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<document id="B89F029C5CF3BD8F279EE644D8A4046C" ID-CLB-Dataset="45976" ID-DOI="10.5281/zenodo.205242" ID-GBIF-Dataset="abff3177-6762-47ae-bc19-4503f7d41774" ID-ISSN="1175-5326" ID-Zenodo-Dep="205242" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460410161616" checkinUser="plazi" docAuthor="Cordeiro, Danilo, Bravo, Freddy &amp; De, Claudio J. B." docDate="2011" docId="03A8E828FF8C291DDCB5723FFA9DF936" docLanguage="en" docName="zt03101p037.pdf" docOrigin="Zootaxa 3101" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Psychoda complexa Cordeiro, Bravo &amp; De, 2011, sp. nov." docType="treatment" docVersion="17" lastPageNumber="21" masterDocId="FF919050FF982909DC227120FFDFFF8B" masterDocTitle="Taxonomy of Brazilian Psychoda Latreille, 1796 (Diptera, Psychodidae) with the description of thirteen new species" masterLastPageNumber="37" masterPageNumber="1" pageNumber="21" updateTime="1698300371931" updateUser="ExternalLinkService">
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<mods:title id="C5663DB98377F8FCC33A9302DBA9BE1D">Taxonomy of Brazilian Psychoda Latreille, 1796 (Diptera, Psychodidae) with the description of thirteen new species</mods:title>
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<mods:name id="7164A74F8FA7419D2FEA86D9D8D02916" type="personal">
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<mods:namePart id="AC4C160C21AFB036309ACBDBC1559404">Cordeiro, Danilo</mods:namePart>
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<mods:namePart id="4C66026F7BAA4D8D31328C77BF1083AB">Bravo, Freddy</mods:namePart>
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<mods:namePart id="486AF0CC07F58696FFCD72771D2118AC">De, Claudio J. B.</mods:namePart>
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<mods:title id="6966A7E021489E5E8D5D62C8B4619790">Zootaxa</mods:title>
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<mods:date id="3D6391E1F881B0984A5E72E8C98CE63A">2011</mods:date>
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<treatment id="03A8E828FF8C291DDCB5723FFA9DF936" ID-DOI="http://doi.org/10.5281/zenodo.6192115" ID-GBIF-Taxon="119423187" ID-Zenodo-Dep="6192115" LSID="urn:lsid:zoobank.org:act:5A4BAAB5-6595-4D50-A6BD-C13201DD3E5C" httpUri="http://treatment.plazi.org/id/03A8E828FF8C291DDCB5723FFA9DF936" lastPageNumber="21" pageId="20" pageNumber="21">
<subSubSection id="C31B0AB5FF8C291DDCB5723FFEF8FCD1" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BBE593EFF8C291DDCB5723FFE2EFCB2" blockId="20.[151,497,799,858]" box="[151,497,799,825]" pageId="20" pageNumber="21">
<heading id="D0F6EE52FF8C291DDCB5723FFE2EFCB2" bold="true" box="[151,497,799,825]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="B975852CFF8C291DDCB5723FFE2EFCB2" bold="true" box="[151,497,799,825]" pageId="20" pageNumber="21">
<taxonomicName id="4C0122BDFF8C291DDCB5723FFE59FCB2" ID-CoL="8DK3T" box="[151,390,799,825]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="complexa" status="sp. nov.">
<emphasis id="B975852CFF8C291DDCB5723FFE59FCB2" bold="true" box="[151,390,799,825]" italics="true" pageId="20" pageNumber="21">Psychoda complexa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2463857FF8C291DDDB6723FFE2EFCB2" box="[404,497,799,825]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE593EFF8C291DDCB57262FEF8FCD1" blockId="20.[151,497,799,858]" box="[151,295,834,858]" pageId="20" pageNumber="21">
(
