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<document id="5D7A384F8C3486EACA19F2075F736554" ID-DOI="10.11646/zootaxa.4134.1.1" ID-GBIF-Dataset="a157eae0-8cb8-4b9a-8ce5-dd7ff7913ba3" ID-ISSN="1175-5326" ID-Zenodo-Dep="399754" ID-ZooBank="92AC0E20-F532-4D21-AE1F-4B056327212F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1467637689624" checkinUser="plazi" docAuthor="Santos-Silva, Antonio, Nearns, Eugenio H. &amp; Swift, Ian P." docDate="2016" docId="03AA87ACFFE76725FF2CC43129BE86DE" docLanguage="en" docName="zootaxa.4134.1.pdf.imf" docOrigin="Zootaxa 4134 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Prionus (Prionus) howdeni Chemsak 1979" docType="treatment" docVersion="12" lastPageNumber="37" masterDocId="FF93FFD4FFC36701FFBBC73C2B4C8170" masterDocTitle="Revision of the American species of the genus Prionus Geoffroy, 1762 (Coleoptera, Cerambycidae, Prioninae, Prionini)" masterLastPageNumber="103" masterPageNumber="1" pageNumber="37" updateTime="1698363407732" updateUser="plazi">
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<mods:title id="B27E27FA9B8653012C9918EFC9F95F86">Revision of the American species of the genus Prionus Geoffroy, 1762 (Coleoptera, Cerambycidae, Prioninae, Prionini)</mods:title>
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<mods:namePart id="B3783D331F2FE7BB7BF13D1B8EA35714">Santos-Silva, Antonio</mods:namePart>
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<mods:namePart id="F767BBE915C8E23521C94AE4FC79450D">Nearns, Eugenio H.</mods:namePart>
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<mods:namePart id="8F8D5371EBF5445EC5411B6B1D38F54F">Swift, Ian P.</mods:namePart>
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<mods:date id="541672AC010D06A70179442D79748014">2016</mods:date>
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<taxonomicName id="4C034D39FFE76725FF2CC43129EA8257" ID-CoL="4MH36" authority="Chemsak, 1979" authorityName="Chemsak" authorityYear="1979" box="[151,678,781,807]" class="Insecta" family="Cerambycidae" genus="Prionus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="howdeni" subGenus="Prionus">
<emphasis id="B977EAA8FFE76725FF2CC43129EA8257" bold="true" box="[151,678,781,807]" pageId="36" pageNumber="37">
<emphasis id="B977EAA8FFE76725FF2CC4312BBB8257" bold="true" box="[151,247,781,807]" italics="true" pageId="36" pageNumber="37">Prionus</emphasis>
(
<emphasis id="B977EAA8FFE76725FEB3C4312A248257" bold="true" box="[264,360,781,807]" italics="true" pageId="36" pageNumber="37">Prionus</emphasis>
)
<emphasis id="B977EAA8FFE76725FEC2C4312A938257" bold="true" box="[377,479,781,807]" italics="true" pageId="36" pageNumber="37">howdeni</emphasis>
<bibRefCitation id="EF924B4BFFE76725FE5CC43129EA8257" author="Chemsak" box="[487,678,781,807]" pageId="36" pageNumber="37" refString="Chemsak, J. A. (1979) New species of Neotropical Prioninae (Coleoptera: Cerambycidae). The Coleopterists Bulletin, 33 (1), 125 - 128." type="journal article" year="1979">Chemsak, 1979</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BBC36BAFFE76725FF2CC4132A628238" blockId="36.[151,678,781,840]" box="[151,302,815,840]" pageId="36" pageNumber="37">
(
<figureCitation id="13382A3FFFE76725FF24C4132A698238" box="[159,293,815,840]" captionStart="FIGURES 91 94. 91 93" captionStartId="37.[151,264,1953,1975]" captionTargetBox="[249,1341,226,1913]" captionTargetId="figure@37.[237,1350,193,1932]" captionTargetPageId="37" captionText="FIGURES 91 94. 91 93, Prionus (Prionus) howdeni, holotype male: 91, dorsal habitus; 92, lateral habitus; 93, head, frontal view. 94, Prionus (Prionus) evoluticornis, paratype male, dorsal habitus. Photographs 91 93 by Serge Laplante; photograph 94 by Ziro Komiya." httpUri="https://zenodo.org/record/399978/files/figure.png" pageId="36" pageNumber="37">Figs. 9193</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3196531FFE76725FF2CC4452BB5871A" pageId="36" pageNumber="37" type="reference_group">
<paragraph id="8BBC36BAFFE76725FF2CC445297782BB" blockId="36.[151,1436,888,1032]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="AB131194FFE76725FF2CC445297782BB" pageId="36" pageNumber="37">
<taxonomicName id="4C034D39FFE76725FF2CC44529D482FE" ID-CoL="4MH36" authority="Chemsak, 1979: 127" authorityName="Chemsak" authorityPageNumber="127" authorityYear="1979" box="[151,664,888,910]" class="Insecta" family="Cerambycidae" genus="Prionus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="howdeni" subGenus="Prionus">
<emphasis id="B977EAA8FFE76725FF2CC4452BAB82FE" box="[151,231,889,910]" italics="true" pageId="36" pageNumber="37">Prionus</emphasis>
(
<emphasis id="B977EAA8FFE76725FF41C4452A0682FE" box="[250,330,889,910]" italics="true" pageId="36" pageNumber="37">Prionus</emphasis>
)
<emphasis id="B977EAA8FFE76725FEE7C4442AFD82FE" box="[348,433,888,910]" italics="true" pageId="36" pageNumber="37">howdeni</emphasis>
<treatmentCitation id="0AA210ABFFE76725FE07C44429D482FE" author="Chemsak" box="[444,664,888,910]" page="127" pageId="36" pageNumber="37" year="1979">
<bibRefCitation id="EF924B4BFFE76725FE07C444292D82FE" author="Chemsak" box="[444,609,888,910]" pageId="36" pageNumber="37" refString="Chemsak, J. A. (1979) New species of Neotropical Prioninae (Coleoptera: Cerambycidae). The Coleopterists Bulletin, 33 (1), 125 - 128." type="journal article" year="1979">Chemsak, 1979</bibRefCitation>
: 127
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AA210ABFFE76725FD1CC44528DF82FE" author="McNamara" box="[679,915,888,910]" page="734" pageId="36" pageNumber="37" year="1984">
<bibRefCitation id="EF924B4BFFE76725FD1CC445281382FE" author="McNamara" box="[679,863,888,910]" pageId="36" pageNumber="37" refString="McNamara, J. (1984) A catalogue of types of Coleoptera in the Canadian National Collection of insects, supplement II. The Canadian Entomologist, 116, 725 - 772." type="journal article" year="1984">McNamara, 1984</bibRefCitation>
: 734
</treatmentCitation>
(
<typeStatus id="54B88818FFE76725FC1EC444289982FE" box="[933,981,888,910]" pageId="36" pageNumber="37">type</typeStatus>
);
<treatmentCitation id="0AA210ABFFE76725FC50C4442E5382FE" author="Monne" box="[1003,1311,888,910]" page="15" pageId="36" pageNumber="37" year="1994">
<bibRefCitation id="EF924B4BFFE76725FC50C4442FB482FE" author="Monne" box="[1003,1272,888,910]" pageId="36" pageNumber="37" refString="Monne, M. A. &amp; Giesbert, E. F. (1994) Checklist of the Cerambycidae and Disteniidae (Coleoptera) of the Western Hemisphere. Wolfsgarden Books, Burbank, 409 pp." type="book" year="1994">Monné &amp; Giesbert, 1994</bibRefCitation>
: 15
</treatmentCitation>
(checklist); Monné, 1995: 52 (cat.);
<treatmentCitation id="0AA210ABFFE76725FE69C4AA284C82DC" author="Monne" box="[466,768,918,940]" page="20" pageId="36" pageNumber="37" year="2005">
<bibRefCitation id="EF924B4BFFE76725FE69C4AA299482DC" author="Monne" box="[466,728,918,940]" pageId="36" pageNumber="37" refString="Monne, M. A. &amp; Hovore, F. T. (2005) Checklist of the Cerambycidae, or longhorned wood-boring beetles of the Western Hemisphere. Bio Quip Publications, Rancho Dominguez, 393 pp." type="book" year="2005">Monné &amp; Hovore, 2005</bibRefCitation>
: 20
</treatmentCitation>
(checklist); 2006: 19 (checklist);
<treatmentCitation id="0AA210ABFFE76725FBCAC4AA2ED782DC" author="Bezark" box="[1137,1435,918,940]" page="28" pageId="36" pageNumber="37" year="2013">
<bibRefCitation id="EF924B4BFFE76725FBCAC4AA2E3F82DC" author="Bezark" box="[1137,1395,918,940]" pageId="36" pageNumber="37" refString="Bezark, L. G. &amp; Monne, M. A. (2013) Checklist of the Oxypeltidae, Vesperidae, Disteniidae and Cerambycidae, (Coleoptera) of the Western Hemisphere. Bio Quip Publications, Rancho Dominguez, 484 pp." type="book" year="2013">Bezark &amp; Monné, 2013</bibRefCitation>
: 28
</treatmentCitation>
(checklist); Monné, 2015: 176 (cat.).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBC36BAFFE76725FF2CC4E929428578" blockId="36.[151,1436,888,1032]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="AB131194FFE76725FF2CC4E929428578" pageId="36" pageNumber="37">
<taxonomicName id="4C034D39FFE76725FF2CC4E92A0B829A" ID-CoL="4MH36" box="[151,327,980,1002]" class="Insecta" family="Cerambycidae" genus="Prionus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="howdeni">
<emphasis id="B977EAA8FFE76725FF2CC4E92A0B829A" box="[151,327,980,1002]" italics="true" pageId="36" pageNumber="37">Prionus howdeni</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AA210ABFFE76725FEECC4E8292E829A" author="Chemsak" box="[343,610,980,1002]" page="21" pageId="36" pageNumber="37" year="1992">
<bibRefCitation id="EF924B4BFFE76725FEECC4E82970829A" author="Chemsak" box="[343,572,980,1002]" pageId="36" pageNumber="37" refString="Chemsak, J. A., Linsley, E. G. &amp; Noguera, F. A. (1992) Listados faunisticos de Mexico. II. Los Cerambycidae y Disteniidae de Norteamerica, Centroamerica y las Indias Occidentales (Coleoptera). Universidad Nacional Autonoma, Mexico, D. F., 204 pp." type="book" year="1992">
Chemsak
<emphasis id="B977EAA8FFE76725FE04C4E92ABA829A" box="[447,502,980,1002]" italics="true" pageId="36" pageNumber="37">et al.</emphasis>
, 1992
</bibRefCitation>
: 21
</treatmentCitation>
(checklist);
<treatmentCitation id="0AA210ABFFE76725FD53C4E92F72829A" author="Noguera" box="[744,1086,980,1002]" page="396" pageId="36" pageNumber="37" year="1996">
<bibRefCitation id="EF924B4BFFE76725FD53C4E92F46829A" author="Noguera" box="[744,1034,980,1002]" pageId="36" pageNumber="37" refString="Noguera, F. A. &amp; Chemsak, J. A. (1996) Cerambycidae (Coleoptera). In: Biodiversidad, Taxonomia y Biogeografia de Artropodos de Mexico: Hacia una Sintesis de su conocimiento. Universidad Nacional Autonoma de Mexico, pp. 381 - 409." type="book chapter" year="1996">Noguera &amp; Chemsak, 1996</bibRefCitation>
: 396
</treatmentCitation>
(distr.); Monné, 2002: 24 (host);
<treatmentCitation id="0AA210ABFFE76725FF7CC4CE29468578" author="Ozdikmen" box="[199,522,1010,1032]" page="410" pageId="36" pageNumber="37" year="2009">
<bibRefCitation id="EF924B4BFFE76725FF7CC4CE2A948578" author="Ozdikmen" box="[199,472,1010,1032]" pageId="36" pageNumber="37" refString="Ozdikmen, H. &amp; Turgut, S. (2009) A synopsis of Turkish Vesperinae Mulsant, 1839 and Prioninae Latreille, 1802 (Coleoptera: Cerambycidae). Munis Entomology &amp; Zoology, 4 (2), 402 - 423." type="journal article" year="2009">Özdikmen &amp; Turgut, 2009</bibRefCitation>
: 410
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBC36BAFFE76725FF2CC3092BB5871A" blockId="36.[151,1436,1077,1966]" pageId="36" pageNumber="37">
Original description (
<bibRefCitation id="EF924B4BFFE76725FE3DC3092978853E" author="Chemsak" box="[390,564,1077,1102]" pageId="36" pageNumber="37" refString="Chemsak, J. A. (1979) New species of Neotropical Prioninae (Coleoptera: Cerambycidae). The Coleopterists Bulletin, 33 (1), 125 - 128." type="journal article" year="1979">Chemsak 1979</bibRefCitation>
): “
<emphasis id="B977EAA8FFE76725FDE8C30929C1853E" box="[595,653,1077,1102]" italics="true" pageId="36" pageNumber="37">Male</emphasis>
: Form moderate sized; integument piceous, elytra and lateral margins of pronotum orange brown. Head with front very short, deeply impressed; mandibles moderately long, finely punctate; median impression deep, extending onto neck; punctures rather fine, confluent with rounded pits on vertex; eyes separated on vertex by about length of fourth antennal segment; antennae 12-segmented, reaching to a little beyond middle of elytra, segments slightly produced at apices, segments to sixth finely, sparsely punctate, outer segments finely striate. Pronotum narrower than base of elytra; lateral teeth moderate, anterior pair short, median pair larger, basal margin almost right angles; disk very irregularly, densely punctate, punctures consisting of rounded, elevated protuberances of varying sizes; prosternum barely punctate, very sparsely pubescent; mesosternum very finely, shallowly punctate; metasternum finely, densely punctate, densely clothed with long, erect, yellowish hairs. Scutellum glabrous, finely, moderately densely punctate, rounded behind. Elytra less than twice as long as broad; punctures dense, rugose, confluent, becoming finer toward apex; pubescence absent; each elytron bicostate, costae extending from base and uniting at apical one fourth; apices rounded, sutural angles lightly dentate. Legs slender, sparsely punctate; front tibiae excavated and asperate beneath; posterior tarsi slender, segments finely punctate and sparsely pubescent dorsally, third segment with lobes dentate at apices. Abdomen shining, glabrous, finely, rather sparsely punctate; last sternite emarginate at apex. Length, exclusive of mandibles,
<quantity id="4CFB9B5FFFE76725FF2CC16E2BA7871A" box="[151,235,1618,1642]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="36" pageNumber="37" unit="mm" value="31.0">31 mm</quantity>
.”
</paragraph>
</subSubSection>
<subSubSection id="C3196531FFE76725FF7CC14929A887FE" box="[199,740,1653,1678]" pageId="36" pageNumber="37" type="description">
<paragraph id="8BBC36BAFFE76725FF7CC14929A887FE" blockId="36.[151,1436,1077,1966]" box="[199,740,1653,1678]" pageId="36" pageNumber="37">
<emphasis id="B977EAA8FFE76725FF7CC149294887FE" bold="true" box="[199,516,1653,1678]" pageId="36" pageNumber="37">Geographical distribution.</emphasis>
<collectingCountry id="F314762AFFE76725FDB0C149292C87FE" box="[523,608,1653,1678]" name="Mexico" pageId="36" pageNumber="37">Mexico</collectingCountry>
(Durango).
</paragraph>
</subSubSection>
<subSubSection id="C3196531FFE76725FF7CC1A52F0987C2" box="[199,1093,1689,1714]" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="8BBC36BAFFE76725FF7CC1A52F0987C2" blockId="36.[151,1436,1077,1966]" box="[199,1093,1689,1714]" pageId="36" pageNumber="37">
<emphasis id="B977EAA8FFE76725FF7CC1A52AEB87C2" bold="true" box="[199,423,1689,1714]" pageId="36" pageNumber="37">
<typeStatus id="54B88818FFE76725FF7CC1A52A4F87C2" box="[199,259,1689,1714]" pageId="36" pageNumber="37">Type</typeStatus>
,
<typeStatus id="54B88818FFE76725FEB4C1A52A0E87C2" box="[271,322,1689,1714]" pageId="36" pageNumber="37">type</typeStatus>
locality.
</emphasis>
<typeStatus id="54B88818FFE76725FE16C1A5295987C2" box="[429,533,1689,1714]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
male from
<collectingCountry id="F314762AFFE76725FD2CC1A529A087C2" box="[663,748,1689,1714]" name="Mexico" pageId="36" pageNumber="37">Mexico</collectingCountry>
(Durango), deposited at
<collectionCode id="ED12AE7FFFE76725FBBFC1A62F0E87C2" box="[1028,1090,1690,1714]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="36" pageNumber="37">CNC</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3196531FFE76725FF7CC18129BE86DE" pageId="36" pageNumber="37" type="discussion">
<paragraph id="8BBC36BAFFE76725FF7CC1812878878A" blockId="36.[151,1436,1077,1966]" pageId="36" pageNumber="37">
<emphasis id="B977EAA8FFE76725FF7CC1812A7787A6" bold="true" box="[199,315,1725,1750]" pageId="36" pageNumber="37">Remarks.</emphasis>
We could not examine specimens of this species, but we examined photographs of the male
<typeStatus id="54B88818FFE76725FF2CC1DD2BB5878A" box="[151,249,1761,1786]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
and a female (this latter figured at
<bibRefCitation id="EF924B4BFFE76725FD39C1DD287D878A" author="Bezark" box="[642,817,1761,1786]" pageId="36" pageNumber="37" refString="Bezark, L. G. (2016) New World Cerambycidae Catalog. Available from: https: // apps 2. cdfa. ca. gov / publicApps / plant / bycidDB / wdefault. asp? w = n (accessed on 18 January 2016)" type="journal article" year="2016">Bezark (2016))</bibRefCitation>
.
</paragraph>
<paragraph id="8BBC36BAFFE76725FF7CC0392A198615" blockId="36.[151,1436,1077,1966]" pageId="36" pageNumber="37">
According to
<bibRefCitation id="EF924B4BFFE76725FED7C0392963866E" author="Chemsak" box="[364,559,1797,1822]" pageId="36" pageNumber="37" refString="Chemsak, J. A. (1979) New species of Neotropical Prioninae (Coleoptera: Cerambycidae). The Coleopterists Bulletin, 33 (1), 125 - 128." type="journal article" year="1979">Chemsak (1979)</bibRefCitation>
: “The coloration and sculpturing of the elytra and the unusual protruding punctures of the pronotum make this one of the most distinctive New World species of
<taxonomicName id="4C034D39FFE76725FBD2C0162F8E8631" box="[1129,1218,1834,1857]" class="Insecta" family="Cerambycidae" genus="Prionus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B977EAA8FFE76725FBD2C0162F8E8631" box="[1129,1218,1834,1857]" italics="true" pageId="36" pageNumber="37">Prionus</emphasis>
</taxonomicName>
.” That statement is certainly correct.
</paragraph>
<paragraph id="8BBC36BAFFE76725FF7CC04E29BE86DE" blockId="36.[151,1436,1077,1966]" pageId="36" pageNumber="37">As usual, ventrite I is not totally glabrous, having short, sparse setae laterally. Also ventrites IIV are coarser and more abundantly punctate laterally than centrally.</paragraph>
</subSubSection>
</treatment>
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