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<document id="BACA2A1D902BEC32809502A76F85B1AA" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFEAE51E531A87FB61C8F895" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) densissimus Assing 2017, spec. nov." docType="treatment" docUuid="C82B20D3-1259-446E-B077-0D026B057901" docUuidSource="ZooBank" docVersion="14" lastPageNumber="161" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="160" updateTime="1699471407760" updateUser="ExternalLinkService">
<mods:mods id="B8743AB0BA6E85658CF7D96C1C9E536F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CAB4216CC5AA9ED85B5B9B888BF3E4B9">
<mods:title id="451C55046FA5110166FE979D3E97AB3C">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:role id="6AF3B7B22CEB03B95A5388BD723AE320">
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</mods:role>
<mods:namePart id="1C1C70B33CAD5AE67D95C89459FFCCBB">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="A1E9AE8BC5629389DD865AD69108F5BB">text</mods:typeOfResource>
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<mods:title id="C8742D0E90D276FACBC283C97BD2F12F">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="4A14561ADB78D34E040ACF6B2FDE04C6">
<mods:date id="EF584165DA5D77D3990CAB5C9F37D757">2017</mods:date>
<mods:detail id="B01E92407AA49BCD924E246FC00B1E55" type="pubDate">
<mods:number id="D88894DEFDB2113D2BB5F85E815F8718">2017-06-30</mods:number>
</mods:detail>
<mods:detail id="FCB8DFBA439887E894FC5F3E5BA43658" type="volume">
<mods:number id="BE5338F25E8D008283A0D01D630507EF">67</mods:number>
</mods:detail>
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<mods:number id="86534FE1EF851F56C961A7E2F06A9B7D">1</mods:number>
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<mods:start id="20B61CBF17348E0E1FF4551780E1559F">117</mods:start>
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<mods:classification id="89AB74DB7E34114196DD000D87572B38">journal article</mods:classification>
<mods:identifier id="5284C8AE25D62F8C6773D4B7BA06D167" type="CLB-Dataset">2472</mods:identifier>
<mods:identifier id="3DEE8F717ECA67EB1DA8864D7CE68CB6" type="DOI">10.21248/contrib.entomol.67.1.117-192</mods:identifier>
<mods:identifier id="A946C451B8B89F5697BBD90ED0E136DF" type="GBIF-Dataset">39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43</mods:identifier>
<mods:identifier id="338270B92F699EF699CA597CCE63E141" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="31CCE6BD6DBC9286419DABD13DB69B0C" type="Zenodo-Dep">5742363</mods:identifier>
<mods:identifier id="A7856D4240EFA7417FF227DF95FD77A7" type="ZooBank">FD33C1AE-F7D9-4E3A-A053-A2CAA7261CFE</mods:identifier>
</mods:mods>
<treatment id="03A787BAFFEAE51E531A87FB61C8F895" ID-DOI="http://doi.org/10.5281/zenodo.5888493" ID-GBIF-Taxon="192498151" ID-Zenodo-Dep="5888493" LSID="urn:lsid:zoobank.org:act:C82B20D3-1259-446E-B077-0D026B057901" httpUri="http://treatment.plazi.org/id/03A787BAFFEAE51E531A87FB61C8F895" lastPageId="44" lastPageNumber="161" pageId="43" pageNumber="160">
<subSubSection id="C3146527FFEAE519531A87FB67BFFEBA" box="[831,1204,243,270]" pageId="43" pageNumber="160" type="nomenclature">
<paragraph id="8BB136ACFFEAE519531A87FB67BFFEBA" blockId="43.[831,1349,243,335]" box="[831,1204,243,270]" pageId="43" pageNumber="160">
<heading id="D0F981C0FFEAE519531A87FB67BFFEBA" box="[831,1204,243,270]" fontSize="12" level="2" pageId="43" pageNumber="160" reason="2">
