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<document id="F438A78244D7C079C95F51A5081E6CB7" ID-DOI="10.11646/zootaxa.3664.2.1" ID-GBIF-Dataset="a50b71f5-48a6-464f-aea1-f29bbc8ee322" ID-ISSN="1175-5326" ID-Zenodo-Dep="220503" ID-ZooBank="105D178E-6DBF-401D-85BB-CDB041B67E91" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460615203542" checkinUser="plazi" docAuthor="Shi, Hongliang, Sciaky, Riccardo, Liang, Hongbin &amp; Zhou, Hongzhang" docDate="2013" docId="03AA10085650FFE5C1D8FC27FDFDFD52" docLanguage="en" docName="zt03664p135.pdf" docOrigin="Zootaxa 3664 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Wraseiellus) comatus Shi &amp; Sciaky, new species" docType="treatment" docVersion="9" lastPageNumber="113" masterDocId="FF936870565BFFE9C14FFF95FFA9FF9E" masterDocTitle="A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions" masterLastPageNumber="135" masterPageNumber="101" pageNumber="112" updateTime="1698330049886" updateUser="plazi">
<mods:mods id="6020D79043C04CBE518D2F3D8EC94CB4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4578070761AC54D315B02FB389FB4615">A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions</mods:title>
</mods:titleInfo>
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<mods:namePart id="FD467814B043B9A0617443DECAEDAF7B">Shi, Hongliang</mods:namePart>
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<mods:namePart id="17693D63D73B4F61616723E7DC86DE25">Sciaky, Riccardo</mods:namePart>
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<mods:namePart id="B98156D54F996D010DE2A0B95717F703">Liang, Hongbin</mods:namePart>
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<mods:namePart id="3C1F49C576DB529865CF722CA76C8B65">Zhou, Hongzhang</mods:namePart>
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<mods:title id="25E3671690A69ACCC15F8C3741157E74">Zootaxa</mods:title>
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<mods:part id="17D5168A7ED63976AD71580D5A892A66">
<mods:date id="1AF43CBEFCB4FEE3D0F4DF221E6673D8">2013</mods:date>
<mods:detail id="1386EAA6F7E360E1A30149C51E2611BE" type="volume">
<mods:number id="BB928247E56D053509CBF51DBDAFD3BD">3664</mods:number>
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<mods:identifier id="BDC706179428B887BFCFFF7C8F7D4447" type="DOI">10.11646/zootaxa.3664.2.1</mods:identifier>
<mods:identifier id="2829EEBBFCC625086C87B451BB348B0B" type="GBIF-Dataset">a50b71f5-48a6-464f-aea1-f29bbc8ee322</mods:identifier>
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<treatment id="03AA10085650FFE5C1D8FC27FDFDFD52" ID-DOI="http://doi.org/10.5281/zenodo.6154509" ID-GBIF-Taxon="119568525" ID-Zenodo-Dep="6154509" LSID="urn:lsid:plazi:treatment:03AA10085650FFE5C1D8FC27FDFDFD52" httpUri="http://treatment.plazi.org/id/03AA10085650FFE5C1D8FC27FDFDFD52" lastPageId="12" lastPageNumber="113" pageId="11" pageNumber="112">
<subSubSection id="C319F2955650FFE2C1D8FC27FDB4FC73" pageId="11" pageNumber="112" type="nomenclature">
<paragraph id="8BBCA11E5650FFE2C1D8FC27FC2FFC52" blockId="11.[151,902,946,1005]" box="[151,902,946,972]" pageId="11" pageNumber="112">
<heading id="D0F416725650FFE2C1D8FC27FC2FFC52" bold="true" box="[151,902,946,972]" fontSize="11" level="1" pageId="11" pageNumber="112" reason="1">
<emphasis id="B9777D0C5650FFE2C1D8FC27FC2FFC52" bold="true" box="[151,902,946,972]" pageId="11" pageNumber="112">
