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<document id="F69BC3D4FA8AF423D5BDA1E01691E9E5" ID-DOI="10.11646/zootaxa.2598.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="10095059" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629698685068" checkinUser="felipe" docAuthor="Ruta, Rafał &amp; Yoshitomi, Hiroyuki" docDate="2010" docId="03B087BA681ABF14FF01FE1D1776FC7A" docLanguage="en" docName="zt02598p080.pdf" docOrigin="Zootaxa 2598 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2598.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Exochomoscirtes longispinus Ruta &amp; Yoshitomi 2010, sp. nov." docType="treatment" docUuid="2761D4CD-61FE-40FF-96D3-D926A45D9E2B" docUuidSource="ZooBank" docVersion="4" lastPageNumber="70" masterDocId="FF89FFC2685EBF51FF96FF9A130CFF80" masterDocTitle="Revision of the genus Exochomoscirtes Pic (Coleoptera: Scirtidae: Scirtinae) 2598" masterLastPageNumber="80" masterPageNumber="1" pageNumber="69" updateTime="1699569420699" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D975F54B39A0BAA4C89C9BA01498351B">Revision of the genus Exochomoscirtes Pic (Coleoptera: Scirtidae: Scirtinae) 2598</mods:title>
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<mods:namePart id="5DC51AAC8F47475D3346B5BBEDE041E3">Ruta, Rafał</mods:namePart>
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<mods:namePart id="E7A4247A9CAF7CE171D1DFC6EC317CD4">Yoshitomi, Hiroyuki</mods:namePart>
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<mods:title id="829A21F216152D7E5494FA60C0C1DB17">Zootaxa</mods:title>
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<mods:date id="97A473FE48D5D7DAB815B1DA9EAF983D">2010</mods:date>
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<mods:number id="CA4EB05F9B548BAFD2378D1B997FDC8A">2010-08-31</mods:number>
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<treatment id="03B087BA681ABF14FF01FE1D1776FC7A" LSID="urn:lsid:zoobank.org:act:2761D4CD-61FE-40FF-96D3-D926A45D9E2B" httpUri="http://treatment.plazi.org/id/03B087BA681ABF14FF01FE1D1776FC7A" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69">
<subSubSection id="C3036527681ABF15FF01FE1D1151FE21" box="[151,605,391,417]" pageId="68" pageNumber="69" type="nomenclature">
<paragraph id="8BA636AC681ABF15FF01FE1D1151FE21" blockId="68.[151,605,391,452]" box="[151,605,391,417]" pageId="68" pageNumber="69">
<heading id="D0EE81C0681ABF15FF01FE1D1151FE21" bold="true" box="[151,605,391,417]" fontSize="11" level="1" pageId="68" pageNumber="69" reason="1">
<emphasis id="B96DEABE681ABF15FF01FE1D1151FE21" bold="true" box="[151,605,391,417]" pageId="68" pageNumber="69">
<taxonomicName id="4C194D2F681ABF15FF01FE1D12F5FE21" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[151,505,391,417]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="longispinus" status="sp. nov.">
<emphasis id="B96DEABE681ABF15FF01FE1D12F5FE21" bold="true" box="[151,505,391,417]" italics="true" pageId="68" pageNumber="69">Exochomoscirtes longispinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C5681ABF15FD96FE1D1151FE21" box="[512,605,391,417]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3036527681ABF15FF01FE301275FE44" box="[151,377,426,452]" pageId="68" pageNumber="69" type="description">
<paragraph id="8BA636AC681ABF15FF01FE301275FE44" blockId="68.[151,605,391,452]" box="[151,377,426,452]" pageId="68" pageNumber="69">
(
<figureCitation id="13222A29681ABF15FF09FE30120AFE44" box="[159,262,426,452]" captionStart="FIGURE 3" captionStartId="8.[151,255,1079,1103]" captionTargetBox="[153,1433,194,1055]" captionTargetId="figure-129@8.[153,1433,194,1055]" captionTargetPageId="8" captionText="FIGURE 3. Metatibial spurs in Exochomoscirtes. A) E. dilaticornis sp. nov.; B) E. discoidalis (Pic); C) E. longispinus sp. nov.; D) E. nigromarginatus sp. nov.; E) E. picturatus sp. nov.; F) E. takizawai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10095065" httpUri="https://zenodo.org/record/10095065/files/figure.png" pageId="68" pageNumber="69">Figs. 3C</figureCitation>
,
