plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/B0/87/03B0879DFF93FFEBFE32D248FD0EFA08.xml

340 lines
39 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="D629AA61E8D704A4FDEF97C3D07A17BE" ID-DOI="10.5852/ejt.2024.926.2469" ID-GBIF-Dataset="4511b4bd-f026-49d3-9d4d-a39875e75cc5" ID-ISSN="2118-9773" ID-Zenodo-Dep="10847806" ID-ZooBank="A00D7669-2151-46AE-A066-9AF4D0F2BEEE" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1711011351298" checkinUser="plazi" docAuthor="Jochum, Adrienne, Michalik, Peter, Inäbnit, Thomas, Kneubühler, Jeannette, Slapnik, Rajko, Vrabec, Marko, Schilthuizen, Menno &amp; Ruthensteiner, Bernhard" docDate="2024" docId="03B0879DFF93FFEBFE32D248FD0EFA08" docLanguage="en" docName="EJT.2024.926.1-62.pdf" docOrigin="European Journal of Taxonomy 926" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2469/10973" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zospeum biokovoense Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner 2024, sp. nov." docType="treatment" docUuid="A976D460-68C4-460E-9023-E32539BE2C5B" docUuidSource="ZooBank" docVersion="11" lastPageNumber="44" masterDocId="FF89FFE5FFBAFFC0FFFFD700FFD4FFD4" masterDocTitle="3 D X-ray microscopy (Micro-CT) and SEM reveal Zospeum troglobalcanicum Absolon, 1916 and allied species from the Western Balkans (Ellobioidea: Carychiidae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="42" updateTime="1722640713706" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="35CF5CF88A9B8306B08003C5391D7C17" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8F5DDC81D38A247781D80E129B3138E8">
<mods:title id="3FF32517A5EAF22D28E8E575AD9E387A">3 D X-ray microscopy (Micro-CT) and SEM reveal Zospeum troglobalcanicum Absolon, 1916 and allied species from the Western Balkans (Ellobioidea: Carychiidae)</mods:title>
</mods:titleInfo>
<mods:name id="708E849019163A9A2DC5D823F2A7A79C" type="personal">
<mods:role id="EA54A9DA759882E16D917137A1E5B032">
<mods:roleTerm id="C748178521389CF3CB5EF3D9DE399E36">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="69A0EA504AABD6428B8509AC1F8968CE">Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier id="5C5F57FBB6BDF84D5317E1362E45724D" type="LSID">E2AD3AF2-ECC1-4DC3-BF5F-5C411AA946B3</mods:nameIdentifier>
<mods:affiliation id="C002E5BAAB1B017B896408630CEB133C">Senckenberg Research Institute and Natural History Museum, Senckenberganlage 25, D- 60325 Frankfurt / M., Germany.</mods:affiliation>
<mods:nameIdentifier id="AE4918F4951E1205E5C50660B2222DB7" type="email">nne.jochum@senckenberg.de</mods:nameIdentifier>
</mods:name>
<mods:name id="907F5039B53A05DD481C1759841A97D5" type="personal">
<mods:role id="BBBF859DC9F699D896BC73EA1C3BA4D4">
<mods:roleTerm id="C71AF6DC6F83C78F40A83D40F868763B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F7A212F7AB6538B42F667E486550073C">Michalik, Peter</mods:namePart>
<mods:nameIdentifier id="CEB8982FE5C54FD167C9B1651B991604" type="LSID">FAEE4B6B-80F5-4452-9ED2-08CA38DFC1F9</mods:nameIdentifier>
<mods:affiliation id="3F228086644F49AE556CE17BBFDA27B5">Zoological Institute and Museum, University of Greifswald, Loitzer Strasse 26, D- 17489 Greifswald, Germany.</mods:affiliation>
<mods:nameIdentifier id="4DCC363C34C067D7C0D0CC5DB3B7E545" type="email">michalik@uni-greifswald.de</mods:nameIdentifier>
</mods:name>
<mods:name id="EB2E8CE25787E9FEBC5263EA6C91F883" type="personal">
<mods:role id="A2C626FBE519D92F6860AA57D29261E8">
<mods:roleTerm id="561F4349DAD94D164BAB2D7D011CF90D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9945683B36B9E0C62040542C1FD66ADA">Inäbnit, Thomas</mods:namePart>
<mods:nameIdentifier id="6EB3071E0AF781CE62C047D9F1DD91D1" type="LSID">95F0148B-E147-4EB9-903A-6EEA7F55EB47</mods:nameIdentifier>
<mods:affiliation id="110FFBFE3D8EA17AA524841E26D57609">Institute for Biochemistry &amp; Biology, University of Potsdam, Karl-Liebknecht-Strasse 24 - 25, House 26, D- 14476 Potsdam, Germany.</mods:affiliation>
<mods:nameIdentifier id="3142C261B94E159FA2C3FCE56FE89BB1" type="email">inaebnit.thomas@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="8FC75247F0ADA34257855B4BD8E8883D" type="personal">
<mods:role id="2038DC15B4BD42F8E8830870EC1B8F8F">
<mods:roleTerm id="F672055D8CE1ABE700769CD9B57A0BBD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="14DDDBA0EE2F97A15EACD217D0561B43">Kneubühler, Jeannette</mods:namePart>
<mods:nameIdentifier id="D3B46D84EB927E5ABB8143CE48B7E2CE" type="LSID">A42A773F-9C7C-4681-B861-B354A330DC4F</mods:nameIdentifier>
<mods:affiliation id="0152DE1A3A1B4035D17A245A3FE218E1">Naturhistorisches Museum Bern, Bernastrasse 15, CH- 3005 Bern, Switzerland. &amp; Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, CH- 3012 Bern, Switzerland.</mods:affiliation>
<mods:nameIdentifier id="0BDBD127AF1A4D924E8688A4F1DF5305" type="email">jeannette.kneubuehler@nmbe.ch</mods:nameIdentifier>
</mods:name>
<mods:name id="7CDD4295AFE765B2AD0615E9EFEE2550" type="personal">
<mods:role id="CFA636786579F00878A0AB8A2238A666">
<mods:roleTerm id="33763254E8304A184F021A1E693B5340">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1AD51B83026E20BD010A186085AD9F66">Slapnik, Rajko</mods:namePart>
<mods:nameIdentifier id="9198E9F99AF1112295086DB443E8B767" type="LSID">09E86269-966B-49E6-AB92-31B66968DC0D</mods:nameIdentifier>
<mods:affiliation id="337D99B551F35AE28F0CB95EC60A1CAF">ZOSPEUM, Molluscs, Cave &amp; Karst Biological Consulting, Drnovškova pot 2, Mekinje, SI 1240 Kamnik, Slovenia.</mods:affiliation>
<mods:nameIdentifier id="4552E8A75E97620DD06754A72E62CB18" type="email">rajko.slapnik@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="E31BB9D8441F1D0593EA8D7CD9DC89C6" type="personal">
<mods:role id="6DA36A1788F24F768873DE58D075E233">
<mods:roleTerm id="6272FD82336DA0DCCFDAD77F0FFAF97E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9116D0A0730B324FB45A24D0DB8A315D">Vrabec, Marko</mods:namePart>
<mods:nameIdentifier id="312A62F68661549F3AF77AC348AE0741" type="LSID">00C5A26B-6E4A-41D9-BC9B-F36A2799E1AF</mods:nameIdentifier>
<mods:affiliation id="B4A1E437A0E364E055813C4CD8FB2197">Department of Geology, Faculty of Natural Sciences and Engineering, Aškerčeva 12, University of Ljubljana, 1000 Ljubljana, Slovenia.</mods:affiliation>
<mods:nameIdentifier id="522D68824D97A2FB5B405ABD0D84DC2E" type="email">marko.vrabec@geo.ntf.uni-lj.si</mods:nameIdentifier>
</mods:name>
<mods:name id="2BCB0B5D8E0B0AE69B5F47EC879DEE26" type="personal">
<mods:role id="A5D4D443BF36D6925099D22378B9F507">
<mods:roleTerm id="FB15E307202E53BAF9DC68A3D959C9C9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5ACF4567B0C326151FC796D41D5585F9">Schilthuizen, Menno</mods:namePart>
<mods:nameIdentifier id="E5A5DA770075E39CDB3E7B52E1FF084B" type="LSID">683D0AB7-CDD8-4FAA-94B6-436F3BFB8873</mods:nameIdentifier>
<mods:affiliation id="AED3D3E1705935EF05839F7460E49933">Naturalis Biodiversity Center and Leiden University, Vondellaan 55, NL- 2332 AA Leiden, The Netherlands.</mods:affiliation>
<mods:nameIdentifier id="7F37B8085CD35100D68BDB1A48290966" type="email">menno.schilthuizen@naturalis.nl</mods:nameIdentifier>
</mods:name>
<mods:name id="780C43909E499C87DA7ADFB2F7D04554" type="personal">
<mods:role id="DB532BEED232E2FD937B9CE40951E1FA">
<mods:roleTerm id="30568FDCE819360FD166F04B756998A9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B2C3D5F18088FDC9C4E8F91CA2020094">Ruthensteiner, Bernhard</mods:namePart>
<mods:nameIdentifier id="7B37D4BFC5E854F0181BF7BF20891646" type="LSID">9A8A56B0-19FA-42AB-A85E-4FCA98BF4A6F</mods:nameIdentifier>
<mods:affiliation id="1852631189CEEA4BE2C3D89DEB314A54">SNSB - Zoologische Staatssammlung München (ZSM), 81247 München, Germany.</mods:affiliation>
<mods:nameIdentifier id="5F9ADA519B3C1BC724A168B97DDCE1EC" type="email">ruthensteiner@snsb.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="15CBA8247CDAE5C28C8EB9D43DB96D17">text</mods:typeOfResource>
<mods:relatedItem id="AE58C7B8DC288DEEB7516B2C40CFF7C5" type="host">
<mods:titleInfo id="CA3FDE686D21A67DBD5D6318ACFC6405">
<mods:title id="4E78C29A17E423A7DAF3051F94250CF9">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="E28D670CB854C2A93A51173269A56B69">
<mods:date id="683D7FF509BDEED06BE15838EB29F60E">2024</mods:date>
<mods:detail id="852BBAA1BAEEA635E4A2411C9D92E0E1" type="pubDate">
<mods:number id="A7F50B77116F9D32EF2F88F0CC8BB023">2024-03-20</mods:number>
</mods:detail>
<mods:detail id="0A3E6990F49B04C5D828CBF451E0CCDC" type="volume">
<mods:number id="11894A47B3E5E55CC2D8248047DA69FE">926</mods:number>
</mods:detail>
<mods:extent id="18E852755D87F4BE7F689C233018127A" unit="page">
<mods:start id="1DD8FE3905572B19EFED1A8106AD29EB">1</mods:start>
<mods:end id="02E37378ED9320610626DEB52F8882AC">62</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="1ACC4E861DD1687C27C49F422A61ED71">
<mods:url id="775FFB96966CBA01D37923E87FB64FBD">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2469/10973</mods:url>
</mods:location>
<mods:classification id="073C1AE9F91AA874B5398BCF678B5FDB">journal article</mods:classification>
<mods:identifier id="E62080DB31822045F46879F041029E1D" type="DOI">10.5852/ejt.2024.926.2469</mods:identifier>
<mods:identifier id="638F9E3A6301A0036AE010AD0FFE8495" type="GBIF-Dataset">4511b4bd-f026-49d3-9d4d-a39875e75cc5</mods:identifier>
<mods:identifier id="2BE3809822EE5F84A004FCEDDC5E2F8C" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="B01F2577D11332FCD74CD04BFC85F25E" type="Zenodo-Dep">10847806</mods:identifier>
<mods:identifier id="61088E84B22FFD323E0A91120B3C6593" type="ZooBank">A00D7669-2151-46AE-A066-9AF4D0F2BEEE</mods:identifier>
</mods:mods>
<treatment id="03B0879DFF93FFEBFE32D248FD0EFA08" ID-DOI="http://doi.org/10.5281/zenodo.10847862" ID-GBIF-Taxon="224067519" ID-Zenodo-Dep="10847862" LSID="urn:lsid:zoobank.org:act:A976D460-68C4-460E-9023-E32539BE2C5B" httpUri="http://treatment.plazi.org/id/03B0879DFF93FFEBFE32D248FD0EFA08" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42">
<subSubSection id="C3036500FF93FFE9FE32D248FBB3FAB6" box="[461,1127,1351,1378]" pageId="41" pageNumber="42" type="nomenclature">
<paragraph id="8BA6368BFF93FFE9FE32D248FBB3FAB6" blockId="41.[372,1216,1351,1449]" box="[461,1127,1351,1378]" pageId="41" pageNumber="42">
<heading id="D0EE81E7FF93FFE9FE32D248FBB3FAB6" box="[461,1127,1351,1378]" centered="true" fontSize="11" level="2" pageId="41" pageNumber="42" reason="2">
<taxonomicName id="4C194D08FF93FFE9FE32D248FBD1FAB5" ID-CoL="C8NT4" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[461,1029,1351,1378]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="biokovoense" status="sp. nov.">
<emphasis id="B96DEA99FF93FFE9FE32D248FD0EFAB6" bold="true" box="[461,730,1351,1378]" italics="true" pageId="41" pageNumber="42">Zospeum biokovoense</emphasis>
Jochum &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FF93FFE9FBF2D248FBB3FAB6" box="[1037,1127,1352,1378]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF93FFE9FE8BD26AFC97FA7D" pageId="41" pageNumber="42" type="description">
<paragraph id="8BA6368BFF93FFE9FE8BD26AFB14FA51" blockId="41.[372,1216,1351,1449]" box="[372,1216,1386,1413]" pageId="41" pageNumber="42">
<uri id="FF883A89FF93FFE9FE8BD26AFB14FA51" box="[372,1216,1386,1413]" pageId="41" pageNumber="42">
urn:lsid:zoobank.org:act:
<uuid id="FFBF0C5EFF93FFE9FD6DD26AFB14FA51" box="[658,1216,1386,1413]" pageId="41" pageNumber="42">A976D460-68C4-460E-9023-E32539BE2C5B</uuid>
</uri>
</paragraph>
<paragraph id="8BA6368BFF93FFE9FD0FD28EFC97FA7D" blockId="41.[372,1216,1351,1449]" box="[752,835,1422,1449]" pageId="41" pageNumber="42">
<figureCitation id="13222A0EFF93FFE9FD0FD28EFC97FA7D" box="[752,835,1422,1449]" captionStart="Fig" captionStartId="42.[189,232,1822,1848]" captionTargetBox="[186,1395,842,1781]" captionTargetId="figure-263@42.[186,1395,842,1781]" captionTargetPageId="42" captionText="Fig. 15. Zospeum biokovoense Jochum &amp; Ruthensteiner sp. nov. (NHMW Mol.Coll.Edlauer 16.390). A–B. Light microscopic images showing aperture and dorsal views and collection labels. A. Juvenile shell. B. Holotype. C–H. 3D visualizations of Micro-CT data of holotype. C. Aperture view showing gnarled deformation of the columella. D. Aperture facing right view showing fat columella and lamella. E. Apical view. F. Dorsal view showing lopsided spire and incomplete lamella on top of fat columella. G. Aperture facing left view. H. Ventral view showing puckered umbilical depression." figureDoi="http://doi.org/10.5281/zenodo.10847848" httpUri="https://zenodo.org/record/10847848/files/figure.png" pageId="41" pageNumber="42">Fig. 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF93FFE9FF42D2CFFC19F955" pageId="41" pageNumber="42" type="diagnosis">
<paragraph id="8BA6368BFF93FFE9FF42D2CFFEE2FA3D" blockId="41.[189,310,1487,1513]" box="[189,310,1487,1513]" pageId="41" pageNumber="42">
<heading id="D0EE81E7FF93FFE9FF42D2CFFEE2FA3D" bold="true" box="[189,310,1487,1513]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="B96DEA99FF93FFE9FF42D2CFFEE2FA3D" bold="true" box="[189,310,1487,1513]" pageId="41" pageNumber="42">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF93FFE9FF42D2FEFC19F955" blockId="41.[189,1399,1532,1665]" pageId="41" pageNumber="42">
Shell ca
<quantity id="4CE19B6EFF93FFE9FED7D2FEFE5CF9CC" box="[296,392,1534,1560]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="41" pageNumber="42" unit="mm" value="1.4">1.4 mm</quantity>
