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<document id="CCEC00BE8AA720C6F9D8EA9E0B1AD508" ID-CLB-Dataset="5666" ID-DOI="10.11646/zootaxa.3785.3.7" ID-GBIF-Dataset="8eddca4b-08a5-433a-8e3a-c89fb483818e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4913363" ID-ZooBank="D6E3D055-4390-4DC2-A9AC-0DBC40C2740F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623170138633" checkinUser="felipe" docAuthor="Neubauer, Thomas A., Harzhauser, Mathias, Georgopoulou, Elisavet, Mandic, Oleg &amp; Kroh, Andreas" docDate="2014" docId="03A7D746F079FFACFF3DFBB5FD8FFD6A" docLanguage="en" docName="zt03785p468.pdf" docOrigin="Zootaxa 3785 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Planorbis varians Fuchs 1870" docType="treatment" docVersion="7" lastPageNumber="465" masterDocId="FF9EAF3EF072FFA0FFAAFFE8FFBBFF9A" masterDocTitle="Replacement names and nomenclatural comments for problematic species-group names in Europe's Neogene freshwater Gastropoda" masterLastPageNumber="468" masterPageNumber="453" pageNumber="464" updateTime="1699005021158" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="061DB416387FA93BA715F66E70B48583">Replacement names and nomenclatural comments for problematic species-group names in Europe's Neogene freshwater Gastropoda</mods:title>
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<mods:namePart id="0008DE355017AEF08C459DD9F7263B3B">Neubauer, Thomas A.</mods:namePart>
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<mods:namePart id="7C9EC8F5B64FAB364FB6493CB1E75686">Harzhauser, Mathias</mods:namePart>
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<mods:namePart id="CA2C553735B85B78E1D2379FE41AC7A3">Georgopoulou, Elisavet</mods:namePart>
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<mods:namePart id="D2A0BF653239E66203B6E840EF9FABB1">Mandic, Oleg</mods:namePart>
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<treatment id="03A7D746F079FFACFF3DFBB5FD8FFD6A" ID-DOI="http://doi.org/10.5281/zenodo.5517085" ID-GBIF-Taxon="183399399" ID-Zenodo-Dep="5517085" LSID="urn:lsid:plazi:treatment:03A7D746F079FFACFF3DFBB5FD8FFD6A" httpUri="http://treatment.plazi.org/id/03A7D746F079FFACFF3DFBB5FD8FFD6A" lastPageId="12" lastPageNumber="465" pageId="11" pageNumber="464">
<subSubSection id="C31435DBF079FFABFF3DFBB5FCA8FBED" box="[151,787,1117,1143]" pageId="11" pageNumber="464" type="nomenclature">
<paragraph id="8BB16650F079FFABFF3DFBB5FCA8FBED" blockId="11.[151,787,1117,1143]" box="[151,787,1117,1143]" pageId="11" pageNumber="464">
<heading id="D0F9D13CF079FFABFF3DFBB5FCA8FBED" bold="true" box="[151,787,1117,1143]" fontSize="11" level="1" pageId="11" pageNumber="464" reason="1">
<emphasis id="B97ABA42F079FFABFF3DFBB5FCA8FBED" bold="true" box="[151,787,1117,1143]" pageId="11" pageNumber="464">
<taxonomicName id="4C0E1DD3F079FFABFF3DFBB5FDBCFBED" ID-CoL="4JL3R" authority="Fuchs, 1870" authorityName="Fuchs" authorityYear="1870" box="[151,519,1117,1143]" class="Gastropoda" family="Planorbidae" genus="Planorbis" kingdom="Animalia" order="Hygrophila" pageId="11" pageNumber="464" phylum="Mollusca" rank="species" species="varians">
<emphasis id="B97ABA42F079FFABFF3DFBB5FED6FBED" bold="true" box="[151,365,1117,1143]" italics="true" pageId="11" pageNumber="464">Planorbis varians</emphasis>
Fuchs, 1870
</taxonomicName>
—how to recombine?
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31435DBF079FFABFF3DFB76FB99F994" pageId="11" pageNumber="464" type="reference_group">
<paragraph id="8BB16650F079FFABFF3DFB76FCF4FB2E" blockId="11.[151,1339,1182,1296]" box="[151,847,1182,1204]" pageId="11" pageNumber="464">
<treatmentCitationGroup id="AB1E417EF079FFABFF3DFB76FCF4FB2E" box="[151,847,1182,1204]" pageId="11" pageNumber="464">
<taxonomicName id="4C0E1DD3F079FFABFF3DFB76FE74FB2E" ID-CoL="4JL3R" authority="Fuchs, 1870" authorityName="Fuchs" authorityYear="1870" box="[151,463,1182,1204]" class="Gastropoda" family="Planorbidae" genus="Planorbis" kingdom="Animalia" order="Hygrophila" pageId="11" pageNumber="464" phylum="Mollusca" rank="species" species="varians">
<emphasis id="B97ABA42F079FFABFF3DFB76FEF6FB2E" box="[151,333,1182,1204]" italics="true" pageId="11" pageNumber="464">Planorbis varians</emphasis>
Fuchs,
<quantity id="4CF6CBB5F079FFABFE36FB76FE74FB2E" box="[412,463,1182,1204]" metricMagnitude="1" metricUnit="m" metricValue="4.7498" pageId="11" pageNumber="464" unit="in" value="1870.0">1870</quantity>
</taxonomicName>
in
<treatmentCitation id="0AAF4041F079FFABFE45FB76FD1EFB2E" author="Fuchs, T." box="[495,677,1182,1204]" page="356" pageId="11" pageNumber="464" year="1870">
<bibRefCitation id="EF9F1BA1F079FFABFE45FB76FD1EFB2E" author="Fuchs, T." box="[495,677,1182,1204]" pageId="11" pageNumber="464" pagination="343 - 364" refId="ref9397" refString="Fuchs, T. (1870 a) III. Beitrage zur Kenntniss fossiler Binnenfaunen. III. Die Fauna der Congerienschichten von Radmanest im Banate. Jahrbuch der k. k. geologischen Reichsanstalt, 20, 343 - 364." type="journal article" year="1870">Fuchs 1870a: 356</bibRefCitation>
</treatmentCitation>
, pl. 14, figs 1–9.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB16650F079FFABFF3DFB55FCF2FB49" blockId="11.[151,1339,1182,1296]" box="[151,841,1213,1235]" pageId="11" pageNumber="464">
<treatmentCitationGroup id="AB1E417EF079FFABFF3DFB55FCF2FB49" box="[151,841,1213,1235]" pageId="11" pageNumber="464">
<taxonomicName id="4C0E1DD3F079FFABFF3DFB55FDAFFB49" ID-CoL="3HYFC" authority="(Fuchs)" baseAuthorityName="Fuchs" baseAuthorityYear="1870" box="[151,532,1213,1235]" class="Gastropoda" family="Planorbidae" genus="Gyraulus" kingdom="Animalia" order="Hygrophila" pageId="11" pageNumber="464" phylum="Mollusca" rank="species" species="varians" subGenus="Gyraulus">
<emphasis id="B97ABA42F079FFABFF3DFB55FE06FB49" box="[151,445,1213,1235]" italics="true" pageId="11" pageNumber="464">Gyraulus (Gyraulus) varians</emphasis>
(Fuchs)
</taxonomicName>
:
<treatmentCitation id="0AAF4041F079FFABFDB4FB56FD6BFB49" author="Wenz, W." box="[542,720,1213,1235]" page="1622" pageId="11" pageNumber="464" year="1923">
<bibRefCitation id="EF9F1BA1F079FFABFDB4FB56FD6BFB49" author="Wenz, W." box="[542,720,1213,1235]" pageId="11" pageNumber="464" pagination="3015 - 3387" refId="ref12109" refString="Wenz, W. (1923 - 1930) Fossilium Catalogus I: Animalia. Gastropoda extramarina tertiaria. W. Junk, Berlin, I: pp. 1 - 352 (1923), II: pp. 353 - 736 (1923), III: pp. 737 - 1068 (1923), IV: pp. 1069 - 1420 (1923), V: pp. 1421 - 1734 (1923), VI: pp. 1735 - 1862 (1923), VII: pp. 1863 - 2230 (1926), VIII: pp. 2231 - 2502 (1928), IX: pp. 2503 - 2886 (1929), X: pp. 2887 - 3014 (1929), XI: pp. 3015 - 3387 (1930)." type="journal article" year="1923">Wenz 1923: 1622</bibRefCitation>
</treatmentCitation>
(cum syn.).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB16650F079FFABFF3DFB34FC0EFA8A" blockId="11.[151,1339,1182,1296]" pageId="11" pageNumber="464">
<taxonomicName id="4C0E1DD3F079FFABFF3DFB34FDB9FB68" authority="(Fuchs, 1870)" baseAuthorityName="Fuchs" baseAuthorityYear="1870" box="[151,514,1244,1266]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="464" phylum="Mollusca" rank="species" species="varians">
<emphasis id="B97ABA42F079FFABFF3DFB34FED6FB6B" box="[151,365,1244,1266]" italics="true" pageId="11" pageNumber="464">Muellerpalia varians</emphasis>
(Fuchs, 1870)
</taxonomicName>
;
<bibRefCitation id="EF9F1BA1F079FFABFDA7FB34FD7AFB68" author="Bandel, K." box="[525,705,1244,1266]" pageId="11" pageNumber="464" pagination="91 - 155" refId="ref8379" refString="Bandel, K. (2010) Valvatiform Gastropoda (Heterostropha and Caenogastropoda) from the Paratethys Basin compared to living relatives, with description of several new genera and species. Freiberger Forschungshefte: Palaontologie, Stratigraphie, Fazies, C 536, 91 - 155." type="journal article" year="2010">Bandel 2010: 104</bibRefCitation>
[not pl. 8, figs 90–94, =
<taxonomicName id="4C0E1DD3F079FFABFC17FB34FA8AFB68" ID-CoL="44HW8" authority="(Fuchs, 1870)" baseAuthorityName="Fuchs" baseAuthorityYear="1870" box="[957,1329,1244,1266]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="464" phylum="Mollusca" rank="species" species="bicincta">
<emphasis id="B97ABA42F079FFABFC17FB34FB21FB68" box="[957,1178,1244,1266]" italics="true" pageId="11" pageNumber="464">Muellerpalia bicincta</emphasis>
(Fuchs, 1870)
</taxonomicName>
].
<emphasis id="B97ABA42F079FFABFF3DFB13FE89FA8A" box="[151,306,1275,1296]" italics="true" pageId="11" pageNumber="464">Marinescugyra</emphasis>
<taxonomicName id="4C0E1DD3F079FFABFE92FB13FDA2FA8A" authority="(Fuchs, 1870)" baseAuthorityName="Fuchs" baseAuthorityYear="1870" box="[312,537,1274,1296]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="464" phylum="Mollusca" rank="species" species="varians">
<emphasis id="B97ABA42F079FFABFE92FB13FE3FFA8A" box="[312,388,1275,1296]" italics="true" pageId="11" pageNumber="464">varians</emphasis>
(Fuchs, 1870)
</taxonomicName>
;
<bibRefCitation id="EF9F1BA1F079FFABFD89FB12FD63FA8A" author="Bandel, K." box="[547,728,1274,1296]" pageId="11" pageNumber="464" pagination="91 - 155" refId="ref8379" refString="Bandel, K. (2010) Valvatiform Gastropoda (Heterostropha and Caenogastropoda) from the Paratethys Basin compared to living relatives, with description of several new genera and species. Freiberger Forschungshefte: Palaontologie, Stratigraphie, Fazies, C 536, 91 - 155." type="journal article" year="2010">Bandel 2010: 112</bibRefCitation>
, pl. 13, figs 153–158.
</paragraph>
<paragraph id="8BB16650F079FFABFF3DFAD6FB99F994" blockId="11.[151,1436,1341,2018]" pageId="11" pageNumber="464">
In his revision about the valvatiform gastropods of the Paratethys realm,
<bibRefCitation id="EF9F1BA1F079FFABFC60FAD5FBD0FACC" author="Bandel, K." box="[970,1131,1341,1366]" pageId="11" pageNumber="464" pagination="91 - 155" refId="ref8379" refString="Bandel, K. (2010) Valvatiform Gastropoda (Heterostropha and Caenogastropoda) from the Paratethys Basin compared to living relatives, with description of several new genera and species. Freiberger Forschungshefte: Palaontologie, Stratigraphie, Fazies, C 536, 91 - 155." type="journal article" year="2010">Bandel (2010)</bibRefCitation>
introduced amongst others the genera
<taxonomicName id="4C0E1DD3F079FFABFEB4FA89FDE7FAE0" ID-CoL="7P9BC" authorityName="Bandel" authorityYear="2010" box="[286,604,1377,1402]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="464" phylum="Mollusca" rank="genus">
<emphasis id="B97ABA42F079FFABFEB4FA89FE0BFAE0" box="[286,432,1377,1402]" italics="true" pageId="11" pageNumber="464">Muellerpalia</emphasis>
(Hydrobiidae)
</taxonomicName>
and
<emphasis id="B97ABA42F079FFABFD0AFA8AFCF0FAE3" box="[672,843,1378,1401]" italics="true" pageId="11" pageNumber="464">Marinescugyra</emphasis>
(
<taxonomicName id="4C0E1DD3F079FFABFCCAFA89FC57FAE0" ID-CoL="7NKHX" authorityName="Rafinesque" authorityYear="1815" box="[864,1004,1377,1402]" class="Gastropoda" family="Planorbidae" kingdom="Animalia" order="Hygrophila" pageId="11" pageNumber="464" phylum="Mollusca" rank="family">Planorbidae</taxonomicName>
), based on characteristic protoconch features. One of the discussed species is
<taxonomicName id="4C0E1DD3F079FFABFDDDFA6FFC61FA3A" ID-CoL="4JL3R" authority="Fuchs, 1870" authorityName="Fuchs" authorityYear="1870" box="[631,986,1415,1440]" class="Gastropoda" family="Planorbidae" genus="Planorbis" kingdom="Animalia" order="Hygrophila" pageId="11" pageNumber="464" phylum="Mollusca" rank="species" species="varians">
<emphasis id="B97ABA42F079FFABFDDDFA6FFCFFFA05" box="[631,836,1415,1440]" italics="true" pageId="11" pageNumber="464">Planorbis varians</emphasis>
Fuchs, 1870
</taxonomicName>
from Rădmăneşti in
<collectingCountry id="F31926C0F079FFABFB7CFA6FFA84FA3A" box="[1238,1343,1415,1440]" name="Romania" pageId="11" pageNumber="464">Romania</collectingCountry>
. On the grounds of different protoconch properties, Bandel assigned specimens, putatively belonging to the same species, to either
<taxonomicName id="4C0E1DD3F079FFABFF53FA39FE30FA70" ID-CoL="7P9BC" authorityName="Bandel" authorityYear="2010" box="[249,395,1489,1514]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="464" phylum="Mollusca" rank="genus">
<emphasis id="B97ABA42F079FFABFF53FA39FE30FA70" box="[249,395,1489,1514]" italics="true" pageId="11" pageNumber="464">Muellerpalia</emphasis>
</taxonomicName>
(p. 104) or
<emphasis id="B97ABA42F079FFABFDA4FA3AFD02FA73" box="[526,697,1490,1513]" italics="true" pageId="11" pageNumber="464">Marinescugyra</emphasis>
(p. 112). For the latter, it even constitutes the
<typeStatus id="54B5D8F2F079FFABFB12FA3AFB53FA70" box="[1208,1256,1490,1514]" pageId="11" pageNumber="464">type</typeStatus>
species. Hence, Bandel used two different combinations based on the same original combination.
</paragraph>
</subSubSection>
<subSubSection id="C31435DBF079FFACFF6DF9F2FD8FFD6A" lastPageId="12" lastPageNumber="465" pageId="11" pageNumber="464" type="description">
<paragraph id="8BB16650F079FFABFF6DF9F2FE27F97F" blockId="11.[151,1436,1341,2018]" pageId="11" pageNumber="464">
From a nomenclatural as well as a taxonomic view this is of course not tenable. It means that either the various specimens belong to two different species or the diagnostic features of the two genera are not meaningful. The associated problems were clear to the author as became evident from personal correspondence: Bandel (pers. comm.) stated the possibility that &quot;
<taxonomicName id="4C0E1DD3F079FFABFDB5F96DFD52F907" authorityName="Fuchs" authorityYear="1870" box="[543,745,1669,1694]" class="Gastropoda" family="Planorbidae" genus="Planorbis" kingdom="Animalia" order="Hygrophila" pageId="11" pageNumber="464" phylum="Mollusca" rank="species" species="varians">
<emphasis id="B97ABA42F079FFABFDB5F96DFD52F907" box="[543,745,1669,1694]" italics="true" pageId="11" pageNumber="464">Planorbis varians</emphasis>
</taxonomicName>
&quot; is actually represented by two species, which would have to be assigned variably to
<taxonomicName id="4C0E1DD3F079FFABFE09F941FD8EF958" authorityName="Bandel" authorityYear="2010" box="[419,565,1705,1730]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="464" phylum="Mollusca" rank="genus">
<emphasis id="B97ABA42F079FFABFE09F941FD8EF958" box="[419,565,1705,1730]" italics="true" pageId="11" pageNumber="464">Muellerpalia</emphasis>
</taxonomicName>
or
<emphasis id="B97ABA42F079FFABFDF6F942FCBCF95B" box="[604,775,1706,1729]" italics="true" pageId="11" pageNumber="464">Marinescugyra</emphasis>
based on their protoconchs. Unfortunately, this discussion is wanting in the paper.
</paragraph>
<paragraph id="8BB16650F079FFACFF6DF91AFE90FEFF" blockId="11.[151,1436,1341,2018]" lastBlockId="12.[151,1437,151,752]" lastPageId="12" lastPageNumber="465" pageId="11" pageNumber="464">
However, it turned out that the two combinations indeed refer to different species. The specimens illustrated by
<bibRefCitation id="EF9F1BA1F079FFABFF3DF8FDFE85F8B4" author="Bandel, K." box="[151,318,1813,1838]" pageId="11" pageNumber="464" pagination="91 - 155" refId="ref8379" refString="Bandel, K. (2010) Valvatiform Gastropoda (Heterostropha and Caenogastropoda) from the Paratethys Basin compared to living relatives, with description of several new genera and species. Freiberger Forschungshefte: Palaontologie, Stratigraphie, Fazies, C 536, 91 - 155." type="journal article" year="2010">Bandel (2010)</bibRefCitation>
as &quot;
<emphasis id="B97ABA42F079FFABFED1F8FEFD3CF8B7" box="[379,647,1814,1837]" italics="true" pageId="11" pageNumber="464">
Marinescugyra
<taxonomicName id="4C0E1DD3F079FFABFD99F8FEFD3CF8B7" authorityName="Fuchs" authorityYear="1870" box="[563,647,1814,1837]" class="Gastropoda" family="Planorbidae" genus="Planorbis" kingdom="Animalia" order="Hygrophila" pageId="11" pageNumber="464" phylum="Mollusca" rank="species" species="varians">varians</taxonomicName>
</emphasis>
&quot; (pl. 13, figs 153–158) undoubtedly refer to the same species as described by
<bibRefCitation id="EF9F1BA1F079FFABFE98F8D1FE62F8C8" author="Fuchs, T." box="[306,473,1849,1874]" pageId="11" pageNumber="464" pagination="343 - 364" refId="ref9397" refString="Fuchs, T. (1870 a) III. Beitrage zur Kenntniss fossiler Binnenfaunen. III. Die Fauna der Congerienschichten von Radmanest im Banate. Jahrbuch der k. k. geologischen Reichsanstalt, 20, 343 - 364." type="journal article" year="1870">Fuchs (1870a)</bibRefCitation>
. They correspond in the two strong keels, the intermediate spiral lines, the slightly concave umbilical part, and the weakly stepped funnel-like apical part. In contrast, the individuals depicted as &quot;
<taxonomicName id="4C0E1DD3F079FFABFF08F869FE2BF803" baseAuthorityName="Fuchs" baseAuthorityYear="1870" box="[162,400,1921,1946]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="464" phylum="Mollusca" rank="species" species="varians">
<emphasis id="B97ABA42F079FFABFF08F869FE2BF803" box="[162,400,1921,1946]" italics="true" pageId="11" pageNumber="464">Muellerpalia varians</emphasis>
</taxonomicName>
&quot; (
<bibRefCitation id="EF9F1BA1F079FFABFE01F869FD84F800" author="Bandel, K." box="[427,575,1921,1946]" pageId="11" pageNumber="464" pagination="91 - 155" refId="ref8379" refString="Bandel, K. (2010) Valvatiform Gastropoda (Heterostropha and Caenogastropoda) from the Paratethys Basin compared to living relatives, with description of several new genera and species. Freiberger Forschungshefte: Palaontologie, Stratigraphie, Fazies, C 536, 91 - 155." type="journal article" year="2010">Bandel 2010</bibRefCitation>
, pl. 8, figs 90–95) belong to a different species: they have both keels on the umbilical side, apparently lack additional spiral lines, have highly convex whorls, and more distinct growth lines. Direct comparison of the descriptions and illustrations of
<bibRefCitation id="EF9F1BA1F079FFABFC8BF821FC07F878" author="Fuchs, T." box="[801,956,1993,2018]" pageId="11" pageNumber="464" pagination="343 - 364" refId="ref9397" refString="Fuchs, T. (1870 a) III. Beitrage zur Kenntniss fossiler Binnenfaunen. III. Die Fauna der Congerienschichten von Radmanest im Banate. Jahrbuch der k. k. geologischen Reichsanstalt, 20, 343 - 364." type="journal article" year="1870">Fuchs (1870a</bibRefCitation>
, b) and
<bibRefCitation id="EF9F1BA1F079FFABFBBCF821FB03F878" author="Bandel, K." box="[1046,1208,1993,2018]" pageId="11" pageNumber="464" pagination="91 - 155" refId="ref8379" refString="Bandel, K. (2010) Valvatiform Gastropoda (Heterostropha and Caenogastropoda) from the Paratethys Basin compared to living relatives, with description of several new genera and species. Freiberger Forschungshefte: Palaontologie, Stratigraphie, Fazies, C 536, 91 - 155." type="journal article" year="2010">Bandel (2010)</bibRefCitation>
suggest that Bandel confused several species. Firstly, &quot;
<taxonomicName id="4C0E1DD3F07EFFACFD80FF7FFC70FF2A" authority="&quot; sensu Bandel" authorityName="Bandel" box="[554,971,151,176]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="varians">
<emphasis id="B97ABA42F07EFFACFD80FF7FFCA7FF35" box="[554,796,151,176]" italics="true" pageId="12" pageNumber="465">Muellerpalia varians</emphasis>
&quot; sensu Bandel
</taxonomicName>
rather corresponds to &quot;
<taxonomicName id="4C0E1DD3F07EFFACFB48FF7FFE92FF4F" authority="Fuchs, 1870" authorityName="Fuchs" authorityYear="1870" class="Gastropoda" family="Valvatidae" genus="Valvata" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="bicincta">
<emphasis id="B97ABA42F07EFFACFB48FF7FFA27FF2A" box="[1250,1436,151,176]" italics="true" pageId="12" pageNumber="465">Valvata bicincta</emphasis>
Fuchs, 1870
</taxonomicName>
&quot; in
<bibRefCitation id="EF9F1BA1F07EFFACFEC9FF54FDB6FF4F" author="Fuchs, T." box="[355,525,188,213]" pageId="12" pageNumber="465" pagination="531 - 548" refId="ref9439" refString="Fuchs, T. (1870 b) VII. Beitrage zur Kenntniss fossiler Binnenfaunen. IV. und V. Die Fanna der Congerienschichten von Tihany am Plattensee und Kup bei Papa in Ungarn. Jahrbuch der k. k. geologischen Reichsanstalt, 20, 531 - 548." type="journal article" year="1870">Fuchs (1870b)</bibRefCitation>
[=
<taxonomicName id="4C0E1DD3F07EFFACFDEBFF54FC4AFF4F" authority="(Fuchs, 1870)" baseAuthorityName="Fuchs" baseAuthorityYear="1870" box="[577,1009,188,213]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="bicincta">
<emphasis id="B97ABA42F07EFFACFDEBFF54FC80FF4F" box="[577,827,188,213]" italics="true" pageId="12" pageNumber="465">Muellerpalia bicincta</emphasis>
(Fuchs, 1870)
</taxonomicName>
]. Secondly, specimens illustrated as &quot;
<taxonomicName id="4C0E1DD3F07EFFACFF08FF37FE24FF62" baseAuthorityName="Fuchs" baseAuthorityYear="1870" box="[162,415,223,248]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="bicincta">
<emphasis id="B97ABA42F07EFFACFF08FF37FE24FF62" box="[162,415,223,248]" italics="true" pageId="12" pageNumber="465">Muellerpalia bicincta</emphasis>
</taxonomicName>
&quot; by Bandel rather resemble &quot;
<taxonomicName id="4C0E1DD3F07EFFACFCB6FF37FB3DFF62" authority="Fuchs, 1870" authorityName="Fuchs" authorityYear="1870" box="[796,1158,223,248]" class="Gastropoda" family="Valvatidae" genus="Valvata" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="carinata">
<emphasis id="B97ABA42F07EFFACFCB6FF37FC5BFF6D" box="[796,992,223,248]" italics="true" pageId="12" pageNumber="465">Valvata carinata</emphasis>
Fuchs, 1870
</taxonomicName>
&quot; in
<bibRefCitation id="EF9F1BA1F07EFFACFB6CFF37FAC9FF62" author="Fuchs, T." box="[1222,1394,223,248]" pageId="12" pageNumber="465" pagination="531 - 548" refId="ref9439" refString="Fuchs, T. (1870 b) VII. Beitrage zur Kenntniss fossiler Binnenfaunen. IV. und V. Die Fanna der Congerienschichten von Tihany am Plattensee und Kup bei Papa in Ungarn. Jahrbuch der k. k. geologischen Reichsanstalt, 20, 531 - 548." type="journal article" year="1870">Fuchs (1870b)</bibRefCitation>
[=
<taxonomicName id="4C0E1DD3F07EFFACFF3DFEECFD87FE87" authority="(Fuchs, 1870)" authorityName="sensu Bandel" authorityYear="2010" baseAuthorityName="Fuchs" baseAuthorityYear="1870" box="[151,572,260,285]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="carinata">
<emphasis id="B97ABA42F07EFFACFF3DFEECFE2AFE86" box="[151,401,260,285]" italics="true" pageId="12" pageNumber="465">Muellerpalia carinata</emphasis>
(Fuchs, 1870)
</taxonomicName>
]. Conversely,
<taxonomicName id="4C0E1DD3F07EFFACFD77FEECFAD8FE87" authority="(Fuchs, 1870) sensu Bandel, 2010" authorityName="sensu Bandel" authorityYear="2010" baseAuthorityName="Fuchs" baseAuthorityYear="1870" box="[733,1379,260,285]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="carinata">
<emphasis id="B97ABA42F07EFFACFD77FEECFC6CFE86" box="[733,983,260,285]" italics="true" pageId="12" pageNumber="465">Muellerpalia carinata</emphasis>
(Fuchs, 1870) sensu
<bibRefCitation id="EF9F1BA1F07EFFACFB66FEECFAD8FE87" author="Bandel, K." box="[1228,1379,260,285]" pageId="12" pageNumber="465" pagination="91 - 155" refId="ref8379" refString="Bandel, K. (2010) Valvatiform Gastropoda (Heterostropha and Caenogastropoda) from the Paratethys Basin compared to living relatives, with description of several new genera and species. Freiberger Forschungshefte: Palaontologie, Stratigraphie, Fazies, C 536, 91 - 155." type="journal article" year="2010">Bandel, 2010</bibRefCitation>
</taxonomicName>
may represent
<taxonomicName id="4C0E1DD3F07EFFACFEACFECFFE47FEDA" baseAuthorityName="Fuchs" baseAuthorityYear="1870" box="[262,508,295,320]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="bicincta">
<emphasis id="B97ABA42F07EFFACFEACFECFFE47FEDA" box="[262,508,295,320]" italics="true" pageId="12" pageNumber="465">Muellerpalia bicincta</emphasis>
</taxonomicName>
. The new combinations of both latter taxa with the new genus
<taxonomicName id="4C0E1DD3F07EFFACFB7CFECFFAD2FEDA" authorityName="Bandel" authorityYear="2010" box="[1238,1385,295,320]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="465" phylum="Mollusca" rank="genus">
<emphasis id="B97ABA42F07EFFACFB7CFECFFAD2FEDA" box="[1238,1385,295,320]" italics="true" pageId="12" pageNumber="465">Muellerpalia</emphasis>
</taxonomicName>
still remain valid.
</paragraph>
<paragraph id="8BB16650F07EFFACFF6DFE87FF49FDC5" blockId="12.[151,1437,151,752]" pageId="12" pageNumber="465">
The presence of a second, fainter keel on the umbilical side in &quot;
<taxonomicName id="4C0E1DD3F07EFFACFC3EFE87FAC0FE12" authority="&quot; sensu Bandel (2010)" authorityName="Bandel" authorityYear="2010" box="[916,1403,367,392]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="varians">
<emphasis id="B97ABA42F07EFFACFC3EFE87FB3AFE1D" box="[916,1153,367,392]" italics="true" pageId="12" pageNumber="465">Muellerpalia varians</emphasis>
&quot; sensu
<bibRefCitation id="EF9F1BA1F07EFFACFB70FE87FAC0FE12" author="Bandel, K." box="[1242,1403,367,392]" pageId="12" pageNumber="465" pagination="91 - 155" refId="ref8379" refString="Bandel, K. (2010) Valvatiform Gastropoda (Heterostropha and Caenogastropoda) from the Paratethys Basin compared to living relatives, with description of several new genera and species. Freiberger Forschungshefte: Palaontologie, Stratigraphie, Fazies, C 536, 91 - 155." type="journal article" year="2010">Bandel (2010)</bibRefCitation>
</taxonomicName>
[=
<taxonomicName id="4C0E1DD3F07EFFACFF3DFE7CFD94FE37" authority="(Fuchs, 1870)" baseAuthorityName="Fuchs" baseAuthorityYear="1870" box="[151,559,404,429]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="bicincta">
<emphasis id="B97ABA42F07EFFACFF3DFE7CFE32FE37" box="[151,393,404,429]" italics="true" pageId="12" pageNumber="465">Muellerpalia bicincta</emphasis>
(Fuchs, 1870)
</taxonomicName>
], which is not discernible in the original material of
<bibRefCitation id="EF9F1BA1F07EFFACFBD9FE7CFAADFE37" author="Fuchs, T." box="[1139,1302,404,429]" pageId="12" pageNumber="465" pagination="343 - 364" refId="ref9397" refString="Fuchs, T. (1870 a) III. Beitrage zur Kenntniss fossiler Binnenfaunen. III. Die Fauna der Congerienschichten von Radmanest im Banate. Jahrbuch der k. k. geologischen Reichsanstalt, 20, 343 - 364." type="journal article" year="1870">Fuchs (1870a)</bibRefCitation>
, is regarded as intraspecific variability herein, as all other features correspond quite well. Unfortunately, Bandel did not provide an illustration of the apical region, which would show a distinct keel if our determination as
<taxonomicName id="4C0E1DD3F07EFFACFB0CFE35FA9DFE6F" baseAuthorityName="Fuchs" baseAuthorityYear="1870" box="[1190,1318,476,501]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="bicincta">
<emphasis id="B97ABA42F07EFFACFB0CFE35FA9DFE6F" box="[1190,1318,476,501]" italics="true" pageId="12" pageNumber="465">M. bicincta</emphasis>
</taxonomicName>
is correct. Nevertheless, the endpoint of keel-like structure is visible in the apertural view (
<bibRefCitation id="EF9F1BA1F07EFFACFB8AFE17FB0FFD82" author="Bandel, K." box="[1056,1204,511,536]" pageId="12" pageNumber="465" pagination="91 - 155" refId="ref8379" refString="Bandel, K. (2010) Valvatiform Gastropoda (Heterostropha and Caenogastropoda) from the Paratethys Basin compared to living relatives, with description of several new genera and species. Freiberger Forschungshefte: Palaontologie, Stratigraphie, Fazies, C 536, 91 - 155." type="journal article" year="2010">Bandel 2010</bibRefCitation>
, pl. 8, fig. 94). Even if it might turn out that this tentative placement is wrong, it still remains well separated from
<emphasis id="B97ABA42F07EFFACFB5BFDCDFF50FDC5" italics="true" pageId="12" pageNumber="465">
Marinescugyra
<taxonomicName id="4C0E1DD3F07EFFACFF3DFDA0FF50FDC5" box="[151,235,584,607]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="varians">varians</taxonomicName>
</emphasis>
.
</paragraph>
<paragraph id="8BB16650F07EFFACFF6DFD84FD8FFD6A" blockId="12.[151,1437,151,752]" pageId="12" pageNumber="465">
In conclusion, the genus
<emphasis id="B97ABA42F07EFFACFE4DFD85FD29FD1E" box="[487,658,621,644]" italics="true" pageId="12" pageNumber="465">Marinescugyra</emphasis>
, with
<taxonomicName id="4C0E1DD3F07EFFACFD77FD84FB86FD1F" authority="Fuchs, 1870" authorityName="Fuchs" authorityYear="1870" box="[733,1085,620,645]" class="Gastropoda" family="Planorbidae" genus="Planorbis" kingdom="Animalia" order="Hygrophila" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="varians">
<emphasis id="B97ABA42F07EFFACFD77FD84FC12FD1E" box="[733,937,620,645]" italics="true" pageId="12" pageNumber="465">Planorbis varians</emphasis>
Fuchs, 1870
</taxonomicName>
as
<typeStatus id="54B5D8F2F07EFFACFBCDFD84FB2CFD1E" box="[1127,1175,620,644]" pageId="12" pageNumber="465">type</typeStatus>
species, remains valid and the combination
<emphasis id="B97ABA42F07EFFACFE2AFD78FD90FD3D" box="[384,555,656,679]" italics="true" pageId="12" pageNumber="465">Marinescugyra</emphasis>
<taxonomicName id="4C0E1DD3F07EFFACFD98FD78FC9CFD32" authority="(Fuchs, 1870)" baseAuthorityName="Fuchs" baseAuthorityYear="1870" box="[562,807,655,680]" class="Gastropoda" family="Planorbidae" genus="Planorbis" kingdom="Animalia" order="Hygrophila" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="varians">
<emphasis id="B97ABA42F07EFFACFD98FD78FD3DFD3D" box="[562,646,656,679]" italics="true" pageId="12" pageNumber="465">varians</emphasis>
(Fuchs, 1870)
</taxonomicName>
is the only accepted one. The combination
<taxonomicName id="4C0E1DD3F07EFFACFAA0FD67FE2AFD57" authority="(Fuchs, 1870)" baseAuthorityName="Fuchs" baseAuthorityYear="1870" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="varians">
<emphasis id="B97ABA42F07EFFACFAA0FD67FF50FD56" italics="true" pageId="12" pageNumber="465">Muellerpalia varians</emphasis>
(Fuchs, 1870)
</taxonomicName>
is based on a misidentification of the species. We tentatively synonymize this record with
<taxonomicName id="4C0E1DD3F07EFFACFF3DFD3FFD8BFD6A" authority="(Fuchs, 1870)" baseAuthorityName="Fuchs" baseAuthorityYear="1870" box="[151,560,727,752]" class="Gastropoda" family="Hydrobiidae" genus="Muellerpalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="465" phylum="Mollusca" rank="species" species="bicincta">
<emphasis id="B97ABA42F07EFFACFF3DFD3FFE31FD6A" box="[151,394,727,752]" italics="true" pageId="12" pageNumber="465">Muellerpalia bicincta</emphasis>
(Fuchs, 1870)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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