="54A2884EDC15FFC5C621FC8AFE442F12" box="[302,398,801,827]" pageId="11" pageNumber="344" type="paratype">paratype</typeStatus>
|
||
yet to reach full size, reaching to, or almost to, but not covering coxa of walking legs; of 6 free somites and telson.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3036567DC15FFC5C7C8FCC2FCAC2FAD" box="[199,870,873,900]" pageId="11" pageNumber="344" type="biology_ecology">
|
||
<paragraph id="8BA636ECDC15FFC5C7C8FCC2FCAC2FAD" blockId="11.[151,1437,153,2015]" box="[199,870,873,900]" pageId="11" pageNumber="344">
|
||
<emphasis id="B96DEAFEDC15FFC5C7C8FCC2FF352FAA" bold="true" box="[199,255,873,899]" pageId="11" pageNumber="344">Host</emphasis>
|
||
.
|
||
<emphasis id="B96DEAFEDC15FFC5C602FCC2FE752FAA" box="[269,447,873,899]" italics="true" pageId="11" pageNumber="344">Fulvia australis</emphasis>
|
||
(G.B. Sowerby II, 1834) (
|
||
<taxonomicName id="4C194D6FDC15FFC5C5E8FCC2FC912FAD" authorityName="Lamarck" authorityYear="1809" box="[743,859,873,900]" class="Bivalvia" family="Cardiidae" kingdom="Animalia" order="Veneroida" pageId="11" pageNumber="344" phylum="Mollusca" rank="family">Cardiidae</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3036567DC15FFC5C7C8FC26FB7F2F8E" box="[199,1205,909,935]" pageId="11" pageNumber="344" type="etymology">
|
||
<paragraph id="8BA636ECDC15FFC5C7C8FC26FB7F2F8E" blockId="11.[151,1437,153,2015]" box="[199,1205,909,935]" pageId="11" pageNumber="344">
|
||
<emphasis id="B96DEAFEDC15FFC5C7C8FC26FE822F8E" bold="true" box="[199,328,909,935]" pageId="11" pageNumber="344">Etymology</emphasis>
|
||
. Named
|
||
<taxonomicName id="4C194D6FDC15FFC5C6A1FC26FE242F8E" authorityName="Ahyong & Ng" authorityYear="2020" box="[430,494,909,935]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="fulvia">
|
||
<emphasis id="B96DEAFEDC15FFC5C6A1FC26FE242F8E" box="[430,494,909,935]" italics="true" pageId="11" pageNumber="344">fulvia</emphasis>
|
||
</taxonomicName>
|
||
, after the genus name of the host; used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3036567DC15FFC3C7C8FC1AFEB22B55" lastPageId="13" lastPageNumber="346" pageId="11" pageNumber="344" type="discussion">
|
||
<paragraph id="8BA636ECDC15FFC5C7C8FC1AFE062B66" blockId="11.[151,1437,153,2015]" pageId="11" pageNumber="344">
|
||
<emphasis id="B96DEAFEDC15FFC5C7C8FC1AFEFE2FE2" bold="true" box="[199,308,945,971]" pageId="11" pageNumber="344">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C194D6FDC15FFC5C64CFC1AFDE42FE2" authority="Ahyong & Ng, 2020" authorityName="Ahyong & Ng" authorityYear="2020" box="[323,558,945,971]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="fulvia" status="sp. nov.">
|
||
<emphasis id="B96DEAFEDC15FFC5C64CFC1AFDE42FE2" box="[323,558,945,971]" italics="true" pageId="11" pageNumber="344">Nepinnotheres fulvia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96DEAFEDC15FFC5C539FC1AFD5A2FE2" bold="true" box="[566,656,945,971]" pageId="11" pageNumber="344">
|
||
<taxonomicNameLabel id="A25E5785DC15FFC5C539FC1AFD5A2FE2" box="[566,656,945,971]" pageId="11" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is readily distinguished from the majority of its congeners in having the maxilliped 3 dactylus distinctly under-reaching (
|
||
<figureCitation id="13222A69DC15FFC5C5D2FC7EFCFE2FC6" box="[733,820,981,1007]" captionStart="FIGURE 6" captionStartId="12.[151,250,1811,1836]" captionTargetBox="[162,1429,245,1773]" captionTargetId="figure@12.[151,1436,237,1786]" captionTargetPageId="12" captionText="FIGURE 6. Nepinnotheres fulvia sp. nov., male holotype, cl 2.8 mm, cw 2.9 mm, ZRC 2018.1391. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum and anterior pleon; E, right chela; F–I, right P2–5; J, right P2 proximal articles, ventral view; K, pleon; L, left G1, pleonal view; M, left G2, pleonal view. Scales: A = 1.0 mm; B, L, M = 0.25 mm; C–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954459" httpUri="https://zenodo.org/record/3954459/files/figure.png" pageId="11" pageNumber="344">Fig. 6B</figureCitation>
|
||
,
|
||
<figureCitation id="13222A69DC15FFC5C430FC7EFCAA2FC6" box="[831,864,981,1007]" captionStart="FIGURE 7" captionStartId="13.[151,250,1530,1555]" captionTargetBox="[155,1435,190,1494]" captionTargetId="figure@13.[151,1436,181,1505]" captionTargetPageId="13" captionText="FIGURE 7. Nepinnotheres fulvia sp. nov., female paratype, cl 3.0 mm, cw 3.1 mm, ZRC 2018.1392. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, right P2–5; J, right P2 proximal articles, ventral view; K, pleon. Scales: A, K = 1.0 mm; B = 0.25 mm; C–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954461" httpUri="https://zenodo.org/record/3954461/files/figure.png" pageId="11" pageNumber="344">7B</figureCitation>
|
||
) rather than reaching to, or over-reaching, the apex of the propodus. Six species, however, also have a maxilliped 3 dactylus that distinctly under-reaches the propodus:
|
||
<taxonomicName id="4C194D6FDC15FFC5C798FBB5FE6C281E" authority="(Burger, 1895)" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[151,422,1052,1079]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="cardii">
|
||
<emphasis id="B96DEAFEDC15FFC5C798FBB5FF30281E" box="[151,250,1053,1079]" italics="true" pageId="11" pageNumber="344">N. cardii</emphasis>
|
||
(
|
||
<bibRefCitation id="EF884B1DDC15FFC5C606FBB6FE54281E" author="Burger, O." box="[265,414,1052,1079]" pageId="11" pageNumber="344" pagination="361 - 390" refId="ref9358" refString="Burger, O. (1895) Ein Beitrag zur kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article" year="1895">Bürger, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(type locality:
