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<document id="303D404EC3C51B09C8A5BA0896059318" ID-DOI="10.5252/cryptogamie-bryologie2022v43a2" ID-ISSN="1776-0992" ID-Zenodo-Dep="12215089" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719003595154" checkinUser="felipe" docAuthor="Bruggeman-Nannenga, Maria Alida" docDate="2022" docId="03AB87C7FF94FFB552A5F982DD8CF870" docLanguage="en" docName="CryptogamieBryologie.43.2.9-36.pdf" docOrigin="Cryptogamie, Bryologie 20 (2)" docSource="http://dx.doi.org/10.5252/cryptogamie-bryologie2022v43a2" docStyle="DocumentStyle:3F4A000F5B047ABA0EAB2943AA9373AE.3:CryptogamieBryologie.2019-.journal_article" docStyleId="3F4A000F5B047ABA0EAB2943AA9373AE" docStyleName="CryptogamieBryologie.2019-.journal_article" docStyleVersion="3" docTitle="Fissidens ah-pengae Brugg." docType="treatment" docVersion="2" lastPageNumber="13" masterDocId="FF92FFBFFF90FFB35110FFCEDE26FF9B" masterDocTitle="On the peristomes of the corticolous African species of Fissidens Hedw. (Fissidentaceae, Bryophyta)" masterLastPageNumber="36" masterPageNumber="9" pageNumber="11" updateTime="1719018102003" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="99F36A2314C7744AED6378A759F142CB">On the peristomes of the corticolous African species of Fissidens Hedw. (Fissidentaceae, Bryophyta)</mods:title>
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<mods:namePart id="AAC10084CB8AB1AEEDCDA077D2D607FD">Bruggeman-Nannenga, Maria Alida</mods:namePart>
<mods:affiliation id="6BA165FF90A2DB02437FED199D21B87D">Griffensteijnseplein 23 NL- 3703 BE Zeist (The Netherlands)</mods:affiliation>
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<mods:title id="0F91BBFD27FE56FFD9D024D868187B32">Cryptogamie, Bryologie</mods:title>
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<mods:date id="A2B8A7E251C189F49055C59205988047">2022</mods:date>
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<mods:identifier id="989474D74F27406BD4385774239862E4" type="DOI">10.5252/cryptogamie-bryologie2022v43a2</mods:identifier>
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<treatment id="03AB87C7FF94FFB552A5F982DD8CF870" ID-DOI="http://doi.org/10.5281/zenodo.12215105" ID-Zenodo-Dep="12215105" LSID="urn:lsid:plazi:treatment:03AB87C7FF94FFB552A5F982DD8CF870" httpUri="http://treatment.plazi.org/id/03AB87C7FF94FFB552A5F982DD8CF870" lastPageId="6" lastPageNumber="13" pageId="4" pageNumber="11">
<subSubSection id="C318655AFF94FFB752A5F982DB01F9FC" box="[949,1319,1612,1639]" pageId="4" pageNumber="11" type="nomenclature">
<paragraph id="8BBD36D1FF94FFB752A5F982DB01F9FC" blockId="4.[949,1319,1612,1639]" box="[949,1319,1612,1639]" pageId="4" pageNumber="11">
<heading id="D0F581BDFF94FFB752A5F982DB01F9FC" box="[949,1319,1612,1639]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="11" reason="4">
<taxonomicName id="4C024D52FF94FFB752A5F982DAFAF9FC" authority="Brugg." authorityName="Brugg." box="[949,1244,1612,1639]" class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="4" pageNumber="11" phylum="Bryophyta" rank="species" species="ah-pengae">
<emphasis id="B976EAC3FF94FFB752A5F982DAAEF9FC" bold="true" box="[949,1160,1612,1639]" italics="true" pageId="4" pageNumber="11">Fissidens ah-pengae</emphasis>
Brugg.
