548 lines
75 KiB
XML
548 lines
75 KiB
XML
<document id="FC7F33DD9172282D397C56F9F4D60E9A" ID-CLB-Dataset="270810" ID-DOI="10.11646/zootaxa.5352.1.5" ID-GBIF-Dataset="0d2702de-5255-4f93-b43f-35d32ba1b042" ID-ISSN="1175-5326" ID-Zenodo-Dep="8407019" ID-ZooBank="3A7DEF13-AEE8-48C5-A281-69362CD3B3AE" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1696427592721" checkinUser="plazi" docAuthor="Gurung, Mer Man & Phan, Quoc Toan" docDate="2023" docId="03AA7D6EFFE9FFF4FF74736DFB7207CB" docLanguage="en" docName="zootaxa.5352.1.5.pdf" docOrigin="Zootaxa 5352 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5352.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protosticta samtsensis Gurung & Phan 2023, sp. nov." docType="treatment" docVersion="6" lastPageNumber="142" masterDocId="FF930516FFE8FFF1FFE3767AFFF40001" masterDocTitle="Description of Protosticta samtsensis sp. nov. (Odonata: Zygoptera: Platystictidae) from Samtse district, Bhutan" masterLastPageNumber="144" masterPageNumber="137" pageNumber="138" updateTime="1699693814341" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="86F5CBFBF7A3DF513B380737B2868238">Description of Protosticta samtsensis sp. nov. (Odonata: Zygoptera: Platystictidae) from Samtse district, Bhutan</mods:title>
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<mods:name id="B120AD61A598B9CCC4215E63AFEAA6C2" type="personal">
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<mods:namePart id="895B0DEB0709F1B6D469056815C44ABB">Gurung, Mer Man</mods:namePart>
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<mods:affiliation id="98DF834499C9353B79E6520A2959CAA0">College of Natural Resources, Department of Forest Science, Royal University of Bhutan</mods:affiliation>
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<mods:namePart id="509FB0F775929B932AEC380384BD288B">Phan, Quoc Toan</mods:namePart>
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<mods:affiliation id="5F0C7759B51406DA57D0A34827A3FC8A">The Center for Entomology & Parasitology Research, College of Medicine and Pharmacy, Duy Tan University, Da Nang, Vietnam pqtoan 84 @ gmail. com; phanquoctoan 1 @ dtu. edu. vn; https: // orcid. org / 0000 - 0002 - 3154 - 6546</mods:affiliation>
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<mods:nameIdentifier id="8EDEAECD0D931384D8F98A59389458CF" type="email">pqtoan84@gmail.com</mods:nameIdentifier>
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<mods:title id="046EF86DB9D8D2C81D40875EC24250CF">Zootaxa</mods:title>
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<mods:date id="E24AFB9132AD1E36EAD5BACC96E42919">2023</mods:date>
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<mods:number id="120F66B2EC3B077C3B1022616DDF757D">2023-10-02</mods:number>
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<treatment id="03AA7D6EFFE9FFF4FF74736DFB7207CB" ID-DOI="http://doi.org/10.5281/zenodo.8412178" ID-GBIF-Taxon="213730658" ID-Zenodo-Dep="8412178" LSID="urn:lsid:plazi:treatment:03AA7D6EFFE9FFF4FF74736DFB7207CB" httpUri="http://treatment.plazi.org/id/03AA7D6EFFE9FFF4FF74736DFB7207CB" lastPageId="5" lastPageNumber="142" pageId="1" pageNumber="138">
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<paragraph id="8BBCCC78FFE9FFF0FF74736DFDF30530" blockId="1.[151,519,1303,1366]" box="[151,519,1303,1329]" pageId="1" pageNumber="138">
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<heading id="D0F47B14FFE9FFF0FF74736DFDF30530" bold="true" box="[151,519,1303,1329]" fontSize="11" level="1" pageId="1" pageNumber="138" reason="1">
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<emphasis id="B977106AFFE9FFF0FF74736DFDF30530" bold="true" box="[151,519,1303,1329]" pageId="1" pageNumber="138">
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<taxonomicName id="4C03B7FBFFE9FFF0FF74736DFE560530" ID-CoL="BTLXF" authority="Gurung & Phan, 2023" authorityName="Gurung & Phan" authorityYear="2023" box="[151,418,1303,1329]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="samtsensis" status="sp. nov.">
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<emphasis id="B977106AFFE9FFF0FF74736DFE560530" bold="true" box="[151,418,1303,1329]" italics="true" pageId="1" pageNumber="138">Protosticta samtsensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A244AD11FFE9FFF0FE4A736DFDF30530" box="[425,519,1303,1329]" pageId="1" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3199FF3FFE9FFF0FF747341FE750554" box="[151,385,1339,1366]" pageId="1" pageNumber="138" type="description">
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<paragraph id="8BBCCC78FFE9FFF0FF747341FE750554" blockId="1.[151,519,1303,1366]" box="[151,385,1339,1366]" pageId="1" pageNumber="138">
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(
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<figureCitation id="1338D0FDFFE9FFF0FF7C7341FEDC0557" box="[159,296,1339,1366]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="1.[151,250,1992,2016]" captionStartId-1="2.[151,250,1361,1385]" captionStartId-2="3.[151,250,1204,1228]" captionStartId-3="4.[151,250,1710,1734]" captionTargetBox-1="[151,1436,181,1337]" captionTargetBox-2="[152,1436,181,1181]" captionTargetBox-3="[161,1426,319,1677]" captionTargetId-0="figure-91@1.[340,1247,1389,1969]" captionTargetId-1="figure-18@2.[151,1436,181,1337]" captionTargetId-2="figure-17@3.[151,1436,181,1181]" captionTargetId-3="figure-66@4.[151,1436,309,1687]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 2. Protosticta samtsensis sp. nov., holotype ♁ alive in the wild." captionText-1="FIGURE 3. Habitus of Protosticta samtsensis sp. nov. (A), holotype ♁ & (B), paratype ♀." captionText-2="FIGURE 4. Head and prothorax of Protosticta samtsensis sp. nov. (A, B), holotype ♁ & (C, D), paratype ♀." captionText-3="FIGURE 5. Wings and genital ligula of Protosticta samtsensis sp. nov. (A), wings holotype ♁; (B), wings paratype ♀; (C), genital ligula of holotype ♁." figureDoi-0="http://doi.org/10.5281/zenodo.8412176" figureDoi-1="http://doi.org/10.5281/zenodo.8407023" figureDoi-2="http://doi.org/10.5281/zenodo.8407025" figureDoi-3="http://doi.org/10.5281/zenodo.8407027" httpUri-0="https://zenodo.org/record/8412176/files/figure.png" httpUri-1="https://zenodo.org/record/8407023/files/figure.png" httpUri-2="https://zenodo.org/record/8407025/files/figure.png" httpUri-3="https://zenodo.org/record/8407027/files/figure.png" pageId="1" pageNumber="138">Figures 2–5</figureCitation>
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,
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<figureCitation id="1338D0FDFFE9FFF0FED77341FE8D0554" box="[308,377,1339,1365]" captionStart="FIGURE 6" captionStartId="5.[152,255,745,769]" captionTargetBox="[340,1247,345,722]" captionTargetId="figure-76@5.[340,1247,345,722]" captionTargetPageId="5" captionText="FIGURE 6. Anal appendages of Protosticta samtsensis sp. nov., paratype ♀: (A) lateral view, (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8407029" httpUri="https://zenodo.org/record/8407029/files/figure.png" pageId="1" pageNumber="138">6A–D</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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<caption id="DF7C9CF0FFE9FFF0FF7471B2FC7707E1" ID-DOI="http://doi.org/10.5281/zenodo.8412176" ID-Zenodo-Dep="8412176" box="[151,899,1992,2019]" httpUri="https://zenodo.org/record/8412176/files/figure.png" pageId="1" pageNumber="138" startId="1.[151,250,1992,2016]" targetBox="[340,1247,1389,1969]" targetPageId="1" targetType="figure">
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<paragraph id="8BBCCC78FFE9FFF0FF7471B2FC7707E1" blockId="1.[151,899,1992,2019]" box="[151,899,1992,2019]" pageId="1" pageNumber="138">
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<emphasis id="B977106AFFE9FFF0FF7471B2FEE707E1" bold="true" box="[151,275,1992,2016]" pageId="1" pageNumber="138">FIGURE 2.</emphasis>
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||
<taxonomicName id="4C03B7FBFFE9FFF0FEFA71B3FE0F07E1" authority="Gurung & Phan, 2023" authorityName="Gurung & Phan" authorityYear="2023" box="[281,507,1993,2016]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="samtsensis" status="sp. nov.">
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||
<emphasis id="B977106AFFE9FFF0FEFA71B3FE0F07E1" box="[281,507,1993,2016]" italics="true" pageId="1" pageNumber="138">Protosticta samtsensis</emphasis>
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</taxonomicName>
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||
<emphasis id="B977106AFFE9FFF0FDE171B3FDAD07E0" bold="true" box="[514,601,1993,2017]" pageId="1" pageNumber="138">
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||
<taxonomicNameLabel id="A244AD11FFE9FFF0FDE171B3FDA107E0" box="[514,597,1993,2017]" pageId="1" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
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,
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</emphasis>
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holotype ♁ alive in the wild.
