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<document id="793BF733761F15496E29DE3D17AE9ED1" ID-DOI="10.11646/zootaxa.5155.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="6669176" ID-ZooBank="6E9C5E63-48B6-4D29-B98D-3DA2755A5AB8" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1655713213397" checkinUser="plazi" docAuthor="Golovatch, Sergei I., Bouzan, Rodrigo S., Gallo, Jéssica S. &amp; Bichuette, Maria E." docDate="2022" docId="03AA0A59BB44F176FF487521FE43FC4E" docLanguage="en" docName="zootaxa.5155.1.4.pdf" docOrigin="Zootaxa 5155 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Strongylosomides troglobius Golovatch, Bouzan &amp; Gallo 2022, sp. nov." docType="treatment" docVersion="5" lastPageNumber="95" masterDocId="FF937221BB41F17EFFDF7046FFAAFFCF" masterDocTitle="A new genus and two new species of the millipede family Chelodesmidae from Bahia state, northeastern Brazil, including a likely troglobiont (Diplopoda, Polydesmida)" masterLastPageNumber="104" masterPageNumber="87" pageNumber="92" updateTime="1731957614320" updateUser="juliana" zenodo-license-document="CLOSED">
<mods:mods id="894FF437715209133E0DC5CB927FD1E1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D6357A3B04754D33428407D604F525F1">
<mods:title id="82B7E60809795A4A6018DA087AA19780">A new genus and two new species of the millipede family Chelodesmidae from Bahia state, northeastern Brazil, including a likely troglobiont (Diplopoda, Polydesmida)</mods:title>
</mods:titleInfo>
<mods:name id="4DF870BBBFEB9BC782B245682950079B" type="personal">
<mods:role id="3474EB72CD39470DBEE2A7E3E5A7F702">
<mods:roleTerm id="EC78B826C59526C0C009AC74E818DB85">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AAB20018732D5662D0B01E5979895CE1">Golovatch, Sergei I.</mods:namePart>
<mods:nameIdentifier id="C6CA126E4EC27EB5698E4A35F03B5EB8" type="ORCID">0000-0001-7159-5484</mods:nameIdentifier>
<mods:affiliation id="2B30C7EC41C4FF9E7B1276061EE01D6F">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia.</mods:affiliation>
<mods:nameIdentifier id="F685C2DCD77C98629477573DF8E6CC08" type="email">sgolovatch@yandex.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="5843649C77F48B94A2A6547CB85B1149" type="personal">
<mods:role id="7DA25C56249EEFFA4077CEB543077AC4">
<mods:roleTerm id="56C8C4E973F1DC477917A3B2E5897C90">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="17C9E20036775F786CA79A45BCA2DCBE">Bouzan, Rodrigo S.</mods:namePart>
<mods:nameIdentifier id="9C0EAED94501C41E7E486EA14DF979ED" type="ORCID">0000-0002-5331-7031</mods:nameIdentifier>
<mods:affiliation id="3E159B46750970F99238C39FE2FB3480">Laboratório de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brasil</mods:affiliation>
<mods:nameIdentifier id="48B405D89564D59E69B853F931AE624E" type="email">rodrigobouzan@outlook.com</mods:nameIdentifier>
</mods:name>
<mods:name id="E2493C4DA2DB361659C95CA9C0909CDB" type="personal">
<mods:role id="357B0DB0194AD4A8711660083BE2EA71">
<mods:roleTerm id="A68143BC6356ED45A07A0F54A3EC3AA7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="170CAB71EFE2323DA10F5EB9D524EF2A">Gallo, Jéssica S.</mods:namePart>
<mods:nameIdentifier id="964BAEF961C6FE3C21AE860D6C49B1D6" type="ORCID">0000-0001-9350-4686</mods:nameIdentifier>
<mods:affiliation id="5DABE539A37F3BBC3D49F6C271C82976">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos (UFSCar), Rodovia Washington Luis, km 235, 13565 - 905, CP 676. São Carlos, São Paulo, Brasil</mods:affiliation>
<mods:nameIdentifier id="A63D65BF65DC16575A881DDD890F36BF" type="email">jessicasgallo@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="B5EF21DBB5C9E6B55C981EE799A5434A" type="personal">
<mods:role id="7EFFFE19E2D56E067C1293210919D4B7">
<mods:roleTerm id="4F59E96FE50B62303661BE74A630062E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="58C0B9C7CE2D803A811F99F156DCC2E9">Bichuette, Maria E.</mods:namePart>
<mods:nameIdentifier id="4DF127C9978BD682D63889CA73E69C90" type="ORCID">0000-0002-9515-4832</mods:nameIdentifier>
<mods:affiliation id="EC2CDE00CAE2BCDCBCE293227E58E070">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos (UFSCar), Rodovia Washington Luis, km 235, 13565 - 905, CP 676. São Carlos, São Paulo, Brasil</mods:affiliation>
<mods:nameIdentifier id="0BCA31CA1A05FCA02AAC6C1A84081EEC" type="email">lina.cave@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="BDCAC69DF655FF407733CEF519E1B0B9">text</mods:typeOfResource>
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<mods:titleInfo id="48D755A2E3B8D1A76B8E0E5E42C17127">
<mods:title id="DCB2C5E226340DD9D9D152C7AD4358BB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CE8FA0389310C6B63FBF33FE22A02100">
<mods:date id="0357B396CAA84D40E6A3C305D90D361B">2022</mods:date>
<mods:detail id="CEEFF9F770D17202474C55B302D19490" type="pubDate">
<mods:number id="1325231607010A51785B6168127087E8">2022-06-20</mods:number>
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<mods:number id="57D8B1D9FA93DC37C0A44EE7163AC363">5155</mods:number>
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<mods:number id="83773B28CA739C94C04029861F58C99D">1</mods:number>
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<mods:classification id="3E326BE48B2B8AB90E6E6F083C960E6B">journal article</mods:classification>
<mods:identifier id="49956BE74F3501D2C3AB9C9274593DE9" type="DOI">10.11646/zootaxa.5155.1.4</mods:identifier>
<mods:identifier id="D46998876739F629D1FEB1649448BD71" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A5AFB20494F957FFEBE1DF267D62F1B7" type="Zenodo-Dep">6669176</mods:identifier>
<mods:identifier id="05F4EFBDF15E3CD60D55C0F3262F55E2" type="ZooBank">6E9C5E63-48B6-4D29-B98D-3DA2755A5AB8</mods:identifier>
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<treatment id="03AA0A59BB44F176FF487521FE43FC4E" ID-DOI="http://doi.org/10.5281/zenodo.6669352" ID-Zenodo-Dep="6669352" LSID="urn:lsid:plazi:treatment:03AA0A59BB44F176FF487521FE43FC4E" httpUri="http://treatment.plazi.org/id/03AA0A59BB44F176FF487521FE43FC4E" lastPageId="8" lastPageNumber="95" pageId="5" pageNumber="92">
<subSubSection id="C319E8C4BB44F17BFF487521FC11FA4D" box="[151,955,1383,1410]" pageId="5" pageNumber="92" type="nomenclature">
<paragraph id="8BBCBB4FBB44F17BFF487521FC11FA4D" blockId="5.[151,955,1383,1446]" box="[151,955,1383,1410]" pageId="5" pageNumber="92">
<heading id="D0F40C23BB44F17BFF487521FC11FA4D" bold="true" box="[151,955,1383,1410]" fontSize="11" level="1" pageId="5" pageNumber="92" reason="1">
<emphasis id="B977675DBB44F17BFF487521FC11FA4D" bold="true" box="[151,955,1383,1410]" pageId="5" pageNumber="92">
<taxonomicName id="4C03C0CCBB44F17BFF487521FCFBFA4E" ID-CoL="a0444644-259b-4a49-90a4-24c376458e35" authority="Golovatch, Bouzan &amp; Gallo" authorityName="Golovatch, Bouzan &amp; Gallo" authorityYear="2022" box="[151,849,1383,1410]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="B977675DBB44F17BFF487521FE44FA4D" bold="true" box="[151,494,1383,1410]" italics="true" pageId="5" pageNumber="92">Strongylosomides troglobius</emphasis>
Golovatch, Bouzan &amp; Gallo
</taxonomicName>
,
<taxonomicNameLabel id="A244DA26BB44F17BFC81752EFC11FA4D" box="[862,955,1384,1410]" pageId="5" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C319E8C4BB44F17BFF4875CDFF52FA69" box="[151,248,1419,1446]" pageId="5" pageNumber="92" type="description">
<paragraph id="8BBCBB4FBB44F17BFF4875CDFF52FA69" blockId="5.[151,955,1383,1446]" box="[151,248,1419,1446]" pageId="5" pageNumber="92">
<figureCitation id="1338A7CABB44F17BFF4875CDFF52FA69" box="[151,248,1419,1446]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStartId-0="7.[151,250,1808,1832]" captionStartId-1="8.[151,250,1914,1938]" captionStartId-2="9.[151,250,1840,1864]" captionStartId-3="10.[151,250,1944,1968]" captionTargetBox-0="[151,1435,266,1797]" captionTargetBox-1="[151,1436,921,1891]" captionTargetBox-2="[151,1436,181,1816]" captionTargetBox-3="[289,1298,181,1920]" captionTargetId-0="figure-145@7.[151,1435,266,1797]" captionTargetId-1="figure-390@8.[151,1436,921,1891]" captionTargetId-2="figure-17@9.[151,1436,181,1816]" captionTargetId-3="figure-22@10.[289,1298,181,1920]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." captionText-1="FIGURE 5. Strongylosomides troglobius sp. nov., male paratype. A &amp; B. Body ring 7 with intact gonopods, ventral and lateral views, respectively. C. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. D. Body ring 7 with left intact gonopod, caudal view. Photographs by K.V. Makarov, taken not to scale." captionText-2="FIGURE 6. Strongylosomides troglobius sp. nov., male paratype. A-F. Right gonopod, submesal, mesal, ventral, dorsal, sublateral and lateral views, respectively. Abbreviations: A, acropodite; AP, acropodital process; c, cannula; Cx, gonocoxa; Pf, prefemoral region; PfP, prefemoral process; Sg, seminal groove; sl, solenomere; Te, telopodite. Photographs by K.V. Makarov, taken not to scale." captionText-3="FIGURE 7. Strongylosomides troglobius sp. nov., male paratype. A-D. Right gonopod with prefemoral process (labeled PfP) highlighted and outlined in red, submesal, mesal, ventral and dorsal views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.6669197" figureDoi-1="http://doi.org/10.5281/zenodo.6669201" figureDoi-2="http://doi.org/10.5281/zenodo.6669205" figureDoi-3="http://doi.org/10.5281/zenodo.6669207" httpUri-0="https://zenodo.org/record/6669197/files/figure.png" httpUri-1="https://zenodo.org/record/6669201/files/figure.png" httpUri-2="https://zenodo.org/record/6669205/files/figure.png" httpUri-3="https://zenodo.org/record/6669207/files/figure.png" pageId="5" pageNumber="92">Figs 4–7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C319E8C4BB44F17BFF487595FAB6F9B1" pageId="5" pageNumber="92" type="materials_examined">
