635 lines
115 KiB
XML
635 lines
115 KiB
XML
<document id="7E12493BC03423CE4F3E4DE297469E93" ID-CLB-Dataset="216781" ID-DOI="10.26107/RBZ-2022-0026" ID-GBIF-Dataset="d9ac69ed-346e-42d3-9f0b-53b1a92bded7" ID-ISSN="2345-7600" ID-Zenodo-Dep="7399900" ID-ZooBank="2E07F63A-50B5-42CE-98FA-027B0BF38756" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1670242937288" checkinUser="felipe" docAuthor="Murniati, Dewi Citra, Asakura, Akira, Nugroho, Dharma Arif, Hernawan, Udhi Eko & Dharmawan, Wayan Eka" docDate="2022" docId="03B087C64E59FF9B613C25AE4D3DFCD5" docLanguage="en" docName="RafflesBZool.70.461–491.pdf" docOrigin="Raffles Bulletin of Zoology 70" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Tmethypocoelis liki Murniati & Asakura & Nugroho & Hernawan & Dharmawan 2022, new species" docType="treatment" docVersion="7" lastPageNumber="489" masterDocId="FF89FFBE4E4DFF87602D231D4C4EFFA0" masterDocTitle="On a collection of thoracotreme crabs (Crustacea: Brachyura: Ocypodidae, Macrophthalmidae, Dotillidae) from two offshore islands of Papua, eastern Indonesia, with descriptions of two new species" masterLastPageNumber="491" masterPageNumber="461" pageNumber="481" updateTime="1699412988745" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B840C7230EBC1568859F8A8424218F96">On a collection of thoracotreme crabs (Crustacea: Brachyura: Ocypodidae, Macrophthalmidae, Dotillidae) from two offshore islands of Papua, eastern Indonesia, with descriptions of two new species</mods:title>
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<mods:namePart id="95086FDCF63A43BA63B3C015E24F48FB">Murniati, Dewi Citra</mods:namePart>
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<mods:namePart id="E1E30C0ED60AE9F89E3EF0A6CF93ABFF">Asakura, Akira</mods:namePart>
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<mods:namePart id="182E57468A4B6DD7EAA1324CB7DB13E6">Nugroho, Dharma Arif</mods:namePart>
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<mods:namePart id="84A30A1AD0BDD3583E9D079D1521C5DA">Hernawan, Udhi Eko</mods:namePart>
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<mods:namePart id="EA5F0E02389085336175B724EA5A95F2">Dharmawan, Wayan Eka</mods:namePart>
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<mods:title id="CEE0A63363379A42B612E7E1FF6ED93D">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="EAEDAAD33D96998AC12C64A41B2E472E">2022</mods:date>
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<treatment id="03B087C64E59FF9B613C25AE4D3DFCD5" ID-DOI="http://doi.org/10.5281/zenodo.7502090" ID-GBIF-Taxon="208533013" ID-Zenodo-Dep="7502090" LSID="urn:lsid:plazi:treatment:03B087C64E59FF9B613C25AE4D3DFCD5" httpUri="http://treatment.plazi.org/id/03B087C64E59FF9B613C25AE4D3DFCD5" lastPageId="28" lastPageNumber="489" pageId="20" pageNumber="481">
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<paragraph id="8BA636D04E59FF93613C25AE4E30F96B" blockId="20.[273,638,1715,1772]" box="[273,638,1715,1740]" pageId="20" pageNumber="481">
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<taxonomicName id="4C194D534E59FF93613C25AE4DA4F96B" ID-CoL="f1a4025f-058c-4ab8-9844-ef0f1c2a5fb9" authority="Murniati & Asakura & Nugroho & Hernawan & Dharmawan, 2022" authorityName="Murniati & Asakura & Nugroho & Hernawan & Dharmawan" authorityYear="2022" box="[273,490,1715,1739]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="481" phylum="Arthropoda" rank="species" species="liki" status="sp. nov.">
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<emphasis id="B96DEAC24E59FF93613C25AE4DA4F96B" bold="true" box="[273,490,1715,1739]" italics="true" pageId="20" pageNumber="481">Tmethypocoelis liki</emphasis>
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,
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<taxonomicNameLabel id="A25E57B94E59FF9361D425A94E30F96B" box="[505,638,1715,1740]" pageId="20" pageNumber="481" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8BA636D04E59FF93615225C94E5EF94C" blockId="20.[273,638,1715,1772]" box="[383,528,1748,1772]" pageId="20" pageNumber="481">
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(
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<figureCitation id="13222A554E59FF9361AB25C94E47F94C" box="[390,521,1748,1772]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStartId-0="21.[113,148,1127,1149]" captionStartId-1="22.[136,171,1700,1722]" captionStartId-2="23.[113,148,1745,1767]" captionStartId-3="24.[136,171,1129,1151]" captionStartId-4="25.[113,148,1285,1307]" captionStartId-5="26.[135,170,2064,2086]" captionStartId-6="27.[113,148,1297,1319]" captionTargetBox-0="[113,1451,189,1112]" captionTargetBox-1="[134,1474,192,1681]" captionTargetBox-2="[114,1450,191,1726]" captionTargetBox-3="[139,1473,192,1116]" captionTargetBox-4="[114,1451,192,1270]" captionTargetBox-5="[235,1375,189,2048]" captionTargetBox-6="[116,1452,191,1284]" captionTargetId-0="figure-10@21.[113,1451,189,1112]" captionTargetId-1="figure-8@22.[122,1477,186,1700]" captionTargetId-2="figure-11@23.[106,1469,167,1737]" captionTargetId-3="figure-429@24.[138,1490,192,1125]" captionTargetId-4="figure-11@25.[113,1451,164,1281]" captionTargetId-5="figure-7@26.[235,1375,189,2048]" captionTargetId-6="figure-11@27.[95,1465,178,1299]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="24" captionTargetPageId-4="25" captionTargetPageId-5="26" captionTargetPageId-6="27" captionText-0="Fig. 15. Tmethypocoelis liki, new species. A, paratype, MZB.Cru.5012, male, 5.3 × 3.1 mm, in situ with life colouration. B, holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm, dorsal habitus, preserved colouration; Papua, Liki Island. C–D, habitat of Tmethypocoelis liki, new species (photograph C by Sumaryanto Bronto)." captionText-1="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." captionText-2="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." captionText-3="Fig. 18. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Left pereopods: A–D, anterior surfaces; E–H, posterior surfaces; A, E, P2; B, F, P3; C, G, P4; D, H, P5." captionText-4="Fig. 19. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Left pereopods: A–D, upper margin; E–H, lower margin; A, E, P2; B, F, P3; C, G, P4; D, H, P5." captionText-5="Fig. 20. Tmethypocoelis liki, new species. Paratypes: A–C, MZB.Cru.5012, male, 5.0 × 3.0 mm; D–E, MZB.Cru.5012, male, 5.1 × 3.2 mm; Papua, Liki Island. Left G1: A, D, whole morphology; apical portion: B, E, lateral; C, mesial." captionText-6="Fig. 21. Successive motion of waving display by male Tmethypocoelis liki, new species. Refer to text for details." figureDoi-0="http://doi.org/10.5281/zenodo.7399939" figureDoi-1="http://doi.org/10.5281/zenodo.7399941" figureDoi-2="http://doi.org/10.5281/zenodo.7399943" figureDoi-3="http://doi.org/10.5281/zenodo.7399945" figureDoi-4="http://doi.org/10.5281/zenodo.7399947" figureDoi-5="http://doi.org/10.5281/zenodo.7399949" figureDoi-6="http://doi.org/10.5281/zenodo.7399951" httpUri-0="https://zenodo.org/record/7399939/files/figure.png" httpUri-1="https://zenodo.org/record/7399941/files/figure.png" httpUri-2="https://zenodo.org/record/7399943/files/figure.png" httpUri-3="https://zenodo.org/record/7399945/files/figure.png" httpUri-4="https://zenodo.org/record/7399947/files/figure.png" httpUri-5="https://zenodo.org/record/7399949/files/figure.png" httpUri-6="https://zenodo.org/record/7399951/files/figure.png" pageId="20" pageNumber="481">Figs. 15–21</figureCitation>
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)
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<emphasis id="B96DEAC24E59FF9360A5240E4D28F88B" bold="true" box="[136,358,1811,1835]" pageId="20" pageNumber="481">Material examined.</emphasis>
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<typeStatus id="54A288724E59FF93614124094D9DF88C" box="[364,467,1812,1836]" pageId="20" pageNumber="481" type="holotype">Holotype</typeStatus>
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: male (6.2 ×
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. Cru.5011),
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<collectingCounty id="62C74E5C4E59FF93612C24294DCBF8EC" box="[257,389,1844,1868]" pageId="20" pageNumber="481">Liki Village</collectingCounty>
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,
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<collectingMunicipality id="6BC2ACAA4E59FF9361A224294E69F8EC" box="[399,551,1844,1868]" pageId="20" pageNumber="481">Sarmi District</collectingMunicipality>
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<location id="8EC6600B4E59FF93621C24294F4DF8EC" LSID="urn:lsid:plazi:treatment:03B087C64E59FF9B613C25AE4D3DFCD5:8EC6600B4E59FF93621C24294F4DF8EC" box="[561,771,1844,1868]" county="Liki Village" latitude="-1.6236917" longLatPrecision="1" longitude="138.7407" municipality="Sarmi District" name="Sarmi Municipality" pageId="20" pageNumber="481" stateProvince="Papua Barat">Sarmi Municipality</location>
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,
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<location id="8EC6600B4E59FF9360A524494CB2F8CC" LSID="urn:lsid:plazi:treatment:03B087C64E59FF9B613C25AE4D3DFCD5:8EC6600B4E59FF9360A524494CB2F8CC" box="[136,252,1876,1900]" county="Liki Village" latitude="-1.6236917" longLatPrecision="1" longitude="138.7407" municipality="Sarmi District" name="Liki Island" pageId="20" pageNumber="481" stateProvince="Papua Barat">Liki Island</location>
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,
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<collectingRegion id="49DDF8324E59FF93612B24494DE3F8CC" box="[262,429,1876,1900]" country="Indonesia" name="Papua Barat" pageId="20" pageNumber="481">Papua Province</collectingRegion>
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,
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<date id="FFA710104E59FF9360A524694D6FF82C" box="[136,289,1908,1932]" pageId="20" pageNumber="481" value="2018-11-21">
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<collectingDate id="EFE3E9F84E59FF9360A524694D6FF82C" box="[136,289,1908,1932]" pageId="20" pageNumber="481" value="2018-11-21">21 Nov. 2018</collectingDate>
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<collectorName id="26EC53064E59FF93610024694D8EF82C" box="[301,448,1908,1932]" pageId="20" pageNumber="481">DC Murniati.</collectorName>
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<typeStatus id="54A288724E59FF9360A524894CB6F80C" box="[136,248,1940,1964]" pageId="20" pageNumber="481" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D1FFD594E59FF93612B24894D22F80C" box="[262,364,1940,1964]" pageId="20" pageNumber="481" type="male">15 males</specimenCount>
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(5.3 ×
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<quantity id="4CE19B354E59FF9361EA24894E50F80B" box="[455,542,1940,1964]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="20" pageNumber="481" unit="mm" value="3.1">3.1 mm</quantity>
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– 5.8 ×
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<quantity id="4CE19B354E59FF9362A524894EADF80C" box="[648,739,1940,1964]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="20" pageNumber="481" unit="mm" value="3.2">3.2 mm</quantity>
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),
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<materialsCitation id="3B713C8D4E59FF9362D724894DEEF78C" ID-GBIF-Occurrence="4010507313" collectingDate="2018-11-21" collectionCode="MZB" collectorName="DC Murniati" county="Cru." latitude="-1.6236917" location="Sarmi District" longLatPrecision="1" longitude="138.7407" municipality="Liki Village" pageId="20" pageNumber="481" specimenCount="3" specimenCount-female="3" stateProvince="Papua Barat" typeStatus="paratype">
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<specimenCount id="9D1FFD594E59FF9362D724894C95F86C" pageId="20" pageNumber="481" type="female">3 females</specimenCount>
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(3.5 ×
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– 3.9 ×
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<quantity id="4CE19B354E59FF9361C824A94E74F86C" box="[485,570,1972,1996]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="481" unit="mm" value="2.5">2.5 mm</quantity>
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<collectionCode id="ED08AE154E59FF93627F24A94EDEF86C" box="[594,656,1972,1996]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="20" pageNumber="481" type="Museum">MZB</collectionCode>
|
||
.