<figureCitation id="13222A29681ABF15FE84FE301249FE44" box="[274,325,426,452]" captionStart="FIGURE 14" captionStartId="20.[151,255,1093,1117]" captionTargetBox="[170,1435,228,1061]" captionTargetId="figure-111@20.[151,1436,197,1067]" captionTargetPageId="20" captionText="FIGURE 14. Habitus illustrations of Exochomoscirtes. A) E. dilaticornis sp. nov., female; B) E. pectinatus sp. nov.; C) E. longispinus sp. nov.; D) E. nipponicus sp. nov.; E) E. nahangensis sp. nov.; F) E. arunachalensis sp. nov.; G) E. meghalayensis sp. nov. Scale bar = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10095087" httpUri="https://zenodo.org/record/10095087/files/figure.png" pageId="68" pageNumber="69">14C</figureCitation>
,
<figureCitation id="13222A29681ABF15FEC4FE30127DFE44" box="[338,369,426,452]" captionStart="FIGURE 56" captionStartId="68.[151,255,1501,1525]" captionTargetBox="[177,1426,801,1467]" captionTargetId="figure-273@68.[151,1436,791,1475]" captionTargetPageId="68" captionText="FIGURE 56. Exochomoscirtes longispinus sp. nov., male. A) sternite 8; B) sternite 9; C) tergite 8; D) tergite 9; E) tegmen; F) penis. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10095176" httpUri="https://zenodo.org/record/10095176/files/figure.png" pageId="68" pageNumber="69">56</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3036527681ABF15FF01FE6013EDFDE4" pageId="68" pageNumber="69" type="materials_examined">
<paragraph id="8BA636AC681ABF15FF01FE6013EDFDE4" blockId="68.[151,1436,506,732]" pageId="68" pageNumber="69">
<emphasis id="B96DEABE681ABF15FF01FE60125CFD94" bold="true" box="[151,336,506,532]" pageId="68" pageNumber="69">Type material.</emphasis>
<materialsCitation id="3B713CF1681ABF15FECDFE6013EDFDE4" collectingDate="1992-10" collectionCode="NHM" collectorName="J H Martin" country="Brunei" elevation="300" location="Belalong" municipality="Ridge" pageId="68" pageNumber="69" specimenCount="300" stateProvince="Temburong" typeStatus="holotype">
<emphasis id="B96DEABE681ABF15FECDFE601187FD94" bold="true" box="[347,651,506,532]" pageId="68" pageNumber="69">
<typeStatus id="54A2880E681ABF15FECDFE6012C2FD94" box="[347,462,506,532]" pageId="68" pageNumber="69" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="ED08AE69681ABF15FDB9FE601171FD94" box="[559,637,506,532]" pageId="68" pageNumber="69">NHM</collectionCode>
):
</emphasis>
<collectingCountry id="F30E763C681ABF15FD37FE601019FD94" box="[673,789,506,532]" name="Brunei" pageId="68" pageNumber="69">BRUNEI</collectingCountry>
;
<date id="FFA7106C681ABF15FCB2FE60107BFD94" box="[804,887,506,532]" pageId="68" pageNumber="69" value="1992-10">
<collectingDate id="EFE3E984681ABF15FCB2FE60107BFD94" box="[804,887,506,532]" pageId="68" pageNumber="69" value="1992-10">x-1992</collectingDate>
</date>
/
<collectingRegion id="49DDF84E681ABF15FC03FE601772FD94" box="[917,1150,506,532]" country="Brunei" name="Temburong" pageId="68" pageNumber="69">Temburong District</collectingRegion>
/
<collectingMunicipality id="6BC2ACD6681ABF15FB0AFE6017EEFD94" box="[1180,1250,506,532]" pageId="68" pageNumber="69">Ridge</collectingMunicipality>
<locationDeviation id="9A142F8E681ABF15FB7BFE601685FD94" box="[1261,1417,506,532]" location="Kuala" pageId="68" pageNumber="69">NE of Kuala</locationDeviation>
/
<location id="8EC66077681ABF15FF01FDB81208FDBC" LSID="urn:lsid:plazi:treatment:03B087BA681ABF14FF01FE1D1776FC7A:8EC66077681ABF15FF01FDB81208FDBC" box="[151,260,546,572]" country="Brunei" municipality="Ridge" name="Belalong" pageId="68" pageNumber="69" stateProvince="Temburong">Belalong</location>
,
<specimenCount id="9D1FFD25681ABF15FE84FDB81252FDBC" box="[274,350,546,572]" pageId="68" pageNumber="69" type="generic">
<quantity id="4CE19B49681ABF15FE84FDB81256FDBC" box="[274,346,546,572]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="68" pageNumber="69" unit="m" value="300.0">
<elevation id="0034D19F681ABF15FE84FDB81256FDBC" box="[274,346,546,572]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="68" pageNumber="69" unit="m" value="300.0">300m</elevation>
</quantity>
.