<paragraph id="8BBCBB4FBB44F17BFF487595FE65F9FA" blockId="5.[151,1436,1491,2022]" pageId="5" pageNumber="92">
<emphasis id="B977675DBB44F17BFF487595FE2EFA22" bold="true" box="[151,388,1491,1517]" pageId="5" pageNumber="92">Material examined.</emphasis>
<materialsCitation id="3B6BB112BB44F17BFE517595FE65F9FA" collectingDate="2011-11-02" collectionCode="R" collectorName="M. E. Bichuette &amp; J. E. Gallao &amp; C. S. Fernandes &amp; D. R. Pedroso" country="Brazil" latitude="-12.382112" location="Gruta da Baixa Fria cave" longLatPrecision="1" longitude="-44.93667" municipality="Sao Desiderio Municipality" pageId="5" pageNumber="92" specimenCode="LES 0027943" specimenCount="1" specimenCount-male="1" stateProvince="Bahia State" typeStatus="holotype">
<typeStatus id="54B805EDBB44F17BFE517595FE5CFA22" box="[398,502,1491,1517]" pageId="5" pageNumber="92" type="holotype">Holotype</typeStatus>
<specimenCount id="9D0570C6BB44F17BFDDF7595FD9CFA22" box="[512,566,1491,1517]" pageId="5" pageNumber="92" type="male">male</specimenCount>
(
<specimenCode id="DBA51334BB44F17BFD967592FD46FA21" box="[585,748,1492,1518]" collectionCode="LES" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13487" lsid="urn:lsid:biocol.org:col:13487" name="Leeds Museums and Galleries" pageId="5" pageNumber="92" type="Museum">LES 0027943</specimenCode>
),
<collectingCountry id="F314FBDFBB44F17BFCDD7595FCE3FA22" box="[770,841,1491,1517]" name="Brazil" pageId="5" pageNumber="92">Brazil</collectingCountry>
,
<collectingRegion id="49C775ADBB44F17BFC877595FC75FA21" box="[856,991,1491,1518]" country="Brazil" name="Bahia" pageId="5" pageNumber="92">Bahia State</collectingRegion>
,
<collectingMunicipality id="6BD82135BB44F17BFC327592FA80FA22" box="[1005,1322,1491,1518]" pageId="5" pageNumber="92">São Desidério Municipality</collectingMunicipality>
,
<location id="8EDCED94BB44F17BFAE77592FEE4F9DD" LSID="urn:lsid:plazi:treatment:03AA0A59BB44F176FF487521FE43FC4E:8EDCED94BB44F17BFAE77592FEE4F9DD" country="Brazil" latitude="-12.382112" longLatPrecision="1" longitude="-44.93667" municipality="Sao Desiderio Municipality" name="Gruta da Baixa Fria cave" pageId="5" pageNumber="92" stateProvince="Bahia State">Gruta da Baixa Fria cave</location>
, aphotic zone,
<geoCoordinate id="EE37DD88BB44F17BFE2275B1FD33F9DD" box="[509,665,1527,1554]" degrees="12" direction="south" minutes="22" orientation="latitude" pageId="5" pageNumber="92" precision="1" seconds="55.6" value="-12.382112">S12°22’55.6”</geoCoordinate>
,
<geoCoordinate id="EE37DD88BB44F17BFD7975B1FCE7F9DD" box="[678,845,1527,1554]" degrees="44" direction="west" minutes="56" orientation="longitude" pageId="5" pageNumber="92" precision="1" seconds="12.0" value="-44.93667">W44°56’12.0”</geoCoordinate>
,
<date id="FFBD9D8FBB44F17BFC8475BEFC71F9DD" box="[859,987,1528,1554]" pageId="5" pageNumber="92" value="2011-11-02">
<collectingDate id="EFF96467BB44F17BFC8475BEFC71F9DD" box="[859,987,1528,1554]" pageId="5" pageNumber="92" value="2011-11-02">02.XI.2011</collectingDate>
</date>
,
<collectorName id="26F6DE99BB44F17BFC3675BEFB33F9DE" box="[1001,1177,1527,1554]" pageId="5" pageNumber="92">M.E. Bichuette</collectorName>
,
<collectorName id="26F6DE99BB44F17BFB7975BEFA8DF9DE" box="[1190,1319,1527,1554]" pageId="5" pageNumber="92">J.E. Gallão</collectorName>
,
<collectorName id="26F6DE99BB44F17BFAEA75BEFF4FF9FA" pageId="5" pageNumber="92">C.S. Fernandes</collectorName>
&amp;
<collectorName id="26F6DE99BB44F17BFED6765AFE35F9FA" box="[265,415,1563,1590]" pageId="5" pageNumber="92">D.R. Pedroso</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BBCBB4FBB44F17BFF187606FB37F996" blockId="5.[151,1436,1491,2022]" box="[199,1181,1599,1626]" pageId="5" pageNumber="92">
<materialsCitation id="3B6BB112BB44F17BFF187606FB30F996" box="[199,1178,1599,1626]" collectionCode="LES" pageId="5" pageNumber="92" specimenCode="LES 0027944" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54B805EDBB44F17BFF187606FE92F995" box="[199,312,1600,1626]" pageId="5" pageNumber="92" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D0570C6BB44F17BFE9C7606FE3BF996" box="[323,401,1599,1626]" count="1" pageId="5" pageNumber="92" type="male">1 male</specimenCount>
,
<specimenCount id="9D0570C6BB44F17BFE437606FDA2F996" box="[412,520,1599,1626]" count="2" pageId="5" pageNumber="92" type="female">2 females</specimenCount>
(
<specimenCode id="DBA51334BB44F17BFDC87606FD12F995" box="[535,696,1600,1626]" collectionCode="LES" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13487" lsid="urn:lsid:biocol.org:col:13487" name="Leeds Museums and Galleries" pageId="5" pageNumber="92" type="Museum">LES 0027944</specimenCode>
), same place, taken together with holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BBCBB4FBB44F17BFF187622FAB6F9B1" blockId="5.[151,1436,1491,2022]" box="[199,1308,1635,1662]" pageId="5" pageNumber="92">
<emphasis id="B977675DBB44F17BFF187622FEA7F9B1" bold="true" box="[199,269,1636,1662]" pageId="5" pageNumber="92">Name</emphasis>
. To emphasize the cave-dwelling habits of this species, masculine adjective in genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C319E8C4BB44F178FF1876C1FD5EFE3A" lastPageId="6" lastPageNumber="93" pageId="5" pageNumber="92" type="diagnosis">
<paragraph id="8BBCBB4FBB44F178FF1876C1FD67FE8E" blockId="5.[151,1436,1491,2022]" lastBlockId="6.[151,1437,150,2014]" lastPageId="6" lastPageNumber="93" pageId="5" pageNumber="92">
<emphasis id="B977675DBB44F17BFF1876C1FE4AF96E" bold="true" box="[199,480,1671,1698]" pageId="5" pageNumber="92">Diagnosis and affinities</emphasis>
. This new
<taxonomicName id="4C03C0CCBB44F17BFDBF76C1FCD5F96E" box="[608,895,1671,1697]" pageId="5" pageNumber="92">
<emphasis id="B977675DBB44F17BFDBF76C1FC8CF96E" box="[608,806,1671,1697]" italics="true" pageId="5" pageNumber="92">Strongylosomides</emphasis>
species
</taxonomicName>
differs clearly from congeners by the light reddish ochre coloration and a rather fragile body with thin tegument (likely troglomorphisms), combined with the absence of modifications on pregonopodal sterna, and of gonocoxal distodorsal apophyses, but above all, by the prefemoral process (
<emphasis id="B977675DBB44F17BFEA176B5FE00F8C2" bold="true" box="[382,426,1779,1805]" pageId="5" pageNumber="92">PfP</emphasis>
) of the gonotelopodite (
<emphasis id="B977675DBB44F17BFD6276B2FD70F8C1" bold="true" box="[701,730,1780,1806]" pageId="5" pageNumber="92">Te</emphasis>
) being fully coalesced in its basal half with the acropodite (
<emphasis id="B977675DBB44F17BFAA076B5FA39F8C2" bold="true" box="[1407,1427,1779,1805]" pageId="5" pageNumber="92">A</emphasis>
) and leaving the distal half acuminate and directed caudad (
<figureCitation id="1338A7CABB44F17BFCF47751FCD9F8FE" box="[811,883,1815,1841]" captionStart="FIGURE 6" captionStartId="9.[151,250,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-17@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 6. Strongylosomides troglobius sp. nov., male paratype. A-F. Right gonopod, submesal, mesal, ventral, dorsal, sublateral and lateral views, respectively. Abbreviations: A, acropodite; AP, acropodital process; c, cannula; Cx, gonocoxa; Pf, prefemoral region; PfP, prefemoral process; Sg, seminal groove; sl, solenomere; Te, telopodite. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669205" httpUri="https://zenodo.org/record/6669205/files/figure.png" pageId="5" pageNumber="92">Figs 6</figureCitation>
&amp;
<figureCitation id="1338A7CABB44F17BFC48775EFC0DF8FD" box="[919,935,1816,1842]" captionStart="FIGURE 7" captionStartId="10.[151,250,1944,1968]" captionTargetBox="[289,1298,181,1920]" captionTargetId="figure-22@10.[289,1298,181,1920]" captionTargetPageId="10" captionText="FIGURE 7. Strongylosomides troglobius sp. nov., male paratype. A-D. Right gonopod with prefemoral process (labeled PfP) highlighted and outlined in red, submesal, mesal, ventral and dorsal views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669207" httpUri="https://zenodo.org/record/6669207/files/figure.png" pageId="5" pageNumber="92">7</figureCitation>
). The light ochre body coloration, as well as gonopodal shape and structure of
<taxonomicName id="4C03C0CCBB44F17BFDCC777AFD0BF89A" authority="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette, 2022" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" box="[531,673,1851,1878]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="B977675DBB44F17BFDCC777AFD0BF89A" box="[531,673,1851,1878]" italics="true" pageId="5" pageNumber="92">S. troglobius</emphasis>
</taxonomicName>
<emphasis id="B977675DBB44F17BFD77777AFCA8F899" bold="true" box="[680,770,1852,1878]" pageId="5" pageNumber="92">
<taxonomicNameLabel id="A244DA26BB44F17BFD77777AFCA8F899" box="[680,770,1852,1878]" pageId="5" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resemble those of
<taxonomicName id="4C03C0CCBB44F17BFC09777AFAE2F899" authority="(Schubart, 1956)" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" baseAuthorityName="Schubart" baseAuthorityYear="1956" box="[982,1352,1851,1878]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="stercoriarius" status="comb. nov.">
<emphasis id="B977675DBB44F17BFC09777AFB28F89A" box="[982,1154,1852,1878]" italics="true" pageId="5" pageNumber="92">S. stercoriarius</emphasis>
(
<bibRefCitation id="EF92C6BEBB44F17BFB4E777DFAEAF899" author="Schubart, O." box="[1169,1344,1851,1878]" pageId="5" pageNumber="92" pagination="355 - 367" refId="ref10067" refString="Schubart, O. 1956. &quot; Leptodesmidae &quot; brasileiras III. Especies de Minas Gerais (Diplopoda, Proterospermophora). Revista Brasileira de Biologia, 16 (3), 355 - 367." type="journal article" year="1956">Schubart, 1956</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B977675DBB44F17BFA8B777AFF62F8B5" bold="true" pageId="5" pageNumber="92">