|
||
<collectingCounty id="62C74E5C4E59FF9362BE24A94E8CF86C" box="[659,706,1972,1996]" pageId="20" pageNumber="481">Cru.</collectingCounty>
|
||
5012),
|
||
<collectingMunicipality id="6BC2ACAA4E59FF9360A524C94D54F84C" box="[136,282,2004,2028]" pageId="20" pageNumber="481">Liki Village</collectingMunicipality>
|
||
,
|
||
<location id="8EC6600B4E59FF93610724C94D9CF84C" LSID="urn:lsid:plazi:treatment:03B087C64E59FF9B613C25AE4D3DFCD5:8EC6600B4E59FF93610724C94D9CF84C" box="[298,466,2004,2028]" county="Cru." latitude="-1.6236917" longLatPrecision="1" longitude="138.7407" municipality="Liki Village" name="Sarmi District" pageId="20" pageNumber="481" stateProvince="Papua Barat">Sarmi District</location>
|
||
,
|
||
<location id="8EC6600B4E59FF9361CF24C94E86F84C" LSID="urn:lsid:plazi:treatment:03B087C64E59FF9B613C25AE4D3DFCD5:8EC6600B4E59FF9361CF24C94E86F84C" box="[482,712,2004,2028]" county="Cru." latitude="-1.6236917" longLatPrecision="1" longitude="138.7407" municipality="Liki Village" name="Sarmi Municipality" pageId="20" pageNumber="481" stateProvince="Papua Barat">Sarmi Municipality</location>
|
||
,
|
||
<location id="8EC6600B4E59FF9362F524C94C83F7AC" LSID="urn:lsid:plazi:treatment:03B087C64E59FF9B613C25AE4D3DFCD5:8EC6600B4E59FF9362F524C94C83F7AC" county="Cru." latitude="-1.6236917" longLatPrecision="1" longitude="138.7407" municipality="Liki Village" name="Liki Island" pageId="20" pageNumber="481" stateProvince="Papua Barat">Liki Island</location>
|
||
,
|
||
<collectingRegion id="49DDF8324E59FF9360F424E94DCDF7AC" box="[217,387,2036,2060]" country="Indonesia" name="Papua Barat" pageId="20" pageNumber="481">Papua Province</collectingRegion>
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||
,
|
||
<geoCoordinate id="EE2D50174E59FF9361A224E94E64F7AB" box="[399,554,2034,2060]" degrees="01" direction="south" minutes="37" orientation="latitude" pageId="20" pageNumber="481" precision="1" seconds="25.29" value="-1.6236917">01º37′25.29″S</geoCoordinate>
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,
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<date id="FFA710104E59FF9362C124E94CB1F78C" pageId="20" pageNumber="481" value="2018-11-21">
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<collectingDate id="EFE3E9F84E59FF9362C124E94CB1F78C" pageId="20" pageNumber="481" value="2018-11-21">21 Nov. 2018</collectingDate>
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</date>
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,
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<collectorName id="26EC53064E59FF9361232B094DEEF78C" box="[270,416,2068,2092]" pageId="20" pageNumber="481">DC Murniati</collectorName>
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</materialsCitation>
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;
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<materialsCitation id="3B713C8D4E59FF9361832B09496FFEB5" ID-GBIF-Occurrence="4010507311" collectingDate="2018-11-21" collectionCode="ZRC" collectorName="DC Murniati" county="Liki Village" latitude="-1.6236917" location="Sarmi Municipality" longLatPrecision="1" longitude="138.7407" municipality="Sarmi District" pageId="20" pageNumber="481" specimenCode="ZRC 2022.0914" specimenCount="5" specimenCount-male="5" stateProvince="Papua Barat" typeStatus="paratype">
|
||
<specimenCount id="9D1FFD594E59FF9361832B094E4AF78C" box="[430,516,2068,2092]" pageId="20" pageNumber="481" type="male">5 males</specimenCount>
|
||
(4.0 × 2.4 – 4.8 ×
|
||
<quantity id="4CE19B354E59FF9362C82B094F20FF75" box="[741,878,189,2092]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="20" pageNumber="481" unit="mm" value="2.8">2.8 mm</quantity>
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||
) (
|
||
<specimenCode id="DBBF9EAB4E59FF9363A823A04876FF75" box="[901,1080,189,213]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="481">ZRC 2022.0914</specimenCode>
|
||
),
|
||
<collectingCounty id="62C74E5C4E59FF93646623A0489EFF75" box="[1099,1232,189,213]" pageId="20" pageNumber="481">Liki Village</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BC2ACAA4E59FF9364F123A04939FF75" box="[1244,1399,189,213]" pageId="20" pageNumber="481">Sarmi District</collectingMunicipality>
|
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,
|
||
<location id="8EC6600B4E59FF9365AF23A04F9EFF55" LSID="urn:lsid:plazi:treatment:03B087C64E59FF9B613C25AE4D3DFCD5:8EC6600B4E59FF9365AF23A04F9EFF55" county="Liki Village" latitude="-1.6236917" longLatPrecision="1" longitude="138.7407" municipality="Sarmi District" name="Sarmi Municipality" pageId="20" pageNumber="481" stateProvince="Papua Barat">Sarmi Municipality</location>
|
||
,
|
||
<location id="8EC6600B4E59FF9363F323C04817FF55" LSID="urn:lsid:plazi:treatment:03B087C64E59FF9B613C25AE4D3DFCD5:8EC6600B4E59FF9363F323C04817FF55" box="[990,1113,221,245]" county="Liki Village" latitude="-1.6236917" longLatPrecision="1" longitude="138.7407" municipality="Sarmi District" name="Liki Island" pageId="20" pageNumber="481" stateProvince="Papua Barat">Liki Island</location>
|
||
,
|
||
<collectingRegion id="49DDF8324E59FF93644A23C3495AFF55" box="[1127,1300,221,245]" country="Indonesia" name="Papua Barat" pageId="20" pageNumber="481">Papua Province</collectingRegion>
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||
,
|
||
<geoCoordinate id="EE2D50174E59FF93650F23C149F0FF55" box="[1314,1470,220,245]" degrees="01" direction="south" minutes="37" orientation="latitude" pageId="20" pageNumber="481" precision="1" seconds="25.29" value="-1.6236917">01º37′25.29″S</geoCoordinate>
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,
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<geoCoordinate id="EE2D50174E59FF93636E23E14FA0FEB5" box="[835,1006,252,277]" degrees="138" direction="east" minutes="44" orientation="longitude" pageId="20" pageNumber="481" precision="1" seconds="26.54" value="138.7407">138º44′26.54″E</geoCoordinate>
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,
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<date id="FFA710104E59FF9363D523E048C5FEB5" box="[1016,1163,253,277]" pageId="20" pageNumber="481" value="2018-11-21">
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||
<collectingDate id="EFE3E9F84E59FF9363D523E048C5FEB5" box="[1016,1163,253,277]" pageId="20" pageNumber="481" value="2018-11-21">21 Nov. 2018</collectingDate>
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</date>
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||
,
|
||
<collectorName id="26EC53064E59FF9364B823E0496FFEB5" box="[1173,1313,253,277]" pageId="20" pageNumber="481">DC Murniati</collectorName>
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</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B713C8D4E59FF93650623E049F0FED5" ID-GBIF-Occurrence="4010507320" collectingDate="2018-11-21" collectionCode="OMNH" collectorName="DC Murniati" county="Liki Village" latitude="-1.6236917" location="Sarmi Municipality" longLatPrecision="1" longitude="138.7407" municipality="Sarmi District" pageId="20" pageNumber="481" specimenCount="5" specimenCount-male="5" stateProvince="Papua Barat" typeStatus="paratype">
|
||
<specimenCount id="9D1FFD594E59FF93650623E04933FEB5" box="[1323,1405,253,277]" pageId="20" pageNumber="481" type="male">5 males</specimenCount>
|
||
(3.7 × 2.4 – 4.9 ×
|
||
<quantity id="4CE19B354E59FF93639422004845FE95" box="[953,1035,285,309]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="20" pageNumber="481" unit="mm" value="2.8">2.8 mm</quantity>
|
||
) (
|
||
<collectionCode id="ED08AE154E59FF9364322200483CFE95" box="[1055,1138,285,309]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="20" pageNumber="481" type="Museum">OMNH</collectionCode>
|
||
Ar-12646),
|
||
<collectingCounty id="62C74E5C4E59FF9364D822004936FE95" box="[1269,1400,285,309]" pageId="20" pageNumber="481">Liki Village</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BC2ACAA4E59FF9365AF22004FD8FEF5" pageId="20" pageNumber="481">Sarmi District</collectingMunicipality>
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,
|
||
<location id="8EC6600B4E59FF93638922204835FEF5" LSID="urn:lsid:plazi:treatment:03B087C64E59FF9B613C25AE4D3DFCD5:8EC6600B4E59FF93638922204835FEF5" box="[932,1147,317,341]" county="Liki Village" latitude="-1.6236917" longLatPrecision="1" longitude="138.7407" municipality="Sarmi District" name="Sarmi Municipality" pageId="20" pageNumber="481" stateProvince="Papua Barat">Sarmi Municipality</location>
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,
|
||
<location id="8EC6600B4E59FF9364A42220494DFEF5" LSID="urn:lsid:plazi:treatment:03B087C64E59FF9B613C25AE4D3DFCD5:8EC6600B4E59FF9364A42220494DFEF5" box="[1161,1283,317,341]" county="Liki Village" latitude="-1.6236917" longLatPrecision="1" longitude="138.7407" municipality="Sarmi District" name="Liki Island" pageId="20" pageNumber="481" stateProvince="Papua Barat">Liki Island</location>
|
||
,
|
||
<collectingRegion id="49DDF8324E59FF93653C222349F1FEF5" box="[1297,1471,317,341]" country="Indonesia" name="Papua Barat" pageId="20" pageNumber="481">Papua Province</collectingRegion>
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||
,
|
||
<geoCoordinate id="EE2D50174E59FF93636E22414F90FED5" box="[835,990,348,373]" degrees="01" direction="south" minutes="37" orientation="latitude" pageId="20" pageNumber="481" precision="1" seconds="25.29" value="-1.6236917">01º37′25.29″S</geoCoordinate>
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,
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<geoCoordinate id="EE2D50174E59FF9363CA224148DFFED5" box="[999,1169,348,373]" degrees="138" direction="east" minutes="44" orientation="longitude" pageId="20" pageNumber="481" precision="1" seconds="26.54" value="138.7407">138º44′26.54″E</geoCoordinate>
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,
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<date id="FFA710104E59FF9364B722404965FED5" box="[1178,1323,349,373]" pageId="20" pageNumber="481" value="2018-11-21">
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||
<collectingDate id="EFE3E9F84E59FF9364B722404965FED5" box="[1178,1323,349,373]" pageId="20" pageNumber="481" value="2018-11-21">21 Nov. 2018</collectingDate>
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</date>
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||
,
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||
<collectorName id="26EC53064E59FF936519224049F0FED5" box="[1332,1470,349,373]" pageId="20" pageNumber="481">DC Murniati</collectorName>
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</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B713C8D4E59FF93636E22604859FE55" ID-GBIF-Occurrence="4010507319" collectingDate="2018-11-21" collectionCode="RMNH, CRUS" collectorName="DC Murniati" county="Liki Village" latitude="-1.6236917" location="Sarmi Municipality" longLatPrecision="1" longitude="138.7407" municipality="Sarmi District" pageId="20" pageNumber="481" specimenCount="5" specimenCount-male="5" stateProvince="Papua Barat" typeStatus="paratype">
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||
<specimenCount id="9D1FFD594E59FF93636E22604FDAFE35" box="[835,916,381,405]" pageId="20" pageNumber="481" type="male">5 males</specimenCount>
|
||
(5.1 × 2.9 – 5.3 × 3.0 mm) (
|
||
<collectionCode id="ED08AE154E59FF93649922604944FE35" box="[1204,1290,381,405]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="481" type="Museum">RMNH</collectionCode>
|
||
.