|
||
<collectingCountry id="F30E767CDC15FFC5C55DFBB6FD1F2811" box="[594,725,1053,1080]" name="Philippines" pageId="11" pageNumber="344">Philippines</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C194D6FDC15FFC5C5E7FBB5FBE8281E" authority="(Guerin, 1832)" baseAuthorityName="Guerin" baseAuthorityYear="1832" box="[744,1058,1053,1079]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="villosulus">
|
||
<emphasis id="B96DEAFEDC15FFC5C5E7FBB5FCBE281E" box="[744,884,1053,1079]" italics="true" pageId="11" pageNumber="344">N. villosulus</emphasis>
|
||
(Guérin, 1832)
|
||
</taxonomicName>
|
||
(type locality: Timor,
|
||
<collectingCountry id="F30E767CDC15FFC5C210FBB6FA5A2811" box="[1311,1424,1053,1080]" name="Indonesia" pageId="11" pageNumber="344">Indonesia</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C194D6FDC15FFC5C798FBE9FE232872" authority="(Burger, 1895)" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[151,489,1088,1115]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="glaberrimus">
|
||
<emphasis id="B96DEAFEDC15FFC5C798FBE9FE8A2872" box="[151,320,1089,1115]" italics="true" pageId="11" pageNumber="344">N. glaberrimus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF884B1DDC15FFC5C642FBEAFE2A2872" author="Burger, O." box="[333,480,1088,1115]" pageId="11" pageNumber="344" pagination="361 - 390" refId="ref9358" refString="Burger, O. (1895) Ein Beitrag zur kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article" year="1895">Bürger, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(type locality:
|
||
<collectingCountry id="F30E767CDC15FFC5C580FBEAFCD82875" box="[655,786,1089,1116]" name="Philippines" pageId="11" pageNumber="344">Philippines</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C194D6FDC15FFC5C42CFBE9FA972872" authority="(Schmitt, McCain & Davidson, 1973)" baseAuthorityName="Schmitt, McCain & Davidson" baseAuthorityYear="1973" box="[803,1373,1089,1115]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="rathbunae">
|
||
<emphasis id="B96DEAFEDC15FFC5C42CFBE9FC7F2872" box="[803,949,1089,1115]" italics="true" pageId="11" pageNumber="344">N. rathbunae</emphasis>
|
||
(
|
||
<bibRefCitation id="EF884B1DDC15FFC5C4CCFBEAFA9F2872" author="Schmitt, W. L. & McCain, J. C. & Davidson, E." box="[963,1365,1089,1115]" pageId="11" pageNumber="344" pagination="1 - 160" refId="ref11806" refString="Schmitt, W. L., McCain, J. C. & Davidson, E. (1973) Decapoda I. Brachyura I. Fam. Pinnotheridae. Crustaceorum Catalogus, 3, 1 - 160." type="journal article" year="1973">Schmitt, McCain & Davidson, 1973</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(type locality:
|
||
<collectingCountry id="F30E767CDC15FFC5C7F3FBCEFEF628A9" box="[252,316,1125,1152]" name="Palau" pageId="11" pageNumber="344">Palau</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C194D6FDC15FFC5C65BFBCDFD2D2856" authority="(H. Milne Edwards, 1853)" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[340,743,1124,1151]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="rouxi">
|
||
<emphasis id="B96DEAFEDC15FFC5C65BFBCDFE7B2856" box="[340,433,1126,1151]" italics="true" pageId="11" pageNumber="344">N. rouxi</emphasis>
|
||
(H.
|
||
<bibRefCitation id="EF884B1DDC15FFC5C6E7FBCEFD152856" author="Milne Edwards, H." box="[488,735,1124,1151]" pageId="11" pageNumber="344" pagination="163 - 228" refId="ref10329" refString="Milne Edwards, H. (1853) Memoire sur la famille des Ocypodides. Suite (1). Deuxieme Tribu Principale. Annales des Sciences Naturelles, 3 e Serie, 20, 163 - 228, pls. 6 - 11." type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(type locality: Indian Ocean), and
|
||
<taxonomicName id="4C194D6FDC15FFC5C374FBCDFF12288A" authority="(Pillai, 1951)" baseAuthorityName="Pillai" baseAuthorityYear="1951" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="sanguinolariae">
|
||
<emphasis id="B96DEAFEDC15FFC5C374FBCDFA822856" box="[1147,1352,1125,1151]" italics="true" pageId="11" pageNumber="344">N. sanguinolariae</emphasis>
|
||
(
|
||
<bibRefCitation id="EF884B1DDC15FFC5C256FBCEFF1A288A" author="Pillai, N. K." pageId="11" pageNumber="344" pagination="1 - 46" refId="ref11340" refString="Pillai, N. K. (1951) Decapoda (Brachyura) from Travancore. Bulletin of the Central Research Institute, University of Travancore, Trivandrum, Series C (Natural Science), 2, 1 - 46." type="journal article" year="1951">Pillai, 1951</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(type locality:
|
||
<collectingCountry id="F30E767CDC15FFC5C687FB22FE0C288D" box="[392,454,1161,1188]" name="India" pageId="11" pageNumber="344">India</collectingCountry>
|
||
). Female
|
||
<taxonomicName id="4C194D6FDC15FFC5C53AFB21FD5D288A" authorityName="Ahyong & Ng" authorityYear="2020" box="[565,663,1161,1187]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="fulvia">
|
||
<emphasis id="B96DEAFEDC15FFC5C53AFB21FD5D288A" box="[565,663,1161,1187]" italics="true" pageId="11" pageNumber="344">N. fulvia</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4C194D6FDC15FFC5C4C0FB21FBF9288A" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[975,1075,1161,1187]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="cardii">
|
||
<emphasis id="B96DEAFEDC15FFC5C4C0FB21FBF9288A" box="[975,1075,1161,1187]" italics="true" pageId="11" pageNumber="344">N. cardii</emphasis>
|
||
</taxonomicName>
|
||
by the absence of the proximal occlusal tooth on the cheliped pollex (
|
||