, conical with 5¼ whorls; suture zone of penultimate and final whorl with irregular indentations succeeding into course growth lines with some crossing over parietal shield into the aperture; columella fat and short, almost 1/3 width of final whorl with lamellar bulge and an incomplete lamella; umbilical depression deep, callused with much puckering.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF93FFE9FF42D1A9FB94F925" box="[189,1088,1705,1778]" pageId="41" pageNumber="42" type="etymology">
<paragraph id="8BA6368BFF93FFE9FF42D1A9FE90F917" blockId="41.[189,324,1705,1731]" box="[189,324,1705,1731]" pageId="41" pageNumber="42">
<heading id="D0EE81E7FF93FFE9FF42D1A9FE90F917" bold="true" box="[189,324,1705,1731]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="B96DEA99FF93FFE9FF42D1A9FE90F917" bold="true" box="[189,324,1705,1731]" pageId="41" pageNumber="42">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF93FFE9FF42D1D7FB94F925" blockId="41.[189,1088,1751,1778]" box="[189,1088,1751,1778]" pageId="41" pageNumber="42">This species is named after the Biokovo mountain range where it was found.</paragraph>
</subSubSection>
<subSubSection id="C3036500FF93FFEAFF42D019FD85FE0D" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="8BA6368BFF93FFE9FF42D019FEB8F8E7" blockId="41.[189,364,1817,1843]" box="[189,364,1817,1843]" pageId="41" pageNumber="42">
<heading id="D0EE81E7FF93FFE9FF42D019FEB8F8E7" bold="true" box="[189,364,1817,1843]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="B96DEA99FF93FFE9FF42D019FEB8F8E7" bold="true" box="[189,364,1817,1843]" pageId="41" pageNumber="42">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF93FFE9FF15D047FE88F8B5" blockId="41.[189,1399,1863,1960]" box="[234,348,1863,1889]" pageId="41" pageNumber="42">
<emphasis id="B96DEA99FF93FFE9FF15D047FE88F8B5" bold="true" box="[234,348,1863,1889]" pageId="41" pageNumber="42">
<typeStatus id="54A28829FF93FFE9FF15D047FE88F8B5" box="[234,348,1863,1889]" pageId="41" pageNumber="42" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA6368BFF93FFEAFF42D06BFB71FE92" blockId="41.[189,1399,1863,1960]" lastBlockId="42.[188,1398,265,327]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<materialsCitation id="3B713CD6FF93FFEAFF42D06BFB71FE92" ID-GBIF-Occurrence="4598261311" collectionCode="NHMW Mol.Coll." collectorName="Edlauer" country="Croatia" county="Heger" lastPageId="42" lastPageNumber="43" latitude="43.3091" location="Biokovo Alpine Club Vosac" longLatPrecision="6" longitude="17.0473" municipality="Alpine Club" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Dalmatia" typeStatus="holotype">
<collectingCountry id="F30E761BFF93FFE9FF42D06BFEEEF851" box="[189,314,1899,1925]" name="Croatia" pageId="41" pageNumber="42">CROATIA</collectingCountry>
• (
<figureCitation id="13222A0EFF93FFE9FE98D06AFE28F851" box="[359,508,1898,1925]" captionStart="Fig" captionStartId="42.[189,232,1822,1848]" captionTargetBox="[186,1395,842,1781]" captionTargetId="figure-263@42.[186,1395,842,1781]" captionTargetPageId="42" captionText="Fig. 15. Zospeum biokovoense Jochum &amp; Ruthensteiner sp. nov. (NHMW Mol.Coll.Edlauer 16.390). A–B. Light microscopic images showing aperture and dorsal views and collection labels. A. Juvenile shell. B. Holotype. C–H. 3D visualizations of Micro-CT data of holotype. C. Aperture view showing gnarled deformation of the columella. D. Aperture facing right view showing fat columella and lamella. E. Apical view. F. Dorsal view showing lopsided spire and incomplete lamella on top of fat columella. G. Aperture facing left view. H. Ventral view showing puckered umbilical depression." figureDoi="http://doi.org/10.5281/zenodo.10847848" httpUri="https://zenodo.org/record/10847848/files/figure.png" pageId="41" pageNumber="42">Fig. 15B–H</figureCitation>
); “Dalmatien, kl. Grotte beim Alpenvereinshaus, Biokovogebirge, kleine Grotte am Weg vom Alpenvereins zum Hegerhaus“ [
<collectingRegion id="49DDF869FF93FFE9FCDED08EFC47F87C" box="[801,915,1934,1960]" pageId="41" pageNumber="42">Dalmatia</collectingRegion>
, small cave by Alpine Club house, (today probably
<location id="8EC66050FF90FFEAFED4D609FDA7FEF7" LSID="urn:lsid:plazi:treatment:03B0879DFF93FFEBFE32D248FD0EFA08:8EC66050FF90FFEAFED4D609FDA7FEF7" box="[299,627,265,291]" country="Croatia" county="Heger" latitude="43.3091" longLatPrecision="6" longitude="17.0473" municipality="Alpine Club" name="Biokovo Alpine Club Vosac" pageId="42" pageNumber="43" stateProvince="Dalmatia">Biokovo Alpine Club Vosac</location>
),
