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<mods:title id="5ECB0635AC889E5F04B78CACE44F5A0F">A catalogue and systematic overview of the shield-tailed snakes (Serpentes: Uropeltidae)</mods:title>
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<mods:namePart id="7D6708F63EB53E68404ACCE02F8CC5DF">Sharma, Vivek</mods:namePart>
<mods:affiliation id="69EF0A62F0A383D8551FE9D09B7CE902">Sanjeevni Nagar, Jabalpur, Madhya Pradesh 482 003 (India)</mods:affiliation>
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<mods:namePart id="95DB67BCD0D4A8E7B068D5F639BAD299">Wallach, Van</mods:namePart>
<mods:affiliation id="4C0E10558E5D7CAEE609ED8EEBBC1AF0">Potter Park, Cambridge, Massachusetts 02138 (United States)</mods:affiliation>
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<mods:namePart id="9822B2215C8B041DE6B5D461516A7ABE">Somaweera, Ruchira</mods:namePart>
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<heading id="D0F62C3D833AFFA2FEC9FF62FD7C7B80" box="[262,645,215,242]" centered="true" fontSize="9" level="2" pageId="39" pageNumber="492" reason="2">
<taxonomicName id="4C01E0D2833AFFA2FEC9FF62FD7C7B80" authority="(Beddome, 1867)" baseAuthorityName="Beddome" baseAuthorityYear="1867" box="[262,645,215,242]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="492" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B9754743833AFFA2FEC9FF62FE467B80" bold="true" box="[262,447,215,242]" italics="true" pageId="39" pageNumber="492">Uropeltis grandis</emphasis>
(
<bibRefCitation id="EF90E6A0833AFFA2FE02FF62FD847B80" author="BEDDOME R. H." box="[461,637,215,242]" pageId="39" pageNumber="492" pagination="14 - 16" refId="ref46715" refString="BEDDOME R. H. 1867. - Descriptions and figures of five new snakes from the Madras Presidency. Madras Quarterly Journal of Medical Science 11: 14 - 16." type="journal article" year="1867">Beddome, 1867</bibRefCitation>
)
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<taxonomicName id="4C01E0D2833AFFA2FF4BFE81FDFE7A3E" authority="Beddome, 1867: 15" authorityName="Beddome" authorityPageNumber="15" authorityYear="1867" box="[132,519,308,332]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="492" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B9754743833AFFA2FF4BFE81FEC87A3E" box="[132,305,308,332]" italics="true" pageId="39" pageNumber="492">Rhinophis grandis</emphasis>
<bibRefCitation id="EF90E6A0833AFFA2FEF6FE81FDFE7A3E" author="BEDDOME R. H." box="[313,519,308,332]" pageId="39" pageNumber="492" pagination="14 - 16" refId="ref46715" refString="BEDDOME R. H. 1867. - Descriptions and figures of five new snakes from the Madras Presidency. Madras Quarterly Journal of Medical Science 11: 14 - 16." type="journal article" year="1867">Beddome, 1867: 15</bibRefCitation>
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. Three
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: BMNH 1946.1.16.82 and MNHN-RA-1895.79 (two specimens). These three specimens are thus the
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of both
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<emphasis id="B9754743833AFFA2FD95FEDCFCFE7AF3" box="[602,775,361,385]" italics="true" pageId="39" pageNumber="492">Rhinophis grandis</emphasis>
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erected by
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, previously considered a
<emphasis id="B9754743833AFFA2FD5FFE31FF157AC7" italics="true" pageId="39" pageNumber="492">nomen praeoccupatum</emphasis>
by
<bibRefCitation id="EF90E6A0833AFFA2FEDBFE2BFE617AC7" author="GANS C." box="[276,408,413,438]" pageId="39" pageNumber="492" pagination="1 - 29" refId="ref48814" refString="GANS C. 1966. - Liste der rezenten Reptilien und Amphibien. Uropeltidae. Das Tierreich 84: 1 - 29." type="journal article" year="1966">Gans (1966)</bibRefCitation>
, and of
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<emphasis id="B9754743833AFFA2FE21FE28FD717AC7" box="[494,648,413,437]" italics="true" pageId="39" pageNumber="492">Uropeltis smithi</emphasis>
<bibRefCitation id="EF90E6A0833AFFA2FD5FFE2BFCFA7AC7" author="GANS C." box="[656,771,413,438]" pageId="39" pageNumber="492" pagination="1 - 29" refId="ref48814" refString="GANS C. 1966. - Liste der rezenten Reptilien und Amphibien. Uropeltidae. Das Tierreich 84: 1 - 29." type="journal article" year="1966">Gans 1966</bibRefCitation>
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. Type locality: Anaimalai hills, eastern
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/western
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,
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.
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<treatmentCitation id="0AA0BD40833AFFA2FF4BFDA6FE3F7959" author="GANS C." box="[132,454,531,555]" page="22" pageId="39" pageNumber="492" year="1966">
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<emphasis id="B9754743833AFFA2FF4BFDA6FEE47959" box="[132,285,531,555]" italics="true" pageId="39" pageNumber="492">Uropeltis smithi</emphasis>
<bibRefCitation id="EF90E6A0833AFFA2FEE9FDA6FE3F7959" author="GANS C." box="[294,454,531,555]" pageId="39" pageNumber="492" pagination="1 - 29" refId="ref48814" refString="GANS C. 1966. - Liste der rezenten Reptilien und Amphibien. Uropeltidae. Das Tierreich 84: 1 - 29." type="journal article" year="1966">Gans, 1966: 22</bibRefCitation>
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. Objective junior synonym (see Remarks).
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DISTRIBUTION. — This species has a relatively small range near the
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locality in the Anamalai hills of
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,
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(
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Wallach
<emphasis id="B9754743833AFFA2FF7DFD16FF1879C8" box="[178,225,674,698]" italics="true" pageId="39" pageNumber="492">et al.</emphasis>
2014
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;
<bibRefCitation id="EF90E6A0833AFFA2FEE9FD17FE5479C8" author="GANESH S. R." box="[294,429,674,698]" pageId="39" pageNumber="492" pagination="13 - 24" refId="ref48505" refString="GANESH S. R. 2015. - Shieldtail snakes (Reptilia: Uropeltidae. - the Darwin's finches of south Indian snake fauna?, in KANNAN, P. (Ed.) Manual on Identification and Preparation of Keys of Snakes with Special Reference to their Venomous Nature in India. Proceedings by Government Arts College, Udhagamandalam, Tamilnadu, India, 13 - 24." type="book chapter" year="2015">Ganesh 2015</bibRefCitation>
).
</paragraph>
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<paragraph id="8BBE9B51833AFFA2FF4BFD43FEEC787C" blockId="39.[130,778,758,1485]" box="[132,277,758,784]" pageId="39" pageNumber="492">DESCRIPTION</paragraph>
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Maximum total size
<emphasis id="B9754743833AFFA2FEBBFCA3FE7C785D" box="[372,389,790,815]" italics="true" pageId="39" pageNumber="492">c.</emphasis>
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, ventrals 190-218, subcaudals 6-12, dorsal scales in 19 rows at midbody (
<bibRefCitation id="EF90E6A0833AFFA2FD0EFC80FF38781D" author="SMITH M. A." pageId="39" pageNumber="492" refId="ref51495" refString="SMITH M. A. 1943. - The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. Vol. 3 (Serpentes). Taylor &amp; Francis, London, 583 p." type="book" year="1943">Smith 1943</bibRefCitation>
;
<bibRefCitation id="EF90E6A0833AFFA2FF03FCE0FE7B781D" author="CONSTABLE J. D." box="[204,386,853,879]" pageId="39" pageNumber="492" pagination="59 - 160" refId="ref47769" refString="CONSTABLE J. D. 1949. - Reptiles from the Indian Peninsula in the Museum of Comparative Zoology. Bulletin of the Museum of Comparative Zoology 103: 59 - 160." type="journal article" year="1949">Constable 1949</bibRefCitation>
). Snout is “Alternate Pointed” with “boomerang” rostral resembling
<taxonomicName id="4C01E0D2833AFFA2FE38FCC0FD9A78FD" authorityName="Hemprich" authorityYear="1820" box="[503,611,885,911]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="B9754743833AFFA2FE38FCC0FD9A78FD" box="[503,611,885,911]" italics="true" pageId="39" pageNumber="492">Rhinophis</emphasis>
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, to which the species was originally assigned, but does not divide the nasals in the specimens examined.
<bibRefCitation id="EF90E6A0833AFFA2FDC7FC00FD6678BD" author="SMITH M. A." box="[520,671,949,975]" pageId="39" pageNumber="492" refId="ref51495" refString="SMITH M. A. 1943. - The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. Vol. 3 (Serpentes). Taylor &amp; Francis, London, 583 p." type="book" year="1943">Smith (1943)</bibRefCitation>
classified the tail as state III, a group which contains a great deal of variation in observed tail morphology. In the
<typeStatus id="54BA25F3833AFFA2FDB0FC40FD277F7D" box="[639,734,1013,1039]" pageId="39" pageNumber="492" type="syntype">syntypes</typeStatus>
examined, the tail is compressed laterally and dorsally, and the expanded multicarinate scales encircle the entire tail. The terminal scute is enlarged, projecting, and rugose, ending in two larger points. We tentatively agree with Smiths classification (our Type III), but it is also possible that this is a
<taxonomicName id="4C01E0D2833AFFA2FF60FB01FEE27FBC" authorityName="Hemprich" authorityYear="1820" box="[175,283,1204,1230]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="B9754743833AFFA2FF60FB01FEE27FBC" box="[175,283,1204,1230]" italics="true" pageId="39" pageNumber="492">Rhinophis</emphasis>
</taxonomicName>
-like Type VI tail, masquerading as Type III. Distinctive color-pattern, with a dark violet dorsum and venter, and alternating yellow spots or crossbands on the sides of the body. The single specimen dissected lacks an intrapulmonary bronchus, unlike any other uropeltid specimen examined, and has a large number of tracheal rings (336), of which only
<taxonomicName id="4C01E0D2833AFFA2FE0DFAC1FDA77EFF" baseAuthorityName="Beddome" baseAuthorityYear="1886" box="[450,606,1395,1421]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="492" phylum="Chordata" rank="species" species="myhendrae">
<emphasis id="B9754743833AFFA2FE0DFAC1FDA77EFF" box="[450,606,1395,1421]" italics="true" pageId="39" pageNumber="492">U. myhendrae</emphasis>
</taxonomicName>
(353) also has more than 300 among the examined specimens of sampled uropeltid taxa.
</paragraph>
</subSubSection>
<subSubSection id="C31BC8DA833AFFA2FF4BFA46FECF7C99" pageId="39" pageNumber="492" type="discussion">
<paragraph id="8BBE9B51833AFFA2FF4BFA46FF117D79" blockId="39.[130,777,1523,2027]" box="[132,232,1523,1549]" pageId="39" pageNumber="492">REMARKS</paragraph>
<paragraph id="8BBE9B51833AFFA2FF4BF9A7FECF7C99" blockId="39.[130,777,1523,2027]" pageId="39" pageNumber="492">
This species bore a replacement name, having originally been described as
<taxonomicName id="4C01E0D2833AFFA2FE9CF987FDED7D3E" authorityName="Beddome" authorityYear="1867" box="[339,532,1586,1612]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="492" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B9754743833AFFA2FE9CF987FDED7D3E" box="[339,532,1586,1612]" italics="true" pageId="39" pageNumber="492">Rhinophis grandis</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF90E6A0833AFFA2FD8EF987FCFD7D3F" author="BEDDOME R. H." box="[577,772,1586,1613]" pageId="39" pageNumber="492" pagination="14 - 16" refId="ref46715" refString="BEDDOME R. H. 1867. - Descriptions and figures of five new snakes from the Madras Presidency. Madras Quarterly Journal of Medical Science 11: 14 - 16." type="journal article" year="1867">Beddome (1867)</bibRefCitation>
, creating an issue of homonymy when transferred to
<taxonomicName id="4C01E0D2833AFFA2FD1CF9E6FF437DFE" authorityName="Cuvier" authorityYear="1829" class="Reptilia" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="B9754743833AFFA2FD1CF9E6FF437DFE" italics="true" pageId="39" pageNumber="492">Uropeltis</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF90E6A0833AFFA2FF27F9C7FE7C7DFE" author="SMITH M. A." box="[232,389,1650,1676]" pageId="39" pageNumber="492" refId="ref51495" refString="SMITH M. A. 1943. - The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. Vol. 3 (Serpentes). Taylor &amp; Francis, London, 583 p." type="book" year="1943">Smith (1943)</bibRefCitation>
,
<taxonomicName id="4C01E0D2833AFFA2FE5CF9C6FD537DFF" authority="Kelaart, 1853" authorityName="Kelaart" authorityYear="1853" box="[403,682,1650,1677]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="492" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B9754743833AFFA2FE5CF9C6FDFF7DFE" box="[403,518,1650,1676]" italics="true" pageId="39" pageNumber="492">U. grandis</emphasis>
<bibRefCitation id="EF90E6A0833AFFA2FDC0F9C7FD537DFF" author="KELAART E. F." box="[527,682,1650,1677]" pageId="39" pageNumber="492" pagination="102 - 116" refId="ref49987" refString="KELAART E. F. 1853. - Description of new or little-known species of reptiles collected in Ceylon. Journal of the Ceylon Branch of the Royal Asiatic Society 22: 102 - 116." type="journal article" year="1853">Kelaart, 1853</bibRefCitation>
</taxonomicName>
being a pre-existing name, though it had already been relegated to the synonymy of
<taxonomicName id="4C01E0D2833AFFA2FEF4F907FDB67DBE" baseAuthorityName="Muller" baseAuthorityYear="1832" box="[315,591,1714,1740]" class="Reptilia" family="Uropeltidae" genus="Pseudotyphlops" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="492" phylum="Chordata" rank="species" species="philippinus">
<emphasis id="B9754743833AFFA2FEF4F907FDB67DBE" box="[315,591,1714,1740]" italics="true" pageId="39" pageNumber="492">Pseudotyphlops philippinus</emphasis>
</taxonomicName>
(now
<taxonomicName id="4C01E0D2833AFFA2FD5DF906FF567D9E" authorityName="Kelaart" authorityYear="1853" class="Reptilia" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="492" phylum="Chordata" rank="species" species="saffragamus">
<emphasis id="B9754743833AFFA2FD5DF906FF567D9E" italics="true" pageId="39" pageNumber="492">R. saffragamus</emphasis>
</taxonomicName>
). However, this species may actually be a
<taxonomicName id="4C01E0D2833AFFA2FD5BF967FCF97D9E" authorityName="Hemprich" authorityYear="1820" box="[660,768,1746,1772]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="B9754743833AFFA2FD5BF967FCF97D9E" box="[660,768,1746,1772]" italics="true" pageId="39" pageNumber="492">Rhinophis</emphasis>
</taxonomicName>
, as it bears
<emphasis id="B9754743833AFFA2FF30F947FE8C7C7E" box="[255,373,1778,1804]" italics="true" pageId="39" pageNumber="492">Rhinophis-</emphasis>
like rostral and shield morphology, in which case
<taxonomicName id="4C01E0D2833AFFA2FEEAF8A7FD8B7C5F" authority="(Gans 1966)" baseAuthorityName="Gans" baseAuthorityYear="1966" box="[293,626,1810,1837]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="492" phylum="Chordata" rank="species" species="smithi">
<emphasis id="B9754743833AFFA2FEEAF8A7FE257C5E" box="[293,476,1810,1836]" italics="true" pageId="39" pageNumber="492">Rhinophis smithi</emphasis>
(
<bibRefCitation id="EF90E6A0833AFFA2FE24F8A7FD937C5F" author="GANS C." box="[491,618,1810,1837]" pageId="39" pageNumber="492" pagination="1 - 29" refId="ref48814" refString="GANS C. 1966. - Liste der rezenten Reptilien und Amphibien. Uropeltidae. Das Tierreich 84: 1 - 29." type="journal article" year="1966">Gans 1966</bibRefCitation>
)
</taxonomicName>
is the proper name if transferred into that genus.
<bibRefCitation id="EF90E6A0833AFFA2FDC8F884FD3B7C3E" author="BEDDOME R. H." box="[519,706,1841,1868]" pageId="39" pageNumber="492" pagination="14 - 16" refId="ref46715" refString="BEDDOME R. H. 1867. - Descriptions and figures of five new snakes from the Madras Presidency. Madras Quarterly Journal of Medical Science 11: 14 - 16." type="journal article" year="1867">Beddome (1867)</bibRefCitation>
stated that it was clearly congeneric with
<taxonomicName id="4C01E0D2833AFFA2FDD9F8E7FD487C19" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[534,689,1873,1899]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="492" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B9754743833AFFA2FDD9F8E7FD487C19" box="[534,689,1873,1899]" italics="true" pageId="39" pageNumber="492">U. pulneyensis</emphasis>
</taxonomicName>
, which also has
<taxonomicName id="4C01E0D2833AFFA2FF2CF8C4FEB67CF9" authorityName="Hemprich" authorityYear="1820" box="[227,335,1905,1931]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="B9754743833AFFA2FF2CF8C4FEB67CF9" box="[227,335,1905,1931]" italics="true" pageId="39" pageNumber="492">Rhinophis</emphasis>
</taxonomicName>
-like rostral and tail-shield morphology. Further analyses using additional characters such as DNAsequence data will be needed to resolve the relationships of these species.
</paragraph>
</subSubSection>
</treatment>
</document>