|
||
<collectionCode id="ED08AE154E59FF93652022604918FE35" box="[1293,1366,381,405]" pageId="20" pageNumber="481">CRUS</collectionCode>
|
||
.D.58042),
|
||
<collectingCounty id="62C74E5C4E59FF93636E22804F88FE15" box="[835,966,413,437]" pageId="20" pageNumber="481">Liki Village</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BC2ACAA4E59FF9363E22280482BFE15" box="[975,1125,413,437]" pageId="20" pageNumber="481">Sarmi District</collectingMunicipality>
|
||
,
|
||
<location id="8EC6600B4E59FF9364422280490EFE15" LSID="urn:lsid:plazi:treatment:03B087C64E59FF9B613C25AE4D3DFCD5:8EC6600B4E59FF9364422280490EFE15" box="[1135,1344,413,437]" county="Liki Village" latitude="-1.6236917" longLatPrecision="1" longitude="138.7407" municipality="Sarmi District" name="Sarmi Municipality" pageId="20" pageNumber="481" stateProvince="Papua Barat">Sarmi Municipality</location>
|
||
,
|
||
<location id="8EC6600B4E59FF936564228049F0FE15" LSID="urn:lsid:plazi:treatment:03B087C64E59FF9B613C25AE4D3DFCD5:8EC6600B4E59FF936564228049F0FE15" box="[1353,1470,413,437]" county="Liki Village" latitude="-1.6236917" longLatPrecision="1" longitude="138.7407" municipality="Sarmi District" name="Liki Island" pageId="20" pageNumber="481" stateProvince="Papua Barat">Liki Island</location>
|
||
,
|
||
<collectingRegion id="49DDF8324E59FF93636E22A34FBCFE75" box="[835,1010,445,469]" country="Indonesia" name="Papua Barat" pageId="20" pageNumber="481">Papua Province</collectingRegion>
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||
,
|
||
<geoCoordinate id="EE2D50174E59FF93642D22A148D0FE75" box="[1024,1182,444,469]" degrees="01" direction="south" minutes="37" orientation="latitude" pageId="20" pageNumber="481" precision="1" seconds="25.29" value="-1.6236917">01º37′25.29″S</geoCoordinate>
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,
|
||
<geoCoordinate id="EE2D50174E59FF93648022A14914FE75" box="[1197,1370,444,469]" degrees="138" direction="east" minutes="44" orientation="longitude" pageId="20" pageNumber="481" precision="1" seconds="26.54" value="138.7407">138º44′26.54″E</geoCoordinate>
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,
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||
<date id="FFA710104E59FF93654522A04F35FE55" pageId="20" pageNumber="481" value="2018-11-21">
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<collectingDate id="EFE3E9F84E59FF93654522A04F35FE55" pageId="20" pageNumber="481" value="2018-11-21">21 Nov. 2018</collectingDate>
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</date>
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,
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||
<collectorName id="26EC53064E59FF9363A522C04859FE55" box="[904,1047,477,501]" pageId="20" pageNumber="481">DC Murniati</collectorName>
|
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</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C303655B4E59FF93636E210049D2FC55" pageId="20" pageNumber="481" type="diagnosis">
|
||
<paragraph id="8BA636D04E59FF93636E210049D2FC55" blockId="20.[835,1475,541,1013]" pageId="20" pageNumber="481">
|
||
<emphasis id="B96DEAC24E59FF93636E21004FFFFD95" bold="true" box="[835,945,541,565]" pageId="20" pageNumber="481">Diagnosis</emphasis>
|
||
. Carapace pentagonal, broad, 1.7 times as broad as long. Front gently narrowed anteriorly; lateral border of carapace distinctly curved, with single row of tubercles and stiff short setae. External orbital angle triangular, posteriorly followed by broad U-shaped sinus. Chelipeds stout, long, subequal. Palm bulky, inner surface coarsely granular on upper half, outer surface distinctly granular over distal-upper half. Fingers not gaping at base; fixed finger short, triangular. Dactylus with outer surface bearing one median ridge of granules; cutting margin with blunt and wide serration; upper margin with median row of granules culminating in one prominent tooth. Male pleon 1.8 times as long as wide. G1 long, slender, curved; apex forming two lobes; apical setae consisting one long and five short setae. Vulva D-shaped, mesial margin of operculum with triangular projection.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C303655B4E59FF9E63682700485CF787" lastPageId="25" lastPageNumber="486" pageId="20" pageNumber="481" type="description">
|
||
<paragraph id="8BA636D04E59FF92636827004EE2FAE6" blockId="20.[835,1475,1053,2101]" lastBlockId="21.[113,752,1262,1350]" lastPageId="21" lastPageNumber="482" pageId="20" pageNumber="481">
|
||
<emphasis id="B96DEAC24E59FF93636827004FABFB95" bold="true" box="[837,997,1053,1077]" pageId="20" pageNumber="481">Description.</emphasis>
|
||
Carapace pentagonal, broad; convex longitudinally, slightly convex transversely; 1.7 times as broad as long, greatest width across external orbital angles (
|
||
<figureCitation id="13222A554E59FF93636627604FFFFB35" box="[843,945,1149,1173]" captionStart="Fig" captionStartId="21.[113,148,1127,1149]" captionTargetBox="[113,1451,189,1112]" captionTargetId="figure-10@21.[113,1451,189,1112]" captionTargetPageId="21" captionText="Fig. 15. Tmethypocoelis liki, new species. A, paratype, MZB.Cru.5012, male, 5.3 × 3.1 mm, in situ with life colouration. B, holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm, dorsal habitus, preserved colouration; Papua, Liki Island. C–D, habitat of Tmethypocoelis liki, new species (photograph C by Sumaryanto Bronto)." figureDoi="http://doi.org/10.5281/zenodo.7399939" httpUri="https://zenodo.org/record/7399939/files/figure.png" pageId="20" pageNumber="481">Fig. 15B</figureCitation>
|
||
,
|
||
<figureCitation id="13222A554E59FF9363ED27604FBDFB35" box="[960,1011,1149,1173]" captionStart="Fig" captionStartId="22.[136,171,1700,1722]" captionTargetBox="[134,1474,192,1681]" captionTargetId="figure-8@22.[122,1477,186,1700]" captionTargetPageId="22" captionText="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." figureDoi="http://doi.org/10.5281/zenodo.7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="20" pageNumber="481">16A</figureCitation>
|
||
). Dorsal surface almost smooth, sparsely tubercular; regions defined by shallow grooves, epigastric lobe just visible (
|
||
<figureCitation id="13222A554E59FF93643827A048CAFB75" box="[1045,1156,1213,1237]" captionStart="Fig" captionStartId="22.[136,171,1700,1722]" captionTargetBox="[134,1474,192,1681]" captionTargetId="figure-8@22.[122,1477,186,1700]" captionTargetPageId="22" captionText="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." figureDoi="http://doi.org/10.5281/zenodo.7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="20" pageNumber="481">Fig. 16A</figureCitation>
|
||
). Cervical groove dividing gastric and cardiac regions, well-marked; cardiac regions with slight central depression. Branchial region sloping, with small, well-spaced, low tubercles, each bearing single seta. Sub-branchial region bulging, bearing regular setae, separated from branchial region by sinuous lateral border fringed with short setae. Front gently narrowed anteriorly, basal and distal widths 0.22 and 0.12 times as wide as external orbital width, respectively; distolateral angles rounded (
|
||
<figureCitation id="13222A554E59FF93638426C04842FA55" box="[937,1036,1501,1525]" captionStart="Fig" captionStartId="22.[136,171,1700,1722]" captionTargetBox="[134,1474,192,1681]" captionTargetId="figure-8@22.[122,1477,186,1700]" captionTargetPageId="22" captionText="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." figureDoi="http://doi.org/10.5281/zenodo.7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="20" pageNumber="481">Fig. 16A</figureCitation>
|
||
); anterior margin with small central blunt prominence. External orbital angle triangular, anterior and lateral margins slightly convex, with microscopic tubercles, one small ridge extending from mid-lateral margin, parallel to anterior margin, as long as anterior margin, posteriorly followed by broad U-shaped sinus (
|
||
<figureCitation id="13222A554E59FF9364FE25604912F935" box="[1235,1372,1661,1685]" captionStart="Fig" captionStartId="22.[136,171,1700,1722]" captionTargetBox="[134,1474,192,1681]" captionTargetId="figure-8@22.[122,1477,186,1700]" captionTargetPageId="22" captionText="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." figureDoi="http://doi.org/10.5281/zenodo.7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="20" pageNumber="481">Fig. 16A, B</figureCitation>
|
||
). Second anterolateral tooth posterior to U-shaped sinus, concave posteriorly, distance between second anterolateral teeth less than between external orbital angles. Lateral margin of carapace distinctly curved, with single row of tubercles and stiff short setae (
|
||
<figureCitation id="13222A554E59FF93642C24004826F895" box="[1025,1128,1821,1845]" captionStart="Fig" captionStartId="22.[136,171,1700,1722]" captionTargetBox="[134,1474,192,1681]" captionTargetId="figure-8@22.[122,1477,186,1700]" captionTargetPageId="22" captionText="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." figureDoi="http://doi.org/10.5281/zenodo.7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="20" pageNumber="481">Fig. 16A</figureCitation>
|
||
). Anterolateral margin convex; posterolateral margin concave, adjoin with posterior margin without sharp angle. Posterior margin slightly concave, c.0.53 distance between external orbital angles; fine ridge parallel with posterior margin forming broad rim (
|
||
<figureCitation id="13222A554E59FF93652824804926F815" box="[1285,1384,1949,1973]" captionStart="Fig" captionStartId="22.[136,171,1700,1722]" captionTargetBox="[134,1474,192,1681]" captionTargetId="figure-8@22.[122,1477,186,1700]" captionTargetPageId="22" captionText="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." figureDoi="http://doi.org/10.5281/zenodo.7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="20" pageNumber="481">Fig. 16A</figureCitation>
|
||
). Supraorbital margin sinuous, diverging backward, microscopically tuberculate (
|
||
<figureCitation id="13222A554E59FF9363E824C0486AF855" box="[965,1060,2013,2037]" captionStart="Fig" captionStartId="22.[136,171,1700,1722]" captionTargetBox="[134,1474,192,1681]" captionTargetId="figure-8@22.[122,1477,186,1700]" captionTargetPageId="22" captionText="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." figureDoi="http://doi.org/10.5281/zenodo.7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="20" pageNumber="481">Fig. 16C</figureCitation>
|
||
). Infraorbital margin with medial notch, below which, on pterygostome, oblique channel running inwardly; inner portion of infraorbital margin consisting of two granular ridges separated by shallow channel; outer portion of infraorbital border granular, concave, ending in broad notch below external orbital angle (
|
||
<figureCitation id="13222A554E58FF92621126334EEEFAE6" box="[572,672,1326,1350]" captionStart="Fig" captionStartId="22.[136,171,1700,1722]" captionTargetBox="[134,1474,192,1681]" captionTargetId="figure-8@22.[122,1477,186,1700]" captionTargetPageId="22" captionText="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." figureDoi="http://doi.org/10.5281/zenodo.7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="21" pageNumber="482">Fig. 16C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF6666584E58FF92605C277A4E0EFB16" ID-DOI="http://doi.org/10.5281/zenodo.7399939" ID-Zenodo-Dep="7399939" httpUri="https://zenodo.org/record/7399939/files/figure.png" pageId="21" pageNumber="482" startId="21.[113,148,1127,1149]" targetBox="[113,1451,189,1112]" targetPageId="21">
|
||
<paragraph id="8BA636D04E58FF92605C277A4E0EFB16" blockId="21.[113,1452,1127,1207]" pageId="21" pageNumber="482">
|
||
Fig. 15.
