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<document id="5FA62B65B81D8C6C13C8633CD948ABA1" ID-DOI="10.11646/zootaxa.3666.2.2" ID-GBIF-Dataset="930d392c-8720-4820-85eb-501985209cf3" ID-ISSN="1175-5326" ID-Zenodo-Dep="220466" ID-ZooBank="1F8D4E8A-2447-4556-9B98-D8229D858194" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460622323338" checkinUser="plazi" docAuthor="Prokin, Alexander A., Petrov, Pyotr N., Wang, Bo &amp; Ponomarenko, Alexander G." docDate="2013" docId="03AB6D60FF97D23CFF440F0EB7BE4BDA" docLanguage="en" docName="zt03666p159.pdf" docOrigin="Zootaxa 3666 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mesoderus punctatus Prokin, Petrov, Wang &amp; Ponomarenko, 2013, sp. n." docType="treatment" docVersion="8" lastPageNumber="147" masterDocId="FF921518FF9FD236FFD30973B6134A15" masterDocTitle="New fossil taxa and notes on the Mesozoic evolution of Liadytidae and Dytiscidae (Coleoptera)" masterLastPageNumber="159" masterPageNumber="137" pageNumber="145" updateTime="1698334052497" updateUser="plazi">
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<mods:titleInfo id="CD39D2715EAD13D8ED9EC4AC272339DE">
<mods:title id="86B1CC60AA48882EC3D863C532741ECA">New fossil taxa and notes on the Mesozoic evolution of Liadytidae and Dytiscidae (Coleoptera)</mods:title>
</mods:titleInfo>
<mods:name id="0A5BEC468659ABBF0644B7C783D5BDB7" type="personal">
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<mods:namePart id="0E050AF993518A38A5C95AD1FA1505CC">Prokin, Alexander A.</mods:namePart>
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<mods:name id="D3872CB9C71EB3E0AA0EC0E14DFA13FD" type="personal">
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<mods:namePart id="D0B4CFC5BD0D2E0105180EA4481BFE59">Petrov, Pyotr N.</mods:namePart>
</mods:name>
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<mods:namePart id="3A0CCDF2C6791484F7EB0C40B19FF798">Wang, Bo</mods:namePart>
</mods:name>
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<mods:namePart id="245B7DB2566531186AD4D6AE83DC784F">Ponomarenko, Alexander G.</mods:namePart>
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<mods:title id="DE87DAC56B99A756A60EBCADACF638E3">Zootaxa</mods:title>
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<mods:date id="CE38F4B86E8082741CFD47E2DCA10A26">2013</mods:date>
<mods:detail id="3B2330FB14EDC455A05851B54ABA77D0" type="volume">
<mods:number id="06ADD4120A0241F8F7B56F62D1C4E657">3666</mods:number>
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<mods:classification id="E2375B328CD1079F5190FB9D37038E38">journal article</mods:classification>
<mods:identifier id="812D30FB8AD1FD7B2794AE8895A5C510" type="DOI">10.11646/zootaxa.3666.2.2</mods:identifier>
<mods:identifier id="97F28B1BA5DC4B493479BAC9F1173421" type="GBIF-Dataset">930d392c-8720-4820-85eb-501985209cf3</mods:identifier>
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<treatment id="03AB6D60FF97D23CFF440F0EB7BE4BDA" ID-DOI="http://doi.org/10.5281/zenodo.5680608" ID-GBIF-Taxon="119569017" ID-Zenodo-Dep="5680608" LSID="urn:lsid:plazi:treatment:03AB6D60FF97D23CFF440F0EB7BE4BDA" httpUri="http://treatment.plazi.org/id/03AB6D60FF97D23CFF440F0EB7BE4BDA" lastPageId="10" lastPageNumber="147" pageId="8" pageNumber="145">
<subSubSection id="C3188FFDFF97D23EFF440F0EB7F14C82" box="[151,482,1661,1687]" pageId="8" pageNumber="145" type="nomenclature">
<paragraph id="8BBDDC76FF97D23EFF440F0EB7F14C82" blockId="8.[151,482,1661,1687]" box="[151,482,1661,1687]" pageId="8" pageNumber="145">
<heading id="D0F56B1AFF97D23EFF440F0EB7F14C82" bold="true" box="[151,482,1661,1687]" fontSize="11" level="1" pageId="8" pageNumber="145" reason="1">
<emphasis id="B9760064FF97D23EFF440F0EB7F14C82" bold="true" box="[151,482,1661,1687]" pageId="8" pageNumber="145">
<taxonomicName id="4C02A7F5FF97D23EFF440F0EB78A4C82" box="[151,409,1661,1687]" class="Insecta" family="Liadytidae" genus="Mesoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="145" phylum="Arthropoda" rank="species" species="punctatus" status="sp. nov.">
<emphasis id="B9760064FF97D23EFF440F0EB78A4C82" bold="true" box="[151,409,1661,1687]" italics="true" pageId="8" pageNumber="145">Mesoderus punctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245BD1FFF97D23EFE730F0EB7F14C82" box="[416,482,1661,1687]" pageId="8" pageNumber="145" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3188FFDFF97D23EFF440FB7B20A4CC9" box="[151,1049,1732,1756]" pageId="8" pageNumber="145" type="etymology">
<paragraph id="8BBDDC76FF97D23EFF440FB7B20A4CC9" blockId="8.[151,1436,1732,2008]" box="[151,1049,1732,1756]" pageId="8" pageNumber="145">
<emphasis id="B9760064FF97D23EFF440FB7B70B4CC9" bold="true" box="[151,280,1732,1756]" pageId="8" pageNumber="145">Etymology</emphasis>
: The Latin
<taxonomicName id="4C02A7F5FF97D23EFE480FB6B41B4CC9" box="[411,520,1733,1756]" class="Insecta" family="Liadytidae" genus="Mesoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="145" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="B9760064FF97D23EFE480FB6B41B4CC9" box="[411,520,1733,1756]" italics="true" pageId="8" pageNumber="145">punctatus</emphasis>
</taxonomicName>
(spotted); adjective in the nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="C3188FFDFF97D23EFF140F94B7004D31" pageId="8" pageNumber="145" type="materials_examined">
<paragraph id="8BBDDC76FF97D23EFF140F94B7004D31" blockId="8.[151,1436,1732,2008]" pageId="8" pageNumber="145">
<emphasis id="B9760064FF97D23EFF140F94B73C4D15" bold="true" box="[199,303,1767,1792]" pageId="8" pageNumber="145">Material</emphasis>
:
<typeStatus id="54B962D4FF97D23EFEEC0F94B7B44CEA" box="[319,423,1767,1791]" pageId="8" pageNumber="145" type="holotype">Holotype</typeStatus>
NIGP 0 5622, counter impression of beetle;
<collectingCountry id="F3159CE6FF97D23EFC700F94B5FA4CEA" box="[931,1001,1767,1791]" name="China" pageId="8" pageNumber="145">China</collectingCountry>
, Liutiaogou; Early
<taxonomicName id="4C02A7F5FF97D23EFB160F94B3564CEA" box="[1221,1349,1767,1791]" class="Insecta" family="Liadytidae" genus="Liadytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="138" phylum="Arthropoda" rank="species" species="cretaceous">Cretaceous</taxonomicName>
, Yixian Formation.
</paragraph>
</subSubSection>
<subSubSection id="C3188FFDFF97D23CFF140E5CB7BE4BDA" lastPageId="10" lastPageNumber="147" pageId="8" pageNumber="145" type="description">
<paragraph id="8BBDDC76FF97D23CFF140E5CB5FD4B2A" blockId="8.[151,1436,1732,2008]" lastBlockId="10.[151,1436,151,463]" lastPageId="10" lastPageNumber="147" pageId="8" pageNumber="145">
<emphasis id="B9760064FF97D23EFF140E5CB7424D5D" bold="true" box="[199,337,1839,1864]" pageId="8" pageNumber="145">Description</emphasis>
(
<figureCitation id="1339C0F3FF97D23EFEB30E5CB7DF4D52" box="[352,460,1839,1863]" captionStart="FIGURES 8 – 11" captionStartId="9.[151,264,1884,1907]" captionTargetBox="[154,1426,193,1863]" captionTargetId="figure@9.[154,1433,193,1863]" captionTargetPageId="9" captionText="FIGURES 8 – 11. Mesoderus punctatus sp. n.: (8) photo of holotype, (9) line drawing of holotype; arrow indicates penis; Mesoderus ovatus sp. n.: (10) photo of holotype, (11) line drawing of holotype." httpUri="https://zenodo.org/record/220469/files/figure.png" pageId="8" pageNumber="145">Figs. 8, 9</figureCitation>
): Light coloured beetle. Maximum body width situated at middle of abdominal ventrite 2. Elytron light, with dark spots, distance between spots twice as great as their diameter. Head length equal to width at posterior margin of eyes. Pronotum three times as wide as long, 1.6 times as wide basally as distally. Profemur narrowed towards apex; protarsal claws equal in length. Mesocoxae slightly separated. Mesofemur with anterior margin straight and posterior margin convex, slightly protruding beyond sides of body; mesotibia straight, with setose anterior margin. Metacoxal plate 1.2 times as long as lateral lobe of metaventrite (LC/LV = 1.2). Metafemur dilated towards apex. Metatibia straight, slightly shorter than metafemur, with swimming setae. Metatarsomere 1 with swimming setae, equal in length to metatarsomere 2; metatarsomere 2 equal in length to metatarsomeres 3 and 4 together; metatarsomere 5 short, only half as long as metatarsomere 4; claws of metatarsus equal in length, twice as long as metatarsomere 5. Penis pointed and curved (see arrow in
<figureCitation id="1339C0F3FF95D23CFC5D0854B5F24B2A" box="[910,993,295,319]" captionStart="FIGURES 8 – 11" captionStartId="9.[151,264,1884,1907]" captionTargetBox="[154,1426,193,1863]" captionTargetId="figure@9.[154,1433,193,1863]" captionTargetPageId="9" captionText="FIGURES 8 – 11. Mesoderus punctatus sp. n.: (8) photo of holotype, (9) line drawing of holotype; arrow indicates penis; Mesoderus ovatus sp. n.: (10) photo of holotype, (11) line drawing of holotype." httpUri="https://zenodo.org/record/220469/files/figure.png" pageId="10" pageNumber="147">Fig. 10</figureCitation>
).
</paragraph>
<caption id="DF7D8CFEFF96D23FFF440E2FB5AE4D84" httpUri="https://zenodo.org/record/220469/files/figure.png" pageId="9" pageNumber="146" targetBox="[154,1426,193,1863]" targetPageId="9">
<paragraph id="8BBDDC76FF96D23FFF440E2FB5AE4D84" blockId="9.[151,1436,1884,1938]" pageId="9" pageNumber="146">
<emphasis id="B9760064FF96D23FFF440E2FB75A4D67" bold="true" box="[151,329,1884,1907]" pageId="9" pageNumber="146">FIGURES 8–11.</emphasis>
<taxonomicName id="4C02A7F5FF96D23FFE800E2EB43C4D66" box="[339,559,1885,1907]" class="Insecta" family="Liadytidae" genus="Mesoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="146" phylum="Arthropoda" rank="species" species="punctatus" status="sp. nov.">
<emphasis id="B9760064FF96D23FFE800E2EB43C4D66" box="[339,559,1885,1907]" italics="true" pageId="9" pageNumber="146">Mesoderus punctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245BD1FFF96D23FFDEB0E2EB4664D67" box="[568,629,1885,1906]" pageId="9" pageNumber="146" rank="species">
<emphasis id="B9760064FF96D23FFDEB0E2EB4664D67" bold="true" box="[568,629,1885,1906]" pageId="9" pageNumber="146">sp. n.</emphasis>
</taxonomicNameLabel>
: (8) photo of holotype, (9) line drawing of holotype; arrow indicates penis;
<taxonomicName id="4C02A7F5FF96D23FFF440E0FB75D4D87" box="[151,334,1916,1938]" class="Insecta" family="Liadytidae" genus="Mesoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="146" phylum="Arthropoda" rank="species" species="ovatus" status="sp. nov.">
<emphasis id="B9760064FF96D23FFF440E0FB75D4D87" box="[151,334,1916,1938]" italics="true" pageId="9" pageNumber="146">Mesoderus ovatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245BD1FFF96D23FFE870E0FB79E4D84" box="[340,397,1916,1937]" pageId="9" pageNumber="146" rank="species">
<emphasis id="B9760064FF96D23FFE870E0FB79E4D84" bold="true" box="[340,397,1916,1937]" pageId="9" pageNumber="146">sp. n.</emphasis>
</taxonomicNameLabel>
: (10) photo of holotype, (11) line drawing of holotype.
</paragraph>
</caption>
<paragraph id="8BBDDC76FF95D23CFF14083FB2864B71" blockId="10.[151,1436,151,463]" box="[199,1173,332,356]" pageId="10" pageNumber="147">
<emphasis id="B9760064FF95D23CFF14083FB7654B71" bold="true" box="[199,374,332,356]" pageId="10" pageNumber="147">Measurements</emphasis>
: Body length,
<quantity id="4CFA7193FF95D23CFDCF083FB4904B71" box="[540,643,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="10" pageNumber="147" unit="mm" value="14.6">14.6 mm</quantity>
; body width,
<quantity id="4CFA7193FF95D23CFCC8083FB5674B71" box="[795,884,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="10" pageNumber="147" unit="mm" value="7.5">7.5 mm</quantity>
; elytron length,
<quantity id="4CFA7193FF95D23CFBF9083FB2824B71" box="[1066,1169,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="10" pageNumber="147" unit="mm" value="10.5">10.5 mm</quantity>
.
</paragraph>
<paragraph id="8BBDDC76FF95D23CFF14081CB7BE4BDA" blockId="10.[151,1436,151,463]" pageId="10" pageNumber="147">
<emphasis id="B9760064FF95D23CFF14081CB7494B9D" bold="true" box="[199,346,367,392]" pageId="10" pageNumber="147">Comparison</emphasis>
: Similar to
<taxonomicName id="4C02A7F5FF95D23CFE390802B5714B92" authority="Prokin &amp; Ren, 2010" authorityName="Prokin &amp; Ren" authorityYear="2010" box="[490,866,367,392]" class="Insecta" family="Liadytidae" genus="Mesoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="147" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="B9760064FF95D23CFE390802B4794B9D" box="[490,618,369,392]" italics="true" pageId="10" pageNumber="147">M. magnus</emphasis>
Prokin &amp; Ren,
<quantity id="4CFA7193FF95D23CFCF9081CB5714B92" box="[810,866,367,391]" metricMagnitude="1" metricUnit="m" metricValue="5.1053999999999995" pageId="10" pageNumber="147" unit="in" value="2010.0">2010</quantity>
</taxonomicName>
in size and in the pointed and curved penis, but differs from it in the dorsal coloration, LC/LV, longer metacoxal processes, shape of meso- and metafemora, and shorter metatarsomere 1.
</paragraph>
</subSubSection>
</treatment>
</document>
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