</specimenCount>
” [printed label]; “125W
<collectingMethod id="52584EBB681ABF15FD1AFDB81048FDBC" box="[652,836,546,572]" pageId="68" pageNumber="69">MV Light Trap</collectingMethod>
/
<collectorName id="26EC537A681ABF15FCC9FDB810EFFDBC" box="[863,995,546,572]" pageId="68" pageNumber="69">J H Martin</collectorName>
coll. / BMNH(E)1992-172” [printed label].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3036527681ABF15FF50FDE812ADFD5C" pageId="68" pageNumber="69" type="diagnosis">
<paragraph id="8BA636AC681ABF15FF50FDE812ADFD5C" blockId="68.[151,1436,506,732]" pageId="68" pageNumber="69">
<emphasis id="B96DEABE681ABF15FF50FDE8124AFD0C" bold="true" box="[198,326,626,652]" pageId="68" pageNumber="69">Diagnosis.</emphasis>
Similar to several uniformly brown species, from which it can be distinguished by the presence of a long, hooked, dorsal metatibial spur (
<figureCitation id="13222A29681ABF15FD57FD001017FD34" box="[705,795,666,692]" captionStart="FIGURE 3" captionStartId="8.[151,255,1079,1103]" captionTargetBox="[153,1433,194,1055]" captionTargetId="figure-129@8.[153,1433,194,1055]" captionTargetPageId="8" captionText="FIGURE 3. Metatibial spurs in Exochomoscirtes. A) E. dilaticornis sp. nov.; B) E. discoidalis (Pic); C) E. longispinus sp. nov.; D) E. nigromarginatus sp. nov.; E) E. picturatus sp. nov.; F) E. takizawai sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10095065" httpUri="https://zenodo.org/record/10095065/files/figure.png" pageId="68" pageNumber="69">Fig. 3C</figureCitation>
) in the males. Another distinctive feature is extraordinarily elongated penis.
</paragraph>
</subSubSection>
<caption id="DF666624681ABF15FF01FA47111CF997" ID-DOI="http://doi.org/10.5281/zenodo.10095176" ID-Zenodo-Dep="10095176" httpUri="https://zenodo.org/record/10095176/files/figure.png" pageId="68" pageNumber="69" startId="68.[151,255,1501,1525]" targetBox="[177,1426,801,1467]" targetPageId="68">
<paragraph id="8BA636AC681ABF15FF01FA47111CF997" blockId="68.[151,1436,1501,1560]" pageId="68" pageNumber="69">
<emphasis id="B96DEABE681ABF15FF01FA47122BFA75" bold="true" box="[151,295,1501,1525]" pageId="68" pageNumber="69">FIGURE 56.</emphasis>
<taxonomicName id="4C194D2F681ABF15FEB9FA44116BFA75" authority="Ruta &amp; Yoshitomi, 2010" authorityName="Ruta &amp; Yoshitomi" authorityYear="2010" box="[303,615,1502,1525]" class="Insecta" family="Scirtidae" genus="Exochomoscirtes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="longispinus" status="sp. nov.">
<emphasis id="B96DEABE681ABF15FEB9FA44116BFA75" box="[303,615,1502,1525]" italics="true" pageId="68" pageNumber="69">Exochomoscirtes longispinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C5681ABF15FDF8FA4411C8FA75" box="[622,708,1502,1525]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
, male.