<figureCitation id="13222A69DC15FFC5C54EFB06FD5C28EE" box="[577,662,1197,1223]" captionStart="FIGURE 7" captionStartId="13.[151,250,1530,1555]" captionTargetBox="[155,1435,190,1494]" captionTargetId="figure@13.[151,1436,181,1505]" captionTargetPageId="13" captionText="FIGURE 7. Nepinnotheres fulvia sp. nov., female paratype, cl 3.0 mm, cw 3.1 mm, ZRC 2018.1392. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, right P2–5; J, right P2 proximal articles, ventral view; K, pleon. Scales: A, K = 1.0 mm; B = 0.25 mm; C–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954461" httpUri="https://zenodo.org/record/3954461/files/figure.png" pageId="11" pageNumber="344">Fig. 7E</figureCitation>
|
||
) (present in
|
||
<taxonomicName id="4C194D6FDC15FFC5C42DFB05FC4E28EE" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[802,900,1197,1223]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="cardii">
|
||
<emphasis id="B96DEAFEDC15FFC5C42DFB05FC4E28EE" box="[802,900,1197,1223]" italics="true" pageId="11" pageNumber="344">N. cardii</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EF884B1DDC15FFC5C49EFB06FB4B28EE" author="Ahyong, S. T. & Ng, P. K. L." box="[913,1153,1197,1223]" pageId="11" pageNumber="344" pagination="191 - 226" refId="ref9096" refString="Ahyong, S. T. & Ng, P. K. L. (2007 a) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong & Ng 2007a</bibRefCitation>
|
||
: fig. 22B; Ng
|
||
<emphasis id="B96DEAFEDC15FFC5C211FB05FA9C28EE" box="[1310,1366,1197,1223]" italics="true" pageId="11" pageNumber="344">et al.</emphasis>
|
||
2019: fig. 1, 2D), and strongly setose walking legs (
|
||
<figureCitation id="13222A69DC15FFC5C5A4FB7AFCCD28C2" box="[683,775,1233,1259]" captionStart="FIGURE 7" captionStartId="13.[151,250,1530,1555]" captionTargetBox="[155,1435,190,1494]" captionTargetId="figure@13.[151,1436,181,1505]" captionTargetPageId="13" captionText="FIGURE 7. Nepinnotheres fulvia sp. nov., female paratype, cl 3.0 mm, cw 3.1 mm, ZRC 2018.1392. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, right P2–5; J, right P2 proximal articles, ventral view; K, pleon. Scales: A, K = 1.0 mm; B = 0.25 mm; C–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954461" httpUri="https://zenodo.org/record/3954461/files/figure.png" pageId="11" pageNumber="344">Fig. 7A</figureCitation>
|
||
, E–H) (glabrous in
|
||
<taxonomicName id="4C194D6FDC15FFC5C4E1FB79FB8428C2" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1006,1102,1233,1259]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="cardii">
|
||
<emphasis id="B96DEAFEDC15FFC5C4E1FB79FB8428C2" box="[1006,1102,1233,1259]" italics="true" pageId="11" pageNumber="344">P. cardii</emphasis>
|
||
</taxonomicName>
|
||
except for flexor margins of propodi and dactyli;
|
||
<bibRefCitation id="EF884B1DDC15FFC5C68DFB5EFDBF2926" author="Ahyong, S. T. & Ng, P. K. L." box="[386,629,1269,1295]" pageId="11" pageNumber="344" pagination="191 - 226" refId="ref9096" refString="Ahyong, S. T. & Ng, P. K. L. (2007 a) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong & Ng 2007a</bibRefCitation>
|
||
: fig. 22A; Ng
|
||
<emphasis id="B96DEAFEDC15FFC5C416FB5DFC992926" box="[793,851,1269,1295]" italics="true" pageId="11" pageNumber="344">et al.</emphasis>
|
||
2019: fig. 1). Male
|
||
<taxonomicName id="4C194D6FDC15FFC5C336FB5DFB512926" authorityName="Ahyong & Ng" authorityYear="2020" box="[1081,1179,1269,1295]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="fulvia">
|
||
<emphasis id="B96DEAFEDC15FFC5C336FB5DFB512926" box="[1081,1179,1269,1295]" italics="true" pageId="11" pageNumber="344">N. fulvia</emphasis>
|
||
</taxonomicName>
|
||
can be separated from male
|
||
<taxonomicName id="4C194D6FDC15FFC5C7DAFAB1FEF2291A" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[213,312,1305,1331]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="cardii">
|
||
<emphasis id="B96DEAFEDC15FFC5C7DAFAB1FEF2291A" box="[213,312,1305,1331]" italics="true" pageId="11" pageNumber="344">N. cardii</emphasis>
|
||
</taxonomicName>
|
||
by the more slender maxilliped 3 propodus (length 3× height versus 2× height in
|
||
<taxonomicName id="4C194D6FDC15FFC5C3C0FAB1FAF8291A" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1231,1330,1305,1331]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="cardii">
|
||
<emphasis id="B96DEAFEDC15FFC5C3C0FAB1FAF8291A" box="[1231,1330,1305,1331]" italics="true" pageId="11" pageNumber="344">N. cardii</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="13222A69DC15FFC5C24EFAB2FA52291A" box="[1345,1432,1305,1331]" captionStart="FIGURE 6" captionStartId="12.[151,250,1811,1836]" captionTargetBox="[162,1429,245,1773]" captionTargetId="figure@12.[151,1436,237,1786]" captionTargetPageId="12" captionText="FIGURE 6. Nepinnotheres fulvia sp. nov., male holotype, cl 2.8 mm, cw 2.9 mm, ZRC 2018.1391. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum and anterior pleon; E, right chela; F–I, right P2–5; J, right P2 proximal articles, ventral view; K, pleon; L, left G1, pleonal view; M, left G2, pleonal view. Scales: A = 1.0 mm; B, L, M = 0.25 mm; C–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954459" httpUri="https://zenodo.org/record/3954459/files/figure.png" pageId="11" pageNumber="344">Fig. 6B</figureCitation>
|
||
,
|
||
<bibRefCitation id="EF884B1DDC15FFC5C798FA96FE46297E" author="Ahyong, S. T. & Ng, P. K. L." box="[151,396,1341,1367]" pageId="11" pageNumber="344" pagination="191 - 226" refId="ref9096" refString="Ahyong, S. T. & Ng, P. K. L. (2007 a) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong & Ng 2007a</bibRefCitation>