|
||
<taxonomicName id="4C194D534E58FF9260EA277A4DCBFBDC" authority="Murniati & Asakura & Nugroho & Hernawan & Dharmawan, 2022" authorityName="Murniati & Asakura & Nugroho & Hernawan & Dharmawan" authorityYear="2022" box="[199,389,1127,1148]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="482" phylum="Arthropoda" rank="species" species="liki" status="sp. nov.">
|
||
<emphasis id="B96DEAC24E58FF9260EA277A4DCBFBDC" box="[199,389,1127,1148]" italics="true" pageId="21" pageNumber="482">Tmethypocoelis liki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25E57B94E58FF9261B927754E43FBDD" box="[404,525,1127,1149]" pageId="21" pageNumber="482" rank="species">new species</taxonomicNameLabel>
|
||
. A, paratype, MZB.Cru.5012, male, 5.3 × 3.1 mm, in situ with life colouration. B, holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm, dorsal habitus, preserved colouration; Papua, Liki Island. C–D, habitat of
|
||
<taxonomicName id="4C194D534E58FF92649427994938FB39" authority="Murniati & Asakura & Nugroho & Hernawan & Dharmawan, 2022" authorityName="Murniati & Asakura & Nugroho & Hernawan & Dharmawan" authorityYear="2022" box="[1209,1398,1156,1177]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="482" phylum="Arthropoda" rank="species" species="liki" status="sp. nov.">
|
||
<emphasis id="B96DEAC24E58FF92649427994938FB39" box="[1209,1398,1156,1177]" italics="true" pageId="21" pageNumber="482">Tmethypocoelis liki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25E57B94E58FF9265AE27984CF6FB17" pageId="21" pageNumber="482" rank="species">new species</taxonomicNameLabel>
|
||
(photograph C by Sumaryanto Bronto).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA636D04E58FF92605C26734CE2FA66" blockId="21.[113,753,1390,1478]" pageId="21" pageNumber="482">
|
||
Epistome forming three lobes posteriorly, distinctly convex laterally and medially, margins without tuberculation (
|
||
<figureCitation id="13222A554E58FF9262E826934CEEFA66" captionStart="Fig" captionStartId="22.[136,171,1700,1722]" captionTargetBox="[134,1474,192,1681]" captionTargetId="figure-8@22.[122,1477,186,1700]" captionTargetPageId="22" captionText="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." figureDoi="http://doi.org/10.5281/zenodo.7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="21" pageNumber="482">Fig. 16E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA636D04E58FF92605C26F34E04F926" blockId="21.[113,753,1518,1670]" pageId="21" pageNumber="482">
|
||
Eyestalks long, widening distally, distal end of cornea reaching slightly beyond external orbital angles, distally with short projection, projection 0.3 times as long as cornea, tip with several short setae; cornea bulging; medial thickening gives twisted appearance (
|
||
<figureCitation id="13222A554E58FF9261BC25734DB1F926" box="[401,511,1646,1670]" captionStart="Fig" captionStartId="21.[113,148,1127,1149]" captionTargetBox="[113,1451,189,1112]" captionTargetId="figure-10@21.[113,1451,189,1112]" captionTargetPageId="21" captionText="Fig. 15. Tmethypocoelis liki, new species. A, paratype, MZB.Cru.5012, male, 5.3 × 3.1 mm, in situ with life colouration. B, holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm, dorsal habitus, preserved colouration; Papua, Liki Island. C–D, habitat of Tmethypocoelis liki, new species (photograph C by Sumaryanto Bronto)." figureDoi="http://doi.org/10.5281/zenodo.7399939" httpUri="https://zenodo.org/record/7399939/files/figure.png" pageId="21" pageNumber="482">Figs. 15A</figureCitation>
|
||
,
|
||
<figureCitation id="13222A554E58FF92622125734E70F926" box="[524,574,1646,1670]" captionStart="Fig" captionStartId="22.[136,171,1700,1722]" captionTargetBox="[134,1474,192,1681]" captionTargetId="figure-8@22.[122,1477,186,1700]" captionTargetPageId="22" captionText="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." figureDoi="http://doi.org/10.5281/zenodo.7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="21" pageNumber="482">16D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA636D04E58FF92605C25B349ECFA86" blockId="21.[112,753,1710,2118]" lastBlockId="21.[812,1451,1262,1318]" pageId="21" pageNumber="482">
|
||
Third maxillipeds distinctly vaulted, almost completely covering buccal cavern. Ischium subquadrate, antero-mesial angle narrowly triangular; mesial and lateral margins with dense short setae; outer surface smooth, with one oblique row of setae. Merus trapezoidal, subequal in size to ischium, about 1.1 times as long as ischium laterally; outer surface covered sparsely by regular short setae, without granules; mesial margin slightly convex, with long feathered setae; lateral margin convex, converged distally, covered with short setae; anterior margin slightly concave, without setae. Carpus trihedral, distal portion with dense long setae, lateral margin with sparse short setae, mesial margin with dense long setae. Propodus, shorter than dactylus, lateral margins with sparse short setae. Dactylus slender, two times as long as propodus, margins serrated, with long stiff setae apically (
|
||
<figureCitation id="13222A554E58FF926518261349DBFA86" box="[1333,1429,1294,1318]" captionStart="Fig" captionStartId="22.[136,171,1700,1722]" captionTargetBox="[134,1474,192,1681]" captionTargetId="figure-8@22.[122,1477,186,1700]" captionTargetPageId="22" captionText="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." figureDoi="http://doi.org/10.5281/zenodo.7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="21" pageNumber="482">Fig. 16F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA636D04E58FF9163002653486EF7E3" blockId="21.[812,1452,1358,2118]" lastBlockId="22.[834,1474,1835,2115]" lastPageId="22" lastPageNumber="483" pageId="21" pageNumber="482">
|
||
Male chelipeds (P1) stout, long, subequal (
|
||
<figureCitation id="13222A554E58FF9264DE2653491BFAC6" box="[1267,1365,1358,1382]" captionStart="Fig" captionStartId="21.[113,148,1127,1149]" captionTargetBox="[113,1451,189,1112]" captionTargetId="figure-10@21.[113,1451,189,1112]" captionTargetPageId="21" captionText="Fig. 15. Tmethypocoelis liki, new species. A, paratype, MZB.Cru.5012, male, 5.3 × 3.1 mm, in situ with life colouration. B, holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm, dorsal habitus, preserved colouration; Papua, Liki Island. C–D, habitat of Tmethypocoelis liki, new species (photograph C by Sumaryanto Bronto)." figureDoi="http://doi.org/10.5281/zenodo.7399939" httpUri="https://zenodo.org/record/7399939/files/figure.png" pageId="21" pageNumber="482">Fig. 15B</figureCitation>
|
||
). Merus with cross-section triangular (
|
||
<figureCitation id="13222A554E58FF9264A52673495CFA26" box="[1160,1298,1390,1414]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="21" pageNumber="482">Fig. 17A–F</figureCitation>
|
||
); higher than external orbital angle when held against external orbital angle; outer margin with two irregular rows of sharp tubercles (
|
||
<figureCitation id="13222A554E58FF9265AD26B34F10FA46" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="21" pageNumber="482">Fig. 17A</figureCitation>
|
||
); lower margin with one row of tubercles (
|
||
<figureCitation id="13222A554E58FF92651726D349D0FA46" box="[1338,1438,1486,1510]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="21" pageNumber="482">Fig. 17B</figureCitation>
|
||
); upper margin covered by scattered granules, granulation on distal portion continuous to outer surface, distal margin with one row of tubercles (
|
||
<figureCitation id="13222A554E58FF92643525334835F9E6" box="[1048,1147,1582,1606]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="21" pageNumber="482">Fig. 17C</figureCitation>
|
||
); oval tympani on outer and upper surfaces (
|
||
<figureCitation id="13222A554E58FF9263CD2553482FF9C6" box="[992,1121,1614,1638]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="21" pageNumber="482">Fig. 17E–F</figureCitation>
|
||
), tympanum on outer surface smaller than that of upper surface, microscopic granules present outside tympanum (
|
||
<figureCitation id="13222A554E58FF92647C259348E0F906" box="[1105,1198,1678,1702]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="21" pageNumber="482">Fig. 17F</figureCitation>
|
||
); lower surface smooth, without tympanum (
|
||
<figureCitation id="13222A554E58FF92642725B3483EF966" box="[1034,1136,1710,1734]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="21" pageNumber="482">Fig. 17D</figureCitation>
|
||
). Carpus shorter than merus; elongate, 2.15 times as long as wide; all margins tubercular (
|
||
<figureCitation id="13222A554E58FF92631825F34FE0F8A6" box="[821,942,1774,1798]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="21" pageNumber="482">Fig. 17J–L</figureCitation>
|
||
); upper surface flattened, with triangular shape (
|
||
<figureCitation id="13222A554E58FF92631824134FD7F886" box="[821,921,1806,1830]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="21" pageNumber="482">Fig. 17G</figureCitation>
|
||
); lower surface smooth (
|
||
<figureCitation id="13222A554E58FF92648B24134945F886" box="[1190,1291,1806,1830]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="21" pageNumber="482">Fig. 17H</figureCitation>
|
||
); outer surface peak with microscopic granules (17I). Palm bulky, 1.4–1.5 times as long as high (excluding fixed finger); inner surface coarsely granular on upper half, granulation extending to sharply cut upper margin (
|
||
<figureCitation id="13222A554E58FF92646C249348E6F806" box="[1089,1192,1934,1958]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="21" pageNumber="482">Fig. 17M</figureCitation>
|
||
); outer surface distinctly granular over distal-upper half, distinct groove extending below granular rows forming clear granular string near upper margin, half-lower part smooth (
|
||
<figureCitation id="13222A554E58FF9264C624F3491BF7A6" box="[1259,1365,2030,2054]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="21" pageNumber="482">Fig. 17N</figureCitation>
|
||
); upper margin with granules (
|
||
<figureCitation id="13222A554E58FF92640A2B1348C0F786" box="[1063,1166,2062,2086]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="21" pageNumber="482">Fig. 17O</figureCitation>
|
||
); lower margin tubercular (
|
||
<figureCitation id="13222A554E58FF9263182B334FD3F7E6" box="[821,925,2094,2118]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="21" pageNumber="482">Fig. 17P</figureCitation>
|
||
). Fingers without gaping proximally; curved inwards, expanded distally to form spooned tip; cutting margins evenly serrated; inner margins at tip of both fingers with short row of stout setae (
|
||
<figureCitation id="13222A554E5BFF9161F624764E25F823" box="[475,619,1899,1923]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="22" pageNumber="483">Fig. 17M, N</figureCitation>
|
||
). Fixed finger short, triangular shape, 0.34 times as long as palm; inner surface with granules near cutting margin; outer surface with granules near cutting margin; cutting margin slightly convex over entire length, but without one large differentiated tooth or lobe; lower margin granular only along middle to base (
|
||
<figureCitation id="13222A554E5BFF9160BD2B364CA1F7E3" box="[144,239,2091,2115]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="22" pageNumber="483">Fig. 17P</figureCitation>
|
||
). Dactylus with outer surface bearing one median ridge of granules extended nearly whole length, scattered granules between ridge and cutting margin, flat depression with microscopic granules between granular ridge and upper margin, dense granulation on proximal portion near upper margin (
|
||
<figureCitation id="13222A554E5BFF9163B624B64FB5F863" box="[923,1019,1963,1987]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="22" pageNumber="483">Fig. 17N</figureCitation>
|
||
); inner surface granular near cutting margin and near upper margin, median portion smooth (
|
||
<figureCitation id="13222A554E5BFF91656024D649FBF843" box="[1357,1461,1995,2019]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="22" pageNumber="483">Fig. 17M</figureCitation>
|
||
); cutting margin with blunt and wide serrates; upper margin with median row of granules culminating into one prominent tooth (
|
||
<figureCitation id="13222A554E5BFF9163A42B36485AF7E3" box="[905,1044,2091,2115]" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="22" pageNumber="483">Fig. 17M–O</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF6666584E5BFF9160A525B94816F954" ID-DOI="http://doi.org/10.5281/zenodo.7399941" ID-Zenodo-Dep="7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="22" pageNumber="483" startId="22.[136,171,1700,1722]" targetBox="[134,1474,192,1681]" targetPageId="22">
|
||
<paragraph id="8BA636D04E5BFF9160A525B94816F954" blockId="22.[136,1475,1700,1780]" pageId="22" pageNumber="483">
|
||
Fig. 16.
|
||
<taxonomicName id="4C194D534E5BFF9160F025B94DD2F919" authority="Murniati & Asakura & Nugroho & Hernawan & Dharmawan, 2022" authorityName="Murniati & Asakura & Nugroho & Hernawan & Dharmawan" authorityYear="2022" box="[221,412,1700,1721]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="483" phylum="Arthropoda" rank="species" species="liki" status="sp. nov.">
|
||
<emphasis id="B96DEAC24E5BFF9160F025B94DD2F919" box="[221,412,1700,1721]" italics="true" pageId="22" pageNumber="483">Tmethypocoelis liki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25E57B94E5BFF91618725B84E6DF91A" box="[426,547,1700,1722]" pageId="22" pageNumber="483" rank="species">new species</taxonomicNameLabel>
|
||
. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6666584E5AFF90605C25CC4E54F89D" ID-DOI="http://doi.org/10.5281/zenodo.7399943" ID-Zenodo-Dep="7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="23" pageNumber="484" startId="23.[113,148,1745,1767]" targetBox="[114,1450,191,1726]" targetPageId="23">
|
||
<paragraph id="8BA636D04E5AFF90605C25CC4E54F89D" blockId="23.[113,1453,1744,1853]" pageId="23" pageNumber="484">
|
||
Fig. 17.
|
||
<taxonomicName id="4C194D534E5AFF9060E925CC4DCFF946" authority="Murniati & Asakura & Nugroho & Hernawan & Dharmawan, 2022" authorityName="Murniati & Asakura & Nugroho & Hernawan & Dharmawan" authorityYear="2022" box="[196,385,1745,1766]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="484" phylum="Arthropoda" rank="species" species="liki" status="sp. nov.">
|
||
<emphasis id="B96DEAC24E5AFF9060E925CC4DCFF946" box="[196,385,1745,1766]" italics="true" pageId="23" pageNumber="484">Tmethypocoelis liki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25E57B94E5AFF9061A325CC4E4BF947" box="[398,517,1745,1767]" pageId="23" pageNumber="484" rank="species">new species</taxonomicNameLabel>
|
||
. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA636D04E5AFF9F605C24694E70FA8C" blockId="23.[113,752,1908,2092]" lastBlockId="24.[136,775,1235,1324]" lastPageId="24" lastPageNumber="485" pageId="23" pageNumber="484">
|
||
Ambulatory legs (P2–P5) slender, elongate. P2 shorter than P3. Merus 2.9 times as long as wide; tympani on anterior and posterior surfaces; anterior surface without granules; posterior surface with scattered granules near upper margin; upper margin narrowed, convex in cross section, with spaced tubercles, tuberculation not forming ridge, sparse setae distributed irregularly; surface of lower margin wider than that of upper margin, smooth, nearly without setae. Carpus shorter than propodus, surfaces smooth; upper margin with sparse tubercles proximally, sparse setae distributed irregularly; lower margin smooth. Propodus shorter than merus, with anterior surface smooth; posterior surface with oval tympanum, tympanum smaller than that of merus; upper margin smooth, with sparse setae; lower margin smooth, without setation (
|
||
<figureCitation id="13222A554E55FF9F616B260E4D9AFA8B" box="[326,468,1299,1324]" captionStart="Fig" captionStartId="24.[136,171,1129,1151]" captionTargetBox="[139,1473,192,1116]" captionTargetId="figure-429@24.[138,1490,192,1125]" captionTargetPageId="24" captionText="Fig. 18. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Left pereopods: A–D, anterior surfaces; E–H, posterior surfaces; A, E, P2; B, F, P3; C, G, P4; D, H, P5." figureDoi="http://doi.org/10.5281/zenodo.7399945" httpUri="https://zenodo.org/record/7399945/files/figure.png" pageId="24" pageNumber="485">Figs. 18A, E</figureCitation>
|
||
,
|
||
<figureCitation id="13222A554E55FF9F61CC26094E7CFA8C" box="[481,562,1300,1324]" captionStart="Fig" captionStartId="25.[113,148,1285,1307]" captionTargetBox="[114,1451,192,1270]" captionTargetId="figure-11@25.[113,1451,164,1281]" captionTargetPageId="25" captionText="Fig. 19. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Left pereopods: A–D, upper margin; E–H, lower margin; A, E, P2; B, F, P3; C, G, P4; D, H, P5." figureDoi="http://doi.org/10.5281/zenodo.7399947" httpUri="https://zenodo.org/record/7399947/files/figure.png" pageId="24" pageNumber="485">19A, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF6666584E55FF9F60A527744FFCFB3C" ID-DOI="http://doi.org/10.5281/zenodo.7399945" ID-Zenodo-Dep="7399945" httpUri="https://zenodo.org/record/7399945/files/figure.png" pageId="24" pageNumber="485" startId="24.[136,171,1129,1151]" targetBox="[139,1473,192,1116]" targetPageId="24">
|
||
<paragraph id="8BA636D04E55FF9F60A527744FFCFB3C" blockId="24.[135,1473,1129,1180]" pageId="24" pageNumber="485">
|
||
Fig. 18.
|
||
<taxonomicName id="4C194D534E55FF9F60F227744DD1FBDE" authority="Murniati & Asakura & Nugroho & Hernawan & Dharmawan, 2022" authorityName="Murniati & Asakura & Nugroho & Hernawan & Dharmawan" authorityYear="2022" box="[223,415,1129,1150]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="485" phylum="Arthropoda" rank="species" species="liki" status="sp. nov.">
|
||
<emphasis id="B96DEAC24E55FF9F60F227744DD1FBDE" box="[223,415,1129,1150]" italics="true" pageId="24" pageNumber="485">Tmethypocoelis liki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25E57B94E55FF9F618227774E64FBDF" box="[431,554,1129,1151]" pageId="24" pageNumber="485" rank="species">new species</taxonomicNameLabel>
|
||
. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Left pereopods: A–D, anterior surfaces; E–H, posterior surfaces; A, E, P2; B, F, P3; C, G, P4; D, H, P5.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA636D04E55FF9F60A526494DCDF96C" blockId="24.[136,776,1363,1740]" pageId="24" pageNumber="485">
|
||
P3 longest. Merus 3.8 times as long as wide; oval tympani present on both surfaces; anterior surface without granules; posterior surface with sparse granules near tympanum; upper and lower margins convex, narrowed medially by two tympani on two surfaces; upper margin with scattered tubercles and setae; lower margin smooth. Carpus shorter than propodus; anterior and posterior surfaces smooth; surface of upper margin narrow, with sparse setae; surface of lower margin wide, smooth. Propodus shorter than merus; anterior surface smooth; posterior surface with one shallow groove, without granules; upper and lower margins with sparse setae (
|
||
<figureCitation id="13222A554E55FF9F60BD25AE4D55F96B" box="[144,283,1715,1740]" captionStart="Fig" captionStartId="24.[136,171,1129,1151]" captionTargetBox="[139,1473,192,1116]" captionTargetId="figure-429@24.[138,1490,192,1125]" captionTargetPageId="24" captionText="Fig. 18. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Left pereopods: A–D, anterior surfaces; E–H, posterior surfaces; A, E, P2; B, F, P3; C, G, P4; D, H, P5." figureDoi="http://doi.org/10.5281/zenodo.7399945" httpUri="https://zenodo.org/record/7399945/files/figure.png" pageId="24" pageNumber="485">Figs. 18B, F</figureCitation>
|
||
,
|
||
<figureCitation id="13222A554E55FF9F610525A94D38F96C" box="[296,374,1716,1740]" captionStart="Fig" captionStartId="25.[113,148,1285,1307]" captionTargetBox="[114,1451,192,1270]" captionTargetId="figure-11@25.[113,1451,164,1281]" captionTargetPageId="25" captionText="Fig. 19. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Left pereopods: A–D, upper margin; E–H, lower margin; A, E, P2; B, F, P3; C, G, P4; D, H, P5." figureDoi="http://doi.org/10.5281/zenodo.7399947" httpUri="https://zenodo.org/record/7399947/files/figure.png" pageId="24" pageNumber="485">19B, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA636D04E55FF9F60A525E94809FAEC" blockId="24.[136,777,1779,2091]" lastBlockId="24.[835,1474,1235,1356]" pageId="24" pageNumber="485">
|
||
P4 shorter than P2. Merus 3.6 times as long as wide; tympani present on anterior and posterior surfaces, without granules on both surfaces; upper and lower margins convex, narrowed medially by tympani on anterior and posterior surfaces; upper margin with one row of spaced microscopic tubercles, one row of spaced short setae parallel to tuberculation; surface of lower margin smooth, with sparse setae. Carpus shorter than propodus; anterior and posterior surfaces smooth; surface of upper margin narrow, lower margin wide, both margins with sparse setae, without tuberculation. Propodus shorter than merus, anterior and posterior surfaces smooth; posterior surface with one shallow groove; upper and lower margins narrow, without tuberculation, with sparse setae (
|
||
<figureCitation id="13222A554E55FF9F6366262E4F95FAEC" box="[843,987,1331,1356]" captionStart="Fig" captionStartId="24.[136,171,1129,1151]" captionTargetBox="[139,1473,192,1116]" captionTargetId="figure-429@24.[138,1490,192,1125]" captionTargetPageId="24" captionText="Fig. 18. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Left pereopods: A–D, anterior surfaces; E–H, posterior surfaces; A, E, P2; B, F, P3; C, G, P4; D, H, P5." figureDoi="http://doi.org/10.5281/zenodo.7399945" httpUri="https://zenodo.org/record/7399945/files/figure.png" pageId="24" pageNumber="485">Figs. 18C, G</figureCitation>
|
||
,
|
||
<figureCitation id="13222A554E55FF9F63C526294875FAEC" box="[1000,1083,1332,1356]" captionStart="Fig" captionStartId="25.[113,148,1285,1307]" captionTargetBox="[114,1451,192,1270]" captionTargetId="figure-11@25.[113,1451,164,1281]" captionTargetPageId="25" captionText="Fig. 19. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Left pereopods: A–D, upper margin; E–H, lower margin; A, E, P2; B, F, P3; C, G, P4; D, H, P5." figureDoi="http://doi.org/10.5281/zenodo.7399947" httpUri="https://zenodo.org/record/7399947/files/figure.png" pageId="24" pageNumber="485">19C, G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA636D04E55FF9F636E266E498CF90C" blockId="24.[835,1475,1395,1708]" pageId="24" pageNumber="485">
|
||
P5 shortest. Merus 3.2 times as long as wide; tympani present on both surfaces, tympanum on anterior surface larger than that on posterior surface; upper and lower margins without tubercles, with sparse short setae. Carpus shorter than merus; upper and lower margins smooth; surface of upper margin narrow, with sparse short setae; lower margin wide, with sparse short setae. Propodus slightly as long as carpus; anterior and posterior surfaces smooth; surfaces of upper and lower margins narrowed, with sparse short setae. Dactylus with all surfaces and margins smooth (
|
||
<figureCitation id="13222A554E55FF9F64F9258E4912F90B" box="[1236,1372,1683,1708]" captionStart="Fig" captionStartId="24.[136,171,1129,1151]" captionTargetBox="[139,1473,192,1116]" captionTargetId="figure-429@24.[138,1490,192,1125]" captionTargetPageId="24" captionText="Fig. 18. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Left pereopods: A–D, anterior surfaces; E–H, posterior surfaces; A, E, P2; B, F, P3; C, G, P4; D, H, P5." figureDoi="http://doi.org/10.5281/zenodo.7399945" httpUri="https://zenodo.org/record/7399945/files/figure.png" pageId="24" pageNumber="485">Figs. 18D, H</figureCitation>
|
||
,
|
||
<figureCitation id="13222A554E55FF9F654B258949FBF90C" box="[1382,1461,1684,1708]" captionStart="Fig" captionStartId="25.[113,148,1285,1307]" captionTargetBox="[114,1451,192,1270]" captionTargetId="figure-11@25.[113,1451,164,1281]" captionTargetPageId="25" captionText="Fig. 19. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Left pereopods: A–D, upper margin; E–H, lower margin; A, E, P2; B, F, P3; C, G, P4; D, H, P5." figureDoi="http://doi.org/10.5281/zenodo.7399947" httpUri="https://zenodo.org/record/7399947/files/figure.png" pageId="24" pageNumber="485">19D, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA636D04E55FF9F636E25CE4889F82B" blockId="24.[834,1474,1747,1931]" pageId="24" pageNumber="485">Tympani on anterior surfaces of all meri larger than tympani on posterior surfaces, smaller from P2 to P5. Dactyli of P2–P5 proximally straight, slightly curved distally, pointed, slightly shorter than propodi; anterior and posterior surfaces with one ridge extending slightly whole length; surfaces without granules, with sparse setae.</paragraph>
|
||
<paragraph id="8BA636D04E55FF9E636824AE4EBFFA07" blockId="24.[835,1476,1971,2092]" lastBlockId="25.[113,753,1391,1799]" lastPageId="25" lastPageNumber="486" pageId="24" pageNumber="485">Male pleon 1.8 times as long as wide. First somite trapezoid, 13 times as wide as long; anterior margin 0.86 times as long as posterior margin; 1.3 times wider than second somite. Second somite distinctly curved downward, 10 times as wide as long. Third somite 3.4 times as wide as long, anterior margin slightly concave, posterior edge convex.</paragraph>
|
||
<caption id="DF6666584E54FF9E605C26184F5BFA98" ID-DOI="http://doi.org/10.5281/zenodo.7399947" ID-Zenodo-Dep="7399947" httpUri="https://zenodo.org/record/7399947/files/figure.png" pageId="25" pageNumber="486" startId="25.[113,148,1285,1307]" targetBox="[114,1451,192,1270]" targetPageId="25">
|
||
<paragraph id="8BA636D04E54FF9E605C26184F5BFA98" blockId="25.[113,1452,1285,1336]" pageId="25" pageNumber="486">
|
||
Fig. 19.