<taxonomicName id="4C0E4D2FFFEAE519531A87FB674FFEBA" ID-CoL="5DDBS" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[831,1092,243,270]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="160" phylum="Arthropoda" rank="species" species="densissimus" status="sp. nov." subGenus="Zyras">
<emphasis id="B97AEABEFFEAE519531A87FB607EFEBA" box="[831,885,243,270]" italics="true" pageId="43" pageNumber="160">Zyras</emphasis>
(
<emphasis id="B97AEABEFFEAE51953A687FB60B2FEBA" box="[899,953,243,270]" italics="true" pageId="43" pageNumber="160">Zyras</emphasis>
)
<emphasis id="B97AEABEFFEAE51953E287FB674FFEBA" box="[967,1092,243,270]" italics="true" pageId="43" pageNumber="160">densissimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957C5FFEAE519546E87FB67BFFEBA" box="[1099,1204,243,270]" pageId="43" pageNumber="160" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEAE519531A8613664EFE9B" box="[831,1349,283,303]" pageId="43" pageNumber="160" type="description">
<paragraph id="8BB136ACFFEAE519531A8613664EFE9B" blockId="43.[831,1349,243,335]" box="[831,1349,283,303]" pageId="43" pageNumber="160">
<uri id="FF9F3AAEFFEAE519531A8613664EFE9B" box="[831,1349,283,303]" pageId="43" pageNumber="160">
urn:lsid:zoobank.org:act:
<uuid id="FFA80C79FFEAE51953D48613664EFE9B" box="[1009,1349,283,303]" pageId="43" pageNumber="160">C82B20D3-1259-446E-B077-0D026B057901</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEAE519531A8633675BFEFB" box="[831,1104,315,335]" pageId="43" pageNumber="160" type="description">
<paragraph id="8BB136ACFFEAE519531A8633675BFEFB" blockId="43.[831,1349,243,335]" box="[831,1104,315,335]" pageId="43" pageNumber="160">
(
<figureCitation id="13352A29FFEAE519536686336076FEFB" box="[835,893,315,335]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="43" pageNumber="160">Figs 36</figureCitation>
,
<figureCitation id="13352A29FFEAE51953A686336093FEFB" box="[899,920,315,335]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="43" pageNumber="160">75</figureCitation>
,
<figureCitation id="13352A29FFEAE51953BA863360B6FEFB" box="[927,957,315,335]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="43" pageNumber="160">111</figureCitation>
,
<figureCitation id="13352A29FFEAE51953E086336701FEFB" box="[965,1034,315,335]" captionStart="Figs 227–244" captionStartId="61.[151,191,1983,2005]" captionTargetBox="[163,1397,240,1949]" captionTargetId="figure-119@61.[151,1398,231,1970]" captionText="Figs 227–244: Zyras parahirtus (227–228, 231–232), Z. matangensis (229–230, 233), Z. gratellus (Sulawesi) (234–238), and Z. densissimus (239–244): median lobe of aedeagus in lateral and in ventral view (227–230, 234–235, 239–241); paramere (231, 236, 242); male tergite VIII (232, 237, 243); male sternite VIII (233, 238, 244). Scale bars: 227–233, 237–238, 243–244: 0.5 mm; 234–236, 239–242: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888378" httpUri="https://zenodo.org/record/5888378/files/figure.png" pageId="43" pageNumber="160">239–244</figureCitation>
,
<figureCitation id="13352A29FFEAE519543586336740FEFB" box="[1040,1099,315,335]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="43" pageNumber="160">Map 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEAE519533F86616088FD15" pageId="43" pageNumber="160" type="materials_examined">
<paragraph id="8BB136ACFFEAE519533F86616088FD15" blockId="43.[794,1399,361,673]" pageId="43" pageNumber="160">
<emphasis id="B97AEABEFFEAE519533F866160B9FE35" bold="true" box="[794,946,361,385]" pageId="43" pageNumber="160">Type material</emphasis>