<location id="8EC66050FF90FFEAFD7AD609FCA5FEF7" LSID="urn:lsid:plazi:treatment:03B0879DFF93FFEBFE32D248FD0EFA08:8EC66050FF90FFEAFD7AD609FCA5FEF7" box="[645,881,265,291]" country="Croatia" county="Heger" latitude="43.3091" longLatPrecision="6" longitude="17.0473" municipality="Alpine Club" name="Biokovo Mountains" pageId="42" pageNumber="43" stateProvince="Dalmatia">Biokovo Mountains</location>
, small cave on the way from the Alpine Club house to Heger house]; [
<geoCoordinate id="EE2D504CFF90FFEAFE26D62CFDB2FE93" box="[473,614,300,327]" degrees="43.3091" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="5" value="43.3091">43.3091° N</geoCoordinate>
,
<geoCoordinate id="EE2D504CFF90FFEAFD8DD62CFD2FFE93" box="[626,763,300,327]" degrees="17.0473" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="5" value="17.0473">17.0473° E</geoCoordinate>
];
<collectionCode id="ED08AE4EFF90FFEAFCEFD62DFC32FE92" box="[784,998,300,327]" pageId="42" pageNumber="43">NHMW Mol.Coll.</collectionCode>
<collectorName id="26EC535DFF90FFEAFC1AD62CFB91FE92" box="[997,1093,300,326]" pageId="42" pageNumber="43">Edlauer</collectorName>
16.390.
</materialsCitation>
</paragraph>
<paragraph id="8BA6368BFF90FFEAFF42D66DFE2DFE53" blockId="42.[189,505,365,391]" box="[189,505,365,391]" pageId="42" pageNumber="43">
<heading id="D0EE81E7FF90FFEAFF42D66DFE2DFE53" bold="true" box="[189,505,365,391]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="B96DEA99FF90FFEAFF42D66DFE2DFE53" bold="true" box="[189,505,365,391]" pageId="42" pageNumber="43">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF90FFEAFF42D69CFD85FE0D" blockId="42.[189,1398,412,474]" pageId="42" pageNumber="43">
<materialsCitation id="3B713CD6FF90FFEAFF42D69CFD99FE0D" ID-GBIF-Occurrence="4598261301" collectionCode="NHMW Mol.Coll." collectorName="Edlauer" country="Croatia" county="Heger" latitude="43.3091" location="CROATIA" longLatPrecision="6" longitude="17.0473" municipality="Alpine Club" pageId="42" pageNumber="43" specimenCount="1">
<collectingCountry id="F30E761BFF90FFEAFF42D69CFEEEFE62" box="[189,314,412,438]" name="Croatia" pageId="42" pageNumber="43">CROATIA</collectingCountry>
•
<specimenCount id="9D1FFD02FF90FFEAFEA4D69CFE67FE63" box="[347,435,412,439]" count="1" pageId="42" pageNumber="43" type="generic">1 spec.</specimenCount>
; juvenile with damaged shell (
<figureCitation id="13222A0EFF90FFEAFCD4D69CFC49FE63" box="[811,925,412,439]" captionStart="Fig" captionStartId="42.[189,232,1822,1848]" captionTargetBox="[186,1395,842,1781]" captionTargetId="figure-263@42.[186,1395,842,1781]" captionTargetPageId="42" captionText="Fig. 15. Zospeum biokovoense Jochum &amp; Ruthensteiner sp. nov. (NHMW Mol.Coll.Edlauer 16.390). A–B. Light microscopic images showing aperture and dorsal views and collection labels. A. Juvenile shell. B. Holotype. C–H. 3D visualizations of Micro-CT data of holotype. C. Aperture view showing gnarled deformation of the columella. D. Aperture facing right view showing fat columella and lamella. E. Apical view. F. Dorsal view showing lopsided spire and incomplete lamella on top of fat columella. G. Aperture facing left view. H. Ventral view showing puckered umbilical depression." figureDoi="http://doi.org/10.5281/zenodo.10847848" httpUri="https://zenodo.org/record/10847848/files/figure.png" pageId="42" pageNumber="43">Fig. 15A</figureCitation>
); same collection data as for holotype;
<collectionCode id="ED08AE4EFF90FFEAFF42D6C0FE45FE0D" box="[189,401,447,474]" pageId="42" pageNumber="43">NHMW Mol.Coll.</collectionCode>
<collectorName id="26EC535DFF90FFEAFE6ED6BFFE25FE0D" box="[401,497,447,473]" pageId="42" pageNumber="43">Edlauer</collectorName>
16.390
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF90FFEBFF42D500FE08FDB4" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="description">
<paragraph id="8BA6368BFF90FFEAFF42D500FE9AFDCE" blockId="42.[189,334,512,538]" box="[189,334,512,538]" pageId="42" pageNumber="43">
<heading id="D0EE81E7FF90FFEAFF42D500FE9AFDCE" bold="true" box="[189,334,512,538]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="B96DEA99FF90FFEAFF42D500FE9AFDCE" bold="true" box="[189,334,512,538]" pageId="42" pageNumber="43">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF90FFEAFF42D530FAA3FDB8" blockId="42.[189,1399,559,621]" pageId="42" pageNumber="43">
<smallCapsWord id="8D40A057FF90FFEAFF42D530FEBAFD9C" baselines="579,579" box="[189,366,560,586]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="42" pageNumber="43">MEASUREMENTS</smallCapsWord>
.
<typeStatus id="54A28829FF90FFEAFE82D52FFE3BFD9D" box="[381,495,559,585]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
: sh:
<quantity id="4CE19B6EFF90FFEAFDDCD52FFD5BFD9E" box="[547,655,559,586]" metricMagnitude="-3" metricUnit="m" metricValue="1.37" pageId="42" pageNumber="43" unit="mm" value="1.37">1.37 mm</quantity>
; sw:
<quantity id="4CE19B6EFF90FFEAFD34D52FFC92FD9E" box="[715,838,559,586]" metricMagnitude="-4" metricUnit="m" metricValue="9.66" pageId="42" pageNumber="43" unit="mm" value="0.966">0.966 mm</quantity>
; ah:
<quantity id="4CE19B6EFF90FFEAFC82D52FFC23FD9E" box="[893,1015,559,586]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="42" pageNumber="43" unit="mm" value="0.68">0.680 mm</quantity>
; aw:
<quantity id="4CE19B6EFF90FFEAFBCAD52FFB64FD9E" box="[1077,1200,559,586]" metricMagnitude="-4" metricUnit="m" metricValue="5.96" pageId="42" pageNumber="43" unit="mm" value="0.596">0.596 mm</quantity>
; hlw:
<quantity id="4CE19B6EFF90FFEAFB08D52FFAA6FD9E" box="[1271,1394,559,586]" metricMagnitude="-4" metricUnit="m" metricValue="7.69" pageId="42" pageNumber="43" unit="mm" value="0.769">0.769 mm</quantity>
; SA: 66.39 deg. Other material: (N = 1): juvenile with damaged shell, not Micro-CT scanned or measured.
</paragraph>
<paragraph id="8BA6368BFF90FFEBFF44D594FE08FDB4" blockId="42.[187,1398,658,792]" lastBlockId="43.[188,1400,265,608]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
Shell ca
<quantity id="4CE19B6EFF90FFEAFEE2D594FEADFD7A" box="[285,377,660,686]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="42" pageNumber="43" unit="mm" value="1.4">1.4 mm</quantity>
, conical, with 5¼ regularly coiled, convex whorls with penultimate and last whorl being irregularly formed and somewhat lopsided; height of last whorl greater than half of shell height; opaque; aperture reniform, slightly higher than wide; teleoconch sculpture with some irregular blunt growth lines and weak radial banding; suture zone of penultimate and final whorl with dense, deep indentations of course growth lines extending at irregular intervals showing tendency towards costateness; peristome thin at upper palatal side, columellar side thickly callused roundish in form and slightly expanded, callosity extends beyond plane of shell (aperture facing left), basal palatal side narrowly reflected with some low ribbing behind palatal side; parietal shield thins at mid-section with some irregular growth lines bisecting it and entering into the aperture on the columellar side; notch at upper parietal and palatal junction; upper rim of peristome recedes ca
<superScript id="7C6C9BC3FF91FFEBFD3FD6B8FD1DFE12" attach="right" box="[704,713,440,454]" fontSize="6" pageId="43" pageNumber="44">1</superScript>
/
<subScript id="179D34CEFF91FFEBFD2ED6C4FD0EFE06" attach="left" box="[721,730,452,466]" fontSize="6" pageId="43" pageNumber="44">7</subScript>
the width of the penultimate whorl (aperture facing left); columella fat and short, almost ⅓ width of final whorl, centrally aligned and bulging at top with a thin, incomplete lamella; ventral side with deep, callused umbilical depression with much puckering and wrinkles; columellar side of peristome positioned directly above umbilical depression; alignment of last
<superScript id="7C6C9BC3FF91FFEBFF42D545FF12FD87" attach="none" box="[189,198,581,595]" fontSize="6" pageId="43" pageNumber="44">1</superScript>
/
<subScript id="179D34CEFF91FFEBFF31D550FF03FD8A" attach="left" box="[206,215,592,606]" fontSize="6" pageId="43" pageNumber="44">5</subScript>
whorl is not compact.
</paragraph>
</subSubSection>
<caption id="DF666603FF90FFEAFF42D01EFB64F83C" ID-DOI="http://doi.org/10.5281/zenodo.10847848" ID-Zenodo-Dep="10847848" httpUri="https://zenodo.org/record/10847848/files/figure.png" pageId="42" pageNumber="43" startId="42.[189,232,1822,1848]" targetBox="[186,1395,842,1781]" targetPageId="42">
<paragraph id="8BA6368BFF90FFEAFF42D01EFB64F83C" blockId="42.[189,1399,1822,2025]" pageId="42" pageNumber="43">
<emphasis id="B96DEA99FF90FFEAFF42D01EFEC8F8EC" bold="true" box="[189,284,1822,1848]" pageId="42" pageNumber="43">Fig. 15.</emphasis>
<taxonomicName id="4C194D08FF90FFEAFEDAD01FFC89F8EC" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[293,861,1822,1848]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="biokovoense" status="sp. nov.">
<emphasis id="B96DEA99FF90FFEAFEDAD01FFDF8F8EC" box="[293,556,1822,1848]" italics="true" pageId="42" pageNumber="43">Zospeum biokovoense</emphasis>
Jochum &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FF90FFEAFC99D01FFC17F8ED" box="[870,963,1823,1849]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
(NHMW Mol.Coll.Edlauer 16.390).
<emphasis id="B96DEA99FF90FFEAFF42D041FF21F88F" bold="true" box="[189,245,1857,1883]" pageId="42" pageNumber="43">A–B</emphasis>
. Light microscopic images showing aperture and dorsal views and collection labels.
<emphasis id="B96DEA99FF90FFEAFB0ED041FAD0F88F" bold="true" box="[1265,1284,1857,1883]" pageId="42" pageNumber="43">A</emphasis>
. Juvenile shell.
<emphasis id="B96DEA99FF90FFEAFEFAD065FECDF8AB" bold="true" box="[261,281,1893,1919]" pageId="42" pageNumber="43">B</emphasis>
. Holotype.
<emphasis id="B96DEA99FF90FFEAFE57D064FE34F8AA" bold="true" box="[424,480,1892,1918]" pageId="42" pageNumber="43">C–H</emphasis>
. 3D visualizations of Micro-CT data of holotype.
<emphasis id="B96DEA99FF90FFEAFBC2D064FB84F8AA" bold="true" box="[1085,1104,1892,1918]" pageId="42" pageNumber="43">C</emphasis>
. Aperture view showing gnarled deformation of the columella.
<emphasis id="B96DEA99FF90FFEAFD82D088FD46F876" bold="true" box="[637,658,1928,1954]" pageId="42" pageNumber="43">D</emphasis>
. Aperture facing right view showing fat columella and lamella.
<emphasis id="B96DEA99FF90FFEAFF42D0ABFF05F811" bold="true" box="[189,209,1963,1989]" pageId="42" pageNumber="43">E</emphasis>
. Apical view.
<emphasis id="B96DEA99FF90FFEAFE86D0ABFE5FF811" bold="true" box="[377,395,1963,1989]" pageId="42" pageNumber="43">F</emphasis>
. Dorsal view showing lopsided spire and incomplete lamella on top of fat columella.
<emphasis id="B96DEA99FF90FFEAFF42D0CEFF07F83C" bold="true" box="[189,211,1998,2024]" pageId="42" pageNumber="43">G</emphasis>
. Aperture facing left view.
<emphasis id="B96DEA99FF90FFEAFDEAD0CEFDF8F83C" bold="true" box="[533,556,1998,2024]" pageId="42" pageNumber="43">H</emphasis>
. Ventral view showing puckered umbilical depression.
</paragraph>
</caption>
<subSubSection id="C3036500FF91FFEBFF42D587FC66FD27" pageId="43" pageNumber="44" type="distribution">
<paragraph id="8BA6368BFF91FFEBFF42D587FE82FD75" blockId="43.[189,342,647,673]" box="[189,342,647,673]" pageId="43" pageNumber="44">
<heading id="D0EE81E7FF91FFEBFF42D587FE82FD75" bold="true" box="[189,342,647,673]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="B96DEA99FF91FFEBFF42D587FE82FD75" bold="true" box="[189,342,647,673]" pageId="43" pageNumber="44">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF91FFEBFF42D5B6FC66FD27" blockId="43.[189,1399,694,755]" pageId="43" pageNumber="44">
This species is only known from its
<typeStatus id="54A28829FF91FFEBFD81D5B6FD65FD04" box="[638,689,694,720]" pageId="43" pageNumber="44">type</typeStatus>
locality cave located in the Biokovo region of the central Dinarides between the Cetina and the Neretva Rivers of
<collectingCountry id="F30E761BFF91FFEBFCACD5D9FC79FD27" box="[851,941,729,755]" name="Croatia" pageId="43" pageNumber="44">Croatia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF91FFEBFF42D41AFD0EFA08" pageId="43" pageNumber="44" type="discussion">
<paragraph id="8BA6368BFF91FFEBFF42D41AFEFBFCE0" blockId="43.[189,303,794,820]" box="[189,303,794,820]" pageId="43" pageNumber="44">
<heading id="D0EE81E7FF91FFEBFF42D41AFEFBFCE0" bold="true" box="[189,303,794,820]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="B96DEA99FF91FFEBFF42D41AFEFBFCE0" bold="true" box="[189,303,794,820]" pageId="43" pageNumber="44">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF91FFEBFF42D449FD0EFA08" blockId="43.[186,1400,841,1500]" pageId="43" pageNumber="44">
<bibRefCitation id="EF884B7AFF91FFEBFF42D449FE55FCB7" author="Absolon K." box="[189,385,841,867]" pageId="43" pageNumber="44" pagination="242 - 309" refId="ref30302" refString="Absolon K. 1916 a. Vysledky vyzkumnych cest po Balkane [Results from the Balkan scientific expeditions]. Casopis Moravskeho Musea Zemskeho 15: 242 - 309." type="journal article" year="1916">Absolon (1916a)</bibRefCitation>
noted that the Neretva River was a distinctive geographical barrier reflected in different morphological features in species of subterranean fauna between the central Dinarides and the southern Balkans. The species
<taxonomicName id="4C194D08FF91FFEBFE39D490FC66FC7D" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[454,946,911,937]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="biokovoense" status="sp. nov.">
<emphasis id="B96DEA99FF91FFEBFE39D490FDAFFC7D" box="[454,635,911,937]" italics="true" pageId="43" pageNumber="44">Z. biokovoense</emphasis>
Jochum &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FF91FFEBFC42D490FBCFFC7E" box="[957,1051,912,938]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
differs from all other so far internally viewed (via Micro-CT) species by its remarkably fat columella. Though the lower side of the penultimate whorl shows some shell deformation directly above the columella, the columella itself bulges and demonstrates a threadlike lamella like that of
<emphasis id="B96DEA99FF91FFEBFCACD4FAFB9FFBC7" box="[851,1099,1017,1043]" italics="true" pageId="43" pageNumber="44">
<taxonomicName id="4C194D08FF91FFEBFCACD4FAFB93FBC7" authorityName="Absolon, 1916 Absolon" authorityYear="1916" box="[851,1095,1017,1043]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="troglobalcanicum">Z. troglobalcanicum</taxonomicName>
.