|
||
: fig. 22F) and absence of a prominent rounded tooth on the distal one-third of the cheliped pollex occlusal margin (
|
||
<figureCitation id="13222A69DC15FFC5C6A2FACAFDCE2955" box="[429,516,1377,1404]" captionStart="FIGURE 6" captionStartId="12.[151,250,1811,1836]" captionTargetBox="[162,1429,245,1773]" captionTargetId="figure@12.[151,1436,237,1786]" captionTargetPageId="12" captionText="FIGURE 6. Nepinnotheres fulvia sp. nov., male holotype, cl 2.8 mm, cw 2.9 mm, ZRC 2018.1391. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum and anterior pleon; E, right chela; F–I, right P2–5; J, right P2 proximal articles, ventral view; K, pleon; L, left G1, pleonal view; M, left G2, pleonal view. Scales: A = 1.0 mm; B, L, M = 0.25 mm; C–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954459" httpUri="https://zenodo.org/record/3954459/files/figure.png" pageId="11" pageNumber="344">Fig. 6E</figureCitation>
|
||
) (present in
|
||
<taxonomicName id="4C194D6FDC15FFC5C596FAC9FD342952" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[665,766,1377,1403]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="cardii">
|
||
<emphasis id="B96DEAFEDC15FFC5C596FAC9FD342952" box="[665,766,1377,1403]" italics="true" pageId="11" pageNumber="344">N. cardii</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EF884B1DDC15FFC5C401FACAFBCF2952" author="Ahyong, S. T. & Ng, P. K. L." box="[782,1029,1377,1403]" pageId="11" pageNumber="344" pagination="191 - 226" refId="ref9096" refString="Ahyong, S. T. & Ng, P. K. L. (2007 a) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong & Ng 2007a</bibRefCitation>
|
||
: fig. 22E). Female
|
||
<taxonomicName id="4C194D6FDC15FFC5C3EAFAC9FA822952" authorityName="Ahyong & Ng" authorityYear="2020" box="[1253,1352,1377,1403]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="fulvia">
|
||
<emphasis id="B96DEAFEDC15FFC5C3EAFAC9FA822952" box="[1253,1352,1377,1403]" italics="true" pageId="11" pageNumber="344">N. fulvia</emphasis>
|
||
</taxonomicName>
|
||
can be readily separated from female
|
||
<taxonomicName id="4C194D6FDC15FFC5C6E2FA2DFDB329B6" baseAuthorityName="Guerin" baseAuthorityYear="1832" box="[493,633,1413,1439]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="villosulus">
|
||
<emphasis id="B96DEAFEDC15FFC5C6E2FA2DFDB329B6" box="[493,633,1413,1439]" italics="true" pageId="11" pageNumber="344">N. villosulus</emphasis>
|
||
</taxonomicName>
|
||
(males presently unknown) by lacking the short tomentum that densely covers the entire external surface of the cephalothorax and pereopods, and from both sexes of
|
||
<taxonomicName id="4C194D6FDC15FFC5C3BFFA01FAB029EA" baseAuthorityName="Pillai" baseAuthorityYear="1951" box="[1200,1402,1449,1475]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="sanguinolariae">
|
||
<emphasis id="B96DEAFEDC15FFC5C3BFFA01FAB029EA" box="[1200,1402,1449,1475]" italics="true" pageId="11" pageNumber="344">N. sanguinolariae</emphasis>
|
||
</taxonomicName>
|
||
by the subcircular versus transversely ovate carapace.
|
||
<taxonomicName id="4C194D6FDC15FFC5C5DDFA66FBC929CE" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[722,1027,1485,1511]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="glaberrimus">
|
||
<emphasis id="B96DEAFEDC15FFC5C5DDFA66FBC929CE" box="[722,1027,1485,1511]" italics="true" pageId="11" pageNumber="344">Nepinnotheres glaberrimus</emphasis>
|
||
</taxonomicName>
|
||
, known with certainty only from the male
|
||
<typeStatus id="54A2884EDC15FFC5C7DBFA5AFEF42A22" box="[212,318,1521,1547]" pageId="11" pageNumber="344" type="lectotype">lectotype</typeStatus>
|
||
, differs from male
|
||
<taxonomicName id="4C194D6FDC15FFC5C51DFA59FDB82A22" authorityName="Ahyong & Ng" authorityYear="2020" box="[530,626,1521,1547]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="fulvia">
|
||
<emphasis id="B96DEAFEDC15FFC5C51DFA59FDB82A22" box="[530,626,1521,1547]" italics="true" pageId="11" pageNumber="344">N. fulvia</emphasis>
|
||
</taxonomicName>
|
||
by the position of the maxilliped 3 dactylus articulation (proximal to the midlength versus at the midlength in
|
||
<taxonomicName id="4C194D6FDC15FFC5C54FF9BDFD682A06" authorityName="Ahyong & Ng" authorityYear="2020" box="[576,674,1557,1583]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="fulvia">
|
||
<emphasis id="B96DEAFEDC15FFC5C54FF9BDFD682A06" box="[576,674,1557,1583]" italics="true" pageId="11" pageNumber="344">N. fulvia</emphasis>
|
||
</taxonomicName>
|
||
) and the presence of two prominent proximal teeth on the occlusal margin of the cheliped pollex (absent in
|
||
<taxonomicName id="4C194D6FDC15FFC5C569F991FD022A7A" authorityName="Ahyong & Ng" authorityYear="2020" box="[614,712,1593,1619]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="fulvia">
|
||
<emphasis id="B96DEAFEDC15FFC5C569F991FD022A7A" box="[614,712,1593,1619]" italics="true" pageId="11" pageNumber="344">N. fulvia</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C194D6FDC15FFC5C5EFF992FC0F2A7A" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[736,965,1593,1619]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="rouxi">
|
||
<emphasis id="B96DEAFEDC15FFC5C5EFF992FC0F2A7A" box="[736,965,1593,1619]" italics="true" pageId="11" pageNumber="344">Nepinnotheres rouxi</emphasis>
|
||
</taxonomicName>
|
||
is known only from the type description, but, based on the maxilliped 3 of the
|
||
<typeStatus id="54A2884EDC15FFC5C54DF9F6FD6E2A5E" box="[578,676,1629,1655]" pageId="11" pageNumber="344" type="holotype">holotype</typeStatus>
|
||
figured by H.
