treatments-xml/data/03/B0/87/03B087E55B15FFE2FF79F43FFB9BFB36.xml
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<document id="D670B5DF1A06BD7197313A8D6E68BF0B" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B15FFE2FF79F43FFB9BFB36" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena pala Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="114" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="113" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F7867385FD389DD5C5447F0A101684FC">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
</mods:titleInfo>
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<mods:namePart id="4C8255A4DADA73C33561AA821F91F0FD">Perkins, Philip D.</mods:namePart>
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<mods:date id="111DA62780EB7F1668C89B953EF8F467">2011</mods:date>
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<mods:number id="85DEAC9B89A39C8616AC8A1ACDF6D083">2011-07-08</mods:number>
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<mods:url id="10CA6558E78D9D623CD1247ED0BE8C8F">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
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<mods:classification id="40C10A9EF46DC1E00BE18E238CD27D6D">journal article</mods:classification>
<mods:identifier id="F1A9AA0D4AA3CECDDCB8EB4457F1E7CC" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
<mods:identifier id="EC662B927517B4BDAB6D25EE466B65BB" type="ISSN">1175­5334</mods:identifier>
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<treatment id="03B087E55B15FFE2FF79F43FFB9BFB36" ID-DOI="http://doi.org/10.5281/zenodo.5291739" ID-Zenodo-Dep="5291739" LSID="urn:lsid:plazi:treatment:03B087E55B15FFE2FF79F43FFB9BFB36" httpUri="http://treatment.plazi.org/id/03B087E55B15FFE2FF79F43FFB9BFB36" lastPageId="113" lastPageNumber="114" pageId="112" pageNumber="113">
<subSubSection id="C30365785B15FFE3FF79F43FFE5FF9BA" box="[151,489,1554,1580]" pageId="112" pageNumber="113" type="nomenclature">
<paragraph id="8BA636F35B15FFE3FF79F43FFE5FF9BA" blockId="112.[151,489,1554,1613]" box="[151,489,1554,1580]" pageId="112" pageNumber="113">
<heading id="D0EE819F5B15FFE3FF79F43FFE5FF9BA" bold="true" box="[151,489,1554,1580]" fontSize="11" level="1" pageId="112" pageNumber="113" reason="1">
<emphasis id="B96DEAE15B15FFE3FF79F43FFE5FF9BA" bold="true" box="[151,489,1554,1580]" pageId="112" pageNumber="113">
<taxonomicName id="4C194D705B15FFE3FF79F43FFEFCF9BA" ID-CoL="6MJC6" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,330,1554,1580]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="pala" status="sp. nov.">
<emphasis id="B96DEAE15B15FFE3FF79F43FFEFCF9BA" bold="true" box="[151,330,1554,1580]" italics="true" pageId="112" pageNumber="113">Hydraena pala</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5B15FFE3FEB7F43FFE5FF9BA" box="[345,489,1554,1580]" pageId="112" pageNumber="113" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B15FFE3FF79F418FE0FF9DB" box="[151,441,1589,1613]" pageId="112" pageNumber="113" type="description">
<paragraph id="8BA636F35B15FFE3FF79F418FE0FF9DB" blockId="112.[151,489,1554,1613]" box="[151,441,1589,1613]" pageId="112" pageNumber="113">
(
<figureCitation id="13222A765B15FFE3FF71F418FEBAF9DB" box="[159,268,1589,1613]" captionStart="FIGURE 202" captionStartId="296.[151,250,1753,1776]" captionTargetBox="[224,1362,193,1732]" captionTargetId="figure-29@296.[224,1362,193,1732]" captionTargetPageId="296" captionText="FIGURE 202. Hydraena pala, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283804" httpUri="https://zenodo.org/record/5283804/files/figure.png" pageId="112" pageNumber="113">Figs. 202</figureCitation>