|
||
<taxonomicName id="4C194D534E54FF9E60EF26184D31FABA" authority="Murniati & Asakura & Nugroho & Hernawan & Dharmawan, 2022" authorityName="Murniati & Asakura & Nugroho & Hernawan & Dharmawan" authorityYear="2022" box="[194,383,1285,1306]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="486" phylum="Arthropoda" rank="species" species="liki" status="sp. nov.">
|
||
<emphasis id="B96DEAC24E54FF9E60EF26184D31FABA" box="[194,383,1285,1306]" italics="true" pageId="25" pageNumber="486">Tmethypocoelis liki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25E57B94E54FF9E61A6261B4E4FFABB" box="[395,513,1285,1307]" pageId="25" pageNumber="486" rank="species">new species</taxonomicNameLabel>
|
||
. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Left pereopods: A–D, upper margin; E–H, lower margin; A, E, P2; B, F, P3; C, G, P4; D, H, P5.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA636D04E54FF9E605F26B24E61F8A7" blockId="25.[113,753,1391,1799]" pageId="25" pageNumber="486">
|
||
Fourth somite 2.8 times as wide as long; lateral margins concave, anterolateral angle rounded, posterolateral angle pointed; anterior margin convex; posterior margin nearly straight. Fifth somite 1.3 times as wide as long (at widest point), markedly constricted subproximally; proximal margin distinctly concave. Sixth somite 1.2 times as wide as long; anterior margin slightly concave, posterior margin slightly convex; lateral margins slightly wider subdistally; total length slightly as long as fifth somite, with slightly concave, parallel sides. Telson rounded; 1.3 times as wide as long; mid-posterior margin convex (
|
||
<figureCitation id="13222A554E54FF9E619025F24E6DF8A7" box="[445,547,1775,1799]" captionStart="Fig" captionStartId="22.[136,171,1700,1722]" captionTargetBox="[134,1474,192,1681]" captionTargetId="figure-8@22.[122,1477,186,1700]" captionTargetPageId="22" captionText="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." figureDoi="http://doi.org/10.5281/zenodo.7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="25" pageNumber="486">Fig. 16G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA636D04E54FF9E605C24324E15F827" blockId="25.[113,753,1839,1927]" pageId="25" pageNumber="486">
|
||
G1 long, very slender, curved; subproximal portion weakly bulging; apex forming two lobes; apical setae consisting one long setae and five short setae (
|
||
<figureCitation id="13222A554E54FF9E61E624724E01F827" box="[459,591,1903,1927]" captionStart="Fig" captionStartId="26.[135,170,2064,2086]" captionTargetBox="[235,1375,189,2048]" captionTargetId="figure-7@26.[235,1375,189,2048]" captionTargetPageId="26" captionText="Fig. 20. Tmethypocoelis liki, new species. Paratypes: A–C, MZB.Cru.5012, male, 5.0 × 3.0 mm; D–E, MZB.Cru.5012, male, 5.1 × 3.2 mm; Papua, Liki Island. Left G1: A, D, whole morphology; apical portion: B, E, lateral; C, mesial." figureDoi="http://doi.org/10.5281/zenodo.7399949" httpUri="https://zenodo.org/record/7399949/files/figure.png" pageId="25" pageNumber="486">Fig. 20A–E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA636D04E54FF9E605C24B249E3FA47" blockId="25.[113,753,1967,2087]" lastBlockId="25.[813,1453,1391,1511]" pageId="25" pageNumber="486">
|
||
Female chelipeds equal, small. Palm with outer surface smooth; upper and lower margins with microscopic tubercles (
|
||
<figureCitation id="13222A554E54FF9E605424F24C93F7A7" box="[121,221,2031,2055]" captionStart="Fig" captionStartId="22.[136,171,1700,1722]" captionTargetBox="[134,1474,192,1681]" captionTargetId="figure-8@22.[122,1477,186,1700]" captionTargetPageId="22" captionText="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." figureDoi="http://doi.org/10.5281/zenodo.7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="25" pageNumber="486">Fig. 16H</figureCitation>
|
||
); inner surface smooth. Fingers longer than palm. Fixed finger straight; cutting margin serrated; lower margin entire; tip with spoon shaped, with setae at tip; inner surface smooth. Dactylus with one furrow on outer surface; cutting margin slightly serrated; upper margin smooth and with setae; tip with spoon shaped and setae; inner surface smooth.
|
||
</paragraph>
|
||
<paragraph id="8BA636D04E54FF9E630025124F9BF867" blockId="25.[812,1452,1551,1991]" pageId="25" pageNumber="486">
|
||
Female pleon rounded, nearly as long as wide (
|
||
<figureCitation id="13222A554E54FF9E656F251249D1F987" box="[1346,1439,1551,1575]" captionStart="Fig" captionStartId="22.[136,171,1700,1722]" captionTargetBox="[134,1474,192,1681]" captionTargetId="figure-8@22.[122,1477,186,1700]" captionTargetPageId="22" captionText="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." figureDoi="http://doi.org/10.5281/zenodo.7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="25" pageNumber="486">Fig. 16I</figureCitation>
|
||
). First somite distinctly curved, 13.6 times as wide as long; wider than second somite. Second somite 8.2 times as wide as long. Third somite 6.7 times as wide as long; anterior margin slightly concave; posterior margin nearly straight; lateral margins wider anteriorly. Fourth somite 6.4 times as wide as long; lateral margins concave; anterior margin convex; posterior margin nearly straight. Fifth somite 4.2 times as wide as long (at widest point), narrower and longer than fourth somite. Sixth somite 3.9 times as wide as long; narrower than fifth somite, anterior margin concave, posterior margin slightly convex; lateral margins wider posteriorly. Telson triangular; 1.7 times as wide as long; mid-posterior margin convex.
|
||
</paragraph>
|
||
<paragraph id="8BA636D04E54FF9E630024F2485CF787" blockId="25.[813,1452,2031,2087]" pageId="25" pageNumber="486">
|
||
Vulva D-shaped, mesial margin of operculum with triangular projection (
|
||
<figureCitation id="13222A554E54FF9E63842B124848F787" box="[937,1030,2063,2087]" captionStart="Fig" captionStartId="22.[136,171,1700,1722]" captionTargetBox="[134,1474,192,1681]" captionTargetId="figure-8@22.[122,1477,186,1700]" captionTargetPageId="22" captionText="Fig. 16. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. A, carapace; B, external orbital angle; C, orbit area; D, eyestalk; E, epistome; F, outer surface of left third maxilliped; G, male pleon. Paratype, MZB.Cru.5012, female, 3.5 mm × 2.1 mm; Papua, Liki Island. H, outer surface of left chela; I, pleon; J, left vulva." figureDoi="http://doi.org/10.5281/zenodo.7399941" httpUri="https://zenodo.org/record/7399941/files/figure.png" pageId="25" pageNumber="486">Fig. 16J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6666584E57FF9D60AA2B0D4817F7E3" ID-DOI="http://doi.org/10.5281/zenodo.7399949" ID-Zenodo-Dep="7399949" httpUri="https://zenodo.org/record/7399949/files/figure.png" pageId="26" pageNumber="487" startId="26.[135,170,2064,2086]" targetBox="[235,1375,189,2048]" targetPageId="26">
|
||
<paragraph id="8BA636D04E57FF9D60AA2B0D4817F7E3" blockId="26.[135,1474,2064,2115]" pageId="26" pageNumber="487">
|
||
Fig. 20.
|
||
<taxonomicName id="4C194D534E57FF9D60F12B0D4DD4F785" authority="Murniati & Asakura & Nugroho & Hernawan & Dharmawan, 2022" authorityName="Murniati & Asakura & Nugroho & Hernawan & Dharmawan" authorityYear="2022" box="[220,410,2064,2085]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="487" phylum="Arthropoda" rank="species" species="liki" status="sp. nov.">
|
||
<emphasis id="B96DEAC24E57FF9D60F12B0D4DD4F785" box="[220,410,2064,2085]" italics="true" pageId="26" pageNumber="487">Tmethypocoelis liki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25E57B94E57FF9D61842B0D4E6FF786" box="[425,545,2064,2086]" pageId="26" pageNumber="487" rank="species">new species</taxonomicNameLabel>
|
||
. Paratypes: A–C, MZB.Cru.5012, male, 5.0 × 3.0 mm; D–E, MZB.Cru.5012, male, 5.1 × 3.2 mm; Papua, Liki Island. Left G1: A, D, whole morphology; apical portion: B, E, lateral; C, mesial.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6666584E56FF9C605C260C4884FA87" ID-DOI="http://doi.org/10.5281/zenodo.7399951" ID-Zenodo-Dep="7399951" box="[113,1226,1297,1319]" httpUri="https://zenodo.org/record/7399951/files/figure.png" pageId="27" pageNumber="488" startId="27.[113,148,1297,1319]" targetBox="[116,1452,191,1284]" targetPageId="27">
|
||
<paragraph id="8BA636D04E56FF9C605C260C4884FA87" blockId="27.[113,1226,1297,1319]" box="[113,1226,1297,1319]" pageId="27" pageNumber="488">
|
||
Fig. 21. Successive motion of waving display by male
|
||
<taxonomicName id="4C194D534E56FF9C62A2260C4F03FA86" authority="Murniati & Asakura & Nugroho & Hernawan & Dharmawan, 2022" authorityName="Murniati & Asakura & Nugroho & Hernawan & Dharmawan" authorityYear="2022" box="[655,845,1297,1318]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="species" species="liki" status="sp. nov.">
|
||
<emphasis id="B96DEAC24E56FF9C62A2260C4F03FA86" box="[655,845,1297,1318]" italics="true" pageId="27" pageNumber="488">Tmethypocoelis liki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25E57B94E56FF9C6377260C4F9CFA87" box="[858,978,1297,1319]" pageId="27" pageNumber="488" rank="species">new species</taxonomicNameLabel>
|
||
. Refer to text for details.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C303655B4E56FF9C605C26434D85FA16" pageId="27" pageNumber="488" type="etymology">
|
||
<paragraph id="8BA636D04E56FF9C605C26434D85FA16" blockId="27.[113,752,1374,1462]" pageId="27" pageNumber="488">
|
||
<emphasis id="B96DEAC24E56FF9C605C26434CBCFAD6" bold="true" box="[113,242,1374,1398]" pageId="27" pageNumber="488">Etymology.</emphasis>
|
||
The specific epithet of the new species is derived from the name of the
|
||
<typeStatus id="54A288724E56FF9C614826634DDDFA36" box="[357,403,1406,1430]" pageId="27" pageNumber="488">type</typeStatus>
|
||
locality, Liki Island. The name is used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C303655B4E56FF9B605E26C34D3DFCD5" lastPageId="28" lastPageNumber="489" pageId="27" pageNumber="488" type="discussion">
|
||
<paragraph id="8BA636D04E56FF9C605E26C3482AF936" blockId="27.[112,754,1502,2070]" lastBlockId="27.[812,1453,1374,1686]" pageId="27" pageNumber="488">
|
||
<emphasis id="B96DEAC24E56FF9C605E26C34CA6FA56" bold="true" box="[115,232,1502,1526]" pageId="27" pageNumber="488">Remarks.</emphasis>
|
||
There were four species of
|
||
<taxonomicName id="4C194D534E56FF9C621026C34EBEFA56" authorityName="Koelbel" authorityYear="1897" box="[573,752,1502,1526]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96DEAC24E56FF9C621026C34EBEFA56" box="[573,752,1502,1526]" italics="true" pageId="27" pageNumber="488">Tmethypocoelis</emphasis>
|
||
</taxonomicName>
|
||
worldwide, all of which are distributed in the Indo-West Pacific,
|
||
<taxonomicName id="4C194D534E56FF9C60EA25024E5BF996" authority="(Koelbel, 1897)" baseAuthorityName="Koelbel" baseAuthorityYear="1897" box="[199,533,1566,1590]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="species" species="ceratophora">
|
||
<emphasis id="B96DEAC24E56FF9C60EA25024C98F996" box="[199,214,1567,1590]" italics="true" pageId="27" pageNumber="488">T</emphasis>
|
||
.
|
||
<emphasis id="B96DEAC24E56FF9C60CF25034D28F996" box="[226,358,1566,1590]" italics="true" pageId="27" pageNumber="488">ceratophora</emphasis>
|
||
(
|
||
<bibRefCitation id="EF884B214E56FF9C615925034E43F996" author="Koelbel K" box="[372,525,1566,1590]" pageId="27" pageNumber="488" pagination="709 - 718" refId="ref18155" refString="Koelbel K (1897) Beschreibung der Krebse. In: Wissenschaftliche Ergebnisse der Reise des Grafen Bela Szechenyi in Ostasien. Volume 2. Pp. 709 - 718." type="book chapter" year="1897">Koelbel, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54A288724E56FF9C620F25034E1FF996" box="[546,593,1566,1590]" pageId="27" pageNumber="488">type</typeStatus>
|
||
locality:
|
||
<collectingCountry id="F30E76404E56FF9C629825034CFEF9F6" name="Hong Kong" pageId="27" pageNumber="488">Hong Kong</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C194D534E56FF9C60E825224E1EF9F6" authority="Davie & Kosuge, 1995" authorityName="Davie & Kosuge" authorityYear="1995" box="[197,592,1598,1622]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="species" species="choreutes">
|
||
<emphasis id="B96DEAC24E56FF9C60E825224C9AF9F6" box="[197,212,1599,1622]" italics="true" pageId="27" pageNumber="488">T</emphasis>
|
||
.
|
||
<emphasis id="B96DEAC24E56FF9C60CE25234D05F9F6" box="[227,331,1598,1622]" italics="true" pageId="27" pageNumber="488">choreutes</emphasis>
|
||
<bibRefCitation id="EF884B214E56FF9C617925234E1EF9F6" author="Davie PJF & Kosuge T" box="[340,592,1598,1622]" pageId="27" pageNumber="488" pagination="207 - 215" refId="ref17242" refString="Davie PJF & Kosuge T (1995) A new species of Tmethypocoelis (Crustacea: Brachyura: Ocypodidae) from Japan. Raffles Bulletin of Zoology, 43 (1): 207 - 215." type="journal article" year="1995">Davie & Kosuge, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54A288724E56FF9C624D25234EC1F9F6" box="[608,655,1598,1622]" pageId="27" pageNumber="488">type</typeStatus>
|
||
locality: Okinawa, Ryukyu Islands),
|
||
<taxonomicName id="4C194D534E56FF9C618425424EE1F9D6" authority="Davie, 1990" authorityName="Davie" authorityYear="1990" box="[425,687,1630,1654]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="species" species="koelbeli">
|
||
<emphasis id="B96DEAC24E56FF9C618425424DF6F9D6" box="[425,440,1631,1654]" italics="true" pageId="27" pageNumber="488">T</emphasis>
|
||
.
|
||
<emphasis id="B96DEAC24E56FF9C61E425434E50F9D6" box="[457,542,1630,1654]" italics="true" pageId="27" pageNumber="488">koelbeli</emphasis>
|
||
<bibRefCitation id="EF884B214E56FF9C620525434EE1F9D6" author="Davie PJF" box="[552,687,1630,1654]" pageId="27" pageNumber="488" pagination="463 - 473" refId="ref17201" refString="Davie PJF (1990) New and rare crabs of the subfamily Dotillinae (Crustacea: Ocypodidae) from northern Australia and Papua New Guinea. Memoirs of the Queensland Museum, 28 (2): 463 - 473." type="journal article" year="1990">Davie, 1990</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54A288724E56FF9C62ED25434EBEF9D6" box="[704,752,1630,1654]" pageId="27" pageNumber="488">type</typeStatus>
|
||
locality:
|
||
<collectingRegion id="49DDF8324E56FF9C60E125634DDEF936" box="[204,400,1662,1686]" country="Australia" name="Northern Territory" pageId="27" pageNumber="488">Northern Territory</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30E76404E56FF9C61B525634DB2F936" box="[408,508,1662,1686]" name="Australia" pageId="27" pageNumber="488">Australia</collectingCountry>
|
||
), and
|
||
<taxonomicName id="4C194D534E56FF9C621B25624CB9F916" authority="Davie, 1990" authorityName="Davie" authorityYear="1990" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="species" species="odontodactylus">
|
||
<emphasis id="B96DEAC24E56FF9C621B25624E0BF936" box="[566,581,1663,1686]" italics="true" pageId="27" pageNumber="488">T</emphasis>
|
||
.
|
||
<emphasis id="B96DEAC24E56FF9C627C25634EBEF936" box="[593,752,1662,1686]" italics="true" pageId="27" pageNumber="488">odontodactylus</emphasis>
|
||
<bibRefCitation id="EF884B214E56FF9C605C25834CB9F916" author="Davie PJF" box="[113,247,1694,1718]" pageId="27" pageNumber="488" pagination="463 - 473" refId="ref17201" refString="Davie PJF (1990) New and rare crabs of the subfamily Dotillinae (Crustacea: Ocypodidae) from northern Australia and Papua New Guinea. Memoirs of the Queensland Museum, 28 (2): 463 - 473." type="journal article" year="1990">Davie, 1990</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54A288724E56FF9C612525834D79F916" box="[264,311,1694,1718]" pageId="27" pageNumber="488">type</typeStatus>
|
||
locality: South of
|
||
<collectingRegion id="49DDF8324E56FF9C622625834E2BF916" box="[523,613,1694,1718]" country="Papua New Guinea" name="Madang" pageId="27" pageNumber="488">Madang</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30E76404E56FF9C625E25834C8DF976" name="Papua New Guinea" pageId="27" pageNumber="488">Papua New Guinea</collectingCountry>
|
||
). The key characters to distinguish those species are on the male cheliped and G1. In
|
||
<taxonomicName id="4C194D534E56FF9C61F625C24E32F956" baseAuthorityName="Koelbel" baseAuthorityYear="1897" box="[475,636,1758,1782]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="species" species="ceratophora">
|
||
<emphasis id="B96DEAC24E56FF9C61F625C24DA6F956" box="[475,488,1759,1782]" italics="true" pageId="27" pageNumber="488">T</emphasis>
|
||
.
|
||
<emphasis id="B96DEAC24E56FF9C61D525C34E32F956" box="[504,636,1758,1782]" italics="true" pageId="27" pageNumber="488">ceratophora</emphasis>
|
||
</taxonomicName>
|
||
, the upper margin of the dactylus has a subdistal tooth formed by a crest without distinct median granules, while in
|
||
<taxonomicName id="4C194D534E56FF9C626924024E86F896" authorityName="Davie & Kosuge" authorityYear="1995" box="[580,712,1822,1846]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="species" species="choreutes">
|
||
<emphasis id="B96DEAC24E56FF9C626924024E1FF896" box="[580,593,1823,1846]" italics="true" pageId="27" pageNumber="488">T</emphasis>
|
||
.
|
||
<emphasis id="B96DEAC24E56FF9C624D24034E86F896" box="[608,712,1822,1846]" italics="true" pageId="27" pageNumber="488">choreutes</emphasis>
|
||
</taxonomicName>
|
||
the upper margin of the dactylus has a subdistal tooth formed by a crest with clear median granules (
|
||
<bibRefCitation id="EF884B214E56FF9C620624434CE5F836" author="Davie PJF & Kosuge T" pageId="27" pageNumber="488" pagination="207 - 215" refId="ref17242" refString="Davie PJF & Kosuge T (1995) A new species of Tmethypocoelis (Crustacea: Brachyura: Ocypodidae) from Japan. Raffles Bulletin of Zoology, 43 (1): 207 - 215." type="journal article" year="1995">Davie & Kosuge, 1995</bibRefCitation>
|
||
: fig. 1A–F). In
|
||
<taxonomicName id="4C194D534E56FF9C616224624E5CF836" authorityName="Davie" authorityYear="1990" box="[335,530,1918,1942]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="species" species="odontodactylus">
|
||
<emphasis id="B96DEAC24E56FF9C616224624D10F836" box="[335,350,1919,1942]" italics="true" pageId="27" pageNumber="488">T</emphasis>
|
||
.
|
||
<emphasis id="B96DEAC24E56FF9C614124634E5CF836" box="[364,530,1918,1942]" italics="true" pageId="27" pageNumber="488">odontodactylus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96DEAC24E56FF9C623C24634E5BF836" box="[529,533,1918,1942]" italics="true" pageId="27" pageNumber="488">,</emphasis>
|
||
the upper margin of the dactylus of the cheliped is granular, which culminates subdistally as a strong upturned tooth (
|
||
<bibRefCitation id="EF884B214E56FF9C622724A34EC1F876" author="Davie PJF & Kosuge T" box="[522,655,1982,2006]" pageId="27" pageNumber="488" pagination="207 - 215" refId="ref17242" refString="Davie PJF & Kosuge T (1995) A new species of Tmethypocoelis (Crustacea: Brachyura: Ocypodidae) from Japan. Raffles Bulletin of Zoology, 43 (1): 207 - 215." type="journal article" year="1995">Davie, 1995</bibRefCitation>
|
||
, fig. 3C). On the other hand, the upper margin of the same portion in
|
||
<taxonomicName id="4C194D534E56FF9C60BC24E24D47F7B6" authorityName="Davie" authorityYear="1990" box="[145,265,2046,2070]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="species" species="koelbeli">
|
||
<emphasis id="B96DEAC24E56FF9C60BC24E24CEEF7B6" box="[145,160,2047,2070]" italics="true" pageId="27" pageNumber="488">T</emphasis>
|
||
.
|
||
<emphasis id="B96DEAC24E56FF9C609F24E34D47F7B6" box="[178,265,2046,2070]" italics="true" pageId="27" pageNumber="488">koelbeli</emphasis>
|
||
</taxonomicName>
|
||
is straight, extending three quarters of the length of the dactylus, which terminates in an overhanging ‘shelf’ (
|
||
<bibRefCitation id="EF884B214E56FF9C63AC26634847FA36" author="Davie PJF" box="[897,1033,1406,1430]" pageId="27" pageNumber="488" pagination="463 - 473" refId="ref17201" refString="Davie PJF (1990) New and rare crabs of the subfamily Dotillinae (Crustacea: Ocypodidae) from northern Australia and Papua New Guinea. Memoirs of the Queensland Museum, 28 (2): 463 - 473." type="journal article" year="1990">Davie, 1990</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C194D534E56FF9C6436266348A2FA36" authority="Murniati & Asakura & Nugroho & Hernawan & Dharmawan, 2022" authorityName="Murniati & Asakura & Nugroho & Hernawan & Dharmawan" authorityYear="2022" box="[1051,1260,1406,1430]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="species" species="liki" status="sp. nov.">
|
||
<emphasis id="B96DEAC24E56FF9C6436266348A2FA36" box="[1051,1260,1406,1430]" italics="true" pageId="27" pageNumber="488">Tmethypocoelis liki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25E57B94E56FF9C64D726634933FA36" box="[1274,1405,1406,1430]" pageId="27" pageNumber="488" rank="species">new species</taxonomicNameLabel>
|
||
, has a small tooth formed by one big granule on the subdistal portion of the dactylus upper margin. This tooth is the last granule on a granular ridge which runs from the proximal to the subdistal portion of the dactylus upper margin (
|
||
<figureCitation id="13222A554E56FF9C65AD26E34FCDF996" captionStart="Fig" captionStartId="23.[113,148,1745,1767]" captionTargetBox="[114,1450,191,1726]" captionTargetId="figure-11@23.[106,1469,167,1737]" captionTargetPageId="23" captionText="Fig. 17. Tmethypocoelis liki, new species. Holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm; Papua, Liki Island. Merus of the left cheliped: A, outer margin; B, lower margin; C, upper margin; D, lower surface; E, upper surface; F, outer surface. Carpus of the left cheliped: G, upper surface; H, lower surface; I, outer surface; J, upper margin; K, lower margin; L, outer margin. Left palm: M, inner surface; N, outer surface; O, upper margin; P, lower margin." figureDoi="http://doi.org/10.5281/zenodo.7399943" httpUri="https://zenodo.org/record/7399943/files/figure.png" pageId="27" pageNumber="488">Fig. 17M–N</figureCitation>
|
||
). The morphology of the projected tooth varied within the samples. Some specimens had a low tooth, whereas the others had a high tooth. There is no correlation between the tooth size and carapace size.