|
||
<bibRefCitation id="EF884B1DDC15FFC5C441F9F6FB832A5E" author="Milne Edwards, H." box="[846,1097,1628,1655]" pageId="11" pageNumber="344" pagination="163 - 228" refId="ref10329" refString="Milne Edwards, H. (1853) Memoire sur la famille des Ocypodides. Suite (1). Deuxieme Tribu Principale. Annales des Sciences Naturelles, 3 e Serie, 20, 163 - 228, pls. 6 - 11." type="journal article" year="1853">Milne Edwards (1853</bibRefCitation>
|
||
: pl. 11 fig. 7), differs from
|
||
<taxonomicName id="4C194D6FDC15FFC5C28CF9F5FF1D2AB2" authorityName="Ahyong & Ng" authorityYear="2020" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="fulvia">
|
||
<emphasis id="B96DEAFEDC15FFC5C28CF9F5FF1D2AB2" italics="true" pageId="11" pageNumber="344">N. fulvia</emphasis>
|
||
</taxonomicName>
|
||
by the comparatively narrow ischiomerus (length about 2.5× width) with a broadly rounded, weakly produced inner distal angle of the ischiomerus (versus length about twice width with distinctly angular inner distal margin in the new species).
|
||
<taxonomicName id="4C194D6FDC15FFC5C670F962FD562ACA" baseAuthorityName="Schmitt, McCain & Davidson" baseAuthorityYear="1973" box="[383,668,1737,1763]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="rathbunae">
|
||
<emphasis id="B96DEAFEDC15FFC5C670F962FD562ACA" box="[383,668,1737,1763]" italics="true" pageId="11" pageNumber="344">Nepinnotheres rathbunae</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C194D6FDC15FFC5C43CF961FC5F2ACA" authorityName="Ahyong & Ng" authorityYear="2020" box="[819,917,1737,1763]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="fulvia">
|
||
<emphasis id="B96DEAFEDC15FFC5C43CF961FC5F2ACA" box="[819,917,1737,1763]" italics="true" pageId="11" pageNumber="344">N. fulvia</emphasis>
|
||
</taxonomicName>
|
||
in a similar way to
|
||
<taxonomicName id="4C194D6FDC15FFC5C377F961FB1E2ACA" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[1144,1236,1738,1763]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="rouxi">
|
||
<emphasis id="B96DEAFEDC15FFC5C377F961FB1E2ACA" box="[1144,1236,1738,1763]" italics="true" pageId="11" pageNumber="344">N. rouxi</emphasis>
|
||
</taxonomicName>
|
||
. Another unusual feature of both sexes of
|
||
<taxonomicName id="4C194D6FDC15FFC5C6AEF945FDCA2B2E" authorityName="Ahyong & Ng" authorityYear="2020" box="[417,512,1773,1799]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="fulvia">
|
||
<emphasis id="B96DEAFEDC15FFC5C6AEF945FDCA2B2E" box="[417,512,1773,1799]" italics="true" pageId="11" pageNumber="344">N. fulvia</emphasis>
|
||
</taxonomicName>
|
||
is the presence of a granular anterior surface of the P2 coxa (
|
||
<figureCitation id="13222A69DC15FFC5C3ABF946FB372B2E" box="[1188,1277,1773,1799]" captionStart="FIGURE 6" captionStartId="12.[151,250,1811,1836]" captionTargetBox="[162,1429,245,1773]" captionTargetId="figure@12.[151,1436,237,1786]" captionTargetPageId="12" captionText="FIGURE 6. Nepinnotheres fulvia sp. nov., male holotype, cl 2.8 mm, cw 2.9 mm, ZRC 2018.1391. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum and anterior pleon; E, right chela; F–I, right P2–5; J, right P2 proximal articles, ventral view; K, pleon; L, left G1, pleonal view; M, left G2, pleonal view. Scales: A = 1.0 mm; B, L, M = 0.25 mm; C–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954459" httpUri="https://zenodo.org/record/3954459/files/figure.png" pageId="11" pageNumber="344">Figs. 6J</figureCitation>
|
||
,
|
||
<figureCitation id="13222A69DC15FFC5C207F946FAE92B2E" box="[1288,1315,1773,1799]" captionStart="FIGURE 7" captionStartId="13.[151,250,1530,1555]" captionTargetBox="[155,1435,190,1494]" captionTargetId="figure@13.[151,1436,181,1505]" captionTargetPageId="13" captionText="FIGURE 7. Nepinnotheres fulvia sp. nov., female paratype, cl 3.0 mm, cw 3.1 mm, ZRC 2018.1392. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, right P2–5; J, right P2 proximal articles, ventral view; K, pleon. Scales: A, K = 1.0 mm; B = 0.25 mm; C–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954461" httpUri="https://zenodo.org/record/3954461/files/figure.png" pageId="11" pageNumber="344">7J</figureCitation>
|
||
) (although more prominent in the male than female); this feature has not been previously reported in pinnotherids, although it may have been overlooked.
|
||
</paragraph>
|
||
<paragraph id="8BA636ECDC15FFC2C7C8F8F2FE092CF1" blockId="11.[151,1437,153,2015]" lastBlockId="12.[151,1436,153,216]" lastPageId="12" lastPageNumber="345" pageId="11" pageNumber="344">
|
||
The male cheliped in
|
||
<taxonomicName id="4C194D6FDC15FFC5C6B6F8F1FDD02B5A" authority="Ahyong & Ng, 2020" authorityName="Ahyong & Ng" authorityYear="2020" box="[441,538,1881,1907]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="fulvia" status="sp. nov.">
|
||
<emphasis id="B96DEAFEDC15FFC5C6B6F8F1FDD02B5A" box="[441,538,1881,1907]" italics="true" pageId="11" pageNumber="344">N. fulvia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96DEAFEDC15FFC5C52FF8F2FDB02B5A" bold="true" box="[544,634,1881,1907]" pageId="11" pageNumber="344">
|
||
<taxonomicNameLabel id="A25E5785DC15FFC5C52FF8F2FDB02B5A" box="[544,634,1881,1907]" pageId="11" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is conspicuous among congeners for the large gape between the fingers and the presence of a denticulate subdistal lobe on the outer occlusal surface dactylus that occludes with a low swelling on the corresponding surface of the pollex, ‘interlocking’ with the curved apices of the fingers. Of known males of
|
||
<taxonomicName id="4C194D6FDC15FFC5C60EF86EFE692BF6" authorityName="Manning" authorityYear="1993" box="[257,419,1989,2015]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96DEAFEDC15FFC5C60EF86EFE692BF6" box="[257,419,1989,2015]" italics="true" pageId="11" pageNumber="344">Nepinnotheres</emphasis>
|
||
</taxonomicName>
|
||
, such an arrangement is otherwise known only in
|
||
<taxonomicName id="4C194D6FDC15FFC5C4ECF86DFB3C2BF6" authority="(Burger, 1895)" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[995,1270,1988,2015]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="344" phylum="Arthropoda" rank="species" species="cardii">
|
||
<emphasis id="B96DEAFEDC15FFC5C4ECF86DFB8D2BF6" box="[995,1095,1989,2015]" italics="true" pageId="11" pageNumber="344">N. cardii</emphasis>
|
||
(
|
||
<bibRefCitation id="EF884B1DDC15FFC5C358F86EFB242BF6" author="Burger, O." box="[1111,1262,1988,2015]" pageId="11" pageNumber="344" pagination="361 - 390" refId="ref9358" refString="Burger, O. (1895) Ein Beitrag zur kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article" year="1895">Bürger, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54A2884EDC15FFC5C209F86EFAF22BF6" box="[1286,1336,1989,2015]" pageId="11" pageNumber="344">type</typeStatus>
|
||
locality:
|
||
<collectingCountry id="F30E767CDC12FFC2C798FF32FED02C9D" box="[151,282,153,180]" name="Philippines" pageId="12" pageNumber="345">Philippines</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C194D6FDC12FFC2C621FF31FDAE2C9A" authority="(Sakai, 1939)" baseAuthorityName="Sakai" baseAuthorityYear="1939" box="[302,612,153,179]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="345" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B96DEAFEDC12FFC2C621FF31FE0E2C9A" box="[302,452,153,179]" italics="true" pageId="12" pageNumber="345">N. bidentatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF884B1DDC12FFC2C6DBFF32FD912C9A" author="Sakai, T." box="[468,603,153,179]" pageId="12" pageNumber="345" pagination="1 - 129" refId="ref11632" refString="Sakai, T. (1939) Studies on the crabs of Japan. IV. Brachygnatha, Brachyrhyncha. s. n., Tokyo, 741 + 23 pp., figs. 1 - 129, pls. 42 - 111." type="book chapter" year="1939">Sakai, 1939</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54A2884EDC12FFC2C57CFF32FD6F2C9A" box="[627,677,153,179]" pageId="12" pageNumber="345">type</typeStatus>
|
||
locality:
|
||
<collectingRegion id="49DDF80EDC12FFC2C41EFF32FC5A2C9A" box="[785,912,153,179]" country="Japan" name="Wakayama" pageId="12" pageNumber="345">Wakayama</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30E767CDC12FFC2C494FF32FC172C9A" box="[923,989,153,179]" name="Japan" pageId="12" pageNumber="345">Japan</collectingCountry>
|
||
) and
|
||
<taxonomicName id="4C194D6FDC12FFC2C312FF31FA912C9A" authority="(Sakai, 2000)" baseAuthorityName="Sakai" baseAuthorityYear="2000" box="[1053,1371,153,179]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="345" phylum="Arthropoda" rank="species" species="taichungae">
|
||
<emphasis id="B96DEAFEDC12FFC2C312FF31FB702C9A" box="[1053,1210,153,179]" italics="true" pageId="12" pageNumber="345">N. taichungae</emphasis>
|
||
(
|
||
<bibRefCitation id="EF884B1DDC12FFC2C3C4FF32FA992C9A" author="Sakai, K." box="[1227,1363,153,179]" pageId="12" pageNumber="345" pagination="1155 - 1162" refId="ref11573" refString="Sakai, K. (2000) On the occurrence of three species of crabs on Shikoku Island, Japan, and a new species, Pinnotheres taichungae nov. spec., from Taiwan (Decapoda, Brachyura). Crustaceana, 73, 1155 - 1162. https: // doi. org / 10.1163 / 156854000505146" type="journal article" year="2000">Sakai, 2000</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54A2884EDC12FFC2C265FF32FA562C9A" box="[1386,1436,153,179]" pageId="12" pageNumber="345">type</typeStatus>
|
||
locality: Kaomei, Taiwan).