</taxonomicName>
-Nann.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C318655AFF94FFB55556F9A3DA22F977" lastPageId="6" lastPageNumber="13" pageId="4" pageNumber="11" type="description">
<paragraph id="8BBD36D1FF94FFB75556F9A3DAB0F91C" blockId="4.[1094,1174,1645,1671]" box="[1094,1174,1645,1671]" pageId="4" pageNumber="11">
<heading id="D0F581BDFF94FFB75556F9A3DAB0F91C" box="[1094,1174,1645,1671]" centered="true" fontSize="11" level="3" pageId="4" pageNumber="11" reason="3">
(
<figureCitation id="13392A54FF94FFB7555EF9A3DAABF91C" box="[1102,1165,1645,1671]" captionStart="FIG" captionStartId="6.[132,143,1444,1462]" captionTargetBox="[279,1307,215,1399]" captionTargetId="figure-205@6.[278,841,761,1310]" captionTargetPageId="6" captionText="FIG. 2. —Fissidens ah-pengae Brugg.-Nann. A, OPL with focus on the thin trabeculae;B, OPL focus on the through-shining 3-4 irregular columns IPL cells;C, two IPL trabeculae with a papillose margin; D, IPL; E, moist capsule with ±erect, gently incurved teeth. All from holotype, La Réunion, Commune de Saint-Philippe, Ah-Peng &amp; Bardat s.n., private herbarium Bruggeman-Nannenga. Scale bars: A-D, 20 µm; E, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.12215095" httpUri="https://zenodo.org/record/12215095/files/figure.png" pageId="4" pageNumber="11">Fig. 2</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BBD36D1FF94FFB7523DF969DA59F8C5" blockId="4.[811,1458,1703,1886]" pageId="4" pageNumber="11">
<taxonomicName id="4C024D52FF94FFB7523DF969DDC0F924" box="[813,998,1703,1727]" class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="4" pageNumber="11" phylum="Bryophyta" rank="species" species="ah-pengae">
<emphasis id="B976EAC3FF94FFB7523DF969DDC0F924" box="[813,998,1703,1727]" italics="true" pageId="4" pageNumber="11">Fissidens ah-pengae</emphasis>
</taxonomicName>
is characterized by wide, elimbate, oblong to oblong-lingulate leaves with rounded-obtuse tips, limbidia weak, restricted to the vaginant laminae of perichaetial leaves, smooth laminal cells, costae that end 2-6 cells below the leaf tips and undivided, densely papillose peristome teeth. It is the only corticolous subgen.
<taxonomicName id="4C024D52FF94FFB7523CF8E2DDCCF8DF" box="[812,1002,1836,1860]" pageId="4" pageNumber="11">
<emphasis id="B976EAC3FF94FFB7523CF8E2DD82F8DF" box="[812,932,1836,1860]" italics="true" pageId="4" pageNumber="11">Polypodiopsis</emphasis>
species
</taxonomicName>
in Africa with smooth laminal cells. Corticolous on tree-ferns in lowland rainforest.
</paragraph>
<paragraph id="8BBD36D1FF94FFB7523DF85EDDA9F832" blockId="4.[813,911,1936,1961]" box="[813,911,1936,1961]" pageId="4" pageNumber="11">
<heading id="D0F581BDFF94FFB7523DF85EDDA9F832" box="[813,911,1936,1961]" fontSize="11" level="3" pageId="4" pageNumber="11" reason="9">
<emphasis id="B976EAC3FF94FFB7523DF85EDDA9F832" box="[813,911,1936,1961]" italics="true" pageId="4" pageNumber="11">Peristome</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36D1FF94FFB5523DF87CDFEFF9F7" blockId="4.[813,1455,1969,2028]" lastBlockId="6.[132,775,1586,1645]" lastPageId="6" lastPageNumber="13" pageId="4" pageNumber="11">
Erect with slightly incurved distal parts (both dry and wet), ±undivided, (seemingly?) perforated (
<figureCitation id="13392A54FF94FFB755A8F81FDB04F870" box="[1208,1314,2001,2027]" captionStart="FIG" captionStartId="6.[132,143,1444,1462]" captionTargetBox="[279,1307,215,1399]" captionTargetId="figure-205@6.[278,841,761,1310]" captionTargetPageId="6" captionText="FIG. 2. —Fissidens ah-pengae Brugg.-Nann. A, OPL with focus on the thin trabeculae;B, OPL focus on the through-shining 3-4 irregular columns IPL cells;C, two IPL trabeculae with a papillose margin; D, IPL; E, moist capsule with ±erect, gently incurved teeth. All from holotype, La Réunion, Commune de Saint-Philippe, Ah-Peng &amp; Bardat s.n., private herbarium Bruggeman-Nannenga. Scale bars: A-D, 20 µm; E, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.12215095" httpUri="https://zenodo.org/record/12215095/files/figure.png" pageId="4" pageNumber="11">Fig. 2A, B</figureCitation>
), densely and coarsely papillose, ±100 µm long,
<quantity id="4CFA9B34FF96FFB550EEF9FCDC55F9D6" box="[510,627,1586,1613]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="6" pageNumber="13" unit="mm" value="28.0" valueMax="30.0" valueMin="26.0">26-30 mm</quantity>
wide at base; distal parts breaking off easily.
</paragraph>
<caption id="DF7D6659FF95FFB65194F8B3DC54F872" ID-DOI="http://doi.org/10.5281/zenodo.12215091" ID-Zenodo-Dep="12215091" httpUri="https://zenodo.org/record/12215091/files/figure.png" pageId="5" pageNumber="12" startId="5.[132,143,1917,1935]" targetBox="[223,1363,215,1840]" targetPageId="5" targetType="figure">
<paragraph id="8BBD36D1FF95FFB65194F8B3DC54F872" blockId="5.[132,1457,1916,2025]" pageId="5" pageNumber="12">
<smallCapsWord id="8D5BA00DFF95FFB65194F8B3DEBBF816" baselines="1930,1930" box="[132,157,1917,1935]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="5" pageNumber="12">FIG</smallCapsWord>
. 1.
<emphasis id="B976EAC3FF95FFB651ACF8B3DEE9F814" box="[188,207,1917,1935]" italics="true" pageId="5" pageNumber="12"></emphasis>
<taxonomicName id="4C024D52FF95FFB651C4F8B3DFD6F815" authority="Cardot. A, OPL" authorityName="Cardot. A, OPL" box="[212,496,1916,1935]" class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="5" pageNumber="12" phylum="Bryophyta" rank="species" species="marthae">
<emphasis id="B976EAC3FF95FFB651C4F8B3DF4DF814" box="[212,363,1917,1935]" italics="true" pageId="5" pageNumber="12">Fissidens marthae</emphasis>
Cardot.
<emphasis id="B976EAC3FF95FFB650A4F8B3DFE7F814" bold="true" box="[436,449,1917,1935]" pageId="5" pageNumber="12">A</emphasis>
, OPL
</taxonomicName>
mid tooth, showing the bifurcation and filaments;
<emphasis id="B976EAC3FF95FFB65282F8B3DD83F814" bold="true" box="[914,933,1917,1935]" pageId="5" pageNumber="12">B,</emphasis>
OPL basal undivided part and bifurcation;
<emphasis id="B976EAC3FF95FFB6541AF8B2DB38F815" bold="true" box="[1290,1310,1916,1934]" pageId="5" pageNumber="12">C,</emphasis>
IPL showing fimbriate trabeculae of the undivided part and high horizontal to oblique IPL (“spiral”) ornamentation of the filaments;
<emphasis id="B976EAC3FF95FFB65550F85DDA75F83E" bold="true" box="[1088,1107,1939,1957]" pageId="5" pageNumber="12">D,</emphasis>
basal OPL of two teeth;
<emphasis id="B976EAC3FF95FFB65433F85DDB6DF83E" bold="true" box="[1315,1355,1939,1957]" pageId="5" pageNumber="12">E1-3</emphasis>
, side-views of basal part of filament showing the low OPL and thicker IPL;
<emphasis id="B976EAC3FF95FFB6539AF864DC87F827" bold="true" box="[650,673,1962,1980]" pageId="5" pageNumber="12">E1</emphasis>
, focus on the vertical walls of the OPL;
<emphasis id="B976EAC3FF95FFB652FAF864DA27F827" bold="true" box="[1002,1025,1962,1980]" pageId="5" pageNumber="12">E3</emphasis>
, stack of E1 and 2;
<emphasis id="B976EAC3FF95FFB655B5F864DA96F827" bold="true" box="[1189,1200,1962,1980]" pageId="5" pageNumber="12">F</emphasis>
, spores.
<emphasis id="B976EAC3FF95FFB655EDF864DB2CF827" bold="true" box="[1277,1290,1962,1980]" pageId="5" pageNumber="12">A</emphasis>
,
<emphasis id="B976EAC3FF95FFB65404F864DB04F827" bold="true" box="[1300,1314,1962,1980]" pageId="5" pageNumber="12">D</emphasis>
,
<emphasis id="B976EAC3FF95FFB6543CF864DB1EF827" bold="true" box="[1324,1336,1962,1980]" pageId="5" pageNumber="12">E</emphasis>
from Uganda, Mupanga Falls,
<emphasis id="B976EAC3FF95FFB6501AF80FDF5BF848" box="[266,381,1985,2003]" italics="true" pageId="5" pageNumber="12">Porley U495A</emphasis>
, private herbarium Bruggeman-Nannenga;
<emphasis id="B976EAC3FF95FFB653F8F80FDCD3F848" bold="true" box="[744,757,1985,2003]" pageId="5" pageNumber="12">B</emphasis>
from Tanzania,
<emphasis id="B976EAC3FF95FFB6526AF80EDDCFF848" box="[890,1001,1984,2003]" italics="true" pageId="5" pageNumber="12">Pócs 8633/M</emphasis>
, PC;
<emphasis id="B976EAC3FF95FFB65508F80EDA00F849" bold="true" box="[1048,1062,1984,2002]" pageId="5" pageNumber="12">C</emphasis>
,
<emphasis id="B976EAC3FF95FFB65521F80FDA1AF848" bold="true" box="[1073,1084,1985,2003]" pageId="5" pageNumber="12">F</emphasis>
from Central African Republic, S. Ippy, Kéyorédé,
<emphasis id="B976EAC3FF95FFB651AFF819DF01F872" box="[191,295,2007,2025]" italics="true" pageId="5" pageNumber="12">Tisserant s.n</emphasis>
., PC. Scale bars: A-E, 20 µm; F, 50 µm.
</paragraph>
</caption>
<caption id="DF7D6659FF96FFB55194FA6ADDE9FA78" ID-DOI="http://doi.org/10.5281/zenodo.12215095" ID-Zenodo-Dep="12215095" httpUri="https://zenodo.org/record/12215095/files/figure.png" pageId="6" pageNumber="13" startId="6.[132,143,1444,1462]" targetBox="[279,1307,215,1399]" targetPageId="6" targetType="figure">
<paragraph id="8BBD36D1FF96FFB55194FA6ADDE9FA78" blockId="6.[132,1457,1444,1507]" pageId="6" pageNumber="13">
<smallCapsWord id="8D5BA00DFF96FFB55194FA6ADEBBFA2E" baselines="1457,1458" box="[132,157,1444,1462]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="6" pageNumber="13">FIG</smallCapsWord>
. 2.
<emphasis id="B976EAC3FF96FFB551ABFA6ADEE9FA2D" box="[187,207,1444,1462]" italics="true" pageId="6" pageNumber="13"></emphasis>
<taxonomicName id="4C024D52FF96FFB551C1FA6ADF9DFA2D" authority="Brugg." authorityName="Brugg." box="[209,443,1444,1462]" class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="6" pageNumber="13" phylum="Bryophyta" rank="species" species="ah-pengae">
<emphasis id="B976EAC3FF96FFB551C1FA6ADF5DFA2D" box="[209,379,1444,1462]" italics="true" pageId="6" pageNumber="13">Fissidens ah-pengae</emphasis>
Brugg.
</taxonomicName>
-Nann.
<emphasis id="B976EAC3FF96FFB550E6FA6ADC25FA2D" bold="true" box="[502,515,1444,1462]" pageId="6" pageNumber="13">A</emphasis>
, OPL with focus on the thin trabeculae;
<emphasis id="B976EAC3FF96FFB55258FA6ADD73FA2D" bold="true" box="[840,853,1444,1462]" pageId="6" pageNumber="13">B</emphasis>
, OPL focus on the through-shining 3-4 irregular columns IPL cells;
<emphasis id="B976EAC3FF96FFB55469FA6ADBA1FA2D" bold="true" box="[1401,1415,1444,1462]" pageId="6" pageNumber="13">C</emphasis>
, two IPL trabeculae with a papillose margin;
<emphasis id="B976EAC3FF96FFB550DCFA74DFFCFA57" bold="true" box="[460,474,1466,1484]" pageId="6" pageNumber="13">D</emphasis>
, IPL;
<emphasis id="B976EAC3FF96FFB5531BFA74DC31FA57" bold="true" box="[523,535,1466,1484]" pageId="6" pageNumber="13">E</emphasis>
, moist capsule with ±erect, gently incurved teeth. All from holotype, La Réunion, Commune de Saint-Philippe,
<emphasis id="B976EAC3FF96FFB55194FA1FDF1EFA78" box="[132,312,1489,1507]" italics="true" pageId="6" pageNumber="13">Ah-Peng &amp; Bardat s.n</emphasis>
., private herbarium Bruggeman-Nannenga. Scale bars: A-D, 20 µm; E, 0.4 mm.
</paragraph>
</caption>
<paragraph id="8BBD36D1FF96FFB55194F95DDF00F936" blockId="6.[132,294,1683,1709]" box="[132,294,1683,1709]" pageId="6" pageNumber="13">
<heading id="D0F581BDFF96FFB55194F95DDF00F936" box="[132,294,1683,1709]" fontSize="11" level="3" pageId="6" pageNumber="13" reason="9">
<emphasis id="B976EAC3FF96FFB55194F95DDF00F936" box="[132,294,1683,1709]" italics="true" pageId="6" pageNumber="13">Ornamentation</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36D1FF96FFB55194F97CDD21F8F7" blockId="6.[130,776,1714,1900]" pageId="6" pageNumber="13">
OPL trabeculae thin, most distinct in basal cells (
<figureCitation id="13392A54FF96FFB5536CF97CDCEDF957" box="[636,715,1714,1740]" captionStart="FIG" captionStartId="6.[132,143,1444,1462]" captionTargetBox="[279,1307,215,1399]" captionTargetId="figure-205@6.[278,841,761,1310]" captionTargetPageId="6" captionText="FIG. 2. —Fissidens ah-pengae Brugg.-Nann. A, OPL with focus on the thin trabeculae;B, OPL focus on the through-shining 3-4 irregular columns IPL cells;C, two IPL trabeculae with a papillose margin; D, IPL; E, moist capsule with ±erect, gently incurved teeth. All from holotype, La Réunion, Commune de Saint-Philippe, Ah-Peng &amp; Bardat s.n., private herbarium Bruggeman-Nannenga. Scale bars: A-D, 20 µm; E, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.12215095" httpUri="https://zenodo.org/record/12215095/files/figure.png" pageId="6" pageNumber="13">Fig. 2A</figureCitation>
); mid and distal part coarsely papillose interspersed with “perforations”; IPL trabeculae of basal IPL cells with a papillose border (
<figureCitation id="13392A54FF96FFB553BDF93CDCDCF897" box="[685,762,1778,1804]" captionStart="FIG" captionStartId="6.[132,143,1444,1462]" captionTargetBox="[279,1307,215,1399]" captionTargetId="figure-205@6.[278,841,761,1310]" captionTargetPageId="6" captionText="FIG. 2. —Fissidens ah-pengae Brugg.-Nann. A, OPL with focus on the thin trabeculae;B, OPL focus on the through-shining 3-4 irregular columns IPL cells;C, two IPL trabeculae with a papillose margin; D, IPL; E, moist capsule with ±erect, gently incurved teeth. All from holotype, La Réunion, Commune de Saint-Philippe, Ah-Peng &amp; Bardat s.n., private herbarium Bruggeman-Nannenga. Scale bars: A-D, 20 µm; E, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.12215095" httpUri="https://zenodo.org/record/12215095/files/figure.png" pageId="6" pageNumber="13">Fig.2C</figureCitation>
), IPL in mid and upper part with irregularly thickened, papillose plaques forming a pattern of 3(-4) irregular columns that disturb the normal IPL structure of two columns (
<figureCitation id="13392A54FF96FFB5539AF89FDCDCF8F7" box="[650,762,1873,1900]" captionStart="FIG" captionStartId="6.[132,143,1444,1462]" captionTargetBox="[279,1307,215,1399]" captionTargetId="figure-205@6.[278,841,761,1310]" captionTargetPageId="6" captionText="FIG. 2. —Fissidens ah-pengae Brugg.-Nann. A, OPL with focus on the thin trabeculae;B, OPL focus on the through-shining 3-4 irregular columns IPL cells;C, two IPL trabeculae with a papillose margin; D, IPL; E, moist capsule with ±erect, gently incurved teeth. All from holotype, La Réunion, Commune de Saint-Philippe, Ah-Peng &amp; Bardat s.n., private herbarium Bruggeman-Nannenga. Scale bars: A-D, 20 µm; E, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.12215095" httpUri="https://zenodo.org/record/12215095/files/figure.png" pageId="6" pageNumber="13">Fig. 2B, D</figureCitation>
).
</paragraph>
<paragraph id="8BBD36D1FF96FFB55194F85FDED5F830" blockId="6.[132,243,1937,1963]" box="[132,243,1937,1963]" pageId="6" pageNumber="13">
<heading id="D0F581BDFF96FFB55194F85FDED5F830" box="[132,243,1937,1963]" fontSize="11" level="3" pageId="6" pageNumber="13" reason="9">
<emphasis id="B976EAC3FF96FFB55194F85FDED5F830" box="[132,243,1937,1963]" italics="true" pageId="6" pageNumber="13">Sporophyte</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36D1FF96FFB55194F87FDDA1F9F7" blockId="6.[130,775,1969,2028]" lastBlockId="6.[814,1455,1586,1644]" pageId="6" pageNumber="13">
Seta ±
<quantity id="4CFA9B34FF96FFB551D7F87FDF20F857" box="[199,262,1969,1996]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="13" unit="mm" value="1.0">1 mm</quantity>
long, smooth; capsule cylindrical, 0.4 ×
<quantity id="4CFA9B34FF96FFB553BDF87FDD25F857" box="[685,771,1969,1996]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="13" unit="mm" value="0.3">0.3 mm</quantity>
, exothecial cell columns ± 32, the cells quadratic-oblong to oblong; operculum not observed; spores ±15 µm in diam., smooth.
</paragraph>
<paragraph id="8BBD36D1FF96FFB5523DF95DDA6FF937" blockId="6.[813,1097,1682,1708]" box="[813,1097,1682,1708]" pageId="6" pageNumber="13">
<heading id="D0F581BDFF96FFB5523DF95DDA6FF937" box="[813,1097,1682,1708]" fontSize="11" level="3" pageId="6" pageNumber="13" reason="9">
<emphasis id="B976EAC3FF96FFB5523DF95DDA6FF937" box="[813,1097,1682,1708]" italics="true" pageId="6" pageNumber="13">Description and illustration</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36D1FF96FFB5523DF97DDA22F977" blockId="6.[810,1453,1714,1772]" pageId="6" pageNumber="13">
Bruggeman-Nannenga (2009: fig.1);
<bibRefCitation id="EF934B20FF96FFB555B8F97DDDB9F977" author="BRUGGEMAN-NANNENGA M. A. &amp; ARTS T." pageId="6" pageNumber="13" pagination="170 - 207" refId="ref13574" refString="BRUGGEMAN-NANNENGA M. A. &amp; ARTS T. 2010. - A revision of the Fissidentaceae (Musci) of La Reunion (including all species known from Mauritius and Rodriguez). Journal of Bryology 32: 170 - 207. https: // doi. org / 10.1179 / 037366810 X 12735734836179" type="journal article" year="2010">Bruggeman-Nannenga &amp; Arts (2010</bibRefCitation>
: fig. 18).
</paragraph>
</subSubSection>
<subSubSection id="C318655AFF96FFB5523DF8DCDD8CF870" pageId="6" pageNumber="13" type="discussion">
<paragraph id="8BBD36D1FF96FFB5523DF8DCDDA3F8B7" blockId="6.[813,901,1810,1836]" box="[813,901,1810,1836]" pageId="6" pageNumber="13">
<heading id="D0F581BDFF96FFB5523DF8DCDDA3F8B7" box="[813,901,1810,1836]" fontSize="11" level="3" pageId="6" pageNumber="13" reason="9">
<emphasis id="B976EAC3FF96FFB5523DF8DCDDA3F8B7" box="[813,901,1810,1836]" italics="true" pageId="6" pageNumber="13">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36D1FF96FFB5523DF8FCDD8CF870" blockId="6.[811,1456,1841,2027]" pageId="6" pageNumber="13">
In this species the OPL is difficult to distinguish from the through-shining IPL. It is not clear whether the perforations are thin areas or true perforations. The peristome of
<taxonomicName id="4C024D52FF96FFB5523CF85CDD93F830" box="[812,949,1937,1963]" class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="6" pageNumber="13" phylum="Bryophyta" rank="species" species="ah-pengae">
<emphasis id="B976EAC3FF96FFB5523CF85CDD93F830" box="[812,949,1937,1963]" italics="true" pageId="6" pageNumber="13">F. ah-pengae</emphasis>
</taxonomicName>
resembles that of
<taxonomicName id="4C024D52FF96FFB55580F85CDB89F837" authority="Sande Lac." authorityName="Sande Lac." box="[1168,1455,1937,1964]" class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="6" pageNumber="13" phylum="Bryophyta" rank="species" species="punctulatus">
<emphasis id="B976EAC3FF96FFB55580F85CDB0DF830" box="[1168,1323,1937,1963]" italics="true" pageId="6" pageNumber="13">F. punctulatus</emphasis>
Sande Lac.
</taxonomicName>
in being densely and coarsely papillose and seemingly perforated.
</paragraph>
</subSubSection>
</treatment>
</document>