:
<materialsCitation id="3B663CF1FFEAE519539A86616074FD95" ID-GBIF-Occurrence="3450162302" collectingDate="1985-02-24" collectingMethod="Flight interception trap" collectionCode="BMNH" country="Indonesia" determinerName="V. Assing" elevation="200" location="Dumoga-Bone N. P." municipality="Lowland forest" pageId="43" pageNumber="160" specimenCount="1" stateProvince="SULAWESI UTARA" typeStatus="holotype">
<typeStatus id="54B5880EFFEAE519539A86616729FE35" box="[959,1058,361,385]" pageId="43" pageNumber="160" type="holotype">Holotype</typeStatus>
<specimenCount id="9D08FD25FFEAE519540F86626734FE35" box="[1066,1087,362,385]" pageId="43" pageNumber="160">♂</specimenCount>
: “
<collectingCountry id="F319763CFFEAE5195470866167EEFE35" box="[1109,1253,361,385]" name="Indonesia" pageId="43" pageNumber="160">INDONESIA</collectingCountry>
:
<collectingRegion id="49CAF84EFFEAE51954DF8661607FFE15" country="Indonesia" name="Sulawesi Utara" pageId="43" pageNumber="160">SULAWESI UTARA</collectingRegion>
,
<location id="8ED16077FFEAE519535A86826758FE15" LSID="urn:lsid:plazi:treatment:03A787BAFFEAE51E531A87FB61C8F895:8ED16077FFEAE519535A86826758FE15" box="[895,1107,394,418]" country="Indonesia" municipality="Lowland forest" name="Dumoga-Bone N. P." pageId="43" pageNumber="160" stateProvince="SULAWESI UTARA">Dumoga-Bone N.P.</location>
,
<date id="FFB0106CFFEAE519547B8681661AFE15" box="[1118,1297,393,417]" pageId="43" pageNumber="160" value="1985-02-24">
<collectingDate id="EFF4E984FFEAE519547B8681661AFE15" box="[1118,1297,393,417]" pageId="43" pageNumber="160" value="1985-02-24">24 Februar 1985</collectingDate>
</date>
. / Plot A, ca
<quantity id="4CF69B49FFEAE519531E86A16074FE75" box="[827,895,425,449]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="43" pageNumber="160" unit="m" value="200.0">
<elevation id="0023D19FFFEAE519531E86A16074FE75" box="[827,895,425,449]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="43" pageNumber="160" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<location id="8ED16077FFEAE51953A886A1673BFE75" LSID="urn:lsid:plazi:treatment:03A787BAFFEAE51E531A87FB61C8F895:8ED16077FFEAE51953A886A1673BFE75" box="[909,1072,425,449]" country="Indonesia" municipality="Lowland forest" name="Lowland forest" pageId="43" pageNumber="160" stateProvince="SULAWESI UTARA">Lowland forest</location>
/
<collectingMethod id="524F4EBBFFEAE519546886A16647FE75" box="[1101,1356,425,449]" pageId="43" pageNumber="160">Flight interception trap</collectingMethod>
2 / R
<location id="8ED16077FFEAE519530886C160C4FE55" LSID="urn:lsid:plazi:treatment:03A787BAFFEAE51E531A87FB61C8F895:8ED16077FFEAE519530886C160C4FE55" box="[813,975,457,481]" country="Indonesia" municipality="Lowland forest" name=". Ent. Soc. Lond." pageId="43" pageNumber="160" stateProvince="SULAWESI UTARA">.Ent.Soc. Lond.</location>
, Project Wallace, B.M. 1985-10 / 29.21 / Holotypus ♂
<taxonomicName id="4C0E4D2FFFEAE519538286E2676BFDB5" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[935,1120,489,513]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="160" phylum="Arthropoda" rank="species" species="densissimus" status="sp. nov.">
<emphasis id="B97AEABEFFEAE519538286E2676BFDB5" box="[935,1120,489,513]" italics="true" pageId="43" pageNumber="160">Zyras densissimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957C5FFEAE519544186E26790FDB5" box="[1124,1179,490,513]" pageId="43" pageNumber="160" rank="species">sp. n.</taxonomicNameLabel>
, det.