<taxonomicName id="4C03DA9D5650FFE2C1D8FC27FD40FC52" ID-CoL="4Q4HL" authority="Shi &amp; Sciaky" authorityName="Shi &amp; Sciaky" box="[151,745,946,972]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="112" phylum="Arthropoda" rank="species" species="comatus" status="sp. nov." subGenus="Wraseiellus">
<emphasis id="B9777D0C5650FFE2C1D8FC27FE84FC52" bold="true" box="[151,301,946,972]" italics="true" pageId="11" pageNumber="112">Pterostichus</emphasis>
(
<emphasis id="B9777D0C5650FFE2C072FC27FE61FC52" bold="true" box="[317,456,946,972]" italics="true" pageId="11" pageNumber="112">Wraseiellus</emphasis>
)
<emphasis id="B9777D0C5650FFE2C095FC27FD96FC52" bold="true" box="[474,575,946,972]" italics="true" pageId="11" pageNumber="112">comatus</emphasis>
Shi &amp; Sciaky
</taxonomicName>
,
<taxonomicNameLabel id="A244C0775650FFE2C3B9FC27FC2FFC52" box="[758,902,946,972]" pageId="11" pageNumber="112" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBCA11E5650FFE2C1D8FC40FDB4FC73" blockId="11.[151,902,946,1005]" box="[151,541,981,1005]" pageId="11" pageNumber="112">
(
<figureCitation id="1338BD9B5650FFE2C1D1FC40FEA2FC73" box="[158,267,981,1005]" captionStart="FIGURES 1 – 6" captionStartId="21.[151,264,1774,1797]" captionTargetBox="[159,1429,200,1748]" captionTargetId="figure@21.[151,1436,195,1751]" captionTargetPageId="21" captionText="FIGURES 1 – 6. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 1, Pt. andrewesi Jedlička lectotype; 2, Pt. andrewesi Jedlička, a female from Jinfoshan; 3, Pt. meyeri Jedlička, holotype; 4, Pt. meyeri Jedlička, holotype of Feronia crebrepunctata Straneo; 5, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 6, Pt. comatus n. sp., holotype." httpUri="https://zenodo.org/record/220504/files/figure.png" pageId="11" pageNumber="112">Figures 6</figureCitation>
,
<figureCitation id="1338BD9B5650FFE2C058FC40FE9FFC73" box="[279,310,981,1005]" captionStart="FIGURES 19 – 28" captionStartId="24.[151,264,1308,1331]" captionTargetBox="[166,1426,208,1270]" captionTargetId="figure@24.[151,1436,193,1299]" captionTargetPageId="24" captionText="FIGURES 19 – 28. Labels of name-bearing type specimen: 19, lectotype of Pterostichus andrewesi Jedlička, 1931; 20, holotype of Pterostichus meyeri Jedlička, 1934; 21, holotype of Feronia crebrepunctata Straneo, 1937; 22, holotype of Pterostichus comatus n. sp.; 23, holotype of Pterostichus crassiapex n. sp.; 24, a syntype of Omaseus diversus Fairmaire, 1886; 25, holotype of Pterostichus stictopleurus cangshanensis n. ssp.; 26, holotype of Pterostichus pseudodiversus n. sp.; 27, holotype of Feronia kambaiti Andrewes, 1947 [photoed by Dr. J. Bergsten]; 28, holotype of Pterostichus kambaiti Jedlička, 1965." httpUri="https://zenodo.org/record/220507/files/figure.png" pageId="11" pageNumber="112">22</figureCitation>
,
<figureCitation id="1338BD9B5650FFE2C00EFC40FEC9FC73" box="[321,352,981,1005]" captionStart="FIGURES 29 – 34" captionStartId="25.[151,264,1854,1877]" captionTargetBox="[176,1418,197,1824]" captionTargetId="figure@25.[151,1436,193,1833]" captionTargetPageId="25" captionText="FIGURES 29 – 34. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe. 29, Pt. andrewesi Jedlička, a male from Jinfoshan; 30, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 31, Pt. crassiapex n. sp., holotype; 32, Pt. comatus n. sp., holotype; 33, Pt. diversus (Fairmaire), a male from Yulongshan; 34, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;." httpUri="https://zenodo.org/record/220508/files/figure.png" pageId="11" pageNumber="112">32</figureCitation>
,