<figureCitation id="133A45BBFF8C291DDCBD7262FEC0FCD1" box="[159,287,834,858]" captionStart="FIGURES 63 70" captionStartId="21.[151,264,1657,1680]" captionTargetBox="[169,1436,202,1603]" captionTargetId="figure@21.[151,1436,193,1636]" captionTargetPageId="21" captionText="FIGURES 63 70. Psychoda complexa sp. nov.: 63. Head. 64. Antenna: scape, pedicel and basal flagellomeres. 65. Palpus. 65. Labellum. 67. Wing. 68. Distitarsus. 69. Subgenital plate and genital chamber. 70. Subgenital plate, internal ornamentation. [Scale 0.5 mm (67), other pictures 0.06 mm]." httpUri="https://zenodo.org/record/205251/files/figure.png" pageId="20" pageNumber="21">Figs 6370</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31B0AB5FF8C291DDCB572ABFED0FC4D" pageId="20" pageNumber="21" type="etymology">
<paragraph id="8BBE593EFF8C291DDCB572ABFED0FC4D" blockId="20.[151,1437,906,1726]" pageId="20" pageNumber="21">
<emphasis id="B975852CFF8C291DDCB572ABFEC1FC28" bold="true" box="[151,286,907,931]" pageId="20" pageNumber="21">Etymology.</emphasis>
The species name,
<taxonomicName id="4C0122BDFF8C291DDDDD72ABFDB5FC28" box="[511,618,907,931]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="complexa">
<emphasis id="B975852CFF8C291DDDDD72ABFDB5FC28" box="[511,618,907,931]" italics="true" pageId="20" pageNumber="21">complexa</emphasis>
</taxonomicName>
, is based on the complex structure of the subgenital plate internal ornamentation.
</paragraph>
</subSubSection>
<subSubSection id="C31B0AB5FF8C291DDCE772F2FDE3FBB9" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="8BBE593EFF8C291DDCE772F2FDE3FBB9" blockId="20.[151,1437,906,1726]" pageId="20" pageNumber="21">
<emphasis id="B975852CFF8C291DDCE772F2FEE0FC60" bold="true" box="[197,319,978,1003]" pageId="20" pageNumber="21">Diagnosis.</emphasis>
Palpal formula 1.0:1.2:1.0:1.25; radial and medial forks incomplete; distitarsus without apical projection; subgenital plate wider than long, with a narrow concavity between lobes; genital digit large, stretching from base to apex of subgenital plate.
</paragraph>
</subSubSection>
<subSubSection id="C31B0AB5FF8C291DDCE7751FFE59FA4A" pageId="20" pageNumber="21" type="description">
<paragraph id="8BBE593EFF8C291DDCE7751FFBD8FA16" blockId="20.[151,1437,906,1726]" pageId="20" pageNumber="21">
<emphasis id="B975852CFF8C291DDCE7751FFEFCFBDC" bold="true" box="[197,291,1087,1111]" pageId="20" pageNumber="21">Female.</emphasis>
Head (
<figureCitation id="133A45BBFF8C291DDD56751EFE17FBDD" box="[372,456,1086,1110]" captionStart="FIGURES 63 70" captionStartId="21.[151,264,1657,1680]" captionTargetBox="[169,1436,202,1603]" captionTargetId="figure@21.[151,1436,193,1636]" captionTargetPageId="21" captionText="FIGURES 63 70. Psychoda complexa sp. nov.: 63. Head. 64. Antenna: scape, pedicel and basal flagellomeres. 65. Palpus. 65. Labellum. 67. Wing. 68. Distitarsus. 69. Subgenital plate and genital chamber. 70. Subgenital plate, internal ornamentation. [Scale 0.5 mm (67), other pictures 0.06 mm]." httpUri="https://zenodo.org/record/205251/files/figure.png" pageId="20" pageNumber="21">Fig. 63</figureCitation>
): vertex, frons and clypeus pilose; hair patch of frons extending to facet row 1 or 2; eye bridge with 4 facets; eyes separated by 1.52.0 facet diameters; 56 supra-ocular setae; interocular suture sometimes present as a weak line; clypeus with 2 stronger lateral alveoli; frontoclypeal suture absent; antenna incomplete; scape cylindrical, 1.5 the length of subspherical pedicel (
<figureCitation id="133A45BBFF8C291DDF78758AFC70FB49" box="[858,943,1194,1218]" captionStart="FIGURES 63 70" captionStartId="21.[151,264,1657,1680]" captionTargetBox="[169,1436,202,1603]" captionTargetId="figure@21.[151,1436,193,1636]" captionTargetPageId="21" captionText="FIGURES 63 70. Psychoda complexa sp. nov.: 63. Head. 64. Antenna: scape, pedicel and basal flagellomeres. 65. Palpus. 65. Labellum. 67. Wing. 68. Distitarsus. 69. Subgenital plate and genital chamber. 70. Subgenital plate, internal ornamentation. [Scale 0.5 mm (67), other pictures 0.06 mm]." httpUri="https://zenodo.org/record/205251/files/figure.png" pageId="20" pageNumber="21">Fig. 64</figureCitation>
); ascoids in Y; palpal formula 1.0:1.2(1.13 1.21):1.0:1.25 (
<figureCitation id="133A45BBFF8C291DDD6775EEFE47FB6D" box="[325,408,1230,1254]" captionStart="FIGURES 63 70" captionStartId="21.[151,264,1657,1680]" captionTargetBox="[169,1436,202,1603]" captionTargetId="figure@21.[151,1436,193,1636]" captionTargetPageId="21" captionText="FIGURES 63 70. Psychoda complexa sp. nov.: 63. Head. 64. Antenna: scape, pedicel and basal flagellomeres. 65. Palpus. 65. Labellum. 67. Wing. 68. Distitarsus. 69. Subgenital plate and genital chamber. 70. Subgenital plate, internal ornamentation. [Scale 0.5 mm (67), other pictures 0.06 mm]." httpUri="https://zenodo.org/record/205251/files/figure.png" pageId="20" pageNumber="21">Fig. 65</figureCitation>
); labellum with 3 apical and 1 subapical teeth, and 1 or 2 lateral setae (
<figureCitation id="133A45BBFF8C291DD89875EEFAD1FB6D" box="[1210,1294,1230,1254]" captionStart="FIGURES 63 70" captionStartId="21.[151,264,1657,1680]" captionTargetBox="[169,1436,202,1603]" captionTargetId="figure@21.[151,1436,193,1636]" captionTargetPageId="21" captionText="FIGURES 63 70. Psychoda complexa sp. nov.: 63. Head. 64. Antenna: scape, pedicel and basal flagellomeres. 65. Palpus. 65. Labellum. 67. Wing. 68. Distitarsus. 69. Subgenital plate and genital chamber. 70. Subgenital plate, internal ornamentation. [Scale 0.5 mm (67), other pictures 0.06 mm]." httpUri="https://zenodo.org/record/205251/files/figure.png" pageId="20" pageNumber="21">Fig. 66</figureCitation>
). Wing (
<figureCitation id="133A45BBFF8C291DD94D75EEFF68FA81" captionStart="FIGURES 63 70" captionStartId="21.[151,264,1657,1680]" captionTargetBox="[169,1436,202,1603]" captionTargetId="figure@21.[151,1436,193,1636]" captionTargetPageId="21" captionText="FIGURES 63 70. Psychoda complexa sp. nov.: 63. Head. 64. Antenna: scape, pedicel and basal flagellomeres. 65. Palpus. 65. Labellum. 67. Wing. 68. Distitarsus. 69. Subgenital plate and genital chamber. 70. Subgenital plate, internal ornamentation. [Scale 0.5 mm (67), other pictures 0.06 mm]." httpUri="https://zenodo.org/record/205251/files/figure.png" pageId="20" pageNumber="21">Fig. 67</figureCitation>
): Sc vein not extending beyond base of vein Rs; radial and medial forks incomplete; vein R5 and costal cell slightly darkened. Distitarsus without apical projection (
<figureCitation id="133A45BBFF8C291DDF017439FCA4FABA" box="[803,891,1305,1329]" captionStart="FIGURES 63 70" captionStartId="21.[151,264,1657,1680]" captionTargetBox="[169,1436,202,1603]" captionTargetId="figure@21.[151,1436,193,1636]" captionTargetPageId="21" captionText="FIGURES 63 70. Psychoda complexa sp. nov.: 63. Head. 64. Antenna: scape, pedicel and basal flagellomeres. 65. Palpus. 65. Labellum. 67. Wing. 68. Distitarsus. 69. Subgenital plate and genital chamber. 70. Subgenital plate, internal ornamentation. [Scale 0.5 mm (67), other pictures 0.06 mm]." httpUri="https://zenodo.org/record/205251/files/figure.png" pageId="20" pageNumber="21">Fig. 68</figureCitation>
). Terminalia: subgenital plate wider than long, bilobed, pilose on distal half and with a narrow concavity between lobes (