<determinerName id="7E6DDA68FFEAE51954F486E2663FFDB5" box="[1233,1332,489,513]" pageId="43" pageNumber="160">V. Assing</determinerName>
2016” (
<collectionCode id="ED1FAE69FFEAE51953048501607CFD95" box="[801,887,521,545]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="43" pageNumber="160">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B663CF1FFEAE51953AE8502667DFD95" ID-GBIF-Occurrence="3455147309" box="[907,1398,521,546]" collectingDate="1985-05" collectionCode="BMNH" country="Indonesia" determinerName="V. Assing" elevation="200" location="Ent. Soc. Lond." municipality="Lowland forest" pageId="43" pageNumber="160" specimenCount="1" stateProvince="A" typeStatus="paratype">Paratypes: 1 ♂: same data as holotype (cAss)</materialsCitation>
<materialsCitation id="3B663CF1FFEAE519533F85216679FDF5" ID-GBIF-Occurrence="3455147318" box="[794,1394,553,577]" collectingDate="1985-05" collectionCode="BMNH" country="Indonesia" determinerName="V. Assing" elevation="200" location="Ent. Soc. Lond." municipality="Lowland forest" pageId="43" pageNumber="160" specimenCount="1" stateProvince="A" typeStatus="paratype">
1 ♂: same data as holotype, but “
<date id="FFB0106CFFEAE51954AF852167F3FDF5" box="[1162,1272,553,577]" pageId="43" pageNumber="160" value="1985-05">
<collectingDate id="EFF4E984FFEAE51954AF852167F3FDF5" box="[1162,1272,553,577]" pageId="43" pageNumber="160" value="1985-05">May 1985</collectingDate>
</date>
.” (
<collectionCode id="ED1FAE69FFEAE519553385216660FDF5" box="[1302,1387,553,577]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="43" pageNumber="160">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B663CF1FFEAE519533F85416074FD15" ID-GBIF-Occurrence="3450162316" collectingMethod="Malaise trap" collectionCode="BMNH" country="Indonesia" determinerName="V. Assing" elevation="664" location="Lowland forest" municipality="Edwards Camp'" pageId="43" pageNumber="160" specimenCount="1" stateProvince="A" typeStatus="paratype">
1 ♀: same data as holotype, but “
<date id="FFB0106CFFEAE5195448854167D6FDD5" box="[1133,1245,585,609]" pageId="43" pageNumber="160" value="1985-04">April 1985</date>
. /
<collectingMethod id="524F4EBBFFEAE51954D18541667DFDD5" box="[1268,1398,585,609]" pageId="43" pageNumber="160">Malaise trap</collectingMethod>
/ ‘
<collectingMunicipality id="6BD5ACD6FFEAE5195314856160D6FD35" box="[817,989,617,641]" pageId="43" pageNumber="160">Edwards Camp’</collectingMunicipality>
,
<location id="8ED16077FFEAE51953CE8561679BFD35" LSID="urn:lsid:plazi:treatment:03A787BAFFEAE51E531A87FB61C8F895:8ED16077FFEAE51953CE8561679BFD35" box="[1003,1168,617,641]" country="Indonesia" municipality="Edwards Camp'" name="Lowland forest" pageId="43" pageNumber="160" stateProvince="A">Lowland forest</location>
,
<quantity id="4CF69B49FFEAE51954BA856167E8FD35" box="[1183,1251,617,641]" metricMagnitude="2" metricUnit="m" metricValue="6.64" pageId="43" pageNumber="160" unit="m" value="664.0">
<elevation id="0023D19FFFEAE51954BA856167E8FD35" box="[1183,1251,617,641]" metricMagnitude="2" metricUnit="m" metricValue="6.64" pageId="43" pageNumber="160" unit="m" value="664.0">664 m</elevation>
</quantity>
,
<date id="FFB0106CFFEAE51954D785616666FD35" box="[1266,1389,617,641]" pageId="43" pageNumber="160">26.IV–28.V</date>
” (
<collectionCode id="ED1FAE69FFEAE51953048581607CFD15" box="[801,887,649,673]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="43" pageNumber="160">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEAE519533F85C16673FC95" pageId="43" pageNumber="160" type="etymology">
<paragraph id="8BB136ACFFEAE519533F85C16673FC95" blockId="43.[794,1400,713,801]" pageId="43" pageNumber="160">
<emphasis id="B97AEABEFFEAE519533F85C1609AFD55" bold="true" box="[794,913,713,737]" pageId="43" pageNumber="160">Etymology</emphasis>
: The specific epithet (adjective) is the superlative of the Latin adjective densus (dense). It alludes to the conspicuously dense and fine punctation of the forebody.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEAE51E533F84416004F9D5" lastPageId="44" lastPageNumber="161" pageId="43" pageNumber="160" type="description">
<paragraph id="8BB136ACFFEAE519533F84416733FB75" blockId="43.[794,1399,841,2081]" pageId="43" pageNumber="160">
<emphasis id="B97AEABEFFEAE519533F84416094FCD5" bold="true" box="[794,927,841,865]" pageId="43" pageNumber="160">Description</emphasis>
: Small species; body length 4.0–
<quantity id="4CF69B49FFEAE519553884416679FCD5" box="[1309,1394,841,865]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="43" pageNumber="160" unit="mm" value="4.7">4.7 mm</quantity>
; length of forebody
<quantity id="4CF69B49FFEAE51954258461678FFC35" box="[1024,1156,873,897]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="43" pageNumber="160" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.9–2.1 mm</quantity>
. Coloration (
<figureCitation id="13352A29FFEAE519550484616679FC35" box="[1313,1394,873,897]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="43" pageNumber="160">Figs 36</figureCitation>
,
<figureCitation id="13352A29FFEAE519533F8481603DFC15" box="[794,822,905,929]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="43" pageNumber="160">75</figureCitation>
,
<figureCitation id="13352A29FFEAE519536284816079FC15" box="[839,882,905,929]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="43" pageNumber="160">111</figureCitation>
): forebody blackish-brown with the anterior and humeral portions, sometimes also the suture, of the elytra more or less extensively reddish-yellow; abdomen dark-brown to blackish-brown with parts or all of tergites II–V and parts of the paratergites reddish to dark-reddish; antennae dark-yellowish with antennomeres IV–VIII more or less distinctly and more or less extensively brown to blackish-brown; maxillary palpi dark-yellowish to pale-brown with the terminal palpomere pale-yellowish.
</paragraph>
<paragraph id="8BB136ACFFEAE519533F83C167C5FA55" blockId="43.[794,1399,841,2081]" pageId="43" pageNumber="160">
Head (
<figureCitation id="13352A29FFEAE519534E83C160BEFB55" box="[875,949,1225,1249]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="43" pageNumber="160">Fig. 75</figureCitation>
) distinctly transverse; punctation very dense and rather fine; along middle without punctation. Eyes large and bulging, more than twice as long as postocular region in dorsal view. Antenna (
<figureCitation id="13352A29FFEAE51954CB82216632FAF5" box="[1262,1337,1321,1345]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="43" pageNumber="160">Fig. 36</figureCitation>
) very slender, approximately 2.0 mm long; antennomeres IV–V approximately twice as long as broad, V–X of gradually decreasing length and decreasingly oblong, IX weakly oblong, X weakly oblong or as long as broad, and XI as long as the combined length of IX and X.
</paragraph>
<paragraph id="8BB136ACFFEAE519533F82E2605EF975" blockId="43.[794,1399,841,2081]" pageId="43" pageNumber="160">
Pronotum (
<figureCitation id="13352A29FFEAE51953BE82E160EDF9B5" box="[923,998,1513,1537]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="43" pageNumber="160">Fig. 75</figureCitation>
) weakly transverse, 1.05–1.08 times as broad as long and approximately 1.1 times as broad as head, broadest near anterior angles, moderately tapering posteriad; lateral margins straight in posterior two-thirds (dorsal view); punctation fine, regular, and conspicuously dense, also along midline; lateral margins without long setae.
</paragraph>
<paragraph id="8BB136ACFFEAE519533F81C167ADF8D5" blockId="43.[794,1399,841,2081]" pageId="43" pageNumber="160">
Elytra (
<figureCitation id="13352A29FFEAE519535781C160B6F955" box="[882,957,1737,1761]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="43" pageNumber="160">Fig. 75</figureCitation>
) approximately 0.85 times as long as pronotum; punctation similar to that of pronotum; scutellum with very coarse and defined punctation. Hind wings fully developed. Metatarsomere I approximately as long as the combined length of II–IV.
</paragraph>
<paragraph id="8BB136ACFFEAE51E533F8061618AFD95" blockId="43.[794,1399,841,2081]" lastBlockId="44.[189,794,233,865]" lastPageId="44" lastPageNumber="161" pageId="43" pageNumber="160">
Abdomen (
<figureCitation id="13352A29FFEAE51953BF806160FAF835" box="[922,1009,1897,1921]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="43" pageNumber="160">Fig. 111</figureCitation>
) narrower than elytra, with deep anterior impressions on tergites III–V; tergites III–V each with a transverse row of 6–8 defined, large and deep non-setiferous punctures in anterior impressions, with a lateral setiferous puncture on either side, but without setiferous punctures at posterior margin; tergite VI with few fine non-setiferous punctures anteriorly, with a lateral setiferous puncture on either side, without setiferous punctures at posterior margin; tergite VII with scattered fine non-setiferous punctures anteriorly and with two transverse rows of four setiferous punctures posteriorly, each composed of only four punctures, posterior margin with palisade fringe; tergite VIII (
<figureCitation id="13352A29FFEDE51E521B86A1619DFE75" box="[574,662,425,449]" captionStart="Figs 227–244" captionStartId="61.[151,191,1983,2005]" captionTargetBox="[163,1397,240,1949]" captionTargetId="figure-119@61.[151,1398,231,1970]" captionText="Figs 227–244: Zyras parahirtus (227–228, 231–232), Z. matangensis (229–230, 233), Z. gratellus (Sulawesi) (234–238), and Z. densissimus (239–244): median lobe of aedeagus in lateral and in ventral view (227–230, 234–235, 239–241); paramere (231, 236, 242); male tergite VIII (232, 237, 243); male sternite VIII (233, 238, 244). Scale bars: 227–233, 237–238, 243–244: 0.5 mm; 234–236, 239–242: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888378" httpUri="https://zenodo.org/record/5888378/files/figure.png" pageId="44" pageNumber="161">Fig. 243</figureCitation>
) with eight long setae at posterior margin, a few setae postero-laterally, and with fine pubescence in posterior half, posterior margin convex, without median concavity.
</paragraph>
<paragraph id="8BB136ACFFEDE51E5098852262E7FC95" blockId="44.[189,794,233,865]" pageId="44" pageNumber="161">
♂: sternite VIII (
<figureCitation id="13352A29FFEDE51E51A4852162D3FDF5" box="[385,472,553,577]" captionStart="Figs 227–244" captionStartId="61.[151,191,1983,2005]" captionTargetBox="[163,1397,240,1949]" captionTargetId="figure-119@61.[151,1398,231,1970]" captionText="Figs 227–244: Zyras parahirtus (227–228, 231–232), Z. matangensis (229–230, 233), Z. gratellus (Sulawesi) (234–238), and Z. densissimus (239–244): median lobe of aedeagus in lateral and in ventral view (227–230, 234–235, 239–241); paramere (231, 236, 242); male tergite VIII (232, 237, 243); male sternite VIII (233, 238, 244). Scale bars: 227–233, 237–238, 243–244: 0.5 mm; 234–236, 239–242: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888378" httpUri="https://zenodo.org/record/5888378/files/figure.png" pageId="44" pageNumber="161">Fig. 244</figureCitation>
) with fine and rather dense pubescence in posterior half, posterior margin weakly to distinctly convex; median lobe of aedeagus (
<figureCitation id="13352A29FFEDE51E50E18581625FFD15" box="[196,340,649,673]" captionStart="Figs 227–244" captionStartId="61.[151,191,1983,2005]" captionTargetBox="[163,1397,240,1949]" captionTargetId="figure-119@61.[151,1398,231,1970]" captionText="Figs 227–244: Zyras parahirtus (227–228, 231–232), Z. matangensis (229–230, 233), Z. gratellus (Sulawesi) (234–238), and Z. densissimus (239–244): median lobe of aedeagus in lateral and in ventral view (227–230, 234–235, 239–241); paramere (231, 236, 242); male tergite VIII (232, 237, 243); male sternite VIII (233, 238, 244). Scale bars: 227–233, 237–238, 243–244: 0.5 mm; 234–236, 239–242: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888378" httpUri="https://zenodo.org/record/5888378/files/figure.png" pageId="44" pageNumber="161">Figs 239–241</figureCitation>
)
<quantity id="4CF69B49FFEDE51E5141858162F6FD15" box="[356,509,649,673]" metricMagnitude="-4" metricUnit="m" metricValue="5.75" metricValueMax="5.8" metricValueMin="5.699999999999999" pageId="44" pageNumber="161" unit="mm" value="0.575" valueMax="0.58" valueMin="0.57">0.57–0.58 mm</quantity>
long, slender, and of very distinctive morphology; ventral process long and slender, with sinuate lateral margins in ventral view; paramere (
<figureCitation id="13352A29FFEDE51E50E085E16217FCB5" box="[197,284,745,769]" captionStart="Figs 227–244" captionStartId="61.[151,191,1983,2005]" captionTargetBox="[163,1397,240,1949]" captionTargetId="figure-119@61.[151,1398,231,1970]" captionText="Figs 227–244: Zyras parahirtus (227–228, 231–232), Z. matangensis (229–230, 233), Z. gratellus (Sulawesi) (234–238), and Z. densissimus (239–244): median lobe of aedeagus in lateral and in ventral view (227–230, 234–235, 239–241); paramere (231, 236, 242); male tergite VIII (232, 237, 243); male sternite VIII (233, 238, 244). Scale bars: 227–233, 237–238, 243–244: 0.5 mm; 234–236, 239–242: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888378" httpUri="https://zenodo.org/record/5888378/files/figure.png" pageId="44" pageNumber="161">Fig. 242</figureCitation>
) approximately as long as median lobe and with long and slender apical lobe.
</paragraph>
<paragraph id="8BB136ACFFEDE51E50988422623CFCD5" blockId="44.[189,794,233,865]" pageId="44" pageNumber="161">♀: posterior margin of sternite VIII weakly concave in the middle.</paragraph>
<paragraph id="8BB136ACFFEDE51E509884816210FA95" blockId="44.[189,794,905,1633]" pageId="44" pageNumber="161">
<emphasis id="B97AEABEFFEDE51E50988481629FFC15" bold="true" box="[189,404,905,929]" pageId="44" pageNumber="161">Comparative notes</emphasis>
: This highly distinctive species is readily identified and distinguished from its consubgeners by the characteristic punctation of the forebody, the punctation pattern of the abdomen (particularly the absence of setiferous punctures at the posterior margins of tergites III–VI, the large and deep punctures in the anterior impressions of tergites III–V, and the scattered fine non-setiferous punctures in the anterior portions of tergites VI and VII), and by the conspicuous shape of the aedeagus. In addition, it is characterized by small body size, long and slender antennae, the coloration of the antennae, and by the absence of long setae at the pronotal margins.