<taxonomicNameLabel id="A244DA26BB44F17BFA8B777AFF62F8B5" pageId="5" pageNumber="92" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, taken from near a cave in
<collectingRegion id="49C775ADBB44F17BFDDE7719FD32F8B6" box="[513,664,1887,1913]" country="Brazil" name="Minas Gerais" pageId="5" pageNumber="92">Minas Gerais</collectingRegion>
(
<bibRefCitation id="EF92C6BEBB44F17BFD777719FCF8F8B5" author="Schubart, O." box="[680,850,1887,1914]" pageId="5" pageNumber="92" pagination="355 - 367" refId="ref10067" refString="Schubart, O. 1956. &quot; Leptodesmidae &quot; brasileiras III. Especies de Minas Gerais (Diplopoda, Proterospermophora). Revista Brasileira de Biologia, 16 (3), 355 - 367." type="journal article" year="1956">Schubart 1956</bibRefCitation>
), so vividly that both these species are undoubtedly very similar. They share the same, rather troglomorphic light ochre coloration, since both have been collected inside or near a cave, respectively. Both share the presence of male tibial chelae as well. More important, however, is that their gonopods are also very similar (
<figureCitation id="1338A7CABB44F17BFDE3778DFD33F829" box="[572,665,1995,2022]" captionStart="FIGURE 3" captionStartId="5.[151,250,1133,1157]" captionTargetBox="[160,1427,186,1102]" captionTargetId="figure-17@5.[151,1436,181,1109]" captionTargetPageId="5" captionText="FIGURE 3. Strongylosomides petropolites (Attems, 1901), male lectotype. A. Left gonopodal telopodite, mesal view (after Hoffman 1979), and Strongylosomides stercoriarius (Schubart, 1956), comb. nov., male syntype. B &amp; C. Right gonopod and its tip, mesal and sublateral views, respectively (after Schubart 1956). Abbreviations: A, acropodite; AP, acropodital process; c, cannula; Cx, gonocoxa; Pf, prefemoral region; PfP, prefemoral process; Sg, seminal groove; sl, solenomere; T, telopodite. Scale bar: 0.1 mm (B) or drawn not to scale (A, C)." figureDoi="http://doi.org/10.5281/zenodo.6669188" httpUri="https://zenodo.org/record/6669188/files/figure.png" pageId="5" pageNumber="92">Figs 3A</figureCitation>
,
<figureCitation id="1338A7CABB44F17BFD79778DFD1EF82A" box="[678,692,1995,2021]" captionStart="FIGURE 6" captionStartId="9.[151,250,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-17@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 6. Strongylosomides troglobius sp. nov., male paratype. A-F. Right gonopod, submesal, mesal, ventral, dorsal, sublateral and lateral views, respectively. Abbreviations: A, acropodite; AP, acropodital process; c, cannula; Cx, gonocoxa; Pf, prefemoral region; PfP, prefemoral process; Sg, seminal groove; sl, solenomere; Te, telopodite. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669205" httpUri="https://zenodo.org/record/6669205/files/figure.png" pageId="5" pageNumber="92">6</figureCitation>
&amp;
<figureCitation id="1338A7CABB44F17BFD06778AFD43F829" box="[729,745,1996,2022]" captionStart="FIGURE 7" captionStartId="10.[151,250,1944,1968]" captionTargetBox="[289,1298,181,1920]" captionTargetId="figure-22@10.[289,1298,181,1920]" captionTargetPageId="10" captionText="FIGURE 7. Strongylosomides troglobius sp. nov., male paratype. A-D. Right gonopod with prefemoral process (labeled PfP) highlighted and outlined in red, submesal, mesal, ventral and dorsal views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669207" httpUri="https://zenodo.org/record/6669207/files/figure.png" pageId="5" pageNumber="92">7</figureCitation>
), the gonocoxae (
<emphasis id="B977675DBB44F17BFC6D778DFC7EF82A" bold="true" box="[946,980,1995,2021]" pageId="5" pageNumber="92">Cx</emphasis>
) being devoid of distodorsal apophyses, the acropodites (
<emphasis id="B977675DBB47F178FE8B70D0FEC2FF7F" bold="true" box="[340,360,150,176]" pageId="6" pageNumber="93">A</emphasis>
) nearly identical, and the prefemoral processes (
<emphasis id="B977675DBB47F178FC4F70D0FC16FF7F" bold="true" box="[912,956,150,176]" pageId="6" pageNumber="93">PfP</emphasis>
) long, sinuous and attenuating distad. The only sharp difference concerns the
<emphasis id="B977675DBB47F178FDFE70FCFDE7FF1B" bold="true" box="[545,589,186,212]" pageId="6" pageNumber="93">PfP</emphasis>
of
<taxonomicName id="4C03C0CCBB47F178FDAD70FDFCABFF1A" authority="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette, 2022" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" box="[626,769,187,213]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="B977675DBB47F178FDAD70FDFCABFF1A" box="[626,769,187,213]" italics="true" pageId="6" pageNumber="93">S. troglobius</emphasis>
</taxonomicName>
<emphasis id="B977675DBB47F178FCD770FDFCC8FF1A" bold="true" box="[776,866,187,213]" pageId="6" pageNumber="93">
<taxonomicNameLabel id="A244DA26BB47F178FCD770FDFCC8FF1A" box="[776,866,187,213]" pageId="6" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which is fused by its basal half to
<emphasis id="B977675DBB47F178FB3070FCFAA9FF1B" bold="true" box="[1263,1283,186,212]" pageId="6" pageNumber="93">A</emphasis>
while its distal part is directed caudad, gradually attenuating and strongly sharpened at the end (
<figureCitation id="1338A7CABB47F178FB907099FB3DFF36" box="[1103,1175,223,249]" captionStart="FIGURE 6" captionStartId="9.[151,250,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-17@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 6. Strongylosomides troglobius sp. nov., male paratype. A-F. Right gonopod, submesal, mesal, ventral, dorsal, sublateral and lateral views, respectively. Abbreviations: A, acropodite; AP, acropodital process; c, cannula; Cx, gonocoxa; Pf, prefemoral region; PfP, prefemoral process; Sg, seminal groove; sl, solenomere; Te, telopodite. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669205" httpUri="https://zenodo.org/record/6669205/files/figure.png" pageId="6" pageNumber="93">Figs 6</figureCitation>
&amp;
<figureCitation id="1338A7CABB47F178FB627099FB67FF36" box="[1213,1229,223,249]" captionStart="FIGURE 7" captionStartId="10.[151,250,1944,1968]" captionTargetBox="[289,1298,181,1920]" captionTargetId="figure-22@10.[289,1298,181,1920]" captionTargetPageId="10" captionText="FIGURE 7. Strongylosomides troglobius sp. nov., male paratype. A-D. Right gonopod with prefemoral process (labeled PfP) highlighted and outlined in red, submesal, mesal, ventral and dorsal views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669207" httpUri="https://zenodo.org/record/6669207/files/figure.png" pageId="6" pageNumber="93">7</figureCitation>
),
<emphasis id="B977675DBB47F178FB3C7099FAA8FF37" box="[1251,1282,223,248]" italics="true" pageId="6" pageNumber="93">vs.</emphasis>
the
<emphasis id="B977675DBB47F178FAEB7098FACAFF37" bold="true" box="[1332,1376,222,248]" pageId="6" pageNumber="93">PfP</emphasis>
of
<taxonomicName id="4C03C0CCBB47F178FA587099FE8DFED3" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" baseAuthorityName="Schubart" baseAuthorityYear="1956" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="stercoriarius" status="comb. nov.">
<emphasis id="B977675DBB47F178FA587099FE8DFED3" italics="true" pageId="6" pageNumber="93">S. stercoriarius</emphasis>
</taxonomicName>
,
<emphasis id="B977675DBB47F178FEEB7145FE11FED2" bold="true" box="[308,443,259,285]" pageId="6" pageNumber="93">
<taxonomicNameLabel id="A244DA26BB47F178FEEB7145FE1FFED2" box="[308,437,259,285]" pageId="6" pageNumber="93" rank="species">comb. nov.</taxonomicNameLabel>
,
</emphasis>
being completely free from
<emphasis id="B977675DBB47F178FD287144FCA1FED3" bold="true" box="[759,779,258,284]" pageId="6" pageNumber="93">A</emphasis>
(as is typical of the genus,
<figureCitation id="1338A7CABB47F178FBE67145FB24FED2" box="[1081,1166,259,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1133,1157]" captionTargetBox="[160,1427,186,1102]" captionTargetId="figure-17@5.[151,1436,181,1109]" captionTargetPageId="5" captionText="FIGURE 3. Strongylosomides petropolites (Attems, 1901), male lectotype. A. Left gonopodal telopodite, mesal view (after Hoffman 1979), and Strongylosomides stercoriarius (Schubart, 1956), comb. nov., male syntype. B &amp; C. Right gonopod and its tip, mesal and sublateral views, respectively (after Schubart 1956). Abbreviations: A, acropodite; AP, acropodital process; c, cannula; Cx, gonocoxa; Pf, prefemoral region; PfP, prefemoral process; Sg, seminal groove; sl, solenomere; T, telopodite. Scale bar: 0.1 mm (B) or drawn not to scale (A, C)." figureDoi="http://doi.org/10.5281/zenodo.6669188" httpUri="https://zenodo.org/record/6669188/files/figure.png" pageId="6" pageNumber="93">Fig. 3B</figureCitation>
), ribbon-shaped, clearly sinuous and not so sharp at the end (
<figureCitation id="1338A7CABB47F178FDF27161FD6AFE8E" box="[557,704,295,321]" captionStart="FIGURE 3" captionStartId="5.[151,250,1133,1157]" captionTargetBox="[160,1427,186,1102]" captionTargetId="figure-17@5.[151,1436,181,1109]" captionTargetPageId="5" captionText="FIGURE 3. Strongylosomides petropolites (Attems, 1901), male lectotype. A. Left gonopodal telopodite, mesal view (after Hoffman 1979), and Strongylosomides stercoriarius (Schubart, 1956), comb. nov., male syntype. B &amp; C. Right gonopod and its tip, mesal and sublateral views, respectively (after Schubart 1956). Abbreviations: A, acropodite; AP, acropodital process; c, cannula; Cx, gonocoxa; Pf, prefemoral region; PfP, prefemoral process; Sg, seminal groove; sl, solenomere; T, telopodite. Scale bar: 0.1 mm (B) or drawn not to scale (A, C)." figureDoi="http://doi.org/10.5281/zenodo.6669188" httpUri="https://zenodo.org/record/6669188/files/figure.png" pageId="6" pageNumber="93">Figs 3B &amp; C</figureCitation>
).
</paragraph>
<paragraph id="8BBCBB4FBB47F178FF18710DFD5EFE3A" blockId="6.[151,1437,150,2014]" pageId="6" pageNumber="93">
In other words, we formally propose the following new transfer:
<taxonomicName id="4C03C0CCBB47F178FC61710DFF71FE46" authority="(Schubart, 1956)" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" baseAuthorityName="Schubart" baseAuthorityYear="1956" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="stercoriarius" status="comb. nov.">
<emphasis id="B977675DBB47F178FC61710DFAB5FEAB" box="[958,1311,331,357]" italics="true" pageId="6" pageNumber="93">Strongylosomides stercoriarius</emphasis>
(
<bibRefCitation id="EF92C6BEBB47F178FAEF710DFF79FE46" author="Schubart, O." pageId="6" pageNumber="93" pagination="355 - 367" refId="ref10067" refString="Schubart, O. 1956. &quot; Leptodesmidae &quot; brasileiras III. Especies de Minas Gerais (Diplopoda, Proterospermophora). Revista Brasileira de Biologia, 16 (3), 355 - 367." type="journal article" year="1956">Schubart, 1956</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B977675DBB47F178FF377129FEC0FE46" bold="true" box="[232,362,367,393]" pageId="6" pageNumber="93">
<taxonomicNameLabel id="A244DA26BB47F178FF377129FEC0FE46" box="[232,362,367,393]" pageId="6" pageNumber="93" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
ex
<taxonomicName id="4C03C0CCBB47F178FE4A7129FCAAFE46" authority="de Saussure, 1859" authorityName="de Saussure" authorityYear="1859" box="[405,768,366,393]" class="Diplopoda" family="Chelodesmidae" genus="Leptodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B977675DBB47F178FE4A7129FD8DFE46" box="[405,551,367,393]" italics="true" pageId="6" pageNumber="93">Leptodesmus</emphasis>
de Saussure, 1859
</taxonomicName>
. Moreover, despite the
<emphasis id="B977675DBB47F178FBD37128FB92FE47" bold="true" box="[1036,1080,366,392]" pageId="6" pageNumber="93">PfP</emphasis>
basal half being conspicuously fused to
<emphasis id="B977675DBB47F178FF2571D4FEA4FE63" bold="true" box="[250,270,402,428]" pageId="6" pageNumber="93">A</emphasis>
in
<taxonomicName id="4C03C0CCBB47F178FEEA71D5FE6FFE62" authority="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette, 2022" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" box="[309,453,403,429]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="B977675DBB47F178FEEA71D5FE6FFE62" box="[309,453,403,429]" italics="true" pageId="6" pageNumber="93">S. troglobius</emphasis>
</taxonomicName>
<emphasis id="B977675DBB47F178FE1271D5FD82FE62" bold="true" box="[461,552,403,429]" pageId="6" pageNumber="93">
<taxonomicNameLabel id="A244DA26BB47F178FE1271D5FD82FE62" box="[461,552,403,429]" pageId="6" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, we are inclined to formally assign this species to
<taxonomicName id="4C03C0CCBB47F178FBB471D5FA9BFE62" authorityName="Attems" authorityYear="1901" box="[1131,1329,403,429]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B977675DBB47F178FBB471D5FA9BFE62" box="[1131,1329,403,429]" italics="true" pageId="6" pageNumber="93">Strongylosomides</emphasis>
</taxonomicName>
, together with and close to
<taxonomicName id="4C03C0CCBB47F178FE8371F1FDA2FE1F" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" baseAuthorityName="Schubart" baseAuthorityYear="1956" box="[348,520,439,465]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="stercoriarius" status="comb. nov.">
<emphasis id="B977675DBB47F178FE8371F1FDA2FE1F" box="[348,520,439,465]" italics="true" pageId="6" pageNumber="93">S. stercoriarius</emphasis>
</taxonomicName>
,
<emphasis id="B977675DBB47F178FDCA71F1FD3CFE1E" bold="true" box="[533,662,439,465]" pageId="6" pageNumber="93">
<taxonomicNameLabel id="A244DA26BB47F178FDCA71F1FD3CFE1E" box="[533,662,439,465]" pageId="6" pageNumber="93" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
As
<bibRefCitation id="EF92C6BEBB47F178FD1D71F1FCD2FE1E" author="Hoffman, R. L." box="[706,888,439,465]" pageId="6" pageNumber="93" pagination="19 - 26" refId="ref9353" refString="Hoffman, R. L. (1979) Chelodesmid studies. XV. The identity and status of the generic name Strongylosomides Attems. Myriapodologica, 1 (3), 19 - 26. [https: // www. semanticscholar. org / paper / Chelodesmid-studies. - XV. - The-identity-andstatus-of-Hoffman / 451 b 267 fe 325 ee 86010 a 44 e 6 ac 812903 d 842 e 8 cb]" type="journal article" year="1979">Hoffman (1979)</bibRefCitation>
predicted, we can reasonably expect further new species and records of
<taxonomicName id="4C03C0CCBB47F178FE49719DFDF6FE3A" authorityName="Attems" authorityYear="1901" box="[406,604,475,501]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B977675DBB47F178FE49719DFDF6FE3A" box="[406,604,475,501]" italics="true" pageId="6" pageNumber="93">Strongylosomides</emphasis>
</taxonomicName>
in the future.
</paragraph>
</subSubSection>
<subSubSection id="C319E8C4BB47F178FF1871B8FB62F8BD" pageId="6" pageNumber="93" type="description">
<paragraph id="8BBCBB4FBB47F178FF1871B8FAF2FDAE" blockId="6.[151,1437,150,2014]" pageId="6" pageNumber="93">
<emphasis id="B977675DBB47F178FF1871B8FEFBFDD7" bold="true" box="[199,337,510,536]" pageId="6" pageNumber="93">Description</emphasis>
. Length of adults ca
<quantity id="4CFB16AABB47F178FDE771B8FD1AFDD6" box="[568,688,510,537]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="6" pageNumber="93" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">15–16 mm</quantity>
(males) or
<quantity id="4CFB16AABB47F178FCF371B9FCD0FDD6" box="[812,890,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="93" unit="mm" value="22.0">22 mm</quantity>
(females), width of midbody pro- and metazonae 2.0 and
<quantity id="4CFB16AABB47F178FF317265FEE9FDF2" box="[238,323,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="93" unit="mm" value="2.3">2.3 mm</quantity>
(males), 3.0 and
<quantity id="4CFB16AABB47F178FDDF7265FDFFFDF2" box="[512,597,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="6" pageNumber="93" unit="mm" value="3.3">3.3 mm</quantity>
or 2.5 and
<quantity id="4CFB16AABB47F178FD107265FC8EFDF2" box="[719,804,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="93" unit="mm" value="2.8">2.8 mm</quantity>
(females), respectively. Coloration of both live and fixed specimens uniformly light reddish ochre; antennae and legs mostly a little lighter, yellow ochre (
<figureCitation id="1338A7CABB47F178FB0F7201FABCFDAE" box="[1232,1302,583,609]" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Figs 4</figureCitation>
&amp;
<figureCitation id="1338A7CABB47F178FAE57200FAE0FDAE" box="[1338,1354,582,609]" captionStart="FIGURE 5" captionStartId="8.[151,250,1914,1938]" captionTargetBox="[151,1436,921,1891]" captionTargetId="figure-390@8.[151,1436,921,1891]" captionTargetPageId="8" captionText="FIGURE 5. Strongylosomides troglobius sp. nov., male paratype. A &amp; B. Body ring 7 with intact gonopods, ventral and lateral views, respectively. C. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. D. Body ring 7 with left intact gonopod, caudal view. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669201" httpUri="https://zenodo.org/record/6669201/files/figure.png" pageId="6" pageNumber="93">5</figureCitation>
).
</paragraph>
<paragraph id="8BBCBB4FBB47F178FF18722DFDE9FC92" blockId="6.[151,1437,150,2014]" pageId="6" pageNumber="93">
Body subcylindrical, with 20 rings (18p+1a+T) (
<figureCitation id="1338A7CABB47F178FD02722DFC8AFD4A" box="[733,800,619,645]" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Fig. 4</figureCitation>
). Tegument mainly poorly shining, texture mostly smooth to very delicately striolate. Head densely pilose all over clypeolabral region, vertex almost bare, epicranial suture faint; isthmus between antennae ca 1.2 times broader than diameter of antennal socket (
<figureCitation id="1338A7CABB47F178FBB872F5FB15FD02" box="[1127,1215,691,717]" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Fig. 4C</figureCitation>
). Mouthparts usual, not modified. Antennae relatively short and slender, poorly clavate due to only a slightly enlarged antennomere 6,
<emphasis id="B977675DBB47F178FF4872BDFF76FCDB" box="[151,220,763,788]" italics="true" pageId="6" pageNumber="93">in situ</emphasis>
projecting past ring 3 dorsally (male) or 2 (female); in length, antennomeres 2 = 6&gt; 3 = 4 = 5&gt; 1&gt; 7 (
<figureCitation id="1338A7CABB47F178FAB072BDFF02FCF6" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Fig. 4</figureCitation>
). Genae regularly rounded laterally (
<figureCitation id="1338A7CABB47F178FD927359FD4EFCF6" box="[589,740,799,825]" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Figs 4B &amp; C</figureCitation>
), incisura lateralis small and round, located beneath antenna; gnathochilarium without peculiarities.
</paragraph>
<paragraph id="8BBCBB4FBB47F178FF187321FB22FAB6" blockId="6.[151,1437,150,2014]" pageId="6" pageNumber="93">
In width, head &lt;collum &lt;ring 2 &lt;3 &lt;4 &lt;5=15 (male, female), thereafter body gradually tapering towards telson (
<figureCitation id="1338A7CABB47F178FF3673CDFE84FC6A" box="[233,302,907,933]" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Fig. 4</figureCitation>
). Dorsum smooth, regularly and strongly convex, at most very faintly rugulose in places, a little more densely and clearly so below paraterga (
<figureCitation id="1338A7CABB47F178FD8273E9FD0EFC06" box="[605,676,943,969]" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Figs 4</figureCitation>
&amp;
<figureCitation id="1338A7CABB47F178FD1573E8FD70FC06" box="[714,730,942,969]" captionStart="FIGURE 5" captionStartId="8.[151,250,1914,1938]" captionTargetBox="[151,1436,921,1891]" captionTargetId="figure-390@8.[151,1436,921,1891]" captionTargetPageId="8" captionText="FIGURE 5. Strongylosomides troglobius sp. nov., male paratype. A &amp; B. Body ring 7 with intact gonopods, ventral and lateral views, respectively. C. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. D. Body ring 7 with left intact gonopod, caudal view. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669201" httpUri="https://zenodo.org/record/6669201/files/figure.png" pageId="6" pageNumber="93">5</figureCitation>
). Paraterga rather poorly developed, only a little smaller in females, set low, mostly at about midheight of metazonae (
<figureCitation id="1338A7CABB47F178FCD37395FCF8FC22" box="[780,850,979,1005]" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Figs 4</figureCitation>
&amp;
<figureCitation id="1338A7CABB47F178FCAB7394FC29FC22" box="[884,899,978,1005]" captionStart="FIGURE 5" captionStartId="8.[151,250,1914,1938]" captionTargetBox="[151,1436,921,1891]" captionTargetId="figure-390@8.[151,1436,921,1891]" captionTargetPageId="8" captionText="FIGURE 5. Strongylosomides troglobius sp. nov., male paratype. A &amp; B. Body ring 7 with intact gonopods, ventral and lateral views, respectively. C. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. D. Body ring 7 with left intact gonopod, caudal view. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669201" httpUri="https://zenodo.org/record/6669201/files/figure.png" pageId="6" pageNumber="93">5</figureCitation>
), clearly declivous, especially narrowly rimmed on collum, thereafter small, regularly rounded, laterally smooth and anteriorly increasingly incomplete ridges on poreless rings,
<emphasis id="B977675DBB47F178FEE0745DFEF4FBFB" box="[319,350,1051,1076]" italics="true" pageId="6" pageNumber="93">vs.</emphasis>
clearly thickened, also increasingly abbreviated, oblong porostele-like calluses caudally on porebearing rings (
<figureCitation id="1338A7CABB47F178FEE47479FE29FB96" box="[315,387,1087,1113]" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Figs 4</figureCitation>
&amp;
<figureCitation id="1338A7CABB47F178FE777478FE67FB96" box="[424,461,1086,1113]" captionStart="FIGURE 5" captionStartId="8.[151,250,1914,1938]" captionTargetBox="[151,1436,921,1891]" captionTargetId="figure-390@8.[151,1436,921,1891]" captionTargetPageId="8" captionText="FIGURE 5. Strongylosomides troglobius sp. nov., male paratype. A &amp; B. Body ring 7 with intact gonopods, ventral and lateral views, respectively. C. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. D. Body ring 7 with left intact gonopod, caudal view. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669201" httpUri="https://zenodo.org/record/6669201/files/figure.png" pageId="6" pageNumber="93">5D</figureCitation>
). Calluses on paraterga delimited by sulci both dorsally and, in posterior part of body, caudally. Caudal corners of paraterga acutangular and mostly rounded to narrowly rounded, but never drawn past rear tergal margin (
<figureCitation id="1338A7CABB47F178FEAE74C1FE1DFB6E" box="[369,439,1159,1185]" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Fig. 4</figureCitation>
). Pore formula normal (5, 7, 9, 10, 12, 13, 15–19); ozopores small, located inside ovoid pits a little before caudal corners of paraterga (
<figureCitation id="1338A7CABB47F178FD7474EDFD59FB0A" box="[683,755,1195,1221]" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Figs 4</figureCitation>
&amp;
<figureCitation id="1338A7CABB47F178FCC674ECFC94FB0A" box="[793,830,1194,1221]" captionStart="FIGURE 5" captionStartId="8.[151,250,1914,1938]" captionTargetBox="[151,1436,921,1891]" captionTargetId="figure-390@8.[151,1436,921,1891]" captionTargetPageId="8" captionText="FIGURE 5. Strongylosomides troglobius sp. nov., male paratype. A &amp; B. Body ring 7 with intact gonopods, ventral and lateral views, respectively. C. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. D. Body ring 7 with left intact gonopod, caudal view. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669201" httpUri="https://zenodo.org/record/6669201/files/figure.png" pageId="6" pageNumber="93">5D</figureCitation>
). Tergal setae absent, no setation pattern traceable on metaterga. A transverse sulcus or impression on metaterga absent. Stricture between pro- and metazonae distinct, but rather narrow, shallow and smooth. Limbus very thin and entirely smooth. Pleurosternal carinae increasingly low, arcuate, longitudinal ridges or bulges traceable from ring 2 until ring 18 (
<figureCitation id="1338A7CABB47F178FC297551FBC5FAFE" box="[1014,1135,1303,1329]" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Figs 4B, C</figureCitation>
&amp;
<figureCitation id="1338A7CABB47F178FB4E7550FB0BFAFE" box="[1169,1185,1302,1329]" captionStart="FIGURE 5" captionStartId="8.[151,250,1914,1938]" captionTargetBox="[151,1436,921,1891]" captionTargetId="figure-390@8.[151,1436,921,1891]" captionTargetPageId="8" captionText="FIGURE 5. Strongylosomides troglobius sp. nov., male paratype. A &amp; B. Body ring 7 with intact gonopods, ventral and lateral views, respectively. C. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. D. Body ring 7 with left intact gonopod, caudal view. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669201" httpUri="https://zenodo.org/record/6669201/files/figure.png" pageId="6" pageNumber="93">5</figureCitation>
). Epiproct rather long, conical, straight, tip subtruncate; pre-apical papillae small (
<figureCitation id="1338A7CABB47F178FCEB757DFCD0FA9A" box="[820,890,1339,1365]" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Fig. 4</figureCitation>
). Hypoproct roundly subtriangular, sharpened at tip; setigerous papillae at caudal corners small and well separated; sides convex (
<figureCitation id="1338A7CABB47F178FBFD7519FBD6FAB6" box="[1058,1148,1375,1401]" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Fig. 4C</figureCitation>
).
</paragraph>
<paragraph id="8BBCBB4FBB47F178FF1875C5FF62F99E" blockId="6.[151,1437,150,2014]" pageId="6" pageNumber="93">
Sterna broad, without notable modifications even before ring 7, nearly glabrous, cross-impressions weak (
<figureCitation id="1338A7CABB47F178FAB075C5FF16FA0E" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Fig. 4C</figureCitation>
). Epigynal ridge very low. Legs relatively short and slender (
<figureCitation id="1338A7CABB47F178FC8175E1FC0CFA0E" box="[862,934,1447,1473]" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Figs 4</figureCitation>
,
<figureCitation id="1338A7CABB47F178FC6E75E0FBA1FA0E" box="[945,1035,1446,1473]" captionStart="FIGURE 5" captionStartId="8.[151,250,1914,1938]" captionTargetBox="[151,1436,921,1891]" captionTargetId="figure-390@8.[151,1436,921,1891]" captionTargetPageId="8" captionText="FIGURE 5. Strongylosomides troglobius sp. nov., male paratype. A &amp; B. Body ring 7 with intact gonopods, ventral and lateral views, respectively. C. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. D. Body ring 7 with left intact gonopod, caudal view. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669201" httpUri="https://zenodo.org/record/6669201/files/figure.png" pageId="6" pageNumber="93">5C &amp; D</figureCitation>
), especially so in female, ca 1.3–1.4 (male) or 0.9–1.0 times (female) as long as midbody height; male legs with chelae (distoventral protrusions on tibiae) gradually disappearing towards ring 14; in length, femur &gt;&gt; prefemur = tarsus&gt; tibia&gt; coxa = postfemur; claw very short (
<figureCitation id="1338A7CABB47F178FE8F7655FE32F9E2" box="[336,408,1555,1581]" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Figs 4</figureCitation>
,
<figureCitation id="1338A7CABB47F178FE7C7654FE54F9E2" box="[419,510,1554,1581]" captionStart="FIGURE 5" captionStartId="8.[151,250,1914,1938]" captionTargetBox="[151,1436,921,1891]" captionTargetId="figure-390@8.[151,1436,921,1891]" captionTargetPageId="8" captionText="FIGURE 5. Strongylosomides troglobius sp. nov., male paratype. A &amp; B. Body ring 7 with intact gonopods, ventral and lateral views, respectively. C. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. D. Body ring 7 with left intact gonopod, caudal view. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669201" httpUri="https://zenodo.org/record/6669201/files/figure.png" pageId="6" pageNumber="93">5C &amp; D</figureCitation>
). Male gonopores located on distinct, distomesal, subtriangular gonapophyses (
<figureCitation id="1338A7CABB47F178FAB07655FF16F99E" captionStart="FIGURE 4" captionStartId="7.[151,250,1808,1832]" captionTargetBox="[151,1435,266,1797]" captionTargetId="figure-145@7.[151,1435,266,1797]" captionTargetPageId="7" captionText="FIGURE 4. Strongylosomides troglobius sp. nov., male paratype. A-C. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="6" pageNumber="93">Fig. 4C</figureCitation>
).
</paragraph>
<paragraph id="8BBCBB4FBB47F178FF18761DFB65F881" blockId="6.[151,1437,150,2014]" pageId="6" pageNumber="93">
Gonopodal aperture ovoid, simple, only a little broader than following sterna (
<figureCitation id="1338A7CABB47F178FBF1761DFB2CF9BA" box="[1070,1158,1626,1653]" captionStart="FIGURE 5" captionStartId="8.[151,250,1914,1938]" captionTargetBox="[151,1436,921,1891]" captionTargetId="figure-390@8.[151,1436,921,1891]" captionTargetPageId="8" captionText="FIGURE 5. Strongylosomides troglobius sp. nov., male paratype. A &amp; B. Body ring 7 with intact gonopods, ventral and lateral views, respectively. C. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. D. Body ring 7 with left intact gonopod, caudal view. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669201" httpUri="https://zenodo.org/record/6669201/files/figure.png" pageId="6" pageNumber="93">Fig. 5C</figureCitation>
). Gonopods (
<figureCitation id="1338A7CABB47F178FAC4761DFF04F956" captionStart="FIGURE 5" captionStartId="8.[151,250,1914,1938]" captionTargetBox="[151,1436,921,1891]" captionTargetId="figure-390@8.[151,1436,921,1891]" captionTargetPageId="8" captionText="FIGURE 5. Strongylosomides troglobius sp. nov., male paratype. A &amp; B. Body ring 7 with intact gonopods, ventral and lateral views, respectively. C. Body ring 7 with gonopods removed to show gonopod aperture, ventral view. D. Body ring 7 with left intact gonopod, caudal view. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669201" httpUri="https://zenodo.org/record/6669201/files/figure.png" pageId="6" pageNumber="93">Figs 5A, B, D</figureCitation>
,
<figureCitation id="1338A7CABB47F178FF657639FF62F956" box="[186,200,1663,1689]" captionStart="FIGURE 6" captionStartId="9.[151,250,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-17@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 6. Strongylosomides troglobius sp. nov., male paratype. A-F. Right gonopod, submesal, mesal, ventral, dorsal, sublateral and lateral views, respectively. Abbreviations: A, acropodite; AP, acropodital process; c, cannula; Cx, gonocoxa; Pf, prefemoral region; PfP, prefemoral process; Sg, seminal groove; sl, solenomere; Te, telopodite. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669205" httpUri="https://zenodo.org/record/6669205/files/figure.png" pageId="6" pageNumber="93">6</figureCitation>
&amp;
<figureCitation id="1338A7CABB47F178FF337639FF51F956" box="[236,251,1663,1689]" captionStart="FIGURE 7" captionStartId="10.[151,250,1944,1968]" captionTargetBox="[289,1298,181,1920]" captionTargetId="figure-22@10.[289,1298,181,1920]" captionTargetPageId="10" captionText="FIGURE 7. Strongylosomides troglobius sp. nov., male paratype. A-D. Right gonopod with prefemoral process (labeled PfP) highlighted and outlined in red, submesal, mesal, ventral and dorsal views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669207" httpUri="https://zenodo.org/record/6669207/files/figure.png" pageId="6" pageNumber="93">7</figureCitation>
) stout and robust, devoid of sternal rudiments; gonocoxa (
<emphasis id="B977675DBB47F178FC537638FC04F957" bold="true" box="[908,942,1662,1688]" pageId="6" pageNumber="93">Cx</emphasis>
) devoid of a distodorsal apophysis, cannula (
<emphasis id="B977675DBB47F178FF7E76E5FF07F972" bold="true" box="[161,173,1699,1725]" pageId="6" pageNumber="93">c</emphasis>
) as usual, a thin curved tube, devoid of setae mesally at its base; prefemoral region (
<emphasis id="B977675DBB47F178FB8876E4FBDBF973" bold="true" box="[1111,1137,1698,1724]" pageId="6" pageNumber="93">Pf</emphasis>
) as usual, short, about half as long as an untwisted acropodite (
<emphasis id="B977675DBB47F178FDFC7680FD9DF92F" bold="true" box="[547,567,1734,1760]" pageId="6" pageNumber="93">A</emphasis>
); caudal prefemoral process (
<emphasis id="B977675DBB47F178FC5D7680FC04F92F" bold="true" box="[898,942,1734,1760]" pageId="6" pageNumber="93">PfP</emphasis>
) long, gradually attenuating distad, fused by its basal half to
<emphasis id="B977675DBB47F178FE9C76ACFEFDF8CB" bold="true" box="[323,343,1770,1796]" pageId="6" pageNumber="93">A</emphasis>
, its distal half directed caudad, attenuating and strongly sharpened apically (
<figureCitation id="1338A7CABB47F178FB7D76ADFB4DF8CA" box="[1186,1255,1771,1797]" captionStart="FIGURE 6" captionStartId="9.[151,250,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-17@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 6. Strongylosomides troglobius sp. nov., male paratype. A-F. Right gonopod, submesal, mesal, ventral, dorsal, sublateral and lateral views, respectively. Abbreviations: A, acropodite; AP, acropodital process; c, cannula; Cx, gonocoxa; Pf, prefemoral region; PfP, prefemoral process; Sg, seminal groove; sl, solenomere; Te, telopodite. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669205" httpUri="https://zenodo.org/record/6669205/files/figure.png" pageId="6" pageNumber="93">Figs 6</figureCitation>
&amp;
<figureCitation id="1338A7CABB47F178FAD776ADFAB2F8CA" box="[1288,1304,1771,1797]" captionStart="FIGURE 7" captionStartId="10.[151,250,1944,1968]" captionTargetBox="[289,1298,181,1920]" captionTargetId="figure-22@10.[289,1298,181,1920]" captionTargetPageId="10" captionText="FIGURE 7. Strongylosomides troglobius sp. nov., male paratype. A-D. Right gonopod with prefemoral process (labeled PfP) highlighted and outlined in red, submesal, mesal, ventral and dorsal views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669207" httpUri="https://zenodo.org/record/6669207/files/figure.png" pageId="6" pageNumber="93">7</figureCitation>
);
<emphasis id="B977675DBB47F178FAF276ACFAEBF8CB" bold="true" box="[1325,1345,1770,1796]" pageId="6" pageNumber="93">A</emphasis>
laminar and membranous, suberect; seminal groove (
<emphasis id="B977675DBB47F178FD4D7748FD1AF8E7" bold="true" box="[658,688,1806,1832]" pageId="6" pageNumber="93">Sg</emphasis>
) running entirely on mesal side of telopodite (
<emphasis id="B977675DBB47F178FB607748FB4CF8E7" bold="true" box="[1215,1254,1806,1832]" pageId="6" pageNumber="93">Te)</emphasis>
, ending up on a short solenomere (
<emphasis id="B977675DBB47F178FEB77775FED1F882" bold="true" box="[360,379,1843,1869]" pageId="6" pageNumber="93">sl</emphasis>
) opposite a similarly short end branch, or acropodital process (
<emphasis id="B977675DBB47F178FBE07774FBCEF883" bold="true" box="[1087,1124,1842,1868]" pageId="6" pageNumber="93">AP</emphasis>
) (
<figureCitation id="1338A7CABB47F178FBA27775FB6BF881" box="[1149,1217,1843,1870]" captionStart="FIGURE 6" captionStartId="9.[151,250,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-17@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 6. Strongylosomides troglobius sp. nov., male paratype. A-F. Right gonopod, submesal, mesal, ventral, dorsal, sublateral and lateral views, respectively. Abbreviations: A, acropodite; AP, acropodital process; c, cannula; Cx, gonocoxa; Pf, prefemoral region; PfP, prefemoral process; Sg, seminal groove; sl, solenomere; Te, telopodite. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669205" httpUri="https://zenodo.org/record/6669205/files/figure.png" pageId="6" pageNumber="93">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8BBCBB4FBB47F178FF187711FB62F8BD" blockId="6.[151,1437,150,2014]" box="[199,1224,1879,1906]" pageId="6" pageNumber="93">
Vulvae not studied:
<specimenCount id="9D0570C6BB47F178FE777711FD89F8BE" box="[424,547,1879,1905]" pageId="6" pageNumber="93" type="female">one female</specimenCount>
<typeStatus id="54B805EDBB47F178FDF57711FD20F8BE" box="[554,650,1879,1905]" pageId="6" pageNumber="93" type="paratype">paratype</typeStatus>
dissected, but no vulvae found (?underdeveloped).
</paragraph>
</subSubSection>
<subSubSection id="C319E8C4BB47F176FF18773DFE43FC4E" lastPageId="8" lastPageNumber="95" pageId="6" pageNumber="93" type="discussion">
<paragraph id="8BBCBB4FBB47F179FF18773DFD10FF36" blockId="6.[151,1437,150,2014]" lastBlockId="7.[151,1437,150,249]" lastPageId="7" lastPageNumber="94" pageId="6" pageNumber="93">
<emphasis id="B977675DBB47F178FF18773DFE9EF85A" bold="true" box="[199,308,1915,1941]" pageId="6" pageNumber="93">Remarks</emphasis>
. Superficially, the gonopodal telopodite of
<taxonomicName id="4C03C0CCBB47F178FCCB773DFC0AF85A" authority="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette, 2022" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" box="[788,928,1915,1941]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="B977675DBB47F178FCCB773DFC0AF85A" box="[788,928,1915,1941]" italics="true" pageId="6" pageNumber="93">S. troglobius</emphasis>
</taxonomicName>
<emphasis id="B977675DBB47F178FC79773DFC55F85A" bold="true" box="[934,1023,1915,1941]" pageId="6" pageNumber="93">
<taxonomicNameLabel id="A244DA26BB47F178FC79773DFC55F85A" box="[934,1023,1915,1941]" pageId="6" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
looks uniramous at the base, because the basal half of the prefemoral process (
<emphasis id="B977675DBB47F178FDBF77D8FD26F877" bold="true" box="[608,652,1950,1976]" pageId="6" pageNumber="93">PfP</emphasis>
) is tightly appressed to and coalesced with the acropodite (
<emphasis id="B977675DBB47F178FAFD77D8FA9CF877" bold="true" box="[1314,1334,1950,1976]" pageId="6" pageNumber="93">A</emphasis>
), the latter branching off at about midway and independent only distally (
<figureCitation id="1338A7CABB47F178FCAE7785FC1FF811" box="[881,949,1987,2014]" captionStart="FIGURE 6" captionStartId="9.[151,250,1840,1864]" captionTargetBox="[151,1436,181,1816]" captionTargetId="figure-17@9.[151,1436,181,1816]" captionTargetPageId="9" captionText="FIGURE 6. Strongylosomides troglobius sp. nov., male paratype. A-F. Right gonopod, submesal, mesal, ventral, dorsal, sublateral and lateral views, respectively. Abbreviations: A, acropodite; AP, acropodital process; c, cannula; Cx, gonocoxa; Pf, prefemoral region; PfP, prefemoral process; Sg, seminal groove; sl, solenomere; Te, telopodite. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669205" httpUri="https://zenodo.org/record/6669205/files/figure.png" pageId="6" pageNumber="93">Fig. 6</figureCitation>
). However, a closer examination of the
<emphasis id="B977675DBB47F178FAAE7784FA37F813" bold="true" box="[1393,1437,1986,2012]" pageId="6" pageNumber="93">PfP</emphasis>
shows clear traces (
<figureCitation id="1338A7CABB46F179FEAD70D1FE1CFF7E" box="[370,438,151,177]" captionStart="FIGURE 7" captionStartId="10.[151,250,1944,1968]" captionTargetBox="[289,1298,181,1920]" captionTargetId="figure-22@10.[289,1298,181,1920]" captionTargetPageId="10" captionText="FIGURE 7. Strongylosomides troglobius sp. nov., male paratype. A-D. Right gonopod with prefemoral process (labeled PfP) highlighted and outlined in red, submesal, mesal, ventral and dorsal views, respectively. Photographs by K.V. Makarov, taken not to scale." figureDoi="http://doi.org/10.5281/zenodo.6669207" httpUri="https://zenodo.org/record/6669207/files/figure.png" pageId="7" pageNumber="94">Fig. 7</figureCitation>
, outlined in red) of the fusion of the
<emphasis id="B977675DBB46F179FC8F70D0FCD6FF7F" bold="true" box="[848,892,150,176]" pageId="7" pageNumber="94">PfP</emphasis>
base to the acropodite base in all meaningful aspects. This betrays the basic bipartite condition typical of the
<taxonomicName id="4C03C0CCBB46F179FC9C70FDFC5AFF1A" authorityName="Cook" authorityYear="1895" box="[835,1008,187,213]" class="Diplopoda" family="Chelodesmidae" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="87" phylum="Arthropoda" rank="subFamily" subFamily="Chelodesminae">Chelodesminae</taxonomicName>
gonopod, including
<taxonomicName id="4C03C0CCBB46F179FB0970FDFA36FF1A" authorityName="Attems" authorityYear="1901" box="[1238,1436,187,213]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B977675DBB46F179FB0970FDFA36FF1A" box="[1238,1436,187,213]" italics="true" pageId="7" pageNumber="94">Strongylosomides</emphasis>
</taxonomicName>
as a genus that remains unassigned to a tribe yet.
</paragraph>
<caption id="DF7CEBC7BB46F179FF487756FD24F883" ID-DOI="http://doi.org/10.5281/zenodo.6669197" ID-Zenodo-Dep="6669197" httpUri="https://zenodo.org/record/6669197/files/figure.png" pageId="7" pageNumber="94" startId="7.[151,250,1808,1832]" targetBox="[151,1435,266,1797]" targetPageId="7">
<paragraph id="8BBCBB4FBB46F179FF487756FD24F883" blockId="7.[151,1437,1808,1868]" pageId="7" pageNumber="94">
<emphasis id="B977675DBB46F179FF487756FEA7F8E7" bold="true" box="[151,269,1808,1832]" pageId="7" pageNumber="94">FIGURE 4</emphasis>
.
<taxonomicName id="4C03C0CCBB46F179FEC67756FD90F8E7" authority="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette, 2022" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" box="[281,570,1808,1832]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="B977675DBB46F179FEC67756FD90F8E7" box="[281,570,1808,1832]" italics="true" pageId="7" pageNumber="94">Strongylosomides troglobius</emphasis>
</taxonomicName>
<emphasis id="B977675DBB46F179FD9F7757FD3BF8E6" bold="true" box="[576,657,1809,1833]" pageId="7" pageNumber="94">
<taxonomicNameLabel id="A244DA26BB46F179FD9F7757FD3BF8E6" box="[576,657,1809,1833]" pageId="7" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype.
<emphasis id="B977675DBB46F179FCE67756FCCCF8E7" bold="true" box="[825,870,1808,1832]" pageId="7" pageNumber="94">A-C</emphasis>
. Habitus, dorsal, lateral and ventral views, respectively. Photographs by K.V. Makarov, taken not to scale.
</paragraph>
</caption>
<paragraph id="8BBCBB4FBB46F176FF18773DFB27FED2" blockId="7.[151,1437,1914,2013]" lastBlockId="8.[151,1436,151,897]" lastPageId="8" lastPageNumber="95" pageId="7" pageNumber="94">
The absence of a
<emphasis id="B977675DBB46F179FE4A773CFE6BF85B" bold="true" box="[405,449,1914,1940]" pageId="7" pageNumber="94">PfP</emphasis>
is characteristic of the
<taxonomicName id="4C03C0CCBB46F179FD0D773DFCD7F85A" authorityName="Cook" authorityYear="1896" box="[722,893,1915,1941]" class="Diplopoda" family="Chelodesmidae" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="87" phylum="Arthropoda" rank="subFamily" subFamily="Prepodesminae">Prepodesminae</taxonomicName>
(
<bibRefCitation id="EF92C6BEBB46F179FC52773DFB3EF85A" author="Hoffman, R. L. &amp; Reid, J. C." box="[909,1172,1915,1941]" pageId="7" pageNumber="94" pagination="29 - 36" refId="ref9704" refString="Hoffman, R. L. &amp; Reid, J. C. (1990) A new species, genus, and tribe of death-feigning millipeds from Nigeria (Polydesmida: Chelodesmidae). Journal of African Zoology, 104, 29 - 36. [https: // www. semanticscholar. org / paper / A-new-species % 2 Cgenus % 2 C-and-tribe-of-death-feigning-Hoffman-Reid / 4 e 63608 d 3 ab 555 d 914 bb 08 a 1 b 2 f 4032 fb 8 d 6 cddb]" type="journal article" year="1990">Hoffman &amp; Reid 1990</bibRefCitation>
) and several genera of
<taxonomicName id="4C03C0CCBB46F179FF4877D9FEEDF876" authorityName="Cook" authorityYear="1895" box="[151,327,1951,1977]" class="Diplopoda" family="Chelodesmidae" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="87" phylum="Arthropoda" rank="subFamily" subFamily="Chelodesminae">Chelodesminae</taxonomicName>
, both with a definite tribal position (e.g., Telonychopodini) and still not classified in accepted tribes. Among the latter genera, such are, e.g.,
<taxonomicName id="4C03C0CCBB46F179FD867785FC4EF812" authority="Hoffman, 2009" authorityName="Hoffman" authorityYear="2009" box="[601,996,1987,2013]" class="Diplopoda" family="Chelodesmidae" genus="Cryptosolenomeris" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B977675DBB46F179FD867785FC84F812" box="[601,814,1987,2013]" italics="true" pageId="7" pageNumber="94">Cryptosolenomeris</emphasis>
<bibRefCitation id="EF92C6BEBB46F179FCE97785FC4EF812" author="Hoffman, R. L." box="[822,996,1987,2013]" pageId="7" pageNumber="94" pagination="557 - 562" refId="ref9614" refString="Hoffman, R. L. (2009) A new, disjunct, diplopod genus from Espirito Santo, Brasil (Polydesmida: Chelodesmidae). Papeis Avulsos de Zoologia, 49 (42), 557 - 562. [https: // www. researchgate. net / publication / 250984079 _ A _ new _ disjunct _ diplopod _ genus _ from _ Espirito _ Santo _ Brasil _ Polydesmida _ Chelodesmidae] https: // doi. org / 10.1590 / S 0031 - 10492009004200001" type="journal article" year="2009">Hoffman, 2009</bibRefCitation>
</taxonomicName>
, monobasic, with one species from
<collectingRegion id="49C775ADBB46F176FAA87785FEF2FF7E" country="Brazil" lastPageId="8" lastPageNumber="95" name="Espirito Santo" pageId="7" pageNumber="94">Espírito Santo state</collectingRegion>
,
<collectingCountry id="F314FBDFBB49F176FEBB70D1FE02FF7E" box="[356,424,151,177]" name="Brazil" pageId="8" pageNumber="95">Brazil</collectingCountry>
(
<bibRefCitation id="EF92C6BEBB49F176FE6870D1FDCBFF7E" author="Hoffman, R. L." box="[439,609,151,177]" pageId="8" pageNumber="95" pagination="557 - 562" refId="ref9614" refString="Hoffman, R. L. (2009) A new, disjunct, diplopod genus from Espirito Santo, Brasil (Polydesmida: Chelodesmidae). Papeis Avulsos de Zoologia, 49 (42), 557 - 562. [https: // www. researchgate. net / publication / 250984079 _ A _ new _ disjunct _ diplopod _ genus _ from _ Espirito _ Santo _ Brasil _ Polydesmida _ Chelodesmidae] https: // doi. org / 10.1590 / S 0031 - 10492009004200001" type="journal article" year="2009">Hoffman 2009</bibRefCitation>
), or
<taxonomicName id="4C03C0CCBB49F176FD4C70D1FC6CFF7E" authority="Pocock, 1894" authorityName="Pocock" authorityYear="1894" box="[659,966,151,177]" class="Diplopoda" family="Chelodesmidae" genus="Odontopeltis" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="B977675DBB49F176FD4C70D1FC8EFF7E" box="[659,804,151,177]" italics="true" pageId="8" pageNumber="95">Odontopeltis</emphasis>
Pocock, 1894
</taxonomicName>
, whose eight presently known species, all in southeastern
<collectingCountry id="F314FBDFBB49F176FE9A70FDFE23FF1A" box="[325,393,187,213]" name="Brazil" pageId="8" pageNumber="95">Brazil</collectingCountry>
(
<collectingRegion id="49C775ADBB49F176FE4870FDFD90FF1A" box="[407,570,187,213]" country="Brazil" name="Espirito Santo" pageId="8" pageNumber="95">Espírito Santo</collectingRegion>
,
<collectingRegion id="49C775ADBB49F176FD9B70FDFD73FF1A" box="[580,729,187,213]" country="Brazil" name="Minas Gerais" pageId="8" pageNumber="95">Minas Gerais</collectingRegion>
and
<collectingRegion id="49C775ADBB49F176FCD270FDFC05FF19" box="[781,943,187,214]" country="Brazil" name="Rio de Janeiro" pageId="8" pageNumber="95">Rio de Janeiro</collectingRegion>
) (
<bibRefCitation id="EF92C6BEBB49F176FC1C70FDFBC1FF1A" author="Hoffman, R. L." box="[963,1131,187,213]" pageId="8" pageNumber="95" pagination="55 - 62" refId="ref9454" refString="Hoffman, R. L. (1981) Chelodesmid studies. XVII. Revalidation of the generic name Odontopeltis Pocock 1894, with the description of a new species and notes on other members of the group. Myriapodologica, 1 (9), 55 - 62. [https: // www. wikidata. org / wiki / Q 109377335]" type="journal article" year="1981">Hoffman 1981</bibRefCitation>
; Pena-Barbosa
<emphasis id="B977675DBB49F176FAC870FDFAE4FF1A" box="[1303,1358,187,213]" italics="true" pageId="8" pageNumber="95">et al.</emphasis>
2013), are among the largest in the family (
<quantity id="4CFB16AABB49F176FDF27099FD1FFF36" box="[557,693,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="9.35" metricValueMax="10.7" metricValueMin="8.0" pageId="8" pageNumber="95" unit="mm" value="93.5" valueMax="107.0" valueMin="80.0">80–107 mm</quantity>
long). However, none of them show a structure resembling a
<emphasis id="B977675DBB49F176FAB57098FA3CFF37" bold="true" box="[1386,1430,222,248]" pageId="8" pageNumber="95">PfP</emphasis>
, even such that is fused with discernible traces to the base of the gonopodal acropodite (
<emphasis id="B977675DBB49F176FBB67144FBD7FED3" bold="true" box="[1129,1149,258,284]" pageId="8" pageNumber="95">A</emphasis>
).
</paragraph>
<paragraph id="8BBCBB4FBB49F176FF187161FC66FDAE" blockId="8.[151,1436,151,897]" pageId="8" pageNumber="95">
Because both
<taxonomicName id="4C03C0CCBB49F176FEB87161FE5CFE8E" authority="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette, 2022" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" box="[359,502,295,321]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="B977675DBB49F176FEB87161FE5CFE8E" box="[359,502,295,321]" italics="true" pageId="8" pageNumber="95">S. troglobius</emphasis>
</taxonomicName>
<emphasis id="B977675DBB49F176FE217161FDF3FE8E" bold="true" box="[510,601,295,321]" pageId="8" pageNumber="95">
<taxonomicNameLabel id="A244DA26BB49F176FE217161FDF3FE8E" box="[510,601,295,321]" pageId="8" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C03C0CCBB49F176FD4E7161FC95FE8F" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" baseAuthorityName="Schubart" baseAuthorityYear="1956" box="[657,831,295,321]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="stercoriarius" status="comb. nov.">
<emphasis id="B977675DBB49F176FD4E7161FC95FE8F" box="[657,831,295,321]" italics="true" pageId="8" pageNumber="95">S. stercoriarius</emphasis>
</taxonomicName>
,
<emphasis id="B977675DBB49F176FC927161FC65FE8E" bold="true" box="[845,975,295,321]" pageId="8" pageNumber="95">
<taxonomicNameLabel id="A244DA26BB49F176FC927161FC65FE8E" box="[845,975,295,321]" pageId="8" pageNumber="95" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, differ from the remaining congeners by the rather pale coloration (
<emphasis id="B977675DBB49F176FE1F710DFE75FEAB" box="[448,479,331,356]" italics="true" pageId="8" pageNumber="95">vs.</emphasis>
dark), the unmodified pregonopodal sternites 4, 5 and 6 (
<emphasis id="B977675DBB49F176FBBB710DFB29FEAB" box="[1124,1155,331,356]" italics="true" pageId="8" pageNumber="95">vs.</emphasis>
with paramedian cones), the gonocoxae devoid of distodorsal apophyses (
<emphasis id="B977675DBB49F176FD667129FD72FE47" box="[697,728,367,392]" italics="true" pageId="8" pageNumber="95">vs.</emphasis>
present), the basal course of the seminal groove running closer to the ventral margin of the acropodite (
<emphasis id="B977675DBB49F176FD8071D5FDD4FE63" box="[607,638,403,428]" italics="true" pageId="8" pageNumber="95">vs.</emphasis>
abruptly reflexed dorsad across the prefemoral region and proceeding to the solenomere along the dorsomedian edge of the acropodite), the prefemoral process attenuating, directed caudomesad and sharp at the end (
<emphasis id="B977675DBB49F176FE24719DFDB8FE3B" box="[507,530,475,500]" italics="true" pageId="8" pageNumber="95">vs</emphasis>
. broad and suberect), and the absence of a midway cingulum on the acropodite (
<emphasis id="B977675DBB49F176FF7E71B9FF15FDD7" box="[161,191,511,536]" italics="true" pageId="8" pageNumber="95">vs.</emphasis>
often present) (cf.
<figureCitation id="1338A7CABB49F176FE4B71B9FE42FDD6" box="[404,488,511,537]" captionStart="FIGURE 3" captionStartId="5.[151,250,1133,1157]" captionTargetBox="[160,1427,186,1102]" captionTargetId="figure-17@5.[151,1436,181,1109]" captionTargetPageId="5" captionText="FIGURE 3. Strongylosomides petropolites (Attems, 1901), male lectotype. A. Left gonopodal telopodite, mesal view (after Hoffman 1979), and Strongylosomides stercoriarius (Schubart, 1956), comb. nov., male syntype. B &amp; C. Right gonopod and its tip, mesal and sublateral views, respectively (after Schubart 1956). Abbreviations: A, acropodite; AP, acropodital process; c, cannula; Cx, gonocoxa; Pf, prefemoral region; PfP, prefemoral process; Sg, seminal groove; sl, solenomere; T, telopodite. Scale bar: 0.1 mm (B) or drawn not to scale (A, C)." figureDoi="http://doi.org/10.5281/zenodo.6669188" httpUri="https://zenodo.org/record/6669188/files/figure.png" pageId="8" pageNumber="95">Fig. 3A</figureCitation>
and
<figureCitation id="1338A7CABB49F176FDC171B9FDDEFDD6" box="[542,628,511,537]" captionStart="FIGURE 3" captionStartId="5.[151,250,1133,1157]" captionTargetBox="[160,1427,186,1102]" captionTargetId="figure-17@5.[151,1436,181,1109]" captionTargetPageId="5" captionText="FIGURE 3. Strongylosomides petropolites (Attems, 1901), male lectotype. A. Left gonopodal telopodite, mesal view (after Hoffman 1979), and Strongylosomides stercoriarius (Schubart, 1956), comb. nov., male syntype. B &amp; C. Right gonopod and its tip, mesal and sublateral views, respectively (after Schubart 1956). Abbreviations: A, acropodite; AP, acropodital process; c, cannula; Cx, gonocoxa; Pf, prefemoral region; PfP, prefemoral process; Sg, seminal groove; sl, solenomere; T, telopodite. Scale bar: 0.1 mm (B) or drawn not to scale (A, C)." figureDoi="http://doi.org/10.5281/zenodo.6669188" httpUri="https://zenodo.org/record/6669188/files/figure.png" pageId="8" pageNumber="95">Fig. 3B</figureCitation>
), another separate species-group of
<taxonomicName id="4C03C0CCBB49F176FBD971B9FB66FDD6" authorityName="Attems" authorityYear="1901" box="[1030,1228,511,537]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="B977675DBB49F176FBD971B9FB66FDD6" box="[1030,1228,511,537]" italics="true" pageId="8" pageNumber="95">Strongylosomides</emphasis>
</taxonomicName>
, the
<taxonomicName id="4C03C0CCBB49F176FADC71B9FA39FDD7" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" baseAuthorityName="Schubart" baseAuthorityYear="1956" box="[1283,1427,511,536]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="stercoriarius">
<emphasis id="B977675DBB49F176FADC71B9FA39FDD7" box="[1283,1427,511,536]" italics="true" pageId="8" pageNumber="95">stercoriarius</emphasis>
</taxonomicName>
- group, seems warranted to encompass both
<taxonomicName id="4C03C0CCBB49F176FDA37265FCA3FDF2" authority="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette, 2022" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" box="[636,777,547,573]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="B977675DBB49F176FDA37265FCA3FDF2" box="[636,777,547,573]" italics="true" pageId="8" pageNumber="95">S. troglobius</emphasis>
</taxonomicName>
<emphasis id="B977675DBB49F176FCD07265FCC2FDF2" bold="true" box="[783,872,547,573]" pageId="8" pageNumber="95">
<taxonomicNameLabel id="A244DA26BB49F176FCD07265FCC2FDF2" box="[783,872,547,573]" pageId="8" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C03C0CCBB49F176FC427265FBE2FDF3" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" baseAuthorityName="Schubart" baseAuthorityYear="1956" box="[925,1096,547,573]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="stercoriarius" status="comb. nov.">
<emphasis id="B977675DBB49F176FC427265FBE2FDF3" box="[925,1096,547,573]" italics="true" pageId="8" pageNumber="95">S. stercoriarius</emphasis>
</taxonomicName>
,
<emphasis id="B977675DBB49F176FB8A7265FB7EFDF2" bold="true" box="[1109,1236,547,573]" pageId="8" pageNumber="95">
<taxonomicNameLabel id="A244DA26BB49F176FB8A7265FB7EFDF2" box="[1109,1236,547,573]" pageId="8" pageNumber="95" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
The latter species thereby clearly bridges
<taxonomicName id="4C03C0CCBB49F176FE407201FD87FDAE" authority="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette, 2022" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" box="[415,557,583,609]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="B977675DBB49F176FE407201FD87FDAE" box="[415,557,583,609]" italics="true" pageId="8" pageNumber="95">S. troglobius</emphasis>
</taxonomicName>
<emphasis id="B977675DBB49F176FDEB7201FD27FDAE" bold="true" box="[564,653,583,609]" pageId="8" pageNumber="95">
<taxonomicNameLabel id="A244DA26BB49F176FDEB7201FD27FDAE" box="[564,653,583,609]" pageId="8" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
to the remaining congeners.
</paragraph>
<paragraph id="8BBCBB4FBB49F176FF18722DFE43FC4E" blockId="8.[151,1436,151,897]" pageId="8" pageNumber="95">
The few supposedly troglomorphic traits of
<taxonomicName id="4C03C0CCBB49F176FD64722DFCE0FD4A" authority="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette, 2022" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" box="[699,842,619,645]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="B977675DBB49F176FD64722DFCE0FD4A" box="[699,842,619,645]" italics="true" pageId="8" pageNumber="95">S. troglobius</emphasis>
</taxonomicName>
<emphasis id="B977675DBB49F176FC8D722DFC07FD4A" bold="true" box="[850,941,619,645]" pageId="8" pageNumber="95">
<taxonomicNameLabel id="A244DA26BB49F176FC8D722DFC07FD4A" box="[850,941,619,645]" pageId="8" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
alone, the more so as the mouthparts being unmodified, as is sometimes the case in troglobionts, and the antennae and legs relatively short, are certainly not enough for conferring the species the status of a troglobiont. Only direct biological observations will allow for this to be evaluated unequivocally. Even though epigean sampling was conducted on the same day near the cave entrance, with no additional material of
<taxonomicName id="4C03C0CCBB49F176FDE272BDFD61FCDA" authority="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette, 2022" authorityName="Golovatch &amp; Bouzan &amp; Gallo &amp; Bichuette" authorityYear="2022" box="[573,715,763,789]" class="Diplopoda" family="Chelodesmidae" genus="Strongylosomides" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="B977675DBB49F176FDE272BDFD61FCDA" box="[573,715,763,789]" italics="true" pageId="8" pageNumber="95">S. troglobius</emphasis>
</taxonomicName>
<emphasis id="B977675DBB49F176FD0D72BDFC81FCDA" bold="true" box="[722,811,763,789]" pageId="8" pageNumber="95">
<taxonomicNameLabel id="A244DA26BB49F176FD0D72BDFC81FCDA" box="[722,811,763,789]" pageId="8" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
revealed, this species was only found inside the cave in the deepest zones. Isolation in the hypogean environment, combined with certain presumably troglomorphic traits such as reduced body pigmentation and a quite fragile body with thin teguments, seems to be evidence of a neotroglobiont status of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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