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<mods:title id="3ADDB2D91A8082CDD50AACF524A83E3D">Primnoidae (Cnidaria: Octocorallia: Calcaxonia) of the Okeanos Explorer expeditions (CAPSTONE) to the central Pacific</mods:title>
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<mods:namePart id="71DB86D0754AC1F4B98EF422A7BCC72E">Cairns, Stephen D.</mods:namePart>
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<paragraph id="8BA6A569FFFCFFD676CC6CD646E8F971" blockId="18.[151,474,1737,1764]" box="[151,474,1737,1764]" pageId="18" pageNumber="19">
<heading id="D0EE1205FFFCFFD676CC6CD646E8F971" bold="true" box="[151,474,1737,1764]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="B96D797BFFFCFFD676CC6CD646E8F971" bold="true" box="[151,474,1737,1764]" pageId="18" pageNumber="19">
Genus
<taxonomicName id="4C19DEEAFFFCFFD676B56CD646E8F971" ID-CoL="84QPH" authority="Gray, 1870" authorityName="Gray" authorityYear="1870" box="[238,474,1737,1764]" class="Anthozoa" family="Primnoidae" genus="Narella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="18" pageNumber="19" phylum="Cnidaria" rank="genus">
<emphasis id="B96D797BFFFCFFD676B56CD6467BF976" bold="true" box="[238,329,1737,1764]" italics="true" pageId="18" pageNumber="19">Narella</emphasis>
<bibRefCitation id="EF88D898FFFCFFD6770A6CD646E8F971" author="Gray, J. E." box="[337,474,1737,1763]" pageId="18" pageNumber="19" refId="ref28426" refString="Gray, J. E. (1870) Catalogue of the Lithophytes or Stony Corals in the Collection of the British Museum. British Museum, London, 51 pp." type="book" year="1870">Gray, 1870</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<treatmentCitationGroup id="AB098247FFFCFFD676CC6D1246FBF8D6" pageId="18" pageNumber="19">
<taxonomicName id="4C19DEEAFFFCFFD676CC6D1246B1F8B7" ID-CoL="84QPH" authority="Gray, 1870: 49" authorityName="Gray" authorityPageNumber="49" authorityYear="1870" box="[151,387,1804,1829]" class="Anthozoa" family="Primnoidae" genus="Narella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="18" pageNumber="19" phylum="Cnidaria" rank="genus">
<emphasis id="B96D797BFFFCFFD676CC6D1247D1F8B7" box="[151,227,1805,1829]" italics="true" pageId="18" pageNumber="19">Narella</emphasis>
<treatmentCitation id="0AB88378FFFCFFD676B16D1246B1F8B7" author="Gray, J. E." box="[234,387,1804,1829]" page="49" pageId="18" pageNumber="19" year="1870">
<bibRefCitation id="EF88D898FFFCFFD676B16D12466CF8B7" author="Gray, J. E." box="[234,350,1805,1829]" pageId="18" pageNumber="19" refId="ref28426" refString="Gray, J. E. (1870) Catalogue of the Lithophytes or Stony Corals in the Collection of the British Museum. British Museum, London, 51 pp." type="book" year="1870">Gray, 1870</bibRefCitation>
: 49
</treatmentCitation>
</taxonomicName>
.—
<treatmentCitation id="0AB88378FFFCFFD677FB6D134594F8B7" author="Cairns, S. D. &amp; Bayer, F. M." box="[416,678,1804,1830]" page="43" pageId="18" pageNumber="19" year="2009">
<bibRefCitation id="EF88D898FFFCFFD677FB6D1345B6F8B7" author="Cairns, S. D. &amp; Bayer, F. M." box="[416,644,1804,1830]" pageId="18" pageNumber="19" pagination="1 - 79" refId="ref28003" refString="Cairns, S. D. &amp; Bayer, F. M. (2009) A generic revision and phylogenetic analysis of the Primnoidae (Cnidaria: Octocorallia). Smithsonian Contributions to Zoology, 629, 1 - 79. https: // doi. org / 10.5479 / si. 00810282.629" type="journal article" year="2009">Cairns &amp; Bayer, 2009</bibRefCitation>
: 43
</treatmentCitation>
(synonymy and discussion).—
<treatmentCitation id="0AB88378FFFCFFD675B86D1343BBF8B7" author="Cairns, S. D." box="[995,1161,1804,1829]" page="14" pageId="18" pageNumber="19" year="2012">