</paragraph>
<paragraph id="8BB136ACFFEDE51E509882216004F9D5" blockId="44.[189,794,905,1633]" pageId="44" pageNumber="161">
As can be inferred from the similarly derived punctation of the forebody and the abdomen, as well as from the similarly slender antennae and habitus,
<taxonomicName id="4C0E4D2FFFEDE51E5283826263E2FA15" authorityName="Assing" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="161" phylum="Arthropoda" rank="species" species="densissimus">
<emphasis id="B97AEABEFFEDE51E5283826263E2FA15" italics="true" pageId="44" pageNumber="161">Z. densissimus</emphasis>
</taxonomicName>
is closely allied to
<taxonomicName id="4C0E4D2FFFEDE51E518E828261CFFA15" authority="CAMERON, 1943" authorityName="CAMERON" authorityYear="1943" box="[427,708,1417,1441]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="161" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="B97AEABEFFEDE51E518E82826118FA15" box="[427,531,1417,1441]" italics="true" pageId="44" pageNumber="161">Z. bryanti</emphasis>
<bibRefCitation id="EF9F4B5DFFEDE51E523C828161CFFA15" author="CAMERON, M." box="[537,708,1417,1441]" pageId="44" pageNumber="161" pagination="139 - 143" refId="ref57563" refString="CAMERON, M. 1943: New species of Staphylinidae (Col.) from Borneo. - Entomologist's Monthly Magazine 79: 39 - 42, 139 - 143, 183." type="journal article" year="1943">CAMERON, 1943</bibRefCitation>
</taxonomicName>
(female
<typeStatus id="54B5880EFFEDE51E509882A16210FA75" box="[189,283,1449,1473]" pageId="44" pageNumber="161" type="holotype">holotype</typeStatus>
examined; male unknown) from Borneo, from which it differs by different coloration (
<taxonomicName id="4C0E4D2FFFEDE51E527982C261CFFA55" authorityName="CAMERON" authorityYear="1943" box="[604,708,1481,1505]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="161" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="B97AEABEFFEDE51E527982C261CFFA55" box="[604,708,1481,1505]" italics="true" pageId="44" pageNumber="161">Z. bryanti</emphasis>
</taxonomicName>
: pronotum pale-reddish; abdominal segments III-V and VIII reddish-yellow), even denser and finer punctation of the forebody, and a more convex (cross-section) and posteriorly more strongly tapering pronotum (cross-section).
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEDE51E5098818161C8F895" pageId="44" pageNumber="161" type="distribution">
<paragraph id="8BB136ACFFEDE51E5098818161C8F895" blockId="44.[189,794,1673,1825]" pageId="44" pageNumber="161">
<emphasis id="B97AEABEFFEDE51E509881816123F915" bold="true" box="[189,552,1673,1697]" pageId="44" pageNumber="161">Distribution and natural history</emphasis>
: The known distribution is confined to Nani Wartabone National Park in
<collectingRegion id="49CAF84EFFEDE51E509881C16257F955" box="[189,348,1737,1761]" country="Indonesia" name="Sulawesi Utara" pageId="44" pageNumber="161">Sulawesi Utara</collectingRegion>
,
<collectingRegion id="49CAF84EFFEDE51E514381C16103F955" box="[358,520,1737,1761]" country="Indonesia" name="Sulawesi Utara" pageId="44" pageNumber="161">North Sulawesi</collectingRegion>
(
<figureCitation id="13352A29FFEDE51E523381C16160F955" box="[534,619,1737,1761]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="44" pageNumber="161">Map 10</figureCitation>
). The
<typeStatus id="54B5880EFFEDE51E528881C261D0F956" box="[685,731,1738,1762]" pageId="44" pageNumber="161">type</typeStatus>
specimens were collected with flight interception and Malaise traps in lowland forest at altitudes of
<quantity id="4CF69B49FFEDE51E5263800161B4F895" box="[582,703,1801,1825]" metricMagnitude="2" metricUnit="m" metricValue="4.3" metricValueMax="6.6" metricValueMin="2.0" pageId="44" pageNumber="161" unit="m" value="430.0" valueMax="660.0" valueMin="200.0">
200–
<specimenCount id="9D08FD25FFEDE51E525F800161B4F895" box="[634,703,1801,1825]" pageId="44" pageNumber="161" type="generic">660 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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