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<document id="3F909AC848F39D5FC26E441F2A5322FE" ID-CLB-Dataset="56236" ID-DOI="10.11646/zootaxa.5259.1.1" ID-GBIF-Dataset="e2c999aa-4046-4fc1-a351-158efbf234cb" ID-ISSN="1175-5326" ID-Zenodo-Dep="7794984" ID-ZooBank="4306F52E-FD24-45B0-B307-66B71173D805" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1680527882783" checkinUser="plazi" docAuthor="Granja-Fernández, Rebeca, Hendrickx, Michel E., Rangel-Solís, Pedro Diego &amp; López-Pérez, Andrés" docDate="2023" docId="03AB87FAFFB0FFC1FF7BF905FCDBFB4E" docLanguage="en" docName="zootaxa.5259.1.1.pdf" docOrigin="Zootaxa 5259 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5259.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ophiosphalma variabile" docType="treatment" docVersion="9" lastPageNumber="21" masterDocId="FF92FF82FFA1FFD5FFECFFC3FFBAFFA7" masterDocTitle="Deep-sea Ophiuroidea (Echinodermata) collected during the TALUD cruises in western Mexico" masterLastPageNumber="71" masterPageNumber="1" pageNumber="18" updateTime="1699481084854" updateUser="plazi">
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<mods:title id="66965368B2443C7C0440A1512B08F18B">Deep-sea Ophiuroidea (Echinodermata) collected during the TALUD cruises in western Mexico</mods:title>
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<mods:namePart id="E27F824E299BBC43CF61DF3DF4610781">Hendrickx, Michel E.</mods:namePart>
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<mods:namePart id="AA229F30181B4B49E0684E38DC87AF77">Rangel-Solís, Pedro Diego</mods:namePart>
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<treatment id="03AB87FAFFB0FFC1FF7BF905FCDBFB4E" ID-DOI="http://doi.org/10.5281/zenodo.7798728" ID-GBIF-Taxon="209487032" ID-Zenodo-Dep="7798728" LSID="urn:lsid:plazi:treatment:03AB87FAFFB0FFC1FF7BF905FCDBFB4E" httpUri="http://treatment.plazi.org/id/03AB87FAFFB0FFC1FF7BF905FCDBFB4E" lastPageId="20" lastPageNumber="21" pageId="17" pageNumber="18">
<subSubSection id="C3186567FFB0FFC4FF7BF905FC92F947" box="[151,808,1734,1761]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BBD36ECFFB0FFC4FF7BF905FC92F947" blockId="17.[151,808,1734,1797]" box="[151,808,1734,1761]" pageId="17" pageNumber="18">
<heading id="D0F58180FFB0FFC4FF7BF905FC92F947" bold="true" box="[151,808,1734,1761]" fontSize="11" level="2" pageId="17" pageNumber="18" reason="4">
<taxonomicName id="4C024D6FFFB0FFC4FF7BF905FC92F947" ID-CoL="49ZQD" authority="(Lutken &amp; Mortensen, 1899)" baseAuthorityName="Lutken &amp; Mortensen" baseAuthorityYear="1899" box="[151,808,1734,1761]" class="Ophiuroidea" family="Ophiosphalmidae" genus="Ophiosphalma" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="variabile">
<emphasis id="B976EAFEFFB0FFC4FF7BF905FC92F947" bold="true" box="[151,808,1734,1761]" pageId="17" pageNumber="18">
<emphasis id="B976EAFEFFB0FFC4FF7BF905FE00F946" bold="true" box="[151,442,1734,1761]" italics="true" pageId="17" pageNumber="18">Ophiosphalma variabile</emphasis>
(Lütken &amp;
<bibRefCitation id="EF934B1DFFB0FFC4FDA2F904FCA5F947" author="Mortensen, T." box="[590,799,1734,1761]" pageId="17" pageNumber="18" pagination="93 - 208" refId="ref56796" refString="Lu ¨ tken, C. F. &amp;. Mortensen, T. (1899) The Ophiuridae. Memoirs of the Museum of Comparative Zoology at Harvard College, 23 (2), 93 - 208." type="journal article" year="1899">Mortensen, 1899</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3186567FFB0FFC4FF7BF928FEB6F8A2" box="[151,268,1771,1797]" pageId="17" pageNumber="18" type="description">
<paragraph id="8BBD36ECFFB0FFC4FF7BF928FEB6F8A2" blockId="17.[151,808,1734,1797]" box="[151,268,1771,1797]" pageId="17" pageNumber="18">
<figureCitation id="13392A69FFB0FFC4FF7BF928FEB6F8A2" box="[151,268,1771,1797]" captionStart="FIGURE 3" captionStartId="19.[151,250,1867,1891]" captionTargetBox="[151,1436,181,1843]" captionTargetId="figure-21@19.[151,1436,181,1843]" captionTargetPageId="19" captionText="FIGURE 3. Ophiosphalma jolliense (McClendon, 1909) (ICML-EMU-11059-B, DD = 11 mm).A, Dorsal disc; B, Ventral disc; C, Mouth; D, Dorsal arm; E, Ventral arm; F, Dorsal view. Ophiosphalma variabile (Lütken &amp; Mortensen, 1899) (ICML-EMU- 11220, DD = 12 mm). G, Dorsal disc; H, Ventral disc; I, Mouth; J, Dorsal arm; K, Ventral arm; L, Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7794994" httpUri="https://zenodo.org/record/7794994/files/figure.png" pageId="17" pageNumber="18">Fig. 3G‒L</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3186567FFB0FFC4FF7BF8ECFBACF8C1" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8BBD36ECFFB0FFC4FF7BF8ECFC17F8E0" blockId="17.[151,1046,1839,1894]" box="[151,941,1839,1863]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="AB1211C2FFB0FFC4FF7BF8ECFC17F8E0" box="[151,941,1839,1863]" pageId="17" pageNumber="18">
<treatmentCitation id="0AA310FDFFB0FFC4FF7BF8ECFCB3F8E0" author="Mortensen, T." box="[151,777,1839,1863]" page="137" pageId="17" pageNumber="18" year="1899">
<taxonomicName id="4C024D6FFFB0FFC4FF7BF8ECFCB3F8E0" ID-CoL="49Y4N" authority="Lutken &amp; Mortensen, 1899: 135 - 137" authorityName="Lutken &amp; Mortensen" authorityPageNumber="135 - 137" authorityYear="1899" box="[151,777,1839,1863]" class="Ophiuroidea" family="Ophiosphalmidae" genus="Ophiomusium" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="variabile">
<emphasis id="B976EAFEFFB0FFC4FF7BF8ECFE3DF8E0" box="[151,391,1839,1863]" italics="true" pageId="17" pageNumber="18">Ophiomusium variabile</emphasis>
Lütken &amp;
<bibRefCitation id="EF934B1DFFB0FFC4FE1AF8F3FCB3F8E0" author="Mortensen, T." box="[502,777,1839,1863]" pageId="17" pageNumber="18" pagination="93 - 208" refId="ref56796" refString="Lu ¨ tken, C. F. &amp;. Mortensen, T. (1899) The Ophiuridae. Memoirs of the Museum of Comparative Zoology at Harvard College, 23 (2), 93 - 208." type="journal article" year="1899">Mortensen, 1899: 135‒137</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 4, figs. 4‒6.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBD36ECFFB0FFC4FF7BF88DFBACF8C1" blockId="17.[151,1046,1839,1894]" box="[151,1046,1870,1894]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="AB1211C2FFB0FFC4FF7BF88DFBACF8C1" box="[151,1046,1870,1894]" pageId="17" pageNumber="18">
<taxonomicName id="4C024D6FFFB0FFC4FF7BF88DFE34F8C1" ID-CoL="49ZQD" baseAuthorityName="Lutken &amp; Mortensen" baseAuthorityYear="1899" box="[151,398,1870,1894]" class="Ophiuroidea" family="Ophiosphalmidae" genus="Ophiosphalma" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="variabile">
<emphasis id="B976EAFEFFB0FFC4FF7BF88DFE34F8C1" box="[151,398,1870,1894]" italics="true" pageId="17" pageNumber="18">Ophiosphalma variabile</emphasis>
</taxonomicName>
.
<treatmentCitation id="0AA310FDFFB0FFC4FE77F88DFDE1F8C1" author="Hendler, G." box="[411,603,1870,1894]" page="166" pageId="17" pageNumber="18" year="1996">
<bibRefCitation id="EF934B1DFFB0FFC4FE77F88DFDE1F8C1" author="Hendler, G." box="[411,603,1870,1894]" pageId="17" pageNumber="18" pagination="113 - 179" refId="ref54708" refString="Hendler, G. (1996) Taxonomic atlas of the benthic fauna of the Santa Maria Basin and Western Santa Barbara Channel. Class Ophiuroidea. Santa Barbara Museum of Natural History, 14, 113 - 179." type="journal article" year="1996">Hendler 1996: 166</bibRefCitation>
</treatmentCitation>
, fig. 7.20c‒d.—
<treatmentCitation id="0AA310FDFFB0FFC4FD11F88DFC27F8C1" ID-DOI="http://doi.org/10.5281/zenodo.6056545" ID-Zenodo-Dep="6056545" author="Baker, A. N." box="[765,925,1870,1894]" httpUri="http://treatment.plazi.org/id/03C57E0CA758ED00B0CDFD56FC70A3B8" page="37" pageId="17" pageNumber="18" year="2016">
<bibRefCitation id="EF934B1DFFB0FFC4FD11F88DFC27F8C1" author="Baker, A. N." box="[765,925,1870,1894]" pageId="17" pageNumber="18" pagination="1 - 40" refId="ref52632" refString="Baker, A. N. (2016) An illustrated catalogue of type specimens of the bathyal brittlestar genera Ophiomusium Lyman and Ophiosphalma H. L. Clark (Echinodermata: Ophiuroidea). Zootaxa, 4097 (1), 1 - 40. https: // doi. org / 10.11646 / zootaxa. 4097.1.1" type="journal article" year="2016">Baker 2016: 37</bibRefCitation>
</treatmentCitation>
, fig. 61a‒b.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3186567FFB0FFC1FF7BF857FDFAFEE6" lastPageId="20" lastPageNumber="21" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BBD36ECFFB0FFC1FF7BF857FB49FF5E" blockId="17.[151,1437,1940,2039]" lastBlockId="20.[151,1443,150,1257]" lastPageId="20" lastPageNumber="21" pageId="17" pageNumber="18">
<emphasis id="B976EAFEFFB0FFC4FF7BF857FEC6F809" bold="true" box="[151,380,1940,1966]" pageId="17" pageNumber="18">Material examined</emphasis>
. 10,938 individuals at eight stations.
<collectionCode id="ED13AE29FFB0FFC4FCDCF857FC0FF808" box="[816,949,1940,1967]" pageId="17" pageNumber="18">TALUD III</collectionCode>
, Sta. 6, 166 ind. (ICML-EMU-9016-A), 2 ind. (ICML-EMU-9016-B), and 3 ind. (ICML-EMU-11067-A).
<collectionCode id="ED13AE29FFB0FFC4FC9AF87BFC40F874" box="[886,1018,1976,2003]" pageId="17" pageNumber="18">TALUD IV</collectionCode>
, Sta. 3, 2 ind. (ICML-EMU-11116).
<collectionCode id="ED13AE29FFB0FFC4FF7BF81FFF4EF851" box="[151,244,2012,2038]" pageId="17" pageNumber="18">TALUD</collectionCode>
<collectionCode id="ED13AE29FFB0FFC4FF15F81EFEB7F850" box="[249,269,2013,2039]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="17" pageNumber="18" type="Museum">V</collectionCode>
, Sta. 3, 8 ind. (ICML-EMU-11067-B).
<collectionCode id="ED13AE29FFB0FFC4FD53F81FFCF8F850" box="[703,834,2012,2039]" pageId="17" pageNumber="18">TALUD IX</collectionCode>
, Sta. 10, 1 ind. (ICML-EMU-11068-B); Sta. 17, 1 ind. (ICML-EMU-11068-A).
<collectionCode id="ED13AE29FFB5FFC1FE5AFF54FDFDFF16" box="[438,583,151,177]" pageId="20" pageNumber="21">TALUD XV</collectionCode>
, Sta. 5D, 161 ind. (ICML-EMU-11070); Sta. 20, 20 ind. (ICML-EMU-11071), 4 ind. (ICML-EMU-11685), and 1 ind. (ICML-EMU-11948); Sta. 23, 1 ind. (ICML-EMU-11069-A), 43 ind. (ICML-EMU-11069-B), 5,862 ind. (ICML-EMU-11220), and 4,663 ind. (ICML-EMU-11221).
</paragraph>
<caption id="DF7D6664FFB3FFC7FF7BF888FE05F868" ID-DOI="http://doi.org/10.5281/zenodo.7794992" ID-Zenodo-Dep="7794992" httpUri="https://zenodo.org/record/7794992/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,250,1867,1891]" targetBox="[151,1436,181,1843]" targetPageId="18">
<paragraph id="8BBD36ECFFB3FFC7FF7BF888FE05F868" blockId="18.[151,1436,1867,1999]" pageId="18" pageNumber="19">
<emphasis id="B976EAFEFFB3FFC7FF7BF888FEAEF8C4" bold="true" box="[151,276,1867,1891]" pageId="18" pageNumber="19">FIGURE 2.</emphasis>
<taxonomicName id="4C024D6FFFB3FFC7FEF7F888FCBAF8C4" authority="(Lutken &amp; Mortensen, 1899)" baseAuthorityName="Lutken &amp; Mortensen" baseAuthorityYear="1899" box="[283,768,1867,1891]" class="Ophiuroidea" family="Asteronychidae" genus="Astrodia" kingdom="Animalia" order="Euryalida" pageId="18" pageNumber="19" phylum="Echinodermata" rank="species" species="excavata">
<emphasis id="B976EAFEFFB3FFC7FEF7F888FE68F8C4" box="[283,466,1867,1891]" italics="true" pageId="18" pageNumber="19">Astrodia excavata</emphasis>
(Lütken &amp;
<bibRefCitation id="EF934B1DFFB3FFC7FDA7F888FD43F8C4" author="Mortensen, T." box="[587,761,1867,1891]" pageId="18" pageNumber="19" pagination="93 - 208" refId="ref56796" refString="Lu ¨ tken, C. F. &amp;. Mortensen, T. (1899) The Ophiuridae. Memoirs of the Museum of Comparative Zoology at Harvard College, 23 (2), 93 - 208." type="journal article" year="1899">Mortensen, 1899</bibRefCitation>
)
</taxonomicName>
(ICML-EMU-11099, DD = 24.7 mm). A, Dorsal disc; B, Ventral disc; C, Mouth; D, Dorsal arm; E, Proximalmost ventral arm; F, Middle ventral arm; G, Dorsal view.
<taxonomicName id="4C024D6FFFB3FFC7FB46F8ACFE04F80C" authority="(Lutken &amp; Mortensen, 1899)" baseAuthorityName="Lutken &amp; Mortensen" baseAuthorityYear="1899" class="Ophiuroidea" family="Ophiosphalmidae" genus="Ophiosphalma" kingdom="Animalia" order="Ophiurida" pageId="18" pageNumber="19" phylum="Echinodermata" rank="species" species="glabrum">
<emphasis id="B976EAFEFFB3FFC7FB46F8ACFA26F820" box="[1194,1436,1903,1927]" italics="true" pageId="18" pageNumber="19">Ophiosphalma glabrum</emphasis>
(Lütken &amp;
<bibRefCitation id="EF934B1DFFB3FFC7FEE5F850FE0DF80C" author="Mortensen, T." box="[265,439,1939,1963]" pageId="18" pageNumber="19" pagination="93 - 208" refId="ref56796" refString="Lu ¨ tken, C. F. &amp;. Mortensen, T. (1899) The Ophiuridae. Memoirs of the Museum of Comparative Zoology at Harvard College, 23 (2), 93 - 208." type="journal article" year="1899">Mortensen, 1899</bibRefCitation>
)
</taxonomicName>
(ICML-EMU-11061, DD = 31 mm). H, Dorsal disc; I, Ventral disc; J, Mouth; K, Dorsal arm; L, Ventral arm; M, Dorsal view.
</paragraph>
</caption>
<caption id="DF7D6664FFB2FFC6FF7BF888FB46F80C" ID-DOI="http://doi.org/10.5281/zenodo.7794994" ID-Zenodo-Dep="7794994" httpUri="https://zenodo.org/record/7794994/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1867,1891]" targetBox="[151,1436,181,1843]" targetPageId="19">
<paragraph id="8BBD36ECFFB2FFC6FF7BF888FB46F80C" blockId="19.[151,1436,1867,1963]" pageId="19" pageNumber="20">
<emphasis id="B976EAFEFFB2FFC6FF7BF888FEA8F8C4" bold="true" box="[151,274,1867,1891]" pageId="19" pageNumber="20">FIGURE 3.</emphasis>
<taxonomicName id="4C024D6FFFB2FFC6FEF4F888FD6FF8C4" authority="(McClendon, 1909)" baseAuthorityName="McClendon" baseAuthorityYear="1909" box="[280,725,1867,1891]" class="Ophiuroidea" family="Ophiosphalmidae" genus="Ophiosphalma" kingdom="Animalia" order="Ophiurida" pageId="19" pageNumber="20" phylum="Echinodermata" rank="species" species="jolliense">
<emphasis id="B976EAFEFFB2FFC6FEF4F888FDB2F8C4" box="[280,520,1867,1891]" italics="true" pageId="19" pageNumber="20">Ophiosphalma jolliense</emphasis>
(
<bibRefCitation id="EF934B1DFFB2FFC6FDF9F888FD74F8C4" author="McClendon, J. F." box="[533,718,1867,1891]" pageId="19" pageNumber="20" pagination="33 - 64" refId="ref57681" refString="McClendon, J. F. (1909) The ophiurans of the San Diego region. University of California Publications in Zoology, 6 (3), 33 - 64." type="journal article" year="1909">McClendon, 1909</bibRefCitation>
)
</taxonomicName>
(ICML-EMU-11059-B, DD = 11 mm). A, Dorsal disc; B, Ventral disc; C, Mouth; D, Dorsal arm; E, Ventral arm; F, Dorsal view.
<taxonomicName id="4C024D6FFFB2FFC6FD0FF8ACFABFF820" authority="(Lutken &amp; Mortensen, 1899)" baseAuthorityName="Lutken &amp; Mortensen" baseAuthorityYear="1899" box="[739,1285,1903,1927]" class="Ophiuroidea" family="Ophiosphalmidae" genus="Ophiosphalma" kingdom="Animalia" order="Ophiurida" pageId="19" pageNumber="20" phylum="Echinodermata" rank="species" species="variabile">
<emphasis id="B976EAFEFFB2FFC6FD0FF8ACFC63F820" box="[739,985,1903,1927]" italics="true" pageId="19" pageNumber="20">Ophiosphalma variabile</emphasis>
(Lütken &amp;
<bibRefCitation id="EF934B1DFFB2FFC6FBBDF8ACFB44F820" author="Mortensen, T." box="[1105,1278,1903,1927]" pageId="19" pageNumber="20" pagination="93 - 208" refId="ref56796" refString="Lu ¨ tken, C. F. &amp;. Mortensen, T. (1899) The Ophiuridae. Memoirs of the Museum of Comparative Zoology at Harvard College, 23 (2), 93 - 208." type="journal article" year="1899">Mortensen, 1899</bibRefCitation>
)
</taxonomicName>
(ICML-EMU- 11220, DD = 12 mm). G, Dorsal disc; H, Ventral disc; I, Mouth; J, Dorsal arm; K, Ventral arm; L, Dorsal view.
</paragraph>
</caption>
<paragraph id="8BBD36ECFFB5FFC1FF2BFEC1FDFAFEE6" blockId="20.[151,1443,150,1257]" pageId="20" pageNumber="21">
<emphasis id="B976EAFEFFB5FFC1FF2BFEC1FE6DFEBB" bold="true" box="[199,471,258,284]" pageId="20" pageNumber="21">Comparative material.</emphasis>
<taxonomicName id="4C024D6FFFB5FFC1FE31FEC0FBA5FEBA" authority="Lutken &amp; Mortensen, 1899" authorityName="Lutken &amp; Mortensen" authorityYear="1899" box="[477,1055,259,285]" class="Ophiuroidea" family="Ophiosphalmidae" genus="Ophiomusium" kingdom="Animalia" order="Ophiurida" pageId="20" pageNumber="21" phylum="Echinodermata" rank="species" species="variabile">
<emphasis id="B976EAFEFFB5FFC1FE31FEC0FD5CFEBA" box="[477,742,259,285]" italics="true" pageId="20" pageNumber="21">Ophiomusium variabile</emphasis>
<materialsCitation id="3B6A3CB1FFB5FFC1FD01FEC0FBA5FEBA" ID-GBIF-Occurrence="4071433324" box="[749,1055,259,285]" collectionCode="MCZ" pageId="20" pageNumber="21" specimenCode="OPH-507, USNM 19502" specimenCount="1" typeStatus="syntype">
Lütken &amp;
<bibRefCitation id="EF934B1DFFB5FFC1FCB3FEC0FBA5FEBA" author="Mortensen, T." box="[863,1055,259,285]" pageId="20" pageNumber="21" pagination="93 - 208" refId="ref56796" refString="Lu ¨ tken, C. F. &amp;. Mortensen, T. (1899) The Ophiuridae. Memoirs of the Museum of Comparative Zoology at Harvard College, 23 (2), 93 - 208." type="journal article" year="1899">Mortensen, 1899</bibRefCitation>
</materialsCitation>
</taxonomicName>
,
<typeStatus id="54B9884EFFB5FFC1FBC6FEC0FB35FEBA" box="[1066,1167,259,285]" pageId="20" pageNumber="21" type="syntype">syntypes</typeStatus>
, 6 ind.:
<collectionCode id="ED13AE29FFB5FFC1FB0BFEC0FA9EFEBA" box="[1255,1316,259,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="20" pageNumber="21" type="Museum">MCZ</collectionCode>
<specimenCode id="DBA49E97FFB5FFC1FAC6FEC1FA22FEBA" box="[1322,1432,258,285]" collectionCode="OPH-" pageId="20" pageNumber="21">OPH-507</specimenCode>
,
<specimenCode id="DBA49E97FFB5FFC1FF7BFEE4FE8FFEE6" box="[151,309,294,321]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM 19502</specimenCode>
(Supplementary file 2).
</paragraph>
</subSubSection>
<subSubSection id="C3186567FFB5FFC1FF2BFE89FCABFC6E" pageId="20" pageNumber="21" type="description">
<paragraph id="8BBD36ECFFB5FFC1FF2BFE89FB58FC9E" blockId="20.[151,1443,150,1257]" pageId="20" pageNumber="21">
<emphasis id="B976EAFEFFB5FFC1FF2BFE89FEEBFEC3" bold="true" box="[199,337,330,356]" pageId="20" pageNumber="21">Description</emphasis>
(ICML-EMU-11220). DD =
<quantity id="4CFA9B09FFB5FFC1FD5CFE88FCBDFEC2" box="[688,775,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="21" unit="mm" value="12.0">12 mm</quantity>
. Whole-body with weakly tuberculous appearance. Disc pentagonal, flat. Dorsal disc covered with imbricated, irregular size scales. Primary plates not evident. RS longer than broad, oval, separated by several elongated scales; several elongated oval scales between each pair of RS, covering ½ of disc radius (
<figureCitation id="13392A69FFB5FFC1FEB3FE74FE01FE76" box="[351,443,439,465]" captionStart="FIGURE 3" captionStartId="19.[151,250,1867,1891]" captionTargetBox="[151,1436,181,1843]" captionTargetId="figure-21@19.[151,1436,181,1843]" captionTargetPageId="19" captionText="FIGURE 3. Ophiosphalma jolliense (McClendon, 1909) (ICML-EMU-11059-B, DD = 11 mm).A, Dorsal disc; B, Ventral disc; C, Mouth; D, Dorsal arm; E, Ventral arm; F, Dorsal view. Ophiosphalma variabile (Lütken &amp; Mortensen, 1899) (ICML-EMU- 11220, DD = 12 mm). G, Dorsal disc; H, Ventral disc; I, Mouth; J, Dorsal arm; K, Ventral arm; L, Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7794994" httpUri="https://zenodo.org/record/7794994/files/figure.png" pageId="20" pageNumber="21">Fig. 3G</figureCitation>
). Ventral interradii covered by oval imbricated scales, larger than dorsal scales. Genital slits with numerous minute quadrangular papillae (
<figureCitation id="13392A69FFB5FFC1FD25FE18FC98FE52" box="[713,802,475,501]" captionStart="FIGURE 3" captionStartId="19.[151,250,1867,1891]" captionTargetBox="[151,1436,181,1843]" captionTargetId="figure-21@19.[151,1436,181,1843]" captionTargetPageId="19" captionText="FIGURE 3. Ophiosphalma jolliense (McClendon, 1909) (ICML-EMU-11059-B, DD = 11 mm).A, Dorsal disc; B, Ventral disc; C, Mouth; D, Dorsal arm; E, Ventral arm; F, Dorsal view. Ophiosphalma variabile (Lütken &amp; Mortensen, 1899) (ICML-EMU- 11220, DD = 12 mm). G, Dorsal disc; H, Ventral disc; I, Mouth; J, Dorsal arm; K, Ventral arm; L, Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7794994" httpUri="https://zenodo.org/record/7794994/files/figure.png" pageId="20" pageNumber="21">Fig. 3H</figureCitation>
). OSh longer than broad, rhombic with a distal lobe, right angles. Madreporite slightly larger than OSh. AdSh longer than broad, elongated, quadrangular, almost meeting or meeting in front of OSh. Jaws bearing 6‒7 oral papillae at each side, quadrangular, merging, two distalmost the largest; one pointed papilla at jaw tips (
<figureCitation id="13392A69FFB5FFC1FD92FD84FD77FDC6" box="[638,717,583,609]" captionStart="FIGURE 3" captionStartId="19.[151,250,1867,1891]" captionTargetBox="[151,1436,181,1843]" captionTargetId="figure-21@19.[151,1436,181,1843]" captionTargetPageId="19" captionText="FIGURE 3. Ophiosphalma jolliense (McClendon, 1909) (ICML-EMU-11059-B, DD = 11 mm).A, Dorsal disc; B, Ventral disc; C, Mouth; D, Dorsal arm; E, Ventral arm; F, Dorsal view. Ophiosphalma variabile (Lütken &amp; Mortensen, 1899) (ICML-EMU- 11220, DD = 12 mm). G, Dorsal disc; H, Ventral disc; I, Mouth; J, Dorsal arm; K, Ventral arm; L, Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7794994" httpUri="https://zenodo.org/record/7794994/files/figure.png" pageId="20" pageNumber="21">Fig. 3I</figureCitation>
). Arms thin, gradually narrowing distally. Dorsal arm base with 3‒4 overlapping DAP, several granules at edges of arm base, projecting ventrally. DAP slightly longer than broad, rhombic, separating and decreasing in size distally (
<figureCitation id="13392A69FFB5FFC1FD02FD4CFCFBFD0E" box="[750,833,655,681]" captionStart="FIGURE 3" captionStartId="19.[151,250,1867,1891]" captionTargetBox="[151,1436,181,1843]" captionTargetId="figure-21@19.[151,1436,181,1843]" captionTargetPageId="19" captionText="FIGURE 3. Ophiosphalma jolliense (McClendon, 1909) (ICML-EMU-11059-B, DD = 11 mm).A, Dorsal disc; B, Ventral disc; C, Mouth; D, Dorsal arm; E, Ventral arm; F, Dorsal view. Ophiosphalma variabile (Lütken &amp; Mortensen, 1899) (ICML-EMU- 11220, DD = 12 mm). G, Dorsal disc; H, Ventral disc; I, Mouth; J, Dorsal arm; K, Ventral arm; L, Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7794994" httpUri="https://zenodo.org/record/7794994/files/figure.png" pageId="20" pageNumber="21">Fig. 3J</figureCitation>
). First VAP smaller than rest; subsequent three VAP the largest (almost covering all arm segment length), separated from each other, the only ones with tentacle pores; posterior VAP triangular, reduced in size (approximately 1/3 LAP in length), separated from each other. First LAP with two ArSp, separated; subsequent LAP with three ArSp. Three tentacle pores with one rounded TSc covering pore almost completely (
<figureCitation id="13392A69FFB5FFC1FE41FCDCFDBDFC9E" box="[429,519,799,825]" captionStart="FIGURE 3" captionStartId="19.[151,250,1867,1891]" captionTargetBox="[151,1436,181,1843]" captionTargetId="figure-21@19.[151,1436,181,1843]" captionTargetPageId="19" captionText="FIGURE 3. Ophiosphalma jolliense (McClendon, 1909) (ICML-EMU-11059-B, DD = 11 mm).A, Dorsal disc; B, Ventral disc; C, Mouth; D, Dorsal arm; E, Ventral arm; F, Dorsal view. Ophiosphalma variabile (Lütken &amp; Mortensen, 1899) (ICML-EMU- 11220, DD = 12 mm). G, Dorsal disc; H, Ventral disc; I, Mouth; J, Dorsal arm; K, Ventral arm; L, Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7794994" httpUri="https://zenodo.org/record/7794994/files/figure.png" pageId="20" pageNumber="21">Fig. 3K</figureCitation>
). Color pattern beige-whitish (ethanol preservation) (
<figureCitation id="13392A69FFB5FFC1FBB0FCDCFB6CFC9E" box="[1116,1238,799,825]" captionStart="FIGURE 3" captionStartId="19.[151,250,1867,1891]" captionTargetBox="[151,1436,181,1843]" captionTargetId="figure-21@19.[151,1436,181,1843]" captionTargetPageId="19" captionText="FIGURE 3. Ophiosphalma jolliense (McClendon, 1909) (ICML-EMU-11059-B, DD = 11 mm).A, Dorsal disc; B, Ventral disc; C, Mouth; D, Dorsal arm; E, Ventral arm; F, Dorsal view. Ophiosphalma variabile (Lütken &amp; Mortensen, 1899) (ICML-EMU- 11220, DD = 12 mm). G, Dorsal disc; H, Ventral disc; I, Mouth; J, Dorsal arm; K, Ventral arm; L, Dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7794994" httpUri="https://zenodo.org/record/7794994/files/figure.png" pageId="20" pageNumber="21">Fig. 3G‒L</figureCitation>
).
</paragraph>
<paragraph id="8BBD36ECFFB5FFC1FF2BFC81FCABFC6E" blockId="20.[151,1443,150,1257]" pageId="20" pageNumber="21">
<emphasis id="B976EAFEFFB5FFC1FF2BFC81FE46FCFB" bold="true" box="[199,508,834,861]" pageId="20" pageNumber="21">Habitat and distribution.</emphasis>
Mexican Pacific, from western
<collectingRegion id="49C6F80EFFB5FFC1FC96FC80FBE7FCFA" box="[890,1117,835,861]" country="Mexico" name="Baja California Sur" pageId="20" pageNumber="21">Baja California Sur</collectingRegion>
to
<collectingRegion id="49C6F80EFFB5FFC1FB66FC80FB49FCFA" box="[1162,1267,835,861]" country="Mexico" name="Guerrero" pageId="20" pageNumber="21">Guerrero</collectingRegion>
; 267‒
<emphasis id="B976EAFEFFB5FFC1FAD6FC81FA26FCFA" bold="true" box="[1338,1436,834,861]" pageId="20" pageNumber="21">
<quantity id="4CFA9B09FFB5FFC1FAD6FC81FA26FCFA" box="[1338,1436,834,861]" metricMagnitude="0" metricUnit="m" metricValue="1.148" pageId="20" pageNumber="21" unit="m" value="1.148">1,148 m</quantity>
</emphasis>
depth, muddy, sandy and rocky substrates (Lütken &amp;
<bibRefCitation id="EF934B1DFFB5FFC1FD1DFCA4FC16FC26" author="Mortensen, T." box="[753,940,871,897]" pageId="20" pageNumber="21" pagination="93 - 208" refId="ref56796" refString="Lu ¨ tken, C. F. &amp;. Mortensen, T. (1899) The Ophiuridae. Memoirs of the Museum of Comparative Zoology at Harvard College, 23 (2), 93 - 208." type="journal article" year="1899">Mortensen 1899</bibRefCitation>
;
<bibRefCitation id="EF934B1DFFB5FFC1FC54FCA4FB85FC26" author="Maluf, L. Y." box="[952,1087,871,897]" pageId="20" pageNumber="21" pagination="1 - 306" refId="ref57226" refString="Maluf, L. Y. (1988) Composition and distribution of the central Eastern Pacific Echinoderms. Natural History Museum of Los Angeles County, Technical Report, 2, 1 - 306." type="journal article" year="1988">Maluf 1988</bibRefCitation>
;
<bibRefCitation id="EF934B1DFFB5FFC1FBA7FCA4FA2DFC26" author="Granja-Fernandez, R. &amp; Herrero-Perezrul, M. D. &amp; Lopez-Perez, R. A. &amp; Hernandez-Morales, A. &amp; Rangel-Solis, P. D." box="[1099,1431,870,897]" pageId="20" pageNumber="21" pagination="37 - 47" refId="ref54478" refString="Granja-Fernandez, R., Herrero-Perezrul, M. D., Lopez-Perez, R. A., Hernandez-Morales, A. &amp; Rangel-Solis, P. D. (2015) A literature review of the Ophiuroidea (Echinodermata) from the Pacific coast of Mexico. Revista de Biologia Tropical, 63 (2), 37 - 47." type="journal article" year="2015">
Granja-Fernández
<emphasis id="B976EAFEFFB5FFC1FAF1FCA4FAEFFC26" box="[1309,1365,871,897]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2015
</bibRefCitation>
; see Remarks). The material examined was collected
<emphasis id="B976EAFEFFB5FFC1FD16FC49FB3AFC03" bold="true" box="[762,1152,906,933]" pageId="20" pageNumber="21">
off western
<collectingRegion id="49C6F80EFFB5FFC1FC60FC49FB3AFC03" box="[908,1152,906,932]" country="Mexico" name="Baja California Sur" pageId="20" pageNumber="21">Baja California Sur</collectingRegion>
</emphasis>
, in the southern Gulf of
<collectingRegion id="49C6F80EFFB5FFC1FF7BFC6CFEB1FC6E" box="[151,267,943,969]" country="United States of America" name="California" pageId="20" pageNumber="21">California</collectingRegion>
, and near Marias Islands; 530‒
<quantity id="4CFA9B09FFB5FFC1FD86FC6DFD7EFC6E" box="[618,708,942,969]" metricMagnitude="0" metricUnit="m" metricValue="1.148" pageId="20" pageNumber="21" unit="m" value="1.148">1,148 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="C3186567FFB5FFC1FF2BFC10FCDBFB4E" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8BBD36ECFFB5FFC1FF2BFC10FCDBFB4E" blockId="20.[151,1443,150,1257]" pageId="20" pageNumber="21">
<emphasis id="B976EAFEFFB5FFC1FF2BFC10FE81FC4A" bold="true" box="[199,315,979,1005]" pageId="20" pageNumber="21">Remarks.</emphasis>
<taxonomicName id="4C024D6FFFB5FFC1FEA9FC10FDE3FC4A" baseAuthorityName="Lutken &amp; Mortensen" baseAuthorityYear="1899" box="[325,601,979,1005]" class="Ophiuroidea" family="Ophiosphalmidae" genus="Ophiosphalma" kingdom="Animalia" order="Ophiurida" pageId="20" pageNumber="21" phylum="Echinodermata" rank="species" species="variabile">
<emphasis id="B976EAFEFFB5FFC1FEA9FC10FDE3FC4A" box="[325,601,979,1005]" italics="true" pageId="20" pageNumber="21">Ophiosphalma variabile</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C024D6FFFB5FFC1FD1AFC10FCC1FC4A" baseAuthorityName="McClendon" baseAuthorityYear="1909" box="[758,891,979,1005]" class="Ophiuroidea" family="Ophiosphalmidae" genus="Ophiosphalma" kingdom="Animalia" order="Ophiurida" pageId="20" pageNumber="21" phylum="Echinodermata" rank="species" species="jolliense">
<emphasis id="B976EAFEFFB5FFC1FD1AFC10FCB0FC4A" box="[758,778,979,1005]" italics="true" pageId="20" pageNumber="21">O</emphasis>
.
<emphasis id="B976EAFEFFB5FFC1FCF0FC10FCC1FC4A" box="[796,891,979,1005]" italics="true" pageId="20" pageNumber="21">jolliense</emphasis>
</taxonomicName>
in overall appearance. However, differences in their morphology (e.g., radial shields size and lateral arm plates shape) are provided by
<bibRefCitation id="EF934B1DFFB5FFC1FB82FC34FAA7FBB6" author="Hendler, G." box="[1134,1309,1015,1041]" pageId="20" pageNumber="21" pagination="113 - 179" refId="ref54708" refString="Hendler, G. (1996) Taxonomic atlas of the benthic fauna of the Santa Maria Basin and Western Santa Barbara Channel. Class Ophiuroidea. Santa Barbara Museum of Natural History, 14, 113 - 179." type="journal article" year="1996">Hendler (1996)</bibRefCitation>
. Moreover, they can be distinguished by: 1) the primary plates are clearly visible in
<taxonomicName id="4C024D6FFFB5FFC1FC29FBD8FBFCFB92" baseAuthorityName="McClendon" baseAuthorityYear="1909" box="[965,1094,1051,1077]" class="Ophiuroidea" family="Ophiosphalmidae" genus="Ophiosphalma" kingdom="Animalia" order="Ophiurida" pageId="20" pageNumber="21" phylum="Echinodermata" rank="species" species="jolliense">
<emphasis id="B976EAFEFFB5FFC1FC29FBD8FC63FB92" box="[965,985,1051,1077]" italics="true" pageId="20" pageNumber="21">O</emphasis>
.
<emphasis id="B976EAFEFFB5FFC1FC0BFBD8FBFCFB92" box="[999,1094,1051,1077]" italics="true" pageId="20" pageNumber="21">jolliense</emphasis>
</taxonomicName>
vs. not so in
<taxonomicName id="4C024D6FFFB5FFC1FB32FBD8FADFFB92" baseAuthorityName="Lutken &amp; Mortensen" baseAuthorityYear="1899" box="[1246,1381,1051,1077]" class="Ophiuroidea" family="Ophiosphalmidae" genus="Ophiosphalma" kingdom="Animalia" order="Ophiurida" pageId="20" pageNumber="21" phylum="Echinodermata" rank="species" species="variabile">
<emphasis id="B976EAFEFFB5FFC1FB32FBD8FB48FB92" box="[1246,1266,1051,1077]" italics="true" pageId="20" pageNumber="21">O</emphasis>
.
<emphasis id="B976EAFEFFB5FFC1FAECFBD8FADFFB92" box="[1280,1381,1051,1077]" italics="true" pageId="20" pageNumber="21">variabile</emphasis>
</taxonomicName>
, and 2) spearhead-shaped with rounded edges oral shields in
<taxonomicName id="4C024D6FFFB5FFC1FCEEFBFCFC38FBFE" baseAuthorityName="McClendon" baseAuthorityYear="1909" box="[770,898,1087,1113]" class="Ophiuroidea" family="Ophiosphalmidae" genus="Ophiosphalma" kingdom="Animalia" order="Ophiurida" pageId="20" pageNumber="21" phylum="Echinodermata" rank="species" species="jolliense">
<emphasis id="B976EAFEFFB5FFC1FCEEFBFCFCACFBFE" box="[770,790,1087,1113]" italics="true" pageId="20" pageNumber="21">O</emphasis>
.
<emphasis id="B976EAFEFFB5FFC1FCCFFBFCFC38FBFE" box="[803,898,1087,1113]" italics="true" pageId="20" pageNumber="21">jolliense</emphasis>
</taxonomicName>
vs. rhombic-shaped with right angle oral shields in
<taxonomicName id="4C024D6FFFB5FFC1FF58FBA0FE86FBDA" baseAuthorityName="Lutken &amp; Mortensen" baseAuthorityYear="1899" box="[180,316,1123,1149]" class="Ophiuroidea" family="Ophiosphalmidae" genus="Ophiosphalma" kingdom="Animalia" order="Ophiurida" pageId="20" pageNumber="21" phylum="Echinodermata" rank="species" species="variabile">
<emphasis id="B976EAFEFFB5FFC1FF58FBA0FF72FBDA" box="[180,200,1123,1149]" italics="true" pageId="20" pageNumber="21">O</emphasis>
.
<emphasis id="B976EAFEFFB5FFC1FF3BFBA0FE86FBDA" box="[215,316,1123,1149]" italics="true" pageId="20" pageNumber="21">variabile</emphasis>
</taxonomicName>
. The previous northernmost distribution limit of
<taxonomicName id="4C024D6FFFB5FFC1FC8FFBA0FC51FBDA" baseAuthorityName="Lutken &amp; Mortensen" baseAuthorityYear="1899" box="[867,1003,1123,1149]" class="Ophiuroidea" family="Ophiosphalmidae" genus="Ophiosphalma" kingdom="Animalia" order="Ophiurida" pageId="20" pageNumber="21" phylum="Echinodermata" rank="species" species="variabile">
<emphasis id="B976EAFEFFB5FFC1FC8FFBA0FC51FBDA" box="[867,1003,1123,1149]" italics="true" pageId="20" pageNumber="21">O. variabile</emphasis>
</taxonomicName>
was in the Gulf of
<collectingRegion id="49C6F80EFFB5FFC1FB29FBA0FA8DFBDA" box="[1221,1335,1123,1149]" country="United States of America" name="California" pageId="20" pageNumber="21">California</collectingRegion>
(GranjaFernández
<emphasis id="B976EAFEFFB5FFC1FEFEFB44FEF3FB06" box="[274,329,1159,1185]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2015). Its presence off western
<collectingRegion id="49C6F80EFFB5FFC1FD44FB44FC3BFB06" box="[680,897,1159,1185]" country="Mexico" name="Baja California Sur" pageId="20" pageNumber="21">Baja California Sur</collectingRegion>
, as reported herein, represents a new distribution record in the Mexican Pacific as well as a general distribution range extension to the north (to 27ºN). The deepest known record (i.e.,
<quantity id="4CFA9B09FFB5FFC1FE9EFB0CFE06FB4E" box="[370,444,1231,1257]" metricMagnitude="2" metricUnit="m" metricValue="9.02" pageId="20" pageNumber="21" unit="m" value="902.0">902 m</quantity>
;
<bibRefCitation id="EF934B1DFFB5FFC1FE24FB0CFDF6FB4E" author="Maluf, L. Y." box="[456,588,1231,1257]" pageId="20" pageNumber="21" pagination="1 - 306" refId="ref57226" refString="Maluf, L. Y. (1988) Composition and distribution of the central Eastern Pacific Echinoderms. Natural History Museum of Los Angeles County, Technical Report, 2, 1 - 306." type="journal article" year="1988">Maluf 1988</bibRefCitation>
) is extended to
<quantity id="4CFA9B09FFB5FFC1FD12FB0CFCE7FB4E" box="[766,861,1231,1257]" metricMagnitude="0" metricUnit="m" metricValue="1.148" pageId="20" pageNumber="21" unit="m" value="1.148">1,148 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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