<figureCitation id="133A45BBFF8C291DDFFF741DFBEDFADE" box="[989,1074,1341,1365]" captionStart="FIGURES 63 70" captionStartId="21.[151,264,1657,1680]" captionTargetBox="[169,1436,202,1603]" captionTargetId="figure@21.[151,1436,193,1636]" captionTargetPageId="21" captionText="FIGURES 63 70. Psychoda complexa sp. nov.: 63. Head. 64. Antenna: scape, pedicel and basal flagellomeres. 65. Palpus. 65. Labellum. 67. Wing. 68. Distitarsus. 69. Subgenital plate and genital chamber. 70. Subgenital plate, internal ornamentation. [Scale 0.5 mm (67), other pictures 0.06 mm]." httpUri="https://zenodo.org/record/205251/files/figure.png" pageId="20" pageNumber="21">Fig. 69</figureCitation>
); genital digit large, going from base to apex of subgenital plate; subgenital plate internally with diagonal lines and a complex, well sclerotized structure next to apex of genital digit (
<figureCitation id="133A45BBFF8C291DDE6674A5FD47FA16" box="[580,664,1413,1437]" captionStart="FIGURES 63 70" captionStartId="21.[151,264,1657,1680]" captionTargetBox="[169,1436,202,1603]" captionTargetId="figure@21.[151,1436,193,1636]" captionTargetPageId="21" captionText="FIGURES 63 70. Psychoda complexa sp. nov.: 63. Head. 64. Antenna: scape, pedicel and basal flagellomeres. 65. Palpus. 65. Labellum. 67. Wing. 68. Distitarsus. 69. Subgenital plate and genital chamber. 70. Subgenital plate, internal ornamentation. [Scale 0.5 mm (67), other pictures 0.06 mm]." httpUri="https://zenodo.org/record/205251/files/figure.png" pageId="20" pageNumber="21">Fig. 70</figureCitation>
); genital chamber oval (
<figureCitation id="133A45BBFF8C291DDF8474A5FC25FA16" box="[934,1018,1413,1437]" captionStart="FIGURES 63 70" captionStartId="21.[151,264,1657,1680]" captionTargetBox="[169,1436,202,1603]" captionTargetId="figure@21.[151,1436,193,1636]" captionTargetPageId="21" captionText="FIGURES 63 70. Psychoda complexa sp. nov.: 63. Head. 64. Antenna: scape, pedicel and basal flagellomeres. 65. Palpus. 65. Labellum. 67. Wing. 68. Distitarsus. 69. Subgenital plate and genital chamber. 70. Subgenital plate, internal ornamentation. [Scale 0.5 mm (67), other pictures 0.06 mm]." httpUri="https://zenodo.org/record/205251/files/figure.png" pageId="20" pageNumber="21">Fig. 69</figureCitation>
).
</paragraph>
<paragraph id="8BBE593EFF8C291DDCE7748AFE59FA4A" blockId="20.[151,1437,906,1726]" box="[197,390,1449,1474]" pageId="20" pageNumber="21">
<emphasis id="B975852CFF8C291DDCE7748AFED7FA49" bold="true" box="[197,264,1450,1474]" pageId="20" pageNumber="21">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31B0AB5FF8C291DDCE774EDFE90F9FE" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BBE593EFF8C291DDCE774EDFE90F9FE" blockId="20.[151,1437,906,1726]" pageId="20" pageNumber="21">
<emphasis id="B975852CFF8C291DDCE774EDFE6DFA6D" bold="true" box="[197,434,1485,1510]" pageId="20" pageNumber="21">Material examined.</emphasis>
<typeStatus id="54BAE79CFF8C291DDD9E74EDFDFBFA6E" box="[444,548,1485,1509]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
Ƥ
<collectingCountry id="F31619AEFF8C291DDE6C74EEFD69FA6D" box="[590,694,1486,1510]" name="Brazil" pageId="20" pageNumber="21">BRAZIL</collectingCountry>
, Amazonas, Silves, Saracá,
<date id="FFBF7FFEFF8C291DDFDC74EDFBA8FA6E" box="[1022,1143,1485,1509]" pageId="20" pageNumber="21" value="1997-06-01">01.vi.1997</date>
, 6cdc
<quantity id="4CF9F4DBFF8C291DD8FC74EDFADAFA6E" box="[1246,1285,1485,1509]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="1.0">1m</quantity>
, no collector name (
<collectionCode id="ED10C1FBFF8C291DDCC474D1FEF3F982" box="[230,300,1521,1545]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="20" pageNumber="21">INPA</collectionCode>
).
<typeStatus id="54BAE79CFF8C291DDD6074D2FE6BF981" box="[322,436,1522,1546]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
: 2 Ƥ same locality and date of
<typeStatus id="54BAE79CFF8C291DDF1274D1FC4DF982" box="[816,914,1521,1545]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
(MZUEFS); 1 Ƥ Amazonas, Silves, Saracá,
<date id="FFBF7FFEFF8C291DDCB57735FECEF9A6" box="[151,273,1557,1581]" pageId="20" pageNumber="21" value="1997-06-30">30.vi.1997</date>
, 6cdc
<quantity id="4CF9F4DBFF8C291DDD4E7735FE4DF9A6" box="[364,402,1557,1581]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="1.0">1m</quantity>
, no collector name (
<collectionCode id="ED10C1FBFF8C291DDE517735FD66F9A6" box="[627,697,1557,1581]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="20" pageNumber="21">INPA</collectionCode>
); 1 Ƥ Amazonas, Silves, Saracá, área alterada,
<date id="FFBF7FFEFF8C291DD8E47735FA98F9A6" box="[1222,1351,1557,1581]" pageId="20" pageNumber="21" value="1997-12-17">17.xii.1997</date>
, 6cdc m, col.
<collectingCountry id="F31619AEFF8C291DDCC87719FED2F9DA" box="[234,269,1593,1617]" name="Russia" pageId="20" pageNumber="21">RF</collectingCountry>
/FLS (
<collectionCode id="ED10C1FBFF8C291DDD717719FE47F9DA" box="[339,408,1593,1617]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="20" pageNumber="21">INPA</collectionCode>
); 1 Ƥ
<collectingCountry id="F31619AEFF8C291DDDF9771AFD9CF9D9" box="[475,579,1594,1618]" name="Brazil" pageId="20" pageNumber="21">BRAZIL</collectingCountry>
, Paraná, Guarapuava, Est. Águas Sta. Clara,
<date id="FFBF7FFEFF8C291DD81A7719FB7BF9DA" box="[1080,1188,1593,1617]" pageId="20" pageNumber="21" value="1987-01-12">12.i.1987</date>
, Malaise trap, Profaupar 3.4 (
<collectionCode id="ED10C1FBFF8C291DDCD1777DFE9CF9FE" box="[243,323,1629,1653]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="20" pageNumber="21">DZUP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B0AB5FF8C291DDCE777A1FD1CF912" box="[197,707,1665,1690]" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8BBE593EFF8C291DDCE777A1FD1CF912" blockId="20.[151,1437,906,1726]" box="[197,707,1665,1690]" pageId="20" pageNumber="21">
<emphasis id="B975852CFF8C291DDCE777A1FE81F911" bold="true" box="[197,350,1665,1690]" pageId="20" pageNumber="21">Distribution.</emphasis>
<collectingCountry id="F31619AEFF8C291DDD4777A1FE76F912" box="[357,425,1665,1689]" name="Brazil" pageId="20" pageNumber="21">Brazil</collectingCountry>
(Amazonas and Paraná).
</paragraph>
</subSubSection>
<subSubSection id="C31B0AB5FF8C291DDCE77785FA9DF936" box="[197,1346,1701,1726]" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8BBE593EFF8C291DDCE77785FA9DF936" blockId="20.[151,1437,906,1726]" box="[197,1346,1701,1726]" pageId="20" pageNumber="21">
<emphasis id="B975852CFF8C291DDCE77785FE92F935" bold="true" box="[197,333,1701,1726]" pageId="20" pageNumber="21">Comments.</emphasis>
As the apical flagellomeres are missing, this species cannot be assigned to any subgenera.
</paragraph>
</subSubSection>
</treatment>
</document>