<figureCitation id="1338BD9B5650FFE2C024FC40FE20FC73" box="[363,393,981,1005]" captionStart="FIGURES 57 – 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 – 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 64, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="11" pageNumber="112">60</figureCitation>
,
<figureCitation id="1338BD9B5650FFE2C0DAFC40FE1AFC73" box="[405,435,981,1005]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="11" pageNumber="112">70</figureCitation>
,
<figureCitation id="1338BD9B5650FFE2C0F0FC40FE74FC73" box="[447,477,981,1005]" captionStart="FIGURES 94 – 97" captionStartId="31.[151,264,1762,1785]" captionText="FIGURES 94 – 97. Male terminal sternum of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; 94, Pt. andrewesi Jedlička, a male from Jinfoshan; 95, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 96, Pt. crassiapex n. sp., holotype; 97, Pt. comatus n. sp., a paratype from Maoershan." pageId="11" pageNumber="112">97</figureCitation>
,
<figureCitation id="1338BD9B5650FFE2C0A6FC40FDBDFC73" box="[489,532,981,1005]" captionStart="FIGURES 98 – 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 – 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="11" pageNumber="112">101</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C319F2955650FFE2C1D8FB88FD25FAA9" pageId="11" pageNumber="112" type="materials_examined">
<paragraph id="8BBCA11E5650FFE2C1D8FB88FBE3FBE3" blockId="11.[151,1436,1053,2019]" pageId="11" pageNumber="112">
<emphasis id="B9777D0C5650FFE2C1D8FB88FE84FBA8" bold="true" box="[151,301,1053,1078]" pageId="11" pageNumber="112">
<typeStatus id="54B81FBC5650FFE2C1D8FB88FF7BFBAB" box="[151,210,1053,1077]" pageId="11" pageNumber="112">Type</typeStatus>
locality
</emphasis>
. Guangxi: Maoershan Mt. (
<geoCoordinate id="EE37C7D95650FFE2C32AFB88FD6CFBAB" box="[613,709,1053,1077]" direction="north" orientation="latitude" pageId="11" pageNumber="112" precision="555" value="25.87">N25.87º</geoCoordinate>
,
<geoCoordinate id="EE37C7D95650FFE2C39FFB88FC92FBAB" box="[720,827,1053,1077]" direction="east" orientation="longitude" pageId="11" pageNumber="112" precision="555" value="110.42">E110.42º</geoCoordinate>
). The label of
<typeStatus id="54B81FBC5650FFE2C294FB88FB94FBAB" box="[987,1085,1053,1077]" pageId="11" pageNumber="112" type="holotype">holotype</typeStatus>
does not indicate its altitude or exact locality. But from the altitude information on the two
<typeStatus id="54B81FBC5650FFE2C20FFBD7FC02FBC4" box="[832,939,1090,1114]" pageId="11" pageNumber="112" type="paratype">paratypes</typeStatus>
(
<quantity id="4CFB0CFB5650FFE2C2F4FBD4FBA7FBC7" box="[955,1038,1089,1113]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="112" unit="m" value="1900.0">1900m</quantity>
), we suppose that this species only lives in the very high altitude of that mountain, which has its highest peak at
<quantity id="4CFB0CFB5650FFE2C2BBFBF0FBEFFBE3" box="[1012,1094,1125,1149]" metricMagnitude="3" metricUnit="m" metricValue="2.142" pageId="11" pageNumber="112" unit="m" value="2142.0">2142m</quantity>
.
</paragraph>
<paragraph id="8BBCA11E5650FFE2C188FB1FFD25FAA9" blockId="11.[151,1436,1053,2019]" pageId="11" pageNumber="112">
<emphasis id="B9777D0C5650FFE2C188FB1FFED4FB3C" bold="true" box="[199,381,1161,1186]" pageId="11" pageNumber="112">
<typeStatus id="54B81FBC5650FFE2C188FB1FFEADFB3C" box="[199,260,1162,1186]" pageId="11" pageNumber="112">Type</typeStatus>
materials
</emphasis>
.
<emphasis id="B9777D0C5650FFE2C0C3FB1FFE50FB3C" bold="true" box="[396,505,1162,1186]" pageId="11" pageNumber="112">
<typeStatus id="54B81FBC5650FFE2C0C3FB1FFE50FB3C" box="[396,505,1162,1186]" pageId="11" pageNumber="112" type="holotype">Holotype</typeStatus>
</emphasis>
(MNHN): male, body length =
<quantity id="4CFB0CFB5650FFE2C224FB1CFC7BFB3F" box="[875,978,1161,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="11" pageNumber="112" unit="mm" value="12.8">12.8 mm</quantity>
, board mounted, genitalia dissected and glued on plastic film pinned under specimen, &quot;
<collectingCountry id="F314E18E5650FFE2C3FDFB38FCA3FB5B" box="[690,778,1197,1221]" name="China" pageId="11" pageNumber="112">CHINE</collectingCountry>
(GUANGXI) / MAO'ER SHAN /
<date id="FFBD87DE5650FFE2C5EFFB38FAA1FB5B" box="[1184,1288,1197,1221]" pageId="11" pageNumber="112" value="1995-07">VII-1995</date>
T.DEUVE&quot;; &quot;Museum Paris&quot;; &quot;
<typeStatus id="54B81FBC5650FFE2C026FB44FDA9FB77" box="[361,512,1233,1257]" pageId="11" pageNumber="112" type="holotype">HOLOTYPE</typeStatus>
3/
<taxonomicName id="4C03DA9D5650FFE2C361FB44FCFFFB77" authority="Wraseiellus" authorityName="Wraseiellus" box="[558,854,1233,1257]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="112" phylum="Arthropoda" rank="genus">Pterostichus (Wraseiellus)</taxonomicName>
/
<taxonomicName id="4C03DA9D5650FFE2C220FB47FC65FB77" box="[879,972,1234,1257]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="112" phylum="Arthropoda" rank="species" species="comatus" status="sp. nov.">comatus</taxonomicName>
<taxonomicNameLabel id="A244C0775650FFE2C29AFB47FBF4FB77" box="[981,1117,1233,1257]" pageId="11" pageNumber="112" rank="species">new species</taxonomicNameLabel>
/ Des. SHI H.L. 2012&quot; [red label] [
<figureCitation id="1338BD9B5650FFE2C1AEFB60FEB8FA9D" box="[225,273,1269,1283]" captionStart="FIGURES 1 – 6" captionStartId="21.[151,264,1774,1797]" captionTargetBox="[159,1429,200,1748]" captionTargetId="figure@21.[151,1436,195,1751]" captionTargetPageId="21" captionText="FIGURES 1 – 6. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 1, Pt. andrewesi Jedlička lectotype; 2, Pt. andrewesi Jedlička, a female from Jinfoshan; 3, Pt. meyeri Jedlička, holotype; 4, Pt. meyeri Jedlička, holotype of Feronia crebrepunctata Straneo; 5, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 6, Pt. comatus n. sp., holotype." httpUri="https://zenodo.org/record/220504/files/figure.png" pageId="11" pageNumber="112">Figs. 6</figureCitation>
,
<figureCitation id="1338BD9B5650FFE2C055FB60FE82FA9D" box="[282,299,1269,1283]" captionStart="FIGURES 19 – 28" captionStartId="24.[151,264,1308,1331]" captionTargetBox="[166,1426,208,1270]" captionTargetId="figure@24.[151,1436,193,1299]" captionTargetPageId="24" captionText="FIGURES 19 – 28. Labels of name-bearing type specimen: 19, lectotype of Pterostichus andrewesi Jedlička, 1931; 20, holotype of Pterostichus meyeri Jedlička, 1934; 21, holotype of Feronia crebrepunctata Straneo, 1937; 22, holotype of Pterostichus comatus n. sp.; 23, holotype of Pterostichus crassiapex n. sp.; 24, a syntype of Omaseus diversus Fairmaire, 1886; 25, holotype of Pterostichus stictopleurus cangshanensis n. ssp.; 26, holotype of Pterostichus pseudodiversus n. sp.; 27, holotype of Feronia kambaiti Andrewes, 1947 [photoed by Dr. J. Bergsten]; 28, holotype of Pterostichus kambaiti Jedlička, 1965." httpUri="https://zenodo.org/record/220507/files/figure.png" pageId="11" pageNumber="112">22</figureCitation>
,
<figureCitation id="1338BD9B5650FFE2C07BFB60FEECFA9D" box="[308,325,1269,1283]" captionStart="FIGURES 29 – 34" captionStartId="25.[151,264,1854,1877]" captionTargetBox="[176,1418,197,1824]" captionTargetId="figure@25.[151,1436,193,1833]" captionTargetPageId="25" captionText="FIGURES 29 – 34. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe. 29, Pt. andrewesi Jedlička, a male from Jinfoshan; 30, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 31, Pt. crassiapex n. sp., holotype; 32, Pt. comatus n. sp., holotype; 33, Pt. diversus (Fairmaire), a male from Yulongshan; 34, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;." httpUri="https://zenodo.org/record/220508/files/figure.png" pageId="11" pageNumber="112">32</figureCitation>
,
<figureCitation id="1338BD9B5650FFE2C001FB60FEF6FA9D" box="[334,351,1269,1283]" captionStart="FIGURES 57 – 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 – 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 64, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="11" pageNumber="112">60</figureCitation>
,
<figureCitation id="1338BD9B5650FFE2C027FB60FED0FA9D" box="[360,377,1269,1283]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="11" pageNumber="112">70</figureCitation>
,
<figureCitation id="1338BD9B5650FFE2C0CDFB60FE33FA9D" box="[386,410,1269,1283]" captionStart="FIGURES 98 – 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 – 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="11" pageNumber="112">101</figureCitation>
].
<emphasis id="B9777D0C5650FFE2C0FEFB6DFD80FA8E" bold="true" box="[433,553,1272,1296]" pageId="11" pageNumber="112">
<typeStatus id="54B81FBC5650FFE2C0FEFB6DFD80FA8E" box="[433,553,1272,1296]" pageId="11" pageNumber="112" type="paratype">Paratypes</typeStatus>
</emphasis>
: 2 teneral males (IZAS and SCAU), &quot;Guangxi, Maoershan, Sanjiangyuan,
<quantity id="4CFB0CFB5650FFE2C1D8FA8AFF4EFAA9" box="[151,231,1311,1335]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="112" unit="m" value="1900.0">1900m</quantity>
, 2003.8.25, TIAN Mingyi leg.&quot; [
<figureCitation id="1338BD9B5650FFE2C31CFA89FD28FAB4" box="[595,641,1308,1322]" captionStart="FIGURES 94 – 97" captionStartId="31.[151,264,1762,1785]" captionText="FIGURES 94 – 97. Male terminal sternum of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; 94, Pt. andrewesi Jedlička, a male from Jinfoshan; 95, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 96, Pt. crassiapex n. sp., holotype; 97, Pt. comatus n. sp., a paratype from Maoershan." pageId="11" pageNumber="112">Fig. 97</figureCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="C319F2955650FFE2C188FAD7FEBEFA3D" pageId="11" pageNumber="112" type="diagnosis">
<paragraph id="8BBCA11E5650FFE2C188FAD7FEBEFA3D" blockId="11.[151,1436,1053,2019]" pageId="11" pageNumber="112">
<emphasis id="B9777D0C5650FFE2C188FAD7FEE8FAC5" bold="true" box="[199,321,1346,1371]" pageId="11" pageNumber="112">Diagnosis.</emphasis>
This new species can be easily distinguished from all the other species of this subgenus by their lateral margins of pronotum with three to five setae on each side, contrasting other species only with a single midlateral seta.
</paragraph>
</subSubSection>
<subSubSection id="C319F2955650FFE5C188FA3BFBABFE51" lastPageId="12" lastPageNumber="113" pageId="11" pageNumber="112" type="description">
<paragraph id="8BBCA11E5650FFE5C188FA3BFBABFE51" blockId="11.[151,1436,1053,2019]" lastBlockId="12.[151,1436,151,716]" lastPageId="12" lastPageNumber="113" pageId="11" pageNumber="112">
<emphasis id="B9777D0C5650FFE2C188FA3BFEF8FA59" bold="true" box="[199,337,1454,1479]" pageId="11" pageNumber="112">Description</emphasis>
. Body length
<quantity id="4CFB0CFB5650FFE2C0BDFA3AFD30FA59" box="[498,665,1455,1479]" metricMagnitude="-2" metricUnit="m" metricValue="1.255" metricValueMax="1.28" metricValueMin="1.23" pageId="11" pageNumber="112" unit="mm" value="12.55" valueMax="12.8" valueMin="12.3">12.3–12.8 mm</quantity>
; dorsal side black or with elytra slightly brownish; legs dark brown, tarsi light brown; ventral side black or dark brown. Males with elytral microsculpture transverse on the basal half, gradually turning to isodiametric after middle.
<emphasis id="B9777D0C5650FFE2C3F9FA62FD5FF991" bold="true" box="[694,758,1527,1551]" pageId="11" pageNumber="112">Head</emphasis>
. Frons finely and sparsely punctate, densely in the frontal furrows; antennae reaching elytra basal fifth; 3rd antennomere glabrous except apical setae; males with the terminal labial palpomere tubular.
<emphasis id="B9777D0C5650FFE2C378F9AAFD19F9C9" bold="true" box="[567,688,1599,1623]" pageId="11" pageNumber="112">Pronotum</emphasis>
cordiform, widest before middle, PW/PL = 1.26–1.32; lateral margins fully rounded near the middle, and strongly sinuate just before hind angles; three to five mid-lateral setae present before middle, hind angles distinct and sharp, rectangular; basal fovea moderately deep, faintly defined; inner groove of basal fovea subparallel to median line, slightly curved to the outside, ended at a distance from the posterior margin of pronotum; outer groove of basal fovea about half length as the inner one, deep and reaching the basal margin of pronotum; ridge between outer groove and lateral margin distinct; area between inner and outer basal foveal grooves rugose and hardly convex; basal fovea area, as well as area between two basal foveae finely and densely punctate; some very fine and sparse punctures also present in the lateral depression and, sometimes, on the lateral area of disc.
<emphasis id="B9777D0C5650FFE2C0E1F8CAFE53F8E9" bold="true" box="[430,506,1887,1911]" pageId="11" pageNumber="112">Elytra</emphasis>
with base slightly depressed between 3rd and 5th intervals; humeral teeth very small; intervals slightly convex; striae moderately deep, without punctures; scutellar striae incomplete, forming short grooves, located between 1st and 2nd striae; umbilical pore series on 9th interval sparse in the middle, composed of 14–15 pores (6, 2, 6–7).
<emphasis id="B9777D0C5650FFE2C309F85EFD71F87D" bold="true" box="[582,728,1994,2019]" pageId="11" pageNumber="112">Ventral side</emphasis>
. Proepisterna and metaepisterna finely punctate, posterior half of proepisterna with punctures sparser; terminal sternum of males with a very faint transverse ridge at about anterior fourth, and slightly depressed after the ridge (
<figureCitation id="1338BD9B5657FFE5C3BCFF29FCE1FF4A" box="[755,840,188,212]" captionStart="FIGURES 94 – 97" captionStartId="31.[151,264,1762,1785]" captionText="FIGURES 94 – 97. Male terminal sternum of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; 94, Pt. andrewesi Jedlička, a male from Jinfoshan; 95, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 96, Pt. crassiapex n. sp., holotype; 97, Pt. comatus n. sp., a paratype from Maoershan." pageId="12" pageNumber="113">Fig. 97</figureCitation>
).
<emphasis id="B9777D0C5657FFE5C214FF29FC3AFF4A" bold="true" box="[859,915,188,212]" pageId="12" pageNumber="113">Legs</emphasis>
. 5th tarsomere with two or three pairs of setae beneath.
<emphasis id="B9777D0C5657FFE5C1B2FF75FE01FF66" bold="true" box="[253,424,223,248]" pageId="12" pageNumber="113">Male genitalia</emphasis>
. Median lobe of male genitalia bent less than 90 degree, apex slightly bent to the ventral side and then the turned to the dorsal side (
<figureCitation id="1338BD9B5657FFE5C3CEFE91FD7DFE82" box="[641,724,260,284]" captionStart="FIGURES 29 – 34" captionStartId="25.[151,264,1854,1877]" captionTargetBox="[176,1418,197,1824]" captionTargetId="figure@25.[151,1436,193,1833]" captionTargetPageId="25" captionText="FIGURES 29 – 34. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe. 29, Pt. andrewesi Jedlička, a male from Jinfoshan; 30, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 31, Pt. crassiapex n. sp., holotype; 32, Pt. comatus n. sp., holotype; 33, Pt. diversus (Fairmaire), a male from Yulongshan; 34, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;." httpUri="https://zenodo.org/record/220508/files/figure.png" pageId="12" pageNumber="113">Fig. 32</figureCitation>
A); ventral margin almost straight in the middle, dorsal margin gradually curved; apical orifice large, slightly turned to the left side; apical lamella thick, apex slightly thickened and turned to the dorsal side (
<figureCitation id="1338BD9B5657FFE5C0A7FED9FDEEFEFA" box="[488,583,332,356]" captionStart="FIGURES 29 – 34" captionStartId="25.[151,264,1854,1877]" captionTargetBox="[176,1418,197,1824]" captionTargetId="figure@25.[151,1436,193,1833]" captionTargetPageId="25" captionText="FIGURES 29 – 34. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe. 29, Pt. andrewesi Jedlička, a male from Jinfoshan; 30, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 31, Pt. crassiapex n. sp., holotype; 32, Pt. comatus n. sp., holotype; 33, Pt. diversus (Fairmaire), a male from Yulongshan; 34, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;." httpUri="https://zenodo.org/record/220508/files/figure.png" pageId="12" pageNumber="113">Figs. 32</figureCitation>
B, 60B); when in dorsal view, apical lamella narrow and long, length about 2.5 times as basal width, twisted to the left (the widest surface of apical lamella orientated to the dorsal-left side of median lobe), strongly bent to the left side (
<figureCitation id="1338BD9B5657FFE5C3C7FE01FD74FE32" box="[648,733,404,428]" captionStart="FIGURES 29 – 34" captionStartId="25.[151,264,1854,1877]" captionTargetBox="[176,1418,197,1824]" captionTargetId="figure@25.[151,1436,193,1833]" captionTargetPageId="25" captionText="FIGURES 29 – 34. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe. 29, Pt. andrewesi Jedlička, a male from Jinfoshan; 30, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 31, Pt. crassiapex n. sp., holotype; 32, Pt. comatus n. sp., holotype; 33, Pt. diversus (Fairmaire), a male from Yulongshan; 34, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;." httpUri="https://zenodo.org/record/220508/files/figure.png" pageId="12" pageNumber="113">Fig. 32</figureCitation>
B); apex of apical lamella oblique to the left (
<figureCitation id="1338BD9B5657FFE5C5AFFE01FA9AFE32" box="[1248,1331,404,428]" captionStart="FIGURES 57 – 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 – 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 64, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="12" pageNumber="113">Fig. 60</figureCitation>
A). Right paramere fine and long, sickle-form, apex sharp (
<figureCitation id="1338BD9B5657FFE5C3F1FE22FCBAFE51" box="[702,787,439,463]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="12" pageNumber="113">Fig. 70</figureCitation>
B). Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C319F2955657FFE5C188FE4EFB35FE6A" box="[199,1180,475,501]" pageId="12" pageNumber="113" type="distribution">
<paragraph id="8BBCA11E5657FFE5C188FE4EFB35FE6A" blockId="12.[151,1436,151,716]" box="[199,1180,475,501]" pageId="12" pageNumber="113">
<emphasis id="B9777D0C5657FFE5C188FE4EFEF0FE6A" bold="true" box="[199,345,475,500]" pageId="12" pageNumber="113">Distribution</emphasis>
. Only known from the
<typeStatus id="54B81FBC5657FFE5C32FFE48FD39FE6B" box="[608,656,477,501]" pageId="12" pageNumber="113">type</typeStatus>
locality, Maoershan Mt. (Guangxi) (
<figureCitation id="1338BD9B5657FFE5C562FE49FB39FE6A" box="[1069,1168,476,500]" captionStart="FIGURES 110 – 111" captionStartId="33.[151,264,1885,1908]" captionTargetBox="[224,1312,207,1827]" captionTargetId="figure@33.[224,1343,193,1844]" captionTargetPageId="33" captionText="FIGURES 110 – 111. Distribution maps for species of subgenus Pterostichus (Wraseiellus): - Pt. andrewesi Jedlička; « Pt. meyeri Jedlička; ˑ Pt. crassiapex n. sp.; ¿ Pt. comatus n. sp.; ● Pt. diversus (Fairmaire); Ο Pt. pseudodiversus n. sp.; ˔ Pt. stictopleurus stictopleurus (Fairmaire); r Pt. stictopleurus cangshanensis n. ssp.; ․ Pt. kambaiti (Andrewes). 111. shows an enlarged scale of the area in figure 110 marked by the dotted line square." httpUri="https://zenodo.org/record/220516/files/figure.png" pageId="12" pageNumber="113">Fig. 110</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C319F2955657FFE5C188FD95FE06FDA2" pageId="12" pageNumber="113" type="etymology">
<paragraph id="8BBCA11E5657FFE5C188FD95FE06FDA2" blockId="12.[151,1436,151,716]" pageId="12" pageNumber="113">
<emphasis id="B9777D0C5657FFE5C188FD95FEE6FD86" bold="true" box="[199,335,512,536]" pageId="12" pageNumber="113">Etymology.</emphasis>
The name &quot;
<taxonomicName id="4C03DA9D5657FFE5C0A6FD94FDEFFD86" box="[489,582,513,536]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="comatus">
<emphasis id="B9777D0C5657FFE5C0A6FD94FDEFFD86" box="[489,582,513,536]" italics="true" pageId="12" pageNumber="113">comatus</emphasis>
</taxonomicName>
&quot; comes from Latin, meaning hairy, referring to the additional setae on pronotum lateral margin.
</paragraph>
</subSubSection>
<subSubSection id="C319F2955657FFE5C188FDD2FDFDFD52" pageId="12" pageNumber="113" type="description">
<paragraph id="8BBCA11E5657FFE5C188FDD2FDFDFD52" blockId="12.[151,1436,151,716]" pageId="12" pageNumber="113">
<emphasis id="B9777D0C5657FFE5C188FDD2FEE8FDFE" bold="true" box="[199,321,583,608]" pageId="12" pageNumber="113">Variation.</emphasis>
The three specimens we studied are different in the number of lateral setae of pronotum: the
<typeStatus id="54B81FBC5657FFE5C1D8FDF9FF50FD1A" box="[151,249,620,644]" pageId="12" pageNumber="113" type="holotype">holotype</typeStatus>
with five lateral setae, but the two
<typeStatus id="54B81FBC5657FFE5C3CAFDF8FD58FD1B" box="[645,753,621,645]" pageId="12" pageNumber="113" type="paratype">paratypes</typeStatus>
with only three. We suppose that the number of lateral setae of pronotum may vary from individual to individual. Such case is also found in other members of
<taxonomicName id="4C03DA9D5657FFE5C599FD1AFACDFD39" box="[1238,1380,655,679]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="tribe" tribe="Pterostichini">Pterostichini</taxonomicName>
with three or more lateral setae of pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>
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