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</paragraph>
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||
</caption>
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||
<caption id="DF7C9CF0FFEAFFF3FF74732BFBC50568" ID-DOI="http://doi.org/10.5281/zenodo.8407023" ID-Zenodo-Dep="8407023" box="[151,1073,1361,1388]" httpUri="https://zenodo.org/record/8407023/files/figure.png" pageId="2" pageNumber="139" startId="2.[151,250,1361,1385]" targetBox="[151,1436,181,1337]" targetPageId="2" targetType="figure">
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<paragraph id="8BBCCC78FFEAFFF3FF74732BFBC50568" blockId="2.[151,1073,1361,1388]" box="[151,1073,1361,1388]" pageId="2" pageNumber="139">
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||
<emphasis id="B977106AFFEAFFF3FF74732BFEE70568" bold="true" box="[151,275,1361,1385]" pageId="2" pageNumber="139">FIGURE 3.</emphasis>
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Habitus of
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<taxonomicName id="4C03B7FBFFEAFFF3FE697328FD980568" authority="Gurung & Phan, 2023" authorityName="Gurung & Phan" authorityYear="2023" box="[394,620,1362,1385]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="samtsensis" status="sp. nov.">
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<emphasis id="B977106AFFEAFFF3FE697328FD980568" box="[394,620,1362,1385]" italics="true" pageId="2" pageNumber="139">Protosticta samtsensis</emphasis>
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</taxonomicName>
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<emphasis id="B977106AFFEAFFF3FD90732BFD370568" bold="true" box="[627,707,1361,1385]" pageId="2" pageNumber="139">
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<taxonomicNameLabel id="A244AD11FFEAFFF3FD90732BFD370568" box="[627,707,1361,1385]" pageId="2" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(A), holotype ♁ & (B), paratype ♀.
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</paragraph>
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||
</caption>
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||
<subSubSection id="C3199FF3FFEAFFF3FF7473E2FA73061C" pageId="2" pageNumber="139" type="materials_examined">
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<paragraph id="8BBCCC78FFEAFFF3FF7473E2FCBD05D4" blockId="2.[151,1437,1431,2033]" pageId="2" pageNumber="139">
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<materialsCitation id="3B6BC625FFEAFFF3FF7473E2FCBD05D4" ID-GBIF-Occurrence="4421512301" collectingDate="2022-08-12" collectorName="Mer Man Gurung" country="Bhutan" elevation="1264" latitude="27.14267" location="Samtse District" pageId="2" pageNumber="139" specimenCode="♁" specimenCount="1" stateProvince="Samtse District" typeStatus="holotype">
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<emphasis id="B977106AFFEAFFF3FF7473E2FEFF05B3" bold="true" box="[151,267,1432,1458]" pageId="2" pageNumber="139">
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<typeStatus id="54B872DAFFEAFFF3FF7473E2FEF205B3" box="[151,262,1432,1458]" pageId="2" pageNumber="139" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="DBA56403FFEAFFF3FEF573E3FEDF05B5" box="[278,299,1433,1460]" pageId="2" pageNumber="139">♁</specimenCode>
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, Norgaygang
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||
<emphasis id="B977106AFFEAFFF3FE3073E2FDE805B0" box="[467,540,1432,1457]" italics="true" pageId="2" pageNumber="139">gewog</emphasis>
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||
(Bara) (
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||
<geoCoordinate id="EE37AABFFFEAFFF3FD6273EDFCE905B3" box="[641,797,1431,1458]" degrees="27.142670" direction="north" orientation="latitude" pageId="2" pageNumber="139" precision="1" value="27.14267">27.142670°N</geoCoordinate>
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,
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<geoCoordinate id="EE37AABFFFEAFFF3FCCF73EDFC3505B3" box="[812,961,1431,1458]" pageId="2" pageNumber="139">88.835 228°E</geoCoordinate>
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, altitude
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||
<quantity id="4CFB619DFFEAFFF3FBD273EDFB6505B3" box="[1073,1169,1431,1458]" metricMagnitude="3" metricUnit="m" metricValue="1.264" pageId="2" pageNumber="139" unit="m" value="1264.0">
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<elevation id="002E2B4BFFEAFFF3FBD273EDFB6505B3" box="[1073,1169,1431,1458]" metricMagnitude="3" metricUnit="m" metricValue="1.264" pageId="2" pageNumber="139" unit="m" value="1264.0">1,264 m</elevation>
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</quantity>
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a.s.l),
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<collectingRegion id="49C7029AFFEAFFF3FB0673E2FA6C05B0" box="[1253,1432,1431,1458]" country="Bhutan" name="Samchi" pageId="2" pageNumber="139">Samtse District</collectingRegion>
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, southern
|
||
<collectingCountry id="F3148CE8FFEAFFF3FF1C73C1FEA405D4" box="[255,336,1467,1493]" name="Bhutan" pageId="2" pageNumber="139">Bhutan</collectingCountry>
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(
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||
<figureCitation id="1338D0FDFFEAFFF3FEBC73C1FE5405D7" box="[351,416,1467,1494]" captionStart="FIGURE 1" captionStartId="1.[151,250,728,752]" captionTargetBox="[350,1238,181,705]" captionTargetId="figure-17@1.[350,1238,181,705]" captionTargetPageId="1" captionText="FIGURE 1. Map showing the type locality of Protosticta samtsensis sp. nov., Samtse District andP.himalaica, Samdrupjongkhar District (previous record) and Punakha District, Bhutan (new record)." figureDoi="http://doi.org/10.5281/zenodo.8407021" httpUri="https://zenodo.org/record/8407021/files/figure.png" pageId="2" pageNumber="139">Fig. 1</figureCitation>
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||
),
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||
<date id="FFBDEAB8FFEAFFF3FE5A73C1FDB305D4" box="[441,583,1467,1493]" pageId="2" pageNumber="139" value="2022-08-12">
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||
<collectingDate id="EFF91350FFEAFFF3FE5A73C1FDB305D4" box="[441,583,1467,1493]" pageId="2" pageNumber="139" value="2022-08-12">12-viii-2022</collectingDate>
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||
</date>
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||
,
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||
<collectorName id="26F6A9AEFFEAFFF3FDB073C6FCEC05D7" box="[595,792,1468,1494]" pageId="2" pageNumber="139">Mer Man Gurung</collectorName>
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||
leg.
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||
</materialsCitation>
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||
</paragraph>
|
||
<paragraph id="8BBCCC78FFEAFFF3FF24739AFC6205F8" blockId="2.[151,1437,1431,2033]" box="[199,918,1503,1530]" pageId="2" pageNumber="139">
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||
<emphasis id="B977106AFFEAFFF3FF24739AFEC905FB" bold="true" box="[199,317,1504,1530]" pageId="2" pageNumber="139">
|
||
<typeStatus id="54B872DAFFEAFFF3FF24739AFECC05FB" box="[199,312,1504,1530]" pageId="2" pageNumber="139" type="paratype">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D0507F1FFEAFFF3FEA7739AFEAD05FB" box="[324,345,1504,1530]" pageId="2" pageNumber="139" type="female">♀</specimenCount>
|
||
, same date, location, and collector as the
|
||
<typeStatus id="54B872DAFFEAFFF3FCCE73A5FC6705F8" box="[813,915,1503,1529]" pageId="2" pageNumber="139" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFEAFFF3FF24707EFA73061C" blockId="2.[151,1437,1431,2033]" box="[199,1415,1539,1568]" pageId="2" pageNumber="139">
|
||
<emphasis id="B977106AFFEAFFF3FF24707EFDFF061F" bold="true" box="[199,523,1539,1566]" pageId="2" pageNumber="139">Type specimens deposition.</emphasis>
|
||
<materialsCitation id="3B6BC625FFEAFFF3FDF17079FA77061C" ID-GBIF-Occurrence="4421512302" box="[530,1411,1539,1568]" collectionCode="NBC" country="Bhutan" location="Serbithang" pageId="2" pageNumber="139" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||
The
|
||
<typeStatus id="54B872DAFFEAFFF3FDA67079FD53061C" box="[581,679,1539,1565]" pageId="2" pageNumber="139" type="holotype">holotype</typeStatus>
|
||
♁ and paratype
|
||
<specimenCount id="9D0507F1FFEAFFF3FC83707EFC81061F" box="[864,885,1540,1566]" pageId="2" pageNumber="139" type="female">♀</specimenCount>
|
||
will be deposited in
|
||
<collectionCode id="ED1254BDFFEAFFF3FB83707EFB68061F" box="[1120,1180,1540,1566]" collectionName="NBC" pageId="2" pageNumber="139">NBC</collectionCode>
|
||
,
|
||
<location id="8EDC9AA3FFEAFFF3FB447079FAD7061C" LSID="urn:lsid:plazi:treatment:03AA7D6EFFE9FFF4FF74736DFB7207CB:8EDC9AA3FFEAFFF3FB447079FAD7061C" box="[1191,1315,1539,1565]" country="Bhutan" name="Serbithang" pageId="2" pageNumber="139">Serbithang</location>
|
||
,
|
||
<collectingCountry id="F3148CE8FFEAFFF3FACD7079FA77061C" box="[1326,1411,1539,1565]" name="Bhutan" pageId="2" pageNumber="139">Bhutan</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3199FF3FFEAFFF3FF247052FB390688" pageId="2" pageNumber="139" type="etymology">
|
||
<paragraph id="8BBCCC78FFEAFFF3FF247052FD0A0664" blockId="2.[151,1437,1431,2033]" pageId="2" pageNumber="139">
|
||
<emphasis id="B977106AFFEAFFF3FF247052FEBA0643" bold="true" box="[199,334,1576,1602]" pageId="2" pageNumber="139">Etymology.</emphasis>
|
||
The species epithet
|
||
<taxonomicName id="4C03B7FBFFEAFFF3FDDD7052FD420640" authorityName="Gurung & Phan" authorityYear="2023" box="[574,694,1576,1601]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="samtsensis">
|
||
<emphasis id="B977106AFFEAFFF3FDDD7052FD420640" box="[574,694,1576,1601]" italics="true" pageId="2" pageNumber="139">samtsensis</emphasis>
|
||
</taxonomicName>
|
||
is an adjective; based on the name of
|
||
<collectingRegion id="49C7029AFFEAFFF3FB9F7052FADA0640" box="[1148,1326,1575,1602]" country="Bhutan" name="Samchi" pageId="2" pageNumber="139">Samtse District</collectingRegion>
|
||
, southern
|
||
<collectingCountry id="F3148CE8FFEAFFF3FF747031FF1C0664" box="[151,232,1611,1637]" name="Bhutan" pageId="2" pageNumber="139">Bhutan</collectingCountry>
|
||
from where the
|
||
<typeStatus id="54B872DAFFEAFFF3FE427036FE250667" box="[417,465,1612,1638]" pageId="2" pageNumber="139">type</typeStatus>
|
||
specimens were collected.