<emphasis id="B96DEABE681ABF15FC87FA471021FA75" bold="true" box="[785,813,1501,1525]" pageId="68" pageNumber="69">A)</emphasis>
sternite 8;
<emphasis id="B96DEABE681ABF15FC31FA4710CDFA75" bold="true" box="[935,961,1501,1525]" pageId="68" pageNumber="69">B)</emphasis>
sternite 9;
<emphasis id="B96DEABE681ABF15FBADFA47175BFA75" bold="true" box="[1083,1111,1501,1525]" pageId="68" pageNumber="69">C)</emphasis>
tergite 8;
<emphasis id="B96DEABE681ABF15FB53FA4717EEFA75" bold="true" box="[1221,1250,1501,1525]" pageId="68" pageNumber="69">D)</emphasis>
tergite 9;
<emphasis id="B96DEABE681ABF15FAC6FA471667FA75" bold="true" box="[1360,1387,1501,1525]" pageId="68" pageNumber="69">E)</emphasis>
tegmen;
<emphasis id="B96DEABE681ABF15FF45F99A13E0F998" bold="true" box="[211,236,1536,1560]" pageId="68" pageNumber="69">F)</emphasis>
penis. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C3036527681ABF14FF50F9E91755FC2A" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69" type="description">
<paragraph id="8BA636AC681ABF15FF50F9E9113EF885" blockId="68.[151,1436,1651,2037]" pageId="68" pageNumber="69">
<emphasis id="B96DEABE681ABF15FF50F9E91252F90D" bold="true" box="[198,350,1651,1677]" pageId="68" pageNumber="69">Description.</emphasis>
<typeStatus id="54A2880E681ABF15FEFDF9E912D7F90D" box="[363,475,1651,1677]" pageId="68" pageNumber="69" type="holotype">Holotype</typeStatus>
, male. Body oval, TL/EW 1.3, slightly convex, covered with yellow suberect setae. Head dark brown, with lightened posterior margin, pronotum, scutellum and elytra brown, elytra with subtly darkened lateral and apical margins. Antennae and mouthparts yellowish-brown, legs yellowish-brown, metafemora darkened, dark brown.
</paragraph>
<paragraph id="8BA636AC681ABF15FF50F8891255F825" blockId="68.[151,1436,1651,2037]" pageId="68" pageNumber="69">Head with distinct punctation, punctures separated by ca. 1.0 diameter. Eyes big, protuberant; head twice width of interocular space. Antennae filiform, reaching basal 1/3 of elytra. Antennomeres 2 and 3 of subequal lengths, antennomeres 411 as long as antennomeres 2 and 3 combined. Palpomeres of maxillary palpi moderately widened.</paragraph>
<paragraph id="8BA636AC681ABF14FF50F82912B0FE82" blockId="68.[151,1436,1651,2037]" lastBlockId="69.[151,1436,152,1018]" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69">Pronotum slightly convex, covered with distinct punctation, punctures separated by 1.02.0 diameters, anterolateral corners almost right-angled, projecting anteriorly; lateral margins straight; posterolateral corners acute; base of pronotum subtrapezoidal, with incomplete margination along basal margin; PW/PL 0.4. Scutellum relatively big, punctured as on pronotum, equilaterally triangular. Angle between pronotum and elytra not marked in dorsal outline.</paragraph>
<paragraph id="8BA636AC681BBF14FF50FE8A10B7FEFA" blockId="69.[151,1436,152,1018]" pageId="69" pageNumber="70">Elytra oval, stronger diverging in posterior half, than in anterior portion, with slightly explanate lateral margins, widest at basal 1/3, punctation stronger and denser than on pronotum, punctures separated by ca. 1.0 diameter; humeri well marked; EL/EW 1.1; EL/PL 4.6; EW/PW 1.4.</paragraph>
<paragraph id="8BA636AC681BBF14FF50FE121776FE4A" blockId="69.[151,1436,152,1018]" pageId="69" pageNumber="70">Legs moderately long. Hind tibial spurs well developed, dorsal one extremely elongated, almost twice as long as ventral one, curved; ventral almost straight, slightly shorter than tarsomere 1.</paragraph>
<paragraph id="8BA636AC681BBF14FF50FE421172FD9A" blockId="69.[151,1436,152,1018]" pageId="69" pageNumber="70">Ventrite 1 with sparse setation in mesal portion, ventrite 2 with basal row of big punctures. Apex of tergite 7 unmodified, apex of ventrite 5 truncate.</paragraph>
<paragraph id="8BA636AC681BBF14FF50FDB211F5FCDA" blockId="69.[151,1436,152,1018]" pageId="69" pageNumber="70">
Sternite 8 (L 0.22,
<geoCoordinate id="EE2D506B681BBF14FE36FDB212F4FDC2" box="[416,504,552,578]" degrees="0.18" direction="west" orientation="longitude" pageId="69" pageNumber="70" precision="555" value="-0.18">W 0.18</geoCoordinate>
) small, widely U-shaped; sternite 9 (L 0.59,
<geoCoordinate id="EE2D506B681BBF14FC69FDB2175AFDC2" box="[1023,1110,552,578]" degrees="0.33" direction="west" orientation="longitude" pageId="69" pageNumber="70" precision="555" value="-0.33">W 0.33</geoCoordinate>
) consisting of two subtriangular hemisternites, with a row of very short apical setae. Tergite 8 (L 1.47,
<geoCoordinate id="EE2D506B681BBF14FBB6FDCA1775FDEA" box="[1056,1145,592,618]" degrees="0.80" direction="west" orientation="longitude" pageId="69" pageNumber="70" precision="555" value="-0.8">W 0.80</geoCoordinate>
) moderately sclerotized, bearing row of short setae on apical margin, with a pair of extraordinarily elongated apodemes diverging basally; tergite 9 (L 0.73,
<geoCoordinate id="EE2D506B681BBF14FE42FD3A1123FD3A" box="[468,559,672,698]" degrees="0.45" direction="west" orientation="longitude" pageId="69" pageNumber="70" precision="555" value="-0.45">W 0.45</geoCoordinate>
) moderately sclerotized, apical portion small, with minute setae at apex, with a pair of moderately long apodemes diverging basally. Tegmen (L 0.70,
<geoCoordinate id="EE2D506B681BBF14FBF2FD5217CDFD62" box="[1124,1217,712,738]" degrees="0.58" direction="west" orientation="longitude" pageId="69" pageNumber="70" precision="555" value="-0.58">W 0.58</geoCoordinate>
) well sclerotized, parameres straight, widening apically, with oval apical portions, mesal process narrow and slightly shorter than parameres. Penis (L 0.75,
<geoCoordinate id="EE2D506B681BBF14FD9BFC82116BFCB2" box="[525,615,792,818]" degrees="0.15" direction="west" orientation="longitude" pageId="69" pageNumber="70" precision="555" value="-0.15">W 0.15</geoCoordinate>
) relatively long, moderately sclerotized, pala oval, transforming apically into elongated narrow process pointed at apex.
</paragraph>
<paragraph id="8BA636AC681BBF14FF50FCF21295FC02" blockId="69.[151,1436,152,1018]" box="[198,409,872,898]" pageId="69" pageNumber="70">Female unknown.</paragraph>
<paragraph id="8BA636AC681BBF14FF50FC151755FC2A" blockId="69.[151,1436,152,1018]" box="[198,1113,911,938]" pageId="69" pageNumber="70">
<emphasis id="B96DEABE681BBF14FF50FC151289FC29" bold="true" box="[198,389,911,937]" pageId="69" pageNumber="70">Measurements.</emphasis>
Male (n = 1): TL 3.78, PL 0.70, PW 2.02, EL 3.20, EW 2.90.
</paragraph>
</subSubSection>
<subSubSection id="C3036527681BBF14FF50FC2D119DFC52" box="[198,657,951,978]" pageId="69" pageNumber="70" type="distribution">
<paragraph id="8BA636AC681BBF14FF50FC2D119DFC52" blockId="69.[151,1436,152,1018]" box="[198,657,951,978]" pageId="69" pageNumber="70">
<emphasis id="B96DEABE681BBF14FF50FC2D126BFC51" bold="true" box="[198,359,951,977]" pageId="69" pageNumber="70">Distribution.</emphasis>
Kuala Belalong, Borneo.
</paragraph>
</subSubSection>
<subSubSection id="C3036527681BBF14FF50FC451776FC7A" box="[198,1146,991,1018]" pageId="69" pageNumber="70" type="etymology">
<paragraph id="8BA636AC681BBF14FF50FC451776FC7A" blockId="69.[151,1436,152,1018]" box="[198,1146,991,1018]" pageId="69" pageNumber="70">
<emphasis id="B96DEABE681BBF14FF50FC45125FFC79" bold="true" box="[198,339,991,1017]" pageId="69" pageNumber="70">Etymology.</emphasis>
Specific epithet refers to extremely elongated dorsal metatibial spur.
</paragraph>
</subSubSection>
</treatment>
</document>