,
<figureCitation id="13222A765B15FFE3FEF8F418FEF5F9DB" box="[278,323,1589,1613]" captionStart="FIGURES 204205" captionStartId="298.[151,264,1650,1673]" captionTargetBox="[293,1301,219,1614]" captionTargetId="figure-32@298.[226,1360,193,1629]" captionTargetPageId="298" captionText="FIGURES 204205. Aedeagi of holotypes. 204. H. pala. 205. H. maculopala." figureDoi="http://doi.org/10.5281/zenodo.5283812" httpUri="https://zenodo.org/record/5283812/files/figure.png" pageId="112" pageNumber="113">204</figureCitation>
,
<figureCitation id="13222A765B15FFE3FEA0F418FECFF9DB" box="[334,377,1589,1613]" captionStart="FIGURES 408411" captionStartId="374.[151,264,1650,1673]" captionTargetBox="[245,1333,206,1610]" captionTargetId="figure-28@374.[231,1355,193,1629]" captionTargetPageId="374" captionText="FIGURES 408411. Female tergite X, gonocoxite, and spermatheca of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284033" httpUri="https://zenodo.org/record/5284033/files/figure.png" pageId="112" pageNumber="113">411</figureCitation>
,
<figureCitation id="13222A765B15FFE3FE6BF418FE06F9DB" box="[389,432,1589,1613]" captionStart="FIGURES 527530" captionStartId="407.[151,264,1650,1673]" captionTargetBox="[165,1421,224,1616]" captionTargetId="figure-20@407.[165,1421,193,1629]" captionTargetPageId="407" captionText="FIGURES 527530. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284135" httpUri="https://zenodo.org/record/5284135/files/figure.png" pageId="112" pageNumber="113">527</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785B15FFE3FF79F450FE4BF8FB" pageId="112" pageNumber="113" type="materials_examined">
<paragraph id="8BA636F35B15FFE3FF79F450FE4BF8FB" blockId="112.[151,1437,1661,2009]" pageId="112" pageNumber="113">
Type Material.
<materialsCitation id="3B713CAE5B15FFE3FEBAF450FBDCF92F" collectingDate="1964-12-30" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1000" latitude="-6.8333335" location="Gurakor Creek" longLatPrecision="1303" longitude="146.61667" municipality="Lae - Bulolo" pageId="112" pageNumber="113" specimenCount="1" stateProvince="Morobe" typeStatus="holotype">
<typeStatus id="54A288515B15FFE3FEBAF450FE77F903" box="[340,449,1661,1685]" pageId="112" pageNumber="113" type="holotype">
<emphasis id="B96DEAE15B15FFE3FEBAF450FE77F903" bold="true" box="[340,449,1661,1685]" pageId="112" pageNumber="113">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15B15FFE3FDC1F450FCBFF900" bold="true" box="[559,777,1661,1686]" pageId="112" pageNumber="113">
<collectingRegion id="49DDF8115B15FFE3FDC1F450FCB3F900" box="[559,773,1661,1686]" country="Papua New Guinea" name="Morobe" pageId="112" pageNumber="113">Morobe Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BC2AC895B15FFE3FCF3F450FC17F903" box="[797,929,1661,1685]" pageId="112" pageNumber="113">LaeBulolo</collectingMunicipality>
road,
<location id="8EC660285B15FFE3FC03F450FB2CF903" LSID="urn:lsid:plazi:treatment:03B087E55B15FFE2FF79F43FFB9BFB36:8EC660285B15FFE3FC03F450FB2CF903" box="[1005,1178,1661,1685]" latitude="-6.8333335" longLatPrecision="1303" longitude="146.61667" municipality="Lae - Bulolo" name="Gurakor Creek" pageId="112" pageNumber="113" stateProvince="Morobe">Gurakor Creek</location>
, ex gravel banks, [GE est.]
<quantity id="4CE19B165B15FFE3FF20F48CFEEDF92F" box="[206,347,1697,1721]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="112" pageNumber="113" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">
<elevation id="0034D1C05B15FFE3FF20F48CFEEDF92F" box="[206,347,1697,1721]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="112" pageNumber="113" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">9001100 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B15FFE3FE89F48CFE77F92F" box="[359,449,1697,1721]" degrees="6" direction="south" minutes="50" orientation="latitude" pageId="112" pageNumber="113" precision="925" value="-6.8333335">6° 50' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B15FFE3FE20F48CFDF3F92F" box="[462,581,1697,1721]" degrees="146" direction="east" minutes="37" orientation="longitude" pageId="112" pageNumber="113" precision="925" value="146.61667">146° 37' E</geoCoordinate>
,
<date id="FFA710335B15FFE3FDBFF48CFD62F92F" box="[593,724,1697,1721]" pageId="112" pageNumber="113" value="1964-12-30">
<collectingDate id="EFE3E9DB5B15FFE3FDBFF48CFD62F92F" box="[593,724,1697,1721]" pageId="112" pageNumber="113" value="1964-12-30">30 xii 1964</collectingDate>
</date>
,
<collectorName id="26EC53255B15FFE3FD31F48FFC32F92F" box="[735,900,1697,1721]" pageId="112" pageNumber="113">M. E. Bacchus</collectorName>
(MEB 133) (
<collectionCode id="ED08AE365B15FFE3FBF5F48CFBD4F92F" box="[1051,1122,1697,1721]" pageId="112" pageNumber="113">NHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B713CAE5B15FFE3FB9BF48CFC9BF8B3" collectingDate="1988-07-01" collectingDateMax="1988-08-30" collectingDateMin="1988-07-01" collectionCode="NMW" collectorName="D. Dudgeon" elevation="160" latitude="-5.55" location="Koje Creek" longLatPrecision="1305" longitude="145.38333" municipality="Ramu Basin" pageId="112" pageNumber="113" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<emphasis id="B96DEAE15B15FFE3FB9BF48CFB5BF92F" bold="true" box="[1141,1261,1697,1721]" pageId="112" pageNumber="113">
<typeStatus id="54A288515B15FFE3FB9BF48CFB5BF92F" box="[1141,1261,1697,1721]" pageId="112" pageNumber="113" type="paratype">Paratypes</typeStatus>
</emphasis>
(14):
<emphasis id="B96DEAE15B15FFE3FAD6F48CFEBFF948" bold="true" pageId="112" pageNumber="113">
<collectingRegion id="49DDF8115B15FFE3FAD6F48CFEB3F948" country="Papua New Guinea" name="Madang" pageId="112" pageNumber="113">Madang Province</collectingRegion>
:
</emphasis>
<collectingRegion id="49DDF8115B15FFE3FEF8F4E8FEE3F94B" box="[278,341,1733,1757]" country="Papua New Guinea" name="West Sepik" pageId="112" pageNumber="113">Sepik</collectingRegion>
<collectingMunicipality id="6BC2AC895B15FFE3FEB2F4EBFE51F94B" box="[348,487,1733,1757]" pageId="112" pageNumber="113">Ramu Basin</collectingMunicipality>
,
<location id="8EC660285B15FFE3FE1CF4E8FDC3F94B" LSID="urn:lsid:plazi:treatment:03B087E55B15FFE2FF79F43FFB9BFB36:8EC660285B15FFE3FE1CF4E8FDC3F94B" box="[498,629,1733,1757]" latitude="-5.55" longLatPrecision="1305" longitude="145.38333" municipality="Ramu Basin" name="Koje Creek" pageId="112" pageNumber="113" stateProvince="Madang">Kojé Creek</location>
, unshaded creek, small to medium cobbles, gravel and fine sand (the latter in areas of slack water), filamentous algae on stony substrates, small accumulations of leaf litter,
<quantity id="4CE19B165B15FFE3FAEBF4C4FAF8F897" box="[1285,1358,1769,1793]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="112" pageNumber="113" unit="m" value="160.0">
<elevation id="0034D1C05B15FFE3FAEBF4C4FAF8F897" box="[1285,1358,1769,1793]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="112" pageNumber="113" unit="m" value="160.0">160 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B15FFE3FAB4F4C4FF1EF8B3" degrees="5" direction="south" minutes="33" orientation="latitude" pageId="112" pageNumber="113" precision="925" value="-5.55">5° 33' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B15FFE3FF58F520FE86F8B3" box="[182,304,1805,1829]" degrees="145" direction="east" minutes="23" orientation="longitude" pageId="112" pageNumber="113" precision="925" value="145.38333">145° 23' E</geoCoordinate>
,
<date id="FFA710335B15FFE3FED3F520FDBAF8B3" box="[317,524,1805,1829]" pageId="112" pageNumber="113" value="1988-07-01" valueMax="1988-08-30" valueMin="1988-07-01">
<collectingDate id="EFE3E9DB5B15FFE3FED3F520FDBAF8B3" box="[317,524,1805,1829]" pageId="112" pageNumber="113" value="1988-07-01" valueMax="1988-08-30" valueMin="1988-07-01">1 vii30 viii 1988</collectingDate>
</date>
,
<collectorName id="26EC53255B15FFE3FDF6F523FD14F8B3" box="[536,674,1805,1829]" pageId="112" pageNumber="113">D. Dudgeon</collectorName>
(10
<collectionCode id="ED08AE365B15FFE3FD36F520FC92F8B3" box="[728,804,1805,1829]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="112" pageNumber="113">NMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B15FFE3FCD7F520FA2EF8B3" box="[825,1432,1805,1830]" collectingDate="1964-12-30" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="1000" latitude="-6.8333335" location="Gurakor Creek" longLatPrecision="1303" longitude="146.61667" municipality="Lae - Bulolo" pageId="112" pageNumber="113" specimenCount="1" stateProvince="Morobe" typeStatus="paratype">
<emphasis id="B96DEAE15B15FFE3FCD7F520FBA4F8B0" bold="true" box="[825,1042,1805,1830]" pageId="112" pageNumber="113">
<collectingRegion id="49DDF8115B15FFE3FCD7F520FBBBF8B0" box="[825,1037,1805,1830]" country="Papua New Guinea" name="Morobe" pageId="112" pageNumber="113">Morobe Province</collectingRegion>
:
</emphasis>
Same data as holotype (3
<collectionCode id="ED08AE365B15FFE3FAA4F520FA26F8B3" box="[1354,1424,1805,1829]" pageId="112" pageNumber="113">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B15FFE3FF79F51CFE4FF8FB" collectingDate="1979-05-01" collectionCode="MHNG" collectorName="W. G. Ullrich" county="Bayer" elevation="1200" latitude="-5.5333333" location="Antenge River" longLatPrecision="1305" longitude="144.3" municipality="Rokina" pageId="112" pageNumber="113" specimenCount="1" stateProvince="Western Highlands" typeStatus="paratype">
<emphasis id="B96DEAE15B15FFE3FF79F51CFE4FF8DC" bold="true" box="[151,505,1841,1866]" pageId="112" pageNumber="113">
<collectingRegion id="49DDF8115B15FFE3FF79F51CFE42F8DC" box="[151,500,1841,1866]" country="Papua New Guinea" name="Western Highlands" pageId="112" pageNumber="113">Western Highlands Province</collectingRegion>
:
</emphasis>
<collectingCounty id="62C74E7F5B15FFE3FDE5F51CFDF8F8DF" box="[523,590,1841,1865]" pageId="112" pageNumber="113">Bayer</collectingCounty>
/
<collectingMunicipality id="6BC2AC895B15FFE3FDB8F51CFD1FF8DF" box="[598,681,1841,1865]" pageId="112" pageNumber="113">Rokina</collectingMunicipality>
,
<location id="8EC660285B15FFE3FD5BF51CFCEBF8DF" LSID="urn:lsid:plazi:treatment:03B087E55B15FFE2FF79F43FFB9BFB36:8EC660285B15FFE3FD5BF51CFCEBF8DF" box="[693,861,1841,1865]" county="Bayer" latitude="-5.5333333" longLatPrecision="1305" longitude="144.3" municipality="Rokina" name="Antenge River" pageId="112" pageNumber="113" stateProvince="Western Highlands">Antenge River</location>
, [GE est.]
<quantity id="4CE19B165B15FFE3FC37F51CFBC1F8DF" box="[985,1143,1841,1865]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="112" pageNumber="113" unit="m" value="1200.0" valueMax="1300.0" valueMin="1100.0">
<elevation id="0034D1C05B15FFE3FC37F51CFBC1F8DF" box="[985,1143,1841,1865]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="112" pageNumber="113" unit="m" value="1200.0" valueMax="1300.0" valueMin="1100.0">11001300 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B15FFE3FB6AF51CFB57F8DF" box="[1156,1249,1841,1865]" degrees="5" direction="south" minutes="32" orientation="latitude" pageId="112" pageNumber="113" precision="925" value="-5.5333333">5° 32' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B15FFE3FB01F51CFADFF8DF" box="[1263,1385,1841,1865]" degrees="144" direction="east" minutes="18" orientation="longitude" pageId="112" pageNumber="113" precision="925" value="144.3">144° 18' E</geoCoordinate>
,
<date id="FFA710335B15FFE3FA99F51CFF67F8FB" pageId="112" pageNumber="113" value="1979-05-01">
<collectingDate id="EFE3E9DB5B15FFE3FA99F51CFF67F8FB" pageId="112" pageNumber="113" value="1979-05-01">1 v 1979</collectingDate>
</date>
,
<collectorName id="26EC53255B15FFE3FF33F57BFEC6F8FB" box="[221,368,1877,1901]" pageId="112" pageNumber="113">W. G. Ullrich</collectorName>
(1
<collectionCode id="ED08AE365B15FFE3FE78F578FE47F8FB" box="[406,497,1877,1901]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="112" pageNumber="113" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B15FFE2FF29F554FED8FF42" lastPageId="113" lastPageNumber="114" pageId="112" pageNumber="113" type="diagnosis">
<paragraph id="8BA636F35B15FFE2FF29F554FED8FF42" blockId="112.[151,1437,1661,2009]" lastBlockId="113.[151,1437,151,1184]" lastPageId="113" lastPageNumber="114" pageId="112" pageNumber="113">
<emphasis id="B96DEAE15B15FFE3FF29F554FE64F804" bold="true" box="[199,466,1913,1938]" pageId="112" pageNumber="113">Differential Diagnosis.</emphasis>
Very similar in color, body form and dorsal sculpture to
<taxonomicName id="4C194D705B15FFE3FBBDF557FB6FF807" authorityName="Perkins" authorityYear="2011" box="[1107,1241,1913,1937]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="palamita">
<emphasis id="B96DEAE15B15FFE3FBBDF557FBD1F807" box="[1107,1127,1914,1937]" italics="true" pageId="112" pageNumber="113">H</emphasis>
.
<emphasis id="B96DEAE15B15FFE3FB9BF554FB6FF807" box="[1141,1241,1913,1937]" italics="true" pageId="112" pageNumber="113">palamita</emphasis>
</taxonomicName>
(
<figureCitation id="13222A765B15FFE3FB07F554FAE3F807" box="[1257,1365,1913,1937]" captionStart="FIGURE 202" captionStartId="296.[151,250,1753,1776]" captionTargetBox="[224,1362,193,1732]" captionTargetId="figure-29@296.[224,1362,193,1732]" captionTargetPageId="296" captionText="FIGURE 202. Hydraena pala, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283804" httpUri="https://zenodo.org/record/5283804/files/figure.png" pageId="112" pageNumber="113">Figs. 202</figureCitation>
,
<figureCitation id="13222A765B15FFE3FA8FF554FA39F807" box="[1377,1423,1913,1937]" captionStart="FIGURE 207" captionStartId="300.[151,250,1753,1776]" captionTargetBox="[252,1338,209,1719]" captionTargetId="figure-29@300.[242,1345,193,1732]" captionTargetPageId="300" captionText="FIGURE 207. Hydraena palamita, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283820" httpUri="https://zenodo.org/record/5283820/files/figure.png" pageId="112" pageNumber="113">207</figureCitation>
); differing therefrom in slightly larger size (ca. 1.58 vs.
<quantity id="4CE19B165B15FFE3FCE4F5B0FCC6F823" box="[778,880,1949,1973]" metricMagnitude="-3" metricUnit="m" metricValue="1.47" pageId="112" pageNumber="113" unit="mm" value="1.47">1.47 mm</quantity>
) and the slightly more widely separated plaques (ratios ca. 6/1/6/6 vs. 6/1/6/5). The male genitalia show a similarity in basic plan, but differ distinctively in several details, most notably the shape of the left paramere (
<figureCitation id="13222A765B14FFE2FD08F2BAFCE4FF39" box="[742,850,151,175]" captionStart="FIGURES 204205" captionStartId="298.[151,264,1650,1673]" captionTargetBox="[293,1301,219,1614]" captionTargetId="figure-32@298.[226,1360,193,1629]" captionTargetPageId="298" captionText="FIGURES 204205. Aedeagi of holotypes. 204. H. pala. 205. H. maculopala." figureDoi="http://doi.org/10.5281/zenodo.5283812" httpUri="https://zenodo.org/record/5283812/files/figure.png" pageId="113" pageNumber="114">Figs. 204</figureCitation>
,
<figureCitation id="13222A765B14FFE2FCB1F2BAFC3BFF39" box="[863,909,151,175]" captionStart="FIGURES 208209" captionStartId="301.[151,264,1650,1673]" captionText="FIGURES 208209. Aedeagi of holotypes. 208. H. nanopala. 209. H. palamita." pageId="113" pageNumber="114">209</figureCitation>
); reliable determinations will require examination of the aedeagi.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B14FFE2FF29F2F2FE32FBA2" pageId="113" pageNumber="114" type="description">
<paragraph id="8BA636F35B14FFE2FF29F2F2FB50FEF2" blockId="113.[151,1437,151,1184]" pageId="113" pageNumber="114">
<emphasis id="B96DEAE15B14FFE2FF29F2F2FEEEFF6E" bold="true" box="[199,344,223,248]" inLineHeading="true" pageId="113" pageNumber="114" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515B14FFE2FE42F2F2FDB8FF61" box="[428,526,223,247]" pageId="113" pageNumber="114" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.58/0.66; head 0.25/0.36; pronotum 0.38/0.49, PA 0.37, PB 0.40; elytra 0.97/0.66. Dorsum of head dark brown to reddish brown; pronotum light brown to testaceous; elytra dark brown to reddish brown, with very faint slightly more lightly colored macula at posterior declivity; legs brown to light reddish brown; maxillary palpi testaceous, tip not darker.
</paragraph>
<paragraph id="8BA636F35B14FFE2FF29F35DFE54FDAA" blockId="113.[151,1437,151,1184]" pageId="113" pageNumber="114">Frons punctures ca. 1xef, slightly larger near eyes than medially; interstices shining, 14xpd. Clypeus microreticulate laterally, very finely sparsely punctate medially. Mentum very sparsely very finely punctulate, shining; postmentum micropunctulate. Genae raised, shining, without posterior ridge. Pronotum subcordiform, ca. median 3/4 of anterior margin emarginate; punctures on disc ca. 1xpd frons punctures, interstices shining, 13xpd, punctures slightly larger and denser at anterior and posterior; PF1 very shallow, obsolete; PF2 very shallow; PF3 moderately deep; PF4 very shallow.</paragraph>
<paragraph id="8BA636F35B14FFE2FF29F06AFF47FD79" blockId="113.[151,1437,151,1184]" pageId="113" pageNumber="114">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 12xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="8BA636F35B14FFE2FF29F0D1FE32FBA2" blockId="113.[151,1437,151,1184]" pageId="113" pageNumber="114">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 6/1/6/6. P1 ca. 1/2 P2; median carina straight in profile; postcoxal process medially concave. P2 transversely concave, length slightly less than width, sides slightly diverging toward blunt apex. Plaques very narrow, carinate, converging slightly anteriorly, at sides of very deep median depression. Metaventrite concave between P2 and plaques. AIS width at straight posterior margin ca. 1.5x P2. All legs moderately long and slender. Profemur (male) with very small tubercle next to trochanter; protibia slightly arcuate, distal 1/3 widened anteriorly, medial margin flat except small sharp tubercle subapically on posteromedial margin. Meso- and metatibia straight. Abdominal apex symmetrical; last tergite (male) deeply notched, each side of notch produced in point. Aedeagus as illustrated (
<figureCitation id="13222A765B14FFE2FD02F1DAFCE7FB99" box="[748,849,1015,1039]" captionStart="FIGURES 204205" captionStartId="298.[151,264,1650,1673]" captionTargetBox="[293,1301,219,1614]" captionTargetId="figure-32@298.[226,1360,193,1629]" captionTargetPageId="298" captionText="FIGURES 204205. Aedeagi of holotypes. 204. H. pala. 205. H. maculopala." figureDoi="http://doi.org/10.5281/zenodo.5283812" httpUri="https://zenodo.org/record/5283812/files/figure.png" pageId="113" pageNumber="114">Fig. 204</figureCitation>
). Female tergite X, gonocoxite, and spermatheca as illustrated (
<figureCitation id="13222A765B14FFE2FEF8F631FEC1FBA2" box="[278,375,1052,1076]" captionStart="FIGURES 408411" captionStartId="374.[151,264,1650,1673]" captionTargetBox="[245,1333,206,1610]" captionTargetId="figure-28@374.[231,1355,193,1629]" captionTargetPageId="374" captionText="FIGURES 408411. Female tergite X, gonocoxite, and spermatheca of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284033" httpUri="https://zenodo.org/record/5284033/files/figure.png" pageId="113" pageNumber="114">Fig. 411</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B14FFE2FF29F66DFB62FBCE" box="[199,1236,1088,1112]" pageId="113" pageNumber="114" type="etymology">
<paragraph id="8BA636F35B14FFE2FF29F66DFB62FBCE" blockId="113.[151,1437,151,1184]" box="[199,1236,1088,1112]" pageId="113" pageNumber="114">
<emphasis id="B96DEAE15B14FFE2FF29F66DFEFBFBCE" bold="true" box="[199,333,1088,1112]" inLineHeading="true" pageId="113" pageNumber="114" reason="1">Etymology.</emphasis>
&quot;Socket of a ring for a jewel, bezel&quot;; named in reference to the prosternal fovea.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B14FFE2FF29F64EFB9BFB36" pageId="113" pageNumber="114" type="distribution">
<paragraph id="8BA636F35B14FFE2FF29F64EFB9BFB36" blockId="113.[151,1437,151,1184]" pageId="113" pageNumber="114">
<emphasis id="B96DEAE15B14FFE2FF29F64EFED6FBEA" bold="true" box="[199,352,1123,1148]" inLineHeading="true" pageId="113" pageNumber="114" reason="1">Distribution.</emphasis>
Currently known from three localities at three quite different elevations: central Area 1 (
<quantity id="4CE19B165B14FFE2FAB8F649FF59FB36" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="113" pageNumber="114" unit="m" value="1200.0" valueMax="1300.0" valueMin="1100.0">1100 1300 m</quantity>
), eastern Area 2 (
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), and far eastern Area 2 (
<quantity id="4CE19B165B14FFE2FCF7F6A5FC11FB36" box="[793,935,1160,1184]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="113" pageNumber="114" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">9001100 m</quantity>
) (
<figureCitation id="13222A765B14FFE2FC51F6A5FBA8FB36" box="[959,1054,1160,1184]" captionStart="FIGURES 527530" captionStartId="407.[151,264,1650,1673]" captionTargetBox="[165,1421,224,1616]" captionTargetId="figure-20@407.[165,1421,193,1629]" captionTargetPageId="407" captionText="FIGURES 527530. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284135" httpUri="https://zenodo.org/record/5284135/files/figure.png" pageId="113" pageNumber="114">Fig. 527</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>