</emphasis>
The teleoconch however, differs in the lopsided formation of the penultimate whorl, the radial banding on the body whorl and the more numerous irregular growth lines extending in part across the parietal shield into the aperture of the species. The shell diameter is narrower, the aperture is taller and narrower, the height of the last whorl is shorter, and the spire is narrower in
<taxonomicName id="4C194D08FF91FFEBFDA6D386FCDDFB74" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[601,777,1158,1184]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="biokovoense">
<emphasis id="B96DEA99FF91FFEBFDA6D386FCDDFB74" box="[601,777,1158,1184]" italics="true" pageId="43" pageNumber="44">Z. biokovoense</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C194D08FF91FFEBFC9BD386FB81FB74" authorityName="Absolon, 1916 Absolon" authorityYear="1916" box="[868,1109,1158,1184]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="troglobalcanicum">
<emphasis id="B96DEA99FF91FFEBFC9BD386FB81FB74" box="[868,1109,1158,1184]" italics="true" pageId="43" pageNumber="44">Z. troglobalcanicum</emphasis>
</taxonomicName>
. Ventrally, the last whorl is less swollen and the final coiling towards the aperture extends high above the umbilical depression such that is not compact versus that in
<taxonomicName id="4C194D08FF91FFEBFD6DD3CDFC57FB32" authorityName="Absolon, 1916 Absolon" authorityYear="1916" box="[658,899,1228,1254]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="troglobalcanicum">
<emphasis id="B96DEA99FF91FFEBFD6DD3CDFC57FB32" box="[658,899,1228,1254]" italics="true" pageId="43" pageNumber="44">Z. troglobalcanicum</emphasis>
</taxonomicName>
, which does not extend as far beyond the depression and is compact in relation to it. The umbilical depression is callused, puckered, and wrinkled in
<taxonomicName id="4C194D08FF91FFEBFF24D213FE59FAF8" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[219,397,1298,1324]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="biokovoense">
<emphasis id="B96DEA99FF91FFEBFF24D213FE59FAF8" box="[219,397,1298,1324]" italics="true" pageId="43" pageNumber="44">Z. biokovoense</emphasis>
</taxonomicName>
and differs significantly from the smooth and relatively uniform umbilical formation in
<emphasis id="B96DEA99FF91FFEBFF22D236FE0CFA84" box="[221,472,1334,1360]" italics="true" pageId="43" pageNumber="44">
<taxonomicName id="4C194D08FF91FFEBFF22D236FE00FA84" authorityName="Absolon, 1916 Absolon" authorityYear="1916" box="[221,468,1334,1360]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="troglobalcanicum">Z. troglobalcanicum</taxonomicName>
.
</emphasis>
Coiling of the final whorl (ventral side) is similar to that of
<taxonomicName id="4C194D08FF91FFEBFB50D236FE0FFAA7" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Ruthensteiner" authorityYear="2024" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="amplioscutum" status="sp. nov.">
<emphasis id="B96DEA99FF91FFEBFB50D236FAA2FA84" box="[1199,1398,1334,1360]" italics="true" pageId="43" pageNumber="44">Z. amplioscutum</emphasis>
Jochum &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FF91FFEBFE1FD259FDECFAA7" box="[480,568,1369,1395]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
but it does not extend as far up the shell as in that species. Though growth lines extend into the parietal shield and into the aperture also in
<taxonomicName id="4C194D08FF91FFEBFC3ED27DFEB6FA6D" authority="Jochum, Schilthuizen &amp; Ruthensteiner" authorityName="Jochum, Schilthuizen &amp; Ruthensteiner" authorityYear="2024" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="43" pageNumber="2" phylum="Mollusca" rank="species" species="tumidum" status="sp. nov.">
Z.
<emphasis id="B96DEA99FF91FFEBFC18D27CFB9BFA42" box="[999,1103,1404,1430]" italics="true" pageId="43" pageNumber="44">tumidum</emphasis>
Jochum, Schilthuizen &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FF91FFEBFE95D2A0FE1CFA6E" box="[362,456,1440,1466]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
, the two species differ in aperture shape, whorl alignment, teleoconch structure, columellar configuration and size dimensions.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink