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<document id="CD6E457F5058D046C06406A3FFAA27FB" ID-DOI="10.5252/z2016n4a2" ID-ISSN="1638-9387" ID-Zenodo-Dep="4578343" ID-ZooBank="urn:lsid:zoobank.org:pub:BFFD82EF-50C9-42BF-8493-DF57591EA4FF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614790962589" checkinUser="felipe" docAuthor="Pyron, Robert Alexander, Ganesh, Sumaithangi Rajagopalan, Sayyed, Amit, Sharma, Vivek, Wallach, Van &amp; Somaweera, Ruchira" docDate="2016" docId="03A82A47833DFFA5FC50FDC0FBCC7C99" docLanguage="en" docName="Zoosystema.38.4.453-506.pdf" docOrigin="Zoosystema 38 (4)" docSource="http://dx.doi.org/10.5252/z2016n4a2" docStyle="DocumentStyle:0AF8C315773078909029C6FC3CC05C6C.3:Zoosystema.2015-2017.journal_article" docStyleId="0AF8C315773078909029C6FC3CC05C6C" docStyleName="Zoosystema.2015-2017.journal_article" docStyleVersion="3" docTitle="Rhinophis sanguineus Beddome 1863" docType="treatment" docVersion="5" lastPageNumber="485" masterDocId="FF91523F831DFF85FFCFFFB5FFF97B72" masterDocTitle="A catalogue and systematic overview of the shield-tailed snakes (Serpentes: Uropeltidae)" masterLastPageNumber="506" masterPageNumber="453" pageNumber="485" updateTime="1698879475557" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="196D2954181C5782B3434C9AD4809342">A catalogue and systematic overview of the shield-tailed snakes (Serpentes: Uropeltidae)</mods:title>
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<mods:namePart id="05C43D963179B5348C91A38AD3F3421A">Pyron, Robert Alexander</mods:namePart>
<mods:affiliation id="6C168891B9C2D6B0D6F76F242761F9F0">Department of Biological Sciences, The George Washington University, 2029 G St. NW, Washington, D. C. 20052 (United States) rpyron @ colubroid. org</mods:affiliation>
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<mods:namePart id="88E6021D85B8686FB2537CC21133898A">Sayyed, Amit</mods:namePart>
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<mods:affiliation id="088E9287A6A2079BEC2B33F101994776">Sanjeevni Nagar, Jabalpur, Madhya Pradesh 482 003 (India)</mods:affiliation>
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<mods:namePart id="0D2A243E382D8D2D2A1BD349FCA4B68B">Wallach, Van</mods:namePart>
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<taxonomicName id="4C01E0D2833DFFA5FC50FDC0FAC479E2" authority="Beddome, 1863" authorityName="Beddome" authorityYear="1863" box="[927,1341,629,656]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="485" phylum="Chordata" rank="species" species="sanguineus">
<emphasis id="B9754743833DFFA5FC50FDC0FB7079E2" bold="true" box="[927,1161,629,656]" italics="true" pageId="32" pageNumber="485">Rhinophis sanguineus</emphasis>
Beddome, 1863
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(
<figureCitation id="133A87D4833DFFA5FB8DFD23FB6079C2" box="[1090,1177,662,688]" captionStart="FIG" captionStartId="30.[132,143,1964,1982]" captionTargetBox="[132,1455,214,1937]" captionTargetId="figure-94@30.[132,1455,190,1944]" captionTargetPageId="30" captionText="FIG. 7. — Some uropeltid species photographed in life: A, Rhinophis homolepis Hemprich, 1820; B, R. melanogaster (Gray, 1858); C, R. oxyrhynchus (Schneider, 1801); D, R. philippinus (Cuvier, 1829); E, R. phillipsi (Nicholls, 1929) n. comb.; F, R. punctatus Müller,1832; G, R. saffragamus (Kelaart, 1853) n. comb.; H, R. sanguineus Beddome, 1863. Photos by RAP, RS, SRG, VS, A. Dey, R. Pethiyagoda, and S. Kehimkar." figureDoi="http://doi.org/10.5281/zenodo.4578357" httpUri="https://zenodo.org/record/4578357/files/figure.png" pageId="32" pageNumber="485">Fig. 7H</figureCitation>
)
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<treatmentCitation id="0AA0BD40833DFFA5FCE2FD5BFB417874" author="BEDDOME R. H." box="[813,1208,750,774]" page="47" pageId="32" pageNumber="485" year="1863">
<taxonomicName id="4C01E0D2833DFFA5FCE2FD5BFB417874" authority="Beddome, 1863 a: 47" authorityName="Beddome" authorityPageNumber="47" authorityYear="1863" box="[813,1208,750,774]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="485" phylum="Chordata" rank="species" species="sanguineus">
<emphasis id="B9754743833DFFA5FCE2FD5BFC147874" box="[813,1005,750,774]" italics="true" pageId="32" pageNumber="485">Rhinophis sanguineus</emphasis>
<bibRefCitation id="EF90E6A0833DFFA5FC3EFD5BFB417874" author="BEDDOME R. H." box="[1009,1208,750,774]" pageId="32" pageNumber="485" pagination="41 - 48" refId="ref46637" refString="BEDDOME R. H. 1863 a. - Further notes upon the snakes of the Madras Presidency; with descriptions of new species. Madras Quarterly Journal of Medical Science 6: 41 - 48." type="journal article" year="1863">Beddome, 1863a: 47</bibRefCitation>
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.
<bibRefCitation id="EF90E6A0833DFFA5FB70FD5BFA907874" author="BEDDOME R. H." box="[1215,1385,750,774]" pageId="32" pageNumber="485" pagination="41 - 48" refId="ref46637" refString="BEDDOME R. H. 1863 a. - Further notes upon the snakes of the Madras Presidency; with descriptions of new species. Madras Quarterly Journal of Medical Science 6: 41 - 48." type="journal article" year="1863">Beddome (1863a)</bibRefCitation>
reports collecting several
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, but
<bibRefCitation id="EF90E6A0833DFFA5FB99FCBCFAF57853" author="BOULENGER G. A." box="[1110,1292,777,801]" pageId="32" pageNumber="485" refId="ref47173" refString="BOULENGER G. A. 1893 a. - Catalogue of the snakes in the British Museum (Natural History). Volume I., containing the families Typhlopidae, Glauconiidae, Boidae, Ilysiidae, Uropeltidae, Xenopeltidae and Colubridae aglyphae, part. British Museum (Natural History), London, 448 p." type="book" year="1893">Boulenger (1893a)</bibRefCitation>
reports only one male with 197 ventrals and nine subcaudals.
<bibRefCitation id="EF90E6A0833DFFA5FB1CFC96FAB57849" author="GANS C." box="[1235,1356,803,827]" pageId="32" pageNumber="485" pagination="1 - 29" refId="ref48814" refString="GANS C. 1966. - Liste der rezenten Reptilien und Amphibien. Uropeltidae. Das Tierreich 84: 1 - 29." type="journal article" year="1966">Gans (1966)</bibRefCitation>
also states that only one
<typeStatus id="54BA25F3833DFFA5FC76FC8BFBFD7824" box="[953,1028,830,854]" pageId="32" pageNumber="485" type="syntype">syntype</typeStatus>
(BMNH 1864.3.9.2 now: 1946.1.16.54), catalogued as “one of the types”, is now known.
<bibRefCitation id="EF90E6A0833DFFA5FB23FCEDFA567802" author="WALLACH V. &amp; WILLIAMS K. L. &amp; BOUNDY J." box="[1260,1455,856,880]" pageId="32" pageNumber="485" refId="ref52220" refString="WALLACH V., WILLIAMS K. L. &amp; BOUNDY J. 2014. - Snakes of the World: a Catalogue of Living and Extinct Species. CRC Press, Boca Raton, 1209 p." type="book" year="2014">
Wallach
<emphasis id="B9754743833DFFA5FAF1FCECFA937802" box="[1342,1386,856,880]" italics="true" pageId="32" pageNumber="485">et al.</emphasis>
(2014)
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also list BMNH 1874.4.29.110 (= 1946.1.16.76) as a
<typeStatus id="54BA25F3833DFFA5FAF7FCC6FA7C78F9" box="[1336,1413,883,907]" pageId="32" pageNumber="485" type="syntype">syntype</typeStatus>
, but this specimen is putatively the
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of
<taxonomicName id="4C01E0D2833DFFA5FB71FC38FC6678CD" authority="Beddome, 1863" authorityName="Beddome" authorityYear="1863" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="485" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B9754743833DFFA5FB71FC38FA8278D7" box="[1214,1403,909,933]" italics="true" pageId="32" pageNumber="485">Rhinophis microlepis</emphasis>
Beddome, 1863
</taxonomicName>
(see
<bibRefCitation id="EF90E6A0833DFFA5FC04FC1DFBCF78CD" author="GANS C." box="[971,1078,935,960]" pageId="32" pageNumber="485" pagination="1 - 29" refId="ref48814" refString="GANS C. 1966. - Liste der rezenten Reptilien und Amphibien. Uropeltidae. Das Tierreich 84: 1 - 29." type="journal article" year="1966">Gans 1966</bibRefCitation>
). We designate BMNH 1946.1.16.54 as the
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, to remove any ambiguity.Type locality: Cherambadi, northern
<collectingRegion id="49C555B3833DFFA5FC47FC69FC057886" box="[904,1020,988,1012]" country="India" name="Kerala" pageId="32" pageNumber="485">Kerala state</collectingRegion>
,
<collectingCountry id="F316DBC1833DFFA5FBCAFC69FBC57886" box="[1029,1084,988,1012]" name="India" pageId="32" pageNumber="485">India</collectingCountry>
.
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</paragraph>
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<treatmentCitation id="0AA0BD40833DFFA5FCE2FBA8FB187F47" author="BEDDOME R. H." box="[813,1249,1053,1077]" page="47" pageId="32" pageNumber="485" year="1863">
<taxonomicName id="4C01E0D2833DFFA5FCE2FBA8FB187F47" authority="Beddome, 1863 a: 47" authorityName="Beddome" authorityPageNumber="47" authorityYear="1863" box="[813,1249,1053,1077]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="485" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B9754743833DFFA5FCE2FBA8FC037F47" box="[813,1018,1053,1077]" italics="true" pageId="32" pageNumber="485">Rhinophis microlepis</emphasis>
<bibRefCitation id="EF90E6A0833DFFA5FBCCFBA8FB187F47" author="BEDDOME R. H." box="[1027,1249,1053,1077]" pageId="32" pageNumber="485" pagination="41 - 48" refId="ref46637" refString="BEDDOME R. H. 1863 a. - Further notes upon the snakes of the Madras Presidency; with descriptions of new species. Madras Quarterly Journal of Medical Science 6: 41 - 48." type="journal article" year="1863">Beddome, 1863a: 47</bibRefCitation>
</taxonomicName>
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.
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: BMNH 1874.4.29.110 (= 1946.1.16.76). Type locality: Minchin estate,
<collectingRegion id="49C555B3833DFFA5FCE1FBE7FC5A7F1B" box="[814,931,1106,1130]" country="India" name="Kerala" pageId="32" pageNumber="485">Kerala state</collectingRegion>
,
<collectingCountry id="F316DBC1833DFFA5FC62FBE7FC1D7F18" box="[941,996,1106,1130]" name="India" pageId="32" pageNumber="485">India</collectingCountry>
. Designated as a junior subjective synonym by
<bibRefCitation id="EF90E6A0833DFFA5FCE2FBD9FC2E7FF6" author="BEDDOME R. H." box="[813,983,1132,1156]" pageId="32" pageNumber="485" pagination="3 - 33" refId="ref46985" refString="BEDDOME R. H. 1886. - An account of the earth-snakes of the peninsula of India and Ceylon. Annals and Magazine of Natural History 17: 3 - 33." type="journal article" year="1886">Beddome (1886)</bibRefCitation>
. See
<bibRefCitation id="EF90E6A0833DFFA5FBC9FBD9FB427FF6" author="BEDDOME R. H." box="[1030,1211,1132,1156]" pageId="32" pageNumber="485" pagination="41 - 48" refId="ref46637" refString="BEDDOME R. H. 1863 a. - Further notes upon the snakes of the Madras Presidency; with descriptions of new species. Madras Quarterly Journal of Medical Science 6: 41 - 48." type="journal article" year="1863">Beddome (1863a)</bibRefCitation>
.
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</paragraph>
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DISTRIBUTION. —
<collectingCountry id="F316DBC1833DFFA5FC26FB19FBE67FB6" box="[1001,1055,1196,1220]" name="India" pageId="32" pageNumber="485">India</collectingCountry>
, known from the Nilgiris northwards to the Agumbe hills,
<emphasis id="B9754743833DFFA5FC2EFB7DFC097FAD" box="[993,1008,1224,1247]" italics="true" pageId="32" pageNumber="485">c.</emphasis>
<quantity id="4CF936B4833DFFA5FC39FB72FB8B7FAC" box="[1014,1138,1223,1247]" metricMagnitude="2" metricUnit="m" metricValue="9.075" metricValueMax="10.65" metricValueMin="7.5" pageId="32" pageNumber="485" unit="m" value="907.5" valueMax="1065.0" valueMin="750.0">750-1065 m</quantity>
(see
<bibRefCitation id="EF90E6A0833DFFA5FB6CFB72FAC17FAD" author="FERGUSON H. S." box="[1187,1336,1223,1247]" pageId="32" pageNumber="485" pagination="68 - 77" refId="ref48336" refString="FERGUSON H. S. 1895. - List of snakes taken in Travancore from 1888 - 1895. Journal of the Bombay Natural History Society 10: 68 - 77." type="journal article" year="1895">Ferguson 1895</bibRefCitation>
;
<bibRefCitation id="EF90E6A0833DFFA5FA8CFB72FA527FAD" author="WALL F." box="[1347,1451,1223,1247]" pageId="32" pageNumber="485" pagination="552 - 584" refId="ref51926" refString="WALL F. 1919. - Notes on a collection of snakes made in the Nilgiri hills and adjacent Wynaad. Journal of the Bombay Natural History Society 26: 552 - 584." type="journal article" year="1919">Wall 1919</bibRefCitation>
;
<bibRefCitation id="EF90E6A0833DFFA5FCE2FB54FBE77F8B" author="AENGALS R. &amp; GANESH S." box="[813,1054,1249,1273]" pageId="32" pageNumber="485" pagination="61 - 65" refId="ref46558" refString="AENGALS R. &amp; GANESH S. 2013. - Rhinophis goweri - A new species of shieldtail snake from the southern Eastern Ghats, India. Russian Journal of Herpetology 20: 61 - 65." type="journal article" year="2013">Aengals &amp; Ganesh 2013</bibRefCitation>
;
<bibRefCitation id="EF90E6A0833DFFA5FBE7FB54FB1F7F8B" author="WALLACH V. &amp; WILLIAMS K. L. &amp; BOUNDY J." box="[1064,1254,1249,1273]" pageId="32" pageNumber="485" refId="ref52220" refString="WALLACH V., WILLIAMS K. L. &amp; BOUNDY J. 2014. - Snakes of the World: a Catalogue of Living and Extinct Species. CRC Press, Boca Raton, 1209 p." type="book" year="2014">
Wallach
<emphasis id="B9754743833DFFA5FBB3FB57FB527F8B" box="[1148,1195,1249,1273]" italics="true" pageId="32" pageNumber="485">et al.</emphasis>
2014
</bibRefCitation>
;
<bibRefCitation id="EF90E6A0833DFFA5FB3FFB54FA8E7F8B" author="GANESH S. R." box="[1264,1399,1249,1273]" pageId="32" pageNumber="485" pagination="13 - 24" refId="ref48505" refString="GANESH S. R. 2015. - Shieldtail snakes (Reptilia: Uropeltidae. - the Darwin's finches of south Indian snake fauna?, in KANNAN, P. (Ed.) Manual on Identification and Preparation of Keys of Snakes with Special Reference to their Venomous Nature in India. Proceedings by Government Arts College, Udhagamandalam, Tamilnadu, India, 13 - 24." type="book chapter" year="2015">Ganesh 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31BC8DA833DFFA5FCE2FA81FB4D7D5E" pageId="32" pageNumber="485" type="description">
<paragraph id="8BBE9B51833DFFA5FCE2FA81FC447E3E" blockId="32.[811,1457,1332,1581]" box="[813,957,1332,1358]" pageId="32" pageNumber="485">DESCRIPTION</paragraph>
<paragraph id="8BBE9B51833DFFA5FCE2FAE6FB4D7D5E" blockId="32.[811,1457,1332,1581]" pageId="32" pageNumber="485">
Maximum total length
<emphasis id="B9754743833DFFA5FBE9FAE1FBCE7E1F" box="[1062,1079,1364,1389]" italics="true" pageId="32" pageNumber="485">c.</emphasis>
<quantity id="4CF936B4833DFFA5FBF3FAE6FB627E1C" box="[1084,1179,1363,1390]" metricMagnitude="-1" metricUnit="m" metricValue="4.6" pageId="32" pageNumber="485" unit="mm" value="460.0">460 mm</quantity>
, ventrals 182-218 though
<bibRefCitation id="EF90E6A0833DFFA5FCE2FAC1FC3F7EFF" author="MASON G. E." box="[813,966,1395,1422]" pageId="32" pageNumber="485" pagination="184 - 186" refId="ref50518" refString="MASON G. E. 1888. - Description of a new earth-snake of the genus Silybura from the Bombay Presidency with remarks on little known Uropeltidae. Annals and Magazine of Natural History 1: 184 - 186." type="journal article" year="1888">Mason (1888)</bibRefCitation>
reported 166-183 from putatively conspecific populations in the Wullingy forests near Palghat, subcaudals 5-11, dorsal scales in 15 rows at midbody (see
<bibRefCitation id="EF90E6A0833DFFA5FAFBFA06FA537EBF" author="WALL F." box="[1332,1450,1459,1485]" pageId="32" pageNumber="485" pagination="552 - 584" refId="ref51926" refString="WALL F. 1919. - Notes on a collection of snakes made in the Nilgiri hills and adjacent Wynaad. Journal of the Bombay Natural History Society 26: 552 - 584." type="journal article" year="1919">Wall 1919</bibRefCitation>
;
<bibRefCitation id="EF90E6A0833DFFA5FCE1FA66FC547E9F" author="SMITH M. A." box="[814,941,1491,1517]" pageId="32" pageNumber="485" refId="ref51495" refString="SMITH M. A. 1943. - The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. Vol. 3 (Serpentes). Taylor &amp; Francis, London, 583 p." type="book" year="1943">Smith 1943</bibRefCitation>
;
<bibRefCitation id="EF90E6A0833DFFA5FC79FA66FB987E9F" author="CONSTABLE J. D." box="[950,1121,1491,1517]" pageId="32" pageNumber="485" pagination="59 - 160" refId="ref47769" refString="CONSTABLE J. D. 1949. - Reptiles from the Indian Peninsula in the Museum of Comparative Zoology. Bulletin of the Museum of Comparative Zoology 103: 59 - 160." type="journal article" year="1949">Constable 1949</bibRefCitation>
). Distinctive color-pattern, with a bluish-black dorsal coloration, bright-red venter with black mottling, and red streaks on the tail.
</paragraph>
</subSubSection>
<subSubSection id="C31BC8DA833DFFA5FCE2F9E6FBCC7C99" pageId="32" pageNumber="485" type="discussion">
<paragraph id="8BBE9B51833DFFA5FCE2F9E6FC687D19" blockId="32.[813,1458,1619,2027]" box="[813,913,1619,1645]" pageId="32" pageNumber="485">REMARKS</paragraph>
<paragraph id="8BBE9B51833DFFA5FCE2F9C7FBCC7C99" blockId="32.[813,1458,1619,2027]" pageId="32" pageNumber="485">
The original description (
<bibRefCitation id="EF90E6A0833DFFA5FB8DF9C7FB037DFF" author="BEDDOME R. H." box="[1090,1274,1650,1677]" pageId="32" pageNumber="485" pagination="41 - 48" refId="ref46637" refString="BEDDOME R. H. 1863 a. - Further notes upon the snakes of the Madras Presidency; with descriptions of new species. Madras Quarterly Journal of Medical Science 6: 41 - 48." type="journal article" year="1863">Beddome 1863a</bibRefCitation>
) reports the collection of several
<typeStatus id="54BA25F3833DFFA5FC22F926FBB47DDF" box="[1005,1101,1683,1709]" pageId="32" pageNumber="485" type="syntype">syntypes</typeStatus>
, but apparently only a few were catalogued, and only one is now known (
<bibRefCitation id="EF90E6A0833DFFA5FB27F907FA9F7DBF" author="GANS C." box="[1256,1382,1714,1741]" pageId="32" pageNumber="485" pagination="1 - 29" refId="ref48814" refString="GANS C. 1966. - Liste der rezenten Reptilien und Amphibien. Uropeltidae. Das Tierreich 84: 1 - 29." type="journal article" year="1966">Gans 1966</bibRefCitation>
).
<bibRefCitation id="EF90E6A0833DFFA5FAB7F906FC377D9E" author="BOULENGER G. A." pageId="32" pageNumber="485" refId="ref47173" refString="BOULENGER G. A. 1893 a. - Catalogue of the snakes in the British Museum (Natural History). Volume I., containing the families Typhlopidae, Glauconiidae, Boidae, Ilysiidae, Uropeltidae, Xenopeltidae and Colubridae aglyphae, part. British Museum (Natural History), London, 448 p." type="book" year="1893">Boulenger (1893a)</bibRefCitation>
reports a large topotypic series collected by Beddome but does not refer to them as types; either they were collected at a later date, or they werent catalogued as types initially. The lower ventral count reported by
<bibRefCitation id="EF90E6A0833DFFA5FAABF887FC837C1E" author="MASON G. E." pageId="32" pageNumber="485" pagination="184 - 186" refId="ref50518" refString="MASON G. E. 1888. - Description of a new earth-snake of the genus Silybura from the Bombay Presidency with remarks on little known Uropeltidae. Annals and Magazine of Natural History 1: 184 - 186." type="journal article" year="1888">Mason (1888)</bibRefCitation>
is of uncertain validity, and may represent a new or cryptic species from the Wullingy forests near Palghat. Records of this species exist from South of Palghat Gap (
<bibRefCitation id="EF90E6A0833DFFA5FCF8F804FC367CB9" author="HUTTON A. F." box="[823,975,1969,1995]" pageId="32" pageNumber="485" pagination="454 - 460" refId="ref49750" refString="HUTTON A. F. 1949. - Notes on the snakes and mammals of the High Wavy Mountains, Madura District, south India. Part I-Snakes. Journal of the Bombay Natural History Society 48: 454 - 460." type="journal article" year="1949">Hutton 1949</bibRefCitation>
), but we did not examine the specimens to determine their validity.
</paragraph>
</subSubSection>
</treatment>
</document>