|
||
</paragraph>
|
||
<caption id="DF666664DC12FFC2C798F8B8FE542BBE" ID-DOI="http://doi.org/10.5281/zenodo.3954459" ID-Zenodo-Dep="3954459" httpUri="https://zenodo.org/record/3954459/files/figure.png" pageId="12" pageNumber="345" startId="12.[151,250,1811,1836]" targetBox="[162,1429,245,1773]" targetPageId="12">
|
||
<paragraph id="8BA636ECDC12FFC2C798F8B8FE542BBE" blockId="12.[151,1436,1811,1944]" pageId="12" pageNumber="345">
|
||
<emphasis id="B96DEAFEDC12FFC2C798F8B8FEC72B05" bold="true" box="[151,269,1811,1836]" pageId="12" pageNumber="345">FIGURE 6</emphasis>
|
||
.
|
||
<taxonomicName id="4C194D6FDC12FFC2C615F8BFFE242B05" authority="Ahyong & Ng, 2020" authorityName="Ahyong & Ng" authorityYear="2020" box="[282,494,1812,1836]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="345" phylum="Arthropoda" rank="species" species="fulvia" status="sp. nov.">
|
||
<emphasis id="B96DEAFEDC12FFC2C615F8BFFE242B05" box="[282,494,1812,1836]" italics="true" pageId="12" pageNumber="345">Nepinnotheres fulvia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96DEAFEDC12FFC2C6FBF8BFFD8F2B05" bold="true" box="[500,581,1812,1836]" pageId="12" pageNumber="345">
|
||
<taxonomicNameLabel id="A25E5785DC12FFC2C6FBF8BFFD8F2B05" box="[500,581,1812,1836]" pageId="12" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype, cl 2.8 mm, cw 2.9 mm, ZRC 2018.1391. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum and anterior pleon; E, right chela; F–I, right P2–5; J, right P2 proximal articles, ventral view; K, pleon; L, left G1, pleonal view; M, left G2, pleonal view. Scales: A = 1.0 mm; B, L, M = 0.25 mm; C–K = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF666664DC13FFC3C798FA51FC302A72" ID-DOI="http://doi.org/10.5281/zenodo.3954461" ID-Zenodo-Dep="3954461" httpUri="https://zenodo.org/record/3954461/files/figure.png" pageId="13" pageNumber="346" startId="13.[151,250,1530,1555]" targetBox="[155,1435,190,1494]" targetPageId="13">
|
||
<paragraph id="8BA636ECDC13FFC3C798FA51FC302A72" blockId="13.[151,1437,1530,1627]" pageId="13" pageNumber="346">
|
||
<emphasis id="B96DEAFEDC13FFC3C798FA51FEC52A3A" bold="true" box="[151,271,1530,1555]" pageId="13" pageNumber="346">FIGURE 7</emphasis>
|
||
.
|
||
<taxonomicName id="4C194D6FDC13FFC3C611FA50FE3E2A3A" authority="Ahyong & Ng, 2020" authorityName="Ahyong & Ng" authorityYear="2020" box="[286,500,1531,1555]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="346" phylum="Arthropoda" rank="species" species="fulvia" status="sp. nov.">
|
||
<emphasis id="B96DEAFEDC13FFC3C611FA50FE3E2A3A" box="[286,500,1531,1555]" italics="true" pageId="13" pageNumber="346">Nepinnotheres fulvia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96DEAFEDC13FFC3C6F3FA50FD852A3A" bold="true" box="[508,591,1531,1555]" pageId="13" pageNumber="346">
|
||
<taxonomicNameLabel id="A25E5785DC13FFC3C6F3FA50FD852A3A" box="[508,591,1531,1555]" pageId="13" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female paratype, cl 3.0 mm, cw 3.1 mm, ZRC 2018.1392. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, right P2–5; J, right P2 proximal articles, ventral view; K, pleon. Scales: A, K = 1.0 mm; B = 0.25 mm; C–J = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA636ECDC13FFC3C7C8F922FEB22B55" blockId="13.[151,1437,1673,1952]" pageId="13" pageNumber="346">
|
||
The male of
|
||
<taxonomicName id="4C194D6FDC13FFC3C659F921FE7C2A8A" authorityName="Ahyong & Ng" authorityYear="2020" box="[342,438,1673,1699]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="346" phylum="Arthropoda" rank="species" species="fulvia">
|
||
<emphasis id="B96DEAFEDC13FFC3C659F921FE7C2A8A" box="[342,438,1673,1699]" italics="true" pageId="13" pageNumber="346">N. fulvia</emphasis>
|
||
</taxonomicName>
|
||
is mature, indicated by the strongly inflated cheliped and well developed gonopods (
|
||
<figureCitation id="13222A69DC13FFC3C260F922FEEC2AE1" captionStart="FIGURE 6" captionStartId="12.[151,250,1811,1836]" captionTargetBox="[162,1429,245,1773]" captionTargetId="figure@12.[151,1436,237,1786]" captionTargetPageId="12" captionText="FIGURE 6. Nepinnotheres fulvia sp. nov., male holotype, cl 2.8 mm, cw 2.9 mm, ZRC 2018.1391. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum and anterior pleon; E, right chela; F–I, right P2–5; J, right P2 proximal articles, ventral view; K, pleon; L, left G1, pleonal view; M, left G2, pleonal view. Scales: A = 1.0 mm; B, L, M = 0.25 mm; C–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954459" httpUri="https://zenodo.org/record/3954459/files/figure.png" pageId="13" pageNumber="346">Fig. 6A, E, L, M</figureCitation>
|
||
). As in
|
||
<taxonomicName id="4C194D6FDC13FFC3C672F905FD592AE1" authority="(Burger, 1895)" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[381,659,1709,1736]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="346" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B96DEAFEDC13FFC3C672F905FE2F2AE1" box="[381,485,1710,1736]" italics="true" pageId="13" pageNumber="346">N. affinis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF884B1DDC13FFC3C6FAF905FD402AE1" author="Burger, O." box="[501,650,1709,1736]" pageId="13" pageNumber="346" pagination="361 - 390" refId="ref9358" refString="Burger, O. (1895) Ein Beitrag zur kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article" year="1895">Bürger, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from the
|
||