|
||
</paragraph>
|
||
<paragraph id="8BA636D04E56FF9C630025A34896F7B6" blockId="27.[813,1453,1726,2070]" pageId="27" pageNumber="488">
|
||
<taxonomicName id="4C194D534E56FF9C630025A34811F976" baseAuthorityName="Koelbel" baseAuthorityYear="1897" box="[813,1119,1726,1750]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="species" species="ceratophora">
|
||
<emphasis id="B96DEAC24E56FF9C630025A34811F976" box="[813,1119,1726,1750]" italics="true" pageId="27" pageNumber="488">Tmethypocoelis ceratophora</emphasis>
|
||
</taxonomicName>
|
||
has 7–8 apical setae on the G1 (Davie & Kosuge: 1995, fig. 2A–B;
|
||
<bibRefCitation id="EF884B214E56FF9C64EC25C34935F956" author="Shih HT & Chan BKK & Teng SJ & Wong KJH" box="[1217,1403,1758,1782]" pageId="27" pageNumber="488" refId="ref19758" refString="Shih HT, Chan BKK, Teng SJ & Wong KJH (2015) Crustacean Fauna of Taiwan. Brachyuran Crabs. Volume 2: Ocypodoidea. National Chung Hsing University, Taichung, 320 pp." type="book" year="2015">Shih et al., 2015</bibRefCitation>
|
||
: fig. 62A–D), while
|
||
<taxonomicName id="4C194D534E56FF9C63F425E2482FF8B6" authorityName="Davie & Kosuge" authorityYear="1995" box="[985,1121,1790,1814]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="species" species="choreutes">
|
||
<emphasis id="B96DEAC24E56FF9C63F425E2482FF8B6" box="[985,1121,1790,1814]" italics="true" pageId="27" pageNumber="488">T. choreutes</emphasis>
|
||
</taxonomicName>
|
||
has 12 apical setae (
|
||
<bibRefCitation id="EF884B214E56FF9C656125E34F88F896" author="Davie PJF & Kosuge T" pageId="27" pageNumber="488" pagination="207 - 215" refId="ref17242" refString="Davie PJF & Kosuge T (1995) A new species of Tmethypocoelis (Crustacea: Brachyura: Ocypodidae) from Japan. Raffles Bulletin of Zoology, 43 (1): 207 - 215." type="journal article" year="1995">Davie & Kosuge, 1995</bibRefCitation>
|
||
: fig. 2C–D). In
|
||
<taxonomicName id="4C194D534E56FF9C645E2402497AF896" authorityName="Davie" authorityYear="1990" box="[1139,1332,1822,1846]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="species" species="odontodactylus">
|
||
<emphasis id="B96DEAC24E56FF9C645E2402497AF896" box="[1139,1332,1822,1846]" italics="true" pageId="27" pageNumber="488">T. odontodactylus</emphasis>
|
||
</taxonomicName>
|
||
, the apical portion of the G1 is unilobed with one row of regular setae, with a simple pattern of setation (
|
||
<bibRefCitation id="EF884B214E56FF9C64B024434968F8D6" author="Davie PJF" box="[1181,1318,1886,1910]" pageId="27" pageNumber="488" pagination="463 - 473" refId="ref17201" refString="Davie PJF (1990) New and rare crabs of the subfamily Dotillinae (Crustacea: Ocypodidae) from northern Australia and Papua New Guinea. Memoirs of the Queensland Museum, 28 (2): 463 - 473." type="journal article" year="1990">Davie, 1990</bibRefCitation>
|
||
: fig. 3E–G). In
|
||
<taxonomicName id="4C194D534E56FF9C636724624FF5F836" authorityName="Davie" authorityYear="1990" box="[842,955,1918,1942]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="species" species="koelbeli">
|
||
<emphasis id="B96DEAC24E56FF9C636724624FF5F836" box="[842,955,1918,1942]" italics="true" pageId="27" pageNumber="488">T. koelbeli</emphasis>
|
||
</taxonomicName>
|
||
, the apical portion of the G1 is symmetrically bilobed, with five short and two long setae (
|
||
<bibRefCitation id="EF884B214E56FF9C6530248349E6F816" author="Davie PJF" box="[1309,1448,1950,1974]" pageId="27" pageNumber="488" pagination="463 - 473" refId="ref17201" refString="Davie PJF (1990) New and rare crabs of the subfamily Dotillinae (Crustacea: Ocypodidae) from northern Australia and Papua New Guinea. Memoirs of the Queensland Museum, 28 (2): 463 - 473." type="journal article" year="1990">Davie, 1990</bibRefCitation>
|
||
: fig. 2). In contrast, the apical portion of the G1 of
|
||
<taxonomicName id="4C194D534E56FF9C654E24A249EBF876" authority="Murniati & Asakura & Nugroho & Hernawan & Dharmawan, 2022" authorityName="Murniati & Asakura & Nugroho & Hernawan & Dharmawan" authorityYear="2022" box="[1379,1445,1982,2006]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="488" phylum="Arthropoda" rank="species" species="liki" status="sp. nov.">
|
||
<emphasis id="B96DEAC24E56FF9C654E24A249EBF876" box="[1379,1445,1982,2006]" italics="true" pageId="27" pageNumber="488">T. liki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25E57B94E56FF9C630024C34FFEF856" box="[813,944,2014,2038]" pageId="27" pageNumber="488" rank="species">new species</taxonomicNameLabel>
|
||
, is asymmetrically bilobed and armed with five short setae and one long seta (
|
||
<figureCitation id="13222A554E56FF9C645424E34885F7B6" box="[1145,1227,2046,2070]" captionStart="Fig" captionStartId="26.[135,170,2064,2086]" captionTargetBox="[235,1375,189,2048]" captionTargetId="figure-7@26.[235,1375,189,2048]" captionTargetPageId="26" captionText="Fig. 20. Tmethypocoelis liki, new species. Paratypes: A–C, MZB.Cru.5012, male, 5.0 × 3.0 mm; D–E, MZB.Cru.5012, male, 5.1 × 3.2 mm; Papua, Liki Island. Left G1: A, D, whole morphology; apical portion: B, E, lateral; C, mesial." figureDoi="http://doi.org/10.5281/zenodo.7399949" httpUri="https://zenodo.org/record/7399949/files/figure.png" pageId="27" pageNumber="488">Fig. 20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA636D04E51FF9B60A523A04F4BFE96" blockId="28.[135,775,189,310]" pageId="28" pageNumber="489">
|
||
<emphasis id="B96DEAC24E51FF9B60A523A04CA9FF75" bold="true" box="[136,231,189,213]" pageId="28" pageNumber="489">Habitat.</emphasis>
|
||
This species inhabits mangrove areas with a sandy substrate, mixed with very small gravel. Around its habitat, there are some long and narrow trenches which are inundated by water remaining from the previous high tide (
|
||
<figureCitation id="13222A554E51FF9B625622004EB7FE96" box="[635,761,285,310]" captionStart="Fig" captionStartId="21.[113,148,1127,1149]" captionTargetBox="[113,1451,189,1112]" captionTargetId="figure-10@21.[113,1451,189,1112]" captionTargetPageId="21" captionText="Fig. 15. Tmethypocoelis liki, new species. A, paratype, MZB.Cru.5012, male, 5.3 × 3.1 mm, in situ with life colouration. B, holotype, MZB.Cru.5011, male, 6.2 × 3.5 mm, dorsal habitus, preserved colouration; Papua, Liki Island. C–D, habitat of Tmethypocoelis liki, new species (photograph C by Sumaryanto Bronto)." figureDoi="http://doi.org/10.5281/zenodo.7399939" httpUri="https://zenodo.org/record/7399939/files/figure.png" pageId="28" pageNumber="489">Fig. 15C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA636D04E51FF9B60A522404D3DFCD5" blockId="28.[136,776,349,885]" pageId="28" pageNumber="489">
|
||
<emphasis id="B96DEAC24E51FF9B60A522404CB0FED5" bold="true" box="[136,254,349,373]" pageId="28" pageNumber="489">Behavior.</emphasis>
|
||
Male waving display of the new species is distinctly different from those of
|
||
<taxonomicName id="4C194D534E51FF9B622622634EFAFE35" baseAuthorityName="Koelbel" baseAuthorityYear="1897" box="[523,692,381,405]" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="489" phylum="Arthropoda" rank="species" species="ceratophora">
|
||
<emphasis id="B96DEAC24E51FF9B622622634EFAFE35" box="[523,692,381,405]" italics="true" pageId="28" pageNumber="489">T. ceratophora</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C194D534E51FF9B62DC22634E53FE15" authority="(Davie & Kosuge, 1995)" baseAuthorityName="Davie & Kosuge" baseAuthorityYear="1995" class="Malacostraca" family="Dotillidae" genus="Tmethypocoelis" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="489" phylum="Arthropoda" rank="species" species="choreutes">
|
||
<emphasis id="B96DEAC24E51FF9B62DC22634CBDFE15" italics="true" pageId="28" pageNumber="489">T. choreutes</emphasis>
|
||
(
|
||
<bibRefCitation id="EF884B214E51FF9B612B22804E5BFE15" author="Davie PJF & Kosuge T" box="[262,533,413,438]" pageId="28" pageNumber="489" pagination="207 - 215" refId="ref17242" refString="Davie PJF & Kosuge T (1995) A new species of Tmethypocoelis (Crustacea: Brachyura: Ocypodidae) from Japan. Raffles Bulletin of Zoology, 43 (1): 207 - 215." type="journal article" year="1995">Davie & Kosuge, 1995</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The display of
|
||
<specimenCount id="9D1FFD594E51FF9B62F322804C84FE75" pageId="28" pageNumber="489" type="male">five males</specimenCount>
|
||
was recorded with a digital camera. The waving (
|
||
<figureCitation id="13222A554E51FF9B60BC22C04CAEFE56" box="[145,224,477,502]" captionStart="Fig" captionStartId="27.[113,148,1297,1319]" captionTargetBox="[116,1452,191,1284]" captionTargetId="figure-11@27.[95,1465,178,1299]" captionTargetPageId="27" captionText="Fig. 21. Successive motion of waving display by male Tmethypocoelis liki, new species. Refer to text for details." figureDoi="http://doi.org/10.5281/zenodo.7399951" httpUri="https://zenodo.org/record/7399951/files/figure.png" pageId="28" pageNumber="489">Fig. 21</figureCitation>
|
||
) is started with both chelipeds positioned in front of the buccal cavern, the finger tips directed downward; they are then swung forward with the inner surface of the chela facing downward (1). The chela are then raised and directed outward, with fingertips also directed outward (2), then raised and directed centrally (3). The maximum height of the fingertips reaches beyond the projecting eyestalks. When reaching maximum height, the cheliped is in a nearly straight position with the fingertips pointing forward slightly (not pointing upward), meanwhile the P2 is raised (4). The chelipeds are then centrally gathered at a similar height to the maximum height, then refolded promptly (5) and returned to the initial position.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |