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<document id="44D9EAE6F2A69A1EF1A56F3417A2CB62" ID-DOI="10.11646/zootaxa.5529.1.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="14022150" ID-ZooBank="060FCC27-FCE2-451F-BAE6-79A0D8FA8A9F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1730452249895" checkinUser="plazi" docAuthor="Ruengsawang, Nisit, Sangpradub, Narumon, Cubeddu, Tiziana, Pronzato, Roberto &amp; Manconi, Renata" docDate="2024" docId="03AA878EFFC4891EFF781586FC1BF9D2" docLanguage="en" docName="zootaxa.5529.1.7.pdf" docOrigin="Zootaxa 5529 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5529.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Corvospongilla lampaoensis Ruengsawang &amp; Sangpradub &amp; Cubeddu &amp; Pronzato &amp; Manconi 2024, sp. nov." docType="treatment" docUuid="0682FDF8-7972-43B8-B42D-F1AE3989CEF0" docUuidSource="ZooBank" docVersion="2" lastPageNumber="148" masterDocId="FF93FFF6FFC6891AFFEF162FFFCBFFA7" masterDocTitle="Biodiversity assessment of the Lower Mekong Basin: evolutionary novelties in gemmular morphotraits of Genus Corvospongilla (Porifera: Spongillida) with description of a new species from Khorat Plateau, and biogeographic notes" masterLastPageNumber="158" masterPageNumber="144" pageNumber="146" updateTime="1730452876164" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="299B1AE638BC96F7367C202A0EFE22D0">Biodiversity assessment of the Lower Mekong Basin: evolutionary novelties in gemmular morphotraits of Genus Corvospongilla (Porifera: Spongillida) with description of a new species from Khorat Plateau, and biogeographic notes</mods:title>
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<mods:namePart id="BB1D94EB999F711530F65B1BB3805742">Ruengsawang, Nisit</mods:namePart>
<mods:affiliation id="14F907EF27935C109F5A9159AAE0C4AC">Biodiversity and Sustainable Utilization Research Unit, Division of Biology, Faculty of Science and Technology, Rajamangala University of Technology Krungthep, Bangkok, 10120, Thailand.</mods:affiliation>
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<mods:namePart id="25442CF22F4978F042B85F6D35768220">Sangpradub, Narumon</mods:namePart>
<mods:affiliation id="273280D662E9C89AED54B1EB06CA7FD8">Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen, 40002, Thailand.</mods:affiliation>
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<mods:namePart id="65B3C5C7C0ADB32083DEFF4F11DF3197">Cubeddu, Tiziana</mods:namePart>
<mods:affiliation id="92B575E56D0C72FD9F18DA67A01FBEEC">Dipartimento di Medicina Veterinaria, Università di Sassari, Via Vienna 2, 07100 Sassari, Italy.</mods:affiliation>
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<mods:namePart id="9C1B4EC5CB214B71B7FA58C550D1E218">Pronzato, Roberto</mods:namePart>
<mods:affiliation id="D4EB76395FA08620A0751100A963650C">Dipartimento di Scienze della Terra, dellAmbiente e della Vita, Università di Genova, Corso Europa 26, 16132 Genova, Italy.</mods:affiliation>
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<mods:affiliation id="A003B4C151C2E39CCB06602A687B4976">Dipartimento di Medicina Veterinaria, Università di Sassari, Via Vienna 2, 07100 Sassari, Italy.</mods:affiliation>
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<taxonomicName id="4C034D1BFFC48918FF781586FCF3FC64" authority="Ruengsawang &amp; Manconi" authorityName="Ruengsawang &amp; Sangpradub &amp; Cubeddu &amp; Pronzato &amp; Manconi" authorityYear="2024" box="[151,824,937,964]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="species" species="lampaoensis" status="sp. nov.">
<emphasis id="B977EA8AFFC48918FF781586FE26FC63" bold="true" box="[151,493,937,964]" italics="true" pageId="2" pageNumber="146">Corvospongilla lampaoensis</emphasis>
Ruengsawang &amp; Manconi
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<figureCitation id="13382A1DFFC48918FF7815DEFED7FBAC" box="[151,284,1009,1035]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="3.[151,250,1532,1556]" captionStartId-1="6.[151,250,1874,1898]" captionStartId-2="7.[151,250,1546,1570]" captionStartId-3="8.[151,250,767,791]" captionStartId-4="9.[151,250,1547,1571]" captionStartId-5="10.[151,250,1648,1672]" captionTargetBox-0="[268,1320,378,1506]" captionTargetBox-1="[249,1339,319,1850]" captionTargetBox-2="[164,1423,181,1522]" captionTargetBox-3="[215,1371,181,743]" captionTargetBox-4="[164,1423,181,1523]" captionTargetBox-5="[177,1411,309,1624]" captionTargetId-0="figure-103@3.[268,1320,378,1506]" captionTargetId-1="figure-18@6.[249,1339,181,1850]" captionTargetId-2="figure-20@7.[164,1423,181,1522]" captionTargetId-3="figure-18@8.[215,1371,181,743]" captionTargetId-4="figure-20@9.[164,1423,181,1523]" captionTargetId-5="figure-70@10.[177,1411,309,1624]" captionTargetPageId-0="3" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionText-0="FIGURE 1. A, Map of the Lower Mekong hydrographic basin showing the major tributaries in Khorat Plateau, northeast Thailand. Study area of freshwater sponges (Spongillida) at the Lam Pao Reservoir, (Lam Pao River basin), indicated by a red dashed line square. B, Four sampling sites (red) at the Lam Pao Reservoir 1) Na Chueak (Yang Talat District), 2) Lam Pao (Mueang Kalasin District), 3) Phu Din (Mueang Kalasin District), and 4) Lam Pao near Thepsuda Bridge (Nong Bua, Nong Kung Si District). C, Type locality of Corvospongilla lampaoensis sp. nov. near Thepsuda Bridge, site 4 (16°4242.1”N, 103°2700.6”E) Lam Pao Reservoir, 163 m asl, Nong Bua, Nong Kung Si District, Kalasin Province, Thailand." captionText-1="FIGURE 2. Habitat of Corvospongilla lampaoensis sp. nov. (Spongillida) from the Lower Mekong Basin at Lam Pao Thepsuda Bridge site 4 (Figure 1) type locality, in the Lam Pao Reservoir (Kalasin Province, Thailand).AB, Manmade substrata on rafts for fish cage culture. C, Encrusting habitus in dry condition of CNRPORFW 120 holotype in situ on a plastic buoy (black dashed line square, Figure 2C) after sampling of a part (see also Figure 3AB), and paratype CNR-POR-FW 121-12 (black asterisk) on a rubber tire." captionText-2="FIGURE 3. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. SEM micrographs (CF). A, Dry encrusting thin specimen (top view). B, Dry encrusting thin specimen with sessile gemmules at the level of basal spongin plate (bottom view). C, Skeletal meshwork of acanthoxeas megascleres. D, Skeletal meshwork of acanthoxeas megascleres (top view). E, Acanthoxeas megascleres with microspiny surface except at abruptly pointed tips. F, Acanthoxeas megascleres, pseudobirotules microscleres (arrows) and probable larval spicule (asterisk)." captionText-3="FIGURE 4. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNRPORFW 120 from Thailand. A, Light Microscopy micrographs. Skeletal microscleres pseudobirotules with smooth, thin shaft and pseudorotules as curved hooks at tips. B, Scanning Electron Micrographs of microscleres." captionText-4="FIGURE 5. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. Scanning Electron Micrographs. A, Gemmules strongly armed by spicules grouped in a carpet. B, Gemmuloscleres in a dense mosaic-like arrangement at gemmular surface (detail of A). C, Gemmuloscleres acanthostrongyles and acanthostrongyloxeas with variably dense small tubercles and spines. D,Aberrant acanthostrongyles. E, Details of gemmuloscleres ornamentations and tips outlines. F, Megascleres acanthoxeas at the level of scantly developed gemmular cage mixed with gemmuloscleres. Acanthoxeas with scattered, small, conical spines except for tips (diverging from abruptly pointed tips of skeletal acanthoxeas)." captionText-5="FIGURE 6. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. Scanning Electron Micrographs. Gemmular theca architecture of sessile gemmules adhering to basal spongin plate. A, Entire gemmule with few, scattered megascleres (top view). B, Sessile gemmule with scattered thin laminae lining upper portion of theca (white arrow) and smooth, compact inner layer (white arrowhead) and chambered pneumatic layer in basal portion of theca (inner view, cross section). C, Entire sessile gemmule with fragments of basal spongin plate (arrow). D, Gemmular theca (white arrowhead) armed by gemmuloscleres in variably oriented multilayers and scantly developed outer layer and pneumatic layer (cross section); thecamoebae (white arrow). E, Outer layer as a thin pellicle covering spicules (white arrowhead), pneumatic layer absent and inner layer scantly developed (white arrow) in a strongly armed gemmular theca (cross section). F, Pneumatic layer scattered among gemmuloscleres (right) and smooth, compact inner layer of gemmular theca (white arrowhead, cross section). G, Pneumatic layer thin and continuous with dense, minute rounded chambers among few gemmuloscleres in gemmular theca with well-developed outer layer (white arrow, cross section)." figureDoi-0="http://doi.org/10.5281/zenodo.14022154" figureDoi-1="http://doi.org/10.5281/zenodo.14022156" figureDoi-2="http://doi.org/10.5281/zenodo.14022160" figureDoi-3="http://doi.org/10.5281/zenodo.14022164" figureDoi-4="http://doi.org/10.5281/zenodo.14022168" figureDoi-5="http://doi.org/10.5281/zenodo.14022172" httpUri-0="https://zenodo.org/record/14022154/files/figure.png" httpUri-1="https://zenodo.org/record/14022156/files/figure.png" httpUri-2="https://zenodo.org/record/14022160/files/figure.png" httpUri-3="https://zenodo.org/record/14022164/files/figure.png" httpUri-4="https://zenodo.org/record/14022168/files/figure.png" httpUri-5="https://zenodo.org/record/14022172/files/figure.png" pageId="2" pageNumber="146">Figures 16</figureCitation>
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<emphasis id="B977EA8AFFC48918FF781216FEDAFBF4" bold="true" box="[151,273,1081,1107]" pageId="2" pageNumber="146">Materials.</emphasis>
<materialsCitation id="3B6B3CC5FFC48918FEF41216FCD2FB3C" collectingDate="2018-06-14" collectorName="N. Ruengsawang &amp; Thepsuda Bridge" country="Thailand" county="Nong Kung Si District" elevation="163" latitude="16.711695" location="Lam Pao Reservoir" longLatPrecision="1" longitude="103.450165" municipality="Nong Bua" pageId="2" pageNumber="146" specimenCount="1" stateProvince="Kalasin" typeStatus="holotype">
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CNR-POR-FW 120,
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<collectingDate id="EFF9E9B0FFC48918FD931216FD3CFBF4" box="[636,759,1081,1107]" pageId="2" pageNumber="146" value="2018-06-14">14.vi.2018</collectingDate>
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, leg.
<collectorName id="26F6534EFFC48918FCD81215FC37FBF3" box="[823,1020,1082,1108]" pageId="2" pageNumber="146">N. Ruengsawang</collectorName>
, type locality near
<collectorName id="26F6534EFFC48918FB351216FA57FBF4" box="[1242,1436,1081,1107]" pageId="2" pageNumber="146">Thepsuda Bridge</collectorName>
(
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,
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) of
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above sea level
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(asl),
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<collectingRegion id="49C7F87AFFC48918FE0D12AEFD6DFB3C" box="[482,678,1153,1179]" country="Thailand" name="Kalasin" pageId="2" pageNumber="146">Kalasin Province</collectingRegion>
,
<collectingCountry id="F3147608FFC48918FD5D12AEFCD2FB3C" box="[690,793,1153,1179]" name="Thailand" pageId="2" pageNumber="146">Thailand</collectingCountry>
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.
<materialsCitation id="3B6B3CC5FFC48918FCCA12ADFE6AFAAF" collectingDate="2018-06-14" collectorName="N. Ruengsawang. Specimens" country="Thailand" location="Bangkok" pageId="2" pageNumber="146" specimenCount="2" stateProvince="Krung Thep Mahanakhon" typeStatus="paratype">
<typeStatus id="54B8883AFFC48918FCCA12ADFC4CFB3B" box="[805,903,1154,1180]" pageId="2" pageNumber="146" type="paratype">Paratype</typeStatus>
CNR-POR-FW 121-12, dry specimen, ibidem,
<date id="FFBD1058FFC48918FF78128AFEDAFB18" box="[151,273,1189,1215]" pageId="2" pageNumber="146" value="2018-06-14">
<collectingDate id="EFF9E9B0FFC48918FF78128AFEDAFB18" box="[151,273,1189,1215]" pageId="2" pageNumber="146" value="2018-06-14">14.vi.2018</collectingDate>
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, leg.
<collectorName id="26F6534EFFC48918FEA41289FD58FB18" box="[331,659,1189,1216]" pageId="2" pageNumber="146">N. Ruengsawang. Specimens</collectorName>
, slides and SEM stubs are preserved in Collection Nisit Ruengsawang (CNR-POR-FW),
<collectingRegion id="49C7F87AFFC48918FE8812E6FE1BFB44" box="[359,464,1225,1251]" country="Thailand" name="Krung Thep Mahanakhon" pageId="2" pageNumber="146">Bangkok</collectingRegion>
. For
<typeStatus id="54B8883AFFC48918FDE212E5FDB1FB43" box="[525,634,1226,1252]" pageId="2" pageNumber="146" type="topotype">topotypes</typeStatus>
collected in
<collectingDate id="EFF9E9B0FFC48918FCFF12E6FB50FB44" box="[784,1179,1225,1252]" isEnumeration="true" pageId="2" pageNumber="146">
<date id="FFBD1058FFC48918FCFF12E6FC03FB44" box="[784,968,1225,1251]" pageId="2" pageNumber="146" value="2023-11">November 2023</date>
and
<date id="FFBD1058FFC48918FBEC12E5FB50FB44" box="[1027,1179,1225,1252]" pageId="2" pageNumber="146" value="2024-01">January 2024</date>
</collectingDate>
the registration CNR-POR-FW is in progress
</materialsCitation>
.
</paragraph>
<paragraph id="8BBC3698FFC48918FF28133EFCD4F828" blockId="2.[151,1437,1081,2008]" pageId="2" pageNumber="146">
<emphasis id="B977EA8AFFC48918FF28133EFE2EFA8C" bold="true" box="[199,485,1297,1323]" pageId="2" pageNumber="146">Comparative materials.</emphasis>
Genus
<emphasis id="B977EA8AFFC48918FDAF133EFD3EFA8C" box="[576,757,1297,1323]" italics="true" pageId="2" pageNumber="146">
<taxonomicName id="4C034D1BFFC48918FDAF133EFD3AFA8C" authorityName="Annandale" authorityYear="1911" box="[576,753,1297,1323]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="genus">Corvospongilla</taxonomicName>
,
</emphasis>
15 species.
<taxonomicName id="4C034D1BFFC48918FC68133DFB5AFA8C" authority="Poirrier, 1978" authorityName="Poirrier" authorityYear="1978" box="[903,1169,1297,1324]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="species" species="becki">
<emphasis id="B977EA8AFFC48918FC68133DFC2EFA8C" box="[903,997,1297,1324]" italics="true" pageId="2" pageNumber="146">C. becki</emphasis>
<bibRefCitation id="EF924B69FFC48918FC00133EFB5AFA8C" author="Poirrier, M. A." box="[1007,1169,1297,1323]" pageId="2" pageNumber="146" pagination="240 - 243" refId="ref9157" refString="Poirrier, M. A. (1978) Corvospongilla becki n. sp., a new freshwater sponge from Louisiana. Transactions of the American Microscopical Society, 97 (2), 240 - 243. https: // doi. org / 10.2307 / 3225599" type="journal article" year="1978">Poirrier, 1978</bibRefCitation>
</taxonomicName>
,
<collectionCode id="ED12AE5DFFC48918FB71133EFB24FA8C" box="[1182,1263,1297,1323]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="146" type="Museum">USNM</collectionCode>
dry
<typeStatus id="54B8883AFFC48918FAC7133DFA53FA8B" box="[1320,1432,1298,1324]" pageId="2" pageNumber="146" type="topotype">topotypes</typeStatus>
,
<collectionCode id="ED12AE5DFFC48918FF78131AFF09FAE8" box="[151,194,1333,1359]" country="Canada" name="McGill University, Redpath Museum" pageId="2" pageNumber="146" type="Museum">RM</collectionCode>
&amp;
<collectionCode id="ED12AE5DFFC48918FF39131AFEAFFAE8" box="[214,356,1333,1359]" pageId="2" pageNumber="146">RP-FWPOR</collectionCode>
919, 920, 921, Duck Lake,
<collectingRegion id="49C7F87AFFC48918FD22131AFCF5FAE8" box="[717,830,1333,1359]" country="United States of America" name="Louisiana" pageId="2" pageNumber="146">Louisiana</collectingRegion>
,
<collectingCountry id="F3147608FFC48918FCBE131AFC46FAE8" box="[849,909,1333,1359]" name="United States of America" pageId="2" pageNumber="146">USA</collectingCountry>
;
<taxonomicName id="4C034D1BFFC48918FC4F1319FAFFFAE8" authority="(Kirkpatrick, 1908)" baseAuthorityName="Kirkpatrick" baseAuthorityYear="1908" box="[928,1332,1333,1360]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="species" species="burmanica">
<emphasis id="B977EA8AFFC48918FC4F1319FB8AFAE8" box="[928,1089,1333,1360]" italics="true" pageId="2" pageNumber="146">C. burmanica</emphasis>
(
<bibRefCitation id="EF924B69FFC48918FBB7131AFAE6FAE8" author="Kirkpatrick, R." box="[1112,1325,1333,1359]" pageId="2" pageNumber="146" pagination="97 - 99" refId="ref7822" refString="Kirkpatrick, R. (1908) Description of a new variety of Spongilla loricata Weltner. Records of the Indian Museum, 2, 97 - 99. https: // doi. org / 10.26515 / rzsi / v 2 / i 1 / 1908 / 163309" type="journal article" year="1908">Kirkpatrick, 1908</bibRefCitation>
)
</taxonomicName>
,
<collectionCode id="ED12AE5DFFC48918FAA71319FA57FAF7" box="[1352,1436,1334,1360]" pageId="2" pageNumber="146">BMNH</collectionCode>
82.3.22.13, box 6, dry, type,
<collectionCode id="ED12AE5DFFC48918FE051376FDDEFAD4" box="[490,533,1369,1395]" country="Canada" name="McGill University, Redpath Museum" pageId="2" pageNumber="146" type="Museum">RM</collectionCode>
&amp;
<collectionCode id="ED12AE5DFFC48918FDC61376FD7CFAD4" box="[553,695,1369,1395]" pageId="2" pageNumber="146">RP-FWPOR</collectionCode>
420 schizotype,
<collectingCountry id="F3147608FFC48918FC981375FC23FAD3" box="[887,1000,1370,1396]" name="Myanmar" pageId="2" pageNumber="146">Myanmar</collectingCountry>
;
<taxonomicName id="4C034D1BFFC48918FC1C1375FB46FAD4" baseAuthorityName="Kirkpatrick" baseAuthorityYear="1908" box="[1011,1165,1369,1396]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="species" species="burmanica">
<emphasis id="B977EA8AFFC48918FC1C1375FB46FAD4" box="[1011,1165,1369,1396]" italics="true" pageId="2" pageNumber="146">C. burmanica</emphasis>
</taxonomicName>
(?),
<collectionCode id="ED12AE5DFFC48918FB2C1375FADCFAD3" box="[1219,1303,1370,1396]" pageId="2" pageNumber="146">BMNH</collectionCode>
86.10.29.1,
<collectionCode id="ED12AE5DFFC48918FF781352FF09FA30" box="[151,194,1405,1431]" country="Canada" name="McGill University, Redpath Museum" pageId="2" pageNumber="146" type="Museum">RM</collectionCode>
&amp;
<collectionCode id="ED12AE5DFFC48918FF391352FEAFFA30" box="[214,356,1405,1431]" pageId="2" pageNumber="146">RP-FWPOR</collectionCode>
636, River Kuano, NE
<collectingCountry id="F3147608FFC48918FD811352FD60FA30" box="[622,683,1405,1431]" name="India" pageId="2" pageNumber="146">India</collectingCountry>
;
<taxonomicName id="4C034D1BFFC48918FD591351FC37FA30" authority="Annandale, 1911" authorityName="Annandale" authorityYear="1911" box="[694,1020,1405,1432]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="species" species="caunteri">
<emphasis id="B977EA8AFFC48918FD591351FCFFFA30" box="[694,820,1406,1432]" italics="true" pageId="2" pageNumber="146">C. caunteri</emphasis>
<bibRefCitation id="EF924B69FFC48918FCD41352FC37FA30" author="Annandale, N." box="[827,1020,1405,1431]" pageId="2" pageNumber="146" pagination="161 - 251" refId="ref6745" refString="Annandale, N. (1911) Freshwater sponges, hydroids and polyzoa. The Fauna of British India, Including Ceylon and Burma, 2 (369), 161 - 251. https: // doi. org / 10.5962 / bhl. title. 1050" type="journal article" year="1911">Annandale, 1911</bibRefCitation>
</taxonomicName>
,
<collectionCode id="ED12AE5DFFC48918FBE71351FB97FA3F" box="[1032,1116,1406,1432]" pageId="2" pageNumber="146">BMNH</collectionCode>
14.11.24.27 ex-ZEV 4776/
<specimenCount id="9D05FD11FFC48918FA7F1352FF31FA1B" count="7" pageId="2" pageNumber="146" type="generic" typeStatus="paratype">7 paratype</specimenCount>
,
<collectionCode id="ED12AE5DFFC48918FEEA138EFEFBFA1C" box="[261,304,1441,1467]" country="Canada" name="McGill University, Redpath Museum" pageId="2" pageNumber="146" type="Museum">RM</collectionCode>
&amp;
<collectionCode id="ED12AE5DFFC48918FEAB138EFE19FA1C" box="[324,466,1441,1467]" pageId="2" pageNumber="146">RP-FWPOR</collectionCode>
637 schizotype, Lucknow, NE
<collectingCountry id="F3147608FFC48918FCC1138EFCA0FA1C" box="[814,875,1441,1467]" name="India" pageId="2" pageNumber="146">India</collectingCountry>
;
<taxonomicName id="4C034D1BFFC48918FC9A138DFB04FA1C" authority="(Annandale, 1908)" baseAuthorityName="Annandale" baseAuthorityYear="1908" box="[885,1231,1441,1468]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="species" species="lapidosa">
<emphasis id="B977EA8AFFC48918FC9A138DFC3EFA1C" box="[885,1013,1441,1468]" italics="true" pageId="2" pageNumber="146">C. lapidosa</emphasis>
(
<bibRefCitation id="EF924B69FFC48918FBEC138EFB03FA1C" author="Annandale, N." box="[1027,1224,1441,1467]" pageId="2" pageNumber="146" pagination="25 - 28" refId="ref6626" refString="Annandale, N. (1908) Notes on Freshwater Sponges. VIII. Preliminary Notice of a Collection from the Western India, with descriptions of two New Species. Records of the Indian Museum, 2 (1), 25 - 28. https: // doi. org / 10.26515 / rzsi / v 2 / i 1 / 1908 / 163302" type="journal article" year="1908">Annandale, 1908</bibRefCitation>
)
</taxonomicName>
,
<collectionCode id="ED12AE5DFFC48918FB35138DFAE5FA1B" box="[1242,1326,1442,1468]" pageId="2" pageNumber="146">BMNH</collectionCode>
08.2.11.
<specimenCount id="9D05FD11FFC48918FA7D138EFF31FA47" count="1" pageId="2" pageNumber="146" type="generic" typeStatus="paratype">1 paratype</specimenCount>
,
<collectionCode id="ED12AE5DFFC48918FEE913EAFEFAFA78" box="[262,305,1477,1503]" country="Canada" name="McGill University, Redpath Museum" pageId="2" pageNumber="146" type="Museum">RM</collectionCode>
&amp;
<collectionCode id="ED12AE5DFFC48918FEAA13EAFE18FA78" box="[325,467,1477,1503]" pageId="2" pageNumber="146">RP-FWPOR</collectionCode>
638 schizotype, River Godavery Nasik, S-India;
<taxonomicName id="4C034D1BFFC48918FBEF13E9FEE0F9A4" authority="Manconi &amp; Pronzato, 2019 MSNG" authorityName="Manconi &amp; Pronzato" authorityYear="2019" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="species" species="lemuriensis">
<emphasis id="B977EA8AFFC48918FBEF13E9FB69FA78" box="[1024,1186,1477,1504]" italics="true" pageId="2" pageNumber="146">C. lemuriensis</emphasis>
Manconi &amp; Pronzato, 2019 MSNG
</taxonomicName>
60893a
<typeStatus id="54B8883AFFC48918FE7B13C6FE32F9A4" box="[404,505,1513,1539]" pageId="2" pageNumber="146" type="holotype">holotype</typeStatus>
,
<collectionCode id="ED12AE5DFFC48918FDE713C6FDF8F9A4" box="[520,563,1513,1539]" country="Canada" name="McGill University, Redpath Museum" pageId="2" pageNumber="146" type="Museum">RM</collectionCode>
&amp;
<collectionCode id="ED12AE5DFFC48918FDA813C6FD1EF9A4" box="[583,725,1513,1539]" pageId="2" pageNumber="146">RP-FWPOR</collectionCode>
807 schizoholotype,
<collectionCode id="ED12AE5DFFC48918FC3E13C6FC37F9A4" box="[977,1020,1513,1539]" country="Canada" name="McGill University, Redpath Museum" pageId="2" pageNumber="146" type="Museum">RM</collectionCode>
&amp;
<collectionCode id="ED12AE5DFFC48918FBFF13C6FB55F9A4" box="[1040,1182,1513,1539]" pageId="2" pageNumber="146">RP-FWPOR</collectionCode>
804
<typeStatus id="54B8883AFFC48918FB3013C5FA8FF9A3" box="[1247,1348,1514,1540]" pageId="2" pageNumber="146" type="topotype">topotype</typeStatus>
, Farihy Amboromalandi Reservoir,NW
<collectingCountry id="F3147608FFC48918FE151022FD4EF980" box="[506,645,1549,1575]" name="Madagascar" pageId="2" pageNumber="146">Madagascar</collectingCountry>
;
<taxonomicName id="4C034D1BFFC48918FD631021FC78F98F" authority="(Weltner, 1895)" baseAuthorityName="Weltner" baseAuthorityYear="1895" box="[652,947,1549,1576]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="species" species="loricata">
<emphasis id="B977EA8AFFC48918FD631021FCC9F980" box="[652,770,1549,1576]" italics="true" pageId="2" pageNumber="146">C.loricata</emphasis>
(
<bibRefCitation id="EF924B69FFC48918FCE31022FC60F98F" author="Weltner, W." box="[780,939,1549,1576]" pageId="2" pageNumber="146" pagination="114 - 144" refId="ref10584" refString="Weltner, W. (1895) Spongillidenstudien III. Katalog und Verbreitung der bekannten Susswasserschwamme. Archiv fur Naturgeschichte, 61 (1), 114 - 144." type="journal article" year="1895">Weltner, 1895</bibRefCitation>
)
</taxonomicName>
,
<collectionCode id="ED12AE5DFFC48918FC541021FC33F98F" box="[955,1016,1550,1576]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="2" pageNumber="146" type="Museum">ZMB</collectionCode>
2093 SE325-SE37
<specimenCount id="9D05FD11FFC48918FB391022FAE3F98F" box="[1238,1320,1549,1576]" count="41" pageId="2" pageNumber="146" type="generic">41 type</specimenCount>
,
<collectionCode id="ED12AE5DFFC48918FAC01022FA90F980" box="[1327,1371,1549,1575]" country="Canada" name="McGill University, Redpath Museum" pageId="2" pageNumber="146" type="Museum">RM</collectionCode>
&amp;
<collectionCode id="ED12AE5DFFC48918FA9F1022FF3CF9EC" pageId="2" pageNumber="146">RP-FWPOR</collectionCode>
511 schizotype, locality unknown, Africa;
<taxonomicName id="4C034D1BFFC48918FD08101DFB24F9EC" authority="Manconi &amp; Pronzato, 2004" authorityName="Manconi &amp; Pronzato" authorityYear="2004" box="[743,1263,1585,1612]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="species" species="mesopotamica">
<emphasis id="B977EA8AFFC48918FD08101DFC67F9EC" box="[743,940,1586,1612]" italics="true" pageId="2" pageNumber="146">C. mesopotamica</emphasis>
<bibRefCitation id="EF924B69FFC48918FC5A101EFB24F9EC" author="Manconi, R. &amp; Pronzato, R." box="[949,1263,1585,1612]" pageId="2" pageNumber="146" pagination="161 - 189" refId="ref8039" refString="Manconi, R. &amp; Pronzato, R. (2004) The genus Corvospongilla Annandale (Haplosclerida, Spongillina, Spongillidae) with description of a new species from eastern Mesopotamia, Iraq. Archiv fur Hydrobiologie. Supplementband. Monographische Studies, 151, 161 - 189." type="journal article" year="2004">Manconi &amp; Pronzato, 2004</bibRefCitation>
</taxonomicName>
,
<collectionCode id="ED12AE5DFFC48918FB13101EFA86F9EC" box="[1276,1357,1585,1611]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="2" pageNumber="146">MSNG</collectionCode>
51766
<typeStatus id="54B8883AFFC48918FF78107AFF37F9C8" box="[151,252,1621,1647]" pageId="2" pageNumber="146" type="holotype">holotype</typeStatus>
,
<collectionCode id="ED12AE5DFFC48918FEE6107AFEFFF9C8" box="[265,308,1621,1647]" country="Canada" name="McGill University, Redpath Museum" pageId="2" pageNumber="146" type="Museum">RM</collectionCode>
&amp;
<collectionCode id="ED12AE5DFFC48918FEA7107AFE1DF9C8" box="[328,470,1621,1647]" pageId="2" pageNumber="146">RP-FWPOR</collectionCode>
574 schizotype, River
<collectingRegion id="49C7F87AFFC48918FD0D107AFCE4F9C8" box="[738,815,1621,1647]" country="Iraq" name="Diyala" pageId="2" pageNumber="146">Diyala</collectingRegion>
, NE
<collectingCountry id="F3147608FFC48918FC851079FC50F9D7" box="[874,923,1622,1648]" name="Iraq" pageId="2" pageNumber="146">Iraq</collectingCountry>
;
<taxonomicName id="4C034D1BFFC48918FC471079FF19F934" authority="Manconi &amp; Erpenbeck, 2021" authorityName="Manconi &amp; Erpenbeck" authorityYear="2021" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="species" species="moochalabrensis">
<emphasis id="B977EA8AFFC48918FC471079FB42F9C8" box="[936,1161,1621,1648]" italics="true" pageId="2" pageNumber="146">C. moochalabrensis</emphasis>
<bibRefCitation id="EF924B69FFC48918FB7D107AFF19F934" author="Manconi, R. &amp; Erpenbeck, D. &amp; Fromont, J. &amp; Worheide, G. &amp; Pronzato, R." pageId="2" pageNumber="146" pagination="73 - 87" refId="ref8714" refString="Manconi, R., Erpenbeck, D., Fromont, J., Worheide, G. &amp; Pronzato, R. (2021) Discovery of the freshwater sponge genus Corvospongilla Annandale (Porifera: Spongillida) in Australia with the description of a new species and phylogeographic implications. Limnology, 23 (1), 73 - 87. https: // doi. org / 10.1007 / s 10201 - 021 - 00670 - 2" type="journal article" year="2021">Manconi &amp; Erpenbeck, 2021</bibRefCitation>
</taxonomicName>
,
<collectionCode id="ED12AE5DFFC48918FF321055FEEAF933" box="[221,289,1658,1684]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="146" type="Museum">WAM</collectionCode>
Z29235
<typeStatus id="54B8883AFFC48918FE671056FE26F934" box="[392,493,1657,1683]" pageId="2" pageNumber="146" type="holotype">holotype</typeStatus>
,
<collectionCode id="ED12AE5DFFC48918FE171056FDE8F934" box="[504,547,1657,1683]" country="Canada" name="McGill University, Redpath Museum" pageId="2" pageNumber="146" type="Museum">RM</collectionCode>
&amp;
<collectionCode id="ED12AE5DFFC48918FDD81056FD0EF934" box="[567,709,1657,1683]" pageId="2" pageNumber="146">RP-FWPOR</collectionCode>
853 schizotype, Moochalabra Dam, NW
<collectingCountry id="F3147608FFC48918FB771056FAC9F934" box="[1176,1282,1657,1683]" name="Australia" pageId="2" pageNumber="146">Australia</collectingCountry>
;
<taxonomicName id="4C034D1BFFC48918FAE11055FDD8F910" authority="Ruengsawang and Manconi, 2012" authorityName="Ruengsawang and Manconi" authorityYear="2012" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="species" species="siamensis">
<emphasis id="B977EA8AFFC48918FAE11055FA57F934" box="[1294,1436,1658,1684]" italics="true" pageId="2" pageNumber="146">C. siamensis</emphasis>
<bibRefCitation id="EF924B69FFC48918FF7810B1FDD8F910" author="Ruengsawang, N. &amp; Sangpradub, N. &amp; Hanjavanit, C. &amp; Manconi, R." box="[151,531,1693,1720]" pageId="2" pageNumber="146" pagination="47 - 55" refId="ref9688" refString="Ruengsawang, N., Sangpradub, N., Hanjavanit, C. &amp; Manconi, R. (2012) Biodiversity assessment of the Lower Mekong Basin: a new species of Corvospongilla (Porifera: Spongillina: Spongillidae) from Thailand. Zootaxa, 3320 (1), 47 - 55. https: // doi. org / 10.11646 / zootaxa. 3320.1.3" type="journal article" year="2012">Ruengsawang and Manconi, 2012</bibRefCitation>
</taxonomicName>
,
<collectionCode id="ED12AE5DFFC48918FDF210B2FDA5F910" box="[541,622,1693,1719]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="2" pageNumber="146">MSNG</collectionCode>
56533
<typeStatus id="54B8883AFFC48918FD2F10B2FCEEF910" box="[704,805,1693,1719]" pageId="2" pageNumber="146" type="holotype">holotype</typeStatus>
, Ban Huai Sai, Pong River, Mekong Basin, NE
<collectingCountry id="F3147608FFC48918FADE10B2FA53F910" box="[1329,1432,1693,1719]" name="Thailand" pageId="2" pageNumber="146">Thailand</collectingCountry>
;
<taxonomicName id="4C034D1BFFC48918FF7810EDFE5BF97C" authority="(Brien, 1968)" baseAuthorityName="Brien" baseAuthorityYear="1968" box="[151,400,1729,1756]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="species" species="thysi">
<emphasis id="B977EA8AFFC48918FF7810EDFF25F97C" box="[151,238,1729,1756]" italics="true" pageId="2" pageNumber="146">C. thysi</emphasis>
(
<bibRefCitation id="EF924B69FFC48918FF1110EEFE42F97C" author="Brien, P." box="[254,393,1729,1755]" pageId="2" pageNumber="146" pagination="113 - 122" refId="ref6978" refString="Brien, P. (1968) Eponge nouvelle du Cameroun: Potamolepis thysi (n. sp.). Revue de Zoologie et de Botanique Africaines, 78 (1 - 2), 113 - 122." type="journal article" year="1968">Brien, 1968</bibRefCitation>
)
</taxonomicName>
,
<collectionCode id="ED12AE5DFFC48918FE7210EEFE24F97C" box="[413,495,1729,1755]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="2" pageNumber="146" type="Museum">MRAC</collectionCode>
1311 type,
<collectionCode id="ED12AE5DFFC48918FD9810EEFD69F97C" box="[631,674,1729,1755]" country="Canada" name="McGill University, Redpath Museum" pageId="2" pageNumber="146" type="Museum">RM</collectionCode>
&amp;
<collectionCode id="ED12AE5DFFC48918FD5910EEFC8FF97C" box="[694,836,1729,1755]" pageId="2" pageNumber="146">RP-FWPOR</collectionCode>
472 schizotype, Lake Barombi-ma-Mbu,
<collectingCountry id="F3147608FFC48918FACE10EEFA52F97C" box="[1313,1433,1729,1755]" name="Cameroon" pageId="2" pageNumber="146">Cameroun</collectingCountry>
, W-Africa;
<taxonomicName id="4C034D1BFFC48918FEFE10C9FD99F958" authority="Annandale (1910)" authorityName="Annandale" authorityYear="1910" box="[273,594,1765,1792]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="species" species="ultima">
<emphasis id="B977EA8AFFC48918FEFE10C9FEB0F958" box="[273,379,1765,1792]" italics="true" pageId="2" pageNumber="146">C. ultima</emphasis>
<bibRefCitation id="EF924B69FFC48918FE6C10CAFD99F958" author="Annandale, N." box="[387,594,1765,1791]" pageId="2" pageNumber="146" refId="ref6692" refString="Annandale, N. (1910) Notes on freshwater sponges. XII. Description of a new species from Cape Comorin. Records of the Indian Museum, 5, 31. https: // doi. org / 10.26515 / rzsi / v 5 / i 1 / 1910 / 163756" type="journal volume" year="1910">Annandale (1910)</bibRefCitation>
</taxonomicName>
,
<collectionCode id="ED12AE5DFFC48918FDB010C9FD78F8A7" box="[607,691,1766,1792]" pageId="2" pageNumber="146">BMNH</collectionCode>
14.11.24.29 ex-ZEV 4906/7,
<collectionCode id="ED12AE5DFFC48918FBE810CAFBF9F958" box="[1031,1074,1765,1791]" country="Canada" name="McGill University, Redpath Museum" pageId="2" pageNumber="146" type="Museum">RM</collectionCode>
&amp;
<collectionCode id="ED12AE5DFFC48918FBA910CAFB1FF958" box="[1094,1236,1765,1791]" pageId="2" pageNumber="146">RP-FWPOR</collectionCode>
639 fragment, SIndia;
<taxonomicName id="4C034D1BFFC48918FF311125FDFAF883" authority=", BMNH" authorityName="BMNH" box="[222,561,1801,1828]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="variety" species="ultima" variety="spinosa">
<emphasis id="B977EA8AFFC48918FF311125FE8DF884" box="[222,326,1801,1828]" italics="true" pageId="2" pageNumber="146">C. ultima</emphasis>
var.
<emphasis id="B977EA8AFFC48918FE951125FE1BF884" box="[378,464,1802,1827]" italics="true" pageId="2" pageNumber="146">spinosa</emphasis>
, BMNH
</taxonomicName>
14.11.24.30 ex-ZEV 5106/7,
<collectionCode id="ED12AE5DFFC48918FC951126FC6EF884" box="[890,933,1801,1827]" country="Canada" name="McGill University, Redpath Museum" pageId="2" pageNumber="146" type="Museum">RM</collectionCode>
&amp;
<collectionCode id="ED12AE5DFFC48918FC561126FB8CF884" box="[953,1095,1801,1827]" pageId="2" pageNumber="146">RP-FWPOR</collectionCode>
640, Satara District, SE-India;
<taxonomicName id="4C034D1BFFC48918FF781101FDD7F8E0" authority="(Kirkpatrick, 1906)" baseAuthorityName="Kirkpatrick" baseAuthorityYear="1906" box="[151,540,1837,1864]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="species" species="zambesiana">
<emphasis id="B977EA8AFFC48918FF781101FEF0F8E0" box="[151,315,1837,1864]" italics="true" pageId="2" pageNumber="146">C. zambesiana</emphasis>
(
<bibRefCitation id="EF924B69FFC48918FEA61102FDDEF8E0" author="Kirkpatrick, R." box="[329,533,1837,1863]" pageId="2" pageNumber="146" pagination="218 - 227" refId="ref7746" refString="Kirkpatrick, R. (1906) Zoological Results of the Third Tanganyika Expedition conducted by Dr. W. A. Cunnington, 1904 - 1905. Report on the Porifera, with notes on species from the Nile and Zambesi. Proceedings of the Zoological Society of London, 1, 218 - 227. https: // doi. org / 10.1111 / j. 1469 - 7998.1906. tb 08433. x" type="journal article" year="1906">Kirkpatrick, 1906</bibRefCitation>
)
</taxonomicName>
,
<collectionCode id="ED12AE5DFFC48918FDC81101FDB0F8EF" box="[551,635,1838,1864]" pageId="2" pageNumber="146">BMNH</collectionCode>
<date id="FFBD1058FFC48918FD6E1102FD31F8E0" box="[641,762,1837,1863]" pageId="2" pageNumber="146" value="1906-02-28">1906.2.28.</date>
2, 13
<collectionCode id="ED12AE5DFFC48918FCC11102FCAAF8E0" box="[814,865,1837,1863]" pageId="2" pageNumber="146">IIIC</collectionCode>
,
<collectionCode id="ED12AE5DFFC48918FC841102FC5DF8E0" box="[875,918,1837,1863]" country="Canada" name="McGill University, Redpath Museum" pageId="2" pageNumber="146" type="Museum">RM</collectionCode>
&amp;
<collectionCode id="ED12AE5DFFC48918FC441102FBF0F8E0" box="[939,1083,1837,1863]" pageId="2" pageNumber="146">RP-FWPOR</collectionCode>
623 (exDTRG), River Zambezi, E Africa;
<taxonomicName id="4C034D1BFFC48918FEED117EFB94F8CC" authority="Pronzato, Manconi and Samant, 2021 PGDG" authorityName="Pronzato, Manconi and Samant" authorityYear="2021" box="[258,1119,1873,1900]" class="Demospongiae" genus="Palaeocorvospongilla" kingdom="Animalia" order="Spongillida" pageId="2" pageNumber="146" phylum="Porifera" rank="species" species="cretacea">
<emphasis id="B977EA8AFFC48918FEED117EFD94F8CC" box="[258,607,1873,1899]" italics="true" pageId="2" pageNumber="146">Palaeocorvospongilla cretacea</emphasis>
Pronzato,
<bibRefCitation id="EF924B69FFC48918FD39117EFBC3F8CC" author="Manconi, R. &amp; Erpenbeck, D. &amp; Fromont, J. &amp; Worheide, G. &amp; Pronzato, R." box="[726,1032,1873,1899]" pageId="2" pageNumber="146" pagination="73 - 87" refId="ref8714" refString="Manconi, R., Erpenbeck, D., Fromont, J., Worheide, G. &amp; Pronzato, R. (2021) Discovery of the freshwater sponge genus Corvospongilla Annandale (Porifera: Spongillida) in Australia with the description of a new species and phylogeographic implications. Limnology, 23 (1), 73 - 87. https: // doi. org / 10.1007 / s 10201 - 021 - 00670 - 2" type="journal article" year="2021">Manconi and Samant, 2021</bibRefCitation>
PGDG
</taxonomicName>
/
<collectionCode id="ED12AE5DFFC48918FB87117EFB71F8CC" box="[1128,1210,1873,1899]" pageId="2" pageNumber="146">BGVN</collectionCode>
/1/18,2/18,3/18,4/18, Upper Cretaceous (Maastrichtian), Deccan, Central
<collectingCountry id="F3147608FFC48918FD31115AFCD0F828" box="[734,795,1909,1935]" name="India" pageId="2" pageNumber="146">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3196513FFC48919FF2811B6FD9EFEC2" lastPageId="3" lastPageNumber="147" pageId="2" pageNumber="146" type="diagnosis">
<paragraph id="8BBC3698FFC48919FF2811B6FD9EFEC2" blockId="2.[151,1437,1081,2008]" lastBlockId="3.[151,1437,151,357]" lastPageId="3" lastPageNumber="147" pageId="2" pageNumber="146">
<emphasis id="B977EA8AFFC48918FF2811B6FE89F814" bold="true" box="[199,322,1945,1971]" pageId="2" pageNumber="146">Diagnosis.</emphasis>
Megascleres exclusively spiny oxeas in skeletal network and gemmular cage. Gemmuloscleres spiny to tubercled strongyles and strongyloxeas. Gemmular theca sessile. Outer layer scantly developed and strongly armed by multilayered gemmuloscleres to quite developed and armed by few gemmuloscleres. Pneumatic layer thin, continuous with dense, minute rounded chambers and few gemmuloscleres, or as scantly chambered patches among dense multilayered gemmuloscleres, to not chambered pneuma as thin scattered laminae lining the upper portion of theca. Inner layer ranging, in the same theca, from well developed, smooth, compact, lining the basal part of theca to thin, scattered, small laminae irregularly arranged directly adhering to gemmuloscleres layer inner surface at thecas middle-upper portion.
</paragraph>
</subSubSection>
<caption id="DF7C6610FFC58919FF7813D3FB35F96E" ID-DOI="http://doi.org/10.5281/zenodo.14022154" ID-Zenodo-Dep="14022154" httpUri="https://zenodo.org/record/14022154/files/figure.png" pageId="3" pageNumber="147" startId="3.[151,250,1532,1556]" targetBox="[268,1320,378,1506]" targetPageId="3" targetType="figure">
<paragraph id="8BBC3698FFC58919FF7813D3FB35F96E" blockId="3.[151,1436,1532,1737]" pageId="3" pageNumber="147">
<emphasis id="B977EA8AFFC58919FF7813D3FEDBF9B2" bold="true" box="[151,272,1532,1557]" pageId="3" pageNumber="147">FIGURE 1</emphasis>
. A, Map of the Lower Mekong hydrographic basin showing the major tributaries in Khorat Plateau, northeast Thailand. Study area of freshwater sponges (
<taxonomicName id="4C034D1BFFC58919FD84100EFD28F99E" authorityName="Manconi &amp; Pronzato" authorityYear="2002" box="[619,739,1569,1593]" class="Demospongiae" kingdom="Animalia" order="Spongillida" pageId="3" pageNumber="147" phylum="Porifera" rank="order">Spongillida</taxonomicName>
) at the Lam Pao Reservoir, (Lam Pao River basin), indicated by a red dashed line square. B, Four sampling sites (red) at the Lam Pao Reservoir 1) Na Chueak (Yang Talat District), 2) Lam Pao (Mueang Kalasin District), 3) Phu Din (Mueang Kalasin District), and 4) Lam Pao near Thepsuda Bridge (Nong Bua, Nong Kung Si District). C, Type locality of
<taxonomicName id="4C034D1BFFC58919FDBA10A2FCB3F902" authorityName="Ruengsawang &amp; Sangpradub &amp; Cubeddu &amp; Pronzato &amp; Manconi" authorityYear="2024" box="[597,888,1677,1701]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="3" pageNumber="147" phylum="Porifera" rank="species" species="lampaoensis" status="sp. nov.">
<emphasis id="B977EA8AFFC58919FDBA10A2FCB3F902" box="[597,888,1677,1701]" italics="true" pageId="3" pageNumber="147">Corvospongilla lampaoensis</emphasis>
</taxonomicName>
<emphasis id="B977EA8AFFC58919FC9110A2FC04F902" bold="true" box="[894,975,1677,1701]" pageId="3" pageNumber="147">
<taxonomicNameLabel id="A24457F1FFC58919FC9110A2FC04F902" box="[894,975,1677,1701]" pageId="3" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
near Thepsuda Bridge, site 4 (16°4242.1”N, 103°2700.6”E) Lam Pao Reservoir, 163 m asl, Nong Bua, Nong Kung Si District, Kalasin Province, Thailand.
</paragraph>
</caption>
<subSubSection id="C3196513FFC5891EFF781134FD04FB92" lastPageId="4" lastPageNumber="148" pageId="3" pageNumber="147" type="description">
<paragraph id="8BBC3698FFC58919FF781134FEE3F892" blockId="3.[151,296,1819,1845]" box="[151,296,1819,1845]" pageId="3" pageNumber="147">
<emphasis id="B977EA8AFFC58919FF781134FEE3F892" bold="true" box="[151,296,1819,1845]" pageId="3" pageNumber="147">Description</emphasis>
</paragraph>
<paragraph id="8BBC3698FFC5891EFF78114CFD04FB92" blockId="3.[151,1436,1891,2025]" lastBlockId="4.[151,1437,151,1653]" lastPageId="4" lastPageNumber="148" pageId="3" pageNumber="147">
<emphasis id="B977EA8AFFC58919FF78114CFEF1F8DA" bold="true" box="[151,314,1891,1917]" pageId="3" pageNumber="147">Growth form</emphasis>
encrusting, irregular (~4.5 x
<quantity id="4CFB9B7DFFC58919FD78114CFD23F8DA" box="[663,744,1891,1918]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="147" unit="cm" value="8.5">8.5 cm</quantity>
in diameter, ~
<quantity id="4CFB9B7DFFC58919FC7A114CFC2DF8DA" box="[917,998,1891,1917]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="147" unit="cm" value="0.7">0.7 cm</quantity>
in thickness).
<emphasis id="B977EA8AFFC58919FB79114CFB21F8DA" bold="true" box="[1174,1258,1891,1917]" pageId="3" pageNumber="147">Colour</emphasis>
brown to light brown in dry condition (
<figureCitation id="13382A1DFFC58919FE4711A8FDC2F806" box="[424,521,1927,1953]" captionStart="FIGURE 2" captionStartId="6.[151,250,1874,1898]" captionTargetBox="[249,1339,319,1850]" captionTargetId="figure-18@6.[249,1339,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Habitat of Corvospongilla lampaoensis sp. nov. (Spongillida) from the Lower Mekong Basin at Lam Pao Thepsuda Bridge site 4 (Figure 1) type locality, in the Lam Pao Reservoir (Kalasin Province, Thailand).AB, Manmade substrata on rafts for fish cage culture. C, Encrusting habitus in dry condition of CNRPORFW 120 holotype in situ on a plastic buoy (black dashed line square, Figure 2C) after sampling of a part (see also Figure 3AB), and paratype CNR-POR-FW 121-12 (black asterisk) on a rubber tire." figureDoi="http://doi.org/10.5281/zenodo.14022156" httpUri="https://zenodo.org/record/14022156/files/figure.png" pageId="3" pageNumber="147">Figure 2</figureCitation>
) and green-brownish
<emphasis id="B977EA8AFFC58919FD1011A7FC80F806" box="[767,843,1928,1953]" italics="true" pageId="3" pageNumber="147">in vivo</emphasis>
.
<emphasis id="B977EA8AFFC58919FCB611A8FC23F806" bold="true" box="[857,1000,1927,1953]" pageId="3" pageNumber="147">Consistency</emphasis>
hard, fragile in dry condition.
<emphasis id="B977EA8AFFC58919FAAF11A8FA57F806" bold="true" box="[1344,1436,1927,1953]" pageId="3" pageNumber="147">Oscules</emphasis>
inconspicuous in dry specimens.
<emphasis id="B977EA8AFFC58919FDE81184FDA8F862" bold="true" box="[519,611,1963,1989]" pageId="3" pageNumber="147">Surface</emphasis>
densely to irregular and slightly hispidate (
<figureCitation id="13382A1DFFC58919FBAD1184FB68F862" box="[1090,1187,1963,1989]" captionStart="FIGURE 3" captionStartId="7.[151,250,1546,1570]" captionTargetBox="[164,1423,181,1522]" captionTargetId="figure-20@7.[164,1423,181,1522]" captionTargetPageId="7" captionText="FIGURE 3. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. SEM micrographs (CF). A, Dry encrusting thin specimen (top view). B, Dry encrusting thin specimen with sessile gemmules at the level of basal spongin plate (bottom view). C, Skeletal meshwork of acanthoxeas megascleres. D, Skeletal meshwork of acanthoxeas megascleres (top view). E, Acanthoxeas megascleres with microspiny surface except at abruptly pointed tips. F, Acanthoxeas megascleres, pseudobirotules microscleres (arrows) and probable larval spicule (asterisk)." figureDoi="http://doi.org/10.5281/zenodo.14022160" httpUri="https://zenodo.org/record/14022160/files/figure.png" pageId="3" pageNumber="147">Figure 3</figureCitation>
).
<emphasis id="B977EA8AFFC58919FB571184FA57F862" bold="true" box="[1208,1436,1963,1989]" pageId="3" pageNumber="147">Ectosomal skeleton</emphasis>
hispid with no special architecture supporting the dermal membrane.
<emphasis id="B977EA8AFFC58919FC5511E0FB01F84E" bold="true" box="[954,1226,1999,2025]" pageId="3" pageNumber="147">Choanosomal skeleton</emphasis>
as loose isotropic to anisotropic reticulate network of rounded to polygonal meshes, paucispicular to multispicular fibres/tracts with scanty spongin, and ill-defined, irregular skeletal network towards the basal portion (
<figureCitation id="13382A1DFFC2891EFB831694FB1AFF72" box="[1132,1233,187,213]" captionStart="FIGURE 3" captionStartId="7.[151,250,1546,1570]" captionTargetBox="[164,1423,181,1522]" captionTargetId="figure-20@7.[164,1423,181,1522]" captionTargetPageId="7" captionText="FIGURE 3. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. SEM micrographs (CF). A, Dry encrusting thin specimen (top view). B, Dry encrusting thin specimen with sessile gemmules at the level of basal spongin plate (bottom view). C, Skeletal meshwork of acanthoxeas megascleres. D, Skeletal meshwork of acanthoxeas megascleres (top view). E, Acanthoxeas megascleres with microspiny surface except at abruptly pointed tips. F, Acanthoxeas megascleres, pseudobirotules microscleres (arrows) and probable larval spicule (asterisk)." figureDoi="http://doi.org/10.5281/zenodo.14022160" httpUri="https://zenodo.org/record/14022160/files/figure.png" pageId="4" pageNumber="148">Figure 3</figureCitation>
).
<emphasis id="B977EA8AFFC2891EFB061695FA80FF73" bold="true" box="[1257,1355,186,212]" pageId="4" pageNumber="148">Spongin</emphasis>
scanty in skeletal network and well developed at the level of gemmular theca and basal spongin plate (
<figureCitation id="13382A1DFFC2891EFB0C16F0FA9AFF5E" box="[1251,1361,223,249]" captionStart="FIGURE 3" captionStartId="7.[151,250,1546,1570]" captionTargetBox="[164,1423,181,1522]" captionTargetId="figure-20@7.[164,1423,181,1522]" captionTargetPageId="7" captionText="FIGURE 3. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. SEM micrographs (CF). A, Dry encrusting thin specimen (top view). B, Dry encrusting thin specimen with sessile gemmules at the level of basal spongin plate (bottom view). C, Skeletal meshwork of acanthoxeas megascleres. D, Skeletal meshwork of acanthoxeas megascleres (top view). E, Acanthoxeas megascleres with microspiny surface except at abruptly pointed tips. F, Acanthoxeas megascleres, pseudobirotules microscleres (arrows) and probable larval spicule (asterisk)." figureDoi="http://doi.org/10.5281/zenodo.14022160" httpUri="https://zenodo.org/record/14022160/files/figure.png" pageId="4" pageNumber="148">Figures 3</figureCitation>
,
<figureCitation id="13382A1DFFC2891EFA8F16F1FABBFF5E" box="[1376,1392,222,249]" captionStart="FIGURE 5" captionStartId="9.[151,250,1547,1571]" captionTargetBox="[164,1423,181,1523]" captionTargetId="figure-20@9.[164,1423,181,1523]" captionTargetPageId="9" captionText="FIGURE 5. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. Scanning Electron Micrographs. A, Gemmules strongly armed by spicules grouped in a carpet. B, Gemmuloscleres in a dense mosaic-like arrangement at gemmular surface (detail of A). C, Gemmuloscleres acanthostrongyles and acanthostrongyloxeas with variably dense small tubercles and spines. D,Aberrant acanthostrongyles. E, Details of gemmuloscleres ornamentations and tips outlines. F, Megascleres acanthoxeas at the level of scantly developed gemmular cage mixed with gemmuloscleres. Acanthoxeas with scattered, small, conical spines except for tips (diverging from abruptly pointed tips of skeletal acanthoxeas)." figureDoi="http://doi.org/10.5281/zenodo.14022168" httpUri="https://zenodo.org/record/14022168/files/figure.png" pageId="4" pageNumber="148">5</figureCitation>
,
<figureCitation id="13382A1DFFC2891EFA9116F0FA45FF5E" box="[1406,1422,223,249]" captionStart="FIGURE 6" captionStartId="10.[151,250,1648,1672]" captionTargetBox="[177,1411,309,1624]" captionTargetId="figure-70@10.[177,1411,309,1624]" captionTargetPageId="10" captionText="FIGURE 6. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. Scanning Electron Micrographs. Gemmular theca architecture of sessile gemmules adhering to basal spongin plate. A, Entire gemmule with few, scattered megascleres (top view). B, Sessile gemmule with scattered thin laminae lining upper portion of theca (white arrow) and smooth, compact inner layer (white arrowhead) and chambered pneumatic layer in basal portion of theca (inner view, cross section). C, Entire sessile gemmule with fragments of basal spongin plate (arrow). D, Gemmular theca (white arrowhead) armed by gemmuloscleres in variably oriented multilayers and scantly developed outer layer and pneumatic layer (cross section); thecamoebae (white arrow). E, Outer layer as a thin pellicle covering spicules (white arrowhead), pneumatic layer absent and inner layer scantly developed (white arrow) in a strongly armed gemmular theca (cross section). F, Pneumatic layer scattered among gemmuloscleres (right) and smooth, compact inner layer of gemmular theca (white arrowhead, cross section). G, Pneumatic layer thin and continuous with dense, minute rounded chambers among few gemmuloscleres in gemmular theca with well-developed outer layer (white arrow, cross section)." figureDoi="http://doi.org/10.5281/zenodo.14022172" httpUri="https://zenodo.org/record/14022172/files/figure.png" pageId="4" pageNumber="148">6</figureCitation>
).
<emphasis id="B977EA8AFFC2891EFF78172CFE4EFEBA" bold="true" box="[151,389,258,285]" pageId="4" pageNumber="148">Basal spongin plate</emphasis>
armed by megasclere oxeas loosely arranged.
<emphasis id="B977EA8AFFC2891EFC4B172CFBFDFEBA" bold="true" box="[932,1078,259,285]" pageId="4" pageNumber="148">Megascleres</emphasis>
dominant acanthoxeas (137.6 160.1 x
<quantity id="4CFB9B7DFFC2891EFF1C1709FEB8FEE6" box="[243,371,294,321]" metricMagnitude="0" metricUnit="m" metricValue="9.4" metricValueMax="11.5" metricValueMin="7.3" pageId="4" pageNumber="148" unit="m" value="9.4" valueMax="11.5" valueMin="7.3">7.311.5 m</quantity>
, n = 50), slightly curved with abruptly pointed tips and variably dense microspines. Styles and malformations rarely present. Rare slender, spiny oxeas with larger spines probably belonging to larvae (
<figureCitation id="13382A1DFFC2891EFAA91764FF6FFE2E" captionStart="FIGURE 3" captionStartId="7.[151,250,1546,1570]" captionTargetBox="[164,1423,181,1522]" captionTargetId="figure-20@7.[164,1423,181,1522]" captionTargetPageId="7" captionText="FIGURE 3. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. SEM micrographs (CF). A, Dry encrusting thin specimen (top view). B, Dry encrusting thin specimen with sessile gemmules at the level of basal spongin plate (bottom view). C, Skeletal meshwork of acanthoxeas megascleres. D, Skeletal meshwork of acanthoxeas megascleres (top view). E, Acanthoxeas megascleres with microspiny surface except at abruptly pointed tips. F, Acanthoxeas megascleres, pseudobirotules microscleres (arrows) and probable larval spicule (asterisk)." figureDoi="http://doi.org/10.5281/zenodo.14022160" httpUri="https://zenodo.org/record/14022160/files/figure.png" pageId="4" pageNumber="148">Figures 3</figureCitation>
E-F).
<emphasis id="B977EA8AFFC2891EFF071741FEB4FE2F" bold="true" box="[232,383,366,392]" pageId="4" pageNumber="148">Microscleres</emphasis>
few, scattered in the skeletal meshwork, variably curved micropseudobirotules (14.730.4 x 0.91.5 µm, n = 50) with thin smooth shaft bearing smooth pseudorotules at tips (49 µm in diameter) with 46 long, smooth recurved hooks (
<figureCitation id="13382A1DFFC2891EFE021798FDA3FE76" box="[493,616,439,465]" captionStart="FIGURE 3" captionStartId="7.[151,250,1546,1570]" captionTargetBox="[164,1423,181,1522]" captionTargetId="figure-20@7.[164,1423,181,1522]" captionTargetPageId="7" captionText="FIGURE 3. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. SEM micrographs (CF). A, Dry encrusting thin specimen (top view). B, Dry encrusting thin specimen with sessile gemmules at the level of basal spongin plate (bottom view). C, Skeletal meshwork of acanthoxeas megascleres. D, Skeletal meshwork of acanthoxeas megascleres (top view). E, Acanthoxeas megascleres with microspiny surface except at abruptly pointed tips. F, Acanthoxeas megascleres, pseudobirotules microscleres (arrows) and probable larval spicule (asterisk)." figureDoi="http://doi.org/10.5281/zenodo.14022160" httpUri="https://zenodo.org/record/14022160/files/figure.png" pageId="4" pageNumber="148">Figures 3F</figureCitation>
,
<figureCitation id="13382A1DFFC2891EFD9B1798FD4EFE76" box="[628,645,439,465]" captionStart="FIGURE 4" captionStartId="8.[151,250,767,791]" captionTargetBox="[215,1371,181,743]" captionTargetId="figure-18@8.[215,1371,181,743]" captionTargetPageId="8" captionText="FIGURE 4. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNRPORFW 120 from Thailand. A, Light Microscopy micrographs. Skeletal microscleres pseudobirotules with smooth, thin shaft and pseudorotules as curved hooks at tips. B, Scanning Electron Micrographs of microscleres." figureDoi="http://doi.org/10.5281/zenodo.14022164" httpUri="https://zenodo.org/record/14022164/files/figure.png" pageId="4" pageNumber="148">4</figureCitation>
).
<emphasis id="B977EA8AFFC2891EFD751799FC93FE76" bold="true" box="[666,856,438,465]" pageId="4" pageNumber="148">Gemmular cage</emphasis>
as scattered not abundant acanthoxeas megascleres (
<figureCitation id="13382A1DFFC2891EFF7017F4FEC6FE52" box="[159,269,474,501]" captionStart="FIGURE 5" captionStartId="9.[151,250,1547,1571]" captionTargetBox="[164,1423,181,1523]" captionTargetId="figure-20@9.[164,1423,181,1523]" captionTargetPageId="9" captionText="FIGURE 5. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. Scanning Electron Micrographs. A, Gemmules strongly armed by spicules grouped in a carpet. B, Gemmuloscleres in a dense mosaic-like arrangement at gemmular surface (detail of A). C, Gemmuloscleres acanthostrongyles and acanthostrongyloxeas with variably dense small tubercles and spines. D,Aberrant acanthostrongyles. E, Details of gemmuloscleres ornamentations and tips outlines. F, Megascleres acanthoxeas at the level of scantly developed gemmular cage mixed with gemmuloscleres. Acanthoxeas with scattered, small, conical spines except for tips (diverging from abruptly pointed tips of skeletal acanthoxeas)." figureDoi="http://doi.org/10.5281/zenodo.14022168" httpUri="https://zenodo.org/record/14022168/files/figure.png" pageId="4" pageNumber="148">Figure 5F</figureCitation>
) with irregularly distributed small spines (87.5153 x 6.510 µm, n = 11) diverging for outline and spines from skeletal megascleres, and long acanthostrongyles with tubercles (6788.5 x 7.812.2 µm, n = 4).
<emphasis id="B977EA8AFFC2891EFAF217D1FA56FDBF" bold="true" box="[1309,1437,510,536]" pageId="4" pageNumber="148">Gemmules</emphasis>
hemispherical variably flattened at the lower portion to subspherical (400800 µm in diameter); abundant, sessile in large to small dense groups, variably adhering to the basal spongin plate, and sometimes partly sharing the gemmular thecas/cages (
<figureCitation id="13382A1DFFC2891EFE421444FDD0FD22" box="[429,539,619,645]" captionStart="FIGURE 3" captionStartId="7.[151,250,1546,1570]" captionTargetBox="[164,1423,181,1522]" captionTargetId="figure-20@7.[164,1423,181,1522]" captionTargetPageId="7" captionText="FIGURE 3. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. SEM micrographs (CF). A, Dry encrusting thin specimen (top view). B, Dry encrusting thin specimen with sessile gemmules at the level of basal spongin plate (bottom view). C, Skeletal meshwork of acanthoxeas megascleres. D, Skeletal meshwork of acanthoxeas megascleres (top view). E, Acanthoxeas megascleres with microspiny surface except at abruptly pointed tips. F, Acanthoxeas megascleres, pseudobirotules microscleres (arrows) and probable larval spicule (asterisk)." figureDoi="http://doi.org/10.5281/zenodo.14022160" httpUri="https://zenodo.org/record/14022160/files/figure.png" pageId="4" pageNumber="148">Figures 3</figureCitation>
,
<figureCitation id="13382A1DFFC2891EFDC61445FDF2FD22" box="[553,569,618,645]" captionStart="FIGURE 5" captionStartId="9.[151,250,1547,1571]" captionTargetBox="[164,1423,181,1523]" captionTargetId="figure-20@9.[164,1423,181,1523]" captionTargetPageId="9" captionText="FIGURE 5. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. Scanning Electron Micrographs. A, Gemmules strongly armed by spicules grouped in a carpet. B, Gemmuloscleres in a dense mosaic-like arrangement at gemmular surface (detail of A). C, Gemmuloscleres acanthostrongyles and acanthostrongyloxeas with variably dense small tubercles and spines. D,Aberrant acanthostrongyles. E, Details of gemmuloscleres ornamentations and tips outlines. F, Megascleres acanthoxeas at the level of scantly developed gemmular cage mixed with gemmuloscleres. Acanthoxeas with scattered, small, conical spines except for tips (diverging from abruptly pointed tips of skeletal acanthoxeas)." figureDoi="http://doi.org/10.5281/zenodo.14022168" httpUri="https://zenodo.org/record/14022168/files/figure.png" pageId="4" pageNumber="148">5</figureCitation>
,
<figureCitation id="13382A1DFFC2891EFDA81444FD9CFD22" box="[583,599,619,645]" captionStart="FIGURE 6" captionStartId="10.[151,250,1648,1672]" captionTargetBox="[177,1411,309,1624]" captionTargetId="figure-70@10.[177,1411,309,1624]" captionTargetPageId="10" captionText="FIGURE 6. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. Scanning Electron Micrographs. Gemmular theca architecture of sessile gemmules adhering to basal spongin plate. A, Entire gemmule with few, scattered megascleres (top view). B, Sessile gemmule with scattered thin laminae lining upper portion of theca (white arrow) and smooth, compact inner layer (white arrowhead) and chambered pneumatic layer in basal portion of theca (inner view, cross section). C, Entire sessile gemmule with fragments of basal spongin plate (arrow). D, Gemmular theca (white arrowhead) armed by gemmuloscleres in variably oriented multilayers and scantly developed outer layer and pneumatic layer (cross section); thecamoebae (white arrow). E, Outer layer as a thin pellicle covering spicules (white arrowhead), pneumatic layer absent and inner layer scantly developed (white arrow) in a strongly armed gemmular theca (cross section). F, Pneumatic layer scattered among gemmuloscleres (right) and smooth, compact inner layer of gemmular theca (white arrowhead, cross section). G, Pneumatic layer thin and continuous with dense, minute rounded chambers among few gemmuloscleres in gemmular theca with well-developed outer layer (white arrow, cross section)." figureDoi="http://doi.org/10.5281/zenodo.14022172" httpUri="https://zenodo.org/record/14022172/files/figure.png" pageId="4" pageNumber="148">6</figureCitation>
).
<emphasis id="B977EA8AFFC2891EFD811444FD10FD22" bold="true" box="[622,731,619,645]" pageId="4" pageNumber="148">Foramen</emphasis>
rarely evident, rounded with simple collar and a short porous tube.
<emphasis id="B977EA8AFFC2891EFF3614A1FE6EFD0E" bold="true" box="[217,421,654,681]" pageId="4" pageNumber="148">Gemmular theca</emphasis>
bilayered to trilayered with tangential gemmuloscleres variably embedded and oriented and with layers variably developed and spatially arranged (
<figureCitation id="13382A1DFFC2891EFCA7149CFC73FD6A" box="[840,952,690,717]" captionStart="FIGURE 5" captionStartId="9.[151,250,1547,1571]" captionTargetBox="[164,1423,181,1523]" captionTargetId="figure-20@9.[164,1423,181,1523]" captionTargetPageId="9" captionText="FIGURE 5. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. Scanning Electron Micrographs. A, Gemmules strongly armed by spicules grouped in a carpet. B, Gemmuloscleres in a dense mosaic-like arrangement at gemmular surface (detail of A). C, Gemmuloscleres acanthostrongyles and acanthostrongyloxeas with variably dense small tubercles and spines. D,Aberrant acanthostrongyles. E, Details of gemmuloscleres ornamentations and tips outlines. F, Megascleres acanthoxeas at the level of scantly developed gemmular cage mixed with gemmuloscleres. Acanthoxeas with scattered, small, conical spines except for tips (diverging from abruptly pointed tips of skeletal acanthoxeas)." figureDoi="http://doi.org/10.5281/zenodo.14022168" httpUri="https://zenodo.org/record/14022168/files/figure.png" pageId="4" pageNumber="148">Figures 5</figureCitation>
,
<figureCitation id="13382A1DFFC2891EFC28149CFC1CFD6A" box="[967,983,691,717]" captionStart="FIGURE 6" captionStartId="10.[151,250,1648,1672]" captionTargetBox="[177,1411,309,1624]" captionTargetId="figure-70@10.[177,1411,309,1624]" captionTargetPageId="10" captionText="FIGURE 6. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. Scanning Electron Micrographs. Gemmular theca architecture of sessile gemmules adhering to basal spongin plate. A, Entire gemmule with few, scattered megascleres (top view). B, Sessile gemmule with scattered thin laminae lining upper portion of theca (white arrow) and smooth, compact inner layer (white arrowhead) and chambered pneumatic layer in basal portion of theca (inner view, cross section). C, Entire sessile gemmule with fragments of basal spongin plate (arrow). D, Gemmular theca (white arrowhead) armed by gemmuloscleres in variably oriented multilayers and scantly developed outer layer and pneumatic layer (cross section); thecamoebae (white arrow). E, Outer layer as a thin pellicle covering spicules (white arrowhead), pneumatic layer absent and inner layer scantly developed (white arrow) in a strongly armed gemmular theca (cross section). F, Pneumatic layer scattered among gemmuloscleres (right) and smooth, compact inner layer of gemmular theca (white arrowhead, cross section). G, Pneumatic layer thin and continuous with dense, minute rounded chambers among few gemmuloscleres in gemmular theca with well-developed outer layer (white arrow, cross section)." figureDoi="http://doi.org/10.5281/zenodo.14022172" httpUri="https://zenodo.org/record/14022172/files/figure.png" pageId="4" pageNumber="148">6</figureCitation>
).
<emphasis id="B977EA8AFFC2891EFC1F149DFBB4FD6A" bold="true" box="[1008,1151,690,717]" pageId="4" pageNumber="148">Outer layer</emphasis>
from strongly armed by gemmuloscleres multilayers to scantly armed by few gemmuloscleres (
<figureCitation id="13382A1DFFC2891EFC2D14F8FBFAFD56" box="[962,1073,726,753]" captionStart="FIGURE 5" captionStartId="9.[151,250,1547,1571]" captionTargetBox="[164,1423,181,1523]" captionTargetId="figure-20@9.[164,1423,181,1523]" captionTargetPageId="9" captionText="FIGURE 5. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. Scanning Electron Micrographs. A, Gemmules strongly armed by spicules grouped in a carpet. B, Gemmuloscleres in a dense mosaic-like arrangement at gemmular surface (detail of A). C, Gemmuloscleres acanthostrongyles and acanthostrongyloxeas with variably dense small tubercles and spines. D,Aberrant acanthostrongyles. E, Details of gemmuloscleres ornamentations and tips outlines. F, Megascleres acanthoxeas at the level of scantly developed gemmular cage mixed with gemmuloscleres. Acanthoxeas with scattered, small, conical spines except for tips (diverging from abruptly pointed tips of skeletal acanthoxeas)." figureDoi="http://doi.org/10.5281/zenodo.14022168" httpUri="https://zenodo.org/record/14022168/files/figure.png" pageId="4" pageNumber="148">Figures 5</figureCitation>
,
<figureCitation id="13382A1DFFC2891EFBD014F8FB84FD56" box="[1087,1103,727,753]" captionStart="FIGURE 6" captionStartId="10.[151,250,1648,1672]" captionTargetBox="[177,1411,309,1624]" captionTargetId="figure-70@10.[177,1411,309,1624]" captionTargetPageId="10" captionText="FIGURE 6. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. Scanning Electron Micrographs. Gemmular theca architecture of sessile gemmules adhering to basal spongin plate. A, Entire gemmule with few, scattered megascleres (top view). B, Sessile gemmule with scattered thin laminae lining upper portion of theca (white arrow) and smooth, compact inner layer (white arrowhead) and chambered pneumatic layer in basal portion of theca (inner view, cross section). C, Entire sessile gemmule with fragments of basal spongin plate (arrow). D, Gemmular theca (white arrowhead) armed by gemmuloscleres in variably oriented multilayers and scantly developed outer layer and pneumatic layer (cross section); thecamoebae (white arrow). E, Outer layer as a thin pellicle covering spicules (white arrowhead), pneumatic layer absent and inner layer scantly developed (white arrow) in a strongly armed gemmular theca (cross section). F, Pneumatic layer scattered among gemmuloscleres (right) and smooth, compact inner layer of gemmular theca (white arrowhead, cross section). G, Pneumatic layer thin and continuous with dense, minute rounded chambers among few gemmuloscleres in gemmular theca with well-developed outer layer (white arrow, cross section)." figureDoi="http://doi.org/10.5281/zenodo.14022172" httpUri="https://zenodo.org/record/14022172/files/figure.png" pageId="4" pageNumber="148">6</figureCitation>
).
<emphasis id="B977EA8AFFC2891EFB8814F9FAE7FD56" bold="true" box="[1127,1324,726,753]" pageId="4" pageNumber="148">Pneumatic layer</emphasis>
of dense, minute rounded chambers ranging (also in the same theca) from thin and continuous to as small patches in-between gemmuloscleres, or scantly developed to almost absent (
<figureCitation id="13382A1DFFC2891EFCEB1530FCAEFC9E" box="[772,869,799,825]" captionStart="FIGURE 6" captionStartId="10.[151,250,1648,1672]" captionTargetBox="[177,1411,309,1624]" captionTargetId="figure-70@10.[177,1411,309,1624]" captionTargetPageId="10" captionText="FIGURE 6. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. Scanning Electron Micrographs. Gemmular theca architecture of sessile gemmules adhering to basal spongin plate. A, Entire gemmule with few, scattered megascleres (top view). B, Sessile gemmule with scattered thin laminae lining upper portion of theca (white arrow) and smooth, compact inner layer (white arrowhead) and chambered pneumatic layer in basal portion of theca (inner view, cross section). C, Entire sessile gemmule with fragments of basal spongin plate (arrow). D, Gemmular theca (white arrowhead) armed by gemmuloscleres in variably oriented multilayers and scantly developed outer layer and pneumatic layer (cross section); thecamoebae (white arrow). E, Outer layer as a thin pellicle covering spicules (white arrowhead), pneumatic layer absent and inner layer scantly developed (white arrow) in a strongly armed gemmular theca (cross section). F, Pneumatic layer scattered among gemmuloscleres (right) and smooth, compact inner layer of gemmular theca (white arrowhead, cross section). G, Pneumatic layer thin and continuous with dense, minute rounded chambers among few gemmuloscleres in gemmular theca with well-developed outer layer (white arrow, cross section)." figureDoi="http://doi.org/10.5281/zenodo.14022172" httpUri="https://zenodo.org/record/14022172/files/figure.png" pageId="4" pageNumber="148">Figure 6</figureCitation>
).
<emphasis id="B977EA8AFFC2891EFC961530FC36FC9E" bold="true" box="[889,1021,799,825]" pageId="4" pageNumber="148">Inner layer</emphasis>
ranging (also in the same theca) from scattered trabeculae to compact smooth laminae, to multilayered laminae with smooth surface lining the basal part of theca and partly as thin, small trabeculae directly adhering to gemmuloscleres at the middle-upper part of theca (
<figureCitation id="13382A1DFFC2891EFF7015A4FEC8FC02" box="[159,259,907,933]" captionStart="FIGURE 6" captionStartId="10.[151,250,1648,1672]" captionTargetBox="[177,1411,309,1624]" captionTargetId="figure-70@10.[177,1411,309,1624]" captionTargetPageId="10" captionText="FIGURE 6. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. Scanning Electron Micrographs. Gemmular theca architecture of sessile gemmules adhering to basal spongin plate. A, Entire gemmule with few, scattered megascleres (top view). B, Sessile gemmule with scattered thin laminae lining upper portion of theca (white arrow) and smooth, compact inner layer (white arrowhead) and chambered pneumatic layer in basal portion of theca (inner view, cross section). C, Entire sessile gemmule with fragments of basal spongin plate (arrow). D, Gemmular theca (white arrowhead) armed by gemmuloscleres in variably oriented multilayers and scantly developed outer layer and pneumatic layer (cross section); thecamoebae (white arrow). E, Outer layer as a thin pellicle covering spicules (white arrowhead), pneumatic layer absent and inner layer scantly developed (white arrow) in a strongly armed gemmular theca (cross section). F, Pneumatic layer scattered among gemmuloscleres (right) and smooth, compact inner layer of gemmular theca (white arrowhead, cross section). G, Pneumatic layer thin and continuous with dense, minute rounded chambers among few gemmuloscleres in gemmular theca with well-developed outer layer (white arrow, cross section)." figureDoi="http://doi.org/10.5281/zenodo.14022172" httpUri="https://zenodo.org/record/14022172/files/figure.png" pageId="4" pageNumber="148">Figure 6</figureCitation>
).
<emphasis id="B977EA8AFFC2891EFEF415A5FE2BFC03" bold="true" box="[283,480,906,932]" pageId="4" pageNumber="148">Gemmuloscleres</emphasis>
frequently curved (~158°) acanthostrongyles (blunt tips) and acanthostrongyloxeas (acute tips) with ornamentations as small tubercles/spines, irregularly scattered, grouped or singly, along the shaft and denser at tips (
<figureCitation id="13382A1DFFC2891EFE4215FCFDD0FC4A" box="[429,539,978,1005]" captionStart="FIGURE 5" captionStartId="9.[151,250,1547,1571]" captionTargetBox="[164,1423,181,1523]" captionTargetId="figure-20@9.[164,1423,181,1523]" captionTargetPageId="9" captionText="FIGURE 5. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. Scanning Electron Micrographs. A, Gemmules strongly armed by spicules grouped in a carpet. B, Gemmuloscleres in a dense mosaic-like arrangement at gemmular surface (detail of A). C, Gemmuloscleres acanthostrongyles and acanthostrongyloxeas with variably dense small tubercles and spines. D,Aberrant acanthostrongyles. E, Details of gemmuloscleres ornamentations and tips outlines. F, Megascleres acanthoxeas at the level of scantly developed gemmular cage mixed with gemmuloscleres. Acanthoxeas with scattered, small, conical spines except for tips (diverging from abruptly pointed tips of skeletal acanthoxeas)." figureDoi="http://doi.org/10.5281/zenodo.14022168" httpUri="https://zenodo.org/record/14022168/files/figure.png" pageId="4" pageNumber="148">Figures 5</figureCitation>
,
<figureCitation id="13382A1DFFC2891EFDC715FCFDF3FC4A" box="[552,568,979,1005]" captionStart="FIGURE 6" captionStartId="10.[151,250,1648,1672]" captionTargetBox="[177,1411,309,1624]" captionTargetId="figure-70@10.[177,1411,309,1624]" captionTargetPageId="10" captionText="FIGURE 6. Corvospongilla lampaoensis sp. nov. (Spongillida) holotype CNR-POR-FW 120 from Thailand. Scanning Electron Micrographs. Gemmular theca architecture of sessile gemmules adhering to basal spongin plate. A, Entire gemmule with few, scattered megascleres (top view). B, Sessile gemmule with scattered thin laminae lining upper portion of theca (white arrow) and smooth, compact inner layer (white arrowhead) and chambered pneumatic layer in basal portion of theca (inner view, cross section). C, Entire sessile gemmule with fragments of basal spongin plate (arrow). D, Gemmular theca (white arrowhead) armed by gemmuloscleres in variably oriented multilayers and scantly developed outer layer and pneumatic layer (cross section); thecamoebae (white arrow). E, Outer layer as a thin pellicle covering spicules (white arrowhead), pneumatic layer absent and inner layer scantly developed (white arrow) in a strongly armed gemmular theca (cross section). F, Pneumatic layer scattered among gemmuloscleres (right) and smooth, compact inner layer of gemmular theca (white arrowhead, cross section). G, Pneumatic layer thin and continuous with dense, minute rounded chambers among few gemmuloscleres in gemmular theca with well-developed outer layer (white arrow, cross section)." figureDoi="http://doi.org/10.5281/zenodo.14022172" httpUri="https://zenodo.org/record/14022172/files/figure.png" pageId="4" pageNumber="148">6</figureCitation>
). Dominant acanthostrongyles (56104 x 8.512 µm, n = 50), less abundant acanthostrongyloxeas (80.5113 x 7.511.5 µm, n = 50). Rare acanthoxeas (92119 x 710 µm, n = 11) frequently curved (~162°). Malformations frequently present.
</paragraph>
</subSubSection>
<subSubSection id="C3196513FFC2891EFF281210FC1BF9D2" pageId="4" pageNumber="148" type="etymology">
<paragraph id="8BBC3698FFC2891EFF281210FB1EFBDA" blockId="4.[151,1437,151,1653]" pageId="4" pageNumber="148">
<emphasis id="B977EA8AFFC2891EFF281210FE85FBFE" bold="true" box="[199,334,1087,1113]" pageId="4" pageNumber="148">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C034D1BFFC2891EFDD01210FD00FBFE" authorityName="Ruengsawang &amp; Sangpradub &amp; Cubeddu &amp; Pronzato &amp; Manconi" authorityYear="2024" box="[575,715,1087,1113]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="4" pageNumber="148" phylum="Porifera" rank="species" species="lampaoensis">
<emphasis id="B977EA8AFFC2891EFDD01210FD00FBFE" box="[575,715,1087,1113]" italics="true" pageId="4" pageNumber="148">lampaoensis</emphasis>
</taxonomicName>
is derived from the Lam Pao Reservoir, where
<taxonomicName id="4C034D1BFFC2891EFB011210FEE8FBDA" authorityName="Ruengsawang &amp; Sangpradub &amp; Cubeddu &amp; Pronzato &amp; Manconi" authorityYear="2024" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="4" pageNumber="148" phylum="Porifera" rank="species" species="lampaoensis" status="sp. nov.">
<emphasis id="B977EA8AFFC2891EFB011210FEE8FBDA" italics="true" pageId="4" pageNumber="148">Corvospongilla lampaoensis</emphasis>
</taxonomicName>
<emphasis id="B977EA8AFFC2891EFEC5124CFE4FFBDA" bold="true" box="[298,388,1123,1149]" pageId="4" pageNumber="148">
<taxonomicNameLabel id="A24457F1FFC2891EFEC5124CFE4FFBDA" box="[298,388,1123,1149]" pageId="4" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was discovered near the Thepsuda Bridge (
<typeStatus id="54B8883AFFC2891EFC83124CFC55FBDA" box="[876,926,1123,1149]" pageId="4" pageNumber="148">type</typeStatus>
locality). Gender feminine.
</paragraph>
<paragraph id="8BBC3698FFC2891EFF2812A9FBD4FA42" blockId="4.[151,1437,151,1653]" pageId="4" pageNumber="148">
<emphasis id="B977EA8AFFC2891EFF2812A9FE09FB06" bold="true" box="[199,450,1158,1185]" pageId="4" pageNumber="148">Habitat and ecology.</emphasis>
The population of
<taxonomicName id="4C034D1BFFC2891EFD4C12A8FC2CFB06" authorityName="Ruengsawang &amp; Sangpradub &amp; Cubeddu &amp; Pronzato &amp; Manconi" authorityYear="2024" box="[675,999,1159,1185]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="4" pageNumber="148" phylum="Porifera" rank="species" species="lampaoensis" status="sp. nov.">
<emphasis id="B977EA8AFFC2891EFD4C12A8FC2CFB06" box="[675,999,1159,1185]" italics="true" pageId="4" pageNumber="148">Corvospongilla lampaoensis</emphasis>
</taxonomicName>
<emphasis id="B977EA8AFFC2891EFC1E12A8FB85FB06" bold="true" box="[1009,1102,1159,1185]" pageId="4" pageNumber="148">
<taxonomicNameLabel id="A24457F1FFC2891EFC1E12A8FB85FB06" box="[1009,1102,1159,1185]" pageId="4" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
near the Thepsuda Bridge is dominant in lentic, brown, eutrophic shallow water, as large patches with encrusting habitus of variable thickness on hard manmade substrata e.g., fish traps and fish farming (Tilapia) cages structures i.e., plastic buoy, nylon rope, netting, rubble tire (
<figureCitation id="13382A1DFFC2891EFE6F12DCFE2DFAAA" box="[384,486,1267,1293]" captionStart="FIGURE 2" captionStartId="6.[151,250,1874,1898]" captionTargetBox="[249,1339,319,1850]" captionTargetId="figure-18@6.[249,1339,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Habitat of Corvospongilla lampaoensis sp. nov. (Spongillida) from the Lower Mekong Basin at Lam Pao Thepsuda Bridge site 4 (Figure 1) type locality, in the Lam Pao Reservoir (Kalasin Province, Thailand).AB, Manmade substrata on rafts for fish cage culture. C, Encrusting habitus in dry condition of CNRPORFW 120 holotype in situ on a plastic buoy (black dashed line square, Figure 2C) after sampling of a part (see also Figure 3AB), and paratype CNR-POR-FW 121-12 (black asterisk) on a rubber tire." figureDoi="http://doi.org/10.5281/zenodo.14022156" httpUri="https://zenodo.org/record/14022156/files/figure.png" pageId="4" pageNumber="148">Figure 2</figureCitation>
). Sponges were not found, until now, on the rare hard natural substrata along the coastline. The site is characterised by notable seasonal variations of water level as also suggested by the persistent presence of abundant gemmules during sampling replicas. Rafts for fish cage culture are seasonally managed and moved within the basin by fishermen according to the weather and water level variations (
<figureCitation id="13382A1DFFC2891EFB981370FB12FADE" box="[1143,1241,1375,1401]" captionStart="FIGURE 2" captionStartId="6.[151,250,1874,1898]" captionTargetBox="[249,1339,319,1850]" captionTargetId="figure-18@6.[249,1339,181,1850]" captionTargetPageId="6" captionText="FIGURE 2. Habitat of Corvospongilla lampaoensis sp. nov. (Spongillida) from the Lower Mekong Basin at Lam Pao Thepsuda Bridge site 4 (Figure 1) type locality, in the Lam Pao Reservoir (Kalasin Province, Thailand).AB, Manmade substrata on rafts for fish cage culture. C, Encrusting habitus in dry condition of CNRPORFW 120 holotype in situ on a plastic buoy (black dashed line square, Figure 2C) after sampling of a part (see also Figure 3AB), and paratype CNR-POR-FW 121-12 (black asterisk) on a rubber tire." figureDoi="http://doi.org/10.5281/zenodo.14022156" httpUri="https://zenodo.org/record/14022156/files/figure.png" pageId="4" pageNumber="148">Figure 2</figureCitation>
). Aquatic insects, bryozoans, diatoms, and testate thecamoebae were found associated with sponges. Other sponges, with different growth forms, colour and spiculation, were also collected from the Thepsuda Bridge and from the other three sampling sites along the Lam Pao reservoir coastline (identification in progress).
</paragraph>
<paragraph id="8BBC3698FFC2891EFF2813C1FC1BF9D2" blockId="4.[151,1437,151,1653]" pageId="4" pageNumber="148">
<emphasis id="B977EA8AFFC2891EFF2813C1FE26F9AF" bold="true" box="[199,493,1518,1544]" pageId="4" pageNumber="148">Geographic distribution.</emphasis>
<taxonomicName id="4C034D1BFFC2891EFE1D13C0FCF9F9AE" authorityName="Ruengsawang &amp; Sangpradub &amp; Cubeddu &amp; Pronzato &amp; Manconi" authorityYear="2024" box="[498,818,1519,1545]" class="Demospongiae" family="Spongillidae" genus="Corvospongilla" kingdom="Animalia" order="Spongillida" pageId="4" pageNumber="148" phylum="Porifera" rank="species" species="lampaoensis" status="sp. nov.">
<emphasis id="B977EA8AFFC2891EFE1D13C0FCF9F9AE" box="[498,818,1519,1545]" italics="true" pageId="4" pageNumber="148">Corvospongilla lampaoensis</emphasis>
</taxonomicName>
<emphasis id="B977EA8AFFC2891EFCD813C0FC44F9AE" bold="true" box="[823,911,1519,1545]" pageId="4" pageNumber="148">
<taxonomicNameLabel id="A24457F1FFC2891EFCD813C0FC44F9AE" box="[823,911,1519,1545]" pageId="4" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<materialsCitation id="3B6B3CC5FFC2891EFC7B13C0FC07F9D2" country="Thailand" county="Indochina's" latitude="16.711695" location="Lam Pao Reservoir" longLatPrecision="1" longitude="103.450165" municipality="Mekong Basin" pageId="4" pageNumber="148" specimenCount="1" stateProvince="Kalasin">
is only known, until now, from the
<typeStatus id="54B8883AFFC2891EFAFC13C0FA88F9AE" box="[1299,1347,1519,1545]" pageId="4" pageNumber="148">type</typeStatus>
locality near Thepsuda Bridge (
<geoCoordinate id="EE37505FFFC2891EFE4D103CFD88F98A" box="[418,579,1555,1581]" degrees="16" direction="north" minutes="42" orientation="latitude" pageId="4" pageNumber="148" precision="1" seconds="42.1" value="16.711695">16°4242.1”N</geoCoordinate>
,
<geoCoordinate id="EE37505FFFC2891EFDBF103CFD36F98A" box="[592,765,1555,1581]" degrees="103" direction="east" minutes="27" orientation="longitude" pageId="4" pageNumber="148" precision="1" seconds="00.6" value="103.450165">103°2700.6”E</geoCoordinate>
),
<location id="8EDC6043FFC2891EFCFD103CFC3BF98A" LSID="urn:lsid:plazi:treatment:03AA878EFFC4891EFF781586FC1BF9D2:8EDC6043FFC2891EFCFD103CFC3BF98A" box="[786,1008,1555,1581]" country="Thailand" county="Indochina's" latitude="16.711695" longLatPrecision="1" longitude="103.450165" municipality="Mekong Basin" name="Lam Pao Reservoir" pageId="4" pageNumber="148" stateProvince="Kalasin">Lam Pao Reservoir</location>
,
<location id="8EDC6043FFC2891EFC13103CFB52F98A" LSID="urn:lsid:plazi:treatment:03AA878EFFC4891EFF781586FC1BF9D2:8EDC6043FFC2891EFC13103CFB52F98A" box="[1020,1177,1555,1581]" country="Thailand" county="Indochina's" latitude="16.711695" longLatPrecision="1" longitude="103.450165" municipality="Mekong Basin" name="Korat Plateau" pageId="4" pageNumber="148" stateProvince="Kalasin">Korat Plateau</location>
,
<collectingRegion id="49C7F87AFFC2891EFB49103CFAA1F98A" box="[1190,1386,1555,1581]" country="Thailand" name="Kalasin" pageId="4" pageNumber="148">Kalasin Province</collectingRegion>
, NE
<collectingCountry id="F3147608FFC2891EFF781018FF30F9F6" box="[151,251,1591,1617]" name="Thailand" pageId="4" pageNumber="148">Thailand</collectingCountry>
(
<figureCitation id="13382A1DFFC2891EFEE31018FEA6F9F6" box="[268,365,1591,1617]" captionStart="FIGURE 1" captionStartId="3.[151,250,1532,1556]" captionTargetBox="[268,1320,378,1506]" captionTargetId="figure-103@3.[268,1320,378,1506]" captionTargetPageId="3" captionText="FIGURE 1. A, Map of the Lower Mekong hydrographic basin showing the major tributaries in Khorat Plateau, northeast Thailand. Study area of freshwater sponges (Spongillida) at the Lam Pao Reservoir, (Lam Pao River basin), indicated by a red dashed line square. B, Four sampling sites (red) at the Lam Pao Reservoir 1) Na Chueak (Yang Talat District), 2) Lam Pao (Mueang Kalasin District), 3) Phu Din (Mueang Kalasin District), and 4) Lam Pao near Thepsuda Bridge (Nong Bua, Nong Kung Si District). C, Type locality of Corvospongilla lampaoensis sp. nov. near Thepsuda Bridge, site 4 (16°4242.1”N, 103°2700.6”E) Lam Pao Reservoir, 163 m asl, Nong Bua, Nong Kung Si District, Kalasin Province, Thailand." figureDoi="http://doi.org/10.5281/zenodo.14022154" httpUri="https://zenodo.org/record/14022154/files/figure.png" pageId="4" pageNumber="148">Figure 1</figureCitation>
).
<location id="8EDC6043FFC2891EFE681018FE78F9F6" LSID="urn:lsid:plazi:treatment:03AA878EFFC4891EFF781586FC1BF9D2:8EDC6043FFC2891EFE681018FE78F9F6" box="[391,435,1591,1617]" country="Thailand" county="Indochina's" latitude="16.711695" longLatPrecision="1" longitude="103.450165" municipality="Mekong Basin" name="The" pageId="4" pageNumber="148" stateProvince="Kalasin">The</location>
species is endemic to the
<collectingMunicipality id="6BD8ACE2FFC2891EFD091018FC5BF9F6" box="[742,912,1591,1617]" pageId="4" pageNumber="148">Mekong Basin</collectingMunicipality>
, although more sampling effort is required to verify its geographic range also in other
<collectingCounty id="62DD4E14FFC2891EFDB21074FD2BF9D2" box="[605,736,1627,1653]" pageId="4" pageNumber="148">Indochinas</collectingCounty>
hydrographic basins
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>