<collectingCountry id="F30E767CDC13FFC3C40CF906FC4E2AEE" box="[771,900,1709,1735]" name="Philippines" pageId="13" pageNumber="346">Philippines</collectingCountry>
|
||
, the G2 exopod in
|
||
<taxonomicName id="4C194D6FDC13FFC3C352F905FB752AEE" authorityName="Ahyong & Ng" authorityYear="2020" box="[1117,1215,1709,1735]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="346" phylum="Arthropoda" rank="species" species="fulvia">
|
||
<emphasis id="B96DEAFEDC13FFC3C352F905FB752AEE" box="[1117,1215,1709,1735]" italics="true" pageId="13" pageNumber="346">N. fulvia</emphasis>
|
||
</taxonomicName>
|
||
is of similar length to the endopod (
|
||
<figureCitation id="13222A69DC13FFC3C642F97AFE612AC2" box="[333,427,1745,1771]" captionStart="FIGURE 6" captionStartId="12.[151,250,1811,1836]" captionTargetBox="[162,1429,245,1773]" captionTargetId="figure@12.[151,1436,237,1786]" captionTargetPageId="12" captionText="FIGURE 6. Nepinnotheres fulvia sp. nov., male holotype, cl 2.8 mm, cw 2.9 mm, ZRC 2018.1391. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum and anterior pleon; E, right chela; F–I, right P2–5; J, right P2 proximal articles, ventral view; K, pleon; L, left G1, pleonal view; M, left G2, pleonal view. Scales: A = 1.0 mm; B, L, M = 0.25 mm; C–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954459" httpUri="https://zenodo.org/record/3954459/files/figure.png" pageId="13" pageNumber="346">Fig. 6M</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF884B1DDC13FFC3C6BAF97AFD5B2AC5" author="Ahyong, S. T. & Komai, T. & Watanabe, T." box="[437,657,1745,1772]" pageId="13" pageNumber="346" pagination="35 - 47" refId="ref8937" refString="Ahyong, S. T., Komai, T. & Watanabe, T. (2012) First Viridotheres Manning, 1996, from Japan, with a key to the species (Decapoda, Brachyura, Pinnotheridae). In: Komatsu, H., Okuno, J. & Fukuoka, K. (Eds.), Studies in Malacostraca: a homage to Masatsune Takeda. Crustaceana Monographs 17. Brill, Leiden, pp. 35 - 47. https: // doi. org / 10.1163 / 9789004202894 _ 003" type="book chapter" year="2012">
|
||
Ahyong
|
||
<emphasis id="B96DEAFEDC13FFC3C519F979FD842AC2" box="[534,590,1745,1771]" italics="true" pageId="13" pageNumber="346">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
: fig. 4B, C). The female
|
||
<typeStatus id="54A2884EDC13FFC3C4A9F979FBCC2AC5" box="[934,1030,1746,1772]" pageId="13" pageNumber="346" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C194D6FDC13FFC3C324F979FB462AC2" authorityName="Ahyong & Ng" authorityYear="2020" box="[1067,1164,1745,1771]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="346" phylum="Arthropoda" rank="species" species="fulvia">
|
||
<emphasis id="B96DEAFEDC13FFC3C324F979FB462AC2" box="[1067,1164,1745,1771]" italics="true" pageId="13" pageNumber="346">N. fulvia</emphasis>
|
||
</taxonomicName>
|
||
is apparently a late subadult, with the pleon broad but yet to reach full size to cover the coxa of the walking legs (
|
||
<figureCitation id="13222A69DC13FFC3C38AF95EFB142B39" box="[1157,1246,1781,1808]" captionStart="FIGURE 7" captionStartId="13.[151,250,1530,1555]" captionTargetBox="[155,1435,190,1494]" captionTargetId="figure@13.[151,1436,181,1505]" captionTargetPageId="13" captionText="FIGURE 7. Nepinnotheres fulvia sp. nov., female paratype, cl 3.0 mm, cw 3.1 mm, ZRC 2018.1392. A, dorsal habitus; B, right maxilliped 3; C, anterior cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, right P2–5; J, right P2 proximal articles, ventral view; K, pleon. Scales: A, K = 1.0 mm; B = 0.25 mm; C–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3954461" httpUri="https://zenodo.org/record/3954461/files/figure.png" pageId="13" pageNumber="346">Fig. 7K</figureCitation>
|
||
). Although a late subadult, diagnostic features are already well developed by this stage in other species of
|
||
<taxonomicName id="4C194D6FDC13FFC3C374F8B2FAD72B1A" authorityName="Manning" authorityYear="1993" box="[1147,1309,1817,1843]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="346" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96DEAFEDC13FFC3C374F8B2FAD72B1A" box="[1147,1309,1817,1843]" italics="true" pageId="13" pageNumber="346">Nepinnotheres</emphasis>
|
||
</taxonomicName>
|
||
, notably in the proportions of the maxilliped 3 articles and pereopodal setation, so we have little hesitation in recognising the novelty of
|
||
<taxonomicName id="4C194D6FDC13FFC3C61FF8C9FEBB2B52" authorityName="Ahyong & Ng" authorityYear="2020" box="[272,369,1889,1915]" class="Malacostraca" family="Pinnotheridae" genus="Nepinnotheres" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="346" phylum="Arthropoda" rank="species" species="fulvia">
|
||
<emphasis id="B96DEAFEDC13FFC3C61FF8C9FEBB2B52" box="[272,369,1889,1915]" italics="true" pageId="13" pageNumber="346">N. fulvia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3036567DC13FFC3C7C8F82EFB8E2BB6" box="[199,1092,1925,1952]" pageId="13" pageNumber="346" type="distribution">
|
||
<paragraph id="8BA636ECDC13FFC3C7C8F82EFB8E2BB6" blockId="13.[151,1437,1673,1952]" box="[199,1092,1925,1952]" pageId="13" pageNumber="346">
|
||
<emphasis id="B96DEAFEDC13FFC3C7C8F82EFE932B89" bold="true" box="[199,345,1925,1952]" pageId="13" pageNumber="346">Distribution</emphasis>
|
||
. Presently known only from off Tabad Island,
|
||
<collectingCountry id="F30E767CDC13FFC3C46BF82DFB8A2BB6" box="[868,1088,1925,1952]" name="Papua New Guinea" pageId="13" pageNumber="346">Papua New Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |