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<document id="C48AB98CFD8063C17CB37578AADCBAFC" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B5CFFAAFF79F2BAFD5AF900" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena koma Perkins 2011, new species" docType="treatment" docVersion="6" lastPageNumber="58" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="58" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1124A45357E81ADE54B81159F9A62570">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="67A47FBF2DA3959EFEE629E9BECF2AC5">Perkins, Philip D.</mods:namePart>
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<treatment id="03B087E55B5CFFAAFF79F2BAFD5AF900" ID-DOI="http://doi.org/10.5281/zenodo.5291510" ID-Zenodo-Dep="5291510" LSID="urn:lsid:plazi:treatment:03B087E55B5CFFAAFF79F2BAFD5AF900" httpUri="http://treatment.plazi.org/id/03B087E55B5CFFAAFF79F2BAFD5AF900" lastPageNumber="58" pageId="57" pageNumber="58">
<subSubSection id="C30365785B5CFFAAFF79F2BAFE4EFF27" box="[151,504,151,177]" pageId="57" pageNumber="58" type="nomenclature">
<paragraph id="8BA636F35B5CFFAAFF79F2BAFE4EFF27" blockId="57.[151,504,151,210]" box="[151,504,151,177]" pageId="57" pageNumber="58">
<heading id="D0EE819F5B5CFFAAFF79F2BAFE4EFF27" bold="true" box="[151,504,151,177]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
<emphasis id="B96DEAE15B5CFFAAFF79F2BAFE4EFF27" bold="true" box="[151,504,151,177]" pageId="57" pageNumber="58">
<taxonomicName id="4C194D705B5CFFAAFF79F2BAFEEFFF27" ID-CoL="3N3CF" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,345,151,177]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="koma" status="sp. nov.">
<emphasis id="B96DEAE15B5CFFAAFF79F2BAFEEFFF27" bold="true" box="[151,345,151,177]" italics="true" pageId="57" pageNumber="58">Hydraena koma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5B5CFFAAFE86F2BAFE4EFF27" box="[360,504,151,177]" pageId="57" pageNumber="58" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30365785B5CFFAAFF79F297FED0FF44" box="[151,358,186,210]" pageId="57" pageNumber="58" type="description">
<paragraph id="8BA636F35B5CFFAAFF79F297FED0FF44" blockId="57.[151,504,151,210]" box="[151,358,186,210]" pageId="57" pageNumber="58">
(
<figureCitation id="13222A765B5CFFAAFF71F297FF4BFF44" box="[159,253,186,210]" captionStart="FIGURE 86" captionStartId="209.[151,250,1753,1776]" captionTargetBox="[197,1392,211,1718]" captionTargetId="figure-26@209.[180,1408,193,1732]" captionTargetPageId="209" captionText="FIGURE 86. Hydraena koma, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283579" httpUri="https://zenodo.org/record/5283579/files/figure.png" pageId="57" pageNumber="58">Figs. 86</figureCitation>
,
<figureCitation id="13222A765B5CFFAAFEE6F297FE90FF44" box="[264,294,186,210]" captionStart="FIGURES 8889" captionStartId="211.[151,264,1561,1584]" captionTargetBox="[249,1332,217,1516]" captionTargetId="figure-29@211.[226,1360,193,1540]" captionTargetPageId="211" captionText="FIGURES 8889. Aedeagi of holotypes. 88. H. koma. 89. H. colorata." figureDoi="http://doi.org/10.5281/zenodo.5283585" httpUri="https://zenodo.org/record/5283585/files/figure.png" pageId="57" pageNumber="58">88</figureCitation>
,
<figureCitation id="13222A765B5CFFAAFEDCF297FEE8FF44" box="[306,350,186,210]" captionStart="FIGURES 467470" captionStartId="392.[151,264,1650,1673]" captionTargetBox="[165,1421,238,1599]" captionTargetId="figure-23@392.[165,1421,193,1629]" captionTargetPageId="392" captionText="FIGURES 467470. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5295735" httpUri="https://zenodo.org/record/5295735/files/figure.png" pageId="57" pageNumber="58">469</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785B5CFFAAFF79F32EFDD7FE10" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="8BA636F35B5CFFAAFF79F32EFDD7FE10" blockId="57.[151,1437,258,1686]" pageId="57" pageNumber="58">
Type Material.
<materialsCitation id="3B713CAE5B5CFFAAFEB8F32EFDD7FE10" collectingDate="2004-05-31" collectionCode="ANIC" collectorName="N. Porch" elevation="100" latitude="-6.6666665" location="Fio River" longLatPrecision="1304" longitude="145.2" pageId="57" pageNumber="58" specimenCount="1" stateProvince="Eastern Highlands" typeStatus="holotype">
<typeStatus id="54A288515B5CFFAAFEB8F32EFE75FE8D" box="[342,451,259,283]" pageId="57" pageNumber="58" type="holotype">
<emphasis id="B96DEAE15B5CFFAAFEB8F32EFE75FE8D" bold="true" box="[342,451,259,283]" pageId="57" pageNumber="58">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15B5CFFAAFDDDF32EFC25FE8D" bold="true" box="[563,915,258,283]" pageId="57" pageNumber="58">
<collectingRegion id="49DDF8115B5CFFAAFDDDF32EFC39FE8D" box="[563,911,258,283]" country="Papua New Guinea" name="Eastern Highlands" pageId="57" pageNumber="58">Eastern Highlands Province</collectingRegion>
:
</emphasis>
<taxonomicName id="4C194D705B5CFFAAFC46F32EFB8EFE8C" authority="River" authorityName="River" box="[936,1080,258,282]" class="Insecta" family="Cicadidae" genus="Koma" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">Koma River</taxonomicName>
, tributary of
<location id="8EC660285B5CFFAAFB3DF32FFAF5FE8C" LSID="urn:lsid:plazi:treatment:03B087E55B5CFFAAFF79F2BAFD5AF900:8EC660285B5CFFAAFB3DF32FFAF5FE8C" box="[1235,1347,258,282]" latitude="-6.6666665" longLatPrecision="1304" longitude="145.2" name="Fio River" pageId="57" pageNumber="58" stateProvince="Eastern Highlands">Fio River</location>
,
<quantity id="4CE19B165B5CFFAAFABCF32FFA2AFE8C" box="[1362,1436,258,282]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="57" pageNumber="58" unit="m" value="100.0">
<elevation id="0034D1C05B5CFFAAFABCF32FFA2AFE8C" box="[1362,1436,258,282]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="57" pageNumber="58" unit="m" value="100.0">100 m</elevation>
</quantity>
downstream of rattan bridge crossing, ca.
<quantity id="4CE19B165B5CFFAAFD81F30BFD0BFEA8" box="[623,701,294,318]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="57" pageNumber="58" unit="km" value="3.8">3.8 km</quantity>
S by
<locationDeviation id="9A142FD15B5CFFAAFD10F30AFC1EFEA8" box="[766,936,294,318]" location="Herowana" pageId="57" pageNumber="58">E of Herowana</locationDeviation>
airstrip, washed from margins of small pool of a steep, rocky, forest stream at confluence with river; gravel, cobbles, wet rock microhabitats,
<quantity id="4CE19B165B5CFFAAFB34F367FA86FEF4" box="[1242,1328,330,354]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="57" pageNumber="58" unit="m" value="1100.0">
<elevation id="0034D1C05B5CFFAAFB34F367FA86FEF4" box="[1242,1328,330,354]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="57" pageNumber="58" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B5CFFAAFAD2F367FA20FEF4" box="[1340,1430,330,354]" degrees="6" direction="south" minutes="40" orientation="latitude" pageId="57" pageNumber="58" precision="925" value="-6.6666665">6° 40' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B5CFFAAFF79F343FEB8FE10" box="[151,270,366,390]" degrees="145" direction="east" minutes="12" orientation="longitude" pageId="57" pageNumber="58" precision="925" value="145.2">145° 12' E</geoCoordinate>
,
<date id="FFA710335B5CFFAAFEF4F343FE3AFE10" box="[282,396,366,390]" pageId="57" pageNumber="58" value="2004-05-31">
<collectingDate id="EFE3E9DB5B5CFFAAFEF4F343FE3AFE10" box="[282,396,366,390]" pageId="57" pageNumber="58" value="2004-05-31">31 v 2004</collectingDate>
</date>
,
<collectorName id="26EC53255B5CFFAAFE76F342FE4DFE10" box="[408,507,366,390]" pageId="57" pageNumber="58">N. Porch</collectorName>
(
<collectionCode id="ED08AE365B5CFFAAFDE4F343FDE3FE10" box="[522,597,366,390]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="57" pageNumber="58">ANIC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B5CFFAAFF29F3BFFC19FDAC" pageId="57" pageNumber="58" type="diagnosis">
<paragraph id="8BA636F35B5CFFAAFF29F3BFFC19FDAC" blockId="57.[151,1437,258,1686]" pageId="57" pageNumber="58">
<emphasis id="B96DEAE15B5CFFAAFF29F3BFFE65FE3D" bold="true" box="[199,467,402,427]" pageId="57" pageNumber="58">Differential Diagnosis.</emphasis>
Somewhat similar in habitus to
<taxonomicName id="4C194D705B5CFFAAFCAAF3BEFC57FE3D" authorityName="Perkins" authorityYear="2011" box="[836,993,403,427]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="biundulata">
<emphasis id="B96DEAE15B5CFFAAFCAAF3BEFCEEFE3C" box="[836,856,403,426]" italics="true" pageId="57" pageNumber="58">H</emphasis>
.
<emphasis id="B96DEAE15B5CFFAAFC86F3BEFC57FE3D" box="[872,993,403,427]" italics="true" pageId="57" pageNumber="58">biundulata</emphasis>
</taxonomicName>
(
<taxonomicName id="4C194D705B5CFFAAFC1DF3BEFBF7FE3C" authorityName="Brunner von Wattenwyl" authorityYear="1878" box="[1011,1089,403,426]" class="Insecta" family="Tettigoniidae" genus="Tetana" kingdom="Animalia" order="Orthoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAE15B5CFFAAFC1DF3BEFBF7FE3C" box="[1011,1089,403,426]" italics="true" pageId="57" pageNumber="58">Tetana</emphasis>
</taxonomicName>
group), including the shape of the sides of the pronotum, which are nearly equally sinuate in front of and behind the widest part (
<figureCitation id="13222A765B5CFFAAFB38F39BFA85FE58" box="[1238,1331,438,462]" captionStart="FIGURE 75" captionStartId="201.[151,250,1753,1776]" captionTargetBox="[234,1356,209,1718]" captionTargetId="figure-26@201.[220,1368,193,1732]" captionTargetPageId="201" captionText="FIGURE 75. Hydraena biundulata, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283547" httpUri="https://zenodo.org/record/5283547/files/figure.png" pageId="57" pageNumber="58">Figs. 75</figureCitation>
,
<figureCitation id="13222A765B5CFFAAFAD0F39BFAEBFE58" box="[1342,1373,438,462]" captionStart="FIGURE 86" captionStartId="209.[151,250,1753,1776]" captionTargetBox="[197,1392,211,1718]" captionTargetId="figure-26@209.[180,1408,193,1732]" captionTargetPageId="209" captionText="FIGURE 86. Hydraena koma, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283579" httpUri="https://zenodo.org/record/5283579/files/figure.png" pageId="57" pageNumber="58">86</figureCitation>
); distinguished therefrom by the markedly arcuate anterior margin of the pronotum, the very different metaventral plaques, and the different elytral coloration. Additionally, males have very differently shaped tibiae in the two species. The male genitalia of the two species differ greatly (
<figureCitation id="13222A765B5CFFAAFCF4F00FFCCEFDAC" box="[794,888,546,570]" captionStart="FIGURES 7677" captionStartId="202.[151,264,1650,1673]" captionTargetBox="[300,1270,221,1616]" captionTargetId="figure-32@202.[278,1309,193,1629]" captionTargetPageId="202" captionText="FIGURES 7677. Aedeagi of holotypes. 76. H. insandalia. 77. H. biundulata." figureDoi="http://doi.org/10.5281/zenodo.5283551" httpUri="https://zenodo.org/record/5283551/files/figure.png" pageId="57" pageNumber="58">Figs. 77</figureCitation>
,
<figureCitation id="13222A765B5CFFAAFC6DF00FFC14FDAC" box="[899,930,546,570]" captionStart="FIGURES 8889" captionStartId="211.[151,264,1561,1584]" captionTargetBox="[249,1332,217,1516]" captionTargetId="figure-29@211.[226,1360,193,1540]" captionTargetPageId="211" captionText="FIGURES 8889. Aedeagi of holotypes. 88. H. koma. 89. H. colorata." figureDoi="http://doi.org/10.5281/zenodo.5283585" httpUri="https://zenodo.org/record/5283585/files/figure.png" pageId="57" pageNumber="58">88</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B5CFFAAFF29F06BFF75F9BC" pageId="57" pageNumber="58" type="description">
<paragraph id="8BA636F35B5CFFAAFF29F06BFE6FFC84" blockId="57.[151,1437,258,1686]" pageId="57" pageNumber="58">
<emphasis id="B96DEAE15B5CFFAAFF29F06BFEEEFDC9" bold="true" box="[199,344,582,607]" inLineHeading="true" pageId="57" pageNumber="58" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515B5CFFAAFE42F06BFDB8FDC8" box="[428,526,582,606]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.64/0.63; head 0.29/0.36; pronotum 0.38/0.45, PA 0.38, PB 0.36; elytra 0.96/0.63. Dorsum of head with frons and clypeus piceous; labrum dark brown; pronotum light brown to testaceous in front of and behind piceous fascia, ratios of color bands, as measured in midline, ca. 6/15/5; elytra dark brown to piceous in ca. anterior 2/5 and narrowly along explanate margin, otherwise elytra light brown to testaceous; legs light brown; maxillary palpi testaceous, tip not darker. Dorsum moderately strongly shining.
</paragraph>
<paragraph id="8BA636F35B5CFFAAFF29F132FC78FBA4" blockId="57.[151,1437,258,1686]" pageId="57" pageNumber="58">Frons punctures ca. 1xef, only slightly larger near eyes, sparser medially; interstices shining, medially 25xpd. Clypeus effacedly microreticulate laterally, very finely sparsely punctulate and shining medially. Mentum and postmentum conjointly weakly concave. Mentum effacedly microreticulate, very sparsely very finely punctulate, weakly shining. Postmentum microreticulate, dull. Genae very slightly raised, weakly shining, with small median impression, without posterior ridge. Pronotum subcordiform, distinctively narrowed both in front of and behind widest area; ca. median 1/2 of anterior margin rather markedly arcuate to posterior; punctures on disc ca. 1xpd largest frons punctures, interstices shining, ca. 25xpd, punctures anteriorly and posteriorly ca. 2xpd those on disc, and denser; PF1 and PF4 absent; PF2 very shallow, obsolete; PF3 deep, wide.</paragraph>
<paragraph id="8BA636F35B5CFFAAFF29F613FCECFB54" blockId="57.[151,1437,258,1686]" pageId="57" pageNumber="58">
Elytra widest at about midlength; summit of posterior declivity at about midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 12xpd, as are interstices between punctures of a row. Apices partly missing in
<typeStatus id="54A288515B5CFFAAFD1EF687FCE3FB54" box="[752,853,1194,1218]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BA636F35B5CFFAAFF29F6E3FF75F9BC" blockId="57.[151,1437,258,1686]" pageId="57" pageNumber="58">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 3/1/6/5. P1 laminate; median carina sinuate in profile, slightly raised between coxae. P2 length/width ca. 5/3, sides very slightly diverging toward blunt apex. Plaques carinate lines, gradually raising from anterior to posterior, at sides of median depression, anteriorly slightly convergent. Metaventrite flat or slightly concave between mesocoxae. AIS width at straight posterior margin ca. 1.5x P2. All legs of moderate length. Profemur (male) with minute tubercle on ventral margin near trochanter; protibia straight, medial margin widened from about midlength to subapically, then narrowed to apex, greatest width ca. 1.5x width at midlength. Mesotibia straight, slender, with many short spine-like setae. Metatibia with medial margin distinctly widened at about midlength, greatest width ca. 1.5x width at apex; lateral margin emarginate. Abdominal apex symmetrical; last tergite (male) with large, deep apicomedian notch. Aedeagus as illustrated (
<figureCitation id="13222A765B5CFFAAFA80F7C3FF00F9BC" captionStart="FIGURES 8889" captionStartId="211.[151,264,1561,1584]" captionTargetBox="[249,1332,217,1516]" captionTargetId="figure-29@211.[226,1360,193,1540]" captionTargetPageId="211" captionText="FIGURES 8889. Aedeagi of holotypes. 88. H. koma. 89. H. colorata." figureDoi="http://doi.org/10.5281/zenodo.5283585" httpUri="https://zenodo.org/record/5283585/files/figure.png" pageId="57" pageNumber="58">Fig. 88</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B5CFFAAFF29F41AFCBBF9D8" box="[199,781,1590,1615]" pageId="57" pageNumber="58" type="etymology">
<paragraph id="8BA636F35B5CFFAAFF29F41AFCBBF9D8" blockId="57.[151,1437,258,1686]" box="[199,781,1590,1615]" pageId="57" pageNumber="58">
<emphasis id="B96DEAE15B5CFFAAFF29F41AFEFBF9D9" bold="true" box="[199,333,1591,1615]" inLineHeading="true" pageId="57" pageNumber="58" reason="1">Etymology.</emphasis>
Named in reference to the
<taxonomicName id="4C194D705B5CFFAAFD90F41AFCBBF9D8" authority="River." authorityName="River." box="[638,781,1590,1614]" class="Insecta" family="Cicadidae" genus="Koma" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">Koma River.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B5CFFAAFF29F477FD5AF900" pageId="57" pageNumber="58" type="distribution">
<paragraph id="8BA636F35B5CFFAAFF29F477FD5AF900" blockId="57.[151,1437,258,1686]" pageId="57" pageNumber="58">
<emphasis id="B96DEAE15B5CFFAAFF29F477FED6F9E5" bold="true" box="[199,352,1626,1651]" inLineHeading="true" pageId="57" pageNumber="58" reason="1">Distribution.</emphasis>
Currently known only from the
<typeStatus id="54A288515B5CFFAAFD24F476FD4CF9E5" box="[714,762,1627,1651]" pageId="57" pageNumber="58">type</typeStatus>
locality, the
<taxonomicName id="4C194D705B5CFFAAFC67F476FBA4F9E4" authority="River" authorityName="River" box="[905,1042,1626,1650]" class="Insecta" family="Cicadidae" genus="Koma" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">Koma River</taxonomicName>
in southeastern Area 1, near where Area 1 borders Area 13; elevation
<quantity id="4CE19B165B5CFFAAFDF4F453FDD8F900" box="[538,622,1662,1686]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="57" pageNumber="58" unit="m" value="1100.0">1100 m</quantity>
(
<figureCitation id="13222A765B5CFFAAFD93F453FD56F900" box="[637,736,1662,1686]" captionStart="FIGURES 467470" captionStartId="392.[151,264,1650,1673]" captionTargetBox="[165,1421,238,1599]" captionTargetId="figure-23@392.[165,1421,193,1629]" captionTargetPageId="392" captionText="FIGURES 467470. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5295735" httpUri="https://zenodo.org/record/5295735/files/figure.png" pageId="57" pageNumber="58">Fig. 469</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>