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFEAFFF3FF247015FB390688" blockId="2.[151,1437,1431,2033]" box="[199,1229,1647,1674]" pageId="2" pageNumber="139">
|
||
<emphasis id="B977106AFFEAFFF3FF247015FE08068B" bold="true" box="[199,508,1647,1674]" pageId="2" pageNumber="139">Suggested common name.</emphasis>
|
||
<collectingCountry id="F3148CE8FFEAFFF3FDE07015FDA00688" box="[515,596,1647,1673]" name="Bhutan" pageId="2" pageNumber="139">Bhutan</collectingCountry>
|
||
Reedtail is the common name suggested for the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3199FF3FFEAFFF4FF2470E9FC260348" lastPageId="5" lastPageNumber="142" pageId="2" pageNumber="139" type="description">
|
||
<paragraph id="8BBCCC78FFEAFFF3FF2470E9FDD106AF" blockId="2.[151,1437,1431,2033]" box="[199,549,1683,1710]" pageId="2" pageNumber="139">
|
||
<emphasis id="B977106AFFEAFFF3FF2470E9FDD106AF" bold="true" box="[199,549,1683,1710]" pageId="2" pageNumber="139">
|
||
Description of male
|
||
<typeStatus id="54B872DAFFEAFFF3FE5470EEFDD406AF" box="[439,544,1684,1710]" pageId="2" pageNumber="139" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFEAFFF3FF2470CDFA78073C" blockId="2.[151,1437,1431,2033]" pageId="2" pageNumber="139">
|
||
<emphasis id="B977106AFFEAFFF3FF2470CDFEF706D0" box="[199,259,1719,1745]" italics="true" pageId="2" pageNumber="139">Head</emphasis>
|
||
(
|
||
<figureCitation id="1338D0FDFFEAFFF3FEF770CDFE8406D3" box="[276,368,1719,1746]" captionStart="FIGURE 4" captionStartId="3.[151,250,1204,1228]" captionTargetBox="[152,1436,181,1181]" captionTargetId="figure-17@3.[151,1436,181,1181]" captionTargetPageId="3" captionText="FIGURE 4. Head and prothorax of Protosticta samtsensis sp. nov. (A, B), holotype ♁ & (C, D), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.8407025" httpUri="https://zenodo.org/record/8407025/files/figure.png" pageId="2" pageNumber="139">Fig. 4A</figureCitation>
|
||
). Dark metallic green; mandibles, labrum, and labium are yellowish white (pale yellow in live individual); movable hooks and upper margin of labrum greenish black; genae brown. Anteclypeus, postclypeus, vertex, and basal segment of antennae black with green metallic sheen; last antennal segments dark brown, and ocelli brown. Eyes (in live individual) dark green upper half and light green below (
|
||
<figureCitation id="1338D0FDFFEAFFF3FC177159FBCC073F" box="[1012,1080,1827,1854]" captionStart="FIGURE 2" captionStartId="1.[151,250,1992,2016]" captionTargetId="figure-91@1.[340,1247,1389,1969]" captionTargetPageId="1" captionText="FIGURE 2. Protosticta samtsensis sp. nov., holotype ♁ alive in the wild." figureDoi="http://doi.org/10.5281/zenodo.8412176" httpUri="https://zenodo.org/record/8412176/files/figure.png" pageId="2" pageNumber="139">Fig. 2</figureCitation>
|
||
); light brown when preserved.
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFEAFFF3FF24713DFE6C07F0" blockId="2.[151,1437,1431,2033]" pageId="2" pageNumber="139">
|
||
<emphasis id="B977106AFFEAFFF3FF24713DFECD0760" box="[199,313,1863,1889]" italics="true" pageId="2" pageNumber="139">Prothorax</emphasis>
|
||
(
|
||
<figureCitation id="1338D0FDFFEAFFF3FEAB713DFE6A0763" box="[328,414,1863,1890]" captionStart="FIGURE 4" captionStartId="3.[151,250,1204,1228]" captionTargetBox="[152,1436,181,1181]" captionTargetId="figure-17@3.[151,1436,181,1181]" captionTargetPageId="3" captionText="FIGURE 4. Head and prothorax of Protosticta samtsensis sp. nov. (A, B), holotype ♁ & (C, D), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.8407025" httpUri="https://zenodo.org/record/8407025/files/figure.png" pageId="2" pageNumber="139">Fig. 4B</figureCitation>
|
||
) dorsally with broad pale yellow lateral stripe in live individual (
|
||
<figureCitation id="1338D0FDFFEAFFF3FB8C713DFB460763" box="[1135,1202,1863,1890]" captionStart="FIGURE 2" captionStartId="1.[151,250,1992,2016]" captionTargetId="figure-91@1.[340,1247,1389,1969]" captionTargetPageId="1" captionText="FIGURE 2. Protosticta samtsensis sp. nov., holotype ♁ alive in the wild." figureDoi="http://doi.org/10.5281/zenodo.8412176" httpUri="https://zenodo.org/record/8412176/files/figure.png" pageId="2" pageNumber="139">Fig. 2</figureCitation>
|
||
) which turns creamy yellow when dried, pale stripe on anterior lobe separated by a black patch in the center, major portion of middle lobe black, which is square shaped with narrow extensions at the four corners, the black marking extends towards center of the posterior lobe which is spherical shaped; small prothoracic spines narrow, yellowish, pointed opposite and curved downward.
|
||
</paragraph>
|
||
<caption id="DF7C9CF0FFEBFFF2FF7472CEFB1C04CD" ID-DOI="http://doi.org/10.5281/zenodo.8407025" ID-Zenodo-Dep="8407025" box="[151,1256,1204,1231]" httpUri="https://zenodo.org/record/8407025/files/figure.png" pageId="3" pageNumber="140" startId="3.[151,250,1204,1228]" targetBox="[152,1436,181,1181]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BBCCC78FFEBFFF2FF7472CEFB1C04CD" blockId="3.[151,1256,1204,1231]" box="[151,1256,1204,1231]" pageId="3" pageNumber="140">
|
||
<emphasis id="B977106AFFEBFFF2FF7472CEFEE704CD" bold="true" box="[151,275,1204,1228]" pageId="3" pageNumber="140">FIGURE 4.</emphasis>
|
||
Head and prothorax of
|
||
<taxonomicName id="4C03B7FBFFEBFFF2FDE772CFFD1204CD" authority="Gurung & Phan, 2023" authorityName="Gurung & Phan" authorityYear="2023" box="[516,742,1205,1228]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="samtsensis" status="sp. nov.">
|
||
<emphasis id="B977106AFFEBFFF2FDE772CFFD1204CD" box="[516,742,1205,1228]" italics="true" pageId="3" pageNumber="140">Protosticta samtsensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B977106AFFEBFFF2FD0E72CFFCCA04CC" bold="true" box="[749,830,1205,1229]" pageId="3" pageNumber="140">
|
||
<taxonomicNameLabel id="A244AD11FFEBFFF2FD0E72CFFCCA04CC" box="[749,830,1205,1229]" pageId="3" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A, B), holotype ♁ & (C, D), paratype ♀.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBCCC78FFEBFFF2FF247281FDA2055C" blockId="3.[151,1437,1275,2021]" pageId="3" pageNumber="140">
|
||
<emphasis id="B977106AFFEBFFF2FF247281FEB90514" box="[199,333,1275,1301]" italics="true" pageId="3" pageNumber="140">Pterothorax</emphasis>
|
||
(
|
||
<figureCitation id="1338D0FDFFEBFFF2FEBC7281FE480514" box="[351,444,1275,1301]" captionStart="FIGURE 3" captionStartId="2.[151,250,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-18@2.[151,1436,181,1337]" captionTargetPageId="2" captionText="FIGURE 3. Habitus of Protosticta samtsensis sp. nov. (A), holotype ♁ & (B), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.8407023" httpUri="https://zenodo.org/record/8407023/files/figure.png" pageId="3" pageNumber="140">Fig. 3A</figureCitation>
|
||
). Dorsum deep metallic green with creamy yellow metepisternum stripe; lateral stripe oblique black with green metallic sheen; metepimeron and ventral surface pale yellow in live individual but turns creamy yellow when preserved (
|
||
<figureCitation id="1338D0FDFFEBFFF2FDE77339FDBD055C" box="[516,585,1347,1373]" captionStart="FIGURE 2" captionStartId="1.[151,250,1992,2016]" captionTargetId="figure-91@1.[340,1247,1389,1969]" captionTargetPageId="1" captionText="FIGURE 2. Protosticta samtsensis sp. nov., holotype ♁ alive in the wild." figureDoi="http://doi.org/10.5281/zenodo.8412176" httpUri="https://zenodo.org/record/8412176/files/figure.png" pageId="3" pageNumber="140">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFEBFFF2FF247312FD9405C8" blockId="3.[151,1437,1275,2021]" pageId="3" pageNumber="140">
|
||
<emphasis id="B977106AFFEBFFF2FF247312FF080580" box="[199,252,1384,1409]" italics="true" pageId="3" pageNumber="140">Legs</emphasis>
|
||
(
|
||
<figureCitation id="1338D0FDFFEBFFF2FEE9731DFE970580" box="[266,355,1383,1409]" captionStart="FIGURE 3" captionStartId="2.[151,250,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-18@2.[151,1436,181,1337]" captionTargetPageId="2" captionText="FIGURE 3. Habitus of Protosticta samtsensis sp. nov. (A), holotype ♁ & (B), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.8407023" httpUri="https://zenodo.org/record/8407023/files/figure.png" pageId="3" pageNumber="140">Fig. 3A</figureCitation>
|
||
). Coxae pale yellow (live individual) and creamy yellow in dried specimen; trochanters yellowish with black posterior spot; femora yellowish; tibio-femoral joints and flexors blackish brown; tibiae yellowish; tarsus along with claws and spines dark brown.