<bibRefCitation id="EF88D898FFFCFFD675B86D13435BF8B7" author="Cairns, S. D." box="[995,1129,1804,1829]" pageId="18" pageNumber="19" pagination="1 - 71" refId="ref27497" refString="Cairns, S. D. (2012) New Zealand Primnoidae (Anthozoa: Alcyonacea) Part 1. Genera Narella, Narelloides, Metanarella, Calyptrophora, and Helicoprimnoa. NIWA Biodiversity Memoirs, 126, 1 - 71." type="journal article" year="2012">Cairns, 2012</bibRefCitation>
: 14
</treatmentCitation>
.—
<treatmentCitation id="0AB88378FFFCFFD672FC6D1347E6F8D6" author="Taylor, M. L. &amp; Rogers, A. D." page="4" pageId="18" pageNumber="19" year="2017">
<bibRefCitation id="EF88D898FFFCFFD672FC6D1342A8F8B7" author="Taylor, M. L. &amp; Rogers, A. D." box="[1191,1434,1804,1830]" pageId="18" pageNumber="19" pagination="70 - 97" refId="ref29067" refString="Taylor, M. L. &amp; Rogers, A. D. (2017) Primnoidae (Cnidaria: Octocorallia) of the SW Indian Ocean: new species, genus revisions and systematics. Zoological Journal of the Linnean Society, 181, 70 - 97. https: // doi. org / 10.1093 / zoolinnean / zlx 003" type="journal article" year="2017">Taylor &amp; Rogers, 2017</bibRefCitation>
: 4
</treatmentCitation>
.—Cairns, 2018: 20–21.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA6A569FFFCFFD676CC6D5544B8F8F1" blockId="18.[151,1437,1804,1921]" box="[151,906,1866,1891]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="AB098247FFFCFFD676CC6D5544B8F8F1" box="[151,906,1866,1891]" pageId="18" pageNumber="19">
<taxonomicName id="4C19DEEAFFFCFFD676CC6D5545B2F8F1" authority="Wright &amp; Studer in Studer, 1887: 49" authorityName="Wright &amp; Studer in Studer" authorityPageNumber="49" authorityYear="1887" box="[151,640,1866,1891]" class="Anthozoa" family="Primnoidae" genus="Stachyodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="18" pageNumber="19" phylum="Cnidaria" rank="genus">
<emphasis id="B96D797BFFFCFFD676CC6D55463BF8F0" box="[151,265,1866,1890]" italics="true" pageId="18" pageNumber="19">Stachyodes</emphasis>
Wright &amp; Studer in
<treatmentCitation id="0AB88378FFFCFFD677826D5545B2F8F1" author="Studer, T." box="[473,640,1866,1891]" page="49" pageId="18" pageNumber="19" year="1887">
<bibRefCitation id="EF88D898FFFCFFD677826D55456FF8F1" author="Studer, T." box="[473,605,1866,1891]" pageId="18" pageNumber="19" pagination="1 - 74" refId="ref29011" refString="Studer, T. (1887) Versuch Eines Systemes der Alcyonaria. Archive fur Naturgeschichte, 53 (1), 1 - 74." type="journal article" year="1887">Studer, 1887</bibRefCitation>
: 49
</treatmentCitation>
</taxonomicName>
.—
<treatmentCitation id="0AB88378FFFCFFD674C56D55446CF8F1" author="Versluys, J." box="[670,862,1866,1891]" page="86" pageId="18" pageNumber="19" year="1906">
<bibRefCitation id="EF88D898FFFCFFD674C56D554408F8F1" author="Versluys, J." box="[670,826,1866,1891]" pageId="18" pageNumber="19" pagination="1 - 187" refId="ref29267" refString="Versluys, J. (1906) Die Gorgoniden der Siboga - Expedition. II. Die Primnoidae. Siboga-Expeditie, 13 a, 1 - 187." type="book chapter" year="1906">Versluys, 1906</bibRefCitation>
: 86
</treatmentCitation>
–88.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA6A569FFFCFFD676CC6D76458CF813" blockId="18.[151,1437,1804,1921]" box="[151,702,1896,1921]" pageId="18" pageNumber="19">
<emphasis id="B96D797BFFFCFFD676CC6D764628F813" box="[151,282,1897,1921]" italics="true" pageId="18" pageNumber="19">Calypternius</emphasis>
Wright &amp; Studer in
<bibRefCitation id="EF88D898FFFCFFD677B16D77455FF813" author="Studer, T." box="[490,621,1896,1921]" pageId="18" pageNumber="19" pagination="1 - 74" refId="ref29011" refString="Studer, T. (1887) Versuch Eines Systemes der Alcyonaria. Archive fur Naturgeschichte, 53 (1), 1 - 74." type="journal article" year="1887">Studer, 1887</bibRefCitation>
: 49–50.
</paragraph>
</subSubSection>
<subSubSection id="C303F6E2FFFCFFD676CC6DB344E1F856" box="[151,979,1963,1988]" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BA6A569FFFCFFD676CC6DB344E1F856" blockId="18.[151,979,1963,1988]" box="[151,979,1963,1988]" pageId="18" pageNumber="19">
<emphasis id="B96D797BFFFCFFD676CC6DB34610F856" bold="true" box="[151,290,1963,1988]" pageId="18" pageNumber="19">
<typeStatus id="54A21BCBFFFCFFD676CC6DB347FEF856" box="[151,204,1964,1988]" pageId="18" pageNumber="19">Type</typeStatus>
Species
</emphasis>
.
<taxonomicName id="4C19DEEAFFFCFFD677756DB2440EF856" authority="Duchassaing &amp; Michelotti, 1860" authorityName="Duchassaing &amp; Michelotti" authorityYear="1860" box="[302,828,1963,1988]" class="Anthozoa" family="Primnoidae" genus="Primnoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="18" pageNumber="19" phylum="Cnidaria" rank="species" species="regularis">
<emphasis id="B96D797BFFFCFFD677756DB246DBF856" box="[302,489,1964,1988]" italics="true" pageId="18" pageNumber="19">Primnoa regularis</emphasis>
Duchassaing &amp; Michelotti, 1860
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<caption id="DF66F5E1FFFDFFD776CC6CC145D2F8AC" httpUri="https://zenodo.org/record/2614983/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1758,1783]" targetBox="[174,1414,151,1738]" targetPageId="19">
<paragraph id="8BA6A569FFFDFFD776CC6CC145D2F8AC" blockId="19.[151,1437,1758,1855]" pageId="19" pageNumber="20">
<emphasis id="B96D797BFFFDFFD776CC6CC14621F965" bold="true" box="[151,275,1758,1783]" pageId="19" pageNumber="20">FIGURE 9.</emphasis>
<taxonomicName id="4C19DEEAFFFDFFD777426CC046C9F964" authorityName="Cairns" authorityYear="2018" box="[281,507,1759,1782]" class="Anthozoa" family="Primnoidae" genus="Macroprimnoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="19" pageNumber="20" phylum="Cnidaria" rank="species" species="ornata">
<emphasis id="B96D797BFFFDFFD777426CC046C9F964" box="[281,507,1759,1782]" italics="true" pageId="19" pageNumber="20">Macroprimnoa ornata</emphasis>
</taxonomicName>
, paratype from USNM 142423:
<emphasis id="B96D797BFFFDFFD7751F6CC14447F965" bold="true" box="[836,885,1758,1783]" pageId="19" pageNumber="20">A–C</emphasis>
, stereo views of lateral and opercular sides of a polyp.
<emphasis id="B96D797BFFFDFFD776CC6D1D479BF888" bold="true" box="[151,169,1794,1818]" pageId="19" pageNumber="20">D</emphasis>
, stereo view of coenenchymal scales,
<emphasis id="B96D797BFFFDFFD774626D1C4548F888" box="[569,634,1795,1818]" italics="true" pageId="19" pageNumber="20">in situ</emphasis>
.
<emphasis id="B96D797BFFFDFFD774D36D1D45ABF888" bold="true" box="[648,665,1794,1818]" pageId="19" pageNumber="20">E</emphasis>
, body wall scales,
<emphasis id="B96D797BFFFDFFD7753A6D1C4490F888" box="[865,930,1795,1818]" italics="true" pageId="19" pageNumber="20">in situ</emphasis>
.
<emphasis id="B96D797BFFFDFFD775EA6D1D44F2F888" bold="true" box="[945,960,1794,1818]" pageId="19" pageNumber="20">F</emphasis>
, highly ridged body wall scales.
<emphasis id="B96D797BFFFDFFD773416D1D421CF889" bold="true" box="[1306,1326,1794,1819]" pageId="19" pageNumber="20">G</emphasis>
, opercular scales.
<emphasis id="B96D797BFFFDFFD776BB6D3947C6F8AC" bold="true" box="[224,244,1830,1854]" pageId="19" pageNumber="20">H</emphasis>
, ridged coenenchymal scales. Scale bars in mm.
</paragraph>
</caption>
<subSubSection id="C303F6E2FFFDFFD076CC6D7347D4FF69" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BA6A569FFFDFFD076CC6D7347D4FF69" blockId="19.[151,1437,1900,2035]" lastBlockId="20.[151,1437,153,539]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<emphasis id="B96D797BFFFDFFD776CC6D734638F815" bold="true" box="[151,266,1900,1927]" pageId="19" pageNumber="20">Diagnosis</emphasis>
. Colonies branched dichotomously (laterally or equal), pinnately, in a lyrate fashion, or unbranched. Polyps arranged in whorls, all polyps facing downward in contracted condition. Each polyp covered with three (rarely four) pairs of abaxial body wall scales (one pair of basals, one or two pairs of medials, and one pair of buccals) and a variable number of pairs of smaller adaxial scales, nonetheless leaving the adaxial face largely naked. Articular ridge not present on basal scales. Paired infrabasal scales usually present. Opercular scales keeled on inner surface. Coenenchymal scales thin and slightly imbricate or thick and mosaic in placement, and sometimes prominently ridged.
</paragraph>
</subSubSection>
<subSubSection id="C303F6E2FFFAFFD0769C6B1B44A2FE65" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8BA6A569FFFAFFD0769C6B1B469EFE3D" blockId="20.[151,1437,153,539]" pageId="20" pageNumber="21">
<emphasis id="B96D797BFFFAFFD0769C6B1B4677FE8D" bold="true" box="[199,325,260,287]" pageId="20" pageNumber="21">Discussion</emphasis>
. Although keys exist to the
<taxonomicName id="4C19DEEAFFFAFFD074DD6B1A45E9FE8D" authorityName="Gray" authorityYear="1870" box="[646,731,261,287]" class="Anthozoa" family="Primnoidae" genus="Narella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="20" pageNumber="21" phylum="Cnidaria" rank="genus">
<emphasis id="B96D797BFFFAFFD074DD6B1A45E9FE8D" box="[646,731,261,287]" italics="true" pageId="20" pageNumber="21">Narella</emphasis>
</taxonomicName>
species from various geographic regions (see Cairns 2018 for a summary), a comprehensive key, preferably a tabular key, is sorely needed for the 50 species in the world, and is currently being constructed by Cairns and Taylor (in prep.). The last comprehensive key to the species was that of K̹kenthal (1919, 1924), which included only 18 species. Most of the currently recognized species are listed by
<bibRefCitation id="EF88D898FFFAFFD076CC6B8A469AFE3D" author="Taylor, M. L. &amp; Rogers, A. D." box="[151,424,405,431]" pageId="20" pageNumber="21" pagination="70 - 97" refId="ref29067" refString="Taylor, M. L. &amp; Rogers, A. D. (2017) Primnoidae (Cnidaria: Octocorallia) of the SW Indian Ocean: new species, genus revisions and systematics. Zoological Journal of the Linnean Society, 181, 70 - 97. https: // doi. org / 10.1093 / zoolinnean / zlx 003" type="journal article" year="2017">Taylor &amp; Rogers (2017)</bibRefCitation>
.
</paragraph>
<paragraph id="8BA6A569FFFAFFD0769C6BA644A2FE65" blockId="20.[151,1437,153,539]" pageId="20" pageNumber="21">The order in which species are treated below are grouped by presence or absence of a ridge on the dorsolateral basal edge, followed by the colony form, simplest to most complex.</paragraph>
</subSubSection>
<subSubSection id="C303F6E2FFFAFFD0769C681F44B6FD89" box="[199,900,512,539]" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8BA6A569FFFAFFD0769C681F44B6FD89" blockId="20.[151,1437,153,539]" box="[199,900,512,539]" pageId="20" pageNumber="21">
<emphasis id="B96D797BFFFAFFD0769C681F466BFD89" bold="true" box="[199,345,512,539]" pageId="20" pageNumber="21">Distribution</emphasis>
. All ocean basins except for Arctic,
<quantity id="4CE1088CFFFAFFD074AB681F44B2FD89" box="[752,896,512,539]" metricMagnitude="3" metricUnit="m" metricValue="2.3609999999999998" metricValueMax="4.593999999999999" metricValueMin="0.128" pageId="20" pageNumber="21" unit="m" value="2361.0" valueMax="4594.0" valueMin="128.0">128–4594 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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