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFEBFFF2FF2473AEFB1305EF" blockId="3.[151,1437,1275,2021]" box="[199,1255,1491,1518]" pageId="3" pageNumber="140">
|
||
<emphasis id="B977106AFFEBFFF2FF2473AEFEFF05EC" box="[199,267,1492,1517]" italics="true" pageId="3" pageNumber="140">Wings</emphasis>
|
||
(
|
||
<figureCitation id="1338D0FDFFEBFFF2FEF873A9FE8605EF" box="[283,370,1491,1518]" captionStart="FIGURE 5" captionStartId="4.[151,250,1710,1734]" captionTargetBox="[161,1426,319,1677]" captionTargetId="figure-66@4.[151,1436,309,1687]" captionTargetPageId="4" captionText="FIGURE 5. Wings and genital ligula of Protosticta samtsensis sp. nov. (A), wings holotype ♁; (B), wings paratype ♀; (C), genital ligula of holotype ♁." figureDoi="http://doi.org/10.5281/zenodo.8407027" httpUri="https://zenodo.org/record/8407027/files/figure.png" pageId="3" pageNumber="140">Fig. 5A</figureCitation>
|
||
) hyaline;
|
||
<emphasis id="B977106AFFEBFFF2FE0073AEFE0805EC" box="[483,508,1492,1517]" italics="true" pageId="3" pageNumber="140">Pt</emphasis>
|
||
black, trapezoid shaped; postnodal crossveins FW-16 and HW-15.
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFEBFFF2FF24738DFF1206A0" blockId="3.[151,1437,1275,2021]" pageId="3" pageNumber="140">
|
||
<emphasis id="B977106AFFEBFFF2FF24738DFEC50610" box="[199,305,1527,1553]" italics="true" pageId="3" pageNumber="140">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="1338D0FDFFEBFFF2FEA1738DFE680610" box="[322,412,1527,1553]" captionStart="FIGURE 3" captionStartId="2.[151,250,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-18@2.[151,1436,181,1337]" captionTargetPageId="2" captionText="FIGURE 3. Habitus of Protosticta samtsensis sp. nov. (A), holotype ♁ & (B), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.8407023" httpUri="https://zenodo.org/record/8407023/files/figure.png" pageId="3" pageNumber="140">Fig. 3A</figureCitation>
|
||
) Blackish brown with creamy yellow markings as follows: S1–2 dark brown and creamy yellow at lateral lower half, S3–6 blackish brown dorsum which gets darker at the posterior end with dull yellow anterior end, S7 light brown with yellow basal end, S8 dorsum brown and venter yellowish throughout, S9 slightly yellowish-brown streaks covering 3/4 part in live individual but turns brown when preserved, S10 completely dark brown.
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFEBFFF5FF2470D1FBBF00D4" blockId="3.[151,1437,1275,2021]" lastBlockId="4.[151,1437,151,285]" lastPageId="4" lastPageNumber="141" pageId="3" pageNumber="140">
|
||
<emphasis id="B977106AFFEBFFF2FF2470D1FE7F06C4" box="[199,395,1707,1733]" italics="true" pageId="3" pageNumber="140">Anal appendages</emphasis>
|
||
(
|
||
<figureCitation id="1338D0FDFFEBFFF2FE7E70D1FDED06C4" box="[413,537,1707,1733]" captionStart="FIGURE 7" captionStartId="6.[151,250,1128,1152]" captionTargetBox="[160,1428,188,1096]" captionTargetId="figure-18@6.[151,1436,181,1104]" captionTargetPageId="6" captionText="FIGURE 7. Anal appendages of Protosticta samtsensis sp. nov., (A-D), holotype ♁; (E-F), P. himalaica (rearranged from Fig. 54a-b in Fraser 1933) & (G-I), P. himalaica collected from Jichulum, Punakha, – (A, E, G), anal appendages in dorsal view; (B, F, H), anal appendages in lateral view; (C), anal appendages in dorso-oblique view; (D, I), paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.8407031" httpUri="https://zenodo.org/record/8407031/files/figure.png" pageId="3" pageNumber="140">Fig. 7A–D</figureCitation>
|
||
) dark brown, in lateral view (
|
||
<figureCitation id="1338D0FDFFEBFFF2FC9070D1FC3F06C4" box="[883,971,1707,1733]" captionStart="FIGURE 7" captionStartId="6.[151,250,1128,1152]" captionTargetBox="[160,1428,188,1096]" captionTargetId="figure-18@6.[151,1436,181,1104]" captionTargetPageId="6" captionText="FIGURE 7. Anal appendages of Protosticta samtsensis sp. nov., (A-D), holotype ♁; (E-F), P. himalaica (rearranged from Fig. 54a-b in Fraser 1933) & (G-I), P. himalaica collected from Jichulum, Punakha, – (A, E, G), anal appendages in dorsal view; (B, F, H), anal appendages in lateral view; (C), anal appendages in dorso-oblique view; (D, I), paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.8407031" httpUri="https://zenodo.org/record/8407031/files/figure.png" pageId="3" pageNumber="140">Fig. 7B</figureCitation>
|
||
) cercus strongly curved downwards, 2.5 times longer than S10, broad at the base, constricts and toward tip on the inner side broadening abruptly forming a depression then eventually narrowing to form a blunt tip, on the outer side, with a slight protrusion towards the tip. Paraprocts dark brown, longer than cerci, with broad base slightly directed towards cerci, towards tip constricted forming an acute angle in the inner side, which then broadens and terminates in a pointed tip directed slightly upwards. In dorsal view (
|
||
<figureCitation id="1338D0FDFFEBFFF2FE4D7125FDF00778" box="[430,516,1887,1913]" captionStart="FIGURE 7" captionStartId="6.[151,250,1128,1152]" captionTargetBox="[160,1428,188,1096]" captionTargetId="figure-18@6.[151,1436,181,1104]" captionTargetPageId="6" captionText="FIGURE 7. Anal appendages of Protosticta samtsensis sp. nov., (A-D), holotype ♁; (E-F), P. himalaica (rearranged from Fig. 54a-b in Fraser 1933) & (G-I), P. himalaica collected from Jichulum, Punakha, – (A, E, G), anal appendages in dorsal view; (B, F, H), anal appendages in lateral view; (C), anal appendages in dorso-oblique view; (D, I), paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.8407031" httpUri="https://zenodo.org/record/8407031/files/figure.png" pageId="3" pageNumber="140">Fig. 7A</figureCitation>
|
||
) cerci curved inward, broad at base which then narrows and again becomes slightly broader and finally ends with blunt tip. In the ventral view (
|
||
<figureCitation id="1338D0FDFFEBFFF2FCCD71F9FC71079C" box="[814,901,1923,1949]" captionStart="FIGURE 7" captionStartId="6.[151,250,1128,1152]" captionTargetBox="[160,1428,188,1096]" captionTargetId="figure-18@6.[151,1436,181,1104]" captionTargetPageId="6" captionText="FIGURE 7. Anal appendages of Protosticta samtsensis sp. nov., (A-D), holotype ♁; (E-F), P. himalaica (rearranged from Fig. 54a-b in Fraser 1933) & (G-I), P. himalaica collected from Jichulum, Punakha, – (A, E, G), anal appendages in dorsal view; (B, F, H), anal appendages in lateral view; (C), anal appendages in dorso-oblique view; (D, I), paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.8407031" httpUri="https://zenodo.org/record/8407031/files/figure.png" pageId="3" pageNumber="140">Fig. 7D</figureCitation>
|
||
) paraprocts are strongly curved inward, broader at the base, which then constricts and has stout wedge shaped inward directed spines at around 3/4 length of the paraprocts, then it smoothly broadens in outward directed curve which forms a spoon-shaped tip. In dorso-oblique view (
|
||
<figureCitation id="1338D0FDFFECFFF5FF3F76EDFEC500B0" box="[220,305,151,177]" captionStart="FIGURE 7" captionStartId="6.[151,250,1128,1152]" captionTargetBox="[160,1428,188,1096]" captionTargetId="figure-18@6.[151,1436,181,1104]" captionTargetPageId="6" captionText="FIGURE 7. Anal appendages of Protosticta samtsensis sp. nov., (A-D), holotype ♁; (E-F), P. himalaica (rearranged from Fig. 54a-b in Fraser 1933) & (G-I), P. himalaica collected from Jichulum, Punakha, – (A, E, G), anal appendages in dorsal view; (B, F, H), anal appendages in lateral view; (C), anal appendages in dorso-oblique view; (D, I), paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.8407031" httpUri="https://zenodo.org/record/8407031/files/figure.png" pageId="4" pageNumber="141">Fig. 7C</figureCitation>
|
||
) cerci are strongly curved downward almost touching the paraprocts, tip of the spine on the paraprocts slightly lifted towards the cerci and paraprocts terminate in scoop-like structured tip.
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFECFFF5FF2476A5FF02011C" blockId="4.[151,1437,151,285]" pageId="4" pageNumber="141">
|
||
<emphasis id="B977106AFFECFFF5FF2476A5FE9E00F8" box="[199,362,223,249]" italics="true" pageId="4" pageNumber="141">Genital ligula</emphasis>
|
||
(
|
||
<figureCitation id="1338D0FDFFECFFF5FE6376A5FE2800F8" box="[384,476,222,249]" captionStart="FIGURE 5" captionStartId="4.[151,250,1710,1734]" captionTargetBox="[161,1426,319,1677]" captionTargetId="figure-66@4.[151,1436,309,1687]" captionTargetPageId="4" captionText="FIGURE 5. Wings and genital ligula of Protosticta samtsensis sp. nov. (A), wings holotype ♁; (B), wings paratype ♀; (C), genital ligula of holotype ♁." figureDoi="http://doi.org/10.5281/zenodo.8407027" httpUri="https://zenodo.org/record/8407027/files/figure.png" pageId="4" pageNumber="141">Fig. 5C</figureCitation>
|
||
) simple structurally, the tip with two short apical flagella each slightly broader apically.
|
||
</paragraph>
|
||
<caption id="DF7C9CF0FFECFFF5FF7470D4FE4206ED" ID-DOI="http://doi.org/10.5281/zenodo.8407027" ID-Zenodo-Dep="8407027" httpUri="https://zenodo.org/record/8407027/files/figure.png" pageId="4" pageNumber="141" startId="4.[151,250,1710,1734]" targetBox="[161,1426,319,1677]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BBCCC78FFECFFF5FF7470D4FE4206ED" blockId="4.[151,1436,1710,1772]" pageId="4" pageNumber="141">
|
||
<emphasis id="B977106AFFECFFF5FF7470D4FEFB06C7" bold="true" box="[151,271,1710,1734]" pageId="4" pageNumber="141">FIGURE 5</emphasis>
|
||
. Wings and genital ligula of
|
||
<taxonomicName id="4C03B7FBFFECFFF5FDDD70D5FCD606C7" authority="Gurung & Phan, 2023" authorityName="Gurung & Phan" authorityYear="2023" box="[574,802,1711,1734]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="samtsensis" status="sp. nov.">
|
||
<emphasis id="B977106AFFECFFF5FDDD70D5FCD606C7" box="[574,802,1711,1734]" italics="true" pageId="4" pageNumber="141">Protosticta samtsensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B977106AFFECFFF5FCC970D4FC8906C7" bold="true" box="[810,893,1710,1734]" pageId="4" pageNumber="141">
|
||
<taxonomicNameLabel id="A244AD11FFECFFF5FCC970D4FC8906C7" box="[810,893,1710,1734]" pageId="4" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A), wings holotype ♁; (B), wings paratype ♀; (C), genital ligula of holotype ♁.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBCCC78FFECFFF5FF247162FC260733" blockId="4.[151,1437,1816,2022]" box="[199,978,1816,1843]" pageId="4" pageNumber="141">
|
||
<emphasis id="B977106AFFECFFF5FF247162FE820733" bold="true" box="[199,374,1816,1842]" pageId="4" pageNumber="141">Measurements</emphasis>
|
||
(in mm). HW 21; abdomen including appendages 40.
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFECFFF5FF247146FB3A0757" blockId="4.[151,1437,1816,2022]" box="[199,1230,1852,1879]" pageId="4" pageNumber="141">
|
||
<emphasis id="B977106AFFECFFF5FF247146FDB40757" bold="true" box="[199,576,1852,1878]" pageId="4" pageNumber="141">
|
||
Description of female
|
||
<typeStatus id="54B872DAFFECFFF5FE2E7146FDCE0757" box="[461,570,1852,1878]" pageId="4" pageNumber="141" type="paratype">paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Looks similar to male with minute differences as follows:
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFECFFF5FF24711AFBCF079E" blockId="4.[151,1437,1816,2022]" pageId="4" pageNumber="141">
|
||
<emphasis id="B977106AFFECFFF5FF24711AFEF0077B" box="[199,260,1888,1914]" italics="true" pageId="4" pageNumber="141">Head</emphasis>
|
||
(
|
||
<figureCitation id="1338D0FDFFECFFF5FEF1711AFE93077B" box="[274,359,1888,1914]" captionStart="FIGURE 4" captionStartId="3.[151,250,1204,1228]" captionTargetBox="[152,1436,181,1181]" captionTargetId="figure-17@3.[151,1436,181,1181]" captionTargetPageId="3" captionText="FIGURE 4. Head and prothorax of Protosticta samtsensis sp. nov. (A, B), holotype ♁ & (C, D), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.8407025" httpUri="https://zenodo.org/record/8407025/files/figure.png" pageId="4" pageNumber="141">Fig. 4C</figureCitation>
|
||
) labrum and upper half of the labium bluish-white and black below separated by thin faint brown margin. Mandibles black with brown at the corners, genae brown. Eyes dark green.
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFECFFF5FF2471D2FEDE07E7" blockId="4.[151,1437,1816,2022]" pageId="4" pageNumber="141">
|
||
<emphasis id="B977106AFFECFFF5FF2471D2FECD07C3" box="[199,313,1960,1986]" italics="true" pageId="4" pageNumber="141">Prothorax</emphasis>
|
||
(
|
||
<figureCitation id="1338D0FDFFECFFF5FEAF71D2FE5307C3" box="[332,423,1960,1986]" captionStart="FIGURE 4" captionStartId="3.[151,250,1204,1228]" captionTargetBox="[152,1436,181,1181]" captionTargetId="figure-17@3.[151,1436,181,1181]" captionTargetPageId="3" captionText="FIGURE 4. Head and prothorax of Protosticta samtsensis sp. nov. (A, B), holotype ♁ & (C, D), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.8407025" httpUri="https://zenodo.org/record/8407025/files/figure.png" pageId="4" pageNumber="141">Fig. 4D</figureCitation>
|
||
) anterior lobe with different black central marking, only posterior portion marked with black border.
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFEDFFF4FF2476EDFCE500B0" blockId="5.[151,1437,151,321]" box="[199,785,151,177]" pageId="5" pageNumber="142">
|
||
<emphasis id="B977106AFFEDFFF4FF2476EDFF0800B1" box="[199,252,151,176]" italics="true" pageId="5" pageNumber="142">Legs</emphasis>
|
||
(
|
||
<figureCitation id="1338D0FDFFEDFFF4FEE876EDFE9500B0" box="[267,353,151,177]" captionStart="FIGURE 3" captionStartId="2.[151,250,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-18@2.[151,1436,181,1337]" captionTargetPageId="2" captionText="FIGURE 3. Habitus of Protosticta samtsensis sp. nov. (A), holotype ♁ & (B), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.8407023" httpUri="https://zenodo.org/record/8407023/files/figure.png" pageId="5" pageNumber="142">Fig. 3B</figureCitation>
|
||
) trochanter posterior dark spot absent.
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFEDFFF4FF2476C1FCA400D4" blockId="5.[151,1437,151,321]" box="[199,848,186,213]" pageId="5" pageNumber="142">
|
||
<emphasis id="B977106AFFEDFFF4FF2476C1FEFF00D5" box="[199,267,187,212]" italics="true" pageId="5" pageNumber="142">Wings</emphasis>
|
||
(
|
||
<figureCitation id="1338D0FDFFEDFFF4FEF876C1FE8600D4" box="[283,370,186,213]" captionStart="FIGURE 5" captionStartId="4.[151,250,1710,1734]" captionTargetBox="[161,1426,319,1677]" captionTargetId="figure-66@4.[151,1436,309,1687]" captionTargetPageId="4" captionText="FIGURE 5. Wings and genital ligula of Protosticta samtsensis sp. nov. (A), wings holotype ♁; (B), wings paratype ♀; (C), genital ligula of holotype ♁." figureDoi="http://doi.org/10.5281/zenodo.8407027" httpUri="https://zenodo.org/record/8407027/files/figure.png" pageId="5" pageNumber="142">Fig. 5A</figureCitation>
|
||
) postnodal crossveins FW-17 and HW-15.
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFEDFFF4FF2476A5FC5C00F8" blockId="5.[151,1437,151,321]" box="[199,936,223,249]" pageId="5" pageNumber="142">
|
||
<emphasis id="B977106AFFEDFFF4FF2476A5FEC500F8" box="[199,305,223,249]" italics="true" pageId="5" pageNumber="142">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="1338D0FDFFEDFFF4FEA376A5FE6C00F8" box="[320,408,223,249]" captionStart="FIGURE 3" captionStartId="2.[151,250,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-18@2.[151,1436,181,1337]" captionTargetPageId="2" captionText="FIGURE 3. Habitus of Protosticta samtsensis sp. nov. (A), holotype ♁ & (B), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.8407023" httpUri="https://zenodo.org/record/8407023/files/figure.png" pageId="5" pageNumber="142">Fig. 3B</figureCitation>
|
||
). S9 with lateral bright brownish yellow patch.
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFEDFFF4FF247779FB220140" blockId="5.[151,1437,151,321]" pageId="5" pageNumber="142">
|
||
<emphasis id="B977106AFFEDFFF4FF247779FE7E011C" box="[199,394,259,285]" italics="true" pageId="5" pageNumber="142">Anal appendages</emphasis>
|
||
(
|
||
<figureCitation id="1338D0FDFFEDFFF4FE7A7779FDE7011C" box="[409,531,259,285]" captionStart="FIGURE 6" captionStartId="5.[152,255,745,769]" captionTargetBox="[340,1247,345,722]" captionTargetId="figure-76@5.[340,1247,345,722]" captionTargetPageId="5" captionText="FIGURE 6. Anal appendages of Protosticta samtsensis sp. nov., paratype ♀: (A) lateral view, (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8407029" httpUri="https://zenodo.org/record/8407029/files/figure.png" pageId="5" pageNumber="142">Fig. 6A–B</figureCitation>
|
||
) dark brown, broad at the base with smooth tip, slightly longer than S10 and bit curved downward. Ovipositor dark brown laterally with creamy yellow sharp spiky tip (
|
||
<figureCitation id="1338D0FDFFEDFFF4FB92775DFB3E0140" box="[1137,1226,295,321]" captionStart="FIGURE 6" captionStartId="5.[152,255,745,769]" captionTargetBox="[340,1247,345,722]" captionTargetId="figure-76@5.[340,1247,345,722]" captionTargetPageId="5" captionText="FIGURE 6. Anal appendages of Protosticta samtsensis sp. nov., paratype ♀: (A) lateral view, (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8407029" httpUri="https://zenodo.org/record/8407029/files/figure.png" pageId="5" pageNumber="142">Fig. 6B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF7C9CF0FFEDFFF4FF7B7493FAF50300" ID-DOI="http://doi.org/10.5281/zenodo.8407029" ID-Zenodo-Dep="8407029" box="[152,1281,745,770]" httpUri="https://zenodo.org/record/8407029/files/figure.png" pageId="5" pageNumber="142" startId="5.[152,255,745,769]" targetBox="[340,1247,345,722]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BBCCC78FFEDFFF4FF7B7493FAF50300" blockId="5.[152,1281,745,770]" box="[152,1281,745,770]" pageId="5" pageNumber="142">
|
||
<emphasis id="B977106AFFEDFFF4FF7B7493FEEC0300" bold="true" box="[152,280,745,769]" pageId="5" pageNumber="142">FIGURE 6.</emphasis>
|
||
Anal appendages of
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FE0D7490FD240300" authority="Gurung & Phan, 2023" authorityName="Gurung & Phan" authorityYear="2023" box="[494,720,746,769]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="samtsensis" status="sp. nov.">
|
||
<emphasis id="B977106AFFEDFFF4FE0D7490FD240300" box="[494,720,746,769]" italics="true" pageId="5" pageNumber="142">Protosticta samtsensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B977106AFFEDFFF4FD347490FCD30303" bold="true" box="[727,807,746,770]" pageId="5" pageNumber="142">
|
||
<taxonomicNameLabel id="A244AD11FFEDFFF4FD347490FCD30303" box="[727,807,746,770]" pageId="5" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype ♀: (A) lateral view, (B) dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBCCC78FFEDFFF4FF24754AFC260348" blockId="5.[151,1437,815,1994]" box="[199,978,815,842]" pageId="5" pageNumber="142">
|
||
<emphasis id="B977106AFFEDFFF4FF24754AFE82034B" bold="true" box="[199,374,816,842]" pageId="5" pageNumber="142">Measurements</emphasis>
|
||
(in mm). HW 27; abdomen including appendages 36.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3199FF3FFEDFFF4FF247529FB7207CB" pageId="5" pageNumber="142" type="diagnosis">
|
||
<paragraph id="8BBCCC78FFEDFFF4FF247529FD4A04FB" blockId="5.[151,1437,815,1994]" pageId="5" pageNumber="142">
|
||
<emphasis id="B977106AFFEDFFF4FF247529FE3B036C" bold="true" box="[199,463,851,877]" pageId="5" pageNumber="142">Differential diagnosis.</emphasis>
|
||
The genus
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FDB7752EFD3B036C" authorityName="Selys" authorityYear="1885" box="[596,719,852,877]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B977106AFFEDFFF4FDB7752EFD3B036C" box="[596,719,852,877]" italics="true" pageId="5" pageNumber="142">Protosticta</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FBE6752EFABA036F" authority="Laidlaw, 1917" authorityName="Laidlaw" authorityYear="1917" box="[1029,1358,852,878]" class="Insecta" family="Platystictidae" genus="Drepanosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B977106AFFEDFFF4FBE6752EFB50036C" box="[1029,1188,852,877]" italics="true" pageId="5" pageNumber="142">Drepanosticta</emphasis>
|
||
<bibRefCitation id="EF92B189FFEDFFF4FB4E752EFABA036F" author="Laidlaw, F. F." box="[1197,1358,852,878]" pageId="5" pageNumber="142" pagination="321 - 348" refId="ref3502" refString="Laidlaw, F. F. (1917) A list of the dragonflies recorded from the Indian Empire with special reference to the collection of the Indian Museum. Part II. The family Agrioninae. Records of the Indian Museum, 13, 321 - 348. https: // doi. org / 10.5962 / bhl. part. 5864" type="journal article" year="1917">Laidlaw, 1917</bibRefCitation>
|
||
</taxonomicName>
|
||
by the absence of nervure
|
||
<emphasis id="B977106AFFEDFFF4FE8F7502FE7C0393" box="[364,392,888,914]" italics="true" pageId="5" pageNumber="142">ab</emphasis>
|
||
(
|
||
<figureCitation id="1338D0FDFFEDFFF4FE767502FDF80393" box="[405,524,887,914]" captionStart="FIGURE 5" captionStartId="4.[151,250,1710,1734]" captionTargetBox="[161,1426,319,1677]" captionTargetId="figure-66@4.[151,1436,309,1687]" captionTargetPageId="4" captionText="FIGURE 5. Wings and genital ligula of Protosticta samtsensis sp. nov. (A), wings holotype ♁; (B), wings paratype ♀; (C), genital ligula of holotype ♁." figureDoi="http://doi.org/10.5281/zenodo.8407027" httpUri="https://zenodo.org/record/8407027/files/figure.png" pageId="5" pageNumber="142">Fig. 5A–B</figureCitation>
|
||
). Among representatives of the genus
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FC537502FBDF0390" authorityName="Selys" authorityYear="1885" box="[944,1067,888,913]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B977106AFFEDFFF4FC537502FBDF0390" box="[944,1067,888,913]" italics="true" pageId="5" pageNumber="142">Protosticta</emphasis>
|
||
</taxonomicName>
|
||
found in the Indian subcontinent, two species, viz.
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FEBC75E6FE0103B4" authorityName="Gurung & Phan" authorityYear="2023" box="[351,501,924,949]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="samtsensis">
|
||
<emphasis id="B977106AFFEDFFF4FEBC75E6FE0103B4" box="[351,501,924,949]" italics="true" pageId="5" pageNumber="142">P. samtsensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FDD375E6FD4A03B7" box="[560,702,924,950]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="himalaica">
|
||
<emphasis id="B977106AFFEDFFF4FDD375E6FD4A03B7" box="[560,702,924,950]" italics="true" pageId="5" pageNumber="142">P. himalaica</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from congeners by their robust paraprocts being longer than the cerci and bearing two inward directed spines (
|
||
<figureCitation id="1338D0FDFFEDFFF4FC7875BAFBFB03DB" box="[923,1039,960,986]" captionStart="FIGURE 7" captionStartId="6.[151,250,1128,1152]" captionTargetBox="[160,1428,188,1096]" captionTargetId="figure-18@6.[151,1436,181,1104]" captionTargetPageId="6" captionText="FIGURE 7. Anal appendages of Protosticta samtsensis sp. nov., (A-D), holotype ♁; (E-F), P. himalaica (rearranged from Fig. 54a-b in Fraser 1933) & (G-I), P. himalaica collected from Jichulum, Punakha, – (A, E, G), anal appendages in dorsal view; (B, F, H), anal appendages in lateral view; (C), anal appendages in dorso-oblique view; (D, I), paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.8407031" httpUri="https://zenodo.org/record/8407031/files/figure.png" pageId="5" pageNumber="142">Fig. 7A–I</figureCitation>
|
||
). The two species can be separated with the following diagnosis: cercus of
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FD8A759EFD0C03FF" box="[617,760,996,1022]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="himalaica">
|
||
<emphasis id="B977106AFFEDFFF4FD8A759EFD0C03FF" box="[617,760,996,1022]" italics="true" pageId="5" pageNumber="142">P. himalaica</emphasis>
|
||
</taxonomicName>
|
||
with a dorsal inward-directed spine (
|
||
<figureCitation id="1338D0FDFFEDFFF4FB4C759EFAD903FF" box="[1199,1325,996,1022]" captionStart="FIGURE 7" captionStartId="6.[151,250,1128,1152]" captionTargetBox="[160,1428,188,1096]" captionTargetId="figure-18@6.[151,1436,181,1104]" captionTargetPageId="6" captionText="FIGURE 7. Anal appendages of Protosticta samtsensis sp. nov., (A-D), holotype ♁; (E-F), P. himalaica (rearranged from Fig. 54a-b in Fraser 1933) & (G-I), P. himalaica collected from Jichulum, Punakha, – (A, E, G), anal appendages in dorsal view; (B, F, H), anal appendages in lateral view; (C), anal appendages in dorso-oblique view; (D, I), paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.8407031" httpUri="https://zenodo.org/record/8407031/files/figure.png" pageId="5" pageNumber="142">Fig. 7E, G</figureCitation>
|
||
) whereas the dorsal cercus of
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FE637272FDE20420" authorityName="Gurung & Phan" authorityYear="2023" box="[384,534,1032,1057]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="samtsensis">
|
||
<emphasis id="B977106AFFEDFFF4FE637272FDE20420" box="[384,534,1032,1057]" italics="true" pageId="5" pageNumber="142">P. samtsensis</emphasis>
|
||
</taxonomicName>
|
||
is smooth, without spine (
|
||
<figureCitation id="1338D0FDFFEDFFF4FCA47272FC570423" box="[839,931,1032,1058]" captionStart="FIGURE 7" captionStartId="6.[151,250,1128,1152]" captionTargetBox="[160,1428,188,1096]" captionTargetId="figure-18@6.[151,1436,181,1104]" captionTargetPageId="6" captionText="FIGURE 7. Anal appendages of Protosticta samtsensis sp. nov., (A-D), holotype ♁; (E-F), P. himalaica (rearranged from Fig. 54a-b in Fraser 1933) & (G-I), P. himalaica collected from Jichulum, Punakha, – (A, E, G), anal appendages in dorsal view; (B, F, H), anal appendages in lateral view; (C), anal appendages in dorso-oblique view; (D, I), paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.8407031" httpUri="https://zenodo.org/record/8407031/files/figure.png" pageId="5" pageNumber="142">Fig. 7A</figureCitation>
|
||
). In
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FC3A7272FB920423" box="[985,1126,1032,1058]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="himalaica">
|
||
<emphasis id="B977106AFFEDFFF4FC3A7272FB920423" box="[985,1126,1032,1058]" italics="true" pageId="5" pageNumber="142">P. himalaica</emphasis>
|
||
</taxonomicName>
|
||
the paraprocts have a long inward directed narrow medial spine. However, in
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FD2C7256FC970444" authorityName="Gurung & Phan" authorityYear="2023" box="[719,867,1068,1093]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="samtsensis">
|
||
<emphasis id="B977106AFFEDFFF4FD2C7256FC970444" box="[719,867,1068,1093]" italics="true" pageId="5" pageNumber="142">P. samtsensis</emphasis>
|
||
</taxonomicName>
|
||
the spine on the paraproct is much shorter, stouter, and wedge-shaped (
|
||
<figureCitation id="1338D0FDFFEDFFF4FE9B722AFE02046B" box="[376,502,1104,1130]" captionStart="FIGURE 7" captionStartId="6.[151,250,1128,1152]" captionTargetBox="[160,1428,188,1096]" captionTargetId="figure-18@6.[151,1436,181,1104]" captionTargetPageId="6" captionText="FIGURE 7. Anal appendages of Protosticta samtsensis sp. nov., (A-D), holotype ♁; (E-F), P. himalaica (rearranged from Fig. 54a-b in Fraser 1933) & (G-I), P. himalaica collected from Jichulum, Punakha, – (A, E, G), anal appendages in dorsal view; (B, F, H), anal appendages in lateral view; (C), anal appendages in dorso-oblique view; (D, I), paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.8407031" httpUri="https://zenodo.org/record/8407031/files/figure.png" pageId="5" pageNumber="142">Fig. 7C, D</figureCitation>
|
||
). In other species known from the northeast of Indian subcontinent, the cerci of
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FA6B722AFF16048F" authorityName="Kennedy" authorityYear="1936" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="fraseri">
|
||
<emphasis id="B977106AFFEDFFF4FA6B722AFF16048F" italics="true" pageId="5" pageNumber="142">P. fraseri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FEF8720EFE4C048F" authorityName="Terzani & Carletti" authorityYear="1998" box="[283,440,1140,1166]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="damacornu">
|
||
<emphasis id="B977106AFFEDFFF4FEF8720EFE4C048F" box="[283,440,1140,1166]" italics="true" pageId="5" pageNumber="142">P. damacornu</emphasis>
|
||
</taxonomicName>
|
||
are broad in apex, form a triangular plate (for instance, see
|
||
<figureCitation id="1338D0FDFFEDFFF4FB85720EFB5C048F" box="[1126,1192,1140,1166]" captionStart="FIGURE 3" captionStartId="2.[151,250,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-18@2.[151,1436,181,1337]" captionTargetPageId="2" captionText="FIGURE 3. Habitus of Protosticta samtsensis sp. nov. (A), holotype ♁ & (B), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.8407023" httpUri="https://zenodo.org/record/8407023/files/figure.png" pageId="5" pageNumber="142">
|
||
Fig.
|
||
<quantity id="4CFB619DFFEDFFF4FB79720EFB5C048F" box="[1178,1192,1140,1166]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="5" pageNumber="142" unit="in" value="3.0">3</quantity>
|
||
</figureCitation>
|
||
in
|
||
<bibRefCitation id="EF92B189FFEDFFF4FB2D720EFF1004B3" author="Terzani, F. & Carletti, B." pageId="5" pageNumber="142" pagination="479 - 485" refId="ref3781" refString="Terzani, F. & Carletti, B. (1998) Protosticta damacornu spec. nov. and other odonate records from northeastern India (Zygoptera: Platystictidae). Odonatologica, 27, 479 - 485." type="journal article" year="1998">Terzani & Carletti (1998)</bibRefCitation>
|
||
, while cerci of
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FE7072E2FDD204B0" authorityName="Gurung & Phan" authorityYear="2023" box="[403,550,1176,1201]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="samtsensis">
|
||
<emphasis id="B977106AFFEDFFF4FE7072E2FDD204B0" box="[403,550,1176,1201]" italics="true" pageId="5" pageNumber="142">P. samtsensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FDBD72E2FD1D04B3" box="[606,745,1176,1202]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="himalaica">
|
||
<emphasis id="B977106AFFEDFFF4FDBD72E2FD1D04B3" box="[606,745,1176,1202]" italics="true" pageId="5" pageNumber="142">P. himalaica</emphasis>
|
||
</taxonomicName>
|
||
are narrow, and blunt at tip (
|
||
<figureCitation id="1338D0FDFFEDFFF4FBD372E2FB5F04B3" box="[1072,1195,1176,1202]" captionStart="FIGURE 7" captionStartId="6.[151,250,1128,1152]" captionTargetBox="[160,1428,188,1096]" captionTargetId="figure-18@6.[151,1436,181,1104]" captionTargetPageId="6" captionText="FIGURE 7. Anal appendages of Protosticta samtsensis sp. nov., (A-D), holotype ♁; (E-F), P. himalaica (rearranged from Fig. 54a-b in Fraser 1933) & (G-I), P. himalaica collected from Jichulum, Punakha, – (A, E, G), anal appendages in dorsal view; (B, F, H), anal appendages in lateral view; (C), anal appendages in dorso-oblique view; (D, I), paraprocts in ventral view." figureDoi="http://doi.org/10.5281/zenodo.8407031" httpUri="https://zenodo.org/record/8407031/files/figure.png" pageId="5" pageNumber="142">Fig. 7A, E</figureCitation>
|
||
). Moreover, unlike
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FA6B72E2FEFB04D4" authorityName="Gurung & Phan" authorityYear="2023" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="samtsensis">
|
||
<emphasis id="B977106AFFEDFFF4FA6B72E2FEFB04D4" italics="true" pageId="5" pageNumber="142">P. samtsensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FEA472C6FE2704D7" box="[327,467,1212,1238]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="himalaica">
|
||
<emphasis id="B977106AFFEDFFF4FEA472C6FE2704D7" box="[327,467,1212,1238]" italics="true" pageId="5" pageNumber="142">P. himalaica</emphasis>
|
||
</taxonomicName>
|
||
, the cercus and paraproct of both
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FCB672C6FC4804D7" authorityName="Kennedy" authorityYear="1936" box="[853,956,1212,1238]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="fraseri">
|
||
<emphasis id="B977106AFFEDFFF4FCB672C6FC4804D7" box="[853,956,1212,1238]" italics="true" pageId="5" pageNumber="142">P. fraseri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FC1772C6FB6404D7" authorityName="Terzani & Carletti" authorityYear="1998" box="[1012,1168,1212,1238]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="damacornu">
|
||
<emphasis id="B977106AFFEDFFF4FC1772C6FB6404D7" box="[1012,1168,1212,1238]" italics="true" pageId="5" pageNumber="142">P. damacornu</emphasis>
|
||
</taxonomicName>
|
||
are of equal length (for instance, see
|
||
<figureCitation id="1338D0FDFFEDFFF4FECF729AFE9904FB" box="[300,365,1248,1274]" captionStart="FIGURE 2" captionStartId="1.[151,250,1992,2016]" captionTargetId="figure-91@1.[340,1247,1389,1969]" captionTargetPageId="1" captionText="FIGURE 2. Protosticta samtsensis sp. nov., holotype ♁ alive in the wild." figureDoi="http://doi.org/10.5281/zenodo.8412176" httpUri="https://zenodo.org/record/8412176/files/figure.png" pageId="5" pageNumber="142">
|
||
Fig.
|
||
<quantity id="4CFB619DFFEDFFF4FEBC729AFE9904FB" box="[351,365,1248,1274]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="142" unit="in" value="2.0">2</quantity>
|
||
</figureCitation>
|
||
in
|
||
<bibRefCitation id="EF92B189FFEDFFF4FE73729AFD4E04FB" author="Terzani, F. & Carletti, B." box="[400,698,1248,1274]" pageId="5" pageNumber="142" pagination="479 - 485" refId="ref3781" refString="Terzani, F. & Carletti, B. (1998) Protosticta damacornu spec. nov. and other odonate records from northeastern India (Zygoptera: Platystictidae). Odonatologica, 27, 479 - 485." type="journal article" year="1998">Terzani & Carletti (1998))</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFEDFFF4FF24737EFE1A06CF" blockId="5.[151,1437,815,1994]" pageId="5" pageNumber="142">
|
||
While the original description of
|
||
<bibRefCitation id="EF92B189FFEDFFF4FDDA737EFD12051F" author="Laidlaw, F. F." box="[569,742,1284,1310]" pageId="5" pageNumber="142" pagination="321 - 348" refId="ref3502" refString="Laidlaw, F. F. (1917) A list of the dragonflies recorded from the Indian Empire with special reference to the collection of the Indian Museum. Part II. The family Agrioninae. Records of the Indian Museum, 13, 321 - 348. https: // doi. org / 10.5962 / bhl. part. 5864" type="journal article" year="1917">Laidlaw (1917)</bibRefCitation>
|
||
is based on immature specimens, we provide the difference in body coloration between
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FE517352FDC90543" box="[434,573,1320,1346]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="himalaica">
|
||
<emphasis id="B977106AFFEDFFF4FE517352FDC90543" box="[434,573,1320,1346]" italics="true" pageId="5" pageNumber="142">P. himalaica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FD907352FCF30540" authorityName="Gurung & Phan" authorityYear="2023" box="[627,775,1320,1345]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="samtsensis">
|
||
<emphasis id="B977106AFFEDFFF4FD907352FCF30540" box="[627,775,1320,1345]" italics="true" pageId="5" pageNumber="142">P. samtsensis</emphasis>
|
||
</taxonomicName>
|
||
based on the description of
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FBA27352FB380543" box="[1089,1228,1320,1346]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="himalaica">
|
||
<emphasis id="B977106AFFEDFFF4FBA27352FB380543" box="[1089,1228,1320,1346]" italics="true" pageId="5" pageNumber="142">P. himalaica</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="EF92B189FFEDFFF4FB0C7352FA680543" author="Laidlaw, F. F." box="[1263,1436,1320,1346]" pageId="5" pageNumber="142" pagination="321 - 348" refId="ref3502" refString="Laidlaw, F. F. (1917) A list of the dragonflies recorded from the Indian Empire with special reference to the collection of the Indian Museum. Part II. The family Agrioninae. Records of the Indian Museum, 13, 321 - 348. https: // doi. org / 10.5962 / bhl. part. 5864" type="journal article" year="1917">Laidlaw (1917)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF92B189FFEDFFF4FF297336FE910567" author="Fraser, F. C." box="[202,357,1356,1382]" pageId="5" pageNumber="142" refId="ref2959" refString="Fraser, F. C. (1933) The fauna of British India, including Ceylon and Burma. Odonata Vol. I. Taylor and Francis, London, XIV + 423 pp." type="book" year="1933">Fraser (1933)</bibRefCitation>
|
||
and the specimens collected from
|
||
<collectingCountry id="F3148CE8FFEDFFF4FCE27336FCA60567" box="[769,850,1356,1382]" name="Bhutan" pageId="5" pageNumber="142">Bhutan</collectingCountry>
|
||
as follows:
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FC067336FB800567" box="[997,1140,1356,1382]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="himalaica">
|
||
<emphasis id="B977106AFFEDFFF4FC067336FB800567" box="[997,1140,1356,1382]" italics="true" pageId="5" pageNumber="142">P. himalaica</emphasis>
|
||
</taxonomicName>
|
||
with pale brown labium; clypeus blue; and vertex black whereas
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FD8B730AFD0A0588" authorityName="Gurung & Phan" authorityYear="2023" box="[616,766,1392,1417]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="samtsensis">
|
||
<emphasis id="B977106AFFEDFFF4FD8B730AFD0A0588" box="[616,766,1392,1417]" italics="true" pageId="5" pageNumber="142">P. samtsensis</emphasis>
|
||
</taxonomicName>
|
||
has whitish-yellow labium; anteclypeus, postclypeus, and vertex are metallic green. Pterothorax of
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FD9273EEFD0B05AF" box="[625,767,1428,1454]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="himalaica">
|
||
<emphasis id="B977106AFFEDFFF4FD9273EEFD0B05AF" box="[625,767,1428,1454]" italics="true" pageId="5" pageNumber="142">P. himalaica</emphasis>
|
||
</taxonomicName>
|
||
with black dorsum and lateral stripe, pale blue above and below lateral stripe. In
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FE7673C2FDDC05D0" authorityName="Gurung & Phan" authorityYear="2023" box="[405,552,1464,1489]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="samtsensis">
|
||
<emphasis id="B977106AFFEDFFF4FE7673C2FDDC05D0" box="[405,552,1464,1489]" italics="true" pageId="5" pageNumber="142">P. samtsensis</emphasis>
|
||
</taxonomicName>
|
||
pterothorax with dorsum and lateral stripe metallic green, creamy yellow above and below a lateral stripe. The abdomen of
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FD9473A6FCF505F7" box="[631,769,1500,1526]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="himalaica">
|
||
<emphasis id="B977106AFFEDFFF4FD9473A6FCF505F7" box="[631,769,1500,1526]" italics="true" pageId="5" pageNumber="142">P. himalaica</emphasis>
|
||
</taxonomicName>
|
||
in live individual S1 has blue sides with a white apical ring, S2 laterally blue marked with a blue dorsal strip, but it is different in
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FC79707AFBD90618" authorityName="Gurung & Phan" authorityYear="2023" box="[922,1069,1536,1561]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="samtsensis">
|
||
<emphasis id="B977106AFFEDFFF4FC79707AFBD90618" box="[922,1069,1536,1561]" italics="true" pageId="5" pageNumber="142">P. samtsensis</emphasis>
|
||
</taxonomicName>
|
||
S1–2 dorsum brown, lateral half creamy yellow. In
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FE84705EFE07063F" box="[359,499,1572,1598]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="himalaica">
|
||
<emphasis id="B977106AFFEDFFF4FE84705EFE07063F" box="[359,499,1572,1598]" italics="true" pageId="5" pageNumber="142">P. himalaica</emphasis>
|
||
</taxonomicName>
|
||
S3 dorsum has white basal triangle which is missing in
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FB8B705EFB0F063C" authorityName="Gurung & Phan" authorityYear="2023" box="[1128,1275,1572,1597]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="samtsensis">
|
||
<emphasis id="B977106AFFEDFFF4FB8B705EFB0F063C" box="[1128,1275,1572,1597]" italics="true" pageId="5" pageNumber="142">P. samtsensis</emphasis>
|
||
</taxonomicName>
|
||
. Furthermore, in
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FF507032FEC90663" box="[179,317,1608,1634]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="himalaica">
|
||
<emphasis id="B977106AFFEDFFF4FF507032FEC90663" box="[179,317,1608,1634]" italics="true" pageId="5" pageNumber="142">P. himalaica</emphasis>
|
||
</taxonomicName>
|
||
, S7 apical half, S8–9 and S10 the basal end is blue in live individual but turns brown when preserved. In contrast
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FEF67016FE5C0684" authorityName="Gurung & Phan" authorityYear="2023" box="[277,424,1644,1669]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="samtsensis">
|
||
<emphasis id="B977106AFFEDFFF4FEF67016FE5C0684" box="[277,424,1644,1669]" italics="true" pageId="5" pageNumber="142">P. samtsensis</emphasis>
|
||
</taxonomicName>
|
||
S7–8 are brown with yellowish venter and S9–10 complete brown. In female
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FAF27016FA680687" box="[1297,1436,1644,1670]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="himalaica">
|
||
<emphasis id="B977106AFFEDFFF4FAF27016FA680687" box="[1297,1436,1644,1670]" italics="true" pageId="5" pageNumber="142">P. himalaica</emphasis>
|
||
</taxonomicName>
|
||
the ventro-lateral sides of S1–3 are bluish and S8 has a large lateral blue spot, while in
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FBB770EAFB1206A8" authorityName="Gurung & Phan" authorityYear="2023" box="[1108,1254,1680,1705]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="samtsensis">
|
||
<emphasis id="B977106AFFEDFFF4FBB770EAFB1206A8" box="[1108,1254,1680,1705]" italics="true" pageId="5" pageNumber="142">P. samtsensis</emphasis>
|
||
</taxonomicName>
|
||
S1–3, including lateral S8 are brown (
|
||
<figureCitation id="1338D0FDFFEDFFF4FE6A70CEFE1406CF" box="[393,480,1716,1742]" captionStart="FIGURE 3" captionStartId="2.[151,250,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-18@2.[151,1436,181,1337]" captionTargetPageId="2" captionText="FIGURE 3. Habitus of Protosticta samtsensis sp. nov. (A), holotype ♁ & (B), paratype ♀." figureDoi="http://doi.org/10.5281/zenodo.8407023" httpUri="https://zenodo.org/record/8407023/files/figure.png" pageId="5" pageNumber="142">Fig. 3B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBCCC78FFEDFFF4FF2470ADFB7207CB" blockId="5.[151,1437,815,1994]" pageId="5" pageNumber="142">
|
||
<emphasis id="B977106AFFEDFFF4FF2470ADFE4F06F3" bold="true" box="[199,443,1751,1778]" pageId="5" pageNumber="142">Habitat and ecology.</emphasis>
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FE2270A2FD4F06F0" authorityName="Gurung & Phan" authorityYear="2023" box="[449,699,1752,1777]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="samtsensis">
|
||
<emphasis id="B977106AFFEDFFF4FE2270A2FD4F06F0" box="[449,699,1752,1777]" italics="true" pageId="5" pageNumber="142">Protosticta samtsensis</emphasis>
|
||
</taxonomicName>
|
||
inhabits moist, shady seepages with thick tree canopies and it was collected from
|
||
<quantity id="4CFB619DFFEDFFF4FEA37081FE6F0717" box="[320,411,1787,1814]" metricMagnitude="3" metricUnit="m" metricValue="1.264" pageId="5" pageNumber="142" unit="m" value="1264.0">1,264 m</quantity>
|
||
a.s.l.
|
||
<taxonomicName id="4C03B7FBFFEDFFF4FE3F7086FD3A0717" box="[476,718,1788,1814]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="himalaica">
|
||
<emphasis id="B977106AFFEDFFF4FE3F7086FD3A0717" box="[476,718,1788,1814]" italics="true" pageId="5" pageNumber="142">Protosticta himalaica</emphasis>
|
||
</taxonomicName>
|
||
also occurs in similar habitat
|
||
<typeStatus id="54B872DAFFEDFFF4FBF87086FBA20717" box="[1051,1110,1788,1814]" pageId="5" pageNumber="142">types</typeStatus>
|
||
within the elevation range of
|
||
<quantity id="4CFB619DFFEDFFF4FF747165FED8073B" box="[151,300,1823,1850]" metricMagnitude="2" metricUnit="m" metricValue="9.25" metricValueMax="15.0" metricValueMin="3.5" pageId="5" pageNumber="142" unit="m" value="925.0" valueMax="1500.0" valueMin="350.0">350–1,500 m</quantity>
|
||
a.s.l in
|
||
<collectingCountry id="F3148CE8FFEDFFF4FE66715AFE2E073B" box="[389,474,1824,1850]" name="Bhutan" pageId="5" pageNumber="142">Bhutan</collectingCountry>
|
||
. The new species was observed taking short flights and devouring small flies. A male and a female were collected from the same habitat. The side vegetation was a mixed-broad-leaf forest with thick fern undergrowth. The stream called
|
||
<emphasis id="B977106AFFEDFFF4FDCC7112FD5E0783" box="[559,682,1896,1922]" italics="true" pageId="5" pageNumber="142">Seti Khola,</emphasis>
|
||
meaning white stream, had a fast current. Along the sides, there was thick riparian vegetation and a spring that originated from the base of a cliff nearby fed into the stream. The forest floor has black soil with decomposed leaf litter, and it remains wet for most of the season.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |